________________________________ 
Subfamily Heliconiinae Swainson, 1822   —   Actinotes, Heliconians and Fritillaries
Tribe Acraeini Boisduval, 1833   —   Actinotes

Altinote ozomene nox

(H. Bates, 1864)

Lamplight Actinote

[= leucomelas], E & S Mexico to
Panama

Altinote stratonice oaxaca

(J. Y. Miller & L. Miller, 1979)

Orange-disked Actinote

S Oaxaca (Sierra Madre del Sur)

Actinote anteas

(Doubleday, [1847]) (Central
American segregate)

Anteas Actinote

S Mexico to Panama

Actinote guatemalena
guerrerensis

J. De la Maza, 1982

Guatemalan Actinote

SW Mexico (Sierra Madre del Sur)

Actinote guatemalena veraecrucis

Jordan, 1913

Guatemalan Actinote

SE Mexico

Actinote guatemalena guatemalena

(H. Bates, 1864)

Guatemalan Actinote

Chiapas to Costa Rica

Actinote melampeplos

Godman & Salvin, 1881

Bow-winged Actinote

S Mexico (S Oaxaca), Ecuador

Actinote lapitha lapitha

(Staudinger, 1885)

Pale Actinote

C Chiapas to Costa Rica & Panama

Actinote lapitha calderoni

Schaus, 1920

Pale Actinote

W Chiapas to El Salvador
Tribe Heliconiini Swainson, 1822   —   Longwings

Agraulis vanillae vanillae

(Linnaeus, 1758)

Gulf Fritillary

S Lesser Antilles (St. Lucia,
Barbados), Trinidad, N South
America, Panama

Agraulis vanillae incarnata

(N. Riley, 1926)

Gulf Fritillary

SW USA, C & S TX (strays far N)
through Central America; Hawaii

Agraulis vanillae nigrior

Michener, 1942

Gulf Fritillary

SE USA (strays far N)

Agraulis vanillae insularis

Maynard, 1889

Gulf Fritillary

Bahamas, West Indies except
southern Lesser Antilles

Dione moneta poeyii

Butler, 1873

Mexican Silverspot

E & W Mexico (strays N into USA)
to Panama

Dione juno huascuma

(Reakirt, 1866)

Juno Longwing

E & W Mexico through Central
America

Dione juno juno

(Cramer, 1779)

Juno Longwing

Lesser Antilles from Martinique
southward, NE South America

Dryadula phaetusa

(Linnaeus, 1758)

Banded Longwing

S TX (rare stray), E & S Mexico
to Brazil

Dryas iulia moderata

(N. Riley, 1926)

Julia

S TX (strays far N), E & W
Mexico through Central America

Dryas iulia largo

Clench, 1975

Julia

S FL (strays N)

Dryas iulia nudeola

(M. Bates, 1934)

Julia

Cuba

Dryas iulia carteri

(Riley, 1926)

Julia

N Bahamas

Dryas iulia zoe

L. Miller & Steinhauser, 1992

Julia

Grand Cayman & Cayman Brac

Dryas iulia delila

(Fabricius, 1775)

Julia

Jamaica

Dryas iulia fucatus

(Boddaert, 1783)

Julia

[= hispaniola], Hispaniola &
satellite islands

Dryas iulia iulia

(Fabricius, 1775)

Julia

Mona, Puerto Rico, Tortola, St.
John, St. Thomas, St. Croix

Dryas iulia warneri

(A. Hall, 1936)

Julia

St. Kitts, Montserrat, Saba

Dryas iulia dominicana

(A. Hall, 1917)

Julia

Guadeloupe & Dominica

Dryas iulia martinica

Enrico & Pinchon, 1969

Julia

Martinique

Dryas iulia lucia

(Riley, 1926)

Julia

St. Lucia

Dryas iulia framptoni

(Riley, 1926)

Julia

St. Vincent, some Grenadines,
Grenada, [Barbados?]

