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Chlosyne nycteis drusius (W. H. Edwards, 1884)
(Silvery Checkerspot)

Pinned Specimens photo collection Page 1:

Pinned Specimens photo collection Page 2:

Live Adults photo collection Page 1:

Live Adults photo collection Page 2:

Larval Foodplants and Habitats photo collection:

Distribution and Larval Foodplants:
 

Asteraceae: Aster  L., Helianthus L. sp., Rudbeckia laciniata var. ampla (A. Nels.) Cronq., Verbesina  L.

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne nycteis (E. Doubleday, 1847)

In: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3 ♀ D; (21): 181, no. 23 (1848).

Original Combination: Melitaea Nycteis

Type Locality: “United States (Middle States)”

Types: Syntype (#Rh8433) in BMNH.

=    oenone (Scudder, 1863)

Proc. Essex Inst. 3: 166, no. 35.

Original Combination: Melitaea Oenone

Type Locality: “Lenox and Williamstown, Mass....Norway, Me.” [Berkshire County, Massachusetts; Oxford County, Maine]

Types: Syntypes possibly in MCZ.

=    ‡”millburni” (C. Rummel, 1926)

Bull. Brooklyn Entomol. Soc. 21(5): 203.

Original Combination: Phyciodes nycteis ab. millburni

Type Locality: “Millburn, N. J.” [Union County, New Jersey]

Types: “Holotype” in AMNH.

=    ‡”lacteus” (Gunder, 1928)

Can. Entomol. 60(7): 166, pl. B, figs. 16 ♂ D, 16a ♂ V (“holotype”).

Original Combination: Phyciodes nycteis tr. f. lacteus

Type Locality: “near St. Louis, Mo.” [St. Louis County, Missouri]

Types: “Holotype” in AMNH.

=    ‡”greyi” (W. D. Field, 1934)

Can. Entomol. 66(11): 256.

Original Combination: Phyciodes nycteis tr. f. greyi

Type Locality: “north west edge of city limits of Lawrence” [Douglas County, Kansas]

Types: “Holotype” in USNM.

 

Chlosyne nycteis drusius (W. H. Edwards, 1884)

Papilio 4(3): 57.

Original Combination: Phyciodes Nycteis var. Drusius

Type Locality: “Colorado and Arizona”; defined as “Turkey Creek Junction, Jefferson County, Colorado” by F. Brown (1966), Trans. Am. Entomol. Soc. 92(3): 401-402.

Types: Lectotype in CMNH, designated by F. Brown (1966), Trans. Am. Entomol. Soc. 92(3): 402, fig. 15 ♂ D&V.

                         =    ‡”hewitsoni” (W. D. Field, 1936)

J. Entomol. & Zool. 28(2): 23.

Original Combination: Phyciodes nycteis race drusius tr. f. hewitsoni

Type Locality: “Greaterville, Arizona” [Pima County]

Types: “Holotype” in USNM.

Bibliography

Brown, Frederick Martin. 1957. Colorado butterflies [Assisted by Donald Eff and Bernard Rotger.] Denver, Denver Museum Natural History: vii + 368 pp.; 314 unnumbered figs. {1957} 

Brown, Frederick Martin. 1966. The types of the nymphalid butterflies described by William Henry Edwards--Part II, Melitaeinae. Transactions of the American Entomological Society 92(3): 357-468, 34 figs. {Sep, 20 Oct 1966} 

Edwards, William Henry. 1884. Descriptions of new species of butterflies from Arizona. Papilio 4(3): 53-58. {[27] Mar 1884; cited in Papilio 4(4): 82} [description of Phyciodes Nycteis var. Drusius

Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figures [contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford] {1981} 

Field, William Dewitt. 1936. New North American Rhopalocera. Journal of Entomology and Zoology 28(2): 17-26. {Jun 1936} [description of Phyciodes nycteis race drusius tr. f. hewitsoni

Fisher, Michael S. 2006. The butterflies of Colorado. Nymphalidae ‑ Part 3. The subfamily Nymphalinae and relatives. The checkerspots, crescentspots and the true nymphs. Fort Collins; Lepidoptera of North America 7.3; Contributions of the C. P. Gillette Museum of Arthropod Diversity, Colorado State University: vi + 120 pp., photos {30 Sep 2006} 

Forbes, William Trowbridge Merrifield. 1945. The genus Phyciodes (Lepidoptera, Nymphalinae). Entomoogica Americana (n.s.)24: 139-207. {Oct 1944, 24 May 1945} 

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Scott, James A. 1992. Hostplant records for butterflies and skippers (mostly from Colorado) 1959-1991, with new life histories and notes on oviposition, immatures, and ecology. Papilio (n.s.) 6: 1-171, 10 tables {5 Jul 1992} 

Scott, James A. 2006. Chlosyne nycteis drusius X nycteis blend zone. p. 37, in: J. Scott (Ed.), Taxonomic studies and new taxa of North American butterflies. Papilio (New Series) 12 {28 Apr 2006} 

Scott, James A. 2006. Butterfly hostplant records, 1992-2005, with a treatise on the evolution of Erynnis, and a note on new terminology for mate-locating behavior. Papilio (New Series) 14: 74 pp., cartoon {28 Apr 2006}

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