________________________________ 
Tribe Argynnini Swainson, 1833   —   Fritillaries

Euptoieta claudia

(Cramer, 1775)

Variegated Fritillary

S N Am. (strays far N) to
Panama, Bahamas, Gr. Antilles,
Barbuda

Euptoieta hegesia meridiania

Stichel, 1938

Mexican Fritillary

[= hoffmanni], S TX, W MX
(strays N), E Mexico to
Argentina

Euptoieta hegesia hegesia

(Cramer, 1779)

Mexican Fritillary

Bahamas, Caymans, Jamaica, Cuba,
Hispaniola, Mona

Euptoieta hegesia watsoni

W. Comstock, 1944

Mexican Fritillary

Puerto Rico

Boloria alaskensis alaskensis

(W. Holland, 1900)

Mountain Fritillary

Nunavut W to AK, S to C BC

Boloria alaskensis halli

Klots, 1940

Mountain Fritillary

Wind River Mountains, NW WY

Boloria eunomia triclaris

(Hübner, [1821])

Bog Fritillary

Labrador & vic.

Boloria eunomia dawsoni

(W. Barnes & McDunnough, 1916)

Bog Fritillary

N WI, MI, MN, S boreal Canada to
Yukon in taiga habitats

Boloria eunomia nichollae

(W. Barnes & Benjamin, 1926)

Bog Fritillary

Rocky Mountains of C BC & ALTA

Boloria eunomia denali

(Klots, 1940)

Bog Fritillary

AK, N montane YK

Boloria eunomia ursadentis

Ferris & Groothuis, 1971

Bog Fritillary

Beartooth Plateau of N WY and S
MT

Boloria eunomia laddi

(Klots, 1940)

Bog Fritillary

Snowy Range of Albany and Carbon
Cos., WY, similar in Big Horn
Mts of N-central WY

Boloria eunomia caelestis

(Hemming, 1933)

Bog Fritillary

north and central Colorado Rocky
Mountains

Boloria selene myrina

(Cramer, 1777)

Silver-bordered Fritillary

 

Boloria selene nebraskensis

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria selene sabulocollis

Kohler, 1977

Silver-bordered Fritillary

 

Boloria selene tollandensis

(W. Barnes & Benjamin, 1925)

Silver-bordered Fritillary

 

Boloria selene

([Denis & Schiffermüller],
1775) (Pacific Northwestern
segregates)

Silver-bordered Fritillary

E OR, E WA, S ID, NW WY

Boloria selene albequina

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria selene atrocostalis

(Huard, 1927)

Silver-bordered Fritillary

 

Boloria selene terraenovae

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria natazhati natazhati

(Gibson, 1920)

Cryptic Fritillary

Wrangell & St. Elias Mts.,
Montana Mt., YK

Boloria natazhati nabokovi

D. Stallings & Turner, 1947

Cryptic Fritillary

N BC, probably S YK, White Mt.,
MacKenzie Mts., Bernard H., NWT

Boloria natazhati bankslandia

J. Shepard, 1998

Cryptic Fritillary

Victoria Island, NWT (and
adjacent mainland)

Boloria freija freija

(Thunberg, 1791)

Freija Fritillary

 

Boloria freija tarquinius

(J. Curtis, 1835)

Freija Fritillary

E Canadian arctic tundra

Boloria freija browni

(Higgins, 1953)

Freija Fritillary

 

Boloria chariclea montinus

(Scudder, 1863)

Purplish Fritillary

 

Boloria chariclea boisduvalii

(Duponchel, [1832])

Purplish Fritillary

SE Canadian arctic (occurrs
between B. c. arctica & B. c.
grandis)

Boloria chariclea arctica

(Zetterstedt, [1839])

Purplish Fritillary

NE Canadian arctic to N YK

Boloria chariclea grandis

(W. Barnes & Benjamin, 1916)

Purplish Fritillary

taiga from Labrador to YK & BC,
[to MT?]

Boloria chariclea butleri

(W. H. Edwards, 1883)

Purplish Fritillary

montane AK & YK to N BC

Boloria chariclea rainieri

(W. Barnes & McDunnough, 1913)

Purplish Fritillary

 

Boloria chariclea ingens

(W. Barnes & McDunnough, 1918)

Purplish Fritillary

 

Boloria chariclea

(Schneider, 1794) (NW Colorado
segregate)

Purplish Fritillary

Moffat and NW Routt counties,
Colorado

Boloria chariclea helena

(W. H. Edwards, 1871)

Purplish Fritillary

 

Boloria kriemhild

(Strecker, 1879)

Relict Fritillary

 

Boloria epithore epithore

(W. H. Edwards, 1864)

Pacific Fritillary

 

Boloria epithore sierra

E. Perkins, 1973

Pacific Fritillary

 

Boloria epithore chermocki

E. Perkins & S. Perkins, 1966

Pacific Fritillary

 

Boloria epithore sigridae

(J. Shepard, 2001)

Pacific Fritillary

 

Boloria epithore

(W. H. Edwards, 1864) (NE Oregon
segregate)