Philaethria diatonica

(Fruhstorfer, 1912)

Northern Green Longwing

[= Central American "dido"
records], S Mexico to Panama
(strays)

Eueides aliphera gracilis

Stichel, 1903

Fine-lined Longwing

E & S Mexico to Panama

Eueides lineata

Salvin & Godman, 1868

Thick-edged Longwing

S Mexico to Panama

Eueides lybia olympia

(Fabricius, 1793)

Sharp-edged Longwing

Atlantic slope from Nicaragua to
Colombia

Eueides lybia lybioides

Staudinger, 1876

Sharp-edged Longwing

Pacific slope of Costa Rica and
Panama

Eueides procula asidia

Schaus, 1920

Darkened Longwing

SE Mexico to Nicaragua

Eueides procula vulgiformis

Butler & H. Druce, 1872

Darkened Longwing

Costa Rica & Panama

Eueides vibilia vialis

Stichel, 1903

Vibilia Longwing

SE Mexico to Panama

Eueides isabella nigricornis

R. G. De la Maza, 1982

Isabella’s Longwing

SW Mexico

Eueides isabella eva

(Fabricius, 1793)

Isabella’s Longwing

[= zoracon], E Mexico (strays
far N) through Central America

Eueides isabella cleobaea

Geyer, 1832

Isabella’s Longwing

Cuba

Eueides isabella melphis

(Godart, 1819)

Isabella’s Longwing

Hispaniola

Heliconius charithonia vazquezae

W. Comstock & F. Brown, 1950

Zebra Longwing

S TX (strays far N), E & W
Mexico through Central America

Heliconius charithonia tuckeri

W. Comstock & F. Brown, 1950

Zebra Longwing

E TX to FL (strays N), Grand
Bahama, Great Abaco

Heliconius charithonia ramsdeni

W. Comstock & F. Brown, 1950

Zebra Longwing

Cuba, Andros, [New Providence?]

Heliconius charithonia simulator

Röber, 1921

Zebra Longwing

Jamaica

Heliconius charithonia churchi

W. Comstock & F. Brown, 1950

Zebra Longwing

Hispaniola & satellite islands

Heliconius charithonia
charithonia

(Linnaeus, 1767)

Zebra Longwing

Mona, Puerto Rico, Virgin
Islands, some Leeward Is. to
Montserrat

Heliconius charithonia antiquus

Lamas, 1988

Zebra Longwing

St Kitts & Antigua; as an
occasional form on other islands

Heliconius hewitsoni

Hewitson, 1875

Hewitson's Longwing

Costa Rica & Panama; Pacific
slope

Heliconius sapho leuce

Doubleday, 1847

Sapho Longwing

SE Mexico to Costa Rica;
Atlantic slope

Heliconius sapho sapho

(D. Drury, 1782)

Sapho Longwing

Panama

Heliconius eleuchia eleuchia

Hewitson, [1854]

White-edged Longwing

Costa Rica to Colombia

Heliconius sara veraepacis

H. Bates, 1864

Sara Longwing

SE Mexico to Costa Rica
(Atlantic Slope)

Heliconius sara fulgidus

Stichel 1906

Sara Longwing

Guatemala to Panama; Atlantic
slope

Heliconius sara theudela

Hewitson, 1874

Sara Longwing

Costa Rica & Panama; Pacific
slope

Heliconius sara magdalena

H. Bates, 1864

Sara Longwing

Panama to Colombia & Venezuela

Heliconius hortense

Guérin-Méneville, [1844]

Mexican Longwing

E & W Mexico to El Salvador &
Honduras

Heliconius clysonymus montanus

Salvin, 1871

Montane Longwing

Costa Rica & Panama

Heliconius erato cruentus

Lamas, 1998

Crimson-patched Longwing

W Mexico

Heliconius erato petiverana

(E. Doubleday, 1847)

