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Chlosyne leanira wrightii (W. H. Edwards, 1886)
(Leanira Checkerspot)

Type Specimens
photo collection:


 

Pinned Specimens
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Live Adults photo collection
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Live Adults photo collection
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Immatures
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Larval Foodplants and Habitats
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Distribution and Larval Foodplants:
 

Larval foodplants
Scrophulariaceae: Castilleja foliolosa  Hook. & Arn., Castilleja applegatei ssp. martinii  (Abrams) Chuang & Heckard

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne leanira (C. Felder & R. Felder, 1860)

Wien. Entomol. Montaschr. 4(4): 106, no. 64; figured by C. Felder and R. Felder (1867), Reise Novara Zool. (3): pl. 50, figs. 13 ♂ D, 14 ♂ V.

Original Combination: Melitaea Leanira

Type Locality: “Californien”; suggested to be “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 153; corrected to “Hwy 70 at Chambers Creek, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 89.

Types: Lectotype in BMNH, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 89, figs. 13 ♂ D, 14 ♂ V.

=    daviesi (Wind, 1947)

Pan-Pac. Entomol. 23(4): 171.

Original Combination: Melitaea leanira daviesi

Type Locality: “Strawberry Lake, Tuolumne County, California, el. 5500 ft.”

Types: Holotype in CAS.

 

Chlosyne leanira wrightii (W. H. Edwards, 1886)

Can. Entomol. 18(4): 64-65, no. 3.

Original Combination: Melitaea Wrightii

Type Locality: “San Bernardino” [San Bernardino County, California]

Types: Lectotype in CMNH, designated by F. Brown (1966), Trans. Am. Entomol. Soc. 92(3): 389, fig. 11 ♂ D&V; figured by W. Holland (1898), Butt. Book: pl. 17, figs. 9 ♂ D (lectotype), 10 ♀ V (paralectotype).

                   =    cerrita (W. G. Wright, 1905)

Butts. W. Coast: 55, no. 189; 161, no. 189, pl. 20, figs. 189, 189a ♂ D&V (lectotype), 189b ♀ V.

Original Combination: [Melitaea] Cerrita; Melitaea Cerrita n. s.

Type Locality: “Southern California”; suggested to be “edge of Mojave Desert” by Bauer (1975), in: Howe, Butts. N. Am.: 168.

Types: Lectotype (#4306) in CAS, designated by Tilden (1985), Occ. Pap. Calif. Acad. Sci. (118): 22.

Austin and Murphy [1989], J. Res. Lepid. 26(1-4): 280-281, determined this synonymy.

                   =    pelona (Gunder, 1930)

Bull. South. Calif. Acad. Sci. 29(2): 61, no. 71.

Original Combination: Melitaea wrightii form pelona

Type Locality: “Mint Canyon” [Sierra Pelona Valley, Los Angeles County, California]

Types: Holotype possibly in LACM; figured by Comstock (1927), Butts. Calif.: pl. 39, fig. 16 ♀ V.

                         =    ‡”carolynae” (Gunder, 1926)

Entomol. News 37(1): 3, no. 3, pl. 1, fig. 3 ♂ D (“holotype”).

Original Combination: Melitaea wrighti ab. ♂ carolynae

Type Locality: “Mint Canyon, Los Angeles County, California” [Sierra Pelona Valley]

Types: “Holotype” in AMNH.

Bibliography

Austin, George T. and Dennis M. Murphy. 1989. Book Review: A field guide to western butterflies. Tilden J. W. and A. C. Smith, 1986. Journal of Research on the Lepidoptera 26(1-4): 278-283. {Autumn, 1987(1988), 2 Mar 1989} 

Austin, George T. and Michael J. Smith. 1998. 27. Revision of the Thessalia leanira complex (Lepidoptera: Nymphalidae: Melitaeinae): Thessalia leanira (C. & R. Felder), with descriptions of four new subspecies. Pp. 333-358, 67 figs., in: T. C. Emmel (Ed.), Systematics of western North American Butterflies. Gainesville; Mariposa Press. {[18] Dec 1998} 