Pacific Fritillary

NE Oregon and surrounding areas

Boloria improba improba

(Butler, 1877)

Dingy Fritillary

Scandanavia, Nunavut, NWT

Boloria improba youngi

(W. Holland, 1900)

Dingy Fritillary

AK, YK

Boloria improba nunatak

Scott, 1998

Dingy Fritillary

Rocky Mountains of Alberta &
British Columbia

Boloria improba harryi

(Ferris, 1984)

Dingy Fritillary

NW Wyoming

Boloria improba acrocnema

Gall & Sperling, 1980

Dingy Fritillary; Uncompahgre Fritillary

SW Colorado Rocky Mountains

Boloria bellona bellona

(Fabricius, 1775)

Meadow Fritillary

 

Boloria bellona toddi

(W. Holland, 1928)

Meadow Fritillary

 

Boloria bellona jenistai

D. Stallings & Turner, 1947

Meadow Fritillary

boreal habitats from YK to
Labrador

Boloria bellona

(Fabricius, 1775) (S Rocky
Mountain segregate)

Meadow Fritillary

Wyoming, NW Colorado

Boloria frigga saga

(Staudinger, 1861)

Frigga Fritillary

 

Boloria frigga gibsoni

(W. Barnes & McDunnough, 1926)

Frigga Fritillary

arctic; AK to Baffin Island

Boloria frigga sagata

(W. Barnes & Benjamin, 1923)

Frigga Fritillary

 

Boloria alberta

(W. H. Edwards, 1890)

Alberta Fritillary

 

Boloria polaris polaris

(Boisduval, [1828])

Polaris Fritillary

 

Boloria polaris stellata

Masters, 1972

Polaris Fritillary

vic. Churchill, Manitoba

Boloria astarte astarte

(E. Doubleday, [1847])

Astarte Fritillary

far N WA, MT, Rocky Mountains of
BC-Alberta

Boloria astarte distincta

(Gibson, 1920)

Astarte Fritillary

most of AK, YK, NW BC, W NWT

Boloria astarte tschukotkensis

Wyatt, 1961

Astarte Fritillary

Seward Peninsula, AK

Speyeria diana

(Cramer, 1777)

Diana Fritillary

 

Speyeria cybele cybele

(Fabricius, 1775)

Great Spangled Fritillary

 

Speyeria cybele novascotiae

(McDunnough, 1935)

Great Spangled Fritillary

 

Speyeria cybele krautwurmi

(W. Holland, 1931)

Great Spangled Fritillary

 

Speyeria cybele pseudocarpenteri

(F. Chermock & R. Chermock,
1940)

Great Spangled Fritillary

 

Speyeria cybele carpenterii

(W. H. Edwards, 1876)

Great Spangled Fritillary

 

Speyeria cybele charlottii

(W. Barnes, 1897)

Great Spangled Fritillary

 

Speyeria cybele letona

dos Passos & Grey, 1945

Great Spangled Fritillary

 

Speyeria cybele leto

(Behr, 1862)

Great Spangled Fritillary

 

Speyeria cybele pugetensis

F. Chermock & Frechin, 1947

Great Spangled Fritillary

 

Speyeria cybele eileenae

J. Emmel, T. Emmel & Mattoon,
1998

Great Spangled Fritillary

 

Speyeria aphrodite cullasaja

Gatrelle, 1998

Aphrodite Fritillary

 

Speyeria aphrodite aphrodite

(Fabricius, 1787)

Aphrodite Fritillary

 

Speyeria aphrodite alcestis

(W. H. Edwards, [1877])

Aphrodite Fritillary

 

Speyeria aphrodite ethne

(Hemming, 1933)

Aphrodite Fritillary

 

Speyeria aphrodite

(Fabricius, 1789) (W Colorado
segregate)

Aphrodite Fritillary

W Colorado

Speyeria aphrodite byblis

(W. Barnes & Benjamin, 1926)

Aphrodite Fritillary

 

Speyeria aphrodite columbia

(Hy. Edwards, 1877)

Aphrodite Fritillary

 

Speyeria aphrodite whitehousei

(Gunder, 1932)

Aphrodite Fritillary

 

Speyeria aphrodite manitoba

(F. Chermock & R. Chermock,
1940)

Aphrodite Fritillary

 

Speyeria aphrodite winni

(Gunder, 1932)

Aphrodite Fritillary

 

Speyeria idalia idalia

(Drury, 1773)

Regal Fritillary

 

Speyeria idalia occidentalis

B. Williams, 2002

Regal Fritillary

 

Speyeria nokomis wenona

dos Passos & Grey, 1945

Nokomis Fritillary

NE Mexico

Speyeria nokomis melaena

Mooser & García, 1980

Nokomis Fritillary

C Mexico (Aguascalientes)

Speyeria nokomis coerulescens

(W. Holland, 1900)

Nokomis Fritillary

NW Mexico, extirpated from SE AZ

Speyeria nokomis nitocris

(W. H. Edwards, 1874)

Nokomis Fritillary

 

Speyeria nokomis nokomis

(W. H. Edwards, 1862)

Nokomis Fritillary

 

