________________________________ 
Subfamily Hesperiinae Latreille, 1809   —   Grass-Skippers
Tribe Incertae Sedis   —  

Erionota thrax thrax

(Linnaeus, 1767)

Banana Skipper

Introduced to Hawaii

Perichares philetes

(Gmelin, [1790])

Caribbean Ruby-eye

Greater Antilles

Perichares adela

(Hewitson, 1867)

Green-backed Ruby-eye

S TX (uncommon), E & W Mexico to
S Brazil

Perichares aurina

Evans, 1955

Evans' Ruby-eye

[Honduras?], S Brazil & Paraguay

Perichares poaceaphaga

Burns, 2008

Grass-feeding Ruby-eye

Costa Rica

Perichares geonomaphaga

Burns, 2008

Geonoma-feeding Ruby-eye

Costa Rica

Perichares prestoeaphaga

Burns, 2008

Prestoea-feeding Ruby-eye

Costa Rica

Perichares lotus

(Butler, 1870)

Lotus Ruby-eye

SE Mexico S Brazil & Paraguay

Perichares deceptus deceptus

(Butler & H. Druce, 1872)

Flasher Ruby-eye

El Salvador to Costa Rica

Perichares seneca crotona

(Hewitson, 1866)

Seneca Ruby-eye

Panama, Venezuela

Orses cynisca

(Swainson, 1821)

Yellow-edged Ruby-eye

E & W Mexico to S Brazil &
Paraguay

Alera haworthiana

(Swainson, 1821)

Green-banded Ruby-eye

Panama to S Brazil

Lycas argentea

(Hewitson, 1866)

Silvered Ruby-eye

E & W Mexico to Argentina

Pyrrhopygopsis agaricon agaricon

H. Druce, 1908

Agaricon Skipper

E Costa Rica, Colombia
Tribe Megathymini J. H. Comstock & A. Comstock, 1895   —   Giant-Skippers

Megathymus yuccae yuccae

(Boisduval & Le Conte, [1837])

Yucca Giant-Skipper

[= buchholzi], SE USA, E of
Mississippi River

Megathymus yuccae reinthali

H. Freeman, 1963

Yucca Giant-Skipper

[= kendalli], C to NE TX, C & E
OK, SW AR, N & C LA

Megathymus yuccae wilsonorum

D. Stallings & Turner, 1958

Yucca Giant-Skipper

Tamaulipas & far S TX (lower Rio
Grande Valley)

Megathymus yuccae louiseae

H. Freeman, 1963

Yucca Giant-Skipper

S-C TX (Val Verde, Kinney Cos)
to NE Mexico (COAH, NL)

Megathymus yuccae coloradensis

C. Riley, 1878

Yucca Giant-Skipper

SE WY, W NE, W KS, E CO, E NM, W
OK, TX panhandle

Megathymus yuccae reubeni

D. Stallings, Turner & V.
Stallings, 1963

Yucca Giant-Skipper

Hueco Mts., Hudspeth Co., TX, to
N Chihuahua

Megathymus yuccae navajo

Skinner, 1911

Yucca Giant-Skipper

[= arizonae], W CO, S UT, W NM,
AZ, N Sonora

Megathymus yuccae martini

D. Stallings & Turner, 1956

Yucca Giant-Skipper

Mohave Desert, S CA, W NV, far
NW AZ

Megathymus yuccae harbisoni

J. Emmel & T. Emmel, 1998

Yucca Giant-Skipper

Beaumont, Riverside Co. CA, S to
Baja California Norte

Megathymus beulahae beulahae

D. Stallings & Turner, 1958

Broad-banded Giant-Skipper

Hidalgo

Megathymus beulahae gayleae

D Stallings, Turner & V.
Stallings, 1963

Broad-banded Giant-Skipper

COAH, NL, TAMPS, SLP

Megathymus ursus ursus

Poling, 1902

Ursine Giant-Skipper

S AZ (Pinal, Pima, Sta. Cruz,
Cochise), SW NM (Hidalgo), SON

Megathymus ursus deserti

R. Wielgus, J. Wielgus & D.
Wielgus, 1972

Ursine Giant-Skipper

E Maricopa Co., AZ

Megathymus ursus violae

D. Stallings & Turner, 1956

Ursine Giant-Skipper

C & S NM (E of Rio Grande), W
TX, COAH, NL

Megathymus cofaqui cofaqui

(Strecker, 1876)

