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Family HESPERIIDAE Latreille, 1809 – SKIPPERS

  Subfamily Eudaminae

Phocides  Hübner, [1819]

Phocides polybius (Fabricius, 1793)  Guava Skipper  TL: Indiis

Phocides polybius lilea (Reakirt, [1867])  S Texas to Costa Rica  M T S1 S2 L1 L2 I1 I2 G

Phocides polybius polybius (Fabricius, 1793)  Panama, Colombia, Venezuela, Guianas, Trinidad  M T S L G

Phocides polybius phanias (Burmeister, 1880)  TL: Argentina  M S L

Phocides charon (C. Felder & R. Felder, 1859)  TL: Brazil  M T S IL

Phocides distans (Herrich-Schäffer, 1869)  TL: ?

Phocides distans licinus (Möschler, 1879)  Honduras, Costa Rica, Panama  M T S L G

Phocides distans distans (Herrich-Schäffer, 1869)  TL: ?  M T S IL

Phocides distans silva Evans, 1952  TL: Peru  M T S

Phocides perillus (Mabille, 1888)  TL: Colombia  M T OI S

Phocides metrodorus E. Bell, 1932  Metrodorus Skipper  TL: Colombia

Phocides metrodorus nigrescens E. Bell, 1938  Costa Rica, Colombia, Venezuela, Ecuador, Peru; species?  M T S1 S2 I1 I2

Phocides metrodorus metrodorus E. Bell, 1932  Panama to Bolivia and Brazil  M T S G

Phocides metrodorus metron Evans, 1952  TL: Paraguay  M T OD G

Phocides vulcanides Röber, 1925  TL: Colombia  M T S

Phocides novalis Evans, 1952  TL: Peru  M T S G

Phocides thermus (Mabille, 1883)  Thermus Skipper  TL: Colombia

Phocides thermus thermus (Mabille, 1883)  SE Mexico to Venezuela and Ecuador  M T S G

Phocides thermus bellina Evans, 1952  TL: Ecuador  M T S

Phocides thermus valgus (Mabille, 1883)  TL: French Guiana  M T S

Phocides partia Evans, 1952  TL: Peru  M T S G

Phocides padrona Evans, 1952  TL: Peru  M T S G

Phocides belus Godman & Salvin, 1893  Belus Skipper  S Texas (one-time stray) to Costa Rica  M T S1 S2 L I

Phocides pigmalion (Cramer, 1779)  Pigmalion Skipper  TL: Surinam

Phocides pigmalion pigmalion (Cramer, 1779)  TL: Surinam  M T OI G

Phocides pigmalion hewitsonius (Mabille, 1883)  TL: Brazil (Am)  M T TI IL

Phocides pigmalion okeechobee (Worthington, 1881)  S Florida; mangroves  M T S L1 L2 I G

Phocides pigmalion batabanoides (W. Holland, 1902)  most Bahamian Islands  M T S

Phocides pigmalion batabano (Lucas, 1857)  Cuba, I. de Juventud, Andros, Little Cayman  M T S G

Phocides pigmalion bicolora (Boddaert, 1783)  Hispaniola  M T S

Phocides johnsoni E. Bell, 1947  TL: Colombia  M T S

Phocides urania (Westwood, 1852)  Urania Skipper  TL: Mexico ([Oax])

Phocides urania urania (Westwood, 1852)  Mexico, Guatemala; usually in montane forest  M T S L G

Phocides urania vida (A. Butler, 1872)  Costa Rica  M T S

Phocides pialia (Hewitson, 1857)  TL: Brazil

Phocides pialia maximus (Mabille, 1888)  TL: Brazil  M T OI S

Phocides pialia intermedia O. Mielke, 1992  TL: Brazil (MG)  M OI S

Phocides pialia pialia (Hewitson, 1857)  TL: Brazil  M T OI S G

Phocides oreides (Hewitson, [1875])  TL: Peru

Phocides oreides colombiana E. Bell, 1938  TL: Colombia  M T

Phocides oreides oreides (Hewitson, [1875])  TL: Peru  M T OI S IL

Phocides yokhara (A. Butler, 1870)  TL: Peru

Phocides yokhara yokhara (A. Butler, 1870)  TL: Peru  M T S L

Phocides yokhara inca Le Cerf, 1922  TL: Peru  M T

Phocides yokhara dryas Le Cerf, 1922  TL: Peru  M T

Phocides yokhara charonotis (Hewitson, 1874)  TL: Bolivia  M T TI

Phocides lincea (Herrich-Schäffer, 1869)  TL: ?

Phocides lincea lincea (Herrich-Schäffer, 1869)  TL: ?  M S

Phocides lincea perkinsi (Kaye, 1931)  Jamaica  M T S

Hypocryptothrix  E. Watson, 1893

Hypocryptothrix teutas (Hewitson, 1876)  TL: Brazil (Am)  M T S

Tarsoctenus  E. Watson, 1893

Tarsoctenus papias (Hewitson, 1857)  TL: Brazil (Am)  M T OI S L

Tarsoctenus corytus (Cramer, 1777)  TL: Surinam

Tarsoctenus corytus gaudialis (Hewitson, 1876)  Nicaragua to Colombia  M T OI S L G

Tarsoctenus corytus corytus (Cramer, 1777)  TL: Surinam  M T OI OD S

Tarsoctenus corytus corba Evans, 1952  TL: Peru  M T S

Tarsoctenus praecia (Hewitson, 1857)  TL: Brazil (Pa)

Tarsoctenus praecia praecia (Hewitson, 1857)  TL: Brazil (Pa)  M T OI S

Tarsoctenus praecia rufibasis Boullet, 1910  TL: French Guiana  M T IL

Tarsoctenus praecia plutia (Hewitson, 1857)  TL: Brazil (Am)  M T OI S L

Tarsoctenus praecia luna Evans, 1952  TL: Bolivia  M T

Phanus  Hübner, [1819]

Phanus australis L. Miller, 1965  TL: Brazil (SC)  M T S

Phanus vitreus (Stoll, 1781)  Widespread Phanus  Oaxaca and Chiapas to southern Brazil  M T S1 S2 L I G

Phanus ecitonorum Austin, 1993  Army-ant Phanus  Costa Rica to southern Brazil  M T S L

Phanus confusis Austin, 1993  Confusing Phanus  eastern Mexico to El Salvador  M T S

Phanus albiapicalis Austin, 1993  White-tipped Phanus  E & W Mexico to Panama, W Ecuador and Peru  M T S L

Phanus rilma Evans, 1952  West-Mexican Phanus  Pacific coast, Nayarit to Guatemala  M T S G

Phanus grandis Austin, 1993  Grand Phanus  Nicaragua, Costa Rica, Panama, Colombia, Venezuela  M T S L

Phanus obscurior Kaye, 1925  Dark Phanus  TL: Trinidad

Phanus obscurior obscurior Kaye, 1925  SE Mexico to Colombia and Trinidad  M T S1 S2 I G

Phanus obscurior prestoni L. Miller, 1965  TL: Brazil (Am)  M T S

Phanus marshalli (W. F. Kirby, 1880)  Common Phanus  E & W Mexico to Bolivia  M S1 S2 L I G

Udranomia  A. Butler, 1870

Udranomia orcinus (C. Felder & R. Felder, 1867)  Orcinus Skipper  SE Mexico to Bolivia  M T S1 S2 IL I G

Udranomia kikkawai (Weeks, 1906)  Nervous Skipper  E & W Mexico to Brazil  M T TI S1 S2 S3 L I1 I2

Udranomia eurus (Mabille & Boullet, 1919)  Eurus Skipper  Costa Rica to Venezuela & Amazon basin  M S I

Udranomia spitzi (Hayward, 1942)  TL: Brazil (MT)  M S

Drephalys  E. Watson, 1893

Drephalys alcmon (Cramer, 1780)  Alcmon Skipper  Guatemala to Guianas and Brazil  M T S1 S2 S3 IL L I G

Drephalys mourei O. Mielke, 1968  TL: Brazil (SC)  M T IL

Drephalys helixus (Hewitson, 1877)  Helixus Skipper  Panama  M T OD S IL G

Drephalys kidonoi Burns, 2000  Kidono's Skipper  W Costa Rica  M T S1 S2 I G

Drephalys phoenicoides (Mabille & Boullet, 1919)  Curious Skipper  Costa Rica to Guianas and NE Brazil  M S

Drephalys phoenice (Hewitson, 1867)  TL: Brazil (Am)  M T S IL

Drephalys heraclides E. Bell, 1942  TL: Peru  M T S

Drephalys miersi O. Mielke, 1968  TL: Brazil (SC)  M T S IL

Drephalys opifex Evans, 1952  TL: Surinam  M OD

Drephalys olvina Evans, 1952  TL: Guyana  M S

Drephalys olva Evans, 1952  TL: Brazil (Pa)  M T OD

Drephalys eous (Hewitson, 1867)  TL: Brazil (Pa)  M T S

Drephalys croceus Austin, 1995  TL: Brazil (Rd)  M T OI G

Drephalys dumeril (Latreille, [1824])  Dumeril Skipper  Mexico to Ecuador and Brazil  M T OI OD S1 S2 IL G

Drephalys oria Evans, 1952  Oria Skipper  Mexico to Honduras  M T S G

Drephalys oriander (Hewitson, 1867)  Oriander Skipper  Mexico, Costa Rica to Guianas and N Brazil  M T S1 S2 IL G

Drephalys talboti (Le Cerf, 1922)  TL: French Guiana  M T S G

Drephalys tortus Austin, 1995  TL: Brazil (Rd)  M T OI G

Augiades  Hübner, [1819]

Augiades crinisus (Cramer, 1780)  Crinisus Skipper  Costa Rica to Bolivia and Brazil  M T S L

Augiades epimethea (Plötz, 1883)  TL: Brazil

Augiades epimethea bicolor (Mabille & Boullet, 1919)  Costa Rica to Bolivia, Guianas and N Brazil  M T S

Augiades epimethea epimethea (Plötz, 1883)  TL: Brazil  M T

Hyalothyrus  Mabille, 1878

Hyalothyrus infernalis (Möschler, 1877)  TL: French Guiana; Surinam

Hyalothyrus infernalis infernalis (Möschler, 1877)  TL: French Guiana; Surinam  M T S

Hyalothyrus infernalis infa Evans, 1952  TL: Colombia  M T S

Hyalothyrus leucomelas (Geyer, 1832)  Leucomelas Skipper  Panama to Brazil and Peru  M S

Hyalothyrus nitocris (Stoll, 1782)  TL: Surinam  M S

Hyalothyrus mimicus Mabille & Boullet, 1919  TL: Peru  M S

Hyalothyrus neleus (Linnaeus, 1758)  TL: Indiis

Hyalothyrus neleus pemphigargyra (Mabille, 1888)  SE Mexico to Colombia and Venezuela  M T S L I G

Hyalothyrus neleus neleus (Linnaeus, 1758)  TL: Indiis  M T S L

Hyalothyrus neleus neda Evans, 1952  TL: Bolivia  M T S

Phareas  Westwood, 1852

Phareas burnsi Grishin, 2013  Burns' Skipper  Costa Rica, Panama, ?Colombia  M T1 T2 S I

Phareas coeleste Westwood, 1852  Magnificent Skipper  Colombia to Bolivia and S Brazil  M T OI OD S1 S2 S3 IL L G

Entheus  Hübner, [1819]

Entheus eumelus (Cramer, 1777)  TL: Surinam  M T OI OD S IL G

Entheus ninyas H. Druce, 1912  TL: Bolivia  M T OI OD S

Entheus eunyas Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M T OI S G

Entheus lemna (A. Butler, 1870)  TL: ?  M T OD TI S IL Txt

Entheus gentius (Cramer, 1777)  TL: Surinam  M T OI OD TI S1 S2 S3 HS IL G Txt

Entheus bombus Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M T OI S G

Entheus aureolus Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M T OI S1 S2 S3 S4 S5 S6 IL G

Entheus huertasae Grishin, 2013  Blanca's Entheus  Panama & Colombia  M T IL G

Entheus telemus Mabille, 1898  TL: Brazil  M OD S IL G

Entheus latebrosus Austin, 1997  TL: Ecuador  M T OI S

Entheus priassus (Linnaeus, 1758)  TL: Indiis

Entheus priassus priassus (Linnaeus, 1758)  TL: Indiis  M OI OD TI S HS IL G Txt

Entheus priassus pralina Evans, 1952  TL: Brazil (ES)  M T IL G

Entheus aureanota Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M T OI S G

Entheus curvus Austin, 1997  TL: Peru  M T OI S G

Entheus matho Godman & Salvin, 1879  Giant Entheus  TL: Guatemala; Nicaragua

Entheus matho matho Godman & Salvin, 1879  SE Mexico to Costa Rica  M T OD S IL L G Txt

Entheus matho latifascius M. Hering, 1925  TL: Colombia  M OI OD S

Entheus matho aequatorius Mabille & Boullet, 1919  Costa Rica to Ecuador  M OD S IL G

Entheus matho dius Mabille, 1898  Panama Guianas, Peru, and Bolivia  M T OD TI S IL G

Entheus crux Steinhauser, 1989  Mexican Entheus  SE Mexico to Nicaragua  M T OD S G

Entheus warreni Grishin, 2012  Warren's Entheus  Ecuador  M T S G

Cabirus  Hübner, [1819]

Cabirus procas (Cramer, 1777)  TL: Surinam

Cabirus procas procas (Cramer, 1777)  TL: Surinam  M OI OD S IL

Cabirus procas junta Evans, 1952  TL: Peru  M T S1 S2 IL

Cabirus procas purda Evans, 1952  TL: Peru  M T S L

Proteides  Hübner, [1819]

Proteides mercurius (Fabricius, 1787)  Mercurial Skipper  TL: French Guiana

Proteides mercurius mercurius (Fabricius, 1787)  continental, S Texas to Argentina, SE Arizona as rare stray  M OI S1 S2 L I1 I2 G

Proteides mercurius sanantonio (Lucas, 1857)  Cuba, [one-time stray to Florida]  M S L

Proteides mercurius sanchesi E. Bell & W. Comstock, 1948  Hispaniola  M T S

Proteides mercurius jamaicensis Skinner, 1920  Jamaica  M T S

Proteides mercurius pedro (Dewitz, 1877)  Puerto Rico  M S

Proteides mercurius vincenti E. Bell & W. Comstock, 1948  St. Vincent  M T S

Proteides mercurius angasi Godman & Salvin, 1884  Dominica, St. Lucia, Tobago  M T S

Proteides mercurius grenadensis Enrico & Pinchon, 1969  Grenada  M S

Proteides maysi (Lucas, 1857)  Mays' Skipper  Cuba and Isla de Juventud  M T S

Epargyreus  Hübner, [1819]

Epargyreus zestos (Geyer, 1832)  Zestos Skipper  TL: "Surinam"

Epargyreus zestos zestos (Geyer, 1832)  Florida, Bahamas, Puerto Rico, Virgin Islands to St. Vincent  M T S L G

Epargyreus zestos inaguarum Clench & Bjorndal, 1980  Great Inagua, Crooked I., Turks and Caicos Islands  M T S

Epargyreus clarus (Cramer, 1775)  Silver-spotted Skipper  TL: [USA (Va)]

Epargyreus clarus californicus MacNeill, 1975  BC to Baja California Norte; Sierra Nevada-Cascades, westward  M S L I H

Epargyreus clarus Rocky Mountain segregate ,  NE OR, E WA, ID, MT, WY, C & N UT, CO, N NM, Black Hills  M S

Epargyreus clarus clarus (Cramer, 1775)  E USA and Canada to far NE Mexico (hypothetical)  M S HS L1 L2 L3 L4 I H G

Epargyreus clarus huachuca Dixon, 1955  S AZ, S NM, far NW Mexico  M T S L I H

Epargyreus clarus profugus Austin, 1998  S NV, N AZ, possibly to central AZ-NM  M T L

Epargyreus antaeus (Hewitson, 1867)  Jamaican Silverdrop  Jamaica  M T S L G

Epargyreus spanna Evans, 1952  Hispaniolan Silverdrop  Hispaniola  M T S G

Epargyreus barisses (Hewitson, 1874)  TL: Bolivia

Epargyreus barisses barisses (Hewitson, 1874)  TL: Bolivia  M T S G

Epargyreus barisses argentina Mabille, 1903  TL: ?  M T S G

Epargyreus tmolis (Burmeister, 1875)  TL: Argentina  M S G

Epargyreus socus (Hübner, [1825])  TL: ?

Epargyreus socus orizaba Scudder, 1872  E Mexico to Guatemala  M T S L G

Epargyreus socus chota Evans, 1952  TL: Trinidad  M T

Epargyreus socus dicta Evans, 1952  TL: Bolivia  M T

Epargyreus socus sinus Evans, 1952  TL: Brazil (Pa)  M T S

Epargyreus socus socus (Hübner, [1825])  TL: ?  M G

Epargyreus socus pseudexadeus Westwood, 1852  TL: Brazil ([MG])  M T S

Epargyreus windi H. Freeman, 1969  Wind’s Silverdrop  W Mexico  M T S L G

Epargyreus exadeus (Cramer, 1779)  TL: Surinam

Epargyreus exadeus cruza Evans, 1952  (S TX? [CA?]as stray), E & W Mexico to Panama  M T IL L G

Epargyreus exadeus exadeus (Cramer, 1779)  TL: Surinam  M S G

Epargyreus nutra Evans, 1952  TL: Colombia  M T G

Epargyreus aspina Evans, 1952  Spineless Silverdrop  E & W Mexico to Venezuela and Ecuador  M T L G

Epargyreus spina Evans, 1952  Spined Silverdrop  TL: Colombia

Epargyreus spina spina Evans, 1952  E & W Mexico to Brazil and Bolivia  M T S G

Epargyreus spina verruga Evans, 1952  TL: Peru  M T

Epargyreus spinta Evans, 1952  TL: Colombia  M T S

Epargyreus spinosa Evans, 1952  Suffused Silverdrop  W Mexico to Panama  M T S G

Epargyreus clavicornis (Herrich-Schäffer, 1869)  TL: ?

Epargyreus clavicornis gaumeri Godman & Salvin, 1893  SE Mexico (?) to Honduras  M T OI G

Epargyreus clavicornis tenda Evans, 1955  SE Mexico to Venezuela and Ecuador  M T S L G

Epargyreus clavicornis clavicornis (Herrich-Schäffer, 1869)  TL: ?  M T IL G

Epargyreus brodkorbi H. Freeman, 1969  Barely-spotted Silverdrop  Chiapas  M T S L G

Epargyreus deleoni H. Freeman, 1977  Long-spotted Silverdrop  Yucatan Peninsula (CAMP, QROO, YUC Mexico) and Belize  M T S G

Epargyreus enispe (Hewitson, 1867)  TL: S. Amer.

Epargyreus enispe enispe (Hewitson, 1867)  TL: S. Amer.  M T S G

Epargyreus enispe elta Evans, 1952  TL: Bolivia  M T

Polygonus  Hübner, [1825]

Polygonus leo (Gmelin, [1790])  Hammock Skipper  TL: America

Polygonus leo arizonensis (Skinner, 1911)  SW USA (mostly as strays) & S TX to Central America  M T S L I1 I2 G

Polygonus leo pallida Röber, 1925  Central America to Argentina  M T S L

Polygonus leo histrio Röber, 1925  Florida and Cuba, some Bahamian islands  M S L I

Polygonus leo leo (Gmelin, [1790])  [= lividus, = ishmael], Hispaniola, Puerto Rico, SE to St. Lucia  M T S IL L G

Polygonus leo hagar Evans, 1952  Jamaica  M T S

Polygonus savigny (Latreille, [1824])  Manuel’s Skipper  TL: Antilles

Polygonus savigny savigny (Latreille, [1824])  [= manueli], continental, (S TX as stray), N Mexico to Argentina  M T S1 S2 HS L I G

Polygonus savigny punctus E. Bell & W. Comstock, 1948  Lesser Antilles; St. Martin S to Grenadines  M T S

Chioides  Lindsey, 1921

Chioides albofasciatus (Hewitson, 1867)  White-striped Longtail  S TX, S NM, SE AZ, to Costa Rica; Baja California Sur  M T S L1 L2 L3 L4 I H G

Chioides catillus (Cramer, 1779)  TL: Surinam

Chioides catillus albius Evans, 1952  Costa Rica and Panama  M T S1 S2 L I G

Chioides catillus catillus (Cramer, 1779)  TL: Surinam  M S1 S2 L

Chioides catillus jethira (A. Butler, 1870)  TL: Peru  M T S G

Chioides churchi E. Bell & W. Comstock, 1948  Jamaican Longtail  Jamaica  M T S

Chioides cinereus (Mabille & Vuillot, 1891)  TL: Brazil  M T S IL

Chioides concinnus (Mabille, 1877)  Peru  M T OI S G

Chioides vintra Evans, 1952  St. Vincent Longtail  St. Vincent, Grenadines, Grenada  M T S

Chioides iverna Evans, 1952  TL: Peru  M T S

Chioides zilpa (A. Butler, 1872)  Zilpa Longtail  S AZ-NM (stray) & S TX, S to Venezuela and Ecuador  M T OD S1 S2 L I1 I2 G

Chioides ixion (Plötz, 1880)  Hispaniolan Longtail  Hispaniola  M T S

Chioides marmorosa (Herrich-Schäffer, 1865)  Cuban Longtail  Cuba  M T S

Aguna  R. Williams, 1927

Aguna asander (Hewitson, 1867)  Gold-spotted Aguna  TL: Brazil (Am)

Aguna asander asander (Hewitson, 1867)  SE AZ & S TX (as stray) to Argentina  M T S L I G

Aguna asander haitensis (Mabille & Boullet, 1912)  Cuba and Hispaniola  M T S

Aguna asander jasper Evans, 1952  Jamaica  M T S L

Aguna megaeles (Mabille, 1888)  TL: [Brazil (SC)]

Aguna megaeles malia Evans, 1952  TL: Venezuela  M T G

Aguna megaeles megaeles (Mabille, 1888)  TL: [Brazil (SC)]  M T OI S IL

Aguna albistria (Plötz, 1880)  TL: Brazil ([Pa])

Aguna albistria albistria (Plötz, 1880)  TL: Brazil ([Pa])  M T TI S L

Aguna albistria leucogramma (Mabille, 1888)  SE Mexico to Colombia and Venezuela (Surinam?)  M T OI S

Aguna metophis (Latreille, [1824])  Long-tailed Aguna  S Texas to S Brazil  M T S L I

Aguna camagura (R. Williams, 1926)  TL: Brazil (MT)  M T S

Aguna claxon Evans, 1952  Emerald Aguna  Cuba, S Texas to Colombia and Surinam  M T S L I G

Aguna clina Evans, 1952  TL: Colombia  M T S

Aguna latifascia Austin & O. Mielke, 1998  TL: Ecuador  M T S

Aguna coelus (Stoll, 1781)  Stoll's Aguna  Costa Rica and Panama to Peru and central Brazil  M T S

Aguna cirrus Evans, 1952  TL: Brazil (SP)  M T S G

Aguna coeloides Austin & O. Mielke, 1998  Austin's Aguna  SE Mexico to southern Brazil  M T S1 S2 HS I1 I2

Aguna nicolayi Austin & O. Mielke, 1998  TL: Brazil (MT)  M T S

Aguna venezuelae O. Mielke, 1971  TL: Venezuela  M T S

Aguna latimacula Austin & O. Mielke, 1998  TL: Brazil (MG)  M T

Aguna aurunce (Hewitson, 1867)  TL: Brazil (Am)

Aguna aurunce aurunce (Hewitson, 1867)  TL: Brazil (Am)  M T S

Aguna aurunce hypozonius (Plötz, 1880)  SE Mexico to Venezuela  M S L I

Aguna glaphyrus (Mabille, 1888)  TL: Brazil (SC)  M T S

Aguna penicillata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna spicata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna longicauda Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T OI

Aguna ganna (Möschler, 1879)  Ganna Aguna  Costa Rica to Colombia and Venezuela  M T S IL

Aguna panama Austin & O. Mielke, 1998  Panamanian Aguna  Honduras to Venezuela  M T1 T2 S I

Aguna spatulata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna similis Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna mesodentata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna squamalba Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna parva Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Typhedanus  A. Butler, 1870

Typhedanus crameri McHenry, 1960  TL: not specified  M T S L

Typhedanus undulatus (Hewitson, 1867)  Mottled Longtail  S Texas (as stray) to Argentina  M T S1 S2 HS L I G

Typhedanus galbula (Plötz, 1880)  TL: Brazil ([Pa])  M T S

Typhedanus optica Evans, 1952  TL: Guyana

Typhedanus optica optica Evans, 1952  TL: Guyana  M T S

Typhedanus optica goya Evans, 1952  TL: Brazil (Go)  M T S

Typhedanus stylites (Herrich-Schäffer, 1869)  TL: [Brazil]  M T S

Typhedanus aziris (Hewitson, 1867)  TL: Brazil (RJ)  M T S

Typhedanus umber (Herrich-Schäffer, 1869)  TL: ?  M T S

Typhedanus ampyx (Godman & Salvin, 1893)  Gold-tufted Skipper  W & NE Mexico to Costa Rica  M T S1 S2 L I G

Typhedanus salas H. Freeman, 1977  Salas Skipper  S Mexico & Belize  M T L

Typhedanus cursinoi O. Mielke, 1979  TL: Brazil (Ba)  M T OI G

Typhedanus eliasi O. Mielke, 1979  TL: Brazil (ES)  M T OI G

Polythrix  E. Watson, 1893

Polythrix octomaculata (Sepp, [1844])  Eight-spotted Longtail  (SE AZ as stray), S Texas to Argentina  M T OI S1 S2 L I G

Polythrix maizae Hellebuyck, 1998  Melanic Longtail  Nicaragua  M T

Polythrix mexicanus H. Freeman, 1969  Mexican Longtail  SE AZ, S TX (as stray), N Mexico to Costa Rica  M T S1 S2 L

Polythrix roma Evans, 1952  TL: Brazil (Pa)  M T

Polythrix asine (Hewitson, 1867)  Asine Longtail  N Mexico to Panama  M T S1 S2 IL L I1 I2

Polythrix hirtius (A. Butler, 1870)  TL: Venezuela  M T

Polythrix gyges Evans, 1952  TL: Peru  M T S

Polythrix minvanes (R. Williams, 1926)  TL: Brazil (MT)  M T G

Polythrix ceculus (Herrich-Schäffer, 1869)  TL: ?  M T

Polythrix caunus (Herrich-Schäffer, 1869)  Four-spotted Longtail  E Mexico to S Brazil and Paraguay  M T S1 S2 L I G

Polythrix auginus (Hewitson, 1867)  Auginus Longtail  SE Mexico to S Brazil  M T S IL L I

Polythrix metallescens (Mabille, 1888)  Metallescens Skipper  SE Mexico to Peru and northern Brazil  M T S L G

Polythrix kanshul Shuey, 1991  Kanshul Longtail  S Mexico, Belize, NE Costa Rica  M S L I1 I2

Polythrix eudoxus (Stoll, 1781)  TL: Surinam  M OI L

Cephise  Evans, 1952

Cephise cephise (Herrich-Schäffer, 1869)  TL: ?  M T S

Cephise impunctus Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise glarus (Mabille, 1888)  TL: Brazil (Pa)  M T OI S

Cephise nuspesez Burns, 1996  Burns' Scarlet-eye  E Mexico to Costa Rica  M T S1 S2 I1 I2

Cephise malesedis Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise maculatus Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise burnsi Austin & O. Mielke, 2000  TL: Brazil (ES)  M T OI

Cephise mexicanus Austin & O. Mielke, 2000  Mexican Scarlet-eye  SLP, OAX (Mexico)  M T

Cephise procerus (Plötz, 1880)  TL: Brazil (Pa)  M T

Cephise aelius (Plötz, 1880)  Longtailed Scarlet-eye  E & S Mexico to Venezuela and Colombia  M T S1 S2 L I1 I2 G

Cephise callias (Mabille, 1888)  TL: [Venezuela]  M T OI S IL

Cephise guatemalaensis (H. Freeman, 1977)  Guatemalan Scarlet-eye  Guatemala, Chiapas, Quintana Roo  M T

Venada  Evans, 1952

Venada [n. sp.] N/A, N/A  Cryptic Scarlet-eye  ["advena"], Mexico, Guatemala, (Nicaragua?)  M S

Venada nevada Burns, 2005  Nevada Scarlet-eye  Costa Rica  M T OD S1 S2 I1 I2 G

Venada advena (Mabille, 1889)  Advena Scarlet-eye  Panama and Colombia, at least  M T OI

Venada daneva Burns, 2005  Daneva Scarlet-eye  Costa Rica  M T OD S1 S2 L I1 I2 G

Venada lamella Burns, 2013  Lamella Scarlet-eye  Mexico, Guatemala, (Nicaragua?), Costa Rica, Panama  M T S1 S2 G

Venada cacao Burns, 2005  Cacao Scarlet-eye  Costa Rica and Panama  M T OD S I G

Venada naranja Burns, 2005  Orange Scarlet-eye  Costa Rica  M T OD S1 S2 I1 I2 G

Heronia  Mabille & Boullet, 1912

Heronia labriaris (A. Butler, 1877)  TL: Brazil (Am)  M T S IL

Chrysoplectrum  E. Watson, 1893

Chrysoplectrum pervivax (Hübner, [1819])  Pervivax Skipper  Costa Rica to Bolivia, Peru and N Brazil  M T OI S1 S2 HS I

Chrysoplectrum bahiana (Herrich-Schäffer, 1869)  TL: [Brazil (Ba)]  M T OI S

Chrysoplectrum orphne (Plötz, 1881)  TL: Brazil (RJ)  M T TI S

Chrysoplectrum otriades (Hewitson, 1867)  TL: Brazil (Am)  M T S

Chrysoplectrum epicincea (A. Butler & H. Druce, 1872)  Stub-tailed Skipper  SE Mexico to Costa Rica  M T S G

Chrysoplectrum perniciosus (Herrich-Schäffer, 1869)  Perniciosus Skipper  Panama to Bolivia and C Brazil  M T S L

Chrysoplectrum cuminaensis R.F. d'Almeida, 1976  TL: Brazil (Pa)  M T

Chrysoplectrum orpheus (Plötz, 1881)  TL: Brazil (Pa)  M T TI

Chrysoplectrum perna Evans, 1952  TL: Colombia  M T

Chrysoplectrum albovenae E. Bell, 1932  TL: Brazil (SC)  M T

Zestusa  Lindsey, 1925

Zestusa dorus (W. H. Edwards, 1882)  Short-tailed Skipper (Northern Zestusa)  far S CO-UT, AZ, NM, W TX, N Mexico  M T S L1 L2 I Ha G

Zestusa elwesi (Godman & Salvin, 1893)  Mexican Zestusa  Durango to Oaxaca  M T S L G

Zestusa staudingeri (Mabille, 1888)  Southern Zestusa  Chiapas to El Salvador  M T OI S L

Zestusa levona Steinhauser, 1972  Levona's Zestusa  Chiapas to El Salvador  M T

Lobocla  Moore, 1884

Lobocla liliana (Atkinson, 1871)  Asian species, shown here for comparison

Lobocla liliana ignatius (Plötz, 1882)  Asian species, shown here for comparison  M T S

Lobocla liliana liliana (Atkinson, 1871)  Asian species, shown here for comparison  M S G

Lobocla aborica (Tytler, 1915)  Asian species, shown here for comparison

Lobocla aborica aborica (Tytler, 1915)  Asian species, shown here for comparison  M T G

Lobocla aborica zesta Evans, 1952  Asian species, shown here for comparison  M T

Lobocla aborica tonka Evans, 1952  Asian species, shown here for comparison  M T

Lobocla quadripunctata Fan & Wang, 2004  Asian species, shown here for comparison  M T S G

Lobocla disparalis Murayama, 1995  Asian species, shown here for comparison  M OI OD

Lobocla bifasciatus (Bremer & Grey, 1853)  Asian species, shown here for comparison  M T S G

Lobocla contractus (Leech, 1894)  Asian species, shown here for comparison  M T S G

Lobocla germanus (Oberthür, 1886)  Asian species, shown here for comparison  M T S G

Lobocla nepos (Oberthür, 1886)  Asian species, shown here for comparison

Lobocla nepos nepos (Oberthür, 1886)  Asian species, shown here for comparison  M T S G

Lobocla nepos phyllis (Hemming, 1933)  Asian species, shown here for comparison  M S

Lobocla proximus (Leech, 1891)  Asian species, shown here for comparison  M T S G

Lobocla simplex (Leech, 1891)  Asian species, shown here for comparison  M T S G

Codatractus  Lindsey, 1921

Codatractus carlos Evans, 1952  Carlos' Mottled-Skipper  TL: Honduras

Codatractus carlos carlos Evans, 1952  C Mexico to Colombia (at least)  M T S L G

Codatractus carlos arguta Evans, 1952  TL: Trinidad  M T

Codatractus carlos rowena Evans, 1952  TL: Venezuela  M T G

Codatractus alcaeus (Hewitson, 1867)  White-crescent Mottled-Skipper  TL: Nicaragua

