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Chlosyne n. nycteis (E. Doubleday, [1847])
(Silvery Checkerspot)

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Live Adults photo collection Page 1:

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Distribution and Larval Foodplants:
 

Asteraceae: Aster  L., Conyza canadensis (L.) Cronq., Helianthus annuus L., Rudbeckia laciniata L., Verbesina alternifolia (L.) Britt. ex Kearney, Verbesina helianthoides Michx.

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne nycteis (E. Doubleday, 1847)

 

Chlosyne nycteis nycteis (E. Doubleday, 1847)

In: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3 ♀ D; (21): 181, no. 23 (1848).

Original Combination: Melitaea Nycteis

Type Locality: “United States (Middle States)”

Types: Syntype (#Rh8433) in BMNH.

                   =    oenone (Scudder, 1863)

Proc. Essex Inst. 3: 166, no. 35.

Original Combination: Melitaea Oenone

Type Locality: “Lenox and Williamstown, Mass....Norway, Me.” [Berkshire County, Massachusetts; Oxford County, Maine]

Types: Syntypes possibly in MCZ.

                         =    ‡”millburni” (C. Rummel, 1926)

Bull. Brooklyn Entomol. Soc. 21(5): 203.

Original Combination: Phyciodes nycteis ab. millburni

Type Locality: “Millburn, N. J.” [Union County, New Jersey]

Types: “Holotype” in AMNH.

                         =    ‡”lacteus” (Gunder, 1928)

Can. Entomol. 60(7): 166, pl. B, figs. 16 ♂ D, 16a ♂ V (“holotype”).

Original Combination: Phyciodes nycteis tr. f. lacteus

Type Locality: “near St. Louis, Mo.” [St. Louis County, Missouri]

Types: “Holotype” in AMNH.

                         =    ‡”greyi” (W. D. Field, 1934)

Can. Entomol. 66(11): 256.

Original Combination: Phyciodes nycteis tr. f. greyi

Type Locality: “north west edge of city limits of Lawrence” [Douglas County, Kansas]

Types: “Holotype” in USNM.

Bibliography

Aaron, Charles E. 1884. On the caterpillars of North American Papilionidae and Nymphalidae. Papilio 4(5/[6]): 83-91. {May-Jun [22 Jul] 1884} cited in Papilio 4(9/10): 189. [Translated from Gruber, August, 1884, Jena Zeitschrift für naturwissenschaft 17 (N.F. 10)

Beutenmüller, William. 1893. Article XVI.-Descriptive catalogue of the butterflies found within fifty miles of New York City, together with a brief account of their life histories and habits. Bulletin of the American Museum of Natural History 4(16): 241-310, 5 pls. {Oct 1893} 

Burns, John McLauren, Donald Lafontaine, Paul A. Opler (chair), Robert K. Robbins, Felix Sperling. 2000. Precis of first meeting. News of the Lepidopterists= Society 42(1): 9, 13, 23. {Spring, 15 Mar 2000} 

Calhoun, John V. 2003. The history and true identity of Melitaea ismeria (Nymphalidae): a remarkable tale of duplication, misinterpretation, and presumption. Journal of the Lepidopterist=s Society 57(3): 204-219, 25 figs. {29 Sep 2003}  

Calhoun, John V. 2004. Histoire Générale et Iconographie des Lépidoptères et des Chenilles de l=Amérique Septentrionale by Boisduval & Le Conte (1829-[1837]): original drawings used for the engraved plates and the true identities of four figured taxa. Journal of the Lepidopterists= Society 58(3): 143-168, 45 figs., 2 tbls. {27 Oct 2004}

Doubleday, Edward. 1846-1852. The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habits and transformations, and a catalogue of the species of each genus; Illustrated with 86 Plates by W. C. Hewitson. London; Longman, Brown, Green and Longmans: xii + ii + 534 pp.; 86 pls.  2 Volumes in 54 parts. [description of Melitaea Nycteis

Edwards, William Henry. 1870. Rearing butterflies from the egg. Canadian Entomologist 2(12): 162-164. {Dec 1870} 

Edwards, William Henry. 1873. Notes on the early stages of some of our butterflies. Canadian Entomologist, 5(12): 223-225. {Dec 1873} 

Edwards, William Henry. 1875. Notes on butterflies. Canadian Entomologist, 7(8): 150-151. {Aug 1875} 

Edwards, William Henry. 1879. Description of preparatory stages of Phyciodes nycteis. Canadian Entomologist, 11(6): 101-105. {Jun 1879} 

