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Phyciodes cocyta pascoensis W. G. Wright, 1905
(Northern Crescent)

Type Specimens photo collection:

Pinned Specimens photo collection:

Distribution and Larval Foodplants:

Asteraceae: Eucephalus ledophyllus var. ledophyllus  (Gray) Greene, Symphyotrichum frondosum (Nutt.) Nesom


(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Phyciodes Hübner, [1819]

Verz. bekannt. Schmett. (2): 29. Type-species: Papilio cocyta Cramer, 1777, Uitl. Kapellen 2(9): 7, pl. 101, figs. A, B; (16): 148 (index), by designation of Scudder (1872), 4th Ann. Rep. Peabody Acad. Sci. (1871): 46. The arrangement of species in this genus follows Wahlberg et al. (2003), Syst. Entomol. 28(2): 257-273.


Phyciodes cocyta (Cramer, 1777)

Uitl. Kapellen 2(9): 7, pl. 101, figs. A ♂ D, B ♂ V (fig. C ♀ V is not the same species); (16): 148 (index).

Original Combination: Pap[ilio]. Nymph[alis]. Phaler[ata]. Cocyta

Type Locality: Not stated for figs. A and B, “Surinamen” “Suriname” for fig. C; neotype from “Black Rock, Cape Breton, Nova Scotia.”

Types: Neotype in AMNH, designated by Scott (1994), Papilio (n.s.) 7: 36, fig. 111 ♂ D.

=    ‡morpheus (Fabricius, 1775)

Syst. Entomol.: 530, no. 370.

Original Combination: P[apilio]. P[lebejus]. R[uralis]. Morpheus

Type Locality: “America boreali”

Types: Type(s) probably lost.

Preoccupied by Papilio morpheus Pallas, 1771, Reise verschied. Prov. Russisich. Reichs 1, Anhang: 471, no. 64. Fabricius (1793), Entomol. Syst. 3(1): 155-156, no. 479, cited this name in association with Papilio cocyta Cramer, 1777.

=    arctica dos Passos, 1935

Can. Entomol. 67(4): 87.

Original Combination: Phyciodes tharos arctica

Type Locality: “Port au Port, Table Mountain, Newfoundland” [Newfoundland and Labrador]

Types: Holotype in AMNH.


Phyciodes cocyta pascoensis W. G. Wright, 1905

Butts. W. Coast: 55, no. 198; 165, no. 198, pl. 21, figs. 198 ♂ D (lectotype), 198a ♂ V.

Original Combination: [Phyciodes Nycteis var.] Pascoensis; Phyciodes Pascoensis n. s.

Type Locality: “Eastern Oregon”; “Pasco, Franklin County, Washington” implied from the formation of the name.

Types: Lectotype (#4308) in CAS, designated by Tilden (1975), Occ. Pap. Calif. Acad. Sci. (118): 23.


Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figures [contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford] {1981} 

Forbes, William Trowbridge Merrifield. 1945. The genus Phyciodes (Lepidoptera, Nymphalinae). Entomologica Americana (n.s.)24: 139-207. {Oct 1944, 24 May 1945} 

Tilden, James William. 1975. An analysis of the W. G. Wright butterfly and skipper plesiotypes in the collection of the California Academy of Sciences. Occasional Papers of the California Academy of Sciences (118): 1-44. {16 Jan 1975} 

Hall, Arthur. 1928-1930. A revision of the genus Phyciodes Hubn. (Lepidoptera, Nymphalidae). Bulletin of the Hill Museum 2(2)(supplement 1): 1-20, pls. 1-3 {18 Jun 1928} 2(3)(supplement): 21-44 {17 Nov 1928}; 3(1)(supplement): 45-94 {30 Apr 1929}; 3(2)(supplement): 95-146 {29 Aug 1929}; 3(3)(supplement): 147-170 {30 Dec 1929}; 4(1)(supplement): 171-204 {28 Jul 1930}; 4(2)(supplement): 205-206; add. Page {29 Dec 1930} 

Higgins, Lionel George. 1981. A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae (Lepidoptera: Nymphalidae). Bulletin of the British Museum (Natural History) (Entomology) 43(3): 77-243, 490 figs. {29 Oct 1981} 

Pyle, Robert Michael. 2002. Butterflies of Cascadia. Seattle; Seattle Audubon Society: 420 pp., figs., drawings, photos. {2002} 

Scott, James A. 1986. The courtship of Phyciodes, and the relationship between Phyciodes tharos tharos and Phyciodes tharos morpheus (= pascoensis) in Colorado. Papilio (n.s.) 5: 1-8, 1 fig. {5 Mar 1986} 

Scott, James A. 1998. Phyciodes (Phyciodes): New discoveries, new subspecies, and convergence. Papilio (New Series) 10: 1-42, 41 figs., 5 tbls. {20 Feb 1998} 

Tilden, James William. [1970]. Concerning the names and status of certain North American members of the genus Phyciodes. Journal of Research on the Lepidoptera 8(3): 94-98 {3 Sep 1969 [21 Dec 1970]} 

Tilden, James William. 1975. An analysis of the W. G. Wright butterfly and skipper plesiotypes in the collection of the California Academy of Sciences. Occasional Papers of the California Academy of Sciences (118): 1-44. {16 Jan 1975}

Wahlberg, Niklas, Rita Oliviera and James A. Scott. 2003. Phylogenetic relationships of Phyciodes butterfly species (Lepidoptera: Nymphalidae): complex mtDNA variations and species delimitations. Systematic Entomology 28(2): 257-273, 4 figs., 2 tbls., 1 app. {Apr 2003} 

Warren, Andrew D. 2005. Butterflies of Oregon: their taxonomy, distribution, and biology. Fort Collins; Contributions of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University: 405 pp., 3 maps. {15 Mar 2005} 

Wright, William Greenwood. 1905. Butterflies of the West Coast of the United States. San Bernardino, California: 257 pp., 31 pls. {1905} [description of Phyciodes pascoensis]

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