Codatractus alcaeus alcaeus (Hewitson, 1867)  S Texas to Costa Rica  M T S L I G

Codatractus alcaeus apulia Evans, 1952  Panama (?) to W Ecuador  M T S G

Codatractus yucatanus H. Freeman, 1977  Yucatan Mottled-Skipper  Yucatan Peninsula  M T

Codatractus aminias (Hewitson, 1867)  TL: Brazil (MG)  M T L

Codatractus arizonensis (Skinner, 1905)  Arizona Mottled-Skipper  S AZ, S NM, W TX to Oaxaca  M T S L1 L2 I H G

Codatractus sallyae A. Warren, 1995  Sally’s Mottled-Skipper  W Mexico, Yucatan Peninsula, Guatemala  M T S L

Codatractus melon (Godman & Salvin, 1893)  Melon Mottled-Skipper  N Mexico to Costa Rica  M T S1 S2 L I

Codatractus bryaxis (Hewitson, 1867)  Golden Mottled-Skipper  N Mexico to Honduras  M T S L G

Codatractus imalena (A. Butler, 1872)  Bluish Mottled-Skipper  Costa Rica to northern Brazil  M T OI S1 S2 L I1 I2

Codatractus cyledis (Dyar, 1912)  Cyledis Mottled-Skipper  S Mexico  M T S L Ha G

Codatractus cyda (Godman, 1901)  Cyda Mottled-Skipper  Chiapas to Nicaragua  M T OI S

Codatractus uvydixa (Dyar, 1914)  Variable Mottled-Skipper  S Mexico: Guerrero to Chiapas

Codatractus uvydixa dark segregate ,  W Mexico: Sonora to Michoacan  M S L I

Codatractus uvydixa uvydixa (Dyar, 1914)  TL: Mexico (Gue)  M T S L Ha

Codatractus valeriana (Plötz, 1881)  Valeriana Skipper  [= mysie], SE Arizona to Oaxaca  M T S HS L I H F Ha G

Codatractus hyster (Dyar, 1916)  Hyster Skipper  Mexico, El Salvador  M T S L G

Ridens  Evans, 1952

Ridens crison (Godman & Salvin, 1893)  Many-spotted Ridens  TL: Guatemala

Ridens crison crison (Godman & Salvin, 1893)  SE Mexico, Guatemala  M T S G

Ridens crison howarthi Steinhauser, 1974  El Salvador  M T G

Ridens crison cachinnans (Godman, 1901)  Costa Rica, Panama  M T OI S I G

Ridens ridens (Hewitson, 1876)  Frosted Skipper  Costa Rica & Panama  M T S G

Ridens fieldi Steinhauser, 1974  Field's Ridens  Guatemala (Volcan Sta. Maria)  M T G

Ridens toddi Steinhauser, 1974  Todd’s Skipper  S Mexico (CHIS) to El Salvador  M T S G

Ridens biolleyi (Mabille, 1900)  Biolley's Ridens  Costa Rica, Panama  M T S1 S2 I1 I2 I3

Ridens mercedes Steinhauser, 1983  White-tailed Ridens  SW Mexico  M T S G

Ridens harpagus (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Ridens philistus (Hopffer, 1874)  TL: Peru

Ridens philistus philistus (Hopffer, 1874)  TL: Peru  M T S

Ridens philistus philia Evans, 1952  TL: Colombia  M T S

Ridens mephitis (Hewitson, 1876)  Hewitson's Ridens  SE Mexico to Peru and Bolivia  M T S1 S2 I1 I2

Ridens fulminans (Herrich-Schäffer, 1869)  TL: ?  M T S

Ridens fulima Evans, 1952  TL: Brazil (ES)  M T S L

Ridens panche (R. Williams, 1927)  Panche Ridens  Costa Rica to Colombia and Ecuador  M T S1 S2 I1 I2

Ridens pacasa (R. Williams, 1927)  TL: Bolivia  M T S

Ridens bidens Austin, 1998  TL: Brazil (Rd)  M OI

Ridens allyni H. Freeman, 1979  Allyn’s Ridens  SE Mexico, Guatemala  M T1 T2 G

Ridens miltas (Godman & Salvin, 1893)  Mexican Ridens  E Mexico  M T S G

Ridens bridgmani (Weeks, 1902)  TL: Bolivia  M T S

Ridens nora Evans, 1952  TL: Peru  M T S

Ridens tristis (Draudt, [1922])  TL: [Bolivia]  M T S G

Ridens telegonoides (Mabille & Boullet, 1912)  TL: Colombia  M T OI

Urbanus  Hübner, [1807]

Urbanus proteus (Linnaeus, 1758)  Long-tailed Skipper  TL: America

Urbanus proteus proteus (Linnaeus, 1758)  continental; E USA, S CA, S AZ, S NM, TX to Argentina  M T S1 S2 HS IL L1 L2 I1 I2 I3 H G

Urbanus proteus domingo (Scudder, 1872)  Bahamas and Antilles  M S

Urbanus viterboana (Ehrmann, 1907)  Bluish Longtail  E & W Mexico to Colombia and Ecuador  M T S1 S2 L I G

Urbanus dubius Steinhauser, 1981  TL: Colombia  M T

Urbanus belli (Hayward, 1935)  Bell's Longtail  W Mexico & S Texas (as stray) to Argentina  M T TI S1 S2 L I1 I2 G

Urbanus huancavillcas (R. Williams, 1926)  TL: Ecuador  M T S

Urbanus pronta Evans, 1952  Spot-banded Longtail  E & W Mexico to Argentina  M T S1 S2 L I1 I2

Urbanus longicaudus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus parvus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus pronus Evans, 1952  Pronus Longtail  (S TX one-time stray), E & S Mexico to Argentina  M T S L

Urbanus esmeraldus (A. Butler, 1877)  Esmeralda Longtail  (S TX as stray), E & W Mexico to Peru & Guianas, Trinidad  M T S1 S2 L I G

Urbanus esma Evans, 1952  Esma Longtail  Panama to Brazil, Trinidad  M T S

Urbanus evona Evans, 1952  Turquoise Longtail  E & W Mexico to Colombia  M T S1 S2 L I G

Urbanus elmina Evans, 1952  Andean Longtail  (Costa Rica?), Colombia, Ecuador  M T S

Urbanus viridis H. Freeman, 1970  Rare Longtail  SE Mexico (VER)  M T

Urbanus esta Evans, 1952  Esta Longtail  E & W Mexico to Ecuador and Brazil  M T S1 S2 L I

Urbanus prodicus E. Bell, 1956  Montane Longtail  E & W Mexico to Costa Rica; often in cloud forest  M T S L

Urbanus magnus Steinhauser, 1981  TL: Ecuador  M T

Urbanus velinus (Plötz, 1880)  Velinus Skipper  (SE Mexico?). Guatemala to Brazil and Trinidad  M T S

Urbanus villus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus dorantes (Stoll, 1790)  Dorantes Longtail  TL: Surinam

Urbanus dorantes calafia (R. Williams, 1926)  Baja California Sur  M T S L

Urbanus dorantes dorantes (Stoll, 1790)  continental; Florida, S AZ, S NM, TX to Argentina  M T S HS L1 L2 L3 I1 I2 H G

Urbanus dorantes cramptoni W. Comstock, 1944  Hispaniola, Puerto Rico, Mona, Culebra, Virgin Is., Gr. Cayman  M T S

Urbanus dorantes santiago (Lucas, 1857)  Cuba, N Andros, Mangrove Cay, Grand Bahama  M T S

Urbanus dorantes galapagensis (F. Williams, 1911)  TL: Galapagos  M T

Urbanus obscurus (Hewitson, 1867)  Dark Longtail  Lesser Antilles; St. Martin S to Grenadines, Barbados  M T S L

Urbanus teleus (Hübner, 1821)  Teleus Longtail  S Texas to Argentina  M T S1 S2 Ab IL L1 L2 I G

Urbanus tanna Evans, 1952  Tanna Longtail  S Texas (as stray) to Ecuador, Guianas, Trinidad  M T S L G

Urbanus ambiguus de Jong, 1983  TL: Surinam  M T

Urbanus cindra Evans, 1952  TL: Bolivia  M T S

Urbanus zagorus (Plötz, 1880)  TL: Brazil (RS)  M T

Urbanus simplicius (Stoll, 1790)  Plain Longtail  S Texas (as stray) to Argentina  M T S L1 L2 I1 I2 G

Urbanus procne (Plötz, 1881)  Brown Longtail  S Texas, SE AZ/SW NM (as stray), to Argentina  M S L1 L2 L3 I G

Urbanus evenus (Ménétriés, 1855)  TL: Brazil  M S IL

Urbanus trebia (Möschler, 1879)  White-capped Longtail  El Salvador to Colombia and Trinidad  M T S G

Urbanus carmelita (Herrich-Schäffer, 1869)  Carmelita Longtail  TL: [Brazil]

Urbanus carmelita carmelita (Herrich-Schäffer, 1869)  Panama to Brazil  M T S IL

Urbanus carmelita barra Evans, 1952  TL: Brazil (Ba)  M T OD G

Urbanus reductus (N. Riley, 1919)  TL: Brazil (Am)  M T S

Urbanus doryssus (Swainson, 1831)  White-tailed Longtail  TL: Brazil (Ba)

Urbanus doryssus chales (Godman & Salvin, 1893)  W Mexico  M T S L G

Urbanus doryssus E Mexican segregate ,  ["d. doryssus"], S Texas to El Salvador  M S L1 L2

Urbanus doryssus doryssus (Swainson, 1831)  Chiapas to S Brazil & Bolivia  M T OI OD S L1 L2 I1 I2

Urbanus doryssus albicuspis (Herrich-Schäffer, 1869)  TL: ?  M T

Urbanus albimargo (Mabille, 1876)  White-edged Longtail  TL: Panama; Colombia?

Urbanus albimargo albimargo (Mabille, 1876)  NE Mexico to Venezuela and Ecuador  M T S1 S2 IL L I

Urbanus albimargo takuta Evans, 1952  TL: Guyana  M T S

Urbanus albimargo rica Evans, 1952  TL: Paraguay  M T

Urbanus virescens (Mabille, 1877)  Green-washed Longtail  Panama to S Brazil and Paraguay  M T S L G

Urbanus chalco (Hübner, 1823)  Great White Longtail  Costa Rica to S Brazil and Paraguay  M S IL L G

Astraptes  Hübner, [1819]

Astraptes talus (Cramer, 1777)  Green Flasher  E & S Mexico to Argentina; Jamaica, Cuba, Hispaniola, etc.  M T S L G

Astraptes fulgerator (Walch, 1775)  TL: ?

Astraptes fulgerator azul (Reakirt, [1867])  This appears to be a complex of continental species-level taxa.  M T OD S L1 L2 I H Txt

Astraptes fulgerator fulgerator (Walch, 1775)  TL: ?  M OI G

Astraptes fulgor (Hayward, 1939)  TL: Argentina  M TI

Astraptes naxos (Hewitson, 1867)  TL: Brazil  M T S

Astraptes xagua (Lucas, 1857)  Antillean Flasher  TL: Cuba

Astraptes xagua xagua (Lucas, 1857)  Cuba, Andros  M T S

Astraptes xagua christyi (Sharpe, 1898)  Hispaniola  M T S

Astraptes apastus (Cramer, 1777)  Broad-banded Flasher  TL: Surinam

Astraptes apastus apastus (Cramer, 1777)  SE Mexico to Guianas and N Brazil  M S L I G

Astraptes apastus pusa Evans, 1952  TL: Brazil (Ba)  M T

Astraptes megalurus (Mabille, 1877)  Long-tailed Flasher  E & W Mexico to Colombia  M T S L G

Astraptes tucuti (R. Williams, 1927)  Tucuti Flasher  SE Mexico to Peru and Bolivia  M T OI S L

Astraptes brevicauda (Plötz, 1886)  Short-tailed Flasher  [= talthybius], SE Mexico to Costa Rica  M T S1 S2 L I

Astraptes palliolum (H. Druce, 1908)  Ochre-stained Flasher  Costa Rica  M T OI S

Astraptes phalaecus (Godman & Salvin, 1893)  Yellow-edged Flasher  E & W Mexico to Colombia  M T S L G

Astraptes egregius (A. Butler, 1870)  Small-spotted Flasher  TL: ?

Astraptes egregius egregius (A. Butler, 1870)  S TX (rare stray) to Colombia and Venezuela  M T S L Ha G

Astraptes egregius coxeyi (R. Williams, 1931)  TL: Ecuador  M T

Astraptes erycina (Plötz, 1881)  TL: Brazil  M S IL G

Astraptes halesius (Hewitson, 1877)  TL: French Guiana  M T S

Astraptes mabillei Steinhauser, 1989  TL: Bolivia  M T S

Astraptes aulus (Plötz, 1881)  Green-tinged Flasher  Nicaragua to Peru and Argentina  M T OI S G

Astraptes enotrus (Stoll, 1781)  White-spotted Flasher  E & W Mexico to S Brazil, Paraguay, Bolivia  M T S L G

Astraptes janeira (Schaus, 1902)  Schaus' Flasher  TL: Brazil (RJ)  M T S L

Astraptes elorus (Hewitson, 1867)  TL: ?  M T S L

Astraptes alardus (Stoll, 1790)  Frosted Flasher  TL: Surinam

Astraptes alardus latia Evans, 1952  S TX (rare stray), E Mexico to Colombia  M T S1 S2 L I Ha

Astraptes alardus aquila Evans, 1952  TL: Colombia  M T

Astraptes alardus alardus (Stoll, 1790)  TL: Surinam  M T OD S L G

Astraptes habana (Lucas, 1857)  Frosty Flasher  TL: Cuba

Astraptes habana habana (Lucas, 1857)  Cuba, Isla de Juventud  M S

Astraptes habana heriul (Mabille & Boullet, 1912)  Hispaniola  M T S

Astraptes alector (C. Felder & R. Felder, 1867)  TL: Colombia

Astraptes alector hopfferi (Plötz, 1881)  [= gilberti], S TX (stray), E & W Mexico to NW Brazil and Bolivia  M T S L

Astraptes alector alector (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Astraptes cretatus (Hayward, 1939)  Cretatus Flasher  TL: Ecuador

Astraptes cretatus cretatus (Hayward, 1939)  [= alfius], Nicaragua, Colombia to Bolivia  M T G

Astraptes cretatus adoba Evans, 1952  TL: Brazil (ES)  M T S

Astraptes creteus (Cramer, 1780)  Whitened Flasher  TL: Surinam

Astraptes creteus crana Evans, 1952  SE Mexico to Panama  M T S

Astraptes creteus crilla Evans, 1952  TL: Ecuador  M T

Astraptes creteus cyprus Evans, 1952  TL: Bolivia  M T

Astraptes creteus creteus (Cramer, 1780)  Panama to Guianas and Bolivia  M S IL G

Astraptes creteus siges (Mabille, 1903)  TL: Brazil  M T

Astraptes bifascia (Herrich-Schäffer, 1869)  Green-headed Flasher  SE Mexico to Panama  M T S

Astraptes latimargo (Herrich-Schäffer, 1869)  White-edged Flasher  Panama, Colombia, Ecuador  M T OI S

Astraptes tinda Evans, 1952  TL: Brazil (Pa)  M T

Astraptes chiriquensis (Staudinger, 1876)  Chiriqui Flasher  TL: Panama

Astraptes chiriquensis chiriquensis (Staudinger, 1876)  E Mexico to Venezuela and Ecuador  M T S IL L

Astraptes chiriquensis erana Evans, 1952  TL: Ecuador  M T

Astraptes chiriquensis meretrix (Hewitson, 1876)  TL: Ecuador  M T

Astraptes chiriquensis oenander (Hewitson, 1876)  TL: Brazil (Pa)  M IL G

Astraptes weymeri (Plötz, 1882)  Weymer’s Flasher  TL: ?  M S HS

Astraptes jaira (A. Butler, 1870)  Jamaican Flasher  Jamaica  M T S L

Astraptes cassander (Fabricius, 1793)  Cuban Flasher  Cuba and Isla de Juventud  M T S G

Astraptes galesus (Mabille, 1888)  TL: Peru

Astraptes galesus cassius Evans, 1952  Costa Rica to Venezuela  M T S L G

Astraptes galesus galesus (Mabille, 1888)  TL: Peru  M T OI

Astraptes anaphus (Cramer, 1777)  TL: Surinam

Astraptes anaphus annetta Evans, 1952  S Texas to Guiana and Peru  M T S1 S2 L1 L2 I G

Astraptes anaphus anaphus (Cramer, 1777)  TL: Surinam  M T

Astraptes anaphus anoma Evans, 1952  TL: Trinidad  M T

Astraptes anaphus aniza Evans, 1952  TL: Peru  M T

Astraptes anaphus anausis (Godman & Salvin, 1896)  Greater Antilles, Virgin Is., Montserrat to Grenada  M T S

Narcosius  Steinhauser, 1986

Narcosius narcosius (Stoll, 1790)  TL: Surinam  M T S G

Narcosius aulina (Evans, 1952)  TL: French Guiana  M T

Narcosius samson (Evans, 1952)  Samson Flasher  SE Mexico to Brazil and Bolivia  M T S1 S2 I1 I2 G

Narcosius mura (R. Williams, 1927)  TL: Brazil (Am)  M T S IL G

Narcosius pseudomura Austin, 1996  TL: Brazil (Rd)  M T OI G

Narcosius colossus (Herrich-Schäffer, 1869)  Colossal Flasher  TL: Venezuela

Narcosius colossus colossus (Herrich-Schäffer, 1869)  SE Mexico to Colombia and Venezuela  M T S1 S2 I1 I2 G

Narcosius colossus granadensis (Möschler, 1879)  TL: Venezuela  M T S G

Narcosius dosula (Evans, 1952)  TL: Brazil (RS)  M T

Narcosius hercules (E. Bell, 1956)  TL: Bolivia  M T

Narcosius parisi (R. Williams, 1927)  TL: Brazil (MT)

Narcosius parisi helen (Evans, 1952)  E & W Mexico to Panama  M T S I G

Narcosius parisi parisi (R. Williams, 1927)  TL: Brazil (MT)  M T

Narcosius odysseus Austin, 1996  TL: Brazil (Rd)  M T OI G

Narcosius nazaraeus Steinhauser, 1986  Nazareus Flasher  SE Mexico to Guianas, Brazil, Peru, Ecuador and Bolivia  M T S L G

Narcosius steinhauseri Austin, 1996  TL: Brazil (Rd)  M T OI G

Calliades  Mabille & Boullet, 1912

Calliades oryx (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T OI S

Calliades zeutus (Möschler, 1879)  Zeutus Banded-Skipper  SE Mexico to Guianas and Bolivia  M T S1 S2 L I G

Autochton  Hübner, 1823

Autochton cellus (Boisduval & Le Conte, [1837])  Golden Banded-Skipper  E-central & SE USA, SE AZ, SW NM, Mexico, S to El Salvador  M T OI OD S1 S2 S3 S4 IL L1 L2 L3 I Ha G Txt

Autochton siermadror Burns, 1984  East-Mexican Banded-Skipper  NE Mexico; cloud forest  M T L G

Autochton pseudocellus (Coolidge & Clémence, [1910])  Sonoran Banded-Skipper  Mexico; apparently extirpated from S Arizona  M T S L Ha G

Autochton cincta (Plötz, 1882)  Chisos Banded-Skipper  W Texas to Guatemala  M T S L I Ha G

Autochton vectilucis (A. Butler, 1872)  Central American Banded-Skipper  Chiapas to Panama  M T OD S L G

Autochton integrifascia (Mabille, 1891)  TL: Brazil (SP/RS)  M S L

Autochton neis (Geyer, 1832)  Broad Banded-Skipper  E & W Mexico to S Brazil and Paraguay  M T S Ab L G

Autochton reflexus (Mabille & Boullet, 1912)  TL: Brazil (SC)  M T S

Autochton longipennis (Plötz, 1882)  Spike Banded-Skipper  E & S Mexico to S Brazil  M T S L I G

Autochton zarex (Hübner, 1818)  Sharp Banded-Skipper  E & S Mexico to Argentina  M T S1 S2 HS L I G

Autochton bipunctatus (Gmelin, [1790])  Gmelin's Banded-Skipper  SE Mexico to Bolivia and Brazil  M T S L I

Autochton sulfureolus (Mabille, 1883)  TL: Brazil  M S G

Autochton itylus Hübner, 1823  TL: Surinam  M OI S L

Thessia  Steinhauser, 1989

Thessia athesis (Hewitson, 1867)  Athesis Cloudywing  Panama, Colombia, Venezuela to Tobago  M T S L G

Thessia jalapus (Plötz, 1881)  Jalapus Cloudywing  S Texas (rare stray) to Colombia  M T S1 S2 L I1 I2 G

Achalarus  Scudder, 1872

Achalarus toxeus (Plötz, 1882)  Coyote Cloudywing  W Mexico and S Texas to Panama  M T S1 S2 L1 L2 I G

Achalarus albociliatus (Mabille, 1877)  White-edged Cloudywing  TL: Colombia; Panama; Guatemala

Achalarus albociliatus albociliatus (Mabille, 1877)  SE AZ & S TX (?) as stray, E & W Mexico to Costa Rica  M T S1 S2 L I G

Achalarus albociliatus leucophrys (Mabille, 1898)  Panama and Colombia  M T S

Achalarus albociliatus nocera (Plötz, 1882)  TL: Colombia  M T OI

Achalarus lyciades (Geyer, 1832)  Hoary Edge  E USA, (Veracruz)  M S1 S2 HS L1 L2 L3 L4 I G

Achalarus casica (Herrich-Schäffer, 1869)  Desert Cloudywing  SW USA (SE AZ to C TX), Mexico  M T S L I H G

Achalarus tehuacana (Draudt, 1922)  Tehuacan Cloudywing  S TX (rare stray), E Mexico to Guatemala  M T OD S L G

Thorybes  Scudder, 1872

Thorybes confusis E. Bell, 1923  Confused Cloudywing  SE USA  M T1 T2 S1 S2 S3 L1 L2 L3 I G

Thorybes bathyllus (J. E. Smith, 1797)  Southern Cloudywing  E USA  M OI TI S1 S2 S3 HS L1 L2 L3 L4 L5 G

Thorybes mexicana (Herrich-Schäffer, 1869)  Mexican Cloudywing  TL: [Mexico]

Thorybes mexicana aemilea (Skinner, 1893)  Cascade Mts. N CA to C OR  M T S L1 L2 Ha

Thorybes mexicana blanca Scott, 1981  White Mts., CA-NV  M S L1 L2

Thorybes mexicana nevada Scudder, 1872  S Rocky Mountains (S WY-ID to NM, AZ), CA-NV Sierra Nevada  M S L1 L2 Ha G

Thorybes mexicana dobra Evans, 1952  NW Mexico, SE Arizona, SW New Mexico  M T S L

Thorybes mexicana mexicana (Herrich-Schäffer, 1869)  C & W Mexico  M T S L Ha G

Thorybes mexicana ducia Evans, 1952  TL: Panama  M OD

Thorybes diversus E. Bell, 1927  Western Cloudywing  C & N Sierra Nevada and Siskiyou Mts., CA, far SW OR  M S L1 L2 L3 I G

Thorybes drusius (W. H. Edwards, [1884])  Drusius Cloudywing, White-fringed Cloudywing  SE AZ, SW NM, W TX, Mexico  M T S L I Ha

Thorybes pylades (Scudder, 1870)  Northern Cloudywing  TL: USA (Ma)

Thorybes pylades indistinctus Austin & J. Emmel, 1998  Baja California (N) to Washington, Cascades (& Ochocos) west  M T S L1 L2 Ha

Thorybes pylades BCS segregate ,  Baja California Sur  M S

Thorybes pylades S Rocky Mountain segregate ,  NE OR, S ID, MT, WY, W SD, W NE, N & C UT, CO, N NM, N AZ  M S

Thorybes pylades southwestern segregate ,  S AZ, S NM, W Mexico to Oaxaca  M S L1 L2 L3 L4 H

Thorybes pylades albosuffusa H. Freeman, 1943  W Texas, NE Mexico  M T S1 S2

Thorybes pylades pylades (Scudder, 1870)  Canada, E USA  M T S1 S2 L1 L2 L3 L4 L5 L6 L7 L8 L9 L10 I G

Cabares  Godman & Salvin, 1894

Cabares potrillo (Lucas, 1857)  Potrillo Skipper  TL: Cuba

Cabares potrillo potrillo (Lucas, 1857)  SE AZ (stray), S TX to Costa Rica; Jamaica, Cuba, Hispaniola  M T S1 S2 L1 L2 I G

Cabares potrillo reducta Mabille & Boullet, 1919  Panama, Colombia, Venezuela, Trinidad  M S

Cabares rinta Evans, 1952  TL: Argentina  M T S

Spathilepia  A. Butler, 1870

Spathilepia clonius (Cramer, 1775)  Falcate Skipper  S TX (sporadic) to N Argentina  M T S1 S2 IL L I G

Oechydrus  E. Watson, 1893

Oechydrus chersis (Herrich-Schäffer, 1869)  TL: ?

Oechydrus chersis chersis (Herrich-Schäffer, 1869)  TL: ?  M T S

Oechydrus chersis ochrilinea (Schaus, 1902)  TL: Peru  M T S

Oechydrus chersis rufus Evans, 1953  TL: Brazil (RJ)  M T OD S

Oechydrus chersis evelinda (A. Butler, 1870)  TL: Brazil (RJ)  M T TI S G

Ectomis  Mabille, 1878

Ectomis cythna (Hewitson, 1878)  TL: ?

Ectomis cythna cythna (Hewitson, 1878)  TL: ?  M T S L

Ectomis cythna ega Evans, 1953  TL: Brazil (Am)  M T OD

Nerula  Mabille, 1888

Nerula fibrena (Hewitson, 1877)  TL: Brazil (Am)  M T OI S

Nerula tuba Evans, 1953  TL: Brazil (Pa)  M T OD

Marela  Mabille, 1903

Marela tamyroides (C. Felder & R. Felder, 1867)  TL: Brazil (Am)  M T S L

Marela tamyris Mabille, 1903  TL: Brazil (Am)

Marela tamyris tamyris Mabille, 1903  TL: Brazil (Am)  M T TI S L

Marela tamyris tamba Evans, 1953  TL: Peru  M T S

Cogia  A. Butler, 1870

Cogia hassan A. Butler, 1870  TL: Brazil (Pa)

Cogia hassan hassan A. Butler, 1870  TL: Brazil (Pa)  M T S

Cogia hassan evansi E. Bell, 1937  TL: Bolivia  M T S

Cogia abdul Hayward, 1947  TL: Argentina  M TI G

Cogia cajeta (Herrich-Schäffer, 1869)  Yellow-haired Skipper  TL: ?

Cogia cajeta cajeta (Herrich-Schäffer, 1869)  SE Mexico to Nicaragua  M T S L G

Cogia cajeta eluina Godman & Salvin, 1894  W Mexico to Costa Rica  M T S1 S2 L I

Cogia troilus Mabille, 1898  TL: Bolivia  M T S

Cogia punctilia Plötz, 1882  TL: Brazil ([Ba])  M T S

Cogia cerradicola (O. Mielke, 1967)  TL: Brazil (Go)  M T OI S G

Cogia outis (Skinner, 1894)  Outis Skipper  NE Mexico (CHIH, COAH), Texas, S-C OK to far SW MO  M T S L I H Ha G

Cogia hippalus (W. H. Edwards, 1882)  Acacia Skipper  TL: USA (Az)

Cogia hippalus peninsularis L. Miller & MacNeill, 1969  Baja California Sur  M T S

Cogia hippalus hippalus (W. H. Edwards, 1882)  AZ, SW NM, W & S TX, W Mexico  M T S L I H G

Cogia hippalus hiska Evans, 1953  E Mexico to Costa Rica  M T S1 S2 L I G

Cogia hippalus hester Evans, 1953  TL: Venezuela  M T OD

Cogia azila Evans, 1953  TL: Bolivia  M T OD

Cogia caicus (Herrich-Schäffer, 1869)  Gold-costa Skipper  TL: ?

Cogia caicus moschus (W. H. Edwards, 1882)  SE AZ. S NM to OAX  M S L I H G

Cogia caicus caicus (Herrich-Schäffer, 1869)  Chiapas to Costa Rica  M T S1 S2 G

Cogia calchas (Herrich-Schäffer, 1869)  Mimosa Skipper  S Texas to N Argentina  M T S1 S2 L1 L2 I G

Cogia grandis N. Riley, 1921  TL: Brazil (MT)  M T S

Cogia aventinus (Godman & Salvin, 1894)  Trimmed Skipper  W Mexico, Yucatan  M T S1 S2 IL L1 L2 Ha

Cogia buena A. Warren, Dolibaina & Hernández-Mejía, 2015  Nice Gogia  S Mexico  M S

Cogia mala Evans, 1953  Poor Gogia  Guatemala  M T HS

Cogia elaites (Hewitson, 1867)  TL: Brazil (MG)  M T

Paracogia  O. Mielke, 1977

Paracogia acanthopoda O. Mielke, 1977  TL: Brazil (Pr)  M T OI G

Telemiades  Hübner, [1819]

Telemiades delalande (Latreille, [1824])  Delalande Skipper  Guatemala to Venezuela, Peru, upper Amazon  M T OI S L

Telemiades litanicus (Hewitson, 1876)  TL: Brazil (Am)  M T S

Telemiades vansa Evans, 1953  TL: Guyana  M T OD

Telemiades gallius (Mabille, 1888)  Centrites Skipper  Costa Rica and Panama, mid-elevation rain forest  M T OI S L I

Telemiades chrysorrhoea (Godman & Salvin, 1893)  Chrysorrhoea Skipper  Costa Rica and Panama, cloud forest  M T S1 S2 L I1 I2

Telemiades centrites (Hewitson, 1870)  TL: Ecuador

Telemiades centrites contra Evans, 1953  TL: Ecuador  M T OD

Telemiades centrites centrites (Hewitson, 1870)  TL: Ecuador  M T S Txt

Telemiades vespasius (Fabricius, 1793)  TL: Indiis  M T

Telemiades squanda Evans, 1953  TL: Brazil (RJ)  M T S IL G

Telemiades trenda Evans, 1953  TL: Surinam  M T OD

Telemiades nicomedes (Möschler, 1879)  Dark Telemiades  SE Mexico to Guianas and upper Amazon  M T S1 S2 I1 I2

Telemiades meris (Plötz, 1886)  Meris Telemiades  TL: Colombia

Telemiades meris meris (Plötz, 1886)  Panama & Colombia  M T S

Telemiades meris brazus E. Bell, 1949  TL: Brazil (SC)  M T TI

Telemiades choricus (Schaus, 1902)  Mexican Telemiades  W & S Mexico, Belize  M T S L G

Telemiades megallus Mabille, 1888  Orange Telemiades  SE Mexico (?) to Costa Rica  M T OI S1 S2 I1 I2 G

Telemiades sila Evans, 1953  Sila Telemiades  Nicaragua to Colombia, Venezuela and Ecuador  M T S

Telemiades epicalus Hübner, [1819]  Hübner's Telemiades  Panama to S Brazil  M T S HS L G

Telemiades penidas (Hewitson, 1867)  Penidas Telemiades  Costa Rica to S Brazil  M T S IL L

Telemiades avitus (Stoll, 1781)  Yellow-spotted Telemiades  SE Mexico to N Amazon  M T OI S L I G

Telemiades antiope (Plötz, 1882)  Plötz's Telemiades  TL: Colombia; Brazil (Pa)

Telemiades antiope antiope (Plötz, 1882)  Nicaragua, Costa Rica, Colombia, S Brazil  M T S1 S2 I1 I2

Telemiades antiope tosca Evans, 1953  TL: Brazil (Am)  M T OD L

Telemiades fides E. Bell, 1949  Small Telemiades  E & W Mexico to Colombia, Ecuador and upper Amazon  M T S1 S2 L I1 I2

Telemiades amphion (Geyer, 1832)  TL: "West Indies"

Telemiades amphion pekahia (Hewitson, 1868)  TL: Venezuela  M T Txt

Telemiades amphion amphion (Geyer, 1832)  TL: "West Indies"  M T S

Telemiades amphion misitheus Mabille, 1888  TL: Peru  M T OI S L

Telemiades amphion marpesus (Hewitson, 1876)  TL: Brazil  M IL G

Telemiades laogonus (Hewitson, 1876)  TL: Brazil

Telemiades laogonus nicola (Plötz, 1882)  TL: Brazil (Pa)  M T

Telemiades laogonus laogonus (Hewitson, 1876)  TL: Brazil  M T

Telemiades corbulo (Stoll, 1781)  Corbulo Skipper  Guianas, Ecuador, Peru, Bolivia, upper and lower Amazon  M T S IL L G

Telemiades oiclus (Mabille, 1889)  Oiclus Skipper  [= cora], Honduras to Panama  M T OI S I1 I2

Bungalotis  E. Watson, 1893

Bungalotis erythus (Cramer, 1775)  Spotted Scarlet-eye  SE Mexico to Guianas, Brazil and Bolivia  M S1 S2 I1 I2 G

Bungalotis diophorus (Möschler, 1883)  Diophorus Scarlet-Eye  Costa Rica to Guianas and N Brazil  M T S1 S2 I1 I2

Bungalotis gagarini O. Mielke, 1967  Gagarin's Scarlet-Eye  TL: Brazil (Go)  M T OI IL G