Cofaqui Giant-Skipper

[= harrisi], Georgia to NC

Megathymus cofaqui slotteni

Gatrelle, 1999

Cofaqui Giant-Skipper

Florida

Megathymus streckeri texana

W. Barnes & McDunnough, 1912

Strecker's Giant-Skipper

S-C Texas

Megathymus streckeri leussleri

W. Holland, 1931

Strecker's Giant-Skipper

TX panhandle & E NM, to E MT, W
ND, (SE SASK?)

Megathymus streckeri streckeri

(Skinner, 1895)

Strecker's Giant-Skipper

NE AZ, NW NM, SW CO, SE UT

Megathymus streckeri

(Skinner, 1895) (S Navajo Co.,
Arizona segregate)

Strecker's Giant-Skipper

S end White Mts., AZ

Stallingsia smithi

(H. Druce, 1896)

Smith's Giant-Skipper

SW MX (JAL, MICH, GRO, DF, MOR)

Stallingsia maculosus

(H. Freeman, 1955)

Manfreda Giant-Skipper

S TX, N NL

Stallingsia jacki

D. Stallings, Turner & V.
Stallings, 1963

Chiapan Giant-Skipper

Chiapas

Aegiale hesperiaris

(Walker, 1856) complex

Tequila Giant-Skipper; Gusano de Maguey

much of N & C Mexico

Turnerina mejicanus

(E. Bell, 1938)

Turner's Giant-Skipper

Durango

Turnerina hazelae

(D. Stallings & Turner, 1958)

Guerrero Giant-Skipper

Guerrero & Oaxaca

Agathymus neumoegeni neumoegeni

(W. H. Edwards, 1882)

Neumoegen’s Giant-Skipper

C AZ (Yavapai Co.) to W-C NM
(Catron Co.)

Agathymus neumoegeni

(W. H. Edwards, 1882) (S NM
segregate)

Neumoegen’s Giant-Skipper

S NM: Organ Mts. (Dona Ana-Otero
Cos.) to Sacramento Mts.

Agathymus neumoegeni
carlsbadensis

(D. Stallings & Turner, 1957)

Neumoegen’s Giant-Skipper

Guadalupe Mts. (Eddy-Otero Cos.,
NM; Culberson Co., TX)

Agathymus neumoegeni judithae

(D. Stallings & Turner, 1957)

Neumoegen’s Giant-Skipper

Hueco Mts., Hudspeth Co., TX

Agathymus neumoegeni diabloensis

H. Freemen, 1962

Neumoegen’s Giant-Skipper

Sierra Diablo, Sierra Blanca,
Hudspeth Co., TX

Agathymus neumoegeni florenceae

(D. Stallings & Turner, 1957)

Neumoegen’s Giant-Skipper

Davis Mts., Jeff Davis Co., TX

Agathymus neumoegeni mcalpinei

(H. Freeman, 1955)

Neumoegen’s Giant-Skipper

E side of Glass Mts., Brewster
Co., TX

Agathymus neumoegeni chisosensis

(H. Freeman, 1952)

Neumoegen’s Giant-Skipper

Chisos Mts., S Brewster Co., TX

Agathymus juliae

(D. Stallings & Turner, 1958)

Julia’s Giant-Skipper

Durango

Agathymus hoffmanni

(H. Freeman, 1952)

Hoffmann’s Giant-Skipper

S-C Mexico (DF, GTO, HGO, MEX,
SLP)

Agathymus polingi

(Skinner, 1905)

Poling's Giant-Skipper

S AZ, SW NM (Hidalgo Co.), NE
Sonora

Agathymus evansi

(H. Freeman, 1950)

Huachuca Giant-Skipper

Huachuca Mts., Cochise Co., AZ

Agathymus aryxna

(Dyar, 1905)

Arizona Giant-Skipper

SE AZ, SW NM, NE SON

Agathymus baueri baueri

(D. Stallings & Turner, 1954)

Bauer's Giant-Skipper

C AZ: E Yavapai, W Gila, NE
Maricopa, N Pinal Cos.

