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Chlosyne palla eremita (W. G. Wright, 1905)
(Northern Checkerspot)

Type Specimens photo collection:

Pinned Specimens photo collection:

Live Adults photo collection Page 1:

Live Adults photo collection Page 2:

Larval Foodplants and Habitats photo collection:

Distribution and Larval Foodplants:
 

Asteraceae:

Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

      =    ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

      =    ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

      =    ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

=    Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

=    Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

      =    ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

=    Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

 

Chlosyne palla (Boisduval, 1852)

Annls. Soc. Entomol. Fr. (2)10(2): 305, no. 48; figured by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 259, figs. 2173 ♂ D&V, 2174 ♀ D&V.

Original Combination: Melitaea palla

Type Locality: “Dans une grande partie de la Californie”; defined as “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 156; corrected to “Hwy. 70 at Chambers Creek, ca. 6 rd. Mi SW of Belden, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18, figs. 99 ♂ D, 100 ♂ V.

=    ‡”wardi” (Oberthür, 1914)

Étud. Lépid. Comp. 9(2): 79, pl. 259, fig. 2175 ♂ D&V (“holotype”).

Original Combination: [Melitaea Palla] aberration Wardi

Type Locality: Not stated; “Californie” implied from the title of the paper.

Types: “Holotype” in USNM; figured by J. A. Comstock (1927), Butts. Calif.: pl. 27, fig. 2 ♂ D&V.

=    ‡”stygiana” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 32; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 19 ♀ D (“holotype”).

Original Combination: Melitaea palla ab. stygiana

Type Locality: “Willow Ranch, Modoc County, Calif.” [California]

Types: “Holotype” in LACM.

 

Chlosyne palla eremita (W. G. Wright, 1905)

Butts. W. Coast: 55, no. 178; 157, no. 178, pl. 19, figs. 178 ♀ D (lectotype), 178b ♀ D, 178c ♀ V.

Original Combination: [Melitaea] Eremita; Melitaea Eremita n. s.

Type Locality: “Lake County, California”; defined as “San Rafael, Marin County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (12): 141, but the original citation is adequate. Populations in Lake County are phenotypically like those in Marin County.

Types: Lectotype (#4296) in CAS, designated by Tilden (1975), Occ. Pap. Calif. Acad. Sci. (118): 21.

J. Emmel et al. (1998), Syst. W. N. Am. Butts. (12): 141, and A. Warren (2005), Butts. Ore.: 227, discussed the status of this taxon.

                         =    ‡”hemifusa” (Gunn, 1930)

Entomol. News 41(1): 17.

Original Combination: Melitaea palla ab. hemifusa

Type Locality: “Cazadero, Sonoma County, California”

Types: “Holotype” in CIS.

Bibliography

Comstock, John Adams. 1927. Butterflies of California. A popular guide to a knowledge of the butterflies of California embracing all of the 477 Species and varieties at present recorded from the state. Author, Los Angeles, California. 334 pp., 63 pls. 

Emmel, John F., Thomas Chadbourne Emmel and Sterling O. Mattoon. 1998. 12. New subspecies of Nymphalidae (Lepidoptera) from California and a neotype designation for Argynnis rupestris Behr (Lepidoptera: Nymphalidae). Pp. 139-158, 50 figs., in: T. C. Emmel (Ed.), Systematics of western North American Butterflies. Gainesville; Mariposa Press; {[18] Dec 1998} 

Gunn, Norman R. 1930. A new butterfly (Lepid.: Nymphalidae). Entomological News 41(1): 17. {[16] Jan 1930} [description of Melitaea palla ab. hemifusa

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Seitz, Adalbert. 1913. 3. Gattung Melitaea. In: A. Seitz (Ed.), Gross-Schmetterlinge der Erde 5(182): 429-432; (187): 433-434; pl. 88 {1913} 

Shapiro, Arthur Maurice, Cheryl A. Palm and Karen L. Wcislo. 1981. The ecology and biogeography of the butterflies of the Trinity Alps and Mount Eddy, northern California. Journal of Research on the Lepidoptera 18(2): 69-151. {Summer 1979(81), 31 Jan 1981} 

Tilden, James William. 1975. An analysis of the W. G. Wright butterfly and skipper plesiotypes in the collection of the California Academy of Sciences. Occasional Papers of the California Academy of Sciences (118): 1-44. {16 Jan 1975}

Warren, Andrew D. 2005. Butterflies of Oregon: their taxonomy, distribution, and biology. Fort Collins; Contributions of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University: 405 pp., 3 maps. {15 Mar 2005}

Wright, William Greenwood. 1905. Butterflies of the West Coast of the United States. San Bernardino, California: 257 pp., 31 pls. {1905} [description of Melitaea Eremita]

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