Bungalotis midas (Cramer, 1775)  White-cheeked Scarlet-eye  SE Mexico to S Brazil and Paraguay  M T OI S1 S2 IL L I1 I2 G

Bungalotis aureus Austin, 2008  Golden Scarlet-Eye  TL: Ecuador  M OI G

Bungalotis astylos (Cramer, 1780)  Dark-cheeked Scarlet-eye  SE Mexico to Guianas, Peru and S Brazil  M T S1 S2 L I1 I2 G

Bungalotis milleri H. Freeman, 1977  Miller's Scarlet-eye  SE Mexico to Costa Rica  M T OD G

Bungalotis quadratum (Sepp, [1845])  Pallid Scarlet-eye  TL: Surinam

Bungalotis quadratum quadratum (Sepp, [1845])  SE Mexico to Guianas  M T OI S1 S2 L I1 I2

Bungalotis quadratum barba Evans, 1952  TL: Brazil (Pa)  M T

Bungalotis sipa de Jong, 1983  Sipa Scarlet-eye  Surinam  M T G

Bungalotis clusia Evans, 1952  Clusia Scarlet-eye  TL: Brazil (Pa)  M T

Bungalotis borax Evans, 1952  Borax Scarlet-eye  TL: French Guiana  M T S G

Bungalotis lactos Evans, 1952  Lactos Scarlet-eye  TL: Peru  M T G

Salatis  Evans, 1952

Salatis canalis (Skinner, 1920)  Canal Scarlet-eye  Costa Rica to Ecuador  M T S1 S2 I1 I2

Salatis salatis (Stoll, 1782)  Variable Scarlet-eye  Nicaragua, Panama to Guianas, Peru and S Brazil  M T S L G

Salatis cebrenus (Cramer, 1777)  TL: Surinam  M T OI

Salatis fulvius (Plötz, 1882)  TL: Brazil (Pa)  M T S HS

Salatis scyrus (E. Bell, 1934)  TL: Peru  M T

Salatis flavomarginatus (Sepp, [1851])  TL: Surinam  M OI I

Sarmientoia  Berg, 1897

Sarmientoia dinka Evans, 1952  TL: ?  M T

Sarmientoia eriopis (Hewitson, 1867)  TL: Brazil (Pa)  M T S

Sarmientoia phaselis (Hewitson, 1867)  TL: Brazil  M T S IL G

Sarmientoia faustinus (Burmeister, 1878)  TL: Argentina  M T S IL G

Sarmientoia haywardi O. Mielke, 1967  TL: Bolivia  M OI S IL G

Sarmientoia similis O. Mielke, 1967  TL: Brazil (Pa)  M OI S IL G

Sarmientoia almeidai O. Mielke, 1967  TL: Brazil (MT)  M OI S G

Sarmientoia browni O. Mielke, 1967  TL: Brazil (DF)  M OI S G

Nicephellus  Austin, 2008

Nicephellus nicephorus (Hewitson, 1876)  Two-spotted Scarlet-eye  SE Mexico to Guianas, N & S Brazil, NE Peru  M T S IL I

Euriphellus  Austin, 2008

Euriphellus euribates (Stoll, 1782)  Euribates Scarlet-eye  TL: Surinam

Euriphellus euribates euribates (Stoll, 1782)  Costa Rica to Brazil and Bolivia  M T OI S G

Euriphellus euribates polygius (Latreille, [1824])  S Brazil  M T IL

Dyscophellus  Godman & Salvin, 1893

Dyscophellus phraxanor (Hewitson, 1876)  Big-spotted Scarlet-eye  TL: Colombia; Panama

Dyscophellus phraxanor lama Evans, 1952  SE Mexico to Honduras  M T S

Dyscophellus phraxanor phraxanor (Hewitson, 1876)  Nicaragua to Ecuador  M T S1 S2 L I G

Dyscophellus phraxanor mena Evans, 1952  TL: Ecuador  M T

Dyscophellus marian Evans, 1952  TL: Ecuador  M T S

Dyscophellus porcius (C. Felder & R. Felder, 1862)  Fiery Scarlet-eye  TL: Brazil (Am)

Dyscophellus porcius porcius (C. Felder & R. Felder, 1862)  S Mexico to Paraguay and S Brazil  M T S L G

Dyscophellus porcius doriscus (Hewitson, 1867)  TL: Brazil ([RJ])  M T S HS

Dyscophellus sebaldus (Stoll, 1781)  Sebaldus Scarlet-eye  Costa Rica to Guianas and N Brazil  M OI S IL

Dyscophellus erythras (Mabille, 1888)  TL: Brazil (Pa)  M T OI S

Dyscophellus mielkei Austin, 2008  TL: Brazil (Rd)  M OI

Dyscophellus diaphorus (Mabille & Boullet, 1912)  Diaphorus Scarlet-eye  Costa Rica to French Guiana and N Brazil  M T OI S

Dyscophellus porsena (E. Bell, 1934)  TL: Peru  M T

Dyscophellus ramon Evans, 1952  Plain Scarlet-eye  SE Mexico to Panama  M T S1 S2 I1 I2 G

Dyscophellus ramusis (Stoll, 1781)  Subtle Scarlet-eye  Costa Rica to S Brazil  M S L

Dyscophellus damias (Plötz, 1882)  TL: Brazil  M T S

Nascus  E. Watson, 1893

Nascus phintias Schaus, 1913  Schaus’ Scarlet-eye  SE Mexico to French Guiana and Peru  M T S1 S2 I

Nascus phocus (Cramer, 1777)  Common Scarlet-eye  E & W Mexico to S Brazil, Paraguay and Argentina  M T S1 S2 L I1 I2 G

Nascus solon (Plötz, 1882)  TL: S. Amer.

Nascus solon corilla Evans, 1952  S Mexico to Colombia and Venezuela  M T S1 S2 I1 I2

Nascus solon solon (Plötz, 1882)  TL: S. Amer.  M S L

Nascus broteas (Cramer, 1780)  Broteas Scarlet-eye  E Mexico to Argentina, S Brazil and Paraguay  M T S1 S2 HS I1 I2 G

Pseudonascus  Austin, 2008

Pseudonascus paulliniae (Sepp, [1842])  Least Scarlet-eye  S Mexico to Bolivia and S Brazil  M T OI S1 S2 I1 I2 G

Pseudonascus prax Evans, 1952  TL: Brazil (Pa)  M T

Porphyrogenes  E. Watson, 1893

Porphyrogenes boliva Evans, 1952  TL: Venezuela  M T

Porphyrogenes ferruginea (Plötz, 1883)  TL: Brazil (Ba)  M T S

Porphyrogenes despecta (A. Butler, 1870)  TL: Brazil (Pa)  M T S

Porphyrogenes omphale (A. Butler, 1871)  Blue Scarlet-eye  Costa Rica, Panama, Venezuela, Guianas to Bolivia  M T OI S1 S2 L G

Porphyrogenes pausias (Hewitson, 1867)  TL: Brazil (Am)  M T S

Porphyrogenes probus (Möschler, 1877)  TL: Surinam  M T

Porphyrogenes vulpecula (Plötz, 1882)  TL: [Brazil (Pa)]  M T S G

Porphyrogenes sororcula (Mabille & Boullet, 1912)  TL: French Guiana  M T OI S

Porphyrogenes specularis Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes spanda Evans, 1952  TL: Brazil (Pa)  M T S

Porphyrogenes stresa Evans, 1952  TL: Peru  M T

Porphyrogenes zohra (Möschler, 1879)  Zohra Scarlet-eye  Honduras, Nicaragua, Venezuela, Guianas, N Brazil (Para)  M T S G

Porphyrogenes sparus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes speciosus Austin & O. Mielke, 2008  Speciosus Scarlet-eye  Costa Rica  M T G

Porphyrogenes peterwegei Burns, 2010  Peter's Scarlet-eye  NW Costa Rica  M T1 T2 T3 S I1 I2 H G

Porphyrogenes convexus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes spadix Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes splendidus Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes simulator Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes sula (R. Williams & E. Bell, 1940)  Sula Scarlet-eye  Honduras, possibly Costa Rica  M T S I G

Porphyrogenes spina Austin & O. Mielke, 2008  Spina Scarlet-eye  Panama  M T G

Porphyrogenes sporta Austin & O. Mielke, 2008  TL: Colombia  M OI

Porphyrogenes glavia Evans, 1952  Glavia Scarlet-eye  Panama  M T G

Porphyrogenes sparta Evans, 1952  TL: Brazil (Pa)  M T

Porphyrogenes spoda Evans, 1952  Spoda Scarlet-eye  Panama  M T G

Porphyrogenes stupa Evans, 1952  TL: ?  M T

Porphyrogenes passalus (Herrich-Schäffer, 1869)  Passalus Scarlet-eye  Panama, Colombia, Bolivia, Brazil (at least)  M T S

Porphyrogenes virgatus (Mabille, 1888)  Virgatus Scarlet-eye  Honduras, Guyana, Peru  M T OI

Porphyrogenes eudemus (Mabille, 1888)  Eudemus Scarlet-eye  Panama, Peru  M T OI TI S G

Porphyrogenes unassociated_unnamed_female_#1 ,  TL: not specified  M S IL

Porphyrogenes unassociated_unnamed_female_#2 ,  TL: not specified  M IL

Porphyrogenes unassociated_unnamed_female_#3 ,  Mystery Scarlet-eye #3  Panama  M S

Porphyrogenes unassociated_unnamed_female_#4 ,  Mystery Scarlet-eye #4  Panama  M S

Porphyrogenes unassociated_unnamed_female_#5 ,  Mystery Scarlet-eye #5  Costa Rica  M S

Porphyrogenes unassociated_unnamed_females ,  Mystery Female Scarlet-eyes  Porphyrogenes females whose identity remains a mystery  M S HS

Porphyrogenes unassociated_unnamed_male ,  Mystery Scarlet-eye #6  Costa Rica  M S G

Ocyba  Lindsey, 1925

Ocyba calathana (Hewitson, 1868)  Yellow-rimmed Scarlet-eye  TL: Colombia

Ocyba calathana calanus (Godman & Salvin, 1894)  Mexico to Costa Rica  M T OI S1 S2 IL L I G

Ocyba calathana calathana (Hewitson, 1868)  Panama and Colombia  M T S IL

Ocyba calathana compusa (Hewitson, 1868)  TL: Brazil (Am)  M T

Oileides  Hübner, [1825]

Oileides azines (Hewitson, 1867)  TL: Brazil (Am)  M T OI S L

Oileides fenestratus (Gmelin, [1790])  TL: [French Guiana]  M T S

Oileides vulpinus Hübner, [1825]  TL: ?

Oileides vulpinus guyanensis (Mabille & Boullet, 1912)  TL: French Guiana  M T S

Oileides vulpinus vulpinus Hübner, [1825]  TL: ?  M OI S

Oileides amazonensis (E. Bell, 1947)  TL: Colombia

Oileides amazonensis amazonensis (E. Bell, 1947)  TL: Colombia  M T S G

Oileides amazonensis renta (Evans, 1952)  TL: Peru  M T G

Aurina  Evans, 1937

Aurina dida Evans, 1937  TL: "Ivory Coast"  M T OI OD S G Txt

  Subfamily Pyrginae

  Tribe Pyrrhopygini

Azonax  Godman & Salvin, 1893

Azonax typhaon (Hewitson, 1877)  Typhaon Skipper  SE Mexico (OAX), Nicaragua, Colombia, Guyana  M T S L

Myscelus  Hübner, [1819]

Myscelus nobilis (Cramer, 1777)  Noble Myscelus  Surinam, Venezuela, Peru, Brazil, Bolivia  M T S HS IL

Myscelus amystis (Hewitson, 1867)  Widespread Myscelus  TL: Colombia

Myscelus amystis hages Godman & Salvin, 1893  E & W Mexico to Panama  M T S1 S2 HS L I Ha G

Myscelus amystis muttra Evans, 1951  Colombia (Valle del Cauca)  M T

Myscelus amystis amystis (Hewitson, 1867)  Trinidad, Venezuela, Colombia  M T TI S

Myscelus amystis distinctus Röber, 1925  Ecuador  M S G

Myscelus amystis mysus Evans, 1951  Peru, Bolivia  M T S

Myscelus amystis epigona Herrich-Schäffer, 1869  S Brazil, Paraguay, Argentina  M T TI S HS IL

Myscelus phoronis (Hewitson, 1867)  Phoronis Myscelus  TL: Colombia

Myscelus phoronis phoronis (Hewitson, 1867)  Guatemala, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia  M T TI S IL L

Myscelus phoronis caucanus Staudinger, 1888  Colombia  M T S

Myscelus belti Godman & Salvin, 1879  Belt's Myscelus  SE Mexico (OAX, VER, CHIS) to Venezuela, Colombia & Ecuador

Myscelus belti belti Godman & Salvin, 1879  SE Mexico (OAX, VER, CHIS) to Venezuela, Colombia & Ecuador  M T TI S I1 I2 I3

Myscelus belti caucaensis Constantino, Salazar & Vargas, 2013  W Colombia

Myscelus perissodora Dyar, 1914  Dyar's Myscelus  S Mexico (GRO, OAX, VER) to Colombia & Venezuela  M T S L I G

Myscelus pegasus Mabille, 1903  Pegasus Myscelus  French Guiana, Venezuela, Ecuador  M T TI S

Myscelus epimachia Herrich-Schäffer, 1869  Red Myscelus  TL: ?

Myscelus epimachia verina Evans, 1951  Ecuador  M T S

Myscelus epimachia epimachia Herrich-Schäffer, 1869  Venezuela, Colombia, Ecuador, Peru, Bolivia, S Brazil  M S IL

Myscelus epimachia edix Evans, 1951  SE Brazil, Paraguay  M T S

Myscelus draudti N. Riley, 1926  Draudt's Myscelus  Peru, Bolivia  M T S

Myscelus santhilarius (Latreille, [1824])  Hilarious Myscelus  SE Brazil  M T S IL

Myscelus assaricus (Cramer, 1779)  Fiery Myscelus  TL: Surinam

Myscelus assaricus assaricus (Cramer, 1779)  French Guiana, Surinam, N Brazil  M T S G

Myscelus assaricus mapirica (Strand, 1921)  S Venezuela, Peru, Brazil (Rd)  M T S IL

Myscelus assaricus michaeli Nicolay, 1975  SE Mexico (OAX, CHIS), Guatemala, Costa Rica, Panama  M T OI S1 S2 L I G

Myscelus pardalina (C. Felder & R. Felder, 1867)  Flaming Myscelus  TL: Colombia

Myscelus pardalina kesselringi O. Mielke & Casagrande, 2011  Peru  M T OI

Myscelus pardalina guarea Evans, 1951  N Brazil (Para)  M T G

Myscelus pardalina pardalina (C. Felder & R. Felder, 1867)  Colombia, Ecuador, Peru, Bolivia, Brazil  M T S IL

Myscelus pardalina yacutinga O. Mielke & Casagrande, 2011  Argentina & Paraguay  M T OI

Granila  Mabille, 1903

Granila paseas (Hewitson, 1857)  Paseas Skipper  SE Brazil, Paraguay  M T OI S IL

Passova  Evans, 1951

Passova nigrocephala (E. Bell, 1934)  Black-headed Firetip  Colombia  M T S

Passova ganymedes (E. Bell, 1931)  Mimetic Firetip  TL: Colombia

Passova ganymedes ganymedes (E. Bell, 1931)  Colombia  M T S

Passova ganymedes gulia Evans, 1951  Ecuador  M T

Passova ganymedes gelina Evans, 1951  Peru  M T S

Passova gellias (Godman & Salvin, 1893)  Gellias Firetip  Belize, Honduras, Costa Rica, Panama & Venezuela  M T S1 S2 I1 I2

Passova polemon (Hopffer, 1874)  Polemon Firetip  S Brazil  M T S L

Passova glacia Evans, 1951  Shining Firetip  Surinam, French Guiana  M T S

Passova gazera (Hewitson, 1866)  Gazera Firetip  Brazil (Amazonas)  M T TI S

Passova greta Evans, 1951  Great Firetip  Peru, Bolivia  M T S

Passova vilna Evans, 1951  Vilna Firetip  TL: Bolivia

Passova vilna vilna Evans, 1951  Bolivia  M T

Passova vilna vanta Evans, 1951  Bolivia  M T

Passova passova (Hewitson, 1866)  Passova Firetip  TL: Brazil (Am); French Guiana

Passova passova gortyna (Hewitson, 1866)  N Brazil  M T TI S G

Passova passova passova (Hewitson, 1866)  French Guiana  M T S

Passova passova rudex Evans, 1951  N Brazil, Peru  M T S

Passova passova styx (Möschler, 1879)  Colombia, Venezuela, Bolivia  M T S

Passova passova practa Evans, 1951  S Brazil, Paraguay  M T S

Passova passova stacer Evans, 1951  Surinam, Guyana  M T

Aspitha  Evans, 1951

Aspitha aspitha (Hewitson, 1866)  Aspitha Firetip  TL: Brazil (Pa/Am)

Aspitha aspitha parima (Plötz, 1886)  Trinidad, French Guiana, Surinam, N Brazil  M TI

Aspitha aspitha aspitha (Hewitson, 1866)  N Brazil (Pa, Am)  M T TI S

Aspitha aspitha rufescens (N. Riley, 1919)  Brazil (Rd)  M T S

Aspitha leander (Boullet, 1912)  Leander Firetip  SE Mexico (OAX), Panama, Colombia  M T S IL G Txt

Aspitha bassleri (E. Bell, 1940)  Bassler's Firetip  Peru  M T S

Aspitha agenoria (Hewitson, 1876)  Red-lobed Firetip  TL: Peru

Aspitha agenoria agenoria (Hewitson, 1876)  Colombia, Peru, Bolivia  M T S

Aspitha agenoria sanies (H. Druce, 1908)  Peru, Bolivia  M T S IL

Aspitha agenoria cruor (H. Druce, 1908)  Peru  M T

Zonia  Evans, 1951

Zonia zonia Evans, 1951  Zonia Skipper  TL: Brazil (Am)

Zonia zonia panamensis Nicolay, 1975  Belize, Panama  M T OI S G

Zonia zonia zonia Evans, 1951  N Brazil (Am)  M T TI

Zonia zonia diabo O. Mielke & Casagrande, 1998  SE Brazil (SP)  M T

Olafia  Nemésio, 2005

Olafia roscius (Hopffer, 1874)  Olaf's Firetip  TL: Brazil

Olafia roscius iphimedia (Plötz, 1886)  SE Brazil (SP)  M T S IL

Olafia roscius flavomaculata (E. Bell, 1937)  SE Brazil (Pr, SC)  M T

Olafia roscius roscius (Hopffer, 1874)  SE Brazil (RS), Paraguay, Argentina  M T S

Oxynetra  C. Felder & R. Felder, 1862

Oxynetra stangelandi Grishin & Burns, 2013  Stangeland's Firetip  currently known only from ACG in Costa Rica  M T1 T2 L I1 I2

Oxynetra hopfferi Staudinger, 1888  Hopffer's Firetip  SE Mexico (VER), Costa Rica, Panama  M T OI OD TI S G

Oxynetra semihyalina C. Felder & R. Felder, 1862  Felder's Firetip  Colombia, Ecuador, Peru, Bolivia  M T OD S L

Oxynetra confusa Staudinger, 1888  Confusing Firetip  Venezuela (E slope Andes), Ecuador, Peru, Bolivia, [French Guiana?]  M T S

Pyrrhopyge  Hübner, [1819]

Pyrrhopyge zenodorus Godman & Salvin, 1893  Red-headed Firetip  SE Mexico to Costa Rica  M T S L I1 I2 G

Pyrrhopyge evansi E. Bell, 1947  Evans' Firetip  TL: Colombia

Pyrrhopyge evansi evansi E. Bell, 1947  Costa Rica, Panama, Colombia, Venezuela (Bolivar), French Guiana, [Ecuador?]  M T S G

Pyrrhopyge evansi borburata Orellana, [2010]  Venezuela (N slope Cordillera de la Costa, Carabobo)

Pyrrhopyge phidias (Linnaeus, 1758)  Original Firetip  TL: "Asia"

Pyrrhopyge phidias hyperici Hübner, 1823  Brazil (Para)  M S

Pyrrhopyge phidias latifasciata A. Butler, 1873  Colombia  M T S

Pyrrhopyge phidias bixae (Linnaeus, 1758)  Surinam, French Guiana, N Brazil to Matto Grosso  M S

Pyrrhopyge phidias rusca Evans, 1951  Ecuador, Peru, SW Brazil  M T

Pyrrhopyge phidias leucoloma Erschoff, 1875  Peru, Bolivia  M S

Pyrrhopyge phidias garata Hewitson, 1866  Guyana, Surinam, [N Venezuela?]  M T S

Pyrrhopyge phidias phidias (Linnaeus, 1758)  Trinidad, Surinam, Guyana, Venezuela, Colombia, Peru, Bolivia, Paraguay, Brazil  M T S

Pyrrhopyge phidias guianae E. Bell, 1932  Trinidad, Guyana, French Guiana  M T

Pyrrhopyge aziza Hewitson, 1866  Aziza Firetip  TL: Colombia

Pyrrhopyge aziza aziza Hewitson, 1866  Colombia  M T TI S

Pyrrhopyge aziza lexos Evans, 1951  Guyana, Colombia, Ecuador, Peru  M T S

Pyrrhopyge aziza araethyrea Hewitson, 1870  Ecuador  M T TI S

Pyrrhopyge aziza troja Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge aziza arbor Evans, 1951  Colombia, Venezuela  M T S

Pyrrhopyge aziza attis E. Bell, 1931  Bolivia  M T

Pyrrhopyge aziza subnubilus Hayward, 1935  Argentina

Pyrrhopyge pusca Evans, 1951  Pusca Firetip  Peru, Bolivia  M T S

Pyrrhopyge proculus Hopffer, 1874  Proculus Firetip  TL: Guyana

Pyrrhopyge proculus lina E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge proculus cardus Mabille, 1891  N Brazil  M T S

Pyrrhopyge proculus hyleus Mabille, 1891  N Brazil  M T

Pyrrhopyge proculus cintra Evans, 1951  French Guiana, Surinam, Guyana, N Brazil, NE Peru  M T S

Pyrrhopyge proculus draudti E. Bell, 1931  Venezuela, Colombia, Ecuador, Peru, Bolivia (Andean)  M T S

Pyrrhopyge proculus proculus Hopffer, 1874  Trinidad, Guyana, Venezuela (E Cordillera de la Costa), Colombia, N Brazil  M T

Pyrrhopyge infantilis H. Druce, 1908  Infant Firetip  TL: Peru

Pyrrhopyge infantilis infantilis H. Druce, 1908  Peru, Bolivia  M T S

Pyrrhopyge infantilis agala Evans, 1951  Bolivia  M T S

Pyrrhopyge thericles Mabille, 1891  Impostor Firetip  TL: Brazil (Am)

Pyrrhopyge thericles orientis E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge thericles thericles Mabille, 1891  NE Brazil  M T S

Pyrrhopyge thericles ponicia Evans, 1951  French Guiana, Surinam, N Brazil  M T S

Pyrrhopyge thericles pseudophidias E. Bell, 1931  Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru, N Brazil  M T S

Pyrrhopyge thericles raymondi Orellana, [2010]  Venezuela (Cordillera de la Costa, Carabobo, Aragua, Guarico, Miranda)

Pyrrhopyge thericles fola Evans, 1951  Colombia, SW Venezuela  M T

Pyrrhopyge thericles rileyi E. Bell, 1931  Ecuador, Bolivia  M T

Pyrrhopyge thericles grinda Evans, 1953  Guyana  M T

Pyrrhopyge thericles ronda Evans, 1953  Trinidad  M T

Pyrrhopyge amythaon E. Bell, 1931  Amythaon Firetip  TL: Brazil (Pe)

Pyrrhopyge amythaon gradens Evans, 1951  N Brazil (Para)  M T S

Pyrrhopyge amythaon peron E. Bell, 1947  Venezuela (Bolivar), Colombia, NE Peru  M T S

Pyrrhopyge amythaon polka Evans, 1951  French Guiana  M T S

Pyrrhopyge amythaon perula Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge amythaon orino Evans, 1951  Colombia, Venezuela (S of Orinoco), Guyana  M T S

Pyrrhopyge amythaon pollio Evans, 1951  Peru  M T

Pyrrhopyge amythaon podina Evans, 1951  Peru, Bolivia  M T

Pyrrhopyge amythaon amythaon E. Bell, 1931  Brazil (Pernambuco)  M T

Pyrrhopyge sergius Hopffer, 1874  Sergius Firetip  TL: Peru

Pyrrhopyge sergius selina Evans, 1951  Brazil (Para)  M T S

Pyrrhopyge sergius andros Evans, 1951  Colombia, Venezuela (Amazonas)  M T

Pyrrhopyge sergius andronicus E. Bell, 1931  Ecuador, Peru, Bolivia  M T S

Pyrrhopyge sergius semana Evans, 1951  Surinam, French Guiana  M T S

Pyrrhopyge sergius sergius Hopffer, 1874  Peru  M T S

Pyrrhopyge sergius ganus E. Bell, 1947  Guyana, Venezuela (Cordillera de la Costa), [Colombia, Ecuador, Peru, Brazil?]  M T

Pyrrhopyge sergius josephina Draudt, 1921  Peru, Bolivia  M T

Pyrrhopyge caribe Orellana, [2010]  Caribbean Firetip  TL: Venezuela

Pyrrhopyge caribe caribe Orellana, [2010]  Venezuela (Aragua, Carabobo, Dtto. Capital, Lara, Trujillo, Vargas)

Pyrrhopyge caribe camachoi Orellana, [2010]  Venezuela (Gran Sabana)

Pyrrhopyge erazoae Orellana, [2010]  Erazo's Firetip  Venezuela (Bolivar)  M S

Pyrrhopyge charybdis Westwood, 1852  Charybdis Firetip  TL: [Brazil (MG)]

Pyrrhopyge charybdis semita Evans, 1951  Bolivia, S Brazil  M T S

Pyrrhopyge charybdis charybdis Westwood, 1852  S Brazil  M T OI S L

Pyrrhopyge cressoni E. Bell, 1932  Cresson's Firetip  Ecuador, Bolivia  M T S

Pyrrhopyge pelota Plötz, 1879  Pelota Firetip  Bolivia, Paraguay, S Brazil  M T S

Pyrrhopyge amyclas (Cramer, 1779)  Yellow-edged Firetip  TL: Surinam

Pyrrhopyge amyclas amyclas (Cramer, 1779)  Trinidad, Venezuela, Guyana, Surinam, French Guiana, Brazil  M S

Pyrrhopyge amyclas denticulata Herrich-Schäffer, 1869  Ecuador, Peru, Brazil  M S L

Pyrrhopyge phylleia Hewitson, 1874  Orange-edged Firetip  TL: Bolivia

Pyrrhopyge phylleia delos Evans, 1951  Peru  M T S L

Pyrrhopyge phylleia phylleia Hewitson, 1874  S Peru, Bolivia  M T TI S L G

Pyrrhopyge arax Evans, 1951  Arax Firetip  SE Peru, Bolivia  M T S L

Pyrrhopyge haemon Godman & Salvin, 1893  Tawny-rimmed Firetip  Costa Rica  M T S

Pyrrhopyge punctata Röber, 1925  Songo Firetip  Bolivia (Rio Songo)

Pyrrhopyge papius Hopffer, 1874  Papius Firetip  TL: Colombia; Bolivia

Pyrrhopyge papius pasca Evans, 1951  Colombia  M T S

Pyrrhopyge papius papius Hopffer, 1874  Venezuela, Colombia, Ecuador, Peru (Andean)  M T S

Pyrrhopyge frona Evans, 1951  Inca Firetip  Peru  M T S

Pyrrhopyge melanomerus Mabille & Boullet, 1908  Melanomerus Firetip  TL: Bolivia

Pyrrhopyge melanomerus patma Evans, 1951  S Ecuador, NE Peru  M T

Pyrrhopyge melanomerus melanomerus Mabille & Boullet, 1908  Peru, Bolivia  M T L

Pyrrhopyge decipiens Mabille, 1903  Red-spotted Firetip  [Ecuador?], Peru, Bolivia  M T S

Pyrrhopyge placeta Evans, 1951  Placeta Firetip  Brazil (ES)  M T

Pyrrhopyge hadassa Hewitson, 1866  Hadassa Firetip  TL: [Ecuador]

Pyrrhopyge hadassa hadassa Hewitson, 1866  Colombia, Ecuador, Peru  M T TI S

Pyrrhopyge hadassa henna Evans, 1951  Peru  M T

Pyrrhopyge hadassa hanga Evans, 1951  Peru  M T

Pyrrhopyge hadassa pseudohadassa Mabille & Boullet, 1908  Peru  M T S L

Pyrrhopyge hadassa halma Evans, 1951  Peru, Bolivia  M T S L

Pyrrhopyge terra Evans, 1951  Terra Firetip  Bolivia  M T S

Pyrrhopyge telassina Staudinger, 1888  Telassina Firetip  TL: Peru; Bolivia

Pyrrhopyge telassina tagra Evans, 1951  Peru  M T S

Pyrrhopyge telassina telassina Staudinger, 1888  Peru  M T S

Pyrrhopyge telassina shiva Evans, 1951  SE Peru, Bolivia  M T S L

Pyrrhopyge telassa Hewitson, 1866  Telassa Firetip  TL: [Ecuador]

Pyrrhopyge telassa telassa Hewitson, 1866  Ecuador  M T TI S IL

Pyrrhopyge telassa phaeax Hopffer, 1874  Peru  M T S L

Pyrrhopyge telassa silex Evans, 1951  Peru  M T

Pyrrhopyge telassa croceimargo Mabille & Boullet, 1908  S Peru, Bolivia  M T S HS L

Pyrrhopyge martena Hewitson, [1869]  Martena Firetip  Ecuador  M T OI S

Pyrrhopyge schausi E. Bell, 1931  Schaus' Firetip  Colombia, Ecuador, N Peru  M T S

Pyrrhopyge sadia Evans, 1951  Sadia Firetip  Ecuador, Peru  M T S

Pyrrhopyge creona H. Druce, 1874  Creona Firetip  Peru, Brazil (Am)  M T S

Pyrrhopyge kelita Hewitson, [1869]  Streaked Firetip  SE Peru, Bolivia  M T OI L

Pyrrhopyge crista Evans, 1951  Crista Firetip  Bolivia  M T

Pyrrhopyge crida Hewitson, 1871  White-banded Firetip  SE Mexico to Colombia and Ecuador  M T TI S1 S2 IL I1 I2 G

Pyrrhopyge mopsus (E. Bell, 1931)  Mopsus Firetip  Peru  M T S

Pyrrhopyge sarpedon (E. Bell, 1931)  Sarpedon Firetip  Peru (Amazonas)  M T

Pyrrhopyge arinas (Cramer, 1777)  Arinas Firetip  TL: Surinam

Pyrrhopyge arinas arinas (Cramer, 1777)  French Guiana, Surinam, N Brazil, NE Peru  M T TI S IL

Pyrrhopyge arinas temenos (E. Bell, 1931)  Peru  M T S

Pyrrhopyge creusae (E. Bell, 1931)  Creusae Firetip  French Guiana  M T

Pyrrhopyge tatei (E. Bell, 1932)  Tate's Firetip  Venezuela  M T

Pyrrhopyge boulleti Le Cerf, 1922  Boullet's Firetip  Colombia, Venezuela (Tachira)  M S

Gunayan  O. Mielke, 2002

Gunayan rhacia (Hewitson, [1875])  Rhacia Skipper  S Brazil  M T OI

Gunayan rubricollis (Sepp, [1841])  Red-collared Skipper  French Guiana, Peru, Bolivia, Brazil  M T TI S IL L

Gunayan timaeus (E. Bell, 1931)  Timaeus Skipper  Colombia, Peru  M T S

Yanguna  E. Watson, 1893

Yanguna tetricus E. Bell, 1931  Tetricus Skipper  Guyana, Peru  M T

Yanguna cometes (Cramer, 1779)  Comet Skipper  TL: Surinam

Yanguna cometes cometes (Cramer, 1779)  French Guiana, N Brazil (Para)  M S

Yanguna cometes staudingeri (Plötz, 1879)  Surinam, Venezuela (Bolivar), S Colombia (upper Putumayo), Ecuador, N Peru  M S IL L

Yanguna cometes angeli Orellana, [2010]  Venezuela (Tachira)

Yanguna cometes cometides Mabille & Boullet, 1908  Peru, Bolivia  M S IL

Yanguna thelersa (Hewitson, 1866)  Thelersa Skipper  French Guiana, N Brazil, Ecuador, Peru  M T OI S

Yanguna cosyra (H. Druce, 1875)  Cosyra Skipper  Guatemala to Colombia and Ecuador  M T S1 S2 IL I1 I2 I3 G

Yanguna spatiosa (Hewitson, 1870)  Spatiosa Skipper  TL: Ecuador

Yanguna spatiosa spatiosa (Hewitson, 1870)  W Colombia, Ecuador  M T TI S IL L

Yanguna spatiosa mabillei H. Druce, 1909  Peru  M T S L

Yanguna spatiosa aspilos Mabille & Boullet, 1908  Bolivia  M T OI IL

Yanguna erebus (Plötz, 1879)  Erebus Skipper  Brazil (Ba)  M T

Chalypyge  O. Mielke, 2002

Chalypyge chalybea (Scudder, 1872)  Orange-rimmed Skipper  TL: Mexico

Chalypyge chalybea chalybea (Scudder, 1872)  C Mexico interior (central JAL, GTO, QRO)  M S L G

Chalypyge chalybea chloris (Evans, 1951)  W and S Mexico, Belize, Guatemala, Honduras  M T S IL L

Chalypyge zereda (Hewitson, 1866)  Zereda Skipper  TL: Ecuador

Chalypyge zereda hygieia (C. Felder & R. Felder, 1867)  Colombia, N Ecuador  M T S

Chalypyge zereda zereda (Hewitson, 1866)  Ecuador  M T

Chalypyge zereda rufipectus (Godman & Salvin, 1879)  Colombia, C Ecuador  M T S

Chalypyge zereda zepha (Evans, 1951)  S Ecuador  M T

Chalypyge zereda insana (Staudinger, 1876)  Peru  M T S IL

Chalypyge zereda rufinucha (Godman & Salvin, 1879)  Peru, Bolivia  M T S L

Ochropyge  O. Mielke, 2002

Ochropyge ruficauda (Hayward, 1932)  Red-tailed Skipper  S Brazil, Argentina  M T

Apyrrothrix  Lindsey, 1921

Apyrrothrix araxes (Hewitson, 1867)  Araxes Skipper  TL: Mexico

Apyrrothrix araxes arizonae (Godman & Salvin, 1893)  SE AZ, SW NM, W TX (local), SON, CHIN, NL, TAMPS, SLP  M T S L I G

Apyrrothrix araxes Sinaloa-Durango segregate ,  NW Mexico (SIN, DGO [probably NW NAY])  M S

Apyrrothrix araxes Nayarit segregate ,  W-central Mexico (SW NAY)  M S

Apyrrothrix araxes Guerrero-Michoacan segregate ,  SW Mexico (GRO, MICH)  M S

Apyrrothrix araxes araxes (Hewitson, 1867)  central Mexico (COL, JAL, DF, MOR, HGO, PUE, VER)  M T S L G

Apyrrothrix araxes Oaxaca segregate ,  S Mexico (OAX)  M S L

Melanopyge  O. Mielke, 2002

Melanopyge mulleri (E. Bell, 1934)  Red-spotted Skipper  SE Mexico (CHIS, OAX, VER)  M T S L G

Melanopyge hoffmanni (H. Freeman, 1977)  Hoffmann's Skipper  SE Mexico (OAX, TAB, CHIS), Belize  M T OD S L G

Melanopyge erythrosticta (Godman & Salvin, 1879)  Spotted Skipper  SE Mexico to Panama  M T OD S I G Txt

Melanopyge [n. sp.] Burns, MS  Dotted Skipper  SE Mexico to Costa Rica  M S1 S2 S3 HS L I1 I2

Melanopyge maculosa (Hewitson, 1866)  Maculosa Skipper  Panama, Colombia, Ecuador, Venezuela (Zulia)  M T S L

Melanopyge cossea (H. Druce, 1875)  Cossea Skipper  Colombia  M T S

Jonaspyge  O. Mielke, 2002

Jonaspyge jonas (C. Felder & R. Felder, 1859)  Scallop-edged Firetip  SE Mexico (HGO, OAX, PUE, VER, CHIS), Guatemala, Nicaragua  M T OD S L G

Jonaspyge tzotzili (H. Freeman, 1969)  Freeman's Firetip  SE Mexico (CHIS, OAX)  M T OD S

Jonaspyge aesculapus (Staudinger, 1876)  Fire-edged Skipper  Costa Rica, Panama, Colombia, Ecuador  M T OD S1 S2 IL I1 I2 G

Creonpyge  O. Mielke, 2002

Creonpyge creon (H. Druce, 1874)  Creon Skipper  TL: Panama

Creonpyge creon creon (H. Druce, 1874)  N Costa Rica to Cocle Prov., Panama, Atlantic & Pacific slopes  M T S1 S2 L I1 I2 G

Creonpyge creon lilliana (Nicolay & Small, 1969)  Cerro Campana, far E Cocle, far W Panama Prov., Panama  M T S

Creonpyge creon taylori (Nicolay & Small, 1981)  Fila Cruces, Puntarenas, SE Costa Rica  M T S

Cyanopyge  O. Mielke, 2002

Cyanopyge sangaris (Skinner, 1921)  Sangaris Skipper  Colombia  M T S

Microceris  E. Watson, 1893

Microceris variicolor (Ménétriés, 1855)  Zigzag Firetip  C & S-central Brazil  M S IL G

Elbella  Evans, 1951

Elbella intersecta (Herrich-Schäffer, 1869)  Variable Firetip  TL: ?