Crimson-patched Longwing

S TX, E Mexico to Panama

Heliconius erato demophoon

Ménétriés, 1855

Crimson-patched Longwing

Nicaragua

Heliconius erato hydara

Hewitson, 1867

Crimson-patched Longwing

Panama to Brazil

Heliconius erato venus

Staudinger, 1882

Crimson-patched Longwing

SE Panama (Darien), W Colombia
(Choco)

Heliconius hecalesia octavia

H. Bates, 1866

Five-spotted Longwing

SE Mexico to Nicaragua

Heliconius hecalesia formosus

H. Bates, 1866

Five-spotted Longwing

Costa Rica & Panama

Heliconius hecale zuleika

Hewitson, 1854

Hecale Longwing

SE Mexico to Panama

Heliconius hecale fornarina

Hewitson, 1854

Hecale Longwing

C & SW Chiapas to Guatemala

Heliconius hecale discomaculatus

Weymer, 1891

Hecale Longwing

Honduras

Heliconius hecale melicerta

H. Bates, 1866

Hecale Longwing

Panama & Colombia

Heliconius ethilla claudia

Godman & Salvin, 1881

Tawny-striped Longwing

Panama

Heliconius ismenius telchinia

Doubleday, 1847

Tiger-striped Longwing

E Mexico to Panama

Heliconius ismenius clarescens

Butler, 1875

Tiger-striped Longwing

Costa Rica & Panama

Heliconius ismenius fasciatus

Godman & Salvin, 1877

Tiger-striped Longwing

Panama

Heliconius cydno galanthus

H. Bates, 1864

Cydno Longwing

SE Mexico to E Costa Rica

Heliconius cydno chioneus

H. Bates, 1864

Cydno Longwing

SE Costa Rica to Colombia

Heliconius pachinus

Salvin, 1871

Pachinus Longwing

Pacific slope of Costa Rica and
Panama

Heliconius melpomene rosina

Boisduval, 1870

Melpomene Longwing

Costa Rica & Panama

Heliconius melpomene vulcanus

Butler, 1865

Melpomene Longwing

Panama & Colombia

Heliconius doris viridis

Staudinger, 1885

Doris Longwing

SE Mexico to Panama
Tribe Argynnini Swainson, 1833   —   Fritillaries

Euptoieta claudia

(Cramer, 1775)

Variegated Fritillary

S N Am. (strays far N) to
Panama, Bahamas, Gr. Antilles,
Barbuda

Euptoieta hegesia meridiania

Stichel, 1938

Mexican Fritillary

[= hoffmanni], S TX, W MX
(strays N), E Mexico to
Argentina

Euptoieta hegesia hegesia

(Cramer, 1779)

Mexican Fritillary

Bahamas, Caymans, Jamaica, Cuba,
Hispaniola, Mona

Euptoieta hegesia watsoni

W. Comstock, 1944

Mexican Fritillary

Puerto Rico

Boloria alaskensis alaskensis

(W. Holland, 1900)

Mountain Fritillary

Nunavut W to AK, S to C BC

Boloria alaskensis halli

Klots, 1940

Mountain Fritillary

Wind River Mountains, NW WY

Boloria eunomia triclaris

(Hübner, [1821])

Bog Fritillary

Labrador & vic.

Boloria eunomia dawsoni

(W. Barnes & McDunnough, 1916)

Bog Fritillary

N WI, MI, MN, S boreal Canada to
Yukon in taiga habitats

Boloria eunomia nichollae

(W. Barnes & Benjamin, 1926)

Bog Fritillary

Rocky Mountains of C BC & ALTA

Boloria eunomia denali

(Klots, 1940)

Bog Fritillary

AK, N montane YK

Boloria eunomia ursadentis

Ferris & Groothuis, 1971

Bog Fritillary

Beartooth Plateau of N WY and S
MT

Boloria eunomia laddi

(Klots, 1940)

Bog Fritillary

Snowy Range of Albany and Carbon
Cos., WY, similar in Big Horn
Mts of N-central WY

