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Chlosyne leanira alma (Strecker, [1878])
(Leanira Checkerspot)

Type Specimens
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Pinned Specimens
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Live Adults photo collection
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Live Adults photo collection
Page 2 (intergrades):

Live Adults photo collection
Page 3 (courtship):

Immatures photo collection
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Immatures photo collection
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Larval Foodplants
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Habitats
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Distribution and Larval Foodplants:
 

Scrophulariaceae: Castilleja applegatei ssp. martinii  (Abrams) Chuang & Heckard

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne leanira (C. Felder & R. Felder, 1860)

Wien. Entomol. Montaschr. 4(4): 106, no. 64; figured by C. Felder and R. Felder (1867), Reise Novara Zool. (3): pl. 50, figs. 13 ♂ D, 14 ♂ V.

Original Combination: Melitaea Leanira

Type Locality: “Californien”; suggested to be “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 153; corrected to “Hwy 70 at Chambers Creek, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 89.

Types: Lectotype in BMNH, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 89, figs. 13 ♂ D, 14 ♂ V.

=    daviesi (Wind, 1947)

Pan-Pac. Entomol. 23(4): 171.

Original Combination: Melitaea leanira daviesi

Type Locality: “Strawberry Lake, Tuolumne County, California, el. 5500 ft.”

Types: Holotype in CAS.

 

Chlosyne leanira alma (Strecker, [1878])

Lepid. Rhop. Het. (15): 135, pl. 15 ♂ D&V.

Original Combination: Melitaea Alma

Type Locality: “Arizona...S. Utah”; defined as “vic. St. George, Washington Co., Utah” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 153; redefined as “northwestern Mohave County” by Austin and M. Smith (1998), Syst. W. N. Am. Butts. (27): 340. Lectotype is labeled “Arizona.”

Types: Lectotype in FMNH, designated by Austin and M. Smith (1998), Syst. W. N. Am. Butts. (27): 340.

                         =    ‡”koebelei” (Gunder, 1927)

Entomol. News 38(5): 136, no. 7, pl. 2, fig. 7 ♀ V (“holotype”).

Original Combination: Lemonias alma koebelei ♀ nov. tr. form

Type Locality: “Argus Mts., Inyo County, California”

Types: “Holotype” in CAS.

Bibliography

Austin, George T. and Anna T. Austin. 1981. Butterflies of Clark County, Nevada. Journal of Research on the Lepidoptera 19(1): 1-63, 5 tbls. {Spring, 1980(81), 1 Dec 1981} 

Austin, George T. and Michael J. Smith. 1998. 27. Revision of the Thessalia leanira complex (Lepidoptera: Nymphalidae: Melitaeinae): Thessalia leanira (C. & R. Felder), with descriptions of four new subspecies. Pp. 333-358, 67 figs., in: T. C. Emmel (Ed.), Systematics of western North American Butterflies. Gainesville; Mariposa Press. {[18] Dec 1998} 

Barnes, William and James Halliday McDunnough. 1913. Illustrations of rare and typical Lepidoptera. Contributions to the Natural History of the Lepidoptera of North America 2(1): 1-44, pls 1-21. {10 Mar 1913} 

Clemence, Victor L. 1912. Notes on Melitaea alma Strecker. Canadian Entomologist 44(4): 102, pl. 3. {9 Apr 1912} 

Coolidge, Karl R. 1910. Melitaea alma Strecker, and its synonymy. Canadian Entomologist 42(12): 425-431. {15 Dec 1910} 

Comstock, John Adams. 1927. Butterflies of California. A popular guide to a knowledge of the butterflies of California embracing all of the 477 Species and varieties at present recorded from the state. Author, Los Angeles, California. 334 pp., 63 pls. {1927} 

Davenport, Kenneth E. 2004a. A concise update of the information provided in The Butterflies of southern California (1973) by Thomas C. Emmel and John F. Emmel. Taxonomic Report of the International Lepidoptera Survey 4(7): 1-23. {10 May 2004} 

Davenport, Kenneth E. 2004b. The Yosemite butterflies. The Taxonomic Report of the International Lepidoptera Survey 5(1): 1-75, figs. {29 Dec 2004} 

Davenport, Kenneth E., Norbert G. Kondla, Charles Grisham and C. Howard Grisham. 2007. The Yosemite butterflies: color plates. The Taxonomic Report of the International Lepidoptera Survey 5(2): 1-83, color figs. {15 Mar 2007} 

Emmel, Thomas Chadbourne and John F. Emmel. 1973b. The butterflies of southern California. Natural History Museum of Los Angeles County, Science Series 26: xi + 148 pp., 10 pls., 77 figs. {30 Nov 1973} 

Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figs. {contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford}

Gunder, Jean Daniel. 1927. New transition forms or "abs" and their classification (Lepid., Rhopalocera). Entomological News 38(5): 129-138, pl. 2 {[4] May 1927} [description of Lemonias alma koebelei & nov. transition form] 

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Miller, Lee Denmar and Frederick Martin Brown. 1981. A catalogue/checklist of the butterflies of America north of Mexico. Memoirs of the Lepidopterists= Society (2): vii + 280 pp. 

Scott, James A. 1992. Hostplant records for butterflies and skippers (mostly from Colorado) 1959-1991, with new life histories and notes on oviposition, immatures, and ecology. Papilio (n.s.) 6: 1-171, 10 tbls. {5 Jul 1992} 

Scott, James A. 2006. Chlosyne leanira/fulvia classification. p. 37, in: J. Scott (Ed.), Taxonomic studies and new taxa of North American butterflies. Papilio (New Series) 12. {28 Apr 2006} 

Seitz, Adalbert. 1913. 3. Gattung Melitaea. In: A. Seitz (Ed.), Gross-Schmetterlinge der Erde 5(182): 429-432; (187): 433-434; pl. 88. {1913} 

Strecker, Ferdinand Heinrich Herman. 1874. 15. Descriptions and (19) figures of sixteen species and varieties of Lep., two diurnals (one new); two (new) Sphingidae; eleven Bombycidae (two species and five varieties new). Pp. 135-139, in: Lepidoptera, rhopaloceres and heteroceres, indigenous and exotic; with descriptions and colored illustrations. Reading, Pennsylvania; Private. {Nov 1877, [16 Jul 1878]} [description of Melitaea Alma]

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