Elbella intersecta intersecta (Herrich-Schäffer, 1869)  French Guiana, Surinam, S Venezuela, Colombia, Peru, Ecuador, Bolivia, Brazil  M T S HS IL

Elbella intersecta peter Evans, 1951  NE Brazil  M T S IL

Elbella intersecta paraensis O. Mielke, 1995  NE Brazil  M T OI

Elbella intersecta rufitegula O. Mielke, 1995  SE Brazil (coastal)  M T OI

Elbella intersecta strova Evans, 1951  Panama, C Venezuela  M T S IL

Elbella intersecta losca Evans, 1951  SE Brazil (interior)  M T IL

Elbella viriditas (Skinner, 1920)  Gold-rimmed Firetip  TL: Honduras

Elbella viriditas viriditas (Skinner, 1920)  Paraguay  M T IL

Elbella viriditas chia Evans, 1951  N Brazil  M T S IL

Elbella viriditas boliviana O. Mielke, 1995  Bolivia  M T OI

Elbella lamprus (Hopffer, 1874)  Lamprus Firetip  TL: Brazil (RJ)

Elbella lamprus lamprus (Hopffer, 1874)  SE Brazil, S Paraguay  M T S IL

Elbella lamprus albociliata O. Mielke, 1995  SE Brazil  M T OI S L

Elbella mariae (E. Bell, 1931)  Maria's Firetip  TL: Brazil (RS)

Elbella mariae mariae (E. Bell, 1931)  SE Brazil  M T IL

Elbella mariae molinai (Hayward, 1941)  SE Brazil  M S IL

Elbella adonis (E. Bell, 1931)  Adonis Firetip  SE Brazil, S Paraguay  M T Ab IL

Elbella scylla (Ménétriés, 1855)  Scylla Firetip  E & W Mexico to Costa Rica  M S IL L I1 I2 G

Elbella dulcinea (Plötz, 1879)  Dulcinea Firetip  Costa Rica, Panama, Colombia, Venezuela, Ecuador  M T S IL

Elbella luteizona (Mabille, 1877)  Zona Firetip  SE Brazil  M T OD TI S HS IL

Elbella hegesippe (Mabille & Boullet, 1908)  Hegesippe Firetip  SE Brazil  M T OI S IL

Elbella iphinous (Latreille, [1824])  Yellow-barred Skipper  SE Brazil  M T OI Ab IL

Elbella miodesmiata (Röber, 1925)  Rober's Skipper  SE Mexico (OAX), Panama, Colombia  M OI S

Elbella rondonia O. Mielke, 1995  Rondonian Skipper  W Brazil, Peru  M T OI S

Elbella madeira O. Mielke, 1995  Madeira Skipper  W Brazil, Peru  M T OI S

Elbella bicuspis de Jong, 1983  Bicuspis Skipper  Surinam  M T IL

Elbella etna Evans, 1951  Etna Skipper  Trinidad, Colombia, Peru, Bolivia, Brazil  M T S IL

Elbella patroclus (Plötz, 1879)  Patroculus Skipper  TL: Peru

Elbella patroclus patroclus (Plötz, 1879)  Panama, Ecuador, Peru, Bolivia  M T TI S HS

Elbella patroclus acala Evans, 1951  SW Venezuela (Tachira), Colombia  M T S IL

Elbella azeta (Hewitson, 1866)  Azeta Skipper  TL: Brazil (Am)

Elbella azeta azeta (Hewitson, 1866)  Venezuela (Bolivar), Colombia, Peru, Bolivia, Brazil  M T IL

Elbella azeta giffordi O. Mielke, 1995  S Brazil  M T OI

Elbella lustra Evans, 1951  Lustra Skipper  SW Venezuela (Tachira), Colombia, N Ecuador  M T IL

Elbella merops (E. Bell, 1934)  Merops Skipper  French Guiana, Venezuela (Tachira), Colombia, Peru, Bolivia, Brazil  M T S IL

Elbella theseus (E. Bell, 1934)  Theseus Skipper  Colombia, Venezuela (Tachira), Ecuador, Peru, Bolivia, N Brazil  M T S IL

Elbella blanda Evans, 1951  Blanda Skipper  French Guiana, Colombia, Peru, Bolivia, Paraguay, Brazil  M T TI S IL

Elbella patrobas (Hewitson, 1857)  Patrobas Skipper  TL: Colombia

Elbella patrobas mexicana O. Mielke, 1995  SE Mexico (QROO, TAB, CHIS, VER, OAX), Belize  M T OI S

Elbella patrobas patrobas (Hewitson, 1857)  Guatemala to Colombia  M T OI S IL I1 I2 G

Elbella patrobas evansi O. Mielke, 1995  Colombia, Venezuela (Tachira)  M T OI

Elbella patrobas melanina (Mabille & Boullet, 1908)  Peru, Bolivia  M T OI

Elbella patrobas tingo O. Mielke, 1995  Peru, W Brazil  M T OI L

Elbella patrobas amazonica O. Mielke, 1995  N Brazil (Amazonas)  M T OI

Parelbella  O. Mielke, 1995

Parelbella polyzona (Latreille, [1824])  Polyzona Skipper  SE Brazil  M T IL

Parelbella ahira (Hewitson, 1866)  Ahira Skipper  TL: Brazil (Pa)

Parelbella ahira ahira (Hewitson, 1866)  French Guiana, Colombia, Peru, Bolivia, Paraguay, N & W Brazil  M T OI TI S IL L

Parelbella ahira extrema (Röber, 1925)  SE Brazil, Paraguay  M T OI S IL

Parelbella peruana O. Mielke, 1995  Peruana Skipper  Peru  M T OI S

Parelbella macleannani (Godman & Salvin, 1893)  Macleannan’s Skipper  SE Mexico (CHIS, VER) to NW Ecuador  M T S1 S2 IL I1 I2 G

Parelbella nigra O. Mielke, Austin & A. Warren, 2008  Mielke’s Skipper  SE Mexico (OAX, VER)  M T

Croniades  Mabille, 1903

Croniades pieria (Hewitson, 1857)  Fire-banded Skipper  TL: Brazil (Am)

Croniades pieria pieria (Hewitson, 1857)  French Guiana, N Brazil  M T OI S IL

Croniades pieria auraria H. Druce, 1908  Peru, Bolivia  M T S

Pseudocroniades  O. Mielke, 1995

Pseudocroniades machaon (Westwood, 1852)  Yellow-banded Skipper  TL: Brazil

Pseudocroniades machaon machaon (Westwood, 1852)  SE Brazil  M T OI S IL

Pseudocroniades machaon seabrai O. Mielke, 1995  SE Brazil (RJ)  M T OI

Protelbella  O. Mielke, 1995

Protelbella alburna (Mabille, 1891)  Alburna Skipper  TL: Peru

Protelbella alburna alburna (Mabille, 1891)  Ecuador, Peru, Bolivia  M T S IL

Protelbella alburna brasiliensis O. Mielke, 1995  French Guiana, N & W Brazil  M OI

Nosphistia  Mabille & Boullet, 1908

Nosphistia zonara (Hewitson, 1866)  Zonara Skipper  Colombia, Ecuador, N Brazil (Amazonas)  M T OD TI S IL

Jemadia  E. Watson, 1893

Jemadia pseudognetus (Mabille, 1878)  Dot-collared Skipper  SE Mexico to Venezuela and upper Amazon  M T OD S1 S2 L I1 I2 G Txt

Jemadia hospita (A. Butler, 1877)  Hospita Skipper  TL: Peru

Jemadia hospita imitator (Mabille, 1891)  Colombia  M T S

Jemadia hospita hospita (A. Butler, 1877)  S Colombia (upper Putumayo), Ecuador, Peru, Bolivia, W Brazil  M T S IL

Jemadia hospita hephaestos (Plötz, 1879)  Surinam  M T S HS L

Jemadia demarmelsi Orellana, [2010]  Demarmels' Skipper  Venezuela (Bolivar)  M OI

Jemadia sosia (Mabille, 1878)  Sosia Skipper  SW Venezuela (Tachira), Colombia, Ecuador, Peru, Bolivia  M T OD S Txt

Jemadia menechmus (Mabille, 1878)  Menechmus Skipper  

Jemadia menechmus menechmus (Mabille, 1878)  Surinam, Venezuela, Colombia, Ecuador, Peru, Bolivia, N & W Brazil  M T OD S

Jemadia menechmus desousai Orellana, [2010]  Venezuela (Amazonas)  M OI

Jemadia scomber H. Druce, 1908  Mammoth Skipper  Peru  M T S L

Jemadia suekentonmiller Grishin, 2014  Speciesex Skipper  Costa Rica & Panama  M T1 T2 T3 Txt

Jemadia ovid Evans, 1951  Ovid Skipper  Colombia, Ecuador  M T S G

Jemadia pater Evans, 1951  Pater Skipper  Panama, Colombia, Venezuela  M T S1 S2 HS G Txt

Jemadia ortizi Orellana, [2010]  Ortiz's Skipper  Venezuela (SE Tachira), Colombia  M OI S

Jemadia albescens Röber, 1925  Ecuador, Peru, Bolivia  M OD S1 S2 Ma L1 L2 G Txt

Jemadia hewitsonii (Mabille, 1878)  Hewitson's Skipper  TL: Brazil ([Am])

Jemadia hewitsonii hewitsonii (Mabille, 1878)  French Guiana, Venezuela (Amazonas, Bolivar), N Brazil, Peru  M T OI OD TI S1 S2 S3 S4 HS L G

Jemadia gnetus (Fabricius, 1781)  Gnetus Skipper  Surinam, Guyana, Venezuela, Coloimbia, Ecuador, Peru, Bolivia, S Brazil  M T OI OD S1 S2 HS L

Jemadia brevipennis Schaus, 1902  Brevipennis Skipper  Brazil (DF, SP, Pr), Paraguay  M T OD Txt

Jemadia fallax (Mabille, 1878)  Fallax Skipper  TL: Brazil

Jemadia fallax fiska Evans, 1951  Colombia: Cauca & Cundinamarca  M T

Jemadia fallax fida Evans, 1951  Ecuador, Peru, Bolivia  M T S1 S2 L

Jemadia fallax solaris Hayward, 1942  Colombia, Venezuela, Guyana, N Brazil, E Peru, Bolivia  M T S1 S2

Jemadia fallax fallax (Mabille, 1878)  Guianas, N Brazil  M T OD S L Txt

Mimoniades  Hübner, 1823

Mimoniades nurscia (Swainson, 1821)  Nursica Skipper  TL: S. Amer.

Mimoniades nurscia malis (Godman & Salvin, 1879)  Colombia  M T S IL

Mimoniades nurscia nurscia (Swainson, 1821)  Colombia, Ecuador, N Peru  M S

Mimoniades nurscia amans Skinner, 1920  Colombia, Peru  M T S IL

Mimoniades ocyalus Hübner, 1823  Ocyalus Skipper  S Brazil  M S L

Mimoniades versicolor (Latreille, [1824])  Versicolor Skipper  TL: Brazil

Mimoniades versicolor eupheme (Godman & Salvin, 1879)  Ecuador, Peru, Bolivia  M T S L

Mimoniades versicolor versicolor (Latreille, [1824])  Brazil  M T S

Mimoniades montana J. Zikán, 1938  Quadricolor Skipper  SE Brazil  M S

Mimoniades baroni (Godman & Salvin, 1895)  Baron's Skipper  Peru  M T S Txt

Mimardaris  O. Mielke, 2002

Mimardaris aerata (Godman & Salvin, 1879)  Aerata Skipper  Colombia  M T S

Mimardaris sela (Hewitson, 1866)  Sela Skipper  TL: Colombia

Mimardaris sela sela (Hewitson, 1866)  Colombia  M T OI S

Mimardaris sela aequatorea (Röber, 1925)  Colombia, Ecuador, N Peru  M T S

Mimardaris sela peruviana (Draudt, 1921)  Peru  M T S G

Mimardaris sela chanchamayonis (Draudt, 1924)  Peru, Bolivia  M T S

Mimardaris sela periphema (Hewitson, [1875])  Peru, Bolivia  M T OI S L

Mimardaris lomax (Evans, 1951)  Lomax Skipper  Peru  M T S

Mimardaris montra (Evans, 1951)  Montra Skipper  Peru  M T S

Mimardaris pityusa (Hewitson, 1857)  Pityusa Skipper  Ecuador, Peru  M T OI S IL L

Mimardaris porus (Plötz, 1879)  Porus Skipper  TL: Colombia

Mimardaris porus porus (Plötz, 1879)  Colombia  M T OI S HS IL

Mimardaris porus mortis (Evans, 1951)  Peru  M T S

Mimardaris minthe (Godman & Salvin, 1879)  Minthe Skipper  Colombia, Ecuador, N Peru  M T S IL

Ardaris  E. Watson, 1893

Ardaris eximia (Hewitson, 1871)  Crystal-banded Skipper  Venezuela (Cordillera de Merida from Tachira to Trujillo & Lara)  M T TI S IL

Ardaris hantra Evans, 1951  Amber-banded Skipper  Venezuela (Cordillera de Merida in Merida)  M T

Amenis  E. Watson, 1893

Amenis pionia (Hewitson, 1857)  Pionia Skipper  TL: Venezuela

Amenis pionia picia Evans, 1951  Colombia  M T S1 S2

Amenis pionia sandra Orellana, [2010]  Colombia, Venezuela (E slope Andes in Tachira, Barinas)  M S

Amenis pionia pionia (Hewitson, 1857)  Colombia, Venezuela (Cordillera de la Costa), disjunct in Bolivia, N Argentina  M T OI S L

Amenis rogeri Orellana, [2010]  Roger's Skipper  Venezuela (Merida, Lara, at least)  M S

Amenis ponina (Herrich-Schäffer, 1869)  Venezuela, Surinam  M S

Sarbia  E. Watson, 1893

Sarbia antias (C. Felder & R. Felder, 1859)  Antias Firetip  S Brazil  M T S IL

Sarbia catomelaena Mabille & Boullet, 1908  Cato Firetip  S Brazil  M T OI

Sarbia oneka (Hewitson, 1866)  Oneka Firetip  [S Brazil]  M T IL

Sarbia soza Evans, 1951  Soza Firetip  S Brazil (MG, RJ, Pr, SC)  M T IL G

Sarbia curitiba O. Mielke & Casagrande, 2002  Curitiba Firetip  S Brazil  M OI S

Sarbia pertyi (Plötz, 1879)  Perty's Firetip  S Brazil  M IL

Sarbia damippe Mabille & Boullet, 1908  Damippe Firetip  S Brazil  M T OI S IL

Sarbia xanthippe (Latreille, [1824])  Xanthippe Firetip  TL: Brazil

Sarbia xanthippe xanthippe (Latreille, [1824])  S Brazil  M T OI IL

Sarbia xanthippe spixii (Plötz, 1879)  S Brazil  M T S HS IL

Metardaris  Mabille, 1903

Metardaris cosinga (Hewitson, 1874)  Cosinga Firetip  TL: Bolivia

Metardaris cosinga catana Evans, 1951  Peru, Bolivia  M T S

Metardaris cosinga circe Evans, 1951  Bolivia  M T S

Metardaris cosinga cedra Evans, 1951  Bolivia  M T S G

Metardaris cosinga cosinga (Hewitson, 1874)  Peru, Bolivia  M T TI S HS IL

Amysoria  O. Mielke, 2002

Amysoria galgala (Hewitson, 1866)  Red-banded Firetip  S Mexico (SW CHIS), Panama, Colombia and Venezuela  M T S L G

Mysoria  E. Watson, 1893

Mysoria ambigua (Mabille & Boullet, 1908)  Royal Firetip  SE Mexico (CHIS, TAB, VER) to Costa Rica  M T S1 S2 L I Txt

Mysoria barcastus (Sepp, [1851])  Barcastus Firetip  TL: Surinam

Mysoria barcastus venezuelae (Scudder, 1872)  Panama, Colombia, Venezuela, Tobago, [Guianas]  M T S HS L G

Mysoria barcastus alta Evans, 1951  Trinidad  M T

Mysoria barcastus barcastus (Sepp, [1851])  Guyana, Durinam, French Guiana  M T OI OD I

Mysoria barcastus pallens (Mabille, 1891)  S Venezuela (Amazonas, Bolivar), Colombia  M T

Mysoria barcastus antila Evans, 1951  N Brazil  M T

Mysoria barcastus barta Evans, 1951  Bolivia, Paraguay, S Brazil, Argentina  M T S L

Mysoria amra (Hewitson, 1871)  Blue-collared Firetip  W Mexico, Guatemala  M T OI S L G

Mysoria affinis (Herrich-Schäffer, 1869)  Red-collared Firetip  [= wilsoni], W Mexico, Guatemala  M T S L G

Mysarbia  O. Mielke, 2002

Mysarbia sejanus (Hopffer, 1874)  TL: [Peru]

Mysarbia sejanus stolli O. Mielke & Casagrande, 2002  Costa Rica, Colombia, Venezuela, French Guiana, Brazil, NE Peru  M T S L

Mysarbia sejanus sejanus (Hopffer, 1874)  Ecuador, Peru  M T

Mysarbia sejanus erythrostigma (Röber, 1925)  Bolivia  M T OD S

  Tribe Celaenorrhinini

Celaenorrhinus  Hübner, [1819]

Celaenorrhinus aegiochus (Hewitson, 1876)  Flashing Flat  Costa Rica and Panama  M T S L G

Celaenorrhinus savia (Evans, 1952)  TL: Peru  M T S IL

Celaenorrhinus orneates Austin, 1996  TL: Brazil (Rd)  M T OI S

Celaenorrhinus monartus (Plötz, 1884)  Dotted Flat  S Mexico to N Brazil  M T S IL L G

Celaenorrhinus astrigera (A. Butler, 1877)  TL: Brazil (Pa)  M T S

Celaenorrhinus fritzgaertneri (Bailey, 1880)  Fritzgaertner’s Flat  SE AZ & S TX (as strays) to Costa Rica  M T S1 S2 IL L I G

Celaenorrhinus stallingsi H. Freeman, 1946  Stallings’ Flat  S Texas (rare stray), E Mexico to Costa Rica  M T S L I Ha

Celaenorrhinus shema (Hewitson, 1877)  TL: French Guiana

Celaenorrhinus shema shema (Hewitson, 1877)  TL: French Guiana  M T S IL

Celaenorrhinus shema ochra Evans, 1952  TL: Brazil (Am)  M T S

Celaenorrhinus shema vox Evans, 1952  TL: Peru  M T S

Celaenorrhinus shema mercedensis O. Mielke, 1992  TL: Peru  M T OI

Celaenorrhinus par Steinhauser & Austin, 1996  TL: Brazil (Rd)  M T OI

Celaenorrhinus disjunctus E. Bell, 1940  TL: Peru  M T S

Celaenorrhinus vagra Evans, 1952  TL: Peru  M T

Celaenorrhinus stola Evans, 1952  Stola Flat  E & W Mexico to Nicaragua  M T S L

Celaenorrhinus approximatus R. Williams & E. Bell, 1940  Approximate Flat  Costa Rica and Panama  M T S L I G

Celaenorrhinus bifurcus E. Bell, 1934  Bifurcate Flat  Costa Rica to Guianas, Trinidad, Ecuador  M T S L

Celaenorrhinus similis Hayward, 1933  TL: Argentina  M S IL G

Celaenorrhinus eligius (Stoll, 1781)  Eligius Flat  TL: Surinam

Celaenorrhinus eligius eligius (Stoll, 1781)  Costa Rica to Guianas, Trinidad, Tobago, to Peru  M T S IL I G

Celaenorrhinus eligius punctiger (Burmeister, 1878)  TL: Argentina; Brazil (RJ)  M T S IL G

Celaenorrhinus songoensis Draudt, 1922  TL: Bolivia

Celaenorrhinus songoensis saroma Evans, 1952  TL: Colombia  M T S

Celaenorrhinus songoensis tonio Evans, 1952  TL: Colombia  M T

Celaenorrhinus songoensis songoensis Draudt, 1922  TL: Bolivia  M T OI S L G

Celaenorrhinus cynapes (Hewitson, 1870)  Small Flat  TL: Ecuador

Celaenorrhinus cynapes darius Evans, 1952  TL: Ecuador  M T S

Celaenorrhinus cynapes cynapes (Hewitson, 1870)  SE Mexico to Ecuador  M T G

Celaenorrhinus tritonae (Weeks, 1901)  TL: Bolivia  M TI S

Celaenorrhinus anchialus (Mabille, 1878)  TL: Brazil  M T S IL

Celaenorrhinus jao (Mabille, 1889)  TL: Peru  M T OI S

Celaenorrhinus autochton Steinhauser & Austin, 1996  TL: Brazil (Rd)  M T S

Celaenorrhinus suthina (Hewitson, 1877)  TL: Ecuador  M T OI S L

Celaenorrhinus syllius (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S1 S2 S3 L

  Tribe Carcharodini

Mimia  Evans, 1953

Mimia phidyle (Godman & Salvin, 1894)  Phidyle Skipper  TL: Panama

Mimia phidyle phidyle (Godman & Salvin, 1894)  CHIS to Panama  M T S L

Mimia phidyle pazana Evans, 1953  TL: Bolivia  M T S G

Mimia chiapaensis H. Freeman, 1969  Chiapan Skipper  CHIS  M T S I G

Arteurotia  A. Butler & H. Druce, 1872

Arteurotia tractipennis A. Butler & H. Druce, 1872  Starred Skipper  TL: Costa Rica

Arteurotia tractipennis tractipennis A. Butler & H. Druce, 1872  S TX to Surinam and Bolivia  M T S1 S2 IL L I G

Arteurotia tractipennis contractipennis Mabille & Boullet, 1916  TL: Venezuela  M T

Mictris  Evans, 1955

Mictris crispus (Herrich-Schäffer, 1870)  TL: Venezuela

Mictris crispus caerula (Mabille, 1877)  CHIS to Ecuador  M T S1 S2 L I1 I2 G

Mictris crispus crispus (Herrich-Schäffer, 1870)  TL: Venezuela  M T S

Iliana  E. Bell, 1937

Iliana heros (Mabille & Boullet, 1917)  TL: French Guiana; Peru

Iliana heros heroica Evans, 1953  TL: Colombia  M T OD G

Iliana heros heros (Mabille & Boullet, 1917)  TL: French Guiana; Peru  M T OD S L G

Iliana purpurascens (Mabille & Boullet, 1912)  TL: Brazil  M OD S Co G Txt

Iliana remus E. Bell, 1937  TL: Peru  M T OD S G

Iliana romulus E. Bell, 1937  TL: Peru  M T OD S G

Sophista  Plötz, 1879

Sophista aristoteles (Westwood, 1852)  TL: Brazil (Am)

Sophista aristoteles aristoteles (Westwood, 1852)  TL: Brazil (Am)  M T S IL L

Sophista aristoteles plinius Plötz, 1882  TL: [Brazil]  M T S

Sophista latifasciata (Spitz, 1930)  TL: Brazil (SP)

Sophista latifasciata matto Evans, 1953  TL: Brazil (MT)  M S

Sophista latifasciata latifasciata (Spitz, 1930)  TL: Brazil (SP)  M S

Polyctor  Evans, 1953

Polyctor cleta Evans, 1953  Cleta Tufted-Skipper  E & W Mexico to Costa Rica  M T S1 S2 L I G

Polyctor polyctor (Prittwitz, 1868)  Polyctor Tufted-Skipper  TL: Brazil (RJ)

Polyctor polyctor polyctor (Prittwitz, 1868)  Costa Rica to Argentina  M T S1 S2 IL L I G

Polyctor polyctor dagua Evans, 1953  TL: Colombia  M T

Polyctor enops (Godman & Salvin, 1894)  Enops Tufted-Skipper  E & S Mexico to Costa Rica  M T S1 S2 L I

Polyctor fera (Weeks, 1901)  TL: Bolivia

Polyctor fera fera (Weeks, 1901)  TL: Bolivia  M T S

Polyctor fera falla Evans, 1953  TL: Venezuela  M T

Polyctor extensa (Mabille, 1891)  TL: Peru  M T S L

Polyctor tensa Evans, 1953  TL: Brazil  M T S

Nisoniades  Hübner, [1819]

Nisoniades godma Evans, 1953  Godma Tufted-Skipper  E & S Mexico to Costa Rica  M T S1 S2 L I G

Nisoniades panama Evans, 1953  Panamanian Tufted-Skipper  Panama to Ecuador  M T S

Nisoniades benda Evans, 1953  TL: Ecuador  M T S

Nisoniades hecale (Hayward, 1940)  TL: Ecuador  M S

Nisoniades cauca Evans, 1953  TL: Colombia  M T

Nisoniades remo Evans, 1953  TL: Peru  M T

Nisoniades bessus (Möschler, 1877)  TL: Surinam  M T S

Nisoniades montana (R. Williams & E. Bell, 1939)  TL: Venezuela  M T

Nisoniades coca Steinhauser, 1989  TL: Ecuador  M T

Nisoniades lata Steinhauser, 1989  TL: Ecuador  M T S

Nisoniades criton (Mabille, 1898)  TL: Bolivia  M T

Nisoniades hesperia (Hayward, 1939)  TL: Paraguay

Nisoniades maura (Mabille & Boullet, 1917)  TL: Brazil  M T S

Nisoniades brazia Evans, 1953  TL: Brazil (RJ)  M T S

Nisoniades laurentina (R. Williams & E. Bell, 1939)  Dark Tufted-Skipper  SE Mexico (VER) to Guianas and Trinidad  M T S

Nisoniades suprapanama Steinhauser, 1989  TL: Colombia  M T

Nisoniades hora (Hayward, 1939)  TL: Ecuador  M T

Nisoniades rubescens (Möschler, 1877)  Purplish Tufted-Skipper  W Mexico & S Texas (as stray) to lower Amazon and Bolivia  M T S1 S2 L I H G

Nisoniades mimas (Cramer, 1775)  TL: Surinam  M T OI S

Nisoniades bipuncta (Schaus, 1902)  TL: Brazil (RJ)  M T S

Nisoniades ephora (Herrich-Schäffer, 1870)  Ephora Tufted-Skipper  E & W Mexico to Guianas, Trinidad, Ecuador and Peru  M T S L I G

Nisoniades rimana (E. Bell, 1942)  TL: Venezuela  M T S

Nisoniades evansi Steinhauser, 1989  TL: Ecuador  M T S

Nisoniades supra Steinhauser, 1989  TL: Peru  M T

Nisoniades indistincta (R. Williams & E. Bell, 1939)  TL: Colombia  M T

Nisoniades brunneata (R. Williams & E. Bell, 1939)  TL: French Guiana  M T

Nisoniades borra (E. Bell, 1947)  TL: Colombia  M T

Nisoniades torta Steinhauser, 1989  Torta Tufted-Skipper  Costa Rica to Panama  M T S G

Nisoniades macarius (Herrich-Schäffer, 1870)  Macarius Tufted-Skipper  (doubtful for Mexico, Honduras), Panama to Argentina  M T TI S G

Nisoniades guianae (R. Williams & E. Bell, 1939)  TL: French Guiana  M T

Nisoniades nyctineme (A. Butler, 1877)  TL: Brazil (Pa)  M T

Nisoniades castolus (Hewitson, 1878)  Castolus Tufted-Skipper  SE Mexico (VER) to S Brazil  M T S1 S2 I G

Nisoniades haywardi (R. Williams & E. Bell, 1939)  TL: Argentina  M T S

Viola  Evans, 1953

Viola alicus (Schaus, 1902)  TL: Brazil ([Pr])  M T S

Viola minor (Hayward, 1933)  TL: Argentina  M S

Viola olla Evans, 1953  TL: Brazil (Am)  M T S

Viola violella (Mabille, 1898)  TL: Bolivia  M T S L

Viola egra Evans, 1953  TL: Colombia  M T S

Viola dagamba Steinhauser, 1989  TL: Guyana  M T

Pachyneuria  Mabille, 1888

Pachyneuria obscura Mabille, 1888  TL: Peru  M T OI TI S

Pachyneuria jaguar Evans, 1953  TL: Peru  M T S

Pachyneuria lineatopunctata (Mabille & Boullet, 1917)  TL: Peru

Pachyneuria lineatopunctata belema Evans, 1953  TL: Brazil (Pa)  M T

Pachyneuria lineatopunctata lineatopunctata (Mabille & Boullet, 1917)  TL: Peru  M T S

Pachyneuria duidae (E. Bell, 1932)  TL: Venezuela

Pachyneuria duidae duidae (E. Bell, 1932)  TL: Venezuela  M T S

Pachyneuria duidae pozuza Evans, 1953  TL: Peru  M T

Pachyneuria inops (Mabille, 1877)  TL: [Brazil]  M T TI S

Pachyneuria damon (E. Bell, 1937)  TL: Brazil (Am)  M T S

Pachyneuria licisca (Plötz, 1882)  Immaculate Tufted-Skipper  E & W Mexico to Colombia and Venezuela  M T S1 S2 L I G

Pachyneuria lista Evans, 1953  TL: French Guiana  M T S

Pachyneuria herophile (Hayward, 1940)  TL: Ecuador  M S

Pachyneuria helena (Hayward, 1939)  TL: Ecuador  M T S

Pachyneuria milleri Steinhauser, 1989  TL: Peru  M T

Pellicia  Herrich-Schäffer, 1870

Pellicia klugi R. Williams & E. Bell, 1939  TL: Peru  M T S

Pellicia santana R. Williams & E. Bell, 1939  TL: Peru  M T

Pellicia simulator R. Williams & E. Bell, 1939  TL: Peru  M T

Pellicia arina Evans, 1953  Glazed Tufted-Skipper  S TX to Panama  M T S1 S2 L I G

Pellicia costimacula Herrich-Schäffer, 1870  Frosted Tufted-Skipper  TL: [Brazil]

Pellicia costimacula costimacula Herrich-Schäffer, 1870  Panama to S Brazil and Paraguay  M T S G

Pellicia costimacula litoralis Biezanko & O. Mielke, 1973  TL: Brazil (RS)  M T S

Pellicia tyana Plötz, 1882  TL: S. Amer.