Bauer, David L. 1975. Tribe Melitaeini. Pp. 139-195, in: W. H. Howe (ed.) The butterflies of North America. Garden City, New York, Doubleday & Co.: xiii + 633 pp., 97 pls., 32 tf. {[7 Nov] 1975} 

Brown, Frederick Martin. 1966. The types of the nymphalid butterflies described by William Henry Edwards--Part II, Melitaeinae. Transactions of the American Entomological Society 92(3): 357-468, 34 figs. {Sep, 20 Oct 1966} 

Comstock, John Adams. 1927. Butterflies of California. A popular guide to a knowledge of the butterflies of California embracing all of the 477 Species and varieties at present recorded from the state. Author, Los Angeles, California. 334 pp., 63 pls. {1927} 

Comstock, John Adams and Charles M. Dammers. 1932. Early stages of Melitaea leanira wrightii Edw. and Calephelis nemesis Edw. (Lepidoptera). Bulletin of the Southern California Academy of Sciences 31(1): 9-15, 5 pls. {Jan-Apr, 30 May 1932} 

Davenport, Kenneth E. 1983. Geographic distribution and checklist of the butterflies of Kern County, California. Journal of the Lepidopterists= Society 37(1): 46-69. {19 Aug 1983} 

Davenport, Kenneth E. 2004a. A concise update of the information provided in The Butterflies of southern California (1973) by Thomas C. Emmel and John F. Emmel. Taxonomic Report of the International Lepidoptera Survey 4(7): 1-23. {10 May 2004} 

Edwards, William Henry. 1886. Description of new species of butterflies found in the United States. Canadian Entomologist 18(4): 61-65. {21 Apr 1886} [description of Melitaea Wrightii]

Emmel, Thomas Chadbourne and John F. Emmel. 1973b. The butterflies of southern California. Natural History Museum of Los Angeles County, Science Series 26: xi + 148 pp., 10 pls., 77 figs. {30 Nov 1973} 

Gunder, Jean Daniel. 1926. Several new aberrant Lepidoptera (Rhopalocera). Entomological News 37(1): 1-9, pl. 1. {[7] Jan 1926} [description of Melitaea wrighti ab. carolynae]

Gunder, Jeane Daniel. 1930. Butterflies of Los Angeles County, California. Bulletin of the Southern California Academy of Science 29(2): 39-95, pls. 18-23. {May-Aug 1930, 15 Aug 1930} [description of Melitaea wrightii form pelona

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Hoffmann, Carlos Christian. 1940. Catálogo systemático y zoogeográphico de los Lepidópteros Mexicanos, Primera parte. Papilionoidea. Anales del Instituto de Biología. Universidad Nacional de México 11(2): 639-739, 2 maps. {[after 28 Dec] 1940} 

Holland, William Jacob. 1898. The butterfly book; a popular guide to a knowledge of the butterflies of North America. Garden City, New York, Doubleday & McClure Co.  xx + 382 pp., 48 pls., 183 + [1] figs. {1898}

Holland, William Jacob. 1931. The butterfly book. New and thoroughly revised edition.  A popular and scientific manual, describing and depicting all the butterflies of the United States and Canada. Garden City, New York; Doubleday, Doran & Co.  xii + 424 pp., 77 pls., 198 + [1] text figures. {1931}

Seitz, Adalbert. 1913. 3. Gattung Melitaea. In: A. Seitz (Ed.), Gross-Schmetterlinge der Erde 5(182): 429-432; (187): 433-434; pl. 88 {1913} 

Tilden, James William. 1975. An analysis of the W. G. Wright butterfly and skipper plesiotypes in the collection of the California Academy of Sciences. Occasional Papers of the California Academy of Sciences (118): 1-44. {16 Jan 1975} 

Wright, William Greenwood. 1905. Butterflies of the West Coast of the United States. San Bernardino, California: 257 pp., 31 pls. {1905} [description of Melitaea Cerrita]

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