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Chlosyne palla calydon (W. Holland, 1931)
(Northern Checkerspot)

Pinned Specimens photo collection Page 1:

Pinned Specimens photo collection Page 2:

Distribution and Larval Foodplants:
 

Asteraceae: Aster L., Erigeron speciosus var. macranthus (Nutt.) Cronq.

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne palla (Boisduval, 1852)

Annls. Soc. Entomol. Fr. (2)10(2): 305, no. 48; figured by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 259, figs. 2173 ♂ D&V, 2174 ♀ D&V.

Original Combination: Melitaea palla

Type Locality: “Dans une grande partie de la Californie”; defined as “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 156; corrected to “Hwy. 70 at Chambers Creek, ca. 6 rd. Mi SW of Belden, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18, figs. 99 ♂ D, 100 ♂ V.

=    ‡”wardi” (Oberthür, 1914)

Étud. Lépid. Comp. 9(2): 79, pl. 259, fig. 2175 ♂ D&V (“holotype”).

Original Combination: [Melitaea Palla] aberration Wardi

Type Locality: Not stated; “Californie” implied from the title of the paper.

Types: “Holotype” in USNM; figured by J. A. Comstock (1927), Butts. Calif.: pl. 27, fig. 2 ♂ D&V.

=    ‡”stygiana” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 32; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 19 ♀ D (“holotype”).

Original Combination: Melitaea palla ab. stygiana

Type Locality: “Willow Ranch, Modoc County, Calif.” [California]

Types: “Holotype” in LACM.

 

Chlosyne palla calydon (W. Holland, 1931)

Butt. Book (rev. ed.): 125, pl. 58, figs. 19 ♂ D, 20 ♀ D (syntypes).

Original Combination: Melitaea calydon

Type Locality: “Turkey Creek Junction, Colorado” [Jefferson County]

Types: Syntypes in CMNH.

This name was listed but not described by Mead (1875), Rep. Coll. diurn. Lepid. 5(Zool.): 760, and Strecker (1878), Butts. & Moths N. Am., Comp. Syn. Cat.: 126; on both occasions it is a nomen nudum.

Bibliography

Barnes, William and Foster Hendrickson Benjamin. 1925. Notes on two butterfly names omitted from recent lists. Entomological News 36(4): 118-119. {Apr, [30 Mar] 1925} 

Brown, Frederick Martin. 1957. Colorado butterflies [Assisted by Donald Eff and Bernard Rotger.] Denver, Denver Museum Natural History: vii + 368 pp.; 314 unnumbered figs. {1957} 

Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figures [contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford] {1981} 

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Holland, William Jacob. 1931. The butterfly book. New and thoroughly revised edition.  A popular and scientific manual, describing and depicting all the butterflies of the United States and Canada. Garden City, New York; Doubleday, Doran & Co.  xii + 424 pp., 77 pls., 198 + [1] text figures. {1931} [description of Melitaea calydon

Mead, Theodore Luqueer. 1875. Chapter VIII. Report upon the collections of diurnal Lepidoptera made in portions of Nevada, Utah, Colorado, New Mexico, and Arizona during the years 1871, 1872, 1873, and 1874, with notes upon all species known to inhabit Colorado. Pp. 739-791, pls. 35-40, in: George M. Wheeler (and A. A. Humphreys), Report upon Geological Explorations and Surveys West of the One Hundredth Meridian. Washington, D.C.; Government Printing Office; Engineer Dept., U. S. Army: [212] pp., [8] leaves of pls. {1875} 

Schrier, Rachel D., Michael J. Cullenward, Paul R. Ehrlich & Raymond R. White. 1976. The structure and genetics of a montane population of the checkerspot butterfly, Chlosyne palla. Oecologia (Berlin) 25(3): 279-289. {Sep 1976} 

Scott, James A. 1992. Hostplant records for butterflies and skippers (mostly from Colorado) 1959-1991, with new life histories and notes on oviposition, immatures, and ecology. Papilio (n.s.) 6: 1-171, 10 tables {5 Jul 1992} 

Scott, James A. 2006. Butterfly hostplant records, 1992-2005, with a treatise on the evolution of Erynnis, and a note on new terminology for mate-locating behavior. Papilio (New Series) 14: 74 pp., cartoon {28 Apr 2006}

Strecker, Ferdinand Heinrich Herman. 1878. Butterflies and moths of North America, with full instructions for collecting, breeding, preparing, classifying, packing for shipment, etc. A complete synonymical catalogue of Macrolepidoptera, with a full bibliography, to which is added a glossary of terms and an alphabetical and descriptive list of localities. Reading; B. F. Owen: [4] + ii + 283 pp., 2 pls. {[Oct] 1878}

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