Pellicia tyana toza Evans, 1953  TL: Colombia  M T

Pellicia tyana tyana Plötz, 1882  TL: S. Amer.  M T S

Pellicia vecina Schaus, 1902  TL: Brazil (RJ)

Pellicia vecina naja Steinhauser, 1989  TL: Peru  M T

Pellicia vecina vecina Schaus, 1902  TL: Brazil (RJ)  M T S

Pellicia najoides Hayward, 1933  TL: Argentina  M S

Pellicia angra Evans, 1953  Rare Tufted-Skipper  TL: Mexico (Ver)

Pellicia angra angra Evans, 1953  SE Mexico (VER) to Panama  M T S G

Pellicia angra axina Evans, 1953  TL: Ecuador  M T

Pellicia tonga Evans, 1953  Tonga Tufted-Skipper  Panama, Colombia, Venezuela  M T S

Pellicia trax Evans, 1953  TL: Ecuador  M T S

Pellicia theon Plötz, 1882  TL: S. Amer.  M T TI S

Pellicia ranta Evans, 1953  TL: Brazil (RJ)

Pellicia ranta ranta Evans, 1953  TL: Brazil (RJ)  M T

Pellicia ranta rancida Evans, 1953  TL: Paraguay  M T S

Pellicia dimidiata Herrich-Schäffer, 1870  Morning Glory Tufted-Skipper  TL: Mexico; Venezuela

Pellicia dimidiata dimidiata Herrich-Schäffer, 1870  W Mexico and S TX (as stray) to Guianas, Trinidad, up. Amazon  M T OI S1 S2 L I G

Pellicia dimidiata meno (Mabille, 1889)  TL: Panama  M T OI S

Pellicia dimidiata zamia Plötz, 1882  TL: S. Amer.  M T TI S

Pellicia hersilia Hayward, 1939  TL: Argentina  M T S

Pellicia demetrius (Plötz, 1882)  TL: Brazil  M T TI

Pellicia alta Steinhauser, 2007  TL: Peru

Burca  E. Bell & W. Comstock, 1948

Burca braco (Herrich-Schäffer, 1865)  Braco Skipper  TL: Cuba

Burca braco braco (Herrich-Schäffer, 1865)  Cuba  M T S L

Burca braco castigata Rindge, 1955  Bahamas; Great Bahama Bank  M T

Burca stillmanni E. Bell & W. Comstock, 1948  Stillman's Skipper  Hispaniola  M T S

Burca hispaniolae E. Bell & W. Comstock, 1948  Hispaniolan Skipper  Hispaniola  M T S

Burca concolor (Herrich-Schäffer, 1865)  Concolorous Skipper  TL: Cuba

Burca concolor concolor (Herrich-Schäffer, 1865)  Cuba  M T S

Burca concolor atrata Rindge, 1955  Bahamas; Great Bahama Bank  M T S

Burca cubensis (Skinner, 1913)  Cuban Skipper  Cuba  M T L

Fuscocimex  Austin, 2008

Fuscocimex caecus Austin, 2008  Blind Skipper  Brazil (Rondonia)  M T S

Ocella  Evans, 1953

Ocella albata (Mabille, 1888)  TL: Peru  M T OI S

Ocella diophthalma (Plötz, 1884)  TL: ?  M S

Ocella monophthalma (Plötz, 1884)  TL: Brazil  M T S

Windia  H. Freeman, 1969

Windia windi H. Freeman, 1969  Wind’s Skipper  SE Arizona (one-time stray), NW Mexico to Costa Rica  M T S L G

Noctuana  E. Bell, 1937

Noctuana lactifera (A. Butler & H. Druce, 1872)  Cryptic Skipper  TL: Costa Rica

Noctuana lactifera lactifera (A. Butler & H. Druce, 1872)  Costa Rica to Colombia  M T S L I G

Noctuana lactifera bipuncta (Plötz, 1884)  E & W Mexico to Nicaragua  M S L

Noctuana noctua (C. Felder & R. Felder, 1867)  Noctua Skipper  Panama to Ecuador and Peru  M T S

Noctuana brunneofusca (Mabille & Boullet, 1917)  TL: Peru  M T

Noctuana diurna (A. Butler, 1870)  TL: ?  M T S

Noctuana stator (Godman, 1899)  Red-studded Skipper  W Mexico & S TX (one-time stray) to Guianas, Trinidad, Peru  M T S L I G

Noctuana haematospila (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia  M T S

Jera  Lindsey, 1925

Jera tricuspidata (Mabille, 1902)  TL: Ecuador  M T TI S IL

Conognathus  C. Felder & R. Felder, 1862

Conognathus platon C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T OI S L

Morvina  Evans, 1953

Morvina morvus (Plötz, 1884)  TL: Brazil

Morvina morvus cyclopa Evans, 1953  TL: Bolivia  M T S

Morvina morvus para Evans, 1953  TL: Brazil (Pa)  M T S

Morvina morvus morvus (Plötz, 1884)  TL: Brazil  M S

Morvina fissimacula (Mabille, 1878)  TL: Brazil

Morvina fissimacula pelarge (Godman & Salvin, 1894)  Nicaragua to Panama  M T S I G

Morvina fissimacula lenia Evans, 1953  TL: Colombia  M T

Morvina fissimacula rema Evans, 1953  TL: Brazil (Pa)  M T S

Morvina fissimacula fissimacula (Mabille, 1878)  TL: Brazil  M T S

Morvina falisca (Hewitson, 1878)  TL: French Guiana

Morvina falisca falisca (Hewitson, 1878)  TL: French Guiana  M T S

Morvina falisca falia Evans, 1953  TL: Peru  M T S

Myrinia  Evans, 1953

Myrinia binoculus (Möschler, 1877)  TL: Surinam  M T S

Myrinia raymundo H. Freeman, 1979  Raymundo's Skipper  SE Mexico (TAB, CHIS)  M T S G

Myrinia myris (Mabille, 1898)  Myris Skipper  Campeche, Belize, Nicaragua to Guianas, Bolivia, S Brazil  M T S

Myrinia santa Evans, 1953  TL: Brazil (ES)

Myrinia santa monka Evans, 1953  TL: Brazil (Pa)  M T S

Myrinia santa santa Evans, 1953  TL: Brazil (ES)  M T

Myrinia laddeyi (E. Bell, 1942)  TL: Ecuador  M T S

Myrinia catua O. Mielke, 1968  TL: Brazil (MG)  M T S

Xispia  Lindsey, 1925

Xispia quadrata (Mabille, 1889)  TL: Brazil (Am)  M T OI S G

Xispia satyrus (Jörgensen, 1935)  TL: Paraguay  M S IL

Cyclosemia  Mabille, 1878

Cyclosemia lathaea (Hewitson, 1878)  TL: Bolivia  M T S

Cyclosemia leppa Evans, 1953  Leppa Eyed-Skipper  Guatemala to Bolivia  M T S IL

Cyclosemia anastomosis Mabille, 1878  Northern Eyed-Skipper  E Mexico to Ecuador  M T S1 S2 L I G

Cyclosemia earina (Hewitson, 1878)  TL: Brazil (Pa)  M T S

Cyclosemia lyrcaea (Hewitson, 1878)  Lyrcaea Eyed-Skipper  Panama to SE Brazil  M T S

Cyclosemia pedro R. Williams & E. Bell, 1940  Pedro Eyed-Skipper  Honduras to N Brazil ("Amazons")  M T G

Cyclosemia caerulea Austin & DeVries, 2001  Caerulea Eyed-Skipper  Costa Rica  M T S

Cyclosemia subcaerulea Schaus, 1913  Subcaerulea Eyed-Skipper  Costa Rica to Ecuador  M T S1 S2 I

Cyclosemia herennius (Stoll, 1782)  TL: Surinam  M S

Cyclosemia elelea (Hewitson, 1878)  TL: French Guiana  M T S

Gorgopas  Godman & Salvin, 1894

Gorgopas trochilus (Hopffer, 1874)  TL: Bolivia  M T S

Gorgopas petale (Mabille, 1888)  TL: Brazil (SP)  M T OI S

Gorgopas chlorocephala (Herrich-Schäffer, 1870)  Green-headed Sootywing  TL: S. Amer.

Gorgopas chlorocephala chlorocephala (Herrich-Schäffer, 1870)  Nicaragua to Bolivia  M T S L G

Gorgopas chlorocephala sneiderni E. Bell, 1947  TL: Colombia  M T

Gorgopas gutta Evans, 1953  TL: Peru  M T S L

Gorgopas agylla (Mabille, 1898)  TL: Bolivia  M T S L

Bolla  Mabille, 1903

Bolla cylindus (Godman & Salvin, 1896)  Checkered Sootywing  SE Mexico (VER) to Costa Rica  M T S G

Bolla cybele Evans, 1953  Veracruzan Sootywing  SE Mexico (VER)  M T IL

Bolla sonda Evans, 1953  TL: Mexico (Ver)  M T

Bolla cyclops (Mabille, 1877)  Cyclops Sootywing  [= sonda], SE Mexico, Guatemala, Colombia  M T S

Bolla mancoi (Lindsey, 1925)  TL: Peru  M S

Bolla atahuallpai (Lindsey, 1925)  Atahuallpai Sootywing  Panama to Argentina  M OI S IL G

Bolla catharina (E. Bell, 1937)  TL: Brazil (SC)  M T S

Bolla cupreiceps (Mabille, 1891)  Copper-headed Sootywing  E & S Mexico to Bolivia and S Brazil  M T S IL L G

Bolla morona (E. Bell, 1940)  TL: Peru

Bolla morona lorea (E. Bell, 1947)  TL: Colombia  M T

Bolla morona morona (E. Bell, 1940)  TL: Peru  M T S

Bolla nigerrima Mabille & Boullet, 1917  TL: Peru  M T

Bolla subapicatus (Schaus, 1902)  Fin-tipped Sootywing  W & S Mexico, Guatemala  M T S L

Bolla orsines (Godman & Salvin, 1896)  Hook-tipped Sootywing  E & W Mexico to El Salvador  M T S G

Bolla evippe (Godman & Salvin, 1896)  Rough-tipped Sootywing  E & W Mexico, Guatemala, Nicaragua  M T S G

Bolla imbras (Godman & Salvin, 1896)  Rounded Sootywing  [= pullata], E & S Mexico to Colombia and Venezuela  M T S G

Bolla phylo Mabille, 1903  TL: Brazil  M T

Bolla antha Evans, 1953  TL: Colombia  M T

Bolla brennus (Godman & Salvin, 1896)  Obscure Sootywing  TL: Panama

Bolla brennus brennus (Godman & Salvin, 1896)  S Texas (one-time stray), E Mexico to Panama  M T S G

Bolla brennus vexta Evans, 1953  TL: Colombia  M T

Bolla oriza Evans, 1953  Orizaba Sootywing  SE Mexico (VER, CHIS?)  M T S

Bolla guerra Evans, 1953  Guerrero Sootywing  SW Mexico to Nicaragua  M T S L

Bolla boliviensis (E. Bell, 1937)  Bolivian Sootywing  Guatemala to Bolivia  M T S IL G

Bolla dorsolaciniae Steinhauser, 1989  TL: Colombia  M T

Bolla giselus (Mabille, 1883)  TL: Colombia

Bolla giselus tornea Evans, 1953  TL: Colombia  M T

Bolla giselus giselus (Mabille, 1883)  TL: Colombia  M T S

Bolla saletas (Godman & Salvin, 1896)  Saletas Sootywing  Guatemala to El Salvador  M T S G

Bolla eusebius (Plötz, 1884)  Spatulate Sootywing  E & W Mexico to Costa Rica  M T S

Bolla fenestra Steinhauser, 1991  Oaxacan Sootywing  S Mexico (OAX)  M T S L Ha G

Bolla solitaria Steinhauser, 1991  Solitary Sootywing  S Mexico (OAX)  M T S G

Bolla zora Evans, 1953  TL: Ecuador  M S

Bolla zorilla (Plötz, 1886)  Zorilla Sootywing  SE Mexico (VER, OAX) to Bolivia  M T S G

Bolla clytius (Godman & Salvin, 1897)  Mottled Sootywing  SE AZ & S TX (rare strays) to Honduras  M T S L G

Bolla litus (Dyar, 1912)  Many-spotted Sootywing  mostly W Mexico  M T S L

Staphylus  Godman & Salvin, 1896

Staphylus tridentis Steinhauser, 1989  TL: Colombia  M T

Staphylus esmeraldus L. Miller, 1966  Esmeraldus Sootywing  E Costa Rica  M T G

Staphylus chlorocephala (Latreille, [1824])  TL: Brazil (RJ)  M T S

Staphylus chlora Evans, 1953  TL: Bolivia  M T S

Staphylus eryx Evans, 1953  TL: Brazil (Pa)  M T

Staphylus ceos (W. H. Edwards, 1882)  Golden-headed Sootywing  far S CA, S NV (rare), S AZ, S NM, S TX, to QRO  M T S HS L1 L2 I1 I2 G

Staphylus putumayo (E. Bell, 1937)  TL: Peru  M T S

Staphylus sambo Evans, 1953  TL: Brazil (Pa)  M T

Staphylus vulgata (Möschler, 1879)  Golden-snouted Sootywing  E & W Mexico to Argentina  M T S1 S2 IL L I G

Staphylus melaina (Hayward, 1947)  TL: Argentina; Paraguay  M S IL G

Staphylus buena (R. Williams & E. Bell, 1940)  TL: Bolivia  M T S

Staphylus melius Steinhauser, 1989  TL: Argentina  M T

Staphylus incanus E. Bell, 1932  TL: Brazil (SP)  M T S

Staphylus musculus (Burmeister, 1875)  TL: Argentina  M T S IL L G

Staphylus imperspicua (Hayward, 1940)  TL: Ecuador  M S

Staphylus caribbea (R. Williams & E. Bell, 1940)  Panamanian Sootywing  Nicaragua to Ecuador  M T S1 S2 I G

Staphylus lizeri (Hayward, 1938)  TL: Bolivia

Staphylus lizeri album Steinhauser, 1989  TL: Brazil (MG)  M T

Staphylus lizeri lizeri (Hayward, 1938)  TL: Bolivia  M T S

Staphylus corumba (R. Williams & E. Bell, 1940)  TL: Brazil (MS)  M T S

Staphylus ascalon (Staudinger, 1876)  TL: Brazil (RJ)  M T TI S IL

Staphylus melangon (Mabille, 1883)  TL: S. Amer.

Staphylus melangon stanga Evans, 1953  TL: Ecuador  M T

Staphylus melangon epicaste Mabille, 1903  TL: Brazil  M S IL G

Staphylus melangon melangon (Mabille, 1883)  TL: S. Amer.  M T S

Staphylus punctiseparatus Hayward, 1933  TL: Argentina  M IL

Staphylus menuda (Weeks, 1902)  TL: Bolivia

Staphylus shola Evans, 1953  TL: ?  M T

Staphylus tepeca (E. Bell, 1942)  Grizzled Sootywing  C & S Mexico (mostly DF to OAX)  M T S L

Staphylus perna Evans, 1953  TL: Brazil (Pe)  M T S

Staphylus hayhurstii (W. H. Edwards, 1870)  Hayhurst's Sootywing  S Ontario, E USA to far SE SD, NB, KS, to S-C TX  M T OD S HS L1 L2 I1 I2 H G

Staphylus mazans (Reakirt, [1867])  Mazans Sootywing  C TX to CHIS  M T OD S L1 L2 G Txt

Staphylus tierra Evans, 1953  West-Mexican Sootywing  W Mexico (SON to CHIS), El Salvador  M T S L

Staphylus ascalaphus (Staudinger, 1876)  Central American Sootywing  SE Mexico to Bolivia, S Brazil  M T OD S1 S2 L G

Staphylus perforata (Möschler, 1879)  Perforata Sootywing  Panama, Colombia, Peru  M T S G

Staphylus unicornis Steinhauser & Austin, 1993  Unicorn Sootywing  Costa Rica  M T1 T2

Staphylus lenis Steinhauser, 1989  Lenis Sootywing  Yucatan Peninsula to Trinidad  M T S G

Staphylus azteca (Scudder, 1872)  Aztec Sootywing  E & W Mexico to Costa Rica  M T S1 S2 L I G

Staphylus tyro (Mabille, 1878)  TL: Brazil; Venezuela  M T

Staphylus tucumanus (Plötz, 1884)  TL: Argentina  M S

Staphylus cartagoa (R. Williams & E. Bell, 1940)  Cartagoa Sootywing  Costa Rica, El Salvador  M T G

Staphylus huigra (R. Williams & E. Bell, 1940)  Huigra Sootywing  TL: Ecuador

Staphylus huigra huigra (R. Williams & E. Bell, 1940)  El Salvador, Ecuador to Bolivia  M T S G

Staphylus huigra canka Evans, 1953  TL: Peru  M T

Staphylus mossi Evans, 1953  TL: Peru  M T S

Staphylus cordillerae (Lindsey, 1925)  TL: Peru  M S

Staphylus coecatus (Mabille, 1891)  TL: Brazil (RJ)  M T S

Staphylus balsa (E. Bell, 1937)  TL: Peru  M T

Staphylus vincula Plötz, 1886  Mountain Sootywing  E & W Mexico to Venezuela  M T TI S L G

Staphylus oeta (Plötz, 1884)  Oeta Sootywing  Costa Rica to Bolivia and Argentina  M T TI S G

Staphylus tingo Steinhauser, 1989  TL: Peru  M T

Staphylus parvus Steinhauser & Austin, 1993  Parvus Sootywing  Costa Rica  M T

Staphylus saxos Evans, 1953  TL: Colombia

Staphylus saxos saxos Evans, 1953  TL: Colombia  M T

Staphylus saxos satrap Evans, 1953  TL: Bolivia  M T S

Staphylus iguala (R. Williams & E. Bell, 1940)  Iguala Sootywing  W & S Mexico  M T S L G

Staphylus astra (R. Williams & E. Bell, 1940)  TL: Peru  M T S

Staphylus minor Schaus, 1902  TL: Peru

Staphylus minor manx Evans, 1953  TL: Colombia  M T

Staphylus minor minor Schaus, 1902  TL: Peru  M T S IL G

Staphylus incisus (Mabille, 1878)  TL: French Guiana; Brazil (Pa)  M T S IL L G

Staphylus fasciatus Hayward, 1933  TL: Argentina  M T S

Staphylus evemerus Godman & Salvin, 1896  Evemerus Sootywing  Nicaragua to Panama  M T S I G

Staphylus insignis O. Mielke, 1980  TL: Brazil (RS)  M OI S G

Staphylus kayei Cock, 1996  TL: Trinidad  M T

Staphylus veytius H. Freeman, 1969  TL: Mexico (Chia)  M T

Pholisora  Scudder, 1872

Pholisora catullus (Fabricius, 1793)  Common Sootywing  widespread in USA and S Canada, S to Colima and Hidalgo...S to Oaxaca  M S HS L1 L2 I G

Pholisora mejicanus (Reakirt, [1867])  Mexican Sootywing  S-C Colorado to Oaxaca  M T S L G

Hesperopsis  Dyar, 1905

Hesperopsis alpheus (W. H. Edwards, 1876)  Saltbush Sootywing  TL: USA (NM)

Hesperopsis alpheus oricus (W. H. Edwards, 1879)  E & S CA, NV, SW UT, NW AZ  M T S L Ha

Hesperopsis alpheus alpheus (W. H. Edwards, 1876)  [= texana], S AZ, NM, CO, far S MT, TX (verification needed for NE Mexico)  M S1 S2 L1 L2 I1 I2 G

Hesperopsis gracielae (MacNeill, 1970)  MacNeill's Sootywing  lower Colorado River drainage, Wa Co., UT to SON + Baja  M T S L I H

Hesperopsis libya (Scudder, 1878)  Mojave Sootywing  TL: USA (Az)

Hesperopsis libya joaquina J. Emmel, T. Emmel & Mattoon, 1998  S San Joaquin Valley, CA  M T S

Hesperopsis libya libya (Scudder, 1878)  Mojave Desert, CA- S NV-AZ  M S L1 L2 H Ha G V

Hesperopsis libya Great Basin segregate ,  SE OR, SW ID, most of NV, E CA  M S L H Ha

Hesperopsis libya confertiblanca Scott, 1992  [perhaps a synonym of lena], W CO, far NW NM, UT?, N AZ?  M T S I

Hesperopsis libya lena (W. H. Edwards, 1882)  W ND, E MT, WY, UT?  M S

  Tribe Erynnini

Gorgythion  Godman & Salvin, 1896

Gorgythion begga (Prittwitz, 1868)  TL: Brazil (RJ)

Gorgythion begga pyralina (Möschler, 1877)  S TX to Guianas and Bolivia  M T OI OD S1 S2 L I1 I2 G

Gorgythion begga begga (Prittwitz, 1868)  TL: Brazil (RJ)  M T OD S IL L G

Gorgythion plautia (Möschler, 1877)  TL: Surinam  M T OI OD S L G

Gorgythion vox Evans, 1953  Crab’s-claw Skipper  E Mexico to Costa Rica  M T OD S L G

Gorgythion beggina Mabille, 1898  Beggina Skipper  Costa Rica to Argentina

Gorgythion beggina escalophoides (Hayward, 1941)  Costa Rica to Argentina  M T OD TI S IL G Txt

Gorgythion beggina beggina Mabille, 1898  TL: Bolivia  M T OD S G

Gorgythion canda Evans, 1953  TL: Paraguay  M T OD S G

Gorgythion alcandra (Dognin, 1891)  TL: Brazil  M T OD G Txt

Sostrata  Godman & Salvin, 1895

Sostrata cronion (C. Felder & R. Felder, 1867)  TL: Brazil  M T S IL L

Sostrata festiva (Erichson, [1849])  TL: Guyana  M T S IL L

Sostrata caerulans (Mabille & Boullet, 1917)  TL: Bolivia; Peru  M T

Sostrata adamantinus (Mabille, 1898)  TL: Bolivia  M T S IL

Sostrata grippa Evans, 1953  Grippa Skipper  Costa Rica to Ecuador and Bolivia  M T S L

Sostrata jinna Evans, 1953  TL: Colombia  M T OD

Sostrata nordica Evans, 1953  Blue-studded Skipper  S TX to Costa Rica  M T S1 S2 L I

Sostrata bifasciata (Ménétriés, 1829)  TL: Brazil (MG)

Sostrata bifasciata leucorrhoa Godman & Salvin, 1895  Costa Rica to Colombia  M T S L

Sostrata bifasciata adamas (Plötz, 1884)  TL: Brazil  M S

Sostrata bifasciata bifasciata (Ménétriés, 1829)  TL: Brazil (MG)  M T S

Sostrata pusilla Godman & Salvin, 1895  Pusilla Skipper  TL: Nicaragua; Panama; Brazil (Am/Pa)

Sostrata pusilla pusilla Godman & Salvin, 1895  Nicaragua to upper Amazon  M T S I

Sostrata pusilla pulsa Evans, 1953  TL: Ecuador  M T OD

Potamanaxas  Lindsey, 1925

Potamanaxas thoria (Hewitson, 1870)  Thoria Skipper  Nicaragua to Peru  M T OI OD S1 S2 S3 L I G

Potamanaxas okroogly Grishin, 2013  Ovalpatch Skipper  E Colombia, Ecuador, Peru & Bolivia  M T S HS IL L G

Potamanaxas tschotky Grishin, 2013  Crisppatch Skipper  Colombia & Ecuador  M T S L G

Potamanaxas louisghilli Grishin, 2015  Hill's Skipper  Costa Rica & Panama  M T S

Potamanaxas melicertes (Godman & Salvin, 1895)  Melicertes Skipper  Costa Rica to Ecuador  M T OI OD S IL

Potamanaxas paphos Evans, 1953  Paphos Skipper  Costa Rica to Ecuador  M T OD S1 S2 S3 G

Potamanaxas hirta (Weeks, 1901)  Hirta Skipper  Colombia to Bolivia  M T OI OD TI S1 S2 S3 S4 L G Txt

Potamanaxas effusa (Draudt, 1922)  Effusa Skipper  TL: Colombia

Potamanaxas effusa effusa (Draudt, 1922)  Colombia to Guianas and Peru  M OI OD S1 S2 G

Potamanaxas effusa confusa (Draudt, 1922)  Panama, Colombia  M OI OD S

Potamanaxas xantholeuce (Mabille, 1888)  Xantholeuce Skipper  Panama  M T OI TI S

Potamanaxas flavofasciata (Hewitson, 1870)  Yellowbanded Skipper  TL: Ecuador

Potamanaxas flavofasciata flavofasciata (Hewitson, 1870)  Ecuador, Peru  M T S1 S2 G

Potamanaxas flavofasciata pantra Evans, 1953  Bolivia  M T OD S IL G

Potamanaxas cranda Evans, 1953  Cranda Skipper  Costa Rica & Panama  M T OD S1 S2 G

Potamanaxas hermieri Grishin, 2013  Hermier's Skipper  Ecuador  M T

Potamanaxas paralus (Godman & Salvin, 1895)  Paralus Skipper  Colombia to Bolivia  M T OD S1 S2 IL L Txt

Potamanaxas latrea (Hewitson, 1875)  Latrea Skipper  TL: Nicaragua

Potamanaxas latrea latrea (Hewitson, 1875)  Nicaragua to Colombia  M T OI OD TI S

Potamanaxas latrea caliadne (Godman & Salvin, 1895)  Costa Rica  M T OI OD S IL G

Potamanaxas latrea tusca Evans, 1953  Ecuador  M T OD S IL L G

Potamanaxas latrea tyndarus Evans, 1953  Peru  M T OD S G

Potamanaxas thestia (Hewitson, 1870)  Thestia Skipper  Ecuador, Peru, Bolivia  M T S IL L G

Potamanaxas zagadka Grishin, 2013  Phylomystery Skipper  Ecuador  M T

Potamanaxas trex Evans, 1953  Trex Skipper  Colombia, Ecuador  M T OD S G

Potamanaxas tunga E. Bell, 1956  Tunga Skipper  Ecuador  M T OI OD G

Potamanaxas quira E. Bell, 1956  Quira Skipper  Colombia  M T OI OD S G

Potamanaxas trigga Evans, 1953  Trigga Skipper  Colombia, Ecuador, Peru  M T OD S G

Potamanaxas laoma (Hewitson, 1870)  Laoma Skipper  TL: Ecuador

Potamanaxas laoma fumida (Draudt, 1922)  Colombia  M OI OD S

Potamanaxas laoma laoma (Hewitson, 1870)  Ecuador  M T S1 S2 G

Potamanaxas laoma violacea (Dognin, 1888)  Ecuador, Peru  M T OI OD S1 S2 S3 S4 S5 IL G

Potamanaxas laoma cosna Evans, 1953  Peru, Bolivia  M T OD S1 S2 G

Potamanaxas bana E. Bell, 1956  Bana skipper  Ecuador  M T OI OD S L G

Potamanaxas serp Grishin, 2013  Sickle-clasp Skipper  Colombia  M T

Potamanaxas frenda Evans, 1953  Frenda Skipper  Peru  M T OD S IL G

Potamanaxas andraemon (Mabille, 1898)  Andraemon Skipper  Colombia (West Andes)  M T OD S L G

Potamanaxas perornatus Hayward, 1940  Perornatus Skipper  Colombia (East Andes), Ecuador, N Peru  M T OD S G

Potamanaxas lamasi Grishin, 2013  Lamas' Skipper  N & central Peru  M T OD S G

Potamanaxas fuma Evans, 1953  Fuma Skipper  central Peru  M T OD G

Potamanaxas forum Evans, 1953  Forum Skipper  S Peru, Bolivia  M T OD S G

Potamanaxas [n. sp.] Grishin, MS  Potamanaxas louisghilli complex  Colombia to Peru  M S

Mylon  Godman & Salvin, 1894

Mylon lassia (Hewitson, 1868)  Bold Mylon  E & W Mexico to Guianas and Trinidad  M T S1 S2 IL L I G

Mylon zephus (A. Butler, 1870)  Zephus Mylon  TL: Venezuela; Colombia

Mylon zephus zephus (A. Butler, 1870)  Colombia & Venezuela, hypothetical for Panama  M T S

Mylon zephus albodiscus Mabille & Boullet, 1917  Peru, Bolivia  M T L

Mylon illineatus Mabille & Boullet, 1917  Illineatus Mylon  TL: Bolivia

Mylon illineatus illineatus Mabille & Boullet, 1917  Ecuador, Peru, Bolivia  M T S IL Co Ma L G

Mylon illineatus toxina Evans, 1953  Upper Amazonian region of Colombia, Ecuador, NE Peru and NW Brazil  M T OD Co L G

Mylon orsa Evans, 1953  Orsa Mylon  Costa Rica and Panama  M T S IL G

Mylon mestor Evans, 1953  Mestor Mylon  Ecuador  M T S IL G

Mylon salvia Evans, 1953  Evans’ Mylon  Mexico, Guatemala to Ecuador  M T S1 S2 IL I G

Mylon ander Evans, 1953  Narrow-winged Mylon  TL: Brazil (RJ)

Mylon ander andrea Evans, 1953  SE Mexico to Colombia  M T OD S L

Mylon ander ander Evans, 1953  Colombia to Bolivia and S Brazil  M T OD S IL L G

Mylon maimon (Fabricius, 1775)  Common Mylon  [= menippus], E & W Mexico to Argentina  M T S1 S2 S3 IL L1 L2 L3 I G Txt

Mylon cajus (Plötz, 1884)  TL: Peru

Mylon cajus hera Evans, 1953  (SE Mexico?), Guatemala to Panama  M T S IL L G

Mylon cajus cajus (Plötz, 1884)  Colombia to S Brazil  M T S L

Mylon pelopidas (Fabricius, 1793)  Pale Mylon  SE AZ (one-time stray), E & W Mexico to Paraguay  M T S1 S2 IL Ma L1 L2 I G

Mylon cristata Austin, 2000  Austin's Mylon  Campeche and Chiapas to Guianas and Brazil  M OI S IL G

Mylon jason (Ehrmann, 1907)  Jason’s Mylon  (SE Mexico?), Guatemala to S Brazil and Paraguay  M TI S L G

Mylon exstincta Mabille & Boullet, 1917  Forgotten Mylon  NW Brazil  M IL L G

Mylon simplex Austin, 2000  Unadorned Mylon  NW Brazil  M T OI L G

Mylon argonautarum Austin, 2000  Argonaut Mylon  NW Brazil  M T OI S G

Grais  Godman & Salvin, 1894

Grais stigmaticus (Mabille, 1883)  Hermit Skipper  TL: [Panama]

Grais stigmaticus stigmaticus (Mabille, 1883)  C & S TX (as stray), E & W Mexico to Argentina  M T S1 S2 L I G

Grais stigmaticus juncta Evans, 1953  Jamaica  M T S

Timochares  Godman & Salvin, 1896

Timochares ruptifasciata (Plötz, 1884)  Brown-banded Skipper  SE AZ / SW NM, S TX to Chiapas and Belize  M T S L1 L2 I1 I2 G

Timochares runia Evans, 1953  Jamaican-banded Skipper  Jamaica  M T S

Timochares trifasciata (Hewitson, 1868)  Many-banded Skipper  TL: ?