Agathymus baueri freemani

D. Stallings & Turner, 1960

Bauer's Giant-Skipper

W-C AZ: SW Yavapai Co., S Mohave
Co.

Agathymus gentryi

Roever, 1998

Gentry's Giant-Skipper

SW AZ (W Pima, C Maricopa, W
Pinal, W Yuma), E SB Co., CA

Agathymus belli

(H. Freeman, 1955)

Bell's Giant-Skipper

Chihuahua, Durango

Agathymus ricei

D. Stallings, Turner & V.
Stallings, 1966

Rice's Giant-Skipper

Puebla

Agathymus indecisa

(Butler & H. Druce, 1872)

Guatemalan Giant-Skipper

Chiapas to Costa Rica

Agathymus mariae mariae

(W. Barnes & Benjamin, 1924)

Mary's Giant-Skipper

S NM, trans-Pecos counties, TX

Agathymus mariae chinatiensis

H. Freeman, 1964

Mary's Giant-Skipper

Chinati Mts., Presidio Co., TX

Agathymus mariae lajitaensis

H. Freeman, 1964

Mary's Giant-Skipper

S Presidio Co., TX

Agathymus mariae rindgei

H. Freeman, 1964

Mary's Giant-Skipper

Val Verde and Kinney Cos., TX

Agathymus mariae

(W. Barnes & Benjamin, 1924)
complex (Mexican taxa)

Mary's Giant-Skipper

various undescribed entities
occur in COAH, NL, and vic.
(MEX)

Agathymus gilberti

H. Freeman, 1964

Gilbert's Giant-Skipper

Brewster, Terrell and Val Verde
Cos., TX

Agathymus micheneri

D. Stallings, Turner & V.
Stallings, 1961 (Typ. segregate)

Michener’s Giant-Skipper

Coahuila

Agathymus micheneri

D. Stallings, Turner & V.
Stallings, 1961 complex

Michener’s Giant-Skipper

various undescribed entities
occur in COAH, NL, and vic.

Agathymus remingtoni

(D. Stallings & Turner, 1958)

Remington's Giant-Skipper

E Mexico: HGO, SLP, TAMPS

Agathymus fieldi

H. Freeman, 1960

Field's Giant-Skipper

W Mexico: JAL, SON

Agathymus estelleae estelleae

(D. Stallings & Turner, 1958)

Estelle's Giant-Skipper

Coahuila, Nuevo Leon

Agathymus estelleae
valverdiensis

H. Freeman, 1966

Estelle's Giant-Skipper

S-C TX (Terrell, Val Verde,
Edwards, Kinney Cos.)

Agathymus escalantei

D. Stallings, Turner & V.
Stallings, 1966

Escalante's Giant-Skipper

Guerrero

Agathymus stephensi

(Skinner, 1912)

California Giant-Skipper

S CA (Riverside, E San Diego, W
Imperial Cos.), Baja (N)

Agathymus comstocki

(Harbison, 1957)

Comstock's Giant-Skipper

Baja California (N)

Agathymus dawsoni

Harbison, 1963 complex

Dawson’s Giant-Skipper

Baja California (N)

Agathymus sp. undescribed



Undescribed Giant-Skipper

Baja California (N)

Agathymus complex undescribed



Cape Giant-Skipper

Baja California Sur

Agathymus alliae alliae

(D. Stallings & Turner, 1957)