Boloria eunomia caelestis

(Hemming, 1933)

Bog Fritillary

north and central Colorado Rocky
Mountains

Boloria selene myrina

(Cramer, 1777)

Silver-bordered Fritillary

 

Boloria selene nebraskensis

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria selene sabulocollis

Kohler, 1977

Silver-bordered Fritillary

 

Boloria selene tollandensis

(W. Barnes & Benjamin, 1925)

Silver-bordered Fritillary

 

Boloria selene

([Denis & Schiffermüller],
1775) (Pacific Northwestern
segregates)

Silver-bordered Fritillary

E OR, E WA, S ID, NW WY

Boloria selene albequina

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria selene atrocostalis

(Huard, 1927)

Silver-bordered Fritillary

 

Boloria selene terraenovae

(W. Holland, 1928)

Silver-bordered Fritillary

 

Boloria natazhati natazhati

(Gibson, 1920)

Cryptic Fritillary

Wrangell & St. Elias Mts.,
Montana Mt., YK

Boloria natazhati nabokovi

D. Stallings & Turner, 1947

Cryptic Fritillary

N BC, probably S YK, White Mt.,
MacKenzie Mts., Bernard H., NWT

Boloria natazhati bankslandia

J. Shepard, 1998

Cryptic Fritillary

Victoria Island, NWT (and
adjacent mainland)

Boloria freija freija

(Thunberg, 1791)

Freija Fritillary

 

Boloria freija tarquinius

(J. Curtis, 1835)

Freija Fritillary

E Canadian arctic tundra

Boloria freija browni

(Higgins, 1953)

Freija Fritillary

 

Boloria chariclea montinus

(Scudder, 1863)

Purplish Fritillary

 

Boloria chariclea boisduvalii

(Duponchel, [1832])

Purplish Fritillary

SE Canadian arctic (occurrs
between B. c. arctica & B. c.
grandis)

Boloria chariclea arctica

(Zetterstedt, [1839])

Purplish Fritillary

NE Canadian arctic to N YK

Boloria chariclea grandis

(W. Barnes & Benjamin, 1916)

Purplish Fritillary

taiga from Labrador to YK & BC,
[to MT?]

Boloria chariclea butleri

(W. H. Edwards, 1883)

Purplish Fritillary

montane AK & YK to N BC

Boloria chariclea rainieri

(W. Barnes & McDunnough, 1913)

Purplish Fritillary

 

Boloria chariclea ingens

(W. Barnes & McDunnough, 1918)

Purplish Fritillary

 

Boloria chariclea

(Schneider, 1794) (NW Colorado
segregate)

Purplish Fritillary

Moffat and NW Routt counties,
Colorado

Boloria chariclea helena

(W. H. Edwards, 1871)

Purplish Fritillary

 

Boloria kriemhild

(Strecker, 1879)

Relict Fritillary

 

Boloria epithore epithore

(W. H. Edwards, 1864)

Pacific Fritillary

 

Boloria epithore sierra

E. Perkins, 1973

Pacific Fritillary

 

Boloria epithore chermocki

E. Perkins & S. Perkins, 1966

Pacific Fritillary

 

Boloria epithore sigridae

(J. Shepard, 2001)

Pacific Fritillary

 

Boloria epithore

(W. H. Edwards, 1864) (NE Oregon
segregate)

Pacific Fritillary

NE Oregon and surrounding areas

Boloria improba improba

(Butler, 1877)

Dingy Fritillary

Scandanavia, Nunavut, NWT

Boloria improba youngi

(W. Holland, 1900)

Dingy Fritillary

AK, YK

Boloria improba nunatak

Scott, 1998

Dingy Fritillary

Rocky Mountains of Alberta &
British Columbia

Boloria improba harryi

(Ferris, 1984)