Timochares trifasciata trifasciata (Hewitson, 1868)  E & W Mexico to S Brazil and Paraguay  M T S1 S2 HS L G

Timochares trifasciata sanda Evans, 1953  TL: Argentina  M T S G

Anastrus  Hübner, [1824]

Anastrus sempiternus (A. Butler & H. Druce, 1872)  Common Anastrus  TL: Costa Rica

Anastrus sempiternus sempiternus (A. Butler & H. Druce, 1872)  S TX (one-time stray), E & W Mexico to Venezuela and Ecuador  M T S L I1 I2 G

Anastrus sempiternus simplicior (Möschler, 1877)  TL: Surinam; "Cuba"  M T S HS L G

Anastrus sempiternus dilloni (E. Bell & W. Comstock, 1948)  Jamaica, Hispaniola, Cuba  M T S L

Anastrus luctuosus (Godman & Salvin, 1894)  West-Mexican Anastrus  [= Mexican "robigus"], W Mexico  M T S G

Anastrus tolimus (Plötz, 1884)  Blurry Anastrus  TL: Colombia

Anastrus tolimus tolimus (Plötz, 1884)  E Mexico to Ecuador  M S L G

Anastrus tolimus robigus (Plötz, 1884)  TL: S. Amer.  M TI S G

Anastrus petius (Möschler, 1877)  TL: Surinam

Anastrus petius peto Evans, 1953  Guatemala to Costa Rica  M T S G

Anastrus petius petius (Möschler, 1877)  TL: Surinam  M T S L G

Anastrus isidro Grishin, 2012  Isidro's Skipper  Costa Rica and Panama  M T G

Anastrus obliqua (Plötz, 1884)  TL: ?  M T S IL G

Anastrus meliboea (Godman & Salvin, 1894)  Scarce Anastrus  TL: Panama

Anastrus meliboea meliboea (Godman & Salvin, 1894)  (SE Mexico?), Costa Rica to Colombia  M T S L G

Anastrus meliboea bactra Evans, 1953  TL: Bolivia  M T S L

Anastrus neaeris (Möschler, 1879)  Brilliant Anastrus  TL: Colombia

Anastrus neaeris neaeris (Möschler, 1879)  S Mexico to Venezuela and Ecuador  M T S1 S2 L I G

Anastrus neaeris narva Evans, 1953  Panama to Bolivia and E-C Brazil  M T S IL L G

Anastrus chaqua Evans, 1953  TL: Peru  M T S

Anastrus obscurus Hübner, [1824]  TL: ?  M OI S

Anastrus virens Austin, 1999  TL: Brazil (Rd)

Anastrus virens albopannus Austin, 2002  SE Mexico (OAX) to Panama  M T S L G

Anastrus virens virens Austin, 1999  TL: Brazil (Rd)  M T G

Anastrus ulpianus Poey, 1832  TL: not specified  M T S

Anastrus peruvianus (Mabille, 1883)  TL: [Peru]  M T S G

Speculum  Austin, 2008

Speculum speculum Austin, 2008  Hidden Mirror Skipper  French Guiana to Brazil  M OI OD S G

Tosta  Evans, 1953

Tosta tosta Evans, 1953  TL: Peru  M T OD S G

Tosta gorgus (E. Bell, 1937)  Gorgus Skipper  Belize, Costa Rica  M T OD S I G

Tosta platypterus (Mabille, 1895)  Platypterus Skipper  SE Mexico to Panama  M T S I G

Tosta niger (R. Williams & E. Bell, 1940)  Colombian Skipper  Costa Rica to Colombia  M T OD S I1 I2 G

Tosta taurus Evans, 1953  TL: Peru  M T OD S

Tosta sapasoa Nicolay, 1973  TL: Peru  M T OD S G

Tosta capra Austin, 1999  TL: Brazil (Rd)  M T OD G

Tosta saltarana Steinhauser, 2007  TL: Peru

Ebrietas  Godman & Salvin, 1896

Ebrietas osyris (Staudinger, 1876)  Yellow-patched Bent-Skipper  SE Mexico to Venezuela and Ecuador  M T S1 S2 IL L I G

Ebrietas infanda (A. Butler, 1877)  TL: Brazil (Am)  M T S L G

Ebrietas badia (Plötz, 1884)  Badia Bent-Skipper  Costa Rica to Bolivia  M S G

Ebrietas anacreon (Staudinger, 1876)  Common Bent-Skipper  TL: Panama; Colombia; Brazil (RJ)

Ebrietas anacreon anacreon (Staudinger, 1876)  E & W Mexico to Argentina  M T S1 S2 HS L I G

Ebrietas anacreon triseriata (Plötz, 1884)  TL: Venezuela

Ebrietas anacreon leucophaea Mabille & Boullet, 1917  TL: Colombia; Ecuador; Brazil  M T

Ebrietas sappho Steinhauser, 1974  Sappho Bent-Skipper  E Mexico to Costa Rica  M T S G

Ebrietas evanidus Mabille, 1898  Blurred Bent-Skipper  SE Mexico to Peru and upper Amazon  M T S L G

Ebrietas elaudia (Plötz, 1884)  Elaudia Bent-Skipper  TL: Venezuela; Panama

Ebrietas elaudia livius Mabille, 1898  SE Mexico (TAB, CHIS) to Bolivia  M T S IL

Ebrietas elaudia elaudia (Plötz, 1884)  Panama, Colombia, Venezuela  M S

Helias  Fabricius, 1807

Helias cama Evans, 1953  Squared Bent-Skipper  E Mexico to Colombia  M T S1 S2 L I G

Helias godmani (Mabille & Boullet, 1917)  Godman's Bent-Skipper  Costa Rica to Ecuador  M T S

Helias phalaenoides Fabricius, 1807  Fabricius' Bent-Skipper  TL: ?

Helias phalaenoides phalaenoides Fabricius, 1807  Panama to Bolivia  M S L

Helias phalaenoides palpalis (Latreille, [1824])  TL: Brazil  M T S IL G

Camptopleura  Mabille, 1877

Camptopleura theramenes Mabille, 1877  Mabille's Bent-Skipper  SE Mexico to Bolivia and Brazil  M T S IL G

Camptopleura auxo (Möschler, 1879)  Auxo Bent-Skipper  SE Mexico to S Brazil & Paraguay  M T S1 S2 IL L I1 I2 G

Camptopleura oaxaca H. Freeman, 1969  Oaxacan Bent-Skipper  SE Mexico (Oaxaca)  M T

Camptopleura impressus (Mabille, 1889)  Impressive Bent-Skipper  Panama to Minas Gerais  M T OI S L G

Camptopleura termon (Hopffer, 1874)  TL: Peru  M T S G

Camptopleura janthinus (Capronnier, 1874)  TL: Brazil (RJ)  M T S G

Cycloglypha  Mabille, 1903

Cycloglypha thrasibulus (Fabricius, 1793)  Widespread Bent-Skipper  TL: Indiis

Cycloglypha thrasibulus thrasibulus (Fabricius, 1793)  E & W Mexico to S Brazil & Paraguay  M T S1 S2 IL L I G

Cycloglypha thrasibulus flinta Evans, 1953  TL: Ecuador  M T

Cycloglypha tisias (Godman & Salvin, 1896)  Tisias Bent-Skipper  SE Mexico (OAX) to S Brazil  M T OD S G

Cycloglypha caeruleonigra Mabille, 1903  TL: [Ecuador]  M T S G

Cycloglypha stellita J. Zikán, 1938  TL: Brazil (MG)  M S

Cycloglypha enega (Möschler, 1877)  Enega Bent-Skipper  Nicaragua to Bolivia and S Brazil  M T S L

Cycloglypha polax Evans, 1953  TL: Brazil (MT)  M T S G

Theagenes  Godman & Salvin, 1896

Theagenes aegides (Herrich-Schäffer, 1869)  White-centered Bent-Skipper  E & W Mexico to Panama  M S L G

Theagenes albiplaga (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia  M T S IL G

Theagenes dichrous (Mabille, 1878)  TL: Brazil  M T S IL G

Chiomara  Godman & Salvin, 1899

Chiomara georgina (Reakirt, 1868)  White-patched Skipper  TL: Mexico (Ver)

Chiomara georgina pelagica (Weeks, 1891)  Baja California  M T OI S L

Chiomara georgina georgina (Reakirt, 1868)  SE AZ & S TX, E & W Mexico to Costa Rica  M T S1 S2 L1 L2 I G

Chiomara asychis (Stoll, 1780)  TL: Surinam

Chiomara asychis simon Evans, 1953  Costa Rica to Venezuela, Trinidad  M T S1 S2 G

Chiomara asychis vincenta Evans, 1953  St. Vincent  M T S G

Chiomara asychis grenada Evans, 1953  St. Lucia, Grenadines, Grenada  M T S G

Chiomara asychis asychis (Stoll, 1780)  TL: Surinam  M T TI S IL G

Chiomara asychis zania Evans, 1953  TL: Peru  M T S G

Chiomara asychis autander (Mabille, 1891)  TL: Argentina  M T S IL L G

Chiomara khalili N. Riley, 1934  TL: Brazil (Rd)  M T S G

Chiomara basigutta (Plötz, 1884)  TL: S. Amer.  M T S G

Chiomara mithrax (Möschler, 1879)  Mithrax Duskywing  S AZ (rare stray) W & E Mexico to Argentina, Cuba  M T S1 S2 IL L I G

Chiomara crenda Evans, 1953  TL: Brazil (Pr)  M T S HS G

Ephyriades  Hübner, [1819]

Ephyriades arcas (Drury, 1773)  Caribbean Duskywing  TL: St. Christopher's

Ephyriades arcas arcas (Drury, 1773)  northern Leeward Islands  M S

Ephyriades arcas philemon (Fabricius, 1775)  Greater Antilles  M T S IL L

Ephyriades zephodes (Hübner, [1825])  Zephodes Duskywing  Cuba, Hispaniola  M T OI S G

Ephyriades brunnea (Herrich-Schäffer, 1865)  'Cuban' Florida Duskywing  TL: Cuba

Ephyriades brunnea floridensis E. Bell & W. Comstock, 1948  S Florida  M T S L I

Ephyriades brunnea brunnea (Herrich-Schäffer, 1865)  Cuba, Isla de Juventud, Bahamas  M T S L

Ephyriades brunnea jamaicensis (Möschler, 1879)  Jamaica  M T S

Ephyriades brunnea dominicensis E. Bell & W. Comstock, 1948  Dominica and Guadeloupe  M T S

Ephyriades eugramma (Mabille, 1888)  Chiriqui Skipper  Costa Rica and Panama  M T OI S1 S2 I

Gesta  Evans, 1953

Gesta invisus (A. Butler & H. Druce, 1872)  False Duskywing  C & S TX to Costa Rica  M T S L I G

Gesta gesta (Herrich-Schäffer, 1863)  Impostor Duskywing  Greater Antilles, Costa Rica to Argentina  M T S1 S2 IL L I G

Gesta inga Evans, 1953  TL: Brazil (Ba)  M T

Gesta austerus (Schaus, 1902)  TL: Peru  M T S IL G

Gesta heteropterus (Plötz, 1884)  TL: Brazil  M S

Erynnis  Schrank, 1801

Erynnis icelus (Scudder & Burgess, 1870)  Dreamy Duskywing  widespread in USA and Canada  M S1 S2 L1 L2 L3 I Ha G

Erynnis brizo (Boisduval & Le Conte, [1837])  Sleepy Duskywing  TL: N. Amer.

Erynnis brizo somnus (Lintner, 1881)  Peninsular Florida  M T S L

Erynnis brizo brizo (Boisduval & Le Conte, [1837])  SE Canada, E USA, possibly far NE Mexico  M S1 S2 L1 L2 L3 L4 F Ha G

Erynnis brizo burgessi (Skinner, 1914)  S Rocky Mountain states, W TX, far SE CA, USA, far NW Mexico  M T S1 S2 L I Ha G

Erynnis brizo lacustra (W. G. Wright, 1905)  Baja California Norte, S & C CA  M T S L Ha G

Erynnis brizo muelleri (Draudt, 1923)  C & W Mexico  M T S L Ha G

Erynnis mercurius (Dyar, 1926)  Mexican Duskywing  S-C Mexico (MEX, DF)  M T S

Erynnis juvenalis (Fabricius, 1793)  Juvenal's Duskywing  TL: America

Erynnis juvenalis juvenalis (Fabricius, 1793)  E USA, SE Canada, far NE Mexico (COAH)  M T S L1 L2 L3 L4 L5 L6 F G

Erynnis juvenalis clitus (W. H. Edwards, 1883)  SE AZ, SW NM, W TX, E & W Mexico to Chiapas  M T S L I G

Erynnis telemachus Burns, 1960  Rocky Mountain Duskywing  E NV, UT, AZ, NM, CO, far S-C WY, W TX  M T S1 S2 L1 L2 I G

Erynnis propertius (Scudder & Burgess, 1870)  Propertius Duskywing  Cascades-Sierra Nevada, westward, S BC to northern Baja  M S1 S2 L1 L2 L3 Ha G

Erynnis meridianus E. Bell, 1927  Meridian Duskywing  TL: USA (Az)

Erynnis meridianus meridianus E. Bell, 1927  S NV, SW UT, AZ, NM, C & W TX, far N Mexico  M T S L I G

Erynnis meridianus fieldi Burns, 1964  C Mexico  M T S Ha

Erynnis scudderi (Skinner, 1914)  Scudder’s Duskywing  SE AZ, SW NM, W TX to Guatemala  M T S L Ha G

Erynnis horatius (Scudder & Burgess, 1870)  Horace's Duskywing  (far SE UT, far NE AZ), CO, NM, E USA, far NE Mexico (TAMPS)  M S1 S2 L1 L2 L3 L4 L5 I G

Erynnis tristis (Boisduval, 1852)  Mournful Duskywing  TL: USA (Ca)

Erynnis tristis tristis (Boisduval, 1852)  CA and Baja California Norte  M T S L Ha G

Erynnis tristis pattersoni Burns, 1964  Baja California Sur  M T S L

Erynnis tristis tatius (W. H. Edwards, 1883)  AZ, S NM, W & S TX to Colombia  M T S IL L1 L2 I Ha G

Erynnis martialis (Scudder, [1870])  Mottled Duskywing  MN to NH, S to N FL, W to C TX, CO, N NM W NE, W SD  M S1 S2 L H G

Erynnis pacuvius (Lintner, 1878)  Pacuvius Duskywing  TL: USA (NM)

Erynnis pacuvius pacuvius (Lintner, 1878)  S UT, S CO, AZ, NM to Oaxaca  M S L I G

Erynnis pacuvius lilius (Dyar, 1904)  NW CO, N UT, W WY, MT, ID, BC, WA, OR, W NV, N & E CA  M T S L I H

Erynnis pacuvius pernigra (F. Grinnell, 1905)  San Francisco Bay area, C CA  M T S

Erynnis pacuvius callidus (F. Grinnell, 1904)  S CA coast ranges to Baja California (norte)  M T S L H Ha

Erynnis zarucco (Lucas, 1857)  Zarucco Duskywing  SE USA (OK-NC-FL), Cuba, Hispaniola  M S L1 L2 G

Erynnis funeralis (Scudder & Burgess, 1870)  Funereal Duskywing  S CA, S AZ, S NM, TX S to Argentina (as resident); strays widely  M T S IL L1 L2 L3 I Ha G

Erynnis baptisiae (W. Forbes, 1936)  Wild Indigo Duskywing  E USA (far S Ontario) to N FL  M T S L1 L2 L3 L4 I H

Erynnis lucilius (Scudder & Burgess, 1870)  Columbine Duskywing  NE USA, SE Canada  M S G

Erynnis afranius (Lintner, 1878)  Afranius Duskywing  SE ALTA, S SASK to N Mexico

Erynnis afranius BCN segregate ,  SW CA and Baja California (norte)  M S

Erynnis afranius afranius (Lintner, 1878)  TL: USA (Co)  M T S1 S2 L1 L2 L3 L4 I F G

Erynnis afranius SE Mexican segregate ,  SE Mexico (Puebla and Mexico States)  M S

Erynnis persius (Scudder, 1863)  Persius Duskywing  TL: USA (Ma)

Erynnis persius persius (Scudder, 1863)  NE USA  M S L G

Erynnis persius borealis (M. Cary, 1906)  most of W Canada, possibly S along Cascades (& W) to N CA  M T S L

Erynnis persius avinoffi (W. Holland, 1930)  YT, Alaska  M T S

Erynnis persius fredericki H. Freeman, 1943  Rocky Mountain region, USA, possibly S SASK, S ALTA  M T1 T2 T3 T4 S L I1 I2 Ha

  Tribe Achlyodini

Aethilla  Hewitson, 1868

Aethilla eleusinia Hewitson, 1868  TL: Ecuador  M T S L G

Aethilla epicra Hewitson, 1870  TL: Ecuador  M T S HS G

Aethilla lavochrea A. Butler, 1872  Yellow-rimmed Skipper  E & W Mexico to Colombia  M T S1 S2 S3 L I

Aethilla chiapa H. Freeman, 1969  Highlands Skipper  Oaxaca, Chiapas, Guatemala  M T OD S G

Aethilla haber (Mabille, 1891)  TL: Peru  M T S

Aethilla echina Hewitson, 1870  Echina Skipper  TL: Ecuador

Aethilla echina echina Hewitson, 1870  SE Mexico to S Brazil  M T S IL L G

Aethilla echina coracina A. Butler, 1870  TL: Brazil (SP)  M T S

Aethilla memmius A. Butler, 1870  TL: Venezuela  M T OI S G Txt

Aethilla gigas (Mabille, 1877)  TL: Peru  M T S G

Aethilla melas Plötz, 1882  TL: Brazil (RJ)  M OD S G Txt

Aethilla later (Mabille, 1891)  TL: Peru  M T S

Eantis  Boisduval, 1836

Eantis tamenund (W. H. Edwards, 1871)  Northern Sicklewing (Sickle-winged Skipper)  C & S Texas (further N as strays) to N Belize and Guatemala  M T S L1 L2 L3 I1 I2 H G

Eantis thraso (Hübner, [1807])  Southern Sicklewing  SW Chiapas, S Belize, Guatemala to Argentina  M S1 S2 IL L I1 I2 G

Eantis mithridates (Fabricius, 1793)  Jamaican Sicklewing  Jamaica  M T S L

Eantis minor W. Comstock, 1944  Lesser Sicklewing  Puerto Rico to Dominica and Guadeloupe  M T S L

Eantis papinianus (Poey, 1832)  Cuban Sicklewing  [= sagra], Cuba, Hispaniola  M T S L I

Eantis munroei E. Bell, 1956  Munroe's Sicklewing  Cuba,XXXXX  M T OD S G

Eantis minna Evans, 1953  TL: Brazil (Ba)  M T S HS G

Achlyodes  Hübner, [1819]

Achlyodes busirus (Cramer, 1779)  Giant Sicklewing  TL: Surinam

Achlyodes busirus heros Ehrmann, 1909  E & W Mexico to Bolivia  M T S1 S2 HS IL L I1 I2 Txt

Achlyodes busirus busirus (Cramer, 1779)  TL: Surinam  M OI OD S G

Achlyodes busirus negro (Kaye, 1921)  TL: Trinidad  M T S G

Achlyodes busirus rioja Evans, 1953  TL: Brazil (RJ)  M T S HS IL G

Achlyodes pallida (R. Felder, 1869)  Pale Sicklewing  [= selva], W Mexico & S Texas (as stray) to Bolivia  M T S L1 L2 I1 I2 G

Doberes  Godman & Salvin, 1895

Doberes hewitsonius (Reakirt, [1867])  Pale Doberes  S Mexico  M T S L G

Doberes anticus (Plötz, 1884)  Dark Doberes  SE Mexico (VER, OAX, CHIS)

Doberes anticus anticus (Plötz, 1884)  TL: Mexico  M S L G

Doberes anticus Pale segregate ,  S Mexico (Chiapas) to Costa Rica  M S I

Doberes sobrinus (Godman & Salvin, 1895)  West-Mexican Doberes  W Mexico (Guerrero)  M T S

Ouleus  Lindsey, 1925

Ouleus cyrna (Mabille, 1895)  Hidden-yellow Skipper  Chiapas to Panama  M T S I

Ouleus bubaris (Godman & Salvin, 1895)  Bubaris Skipper  SE Mexico, Guatemala  M T S

Ouleus calavius (Godman & Salvin, 1895)  Calavius Skipper  SE Mexico to Panama  M T S

Ouleus juxta (E. Bell, 1934)  TL: Peru

Ouleus juxta juxta (E. Bell, 1934)  TL: Peru  M T S L G

Ouleus juxta dampa Evans, 1953  TL: French Guiana  M T

Ouleus fatinitza (Plötz, 1884)  TL: Colombia  M TI

Ouleus salvina Evans, 1953  Salvin's Skipper  E Mexico to Costa Rica  M T S L

Ouleus panna Evans, 1953  Panna Skipper  Guatemala to Colombia  M T S L

Ouleus fridericus (Geyer, 1832)  TL: Surinam

Ouleus fridericus hilarina (Mabille, 1903)  TL: Brazil (Pa)  M T S

Ouleus fridericus fridericus (Geyer, 1832)  TL: Surinam  M T1 T2 S

Ouleus fridericus sinepunctis (Kaye, 1904)  TL: Trinidad  M T S

Ouleus fridericus candangus O. Mielke, 1968  TL: Brazil (DF)  M T S

Ouleus fridericus riona Evans, 1953  TL: Brazil (RJ)  M T S IL G

Ouleus dilla Evans, 1953  TL: Ecuador

Ouleus dilla baru Nicolay, 1973  Costa Rica and Panama  M T OI S G

Ouleus dilla dilla Evans, 1953  TL: Ecuador  M T

Ouleus accedens (Mabille, 1895)  TL: Brazil (SC)

Ouleus accedens noctis (Lindsey, 1925)  TL: Peru  M S

Ouleus accedens finda Evans, 1953  TL: Brazil (ES)  M T S

Ouleus accedens accedens (Mabille, 1895)  TL: Brazil (SC)  M T S

Ouleus negrus Nicolay, 1980  Negrus Skipper  TL: Panama

Ouleus negrus negrus Nicolay, 1980  Costa Rica and Panama  M T S G

Ouleus negrus cristatus Steinhauser, 1989  TL: Colombia  M T

Ouleus candidus Steinhauser, 1989  TL: Colombia  M T

Ouleus narycus (Mabille, 1889)  TL: [Peru]  M T OI S IL

Zera  Evans, 1953

Zera belti (Godman & Salvin, 1894)  Belti Skipper  Chiapas to Nicaragua  M T OI S IL G

Zera teresa Steinhauser, 1989  TL: Brazil (ES)  M S

Zera zera (A. Butler, 1870)  Zera Skipper  Costa Rica to Venezuela, Ecuador and S Brazil  M T S HS IL L G

Zera difficilis (Weeks, 1901)  Difficult Skipper  Chiapas, Guianas and upper Amazon  M T OI S G

Zera nolckeni (Mabille, 1891)  Nolcken's Skipper  Panama to Bolivia (not in Mexico)  M T S IL L

Zera hosta Evans, 1953  Hosta Skipper  Chiapas to Colombia  M T S I

Zera phila (Godman & Salvin, 1894)  Phila Skipper  Panama to Peru and Bolivia  M T OI S IL L

Zera hyacinthinus (Mabille, 1877)  Bruised Skipper  TL: [Panama; Colombia]

Zera hyacinthinus hyacinthinus (Mabille, 1877)  E & W Mexico to Colombia  M T S IL L

Zera hyacinthinus gellius (Mabille, 1903)  TL: Ecuador  M S

Zera hyacinthinus servius (Plötz, 1884)  TL: Brazil  M S

Zera tetrastigma (Sepp, [1847])  Tetrastigma Skipper  TL: Surinam

Zera tetrastigma tetrastigma (Sepp, [1847])  Chiapas to Guianas, Trinidad, upper Amazon  M T S IL G

Zera tetrastigma erisichthon (Plötz, 1884)  TL: ?  M TI S IL

Zera eboneus (E. Bell, 1947)  Eboneus Skipper  SE Mexico  M T

Quadrus  Lindsey, 1925

Quadrus cerialis (Stoll, 1782)  Common Blue-Skipper  E & W Mexico to Argentina  M OI S1 S2 IL L I G

Quadrus francesius H. Freeman, 1969  Chiapan Blue-Skipper  S Mexico (Chiapas) to Costa Rica  M T S I

Quadrus ineptus (Draudt, 1922)  TL: Peru  M T OI S G

Quadrus jacobus (Plötz, 1884)  TL: Brazil (RJ)  M T S

Quadrus fanda Evans, 1953  TL: Brazil (Pa)  M T S

Quadrus tros Evans, 1953  TL: Brazil (ES)  M T

Quadrus contubernalis (Mabille, 1883)  Striped Blue-Skipper  TL: Brazil; French Guiana

Quadrus contubernalis anicius (Godman & Salvin, 1894)  SE Mexico, Guatemala  M T S IL L G

Quadrus contubernalis contubernalis (Mabille, 1883)  SE Mexico to upper Amazon  M T S1 S2 IL L I

Quadrus deyrollei (Mabille, 1877)  TL: French Guiana

Quadrus deyrollei deyrollei (Mabille, 1877)  TL: French Guiana  M T S IL

Quadrus deyrollei portulana (Plötz, 1884)  TL: S. Amer.  M S

Quadrus deyrollei porta Evans, 1953  TL: Brazil (Am)  M T S IL

Quadrus u-lucida (Plötz, 1884)  TL: Colombia

Quadrus u-lucida u-lucida (Plötz, 1884)  TL: Colombia  M S IL

Quadrus u-lucida mimus (Mabille & Boullet, 1917)  TL: Brazil (RS); Paraguay  M T S

Quadrus u-lucida parabus O. Mielke, 1968  TL: Brazil (MG)  M T

Quadrus lugubris (R. Felder, 1869)  Tanned Blue-Skipper  TL: Mexico (Ver)

Quadrus lugubris lugubris (R. Felder, 1869)  E & W Mexico to Costa Rica  M T S1 S2 IL L I G

Quadrus lugubris ophia (A. Butler, 1870)  TL: Venezuela  M T S

Quadrus truncata (Hewitson, 1870)  TL: Ecuador  M T S IL

Gindanes  Godman & Salvin, 1895

Gindanes brontinus Godman & Salvin, 1895  Straight-edged Skipper  TL: Nicaragua

Gindanes brontinus brontinus Godman & Salvin, 1895  E & W Mexico to Costa Rica  M T S I G

Gindanes brontinus bronta Evans, 1953  TL: Peru  M T S

Gindanes brebisson (Latreille, [1824])  TL: Brazil

Gindanes brebisson panaetius Godman & Salvin, 1895  SE Mexico to Panama  M T S

Gindanes brebisson brebna Evans, 1953  TL: Bolivia  M T S

Gindanes brebisson phagesia (Hewitson, 1868)  TL: Brazil (Pa)  M T S

Gindanes brebisson brebisson (Latreille, [1824])  TL: Brazil  M T S

Gindanes bora Evans, 1953  TL: Brazil (ES)  M T S

Pythonides  Hübner, [1819]

Pythonides lancea (Hewitson, 1868)  TL: Brazil  M S IL

Pythonides tullia Evans, 1953  TL: Brazil (RJ)  M T S

Pythonides jovianus (Stoll, 1782)  Variable Blue-Skipper  TL: Surinam

Pythonides jovianus amaryllis Staudinger, 1876  SE Mexico to Ecuador and Peru  M T S IL L I

Pythonides jovianus jovianus (Stoll, 1782)  Guatemala, Colombia, Guianas, Trinidad, upper Amazon  M S IL L

Pythonides jovianus crameri (Mabille & Boullet, 1917)  TL: Colombia  M T S IL

Pythonides jovianus fabricii W. F. Kirby, 1871  TL: Indiis  M S IL

Pythonides lerina (Hewitson, 1868)  TL: French Guiana; Brazil (Pa)  M T S IL

Pythonides grandis Mabille, 1878  TL: Brazil (Pa)

Pythonides grandis assecla Mabille, 1883  SE Mexico to upper Amazon (Peru)  M T S IL

Pythonides grandis grandis Mabille, 1878  TL: Brazil (Pa)  M T S

Pythonides parallelus Mabille, 1898  TL: Bolivia  M T S

Pythonides proxenus (Godman & Salvin, 1895)  Proxenus Blue-Skipper  SE Mexico to Venezuela and Ecuador  M T S IL L I G

Pythonides herennius Geyer, [1838]  Herennius Blue-Skipper  TL: Brazil

Pythonides herennius herennius Geyer, [1838]  Panama to Peru and lower Amazon  M T S IL L

Pythonides herennius lusorius Mabille, 1891  TL: Brazil (RJ)  M T S

Pythonides braga Evans, 1953  TL: Guyana  M T S

Pythonides nides O. Mielke & Casagrande, 2002  TL: Brazil (ES)  M T

Pythonides homer Evans, 1953  TL: Brazil (MT)  M T S

Pythonides pteras (Godman & Salvin, 1895)  Narrow-winged Blue-Skipper  Chiapas to Colombia  M T S IL I

Pythonides limaea (Hewitson, 1868)  TL: French Guiana  M T S

Pythonides neivai (Hayward, 1940)  TL: Ecuador  M S

Pythonides mundo H. Freeman, 1979  Freeman's Blue-Skipper  S Mexico (OAX)  M T S G

Pythonides eminus E. Bell, 1934  TL: Peru

Pythonides eminus pasha Evans, 1953  TL: Brazil (MT)  M T S

Pythonides eminus eminus E. Bell, 1934  TL: Peru  M T S

Pythonides rosa Steinhauser, 1989  Steinhauser's Blue-Skipper  S Mexico (CHIS)  M T S G

Pythonides hampa Evans, 1953  TL: Brazil (Pa)  M T S

Pythonides maraca O. Mielke & Casagrande, 1992  TL: Brazil (Rr)  M S

Pythonides pescada E. Bell, 1956  TL: Ecuador  M T

Haemactis  Mabille, 1903

Haemactis pyrrhosphenus Lindsey, 1919  TL: Colombia  M S

Haemactis sanguinalis (Westwood, 1852)  TL: Ecuador  M T S

Haemactis albamarita Austin, 1994  TL: Brazil (Rd)  M T S

Haemactis [n. sp.] N/A, N/A  Red-washed Skipper  Nicaragua and Costa Rica  M S L

Atarnes  Godman & Salvin, 1897

Atarnes sallei (C. Felder & R. Felder, 1867)  Orange-spotted Skipper  E & W Mexico to Colombia  M T S1 S2 L I G

Eburuncus  Grishin, 2012

Eburuncus hierax (Hopffer, 1874)  White-veined Tuskyhook  Peru  M T S IL G

Eburuncus unifasciata (C. Felder & R. Felder, 1867)  Single-banded Tuskyhook  SE Mexico to Ecuador and Peru  M T S1 S2 IL L I G

Milanion  Godman & Salvin, 1895

Milanion hemes (Cramer, 1777)  Hemes Skipper  TL: Surinam

Milanion hemes hemes (Cramer, 1777)  Costa Rica to Guianas and Trinidad  M T S L

Milanion hemes pemba Evans, 1953  TL: Peru  M T S

Milanion hemes memba Evans, 1953  TL: Brazil (MT)  M T S

Milanion cramba Evans, 1953  TL: Peru  M T S

Milanion clito (Fabricius, 1787)  TL: French Guiana  M T S Txt

Milanion leucaspis (Mabille, 1878)  TL: French Guiana; Brazil  M T S G

Milanion marciana Godman & Salvin, 1895  Marciana Skipper  Costa Rica to Colombia  M T S1 S2 I G

Milanion alaricus (Plötz, 1884)  TL: Brazil (Ba)  M T S G

Milanion pilumnus Mabille & Boullet, 1917  TL: Bolivia; Peru

Milanion pilumnus albidior Mabille & Boullet, 1917  TL: Venezuela  M T S

Milanion pilumnus pilta Evans, 1953  TL: Peru  M T S

Milanion pilumnus pilumnus Mabille & Boullet, 1917  TL: Bolivia; Peru  M T S G

Paramimus  Hübner, [1819]

Paramimus stigma (C. Felder & R. Felder, 1867)  TL: Colombia

Paramimus stigma stanna Evans, 1953  Panama, (Colombia)  M T S L

Paramimus stigma stigma (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Paramimus scurra (Hübner, [1809])  TL: ?

