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Chlosyne p. palla (Boisduval, 1852)
(Northern Checkerspot)

Pinned Specimens photo collection:

Live Adults photo collection Page 1:

Live Adults photo collection Page 2:

Larval Foodplants and Habitats photo collection:

Distribution and Larval Foodplants:
 

Asteraceae: Eurybia conspicua  (Lindl.) Nesom, Eurybia radulina (Gray) Nesom, Senecio triangularis Hook., Solidago californica Nutt., Symphyotrichum spathulatum var. spathulatum (Lindl.) Nesom 

Scrophulariaceae: Castilleja breviflora ?

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne palla (Boisduval, 1852)

 

Chlosyne palla palla (Boisduval, 1852)

Annls. Soc. Entomol. Fr. (2)10(2): 305, no. 48; figured by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 259, figs. 2173 ♂ D&V, 2174 ♀ D&V.

Original Combination: Melitaea palla

Type Locality: “Dans une grande partie de la Californie”; defined as “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 156; corrected to “Hwy. 70 at Chambers Creek, ca. 6 rd. Mi SW of Belden, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18, figs. 99 ♂ D, 100 ♂ V.

                         =    ‡”wardi” (Oberthür, 1914)

Étud. Lépid. Comp. 9(2): 79, pl. 259, fig. 2175 ♂ D&V (“holotype”).

Original Combination: [Melitaea Palla] aberration Wardi

Type Locality: Not stated; “Californie” implied from the title of the paper.

Types: “Holotype” in USNM; figured by J. A. Comstock (1927), Butts. Calif.: pl. 27, fig. 2 ♂ D&V.

                         =    ‡”stygiana” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 32; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 19 ♀ D (“holotype”).

Original Combination: Melitaea palla ab. stygiana

Type Locality: “Willow Ranch, Modoc County, Calif.” [California]

Types: “Holotype” in LACM.

Bibliography

Barnes, William and James Halliday McDunnough. 1916. Notes on North American diurnal Lepidoptera. Contributions to the Natural History of the Lepidoptera of North America 3(2): 49-156, 8 pls. {15 Dec 1916} 

Boisduval, Jean Baptiste Alphonse d'Echauffour. 1852. Lépidoptères de la Californie. Annales de la Société entomologique de France (2)10(2): 275-324 {[Aug] 1852} [description of Melitaea palla

Boisduval, Jean Baptiste Alphonse d'Echauffour. 1868-1869. Lépidoptères de la Californie. Annales de la Société entomologique de Belgique 12: 1-28 (1868); 37-94 ([1 Nov] 1869) [missing pp. 29-37 through printer's error] {1868-1869} 

Britten, Hugh B., Peter F. Brussard and Dennis D. Murphy. 1996. Isozyme data and the taxonomy of checkerspot butterflies (Euphydryas ). Journal of Research on the Lepidoptera 32: 124-134, 1 fig., 4 tbls. {1993, 1 May 1995} 

Comstock, John Adams. 1926. Studies in Pacific Coast Lepidoptera (Continued). Thirteen new species or aberrations of California butterflies. Bulletin of the Southern California Academy of Sciences 25(1): 29-34, 48. {Jan-Apr, 30 Mar 1926} [description of Melitaea palla ab. stygiana

Comstock, John Adams. 1927. Butterflies of California. A popular guide to a knowledge of the butterflies of California embracing all of the 477 species and varieties at present recorded from the state. Author, Los Angeles, California. 334 pp., 63 pls. {1927} 

Davenport, Ken. 1983. Geographic distribution and checklist of the butterflies of Kern County, California. Journal of the Lepidopterists= Society 37(1): 46-69. {19 Aug 1983} 

Davenport, Kenneth E. 2004. The Yosemite butterflies. The Taxonomic Report of the International Lepidoptera Survey 5(1): 1-75, figs. {29 Dec 2004} 

Davenport, Kenneth E., Norbert G. Kondla, Charles Grisham and C. Howard Grisham. 2007. The Yosemite butterflies: color plates. The Taxonomic Report of the International Lepidoptera Survey 5(2): 1-83, color figs. {15 Mar 2007} 

Edwards, Henry. 1873. Pacific Coast Lepidoptera, No. 2: on the transformation of the diurnal Lepidoptera of California and the adjacent districts. Proceedings of the California Academy of Sciences 5(11): 161-172 {Nov 1873} 

Emmel, John F., Thomas Chadbourne Emmel and Sterling O. Mattoon. 1998a. 2. The types of California butterflies named by Jean Alphonse Boisduval: designation of lectotypes and a neotype, and fixation of type localities. Pp. 3-76, 269 figs., in: T. C. Emmel (Ed.), Systematics of western North American butterflies. Gainesville; Mariposa Press; {[18] Dec 1998}

Emmel, John F., Thomas Chadbourne Emmel and Sterling O. Mattoon. 1998b. 12. New subspecies of Nymphalidae (Lepidoptera) from California and a neotype designation for Argynnis rupestris Behr (Lepidoptera: Nymphalidae). Pp. 139-158, 50 figs., in: T. C. Emmel (Ed.), Systematics of western North American Butterflies. Gainesville; Mariposa Press; {[18] Dec 1998} 

Emmel, J. F., O. Shields & D. E. Breedlove. 1971. Larval foodplant records for North American Rhopalocera. Part 2. Journal of Research on the Lepidoptera 9(4):233-242. {Dec 1970(1971), [13 Dec] 1971} 

Gunder, Jeane Daniel. 1930. Butterflies of Los Angeles County, California. Bulletin of the Southern California Academy of Science 29(2): 39-95, pls. 18-23 {May-Aug 1930, 15 Aug 1930} 

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Miller, Lee Denmar and Frederick Martin Brown. 1981. A catalogue/checklist of the butterflies of America north of Mexico. Memoir.  The Lepidopterists= Society (2): vii + 280 pp. {1981} 

Oberthür, Charles. 1914. VII. Les Lépidoptères de la Californie par le Docteur Boisduval. Études de Lépidopterologie comp. 9(2): 73-90; pls. 258-264. {May 1914} [description of Melitaea Palla aberration Wardi

Seitz, Adalbert. 1913. 3. Gattung Melitaea. In: A. Seitz (Ed.), Gross-Schmetterlinge der Erde 5(182): 429-432; (187): 433-434; pl. 88 {1913}

Warren, Andrew D. 2005. Butterflies of Oregon: their taxonomy, distribution, and biology. Fort Collins; Contributions of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University: 405 pp., 3 maps. {15 Mar 2005}

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