Main Introduction Page Electronic Reference Library Know-how Glossary of acronyms and other terms used on this website A Catalog of the Butterflies of the United States and Canada Illustrated Checklist of Neotropical Butterflies Support the Butterflies of America Foundation Interactive Listing of American Butterflies Photographer Credits Learn about contributing your photos Citation for this website Contact us
Papilionidae (Swallowtails)Pieridae (Whites and Sulphurs)Lycaenidae (Gossamerwings)Riodinidae (Metalmarks)Nymphalidae (Brushfoots)Hesperiidae (Skippers)Links to external lepidoptera websites

Lycaena xanthoides nigromaculata J. Emmel & Pratt, 1998
(Great Copper)

Type Specimens photo collection:


Pinned Specimens photo collection Page 1:

Pinned Specimens photo collection Page 2:

Live Adults photo collection:

Distribution and Larval Foodplants:



(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Lycaena [Fabricius], 1807

In: Illiger Mag. f. Insektenk. 6: 285, no. 32. Type-species: Papilio phlaeas Linnaeus, 1761, Fauna Svecica (ed. 2): 285, no. 1078, by designation of J. Curtis (1824), Brit. Entomol. 1(3): pl. 12, [1]. American species in this genus were reviewed by Pratt and D. Wright (2003), Pan-Pac. Entomol. 78(4): 219-229.


Oliver & Shapiro (2007), Molecular Ecology 16(20): 4308–4320, establish that Lycaena editha and L. xanthoides are distinct species.


Lycaena xanthoides (Boisduval, 1852)

Annls. Soc. Entomol. Fr. (2)10(2): 292-293, no. 26; figured by Oberthür (1913), Étud. Lépid. Comp. 9(1): pl. 236, fig. 1931 ♂ D&V, 1932 ♀ D&V.

Original Combination: Polyommatus xanthoides

Type Locality: “Montagnes de la Californie”; defined as “Sacramento, Sacramento County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 12.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 12, figs. 42 ♂ D, 43 ♂ V.

=    luctuosa (F. Watson & W. Comstock, 1920)

Bull. Am. Mus. Nat. Hist. 42(10): 453-454.

Original Combination: Heodes xanthoides luctuosa

Type Locality: “Tehachapi, California” [Kern County]

Types: Holotype in AMNH.

=    ‡”gunderi” Rudkin, 1933

Entomol. News 44(4): 97.

Original Combination: Lycaena xanthoides tr. f. gunderi

Type Locality: “Bouquet Canyon, Los Angeles Co., California”

Types: “Holotype” in CAS.


Lycaena xanthoides nigromaculata J. Emmel & Pratt, 1998

Syst. W. N. Am. Butts. (52): 674, figs. 41 ♂ D, 42 V ♂ V (holotype); 43 ♀ D, 44 ♀ V.

Original Combination: Lycaena xanthoides nigromaculata

Type Locality: “California: Colusa County; junction of Highway 45 and Highway 20, west of Meridian”

Types: Holotype in LACM.



Top of PageMain PageReference Library Literature ListCitation Interactive Listing

This website is supported by Butterflies of America Foundation, a U.S. registered 501(c)(3) tax-deductible nonprofit 170(b)(1)(A)(vi) public charity.