Paramimus scurra scurra (Hübner, [1809])  TL: ?  M T S G

Paramimus scurra herberti Godman & Salvin, 1896  TL: Brazil (MT)  M T S

Charidia  Mabille, 1903

Charidia lucaria (Hewitson, 1868)  TL: French Guiana

Charidia lucaria pilea Evans, 1953  TL: Colombia  M T S

Charidia lucaria lucaria (Hewitson, 1868)  TL: French Guiana  M T S

Charidia lucaria pocus Evans, 1953  TL: Peru  M T S

Charidia lucaria mayo Evans, 1953  TL: Peru  M T S

Charidia empolaeus (Westwood, 1852)  TL: ?  M T S

  Tribe Pyrgini

Pseudodrephalys  Burns, 1999

Pseudodrephalys sohni Burns, 1999  TL: Brazil (Am)  M T S

Pseudodrephalys atinas (Mabille, 1888)  TL: [Peru]  M T OI S G

Pseudodrephalys hypargus (Mabille, 1891)  TL: Brazil (Am)  M T S

Clito  Evans, 1953

Clito palotchka Grishin, 2014  I-Clito  Ecuador  M T

Clito jonkersi de Jong, 1983  Jonkers's Clito  TL: Surinam  M T OD

Clito litteroides Grishin, 2014  Subscripted Clito  Venezuela  M T

Clito littera (Mabille, 1877)  Scripted Clito  TL: Peru  M T OD S IL Txt

Clito trinidadensis Cock, 1998  Trinidad Clito  TL: Trinidad  M T OD

Clito nebulosa (Draudt, 1924)  Nebulous Clito  TL: Bolivia  M T OD G

Clito anda Evans, 1953  White-patched Clito  TL: French Guiana  M T S IL G

Clito bibulus (N. Riley, 1929)  Mottled Clito  TL: Brazil (MT)  M T S G

Clito aberrans (Draudt, 1924)  Northern Clito  ["clito"], S & W Mexico to S Brazil  M T OD S1 S2 S3 S4 IL L I1 I2 G

Clito sompa Evans, 1953  TL: Brazil (Ba)  M T S G

Clito zelotes (Hewitson, 1873)  Hewitson’s Clito  SE Mexico (VER) to French Guiana and upper Amazon  M T S IL L G

Clito tuva Evans, 1953  TL: Brazil (Pa)  M T G

Clito zenda Evans, 1953  TL: Surinam  M T S G

Clito clada (Evans, 1953)  TL: Brazil (Ma)  M T S IL G

Eracon  Godman & Salvin, 1894

Eracon biternata (Mabille, 1889)  Biternata Skipper  Costa Rica and Panama  M T OI TI S G

Eracon pebana Evans, 1953  Pebana Skipper  TL: Peru  M T S IL G

Eracon asymmetrica Austin, 1997  Asymmetric Skipper  TL: Brazil (Rd)  M T OI G

Eracon sarahburnsae Grishin, 2014  Sarah's Skipper  Costa Rica & Panama  M T1 T2 T3 T4 S I

Eracon clinias (Mabille, 1878)  Clinias Skipper  Costa Rica to upper and lower Amazon  M T S1 S2 IL L G

Eracon paulinus (Stoll, 1782)  Tear-drop Skipper  Costa Rica to Guiana and upper Amazon  M T OI OD S1 S2 S3 S4 IL L I G Txt

Eracon lachesis (Dyar, 1918)  Lachesis Skipper  S Mexico to Panama  M T S1 S2 S3 L I

Eracon mnemon (Schaus, 1913)  Horseshoe-spotted Skipper  Costa Rica  M T S

Cornuphallus  Austin, 1997

Cornuphallus problematica (R. Williams & E. Bell, 1940)  Tricky Hornydick  TL: Argentina  M T G

Cornuphallus albafascia Austin, 1997  White-banded Hornydick  TL: Brazil (Rd)  M T OI G

Cornuphallus onoribo (Möschler, 1883)  Onoribo Hornydick  TL: Surinam  M T S

Spioniades  Hübner, [1819]

Spioniades abbreviata (Mabille, 1888)  White-tipped Skipper  TL: Panama

Spioniades abbreviata abbreviata (Mabille, 1888)  Nicaragua to Colombia  M T OI S1 S2 I

Spioniades abbreviata anta Evans, 1953  TL: Bolivia  M T S

Spioniades artemides (Stoll, 1782)  Artemides Skipper  Panama to S Brazil  M S L I G

Spioniades libethra (Hewitson, 1868)  TL: Brazil (Am)  M T OI S1 S2 S3

Plumbago  Evans, 1953

Plumbago plumbago (Plötz, 1884)  TL: ?  M T OI S L

Plumbago pulverea (Mabille, 1878)  TL: French Guiana  M T TI S

Plumbago serapion (Plötz, 1884)  TL: Brazil (RJ)  M T

Trina  Evans, 1953

Trina geometrina (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia

Trina geometrina geometrina (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia  M T S IL L

Trina geometrina phalaena (Mabille, 1898)  TL: Bolivia  M T S L

Paches  Godman & Salvin, 1895

Paches loxus (Westwood, 1852)  Glorious Blue-Skipper  TL: [Venezuela]

Paches loxus loxus (Westwood, 1852)  Panama to Venezuela, Ecuador, N Brazil  M T OI S IL L G

Paches loxus gloriosus Röber, 1925  [= zonula], SE Mexico to Costa Rica  M T S1 S2 S3 L I

Paches loxus loxana Evans, 1953  TL: Bolivia  M T S

Paches gladiatus (A. Butler, 1870)  TL: Brazil (Pa)  M T S

Paches polla (Mabille, 1888)  Polla Blue-Skipper  E & W Mexico to Costa Rica  M T OI S IL L

Paches trifasciatus Lindsey, 1925  TL: Peru  M S V

Paches era Evans, 1953  TL: Peru  M T S

Paches exosa (A. Butler, 1877)  TL: Brazil (Am)  M T S L

Carrhenes  Godman & Salvin, 1895

Carrhenes fuscescens (Mabille, 1891)  Tanned Hoary-Skipper  TL: Honduras

Carrhenes fuscescens fuscescens (Mabille, 1891)  E & W Mexico to Costa Rica  M T S1 S2 L I G

Carrhenes fuscescens conia Evans, 1953  TL: French Guiana  M T S G

Carrhenes calidius Godman & Salvin, 1895  Rain-forest Hoary-Skipper  SE Mexico to Bolivia  M T S1 S2 I G

Carrhenes sinesinus Steinhauser, 1989  TL: Colombia  M T

Carrhenes bamba Evans, 1953  TL: Peru  M T S IL G

Carrhenes infuscescens Steinhauser, 1989  TL: Argentina  M T S

Carrhenes chaeremon (Mabille, 1891)  TL: Brazil (Am)  M T S IL G

Carrhenes canescens (R. Felder, 1869)  Hoary Skipper  TL: Mexico (Ver)

Carrhenes canescens canescens (R. Felder, 1869)  W Mexico & S TX to Panama  M T S1 S2 IL L I G

Carrhenes canescens pallida Röber, 1925  TL: Brazil (RS)  M T S IL

Carrhenes lilloi Hayward, 1947  TL: Ecuador  M TI IL G

Carrhenes leada (A. Butler, 1870)  Butler's Hoary-Skipper  Costa Rica to Bolivia and S Brazil  M T S HS IL G

Carrhenes recurva Austin, 2000  TL: Brazil (Rd)  M T OI G

Carrhenes callipetes Godman & Salvin, 1895  Cloud-forest Hoary-Skipper  S Mexico and Guatemala  M T OI S L G

Carrhenes meridensis Godman & Salvin, 1895  Costa Rican Hoary-Skipper  Nicaragua to Colombia and Venezuela  M T OI S L G

Carrhenes santes E. Bell, 1940  TL: Peru  M T S

Zobera  H. Freeman, 1970

Zobera albopunctata H. Freeman, 1970  Coliman Zobera  W Mexico (Colima)  M T S G

Zobera marginata H. Freeman, 1979  Western Zobera  W Mexico (NAY, MICH, OAX)  M T S L G

Zobera oaxaquena Steinhauser, 1991  Oaxacan Zobera  S Mexico (OAX)  M T1 T2 S L G

Xenophanes  Godman & Salvin, 1895

Xenophanes tryxus (Stoll, 1780)  Glassy-winged Skipper  S Texas (as stray) to Argentina  M T S1 S2 IL L1 L2 I1 I2 G

Diaeus  Godman & Salvin, 1895

Diaeus varna Evans, 1953  Camouflaged Skipper  E Mexico to Panama  M T S L G

Diaeus lacaena (Hewitson, 1869)  TL: Brazil  M T S L

Diaeus ambata Evans, 1953  TL: Ecuador  M T S

Diaeus variegata (Plötz, 1884)  TL: Brazil (RJ)  M T TI S G

Onenses  Godman & Salvin, 1895

Onenses hyalophora (R. Felder, 1869)  Crystal-winged Skipper  E Mexico to Guatemala  M S L G

Onenses kelso Evans, 1953  TL: Guyana  M T S

Antigonus  Hübner, [1819]

Antigonus nearchus (Latreille, [1817])  Large Spurwing  E & W Mexico to S Brazil and Paraguay  M T S IL L I G

Antigonus erosus (Hübner, [1812])  Common Spurwing  W Mexico & S TX (rare stray) to S Brazil and Paraguay  M T S1 S2 IL L1 L2 I G

Antigonus emorsa (R. Felder, 1869)  White Spurwing  W Mexico (SE AZ as rare stray, SON to OAX)  M T S1 S2 L I G

Antigonus decens A. Butler, 1874  TL: Peru  M T S L

Antigonus funebris (R. Felder, 1869)  West-Mexican Spurwing  W & S Mexico  M T S L G

Antigonus minor O. Mielke, 1980  TL: Brazil (RS)  M T OI S G

Antigonus corrosus Mabille, 1878  Small Spurwing  E Mexico to Panama  M T S L

Antigonus mutilatus (Hopffer, 1874)  TL: Peru  M T S L1 L2 G

Antigonus liborius Plötz, 1884  TL: Brazil (Ba)

Antigonus liborius liborius Plötz, 1884  TL: Brazil (Ba)  M T S IL L G

Antigonus liborius areta Evans, 1953  TL: Brazil (RS)  M T S

Systasea  W. H. Edwards, 1877

Systasea microsticta Dyar, 1923  Rare Spurwing  S Mexico (GRO, MOR, PUE, OAX)  M T S L Ha

Systasea pulverulenta (R. Felder, 1869)  Texas Powdered-Skipper  S AZ, S NM, C & S TX to Guatemala  M T S L1 L2 I H G

Systasea zampa (W. H. Edwards, 1876)  Arizona Powdered-Skipper  Baja, far S CA, S NV, S AZ, S NM, W TX, SON, CHIH, COAH  M T1 T2 TI S L I H G

Celotes  Godman & Salvin, 1899

Celotes nessus (W. H. Edwards, 1877)  Common Streaky-Skipper  far SE CA, S AZ, NM, S OK, TX, N Mexico  M T S1 S2 L I H G

Celotes spurcus A. Warren, Steinhauser, Hernandez-Mejia & Grishin, 2008  Mexican Streaky-Skipper  Central Mexico  M T L Ha G

Celotes limpia Burns, 1974  West-Texas Streaky-Skipper  W TX to Coahuila  M T S L G

Zopyrion  Godman & Salvin, 1896

Zopyrion sandace Godman & Salvin, 1896  Sandy Skipper  E & W Mexico to Costa Rica  M T S L G

Zopyrion satyrina (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia  M T S

Zopyrion evenor Godman, 1901  TL: Brazil (MS)

Zopyrion evenor tressa Evans, 1953  TL: Brazil (Pa)  M T

Zopyrion evenor thania Evans, 1953  TL: Brazil (Pe)  M T S

Zopyrion evenor evenor Godman, 1901  TL: Brazil (MS)  M T S

Zopyrion subvariegata Hayward, 1942  TL: Ecuador

Zopyrion subvariegata subvariegata Hayward, 1942  TL: Ecuador  M T S

Zopyrion subvariegata thyas Evans, 1953  TL: Peru  M T S

Zopyrion reticulata Hayward, 1942  TL: Brazil (Go)  M S

Timochreon  Godman & Salvin, 1896

Timochreon satyrus (C. Felder & R. Felder, 1867)  Satyr Skipper  TL: Colombia

Timochreon satyrus satyrus (C. Felder & R. Felder, 1867)  Costa Rica to upper Amazon  M T S1 S2 L I G

Timochreon satyrus forta Evans, 1953  TL: Brazil (Pa)  M T S

Timochreon satyrus tampa Evans, 1953  TL: Brazil (MT)  M T S IL G

Timochreon doria (Plötz, 1884)  TL: "Mexico" [Brazil]  M S

Anisochoria  Mabille, 1877

Anisochoria nadia Nicolay, 1980  TL: Ecuador  M T

Anisochoria bacchus Evans, 1953  Northern Snout-Skipper  SE Mexico to El Salvador  M T S L G

Anisochoria pedaliodina (A. Butler, 1870)  TL: ?

Anisochoria pedaliodina polysticta Mabille, 1877  Costa Rica to Guianas  M T S I

Anisochoria pedaliodina pedaliodina (A. Butler, 1870)  TL: ?  M T S IL L1 L2 G

Anisochoria pedaliodina extincta Hayward, 1933  TL: Argentina  M S IL G

Anisochoria sublimbata Mabille, 1883  TL: [Brazil (SP)]  M T TI S IL L G

Anisochoria subpicta Schaus, 1902  TL: Brazil (Pr)  M T S

Anisochoria minorella Mabille, 1898  TL: Bolivia

Anisochoria minorella verda Evans, 1953  TL: Ecuador  M T S

Anisochoria minorella minorella Mabille, 1898  TL: Bolivia  M T S L

Anisochoria quadrifenestrata (Bryk, 1953)  TL: Brazil (Ba)  M T S

Anisochoria superior Mabille, 1898  TL: Bolivia  M T S

Anisochoria vianna Evans, 1953  TL: Brazil (Go)  M T S

Anisochoria albida Mabille, 1888  TL: [Peru]

Anisochoria albida albida Mabille, 1888  TL: [Peru]  M T OI S

Anisochoria albida alco Evans, 1953  TL: Bolivia  M T S

Pyrgus  Hübner, [1819]

Pyrgus centaureae (Rambur, [1842])  Grizzled Skipper  TL: not specified

Pyrgus centaureae freija (B. Warren, 1924)  boreal N America  M T S Ha

Pyrgus centaureae loki Evans, 1953  Rocky Mountains  M T S1 S2 S3 L1 L2 I1 I2 Ha

Pyrgus centaureae wyandot (W. H. Edwards, 1863)  E & NE USA  M T S L1 L2 G

Pyrgus limbata (Erschoff, 1876)  TL: Peru

Pyrgus limbata limbata (Erschoff, 1876)  TL: Peru  M T S

Pyrgus limbata nigella (Weeks, 1902)  TL: Bolivia  M S L

Pyrgus fides Hayward, 1940  TL: Argentina; Chile; Peru  M T S IL G

Pyrgus bocchoris (Hewitson, 1874)  TL: Bolivia

Pyrgus bocchoris bocchoris (Hewitson, 1874)  TL: Bolivia  M T S IL G

Pyrgus bocchoris cuzcona (Draudt, 1923)  TL: Peru  M T S L G

Pyrgus bocchoris trisignatus (Mabille, 1876)  TL: Chile  M T S

Pyrgus notatus (Blanchard, 1852)  TL: Chile

Pyrgus notatus notatus (Blanchard, 1852)  TL: Chile  M T S Ab

Pyrgus notatus valdiviana (Reed, 1877)  TL: Chile  M T

Pyrgus barrosi Ureta, 1956  TL: Chile  M S

Pyrgus ruralis (Boisduval, 1852)  Two-banded Checkered-Skipper  TL: USA (Ca)

Pyrgus ruralis ruralis (Boisduval, 1852)  NW CO, UT, W WY, W MT, SW ALTA, S BC to C CA  M T S1 S2 S3 L1 L2 Ha G

Pyrgus ruralis lagunae Scott, 1981  Laguna Mts., San Diego Co., CA  M T1 T2 T3 S L Ha1 Ha2

Pyrgus xanthus W. H. Edwards, 1878  Mountain Checkered-Skipper  S UT, N & E AZ, NM, S-C & SW CO  M T S IL L1 L2 I H G

Pyrgus scriptura (Boisduval, 1852)  Small Checkered-Skipper  TL: USA (Ca)

Pyrgus scriptura scriptura (Boisduval, 1852)  at least San Francisco Bay area, CA  M T S L G

Pyrgus scriptura apertorum Austin, 1998  most of range, ALTA to Zacatecas  M T S L I Ha

Pyrgus crisia Herrich-Schäffer, 1865  Antillean Checkered-Skipper  Cuba and Hispaniola  M T S G

Pyrgus veturius Plötz, 1884  TL: ?  M T S

Pyrgus communis (Grote, 1872)  Common Checkered-Skipper  TL: USA (Al)

Pyrgus communis W Oregon segregate ,  W Oregon, N CA Siskiyous  M S L Ha

Pyrgus communis high-elevation Cascades-Sierra Nevada segregate ,  high elevation C Cascades, OR, & N Sierra Nevada, CA  M L

Pyrgus communis communis (Grote, 1872)  widespread in USA & S Canada, S to Oaxaca  M T S1 S2 Ab L1 L2 L3 L4 I F Ha G

Pyrgus albescens Plötz, 1884  White Checkered-Skipper  S USA (CA to SC), S to Oaxaca  M T S L1 L2 I H G V

Pyrgus adepta Plötz, 1884  TL: Colombia  M T S1 S2 L G

Pyrgus orcynoides (Giacomelli, 1928)  TL: Argentina  M S IL L G

Pyrgus chloe Evans, 1942  TL: Peru  M T S

Pyrgus titicaca (Reverdin, 1921)  TL: ?  M S

Pyrgus philetas W. H. Edwards, 1881  Desert Checkered-Skipper  S AZ to S TX, S to Chiapas  M T S L I H G

Pyrgus oileus (Linnaeus, 1767)  Tropical Checkered-Skipper  SE AZ, SW NM, S TX to GA, S to Costa R., Gr. & Lesser Antilles  M T S1 S2 HS Ma L1 L2 L3 L4 I G

Pyrgus orcus (Stoll, 1780)  Orcus Checkered-Skipper  Oaxaca and Chiapas to Argentina, Trinidad to Dominica  M T S1 S2 IL Co L1 L2 G

Pyrgus brenda Evans, 1942  Brenda Checkered-Skipper  Ecuador & Peru  M T S Co G

Heliopyrgus  Herrera, 1957

Heliopyrgus americanus (Blanchard, 1852)  TL: Chile

Heliopyrgus americanus americanus (Blanchard, 1852)  TL: Chile  M S

Heliopyrgus americanus bellatrix (Plötz, 1884)  TL: Argentina  M T S IL G

Heliopyrgus domicella (Erichson, [1849])  Erichson’s White-Skipper  TL: Guyana

Heliopyrgus domicella domicella (Erichson, [1849])  S CA & S NV to S TX, S to Venezuela  M T S L1 L2 I G

Heliopyrgus domicella margarita (E. Bell, 1937)  TL: Venezuela  M T S

Heliopyrgus domicella willi (Plötz, 1884)  TL: Brazil (MG)  M S IL G

Heliopyrgus sublinea (Schaus, 1902)  East-Mexican White-Skipper  S TX & E Mexico to Yucatan Peninsula  M T S L G

Heliopetes  Billberg, 1820

Heliopetes ericetorum (Boisduval, 1852)  Northern White-Skipper  Baja California (norte), CA, Columbia Basin, Great Basin, AZ  M T S L1 L2 L3 I Ha G

Heliopetes omrina (A. Butler, 1870)  Stained White-Skipper  Panama to Argentina  M T S IL L G

Heliopetes laviana (Hewitson, 1868)  Laviana White-Skipper  TL: Nicaragua

Heliopetes laviana laviana (Hewitson, 1868)  S AZ and Baja California, C & S TX, most of Mexico to Venez.  M T S L1 L2 L3 L4 L5 I H G

Heliopetes laviana leca (A. Butler, 1870)  TL: Venezuela  M T

Heliopetes libra Evans, 1944  TL: Brazil (MT)  M T S IL L G

Heliopetes macaira (Reakirt, [1867])  Turk’s-cap White-Skipper  TL: Mexico (Ver)

Heliopetes macaira macaira (Reakirt, [1867])  S TX, E & W Mexico to Panama  M T S L I G

Heliopetes macaira orbigera (Mabille, 1888)  TL: Brazil  M T OI S

Heliopetes nivella (Mabille, 1883)  TL: Colombia  M T S

Heliopetes purgia Schaus, 1902  TL: Brazil (Pr)  M T S

Heliopetes arsalte (Linnaeus, 1758)  Veined White-Skipper  S TX, E & W Mexico to Argentina  M T S1 S2 IL L1 L2 I G

Heliopetes marginata Hayward, 1940  TL: Ecuador  M T S

Heliopetes leucola (Hewitson, 1868)  TL: Brazil (MG)  M T S

Heliopetes ochroleuca J. Zikán, 1938  TL: Brazil (MG)  M T S IL L G

Heliopetes petrus (Hübner, [1819])  TL: ?  M T S

Heliopetes chimbo Evans, 1953  TL: Ecuador  M T S

Heliopetes alana (Reakirt, 1868)  Alana White-Skipper  E & W Mexico to Argentina  M T S1 S2 IL L1 L2 G

  Subfamily Heteropterinae

Carterocephalus  Lederer, 1853

Carterocephalus palaemon (Pallas, 1771)  Arctic Skipperling  TL: not specified

Carterocephalus palaemon mesapano (Scudder, 1868)  New England, NE Canadian Provinces  M S L G

Carterocephalus palaemon mandan (W. H. Edwards, 1863)  SE SASK through Great Lakes region  M S L

Carterocephalus palaemon S Rocky Mountain segregate ,  NW WY to ALTA  M S L Ha

Carterocephalus palaemon skada (W. H. Edwards, 1870)  Alaska to N CA  M T S L1 L2 Ha

Carterocephalus palaemon magnus Mattoon & Tilden, 1998  coastal Sonoma Co., CA  M T S

Freemaniana  A. Warren, 2001

Freemaniana rawlinsi (A. Warren, 2001)  Rawlins' Skipperling  Ecuador (Zamora)  M S

Dalla  Mabille, 1904

Dalla calaon (Hewitson, 1877)  Calaon Skipperling  Ecuador  M T

Dalla thalia Evans, 1955  Thalia Skipperling  Ecuador  M T G

Dalla orsines (Hewitson, 1877)  Orsines Skipperling  Venezuela, Bolivia  M T S G

Dalla grovius (Mabille, 1898)  Grovius Skipperling  TL: Ecuador

Dalla grovius grovius (Mabille, 1898)  Ecuador (Loja), Peru   M T S

Dalla grovius amba Evans, 1955  Ecuador (Ambato)  M S

Dalla grovius floxa Evans, 1955  Peru (Amazonas, Chachapoyas)  M S

Dalla riza (Mabille, 1889)  Riza Skipperling  Peru, Bolivia  M T S

Dalla agathocles (C. Felder & R. Felder, 1867)  Agathocles Skipperling  TL: Colombia

Dalla agathocles polydesma (Mabille, 1889)  Venezuela  M T S

Dalla agathocles agathocles (C. Felder & R. Felder, 1867)  Colombia (Bogota)  M S

Dalla agathocles lanna Evans, 1955  Colombia (Cauca), Ecuador, Peru  M S

Dalla agathocles lonia Evans, 1955  Peru (Huancabamba)  M S

Dalla lenda Evans, 1955  Lenda Skipperling  Ecuador (Zamora, Loja)  M S

Dalla lorda Evans, 1955  Lorda Skipperling  Ecuador (Zamora, Loja)  M T

Dalla costala Evans, 1955  Costala Skipperling  TL: Bolivia

Dalla costala ascha Evans, 1955  Peru (Huancabamba, Pozuzo, La Merced)  M S

Dalla costala zona Evans, 1955  Peru (Amazonas, Chachapoyas)  M S

Dalla costala costala Evans, 1955  Peru, Bolivia  M S

Dalla huanca Evans, 1955  Huancabamba Skipperling  Peru (Huancabamba)  M S

Dalla sepia Evans, 1955  Falcate Skipperling  Peru  M S

Dalla hesperioides (C. Felder & R. Felder, 1867)  Hesperioides Skipperling  TL: Colombia

Dalla hesperioides hesperioides (C. Felder & R. Felder, 1867)  Colombia  M T S

Dalla hesperioides hister Evans, 1955  Peru (Limbani)  M T S

Dalla polycrates (C. Felder & R. Felder, 1867)  Polycrates Skipperling  TL: Colombia

Dalla polycrates polycrates (C. Felder & R. Felder, 1867)  Colombia  M T

Dalla polycrates ambala Evans, 1955  Ecuador, Peru (Huancabamba)  M S

Dalla polycrates lania Evans, 1955  SE Peru (Aqualani), Bolivia  M S

Dalla superior Draudt, 1923  Superior Skipperling  Colombia, Ecuador  M T S G

Dalla epiphaneus (C. Felder & R. Felder, 1867)  Epiphaneus Skipperling  TL: Colombia

Dalla epiphaneus epiphaneus (C. Felder & R. Felder, 1867)  Colombia  M T S

Dalla epiphaneus gaujoni (Mabille, 1898)  S Ecuador, Peru  M T S

Dalla epiphaneus poya Evans, 1955  Peru (Amazonas)  M S

Dalla epiphaneus anca Evans, 1955  Peru (Huancabamba)  M S

Dalla epiphaneus limba Evans, 1955  Peru  M S

Dalla epiphaneus junga Evans, 1955  Bolivia  M T

Dalla cola E. Bell, 1959  Cola Skipperling  Colombia  M T

Dalla frontinia Evans, 1955  Frontinia Skipperling  TL: Colombia

Dalla frontinia venda Evans, 1955  Venezuela, [Peru ?]  M T S

Dalla frontinia frontinia Evans, 1955  Colombia, Ecuador  M S

Dalla frontinia vanca Evans, 1955  Peru  M S

Dalla taza Evans, 1955  Taza Skipperling  Colombia, Ecuador  M T S

Dalla pota E. Bell, 1959  Pota Skipperling  Ecuador  M T

Dalla bubobon (Dyar, 1921)  Bubobon Skipperling  S-C Mexico (COL, MICH, GRO)  M T S

Dalla freemani A. Warren, 1997  Freeman's Skipperling  Guatemala  M T G

Dalla austini A. Warren & R. G. Maza, 2011  Austin's Skipperling  S Mexico (Chiapas)  M T

Dalla steinhauseri H. Freeman, 1991  Steinhauser's Skipperling  E Oaxaca  M T L

Dalla pulchra (Godman, 1900)  Costa Rican Skipperling  Costa Rica  M T S L

Dalla nubes Steinhauser, 1991  Chiapan Skipperling  S Mexico (OAX & CHIS)  M T S L

Dalla ligilla (Hewitson, 1877)  Ligilla Skipperling  W (JAL) & S Mexico to Costa Rica  M T S L

Dalla dividuum (Dyar, 1913)  Dyar's Skipperling  W Mexico (SON to CHIS)  M T S L Ha

Dalla dimidiatus (C. Felder & R. Felder, 1867)  Dimidiatus Skipperling  TL: Colombia; Venezuela

Dalla dimidiatus dimidiatus (C. Felder & R. Felder, 1867)  Venezuela, Colombia, Ecuador  M T S

Dalla dimidiatus lilla Evans, 1955  S Ecuador, Peru, Bolivia  M S

Dalla dimidiatus pucer Evans, 1955  Peru (Huancabamba)  M S

Dalla cocha Evans, 1955  Cochabamba Skipperling  Peru, Bolivia  M S L

Dalla miser Evans, 1955  Miser Skipperling  Colombia, Peru  M S L

Dalla caicus (Hewitson, 1868)  Caicus Skipperling  TL: Venezuela

Dalla caicus caicus (Hewitson, 1868)  Venezuela  M T S

Dalla caicus inca Draudt, 1923  Colombia, Ecuador, Peru, Bolivia  M T S L G

Dalla hilina (A. Butler, 1870)  Hilina Skipperling  Colombia, Venezuela  M T

Dalla dognini (Mabille, 1889)  Dognin's Skipperling  Ecuador  M T S L

Dalla scylla (Mabille, 1898)  Scylla Skipperling  Peru, Bolivia, N Argentina  M T S1 S2

Dalla genes (Mabille, 1898)  Genes Skipperling  TL: Ecuador

Dalla genes saleca (Mabille, 1898)  Colombia  M T S

Dalla genes genes (Mabille, 1898)  S Ecuador  M T S

Dalla genes golia Evans, 1955  N Peru  M T S

Dalla genes nona Evans, 1955  S Peru, Bolivia  M T S

Dalla cyprius (Mabille, 1898)  Cyprius Skipperling  TL: Bolivia

Dalla cyprius quinka Evans, 1955  Peru  M S

Dalla cyprius cyprius (Mabille, 1898)  Bolivia  M T S

Dalla vinca Evans, 1955  Vinca Skipperling  TL: Peru  M T S

Dalla cypselus (C. Felder & R. Felder, 1867)  Cypselus Skipperling  TL: Colombia

Dalla cypselus cypselus (C. Felder & R. Felder, 1867)  Venezuela, Colombia  M T S

Dalla cypselus evages (Hewitson, 1877)  Ecuador, Peru, Bolivia  M T S

Dalla dora E. Bell, 1947  Dora Skipperling  Ecuador  M T S

Dalla spica Hayward, 1939  Spica Skipperling  TL: Argentina

Dalla spica livia Evans, 1955  Peru, Bolivia  M T S

Dalla spica spica Hayward, 1939  Argentina  M S

Dalla quasca E. Bell, 1947  Quasca Skipperling  TL: Colombia

Dalla quasca quasca E. Bell, 1947  Venezuela, Colombia  M T S

Dalla quasca equatoria E. Bell, 1947  Ecuador  M T S

Dalla pincha Steinhauser, 1991  Pinche Skipperling  Ecuador   M T S

Dalla gelus (Mabille, 1898)  Gelus Skipperling  Ecuador  M T S HS

Dalla cupavia (Mabille, 1898)  Cupavia Skipperling  TL: Bolivia

Dalla cupavia elka Evans, 1955  N & C Peru  M S

Dalla cupavia cupavia (Mabille, 1898)  S Peru, Bolivia  M T S

Dalla frater (Mabille, 1878)  Frater Skipperling  Panama to Bolivia  M T S1 S2 L

Dalla mesoxantha (Plötz, 1884)  Mesoxantha Skipperling  Venezuela, Colombia, Ecuador, Peru, Bolivia  M T S

Dalla wardi Steinhauser, 2002  Ward's Skipperling  Colombia (Magdalena, 8 km E San Pedro, 2550m)  M T OD S G

Dalla merida Evans, 1955  Merida Skipperling  Venezuela, Colombia  M T S

Dalla xantha Steinhauser, 1991  Xantha Skipperling  Colombia  M T L

Dalla geon (Mabille, 1898)  Geon Skipperling  Ecuador  M T S

Dalla octomaculata (Godman, 1900)  Eight-spotted Skipperling  Costa Rica  M S

Dalla oxaites (Hewitson, 1877)  Oxaites Skipperling  Peru, Bolivia  M T S

Dalla eryonas (Hewitson, 1877)  Central American Skipperling  Costa Rica to Colombia  M T S L

Dalla mentor Evans, 1955  Guerrero Skipperling  SW Mexico (Guerrero & Oaxaca)  M S

Dalla kemneri Steinhauser, 1991  Pelon Skipperling  SE Mexico (Oaxaca & Puebla)  M T S L

Dalla pedro Steinhauser, 2002  Pedro Skipperling  Colombia (Magdalena, 8 km E San Pedro, 2550m)  M T OD G

Dalla connexa Draudt, 1923  Connexa Skipperling  Colombia, Ecuador, Peru  M T S G Txt

Dalla disconnexa Steinhauser, 2002  Disconnexa Skipperling  Ecuador  M T OD G

Dalla vista Steinhauser, 2002  Vista Skipperling  Ecuador  M T OD S G

Dalla ticidas (Mabille, 1898)  Ticidas Skipperling  Peru, Bolivia  M T S

Dalla caenides (Hewitson, 1868)  Caenides Skipperling  Venezuela, Colombia, Ecuador  M T S

Dalla bos Steinhauser, 1991  Bos Skipperling  Colombia (Cauca)  M T

Dalla pura Steinhauser, 1991  Pura Skipperling  Colombia (Cauca)  M T

Dalla mora Evans, 1955  Mora Skipperling  Ecuador, N Peru  M T S G

Dalla carnis Evans, 1955  Carnis Skipperling  TL: Bolivia

Dalla carnis sondra Evans, 1955  Peru  M T S

Dalla carnis carnis Evans, 1955  S Peru, Bolivia  M S

Dalla monospila (Mabille, 1898)  TL: Bolivia  M T S

Dalla lalage (Godman, 1900)  Godman’s Skipperling  SW Mexico (CHIS, GRO, OAX)  M S L

Dalla lethaea (Schaus, 1913)  Schaus' Skipperling  S Mexico (CHIS, GRO, OAX, PUE) to Costa Rica  M T OI S L

Dalla faula (Godman, 1900)  West-Mexican Skipperling  W Mexico (SON to MICH & PUE)  M S L

Dalla lysis (Schaus, 1913)  Lysis Skipperling  Costa Rica  M OI

Dalla diraspes (Hewitson, 1877)  Widespread Skipperling  Costa Rica to S Brazil  M T S L

Dalla simplicis Steinhauser, 1991  Simple Skipperling  Colombia  M T S

Dalla seirocastnia Draudt, 1923  Seirocastnia Skipperling  Colombia, Ecuador  M T S G

Dalla celsus Steinhauser, 2002  Celsus Skipperling  Ecuador (Chimborazo)  M T OD G

Dalla pantha Evans, 1955  Pantha Skipperling  N Peru (Huancabamba)  M T

Dalla puracensis Steinhauser, 1991  Purace Skipperling  TL: Colombia

Dalla puracensis puracensis Steinhauser, 1991  Colombia  M T S

Dalla puracensis cotopa Steinhauser, 1991  Ecuador  M T

Dalla puracensis quindio Steinhauser, 1991  Colombia  M T

Dalla ochrolimbata Draudt, 1923  Ochro Skipperling  Peru  M T G

Dalla ramirezi H. Freeman, 1969  Gold-rayed Skipperling  S Mexico (OAX, PUE, VER) to Guatemala  M T S L Ha