Dingy Fritillary

NW Wyoming

Boloria improba acrocnema

Gall & Sperling, 1980

Dingy Fritillary; Uncompahgre Fritillary

SW Colorado Rocky Mountains

Boloria bellona bellona

(Fabricius, 1775)

Meadow Fritillary

 

Boloria bellona toddi

(W. Holland, 1928)

Meadow Fritillary

 

Boloria bellona jenistai

D. Stallings & Turner, 1947

Meadow Fritillary

boreal habitats from YK to
Labrador

Boloria bellona

(Fabricius, 1775) (S Rocky
Mountain segregate)

Meadow Fritillary

Wyoming, NW Colorado

Boloria frigga saga

(Staudinger, 1861)

Frigga Fritillary

 

Boloria frigga gibsoni

(W. Barnes & McDunnough, 1926)

Frigga Fritillary

arctic; AK to Baffin Island

Boloria frigga sagata

(W. Barnes & Benjamin, 1923)

Frigga Fritillary

 

Boloria alberta

(W. H. Edwards, 1890)

Alberta Fritillary

 

Boloria polaris polaris

(Boisduval, [1828])

Polaris Fritillary

 

Boloria polaris stellata

Masters, 1972

Polaris Fritillary

vic. Churchill, Manitoba

Boloria astarte astarte

(E. Doubleday, [1847])

Astarte Fritillary

far N WA, MT, Rocky Mountains of
BC-Alberta

Boloria astarte distincta

(Gibson, 1920)

Astarte Fritillary

most of AK, YK, NW BC, W NWT

Boloria astarte tschukotkensis

Wyatt, 1961

Astarte Fritillary

Seward Peninsula, AK

Speyeria diana

(Cramer, 1777)

Diana Fritillary

 

Speyeria cybele cybele

(Fabricius, 1775)

Great Spangled Fritillary

 

Speyeria cybele novascotiae

(McDunnough, 1935)

Great Spangled Fritillary

 

Speyeria cybele krautwurmi

(W. Holland, 1931)

Great Spangled Fritillary

 

Speyeria cybele pseudocarpenteri

(F. Chermock & R. Chermock,
1940)

Great Spangled Fritillary

 

Speyeria cybele carpenterii

(W. H. Edwards, 1876)

Great Spangled Fritillary

 

Speyeria cybele charlottii

(W. Barnes, 1897)

Great Spangled Fritillary

 

Speyeria cybele letona

dos Passos & Grey, 1945

Great Spangled Fritillary

 

Speyeria cybele leto

(Behr, 1862)

Great Spangled Fritillary

 

Speyeria cybele pugetensis

F. Chermock & Frechin, 1947

Great Spangled Fritillary

 

Speyeria cybele eileenae

J. Emmel, T. Emmel & Mattoon,
1998

Great Spangled Fritillary

 

Speyeria aphrodite cullasaja

Gatrelle, 1998

Aphrodite Fritillary

 

Speyeria aphrodite aphrodite

(Fabricius, 1787)

Aphrodite Fritillary

 

Speyeria aphrodite alcestis

(W. H. Edwards, [1877])

Aphrodite Fritillary

 

Speyeria aphrodite ethne

(Hemming, 1933)

Aphrodite Fritillary

 

Speyeria aphrodite

(Fabricius, 1789) (W Colorado
segregate)

Aphrodite Fritillary

W Colorado

Speyeria aphrodite byblis

(W. Barnes & Benjamin, 1926)

Aphrodite Fritillary

 

Speyeria aphrodite columbia

(Hy. Edwards, 1877)

Aphrodite Fritillary

 

Speyeria aphrodite whitehousei

(Gunder, 1932)

Aphrodite Fritillary

 

Speyeria aphrodite manitoba

(F. Chermock & R. Chermock,
1940)

Aphrodite Fritillary

 

Speyeria aphrodite winni

(Gunder, 1932)

Aphrodite Fritillary

 

Speyeria idalia idalia

(Drury, 1773)

Regal Fritillary