Field, William Dewitt. 1934. On the naming of "transition forms" in Lepidoptera with notes on certain forms captured near Lawrence, Kansas. Canadian Entomologist 66(11): 253-257. {Nov, 1 Dec 1934} [description of Phyciodes nycteis tr. f. greyi]

Forbes, William Trowbridge Merrifield. 1945. The genus Phyciodes (Lepidoptera, Nymphalinae). Entomologica Americana (n.s.)24: 139-207. {Oct 1944, 24 May 1945} 

French, George Hazen. 1886. The butterflies of the eastern United States. Philadelphia, J. B. Lippencott Co.: 408 pp., 93 figs. {1886} 

Gatrelle, Ronald R. 1998. The rediscovery, taxonomy, and biology of Chlosyne gorgone gorgone and Chlosyne ismeria (Nymphalidae) in Burke County, Georgia. The Taxonomic Report of the International Lepidoptera Survey 1(2): 1-8, 7 figs. {1 Nov 1998} 

Gunder, Jean Daniel. 1926. Several new aberrant Lepidoptera (Rhopalocera). Entomological News 37(1): 1-9, pl. 1 {[7] Jan 1926} [description of Phyciodes nycteis transition form lacteus

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Layberry, Ross A., Peter W. Hall, J. Donald Lafontaine (plates by John T. Fowler). 1998. The butterflies of Canada. Toronto, Buffalo & London; University of Toronto Press: 280 pp., 32 pls., 294 maps and illus. {1998} 

Lintner, Joseph Albert. 1872. Entomological contributions- No. 2.  III. Notes on Melitaea Nycteis (Doubl). Pp. 158-159, in: From the twenty-third annual report of the New York State Cabinet of Natural History for the year 1869.  Appendix D. Entomological contributions.  By J. A. Lintner.  Printed in Advance of the Report. Albany; Weed, Parsons and Company, Printers: [4] + 5-90 pp., 8 pls.; pages of Annual Report (published later) and Contributions on each page {1872} 

Maynard, Charles Johnson. 1886. The butterflies of New England; with original descriptions of one hundred and six species, accompanied by eight lithographic plates, in which are given at least two hand-colored figures of each species. Boston, Massachusetts, C. J. Maynard [Bradlee Whidden]: iv + [2] + 66 + [2] pp.; 8 pls. {Feb 1886} 

Riley, Charles Valentine. 1875. On the summer dormancy of the larva of Phyciodes nycteis (Doubleday), with remarks on the natural history of the species. Proceedings of the American Association for the Advancement of Science 23(2B): 108-112. {1874, 1875} 

Röber, Johannes Karl Max 1913-1914. Gattung Phyciodes. In: A. Seitz (Ed.), Die Gross-Schmetterlinge der Erde 5(162): pl. 92 {5 Jun 1913}; (183): 434-440, pl. 90 {10 Dec 1913}; (188): 449-450, pls. 88, 89. {[before 10 Feb] 1914}; (189): pl. 91. {14 Apr 1914} 

Rummel, Charles. 1919. Observations on Physiodes [sic] nycteis. Bulletin of the Brooklyn entomological Society 14(2): 47. {18 Apr 1919}

Rummel, Charles. 1926. Notes on aberrations of New Jersey butterflies. Bulletin of the Brooklyn entomological Society 21(5): 203. {14 Dec 1926} [description of Phyciodes nycteis ab. milburni

Scott, James A. 2006. Chlosyne nycteis drusius X nycteis blend zone. p. 37, in: J. Scott (Ed.), Taxonomic studies and new taxa of North American butterflies. Papilio (New Series) 12 {28 Apr 2006} 

Scudder, Samuel Hubbard. 1863. A list of the butterflies of New England. Proceedings of the Essex Institute 3: 161-179 {pre-print 30 Apr 1863; article [26 Apr] 1864} [description of Melitaea Oenone

Scudder, Samuel Hubbard. 1888-1889. The butterflies of the eastern United States and Canada with special reference to New England. Cambridge; Samuel Hubbard Scudder, 1(introduction, Nymphalidae): xxiv +  766 pp., frtspc. (colored); 2(Lycaenidae, Papilionidae, Hesperiidae): xii + 767-1774, frtspc. (colored), 1 map; 3(appendix, plates): i-viii, 1775-1958, frtspc., pls. 1-89 (14 colored), 3 maps {[Nov 1888], [1Apr] 1889}

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