Dalla jelskyi (Erschoff, 1875)  Jelsky's Skipperling  TL: Peru

Dalla jelskyi jelskyi (Erschoff, 1875)  Peru  M S

Dalla jelskyi aurosa J. Zikán, 1938  S Brazil  M S

Dalla tona Evans, 1955  Tona Skipperling  Peru (Huancabamba)  M T

Dalla decca Evans, 1955  Decca Skipperling  TL: Ecuador

Dalla decca decca Evans, 1955  Colombia, Ecuador  M S

Dalla decca doppa Evans, 1955  Peru  M S

Dalla eburones (Hewitson, 1877)  Eburones Skipperling  TL: Bolivia

Dalla eburones inornata (E. Bell, 1937)  Colombia, Ecuador  M T S

Dalla eburones ena Evans, 1955  Peru (Amazonas, Chachapoyas)  M T S

Dalla eburones eburones (Hewitson, 1877)  SE Peru, Bolivia  M T S

Dalla morva (Mabille, 1898)  Morva Skipperling  Peru, Bolivia  M T S

Dalla ibhara (A. Butler, 1870)  Ibhara Skipperling  Venezuela, Colombia, Ecuador, Peru, Bolivia  M T S

Dalla plancus (Hopffer, 1874)  Plancus Skipperling  Peru, Bolivia  M T S L

Dalla crithote (Hewitson, 1874)  Crithoe Skipperling  Colombia, Ecuador  M T S

Dalla calima Steinhauser, 1991  Calima Skipperling  Colombia (Valle del Cauca)  M T

Dalla rosea Evans, 1955  Rosy Skipperling  Colombia  M T S

Dalla granites (Mabille, 1898)  Granites Skipperling  TL: Ecuador

Dalla granites granites (Mabille, 1898)  Ecuador  M T

Dalla granites privata Draudt, 1923  Bolivia  M OI S

Dalla parma Evans, 1955  Parma Skipperling  Ecuador  M T S G

Dalla mars Evans, 1955  Mars Skipperling  Colombia  M T

Dalla cuadrada (Weeks, 1901)  Bolivian Skipperling  Bolivia  M T OD TI Txt

Dalla quadristriga (Mabille, 1889)  Quadristriga Skipperling  TL: Ecuador

Dalla quadristriga quadristriga (Mabille, 1889)  Colombia, Ecuador  M T S G

Dalla quadristriga regia Evans, 1955  Peru (Amazonas, Chachapoyas)  M T

Dalla quadristriga rota Evans, 1995  SE Peru  M T S

Dalla quadristriga rusa Evans, 1955  Bolivia  M T S

Dalla rubia Evans, 1955  Rubia Skipperling  Peru (Huancabamba)  M T

Dalla xicca (Dyar, 1913)  Xicca Skipperling  TL: Peru

Dalla xicca xicca (Dyar, 1913)  Peru  M T S

Dalla xicca paza Evans, 1955  Bolivia  M T S

Dalla curia Evans, 1955  Curious Skipperling  Colombia (Cordillera Oriental)  M T

Dalla semiargentea (C. Felder & R. Felder, 1867)  Half-silvered Skipperling  Colombia (Cordillera Oriental)  M T S

Dalla superargentea Viloria, A. Warren & Austin, 2008  Silvered Skipperling  Serrania de Valledupar, Sierra de Perija, Colombia-Venezuela  M T

Piruna  Evans, 1955

Piruna pirus (W. H. Edwards, 1878)  Russet Skipperling  SE ID, W & S WY, CO, UT, AZ, NM, (W TX?)  M T S L I Ha G

Piruna cyclosticta (Dyar, 1920)  Plateau Skipperling  Mexican Plateau to DF and MOR  M T S

Piruna haferniki H. Freeman, 1970  Chisos Skipperling  W TX to Hidalgo  M T S L I Ha

Piruna kemneri H. Freeman, 1990  Kemner's Skipperling  S Mexico (Oaxaca, Puebla)  M T L

Piruna dampfi (E. Bell, 1942)  Violet-dusted Skipperling  SW & Transvolcanic Mexico (GRO, MICH, MEX, VER)  M T S

Piruna gyrans (Plötz, 1884)  Variable Skipperling  W, C & S Mexico to Oaxaca  M T OI S L

Piruna ceracates (Hewitson, 1874)  Veracruz Skipperling  S Mexico (Veracruz & Oaxaca)  M S L

Piruna roeveri (L. Miller & J. Miller, 1972)  Roever's Skipperling  E & S Mexico (Coahuila to Oaxaca)  M T1 T2 S L Ha

Piruna brunnea (Scudder, 1872)  Chocolate Skipperling  E and S Mexico to Costa Rica  M S L

Piruna purepecha A. Warren & L. González, 1999  Purepecha Skipperling  SW Mexico (COL, GRO, JAL, MICH, MOR)  M T1 T2 S L

Piruna mullinsi H. Freeman, 1991  Mullins' Skipperling  S Mexico (Oaxaca, Puebla)  M T S L Ha

Piruna sina H. Freeman, 1970  Fine-spotted Skipperling  NW Mexico (DGO, SIN, SON)  M T S L

Piruna microsticta (Godman, 1900)  Southwest-Mexican Skipperling  [= ajijiciensis], SW Mexico (JAL to CHIS)  M T S L Ha

Piruna penaea (Dyar, 1918)  Hour-glass Skipperling  W Mexico to Oaxaca

Piruna penaea penaea (Dyar, 1918)  TL: Mexico (Gue)  M T S L Ha

Piruna penaea E Mexican segregate ,  S TX (one-time stray), E Mexico to Chiapas  M S L

Piruna aea (Dyar, 1912)  Many-spotted Skipperling  TL: Mexico (Gue/Oax)

Piruna aea aea (Dyar, 1912)  C & S Mexico to Guatemala  M T S L Ha

Piruna aea mexicana H. Freeman, 1979  [= cingo], NW Mexico to SE AZ  M T S L I

Piruna millerorum Steinhauser, 1991  Millers' Skipperling  NW Mexico (DGO, SIN, SON, CHIH)  M T S L

Piruna maculata H. Freeman, 1970  Sinaloan Skipperling  NW Mexico (Durango & Sinaloa)  M T S

Piruna polingii (W. Barnes, 1900)  Four-spotted Skipperling  C AZ, S NM to Guerrero and Puebla  M T S L1 L2 I Ha G

Piruna jonka Steinhauser, 1991  Oaxacan Skipperling  S Mexico (Oaxaca, Puebla)  M T S L

Dardarina  Evans, 1937

Dardarina dardaris (Hewitson, 1877)  Dardaris Skipperling  NW Mexico (SON) to Costa Rica  M T S L I Ha

Dardarina amadryas Hayward, 1942  Amadryas Skipperling  Brazil  M S

Dardarina tessellatus (Hayward, 1934)  Argentine Skipperling  Argentina  M S

Dardarina salta Hayward, 1937  Salta Skipperling  Argentina  M S

Dardarina daridaeus (Godman, 1900)  Daridaeus Skipperling  S Brazil, Paraguay, Argentina  M T S

Dardarina rana Evans, 1955  Rana Skipperling  S Brazil (Pr)  M S

Dardarina castra Evans, 1955  Castra Skipperling  S Brazil (Pr)  M S

Dardarina umuarama O. Mielke, 1978  Umuarama Skipperling  S Brazil (SP)  M T

Dardarina jonesi Evans, 1955  Jones' Skipperling  S Brazil (Pr)  M S

Dardarina angeloi O. Mielke & Casagrande, 2005  Angelo's Skipperling  SE & S Brazil (SP)

Dardarina angeloi angeloi O. Mielke & Casagrande, 2005  SE Brazil (SP)  M T

Dardarina angeloi machadoi O. Mielke & Casagrande, 2005  S Brazil (SC)  M T

Dardarina gaucha O. Mielke, 1980  Gaucha Skipperling  S Brazil (RS)  M T

Dardarina para O. Mielke, 1968  Mielke's Skipperling  S Brazil (MG)  M T

Dardarina aspila O. Mielke, 1966  Aspila Skipperling  S Brazil (Pr)  M T S

Butleria  W. F. Kirby, 1871

Butleria flavomaculatus (Blanchard, 1852)  Blanchard's Skipperling  TL: Chile

Butleria flavomaculatus flavomaculatus (Blanchard, 1852)  Chile (widespread)  M S

Butleria flavomaculatus tristriata Bryk, 1944  Chile (Valdivia, Osorno, Llanquihue)  M T

Butleria flavomaculatus valdivianus (Philippi, 1859)  Chile (Chiloe, Coyhaique)  M S

Butleria paniscoides (Blanchard, 1852)  Paniscoides Skipperling  TL: Chile

Butleria paniscoides paniscoides (Blanchard, 1852)  Chile (widespread)  M T S

Butleria paniscoides polyspilus (C. Felder, 1862)  Chile, Argentina (Neuquen, Rio Negro)  M T S

Butleria elwesi Evans, 1939  Elwes' Skipperling  Chile, Argentina (Neuquen, Rio Negro, Chubut)  M T S

Butleria quilla Evans, 1939  Quilla Skipperling  Chile (widespread), Argentina (Neuquen, Rio Negro)  M T S

Butleria fruticolens (A. Butler, 1881)  Butler's Skipperling  Chile, Argentina (Neuquen, Rio Negro)  M T S HS

Butleria sotoi Reed, 1877  Soto's Skipperling  Chile (Valparaiso, Cachapoal, Curico, Cauquenes, Linares, Nuble, Concepcion)  M T S

Butleria philippii (A. Butler, 1881)  Philipp's Skipperling  Chile (Nuble, Concepcion, Arauco, Araucania, Valdivia, Osorno)  M T

Butleria bissexguttatus (Philippi, 1859)  Herrera's Skipperling  Chile (widespread), Argentina (Neuquen, Rio Negro)  M S

Argopteron  E. Watson, 1893

Argopteron aureipennis (Blanchard, 1852)  Gold-plated Skipperling  Chile, Argentina  M T S IL L

Argopteron aureum L. Peña, 1968  Gold-washed Skipperling  Chile  M Ma

Argopteron puelmae (Calvert, 1888)  Gold-winged Skipperling  Chile  M S

  Subfamily Hesperiinae

  Tribe Incertae-sedis

Erionota  Mabille, 1878

Erionota thrax (Linnaeus, 1767)  Banana Skipper  TL: Java

Erionota thrax thrax (Linnaeus, 1767)  Introduced to Hawaii  M S IL

Perichares  Scudder, 1872

Perichares butus (Möschler, 1877)  TL: Surinam  M T S

Perichares philetes (Gmelin, [1790])  Caribbean Ruby-eye  Greater Antilles  M S Txt

Perichares poaceaphaga Burns, 2008  Grass-feeding Ruby-eye  Costa Rica  M T OD G

Perichares adela (Hewitson, 1867)  Green-backed Ruby-eye  S TX (uncommon), E & W Mexico to S Brazil  M T S L I G Txt

Perichares prestoeaphaga Burns, 2008  Prestoea-feeding Ruby-eye  Costa Rica  M T OD G

Perichares geonomaphaga Burns, 2008  Geonoma-feeding Ruby-eye  Costa Rica  M T OD G

Perichares limana Evans, 1955  TL: Peru  M Txt

Perichares aurina Evans, 1955  Evans' Ruby-eye  [Honduras?], S Brazil & Paraguay  M T S Txt

Perichares marmorata Scudder, 1872  TL: Venezuela  M S Txt

Perichares lotus (A. Butler, 1870)  Lotus Ruby-eye  SE Mexico S Brazil & Paraguay  M T S G

Perichares matha Evans, 1955  TL: Ecuador  M T S

Perichares deceptus (A. Butler & H. Druce, 1872)  Flasher Ruby-eye  TL: Costa Rica

Perichares deceptus deceptus (A. Butler & H. Druce, 1872)  El Salvador to Costa Rica  M T S L G

Perichares deceptus drina Evans, 1955  TL: Colombia  M T

Perichares deceptus quadrimaculata Röber, 1925  TL: Colombia

Perichares deceptus fulvimargo (A. Butler, 1873)  TL: Venezuela  M T S

Perichares deceptus luscinia (Plötz, 1882)  TL: Brazil (SC)  M T S L

Perichares colenda (Hewitson, 1866)  TL: Venezuela  M T S

Perichares chima Evans, 1955  TL: Ecuador  M T S

Perichares seneca (Latreille, [1824])  TL: Brazil

Perichares seneca crotona (Hewitson, 1866)  Panama, Venezuela  M T S

Perichares seneca seneca (Latreille, [1824])  TL: Brazil  M T S

Perichares forbesi Biezanko & O. Mielke, 1973  TL: Brazil (SP)  M T S

Orses  Godman, 1901

Orses cynisca (Swainson, 1821)  Yellow-edged Ruby-eye  E & W Mexico to S Brazil & Paraguay  M T OI S IL L

Orses itea (Swainson, 1821)  TL: Brazil (RJ)  M S

Alera  Mabille, 1891

Alera furcata Mabille, 1891  TL: Brazil (SP)  M T S

Alera haworthiana (Swainson, 1821)  Green-banded Ruby-eye  Panama to S Brazil  M T OI

Alera metallica (N. Riley, 1921)  TL: Brazil (SC)  M T

Alera manu O. Mielke & Casagrande, 2004  Peru, Brazil  M T OI S

Alera romeroi O. Mielke & Casagrande, 2004  Venezuela  M T OI

Alera vulpina (C. Felder & R. Felder, 1867)  TL: Colombia  M T S L

Lycas  Godman, 1901

Lycas argentea (Hewitson, 1866)  Silvered Ruby-eye  E & W Mexico to Argentina  M T S L

Lycas godart (Latreille, [1824])  TL: Brazil

Lycas godart godart (Latreille, [1824])  TL: Brazil  M T

Lycas godart boisduvalii (Ehrmann, 1909)  TL: Venezuela  M T S

Pyrrhopygopsis  Godman, 1901

Pyrrhopygopsis agaricon H. Druce, 1908  Agaricon Skipper  TL: Colombia

Pyrrhopygopsis agaricon agaricon H. Druce, 1908  E Costa Rica, Colombia  M T S

Pyrrhopygopsis agaricon nanta Evans, 1955  TL: Ecuador  M T

Pyrrhopygopsis socrates (Ménétriés, 1855)  TL: Brazil (MG)

Pyrrhopygopsis socrates orasus (H. Druce, 1876)  TL: Peru  M T OI S L

Pyrrhopygopsis socrates crates Mabille & Boullet, 1912  TL: Bolivia; [Ecuador]; Peru  M T S IL

Pyrrhopygopsis socrates socrates (Ménétriés, 1855)  TL: Brazil (MG)  M T S

Pyrrhopygopsis romula (H. Druce, 1875)  TL: Colombia

Pyrrhopygopsis romula romula (H. Druce, 1875)  TL: Colombia  M T S

Pyrrhopygopsis romula tenebricosa (Hewitson, 1876)  TL: Peru  M T IL

Pyrrhopygopsis romula lugubris H. Druce, 1908  TL: Bolivia  M T S G

Pyrrhopygopsis quispica (Plötz, 1886)  TL: Peru  M T S IL

Pseudosarbia  Berg, 1897

Pseudosarbia flavofasciata Skinner, 1921  TL: Brazil (MT)  M S

Pseudosarbia phoenicicola Berg, 1897  TL: Argentina  M S

  Tribe Megathymini

Megathymus  Scudder, 1872

Megathymus yuccae (Boisduval & Le Conte, [1837])  Yucca Giant-Skipper  TL: USA (SC)

Megathymus yuccae yuccae (Boisduval & Le Conte, [1837])  [= buchholzi], SE USA, E of Mississippi River  M T1 T2 OI S L1 L2 I Ha

Megathymus yuccae reinthali H. Freeman, 1963  [= kendalli], C to NE TX, C & E OK, SW AR, N & C LA  M T1 T2 S L I1 I2 I3 F Ha

Megathymus yuccae wilsonorum D. Stallings & J. Turner, 1958  Tamaulipas & far S TX (lower Rio Grande Valley)  M T S L I

Megathymus yuccae louiseae H. Freeman, 1963  S-C TX (Val Verde, Kinney Cos) to NE Mexico (COAH, NL)  M T S

Megathymus yuccae coloradensis C. Riley, 1877  SE WY, W NE, W KS, E CO, E NM, W OK, TX panhandle  M T1 T2 T3 T4 S1 S2 L I H

Megathymus yuccae reubeni D. Stallings, J. Turner & V. Stallings, 1963  Hueco Mts., Hudspeth Co., TX, to N Chihuahua  M T S

Megathymus yuccae navajo Skinner, 1911  [= arizonae], W CO, S UT, W NM, AZ, N Sonora  M T1 T2 S Ma L1 L2 I1 I2 Ha

Megathymus yuccae martini D. Stallings & J. Turner, 1956  Mohave Desert, S CA, W NV, far NW AZ  M T S L1 L2 Ha

Megathymus yuccae harbisoni J. Emmel & T. Emmel, 1998  Beaumont, Riverside Co. CA, S to Baja California Norte  M T S Ha

Megathymus beulahae D. Stallings & J. Turner, 1958  Broad-banded Giant-Skipper  TL: Mexico (Hid)

Megathymus beulahae gayleae D. Stallings, J. Turner & V. Stallings, 1963  COAH, NL, TAMPS, SLP  M T1 T2 S L

Megathymus beulahae beulahae D. Stallings & J. Turner, 1958  Hidalgo  M T1 T2 S

Megathymus ursus Poling, 1902  Ursine Giant-Skipper  TL: USA (Az)

Megathymus ursus ursus Poling, 1902  S AZ (Pinal, Pima, Sta. Cruz, Cochise), SW NM (Hidalgo), SON  M T OI S L I Ha

Megathymus ursus deserti R. Wielgus, J. Wielgus & D. Wielgus, 1972  E Maricopa Co., AZ  M T

Megathymus ursus violae D. Stallings & J. Turner, 1956  C & S NM (E of Rio Grande), W TX, COAH, NL  M T S L I Ha

Megathymus streckeri (Skinner, 1895)  Strecker's Giant-Skipper  TL: USA (Az)

Megathymus streckeri streckeri (Skinner, 1895)  NE AZ, NW NM, SW CO, SE UT  M T S L I Ha

Megathymus streckeri S Navajo Co., Arizona segregate ,  S end White Mts., AZ  M S L Ha

Megathymus streckeri leussleri W. Holland, 1931  TX panhandle & E NM, to E MT, W ND, (SE SASK?)  M T S1 S2 L F Ha

Megathymus streckeri texana W. Barnes & McDunnough, 1912  S-C Texas  M T1 T2 L I1 I2 H

Megathymus cofaqui (Strecker, 1876)  Cofaqui Giant-Skipper  TL: USA (Ga)

Megathymus cofaqui cofaqui (Strecker, 1876)  [= harrisi], Georgia to NC  M T TI S L I1 I2 Ha

Megathymus cofaqui slotteni Gatrelle, 1999  Florida  M T S L I H

Stallingsia  H. Freeman, 1959

Stallingsia maculosus (H. Freeman, 1955)  Manfreda Giant-Skipper  S TX, N NL  M T1 T2 S1 S2 S3 S4 H1 H2 H3

Stallingsia smithi (H. Druce, 1896)  Smith's Giant-Skipper  SW MX (JAL, MICH, GRO, DF, MOR)  M T OI S L

Stallingsia jacki D. Stallings, J. Turner & V. Stallings, 1963  Chiapan Giant-Skipper  Chiapas  M T S

Aegiale  C. Felder & R. Felder, 1860

Aegiale hesperiaris (Walker, 1856)  Tequila Giant-Skipper; Gusano de Maguey  much of N & C Mexico  M T S1 S2 S3 S4 S5 IL L

Turnerina  H. Freeman, 1959

Turnerina mejicanus (E. Bell, 1938)  Turner's Giant-Skipper  Durango  M T G

Turnerina hazelae (D. Stallings & J. Turner, 1958)  Guerrero Giant-Skipper  Guerrero & Oaxaca  M T S

Agathymus  H. Freeman, 1959

Agathymus neumoegeni (W. H. Edwards, 1882)  Neumoegen’s Giant-Skipper  TL: USA (Az)

Agathymus neumoegeni neumoegeni (W. H. Edwards, 1882)  C AZ (Yavapai Co.) to W-C NM (Catron Co.)  M T S L

Agathymus neumoegeni S NM segregate ,  S NM: Organ Mts. (Dona Ana-Otero Cos.) to Sacramento Mts.  M S F Ha

Agathymus neumoegeni carlsbadensis (D. Stallings & J. Turner, 1957)  Guadalupe Mts. (Eddy-Otero Cos., NM; Culberson Co., TX)  M T S

Agathymus neumoegeni judithae (D. Stallings & J. Turner, 1957)  Hueco Mts., Hudspeth Co., TX  M T S

Agathymus neumoegeni diabloensis H. Freeman, 1962  Sierra Diablo, Sierra Blanca, Hudspeth Co., TX  M T S

Agathymus neumoegeni florenceae (D. Stallings & J. Turner, 1957)  Davis Mts., Jeff Davis Co., TX  M T S F Ha

Agathymus neumoegeni mcalpinei (H. Freeman, 1955)  E side of Glass Mts., Brewster Co., TX  M T S F Ha

Agathymus chisosensis (H. Freeman, 1952)  Neumoegen’s Giant-Skipper  Chisos Mts., S Brewster Co., TX  M T S

Agathymus juliae (D. Stallings & J. Turner, 1958)  Julia’s Giant-Skipper  Durango  M T S

Agathymus hoffmanni (H. Freeman, 1952)  Hoffmann’s Giant-Skipper  S-C Mexico (DF, GTO, HGO, MEX, SLP)  M T S

Agathymus polingi (Skinner, 1905)  Poling's Giant-Skipper  S AZ, SW NM (Hidalgo Co.), NE Sonora  M T S L1 L2 I H Ha

Agathymus evansi (H. Freeman, 1950)  Huachuca Giant-Skipper  Huachuca Mts., Cochise Co., AZ  M T S HS L1 L2 H

Agathymus aryxna (Dyar, 1905)  Arizona Giant-Skipper  SE AZ, SW NM, NE SON  M T S HS L

Agathymus baueri (D. Stallings & J. Turner, 1954)  Bauer's Giant-Skipper  TL: USA (Az)

Agathymus baueri baueri (D. Stallings & J. Turner, 1954)  C AZ: E Yavapai, W Gila, NE Maricopa, N Pinal Cos.  M T S L H

Agathymus baueri freemani D. Stallings & J. Turner, 1960  W-C AZ: SW Yavapai Co., S Mohave Co.  M T S

Agathymus gentryi Roever, 1998  Gentry's Giant-Skipper  SW AZ (W Pima, C Maricopa, W Pinal, W Yuma), E SB Co., CA  M S

Agathymus belli (H. Freeman, 1955)  Bell's Giant-Skipper  Chihuahua, Durango  M T S

Agathymus fieldi H. Freeman, 1960  Field's Giant-Skipper  W Mexico: JAL, SON  M T S

Agathymus ricei D. Stallings, J. Turner & V. Stallings, 1966  Rice's Giant-Skipper  Puebla  M T S1 S2

Agathymus indecisa (A. Butler & H. Druce, 1872)  Guatemalan Giant-Skipper  Chiapas to Costa Rica  M T S

Agathymus mariae (W. Barnes & Benjamin, 1924)  Mary's Giant-Skipper  TL: USA (Tx)

Agathymus mariae mariae (W. Barnes & Benjamin, 1924)  S NM, trans-Pecos counties, TX  M T S L

Agathymus mariae chinatiensis H. Freeman, 1964  Chinati Mts., Presidio Co., TX  M T S

Agathymus mariae lajitaensis H. Freeman, 1964  S Presidio Co., TX  M T S

Agathymus mariae rindgei H. Freeman, 1964  Val Verde and Kinney Cos., TX  M T S Ha

Agathymus gilberti H. Freeman, 1964  Gilbert's Giant-Skipper  Brewster, Terrell and Val Verde Cos., TX  M T

Agathymus micheneri D. Stallings, J. Turner & V. Stallings, 1961  Michener’s Giant-Skipper  Coahuila  M T S L

Agathymus remingtoni (D. Stallings & J. Turner, 1958)  Remington's Giant-Skipper  E Mexico: HGO, SLP, TAMPS  M T S

Agathymus estelleae (D. Stallings & J. Turner, 1958)  Estelle's Giant-Skipper  TL: Mexico (NL)

Agathymus estelleae valverdiensis H. Freeman, 1966  S-C TX (Terrell, Val Verde, Edwards, Kinney Cos.)  M T S

Agathymus estelleae estelleae (D. Stallings & J. Turner, 1958)  Coahuila, Nuevo Leon  M T S

Agathymus escalantei D. Stallings, J. Turner & V. Stallings, 1966  Escalante's Giant-Skipper  Guerrero  M T Ha

Agathymus stephensi (Skinner, 1912)  California Giant-Skipper  S CA (Riverside, E San Diego, W Imperial Cos.), Baja (N)  M T S L1 L2 I H1 H2

Agathymus comstocki (Harbison, 1957)  Comstock's Giant-Skipper  Baja California (N)  M T1 T2 L1 L2 H Ha

Agathymus dawsoni Harbison, 1963  Dawson’s Giant-Skipper  Baja California (N)  M T1 T2 S L1 L2 H Ha

Agathymus [n. sp.] Roever, MS  Undescribed Giant-Skipper  Baja California (N)  M L H

Agathymus alliae (D. Stallings & J. Turner, 1957)  Mojave Giant-Skipper  TL: USA (Az)

Agathymus alliae alliae (D. Stallings & J. Turner, 1957)  N Coconino Co., AZ  M T S

Agathymus alliae paiute Roever, 1998  SE CA (SB, Inyo), S NV (Clark), SW UT (Wa), NW AZ  M S L Ha

Agathymus rethon (Dyar, 1913)  Black Giant-Skipper  Mexico (COL, GRO, JAL, MICH, MOR, NAY, OAX, PUE, VER), El Salvador  M T S L

  Tribe Thymelicini

Ancyloxypha  C. Felder, 1862

Ancyloxypha arene (W. H. Edwards, 1871)  Tropical Least Skipper  SE AZ, SW NM, W & S TX, S to Costa Rica  M T S L I H G

Ancyloxypha aurea Hayward, 1940  TL: Ecuador  M T

Ancyloxypha dryas Hayward, 1942  TL: Argentina

Ancyloxypha melanoneura C. Felder & R. Felder, 1867  TL: Colombia  M T

Ancyloxypha nitedula (Burmeister, 1878)  TL: Argentina  M S

Ancyloxypha numitor (Fabricius, 1793)  Least Skipper  Great Plains, eastward; SE SASK to Quebec, to S TX, far NE MEX  M T S L1 L2 L3 L4 L5 L6 Ha G

Ancyloxypha ramba Evans, 1955  TL: Ecuador  M T

Oarisma  Scudder, 1872

Oarisma poweshiek (Parker, 1870)  Poweshiek Skipperling  SE MAN, E ND, NE SD, W & S MN, N-central IA, SE WI, S MI  M S G

Oarisma garita (Reakirt, 1866)  Garita Skipperling  TL: USA (Co)

Oarisma garita garita (Reakirt, 1866)  SE BC to S MAN (plus Peace Riv.), S through Rockies to AZ & NM  M T S L1 L2 L3 Ha G

Oarisma garita calega (Godman, 1900)  NW (DGO) to S (OAX) Mexico  M T S

Oarisma edwardsii (W. Barnes, 1897)  Edwards’ Skipperling  S-central CO, NM, C & E AZ, NW Mexico to S-central Mexico  M T S L I G

Oarisma era Dyar, 1927  Bold-veined Skipperling  NW & NE to central Mexico  M T S Ha

Oarisma boeta (Hewitson, 1870)  TL: Ecuador  M T OI L

Oarisma nanus (Herrich-Schäffer, 1865)  Nanus Skipperling  Cuba & Isla de Juventud  M T S L

Oarisma bruneri E. Bell, 1959  Bruner's Skipperling  Cuba  M T

Copaeodes  Speyer, 1877

Copaeodes aurantiaca (Hewitson, 1868)  Orange Skipperling  S CA & NV to E TX (strays N), S to Guatemala  M T S L1 L2 I F G

Copaeodes minima (W. H. Edwards, 1870)  Southern Skipperling  SE AZ, C TX to FL & NC, E & W Mexico to Costa Rica  M T S L1 L2 L3 L4 I1 I2 G

Copaeodes stillmani E. Bell & W. Comstock, 1948  Stillman's Skipperling  Hispaniola including Isla Catalina  M T S

Copaeodes eoa D. Smith, L. Miller & McKenzie, 1991  Anegada Skipperling  Anegada  M OI

Copaeodes jean Evans, 1955  TL: Guyana

Copaeodes jean jean Evans, 1955  TL: Guyana  M T S

Copaeodes jean favor Evans, 1955  TL: Brazil (Pr)  M T S

Copaeodes castanea O. Mielke, 1969  TL: Brazil (Pr)  M T S

Adopaeoides  Godman, 1900

Adopaeoides prittwitzi (Plötz, 1884)  Sunrise Skipper  SE AZ, SW NM, W TX, S to S-central Mexico  M S L I Ha

Adopaeoides bistriata Godman, 1900  TL: Mexico (Dur)  M T

Thymelicus  Hübner, [1819]

Thymelicus lineola (Ochsenheimer, 1808)  European Skipper  TL: Germany

Thymelicus lineola lineola (Ochsenheimer, 1808)  introduced; Newfoundland to VA & TN, W to MAN, S Rocky Mts.  M S L1 L2 I Ha G

  Tribe Calpodini

Turesis  Godman, 1901

Turesis complanula (Herrich-Schäffer, 1869)  Complanula Skipper  SE Mexico to S Brazil  M T S HS L

Turesis tabascoensis H. Freeman, 1979  Tabasco Skipper  SE Mexico  M T

Turesis theste Godman, 1901  Theste Skipper  SE Mexico to Costa Rica  M T S

Turesis basta Evans, 1955  TL: Brazil (Pa)  M T

Turesis osibius Draudt, 1924  TL: ?

Ebusus  Evans, 1955

Ebusus ebusus (Cramer, 1780)  TL: Surinam

Ebusus ebusus ebusus (Cramer, 1780)  TL: Surinam  M T

Ebusus ebusus nigrior L. Miller, 1985  SE Mexico (Veracruz and Chiapas)  M T

Evansiella  Hayward, 1948

Evansiella cordela (Plötz, 1882)  TL: ?  M T S

Argon  Evans, 1955

Argon lota (Hewitson, 1877)  Argon Skipper  [= argus], W & S Mexico to Argentina  M T S L

Cobaloides  Hayward, 1939

Cobaloides argus Hayward, 1939  TL: Paraguay

Sacrator  Evans, 1955

Sacrator polites (Godman & Salvin, 1879)  TL: Colombia

Sacrator polites polites (Godman & Salvin, 1879)  TL: Colombia  M T

Sacrator polites pilla Evans, 1955  TL: Peru  M S L

Sacrator sacrator (Godman & Salvin, 1879)  TL: Colombia  M T L G

Megaleas  Godman, 1901

Megaleas syrna (Godman & Salvin, 1879)  Megaleas Skipper  Costa Rica  M T G

Megaleas angelae Henao & Andrade-C, 2013  Ángela's Megaleas Skipper  SW Colombia

Megaleas chocoensis Salazar & Constantino, 2013  Choco Megaleas Skipper  Choco region in Colombia  M T

Lychnuchus  Hübner, [1831]

Lychnuchus brasta Evans, 1955  TL: Peru

Lychnuchus celsus (Fabricius, 1793)  TL: India  M S

Lychnuchus pelta Evans, 1955  TL: Ecuador  M S

Lychnuchus victa Evans, 1955  TL: Peru  M S

Talides  Hübner, [1819]

Talides sergestus (Cramer, 1775)  Sergestus Ruby-eye  SE Mexico to S Brazil  M S L

Talides cantra Evans, 1955  Cantra Ruby-eye  S Mexico to Colombia, Venezuela & W Ecuador  M S L

Talides sinois Hübner, [1819]  TL: not specified  M T

Talides riosa Evans, 1955  TL: Brazil (RJ)  M S

Talides alternata E. Bell, 1941  Alternate Ruby-eye  S Mexico to S Brazil  M T S L

Talides hispa Evans, 1955  Hispa Skipper  Costa Rica to W Ecuador, W Brazil  M S IL

Tromba  Evans, 1955

Tromba tromba Evans, 1955  TL: Peru  M S

Tromba xanthura (Godman, 1901)  Yellow-washed Ruby-eye  E & W Mexico to Colombia  M T S L

Nyctus  Mabille, 1891

Nyctus crinitus Mabille, 1891  TL: Peru  M T

Nyctus hiarbas (Cramer, 1775)  TL: Surinam  M S

Turmada  Evans, 1955

Turmada camposa (Plötz, 1886)  TL: Brazil  M S

Turmada turmada (H. Druce, 1912) &