Speyeria nokomis tularosa

(R. Holland, 2010)

Nokomis Fritillary

 
 

Speyeria nokomis apacheana

(Skinner, 1918)

Nokomis Fritillary

 

Speyeria nokomis carsonensis

Austin, 1998

Nokomis Fritillary

 

Speyeria edwardsii

(Reakirt, 1866)

Edwards' Fritillary

 

Speyeria coronis semiramis

(W. H. Edwards, 1886)

Coronis Fritillary

 

Speyeria coronis hennei

(Gunder, 1934)

Coronis Fritillary

 

Speyeria coronis coronis

(Behr, 1864)

Coronis Fritillary

 

Speyeria coronis simaetha

dos Passos & Grey, 1945

Coronis Fritillary

 

Speyeria coronis snyderi

(Skinner, 1897)

Coronis Fritillary

 

Speyeria coronis halcyone

(W. H. Edwards, [1869])

Coronis Fritillary

 

Speyeria carolae

(dos Passos & Grey, 1942)

Carol's Fritillary

 

Speyeria zerene sitka

Hammond, Harry & McCorkle, 2001

Zerene Fritillary

 

Speyeria zerene bremnerii

(W. H. Edwards, 1872)

Zerene Fritillary

 

Speyeria zerene

(Boisduval, 1852) (Willamette
Valley, W OR)

Zerene Fritillary

apparently extinct

Speyeria zerene hippolyta

(W. H. Edwards, 1879)

Zerene Fritillary

 

Speyeria zerene gloriosa

Moeck, 1957

Zerene Fritillary

 

Speyeria zerene

(Boisduval, 1852) (S OR coastal
segregate)

Zerene Fritillary

 

Speyeria zerene behrensii

(W. H. Edwards, 1869)

Zerene Fritillary

 

Speyeria zerene sonomensis

J. Emmel, T. Emmel & Mattoon,
1998

Zerene Fritillary

 

Speyeria zerene puntareyes

J. Emmel & T. Emmel, 1998

Zerene Fritillary

 

Speyeria zerene myrtleae

dos Passos & Grey, 1945

Zerene Fritillary

 

Speyeria zerene monticola

(Behr, 1863)

Zerene Fritillary

 

Speyeria zerene malcolmi

(J. A. Comstock, 1920)

Zerene Fritillary

 

Speyeria zerene zerene

(Boisduval, 1852)

Zerene Fritillary

 

Speyeria zerene

(Boisduval, 1852) (E-central OR
Cascades)

Zerene Fritillary

 

Speyeria zerene gunderi

(J. A. Comstock, 1925)

Zerene Fritillary

 

Speyeria zerene picta

(McDunnough, 1924)

Zerene Fritillary

 

Speyeria zerene platina

(Skinner, 1897)

Zerene Fritillary

 

Speyeria zerene sinope

dos Passos & Grey, 1945

Zerene Fritillary

 

Speyeria callippe adiasteoides

J. Emmel, T. Emmel & Mattoon,
1998

Callippe Fritillary

 

Speyeria callippe liliana

(Hy. Edwards, 1877)

Callippe Fritillary

 

Speyeria callippe callippe

(Boisduval, 1852)

Callippe Fritillary

 

Speyeria callippe comstocki

(Gunder, 1925)

Callippe Fritillary

 

Speyeria callippe macaria

(W. H. Edwards, 1877)

Callippe Fritillary

 

Speyeria callippe laurina

(W. G. Wright, 1905)

Callippe Fritillary

 

Speyeria callippe rupestris

(Behr, 1863)

Callippe Fritillary

 

Speyeria callippe inornata

(W. H. Edwards, 1872)

Callippe Fritillary

 

Speyeria callippe juba

(Boisduval, 1869)

Callippe Fritillary

 

Speyeria callippe hagemani

J. Emmel, T. Emmel & Mattoon,
1998

Callippe Fritillary

 

Speyeria callippe shasta

J. Emmel, T. Emmel & Mattoon,
1998

Callippe Fritillary

 

Speyeria callippe elaine

dos Passos & Grey, 1945

Callippe Fritillary

 

Speyeria callippe

(Boisduval, 1852) (N Siskiyous/
W Cascades, low elevations)

Callippe Fritillary

N Siskiyous, W Cascades, Oregon,
below 3500'

Speyeria callippe

(Boisduval, 1852) (W OR
Cascades, high elevations)

Callippe Fritillary

W OR Cascades, above 3500'

Speyeria callippe semivirida

(McDunnough, 1924)

Callippe Fritillary

 

Speyeria callippe chilcotinensis

C. Guppy & J. Shepard, 2001

Callippe Fritillary

 

Speyeria callippe nevadensis

(W. H. Edwards, 1870)

Callippe Fritillary

 

Speyeria callippe harmonia

dos Passos & Grey, 1945

Callippe Fritillary

 

Speyeria callippe calgariana

(McDunnough, 1924)

Callippe Fritillary

 

Speyeria callippe gallatini

(McDunnough, 1929)

Callippe Fritillary

 

Speyeria callippe meadii

(W. H. Edwards, 1872)

Callippe Fr