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Chlosyne gabbii atrifasciata Hawks & J. Emmel, 1998
(Gabb’s Checkerspot)

Distribution and Larval Foodplants:
 

Asteraceae: Corethrogyne filaginifolia var. bernardina (Abrams) Hall, Hazardia squarrosa (Hook. & Arn.) Greene, Haplopappus squarrosus squarrosus Hook. and Arn., Orthocarpus purpurascens var. purpurascens Benth.

Synonymy

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne gabbii (Behr, 1863)

 

Chlosyne gabbii gabbii (Behr, 1863)

Proc. Calif. Acad. Sci. [1]3(6): 88, 89, no. 3.

Original Combination: M[elitaea]. Gabbii

Type Locality: “California” implied from the title of the paper, “the mountains near Los Angeles” from the description; neotype from “La Tuna Canyon, Verdugo Mountains, Los Angeles County, California.”

Types: Type(s) destroyed in the 1906 San Francisco earthquake and fire; neotype in CAS, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (6): 99, figs. 19 ♂ D, 20 ♂ V.

                   =    pola (Boisduval, 1869)

Annls. Soc. Entomol. Belg. 12: 56, no. 44.

Original Combination: Melitaea Pola

Type Locality: “Sonora”; suggested to be “somewhere from Los Angeles southward into extreme northern Baja California” by F. Brown (1967), J. Lepid. Soc. 21(4): 273; defined as “La Tuna Canyon, 1,200' elev., Verdugo Mts., Los Angeles County, California” by Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 32.

Types: Holotype in CMNH, as determined by F. Brown (1965), J. Lepid. Soc. 19(4): 206-207, fig. 2 ♂ D&V; figured by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 32, figs. 243 ♂ D, 244 ♂ V.

Boisduval loaned the holotype to W. H. Edwards, but died before it could be returned. The subsequent figure of a specimen of Poladryas arachne by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 261, fig. 2188, and designation of that figure as the type of “pola” by W. Barnes and McDunnough, (1916), Contr. Nat. Hist. Lepid. N. Am. 3(2): 92, contributed to the general acceptance of this name as a senior synonym of arachne (e.g., dos Passos (1964), Mem. Lepid. Soc. (1): 85, and Bauer (1961), in: P. Ehrlich and A. Ehrlich, How Know Butts.: 130). F. Brown (1965), J. Lepid. Soc. 19(4): 206-207, detailed the particulars of this confusion and figured the holotype of pola, justifying its placement in the synonymy of C. gabbii.

                   =    sonorae (Boisduval, 1869)

Annls. Soc. Entomol. Belg. 12: 56-57, no. 45; figured by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 261, figs. 2189 ♂ D&V, 2190 ♀ D&V.

Original Combination: Melitaea Sonorae

Type Locality: “De la Sonora”; suggested to be “somewhere from Los Angeles southward into extreme northern Baja California” by F. Brown (1967), J. Lepid. Soc. 21(4): 273; stated to be “vic. Los Angeles, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 157; further defined as “La Tuna Canyon, 1,200' elev., Verdugo Mts., Los Angeles County, California” by Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 32.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 32, figs. 246 ♂ D, 247 ♂ V.

                         =    ‡”pasadenae” (Gunder, 1924)

Entomol. News 35(5): 155, pl. 2, fig. C ♂ D (“holotype”).

Original Combination: Euphydryas gabbi ab. ♂ pasadenae

Type Locality: “Pasadena, Los Angeles County, California”

Types: “Holotype” in AMNH; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 4 ♂ D.

                         =    ‡”gunderi” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 33; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 6 ♂ D (“holotype”).

Original Combination: Melitaea gabbii ab. gunderi

Type Locality: “San Fernando, Calif.” [Los Angeles County, California]

Types: “Holotype” in LACM.

                         =    ‡”newcombi” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 33; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 5 ♀ D (“holotype”).

Original Combination: Melitaea gabbii ab. newcombi

Type Locality: “Pasadena, Calif.” [Los Angeles County, California]

Types: “Holotype” in LACM.

 

Chlosyne gabbii atrifasciata Hawks & J. Emmel, 1998

Syst. W. N. Am. Butts. (25): 322-323, figs. 5 ♂ D, 6 ♂ V (holotype); 7 ♀ D, 8 ♀ V.

Original Combination: Chlosyne (Charidryas) gabbii atrifasciata

Type Locality: “California: Santa Barbara County, Santa Cruz Island; La Cascada”

Types: Holotype in LACM.

Bibliography

Davenport, Kenneth E. 2004a. A concise update of the information provided in The Butterflies of southern California (1973) by Thomas C. Emmel and John F. Emmel. Taxonomic Report of the International Lepidoptera Survey 4(7): 1-23 {10 May 2004} 

Hawks, David and John F. Emmel. 1998. 25. New subspecies of Cercyonis sthenele (Boisduval) (Lepidoptera: Satyridae) and Chlosyne (Charidryas) gabbii (Behr) (Nymphalidae) from the California Islands. Pp. 321-326, 8 figs., in: T. C. Emmel (Ed.), Systematics of western North American Butterflies. Gainesville; Mariposa Press; {[18] Dec 1998}

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