Mojave Giant-Skipper

N Coconino Co., AZ

Agathymus alliae paiute

Roever, 1998

Mojave Giant-Skipper

SE CA (SB, Inyo), S NV (Clark),
SW UT (Wa), NW AZ

Agathymus rethon

(Dyar, 1913)

Black Giant-Skipper

Mexico (COL, GRO, JAL, MICH,
MOR, NAY, OAX, PUE, VER), El
Salvador
Tribe Thymelicini Tutt, 1905   —  

Ancyloxypha numitor

(Fabricius, 1793)

Least Skipper

Great Plains, eastward; SE SASK
to Quebec, to S TX, far NE MEX

Ancyloxypha arene

(W. H. Edwards, 1871)

Tropical Least Skipper

SE AZ, SW NM, W & S TX, S to
Costa Rica

Oarisma poweshiek

(Parker, 1870)

Poweshiek Skipperling

SE MAN, E ND, NE SD, W & S MN,
N-central IA, SE WI, S MI

Oarisma garita garita

(Reakirt, 1866)

Garita Skipperling

SE BC to S MAN (plus Peace
Riv.), S through Rockies to AZ &
NM

Oarisma garita calega

(Godman, 1900)

Garita Skipperling

NW (DGO) to S (OAX) Mexico

Oarisma edwardsii

(W. Barnes, 1897)

Edwards’ Skipperling

S-central CO, NM, C & E AZ, NW
Mexico to S-central Mexico

Oarisma era

Dyar, 1927

Bold-veined Skipperling

NW & NE to central Mexico

Oarisma nanus

(Herrich-Schäffer, 1865)

Nanus Skipperling

Cuba & Isla de Juventud

Oarisma bruneri

E. Bell, 1959

Bruner's Skipperling

Cuba

Copaeodes aurantiaca

(Hewitson, 1868)

Orange Skipperling

S CA & NV to E TX (strays N), S
to Guatemala

Copaeodes minima

(W. H. Edwards, 1870)

Southern Skipperling

SE AZ, C TX to FL & NC, E & W
Mexico to Costa Rica

Copaeodes stillmani

E. Bell & W. Comstock, 1948

Stillman's Skipperling

Hispaniola including Isla
Catalina

Copaeodes eoa

D. Smith, L. Miller & McKenzie,
1991

Anegada Skipperling

Anegada

Adopaeoides prittwitzi

(Plötz, 1884)

Sunrise Skipper

SE AZ, SW NM, W TX, S to
S-central Mexico

Thymelicus lineola lineola

(Ochsenheimer, 1808)

European Skipper

introduced; Newfoundland to VA &
TN, W to MAN, S Rocky Mts.
Tribe Calpodini A. Clark, 1948   —  

Turesis complanula

(Herrich-Schäffer, 1869)

Complanula Skipper

SE Mexico to S Brazil

Turesis theste

Godman, 1901

Theste Skipper

SE Mexico to Costa Rica

Turesis tabascoensis

H. Freeman, 1979

Tabasco Skipper

SE Mexico

Ebusus ebusus nigrior

L. Miller, 1985

Ebusus Skipper

SE Mexico (Veracruz and Chiapas)

Argon lota

(Hewitson, 1877)

Argon Skipper

[= argus], W & S Mexico to
Argentina

Megaleas syrna

(Godman & Salvin, 1879)

Megaleas Skipper

Costa Rica

Talides sergestus

(Cramer, 1775)

Sergestus Ruby-eye

SE Mexico to S Brazil

Talides cantra

Evans, 1955

Cantra Ruby-eye

S Mexico to Colombia, Venezuela
& W Ecuador

Talides alternata

E. Bell, 1941

Alternate Ruby-eye

S Mexico to S Brazil

Talides hispa

Evans, 1955

Hispa Skipper

Costa Rica to W Ecuador, W
Brazil

Tromba xanthura

(Godman, 1901)

Yellow-washed Ruby-eye

E & W Mexico to Colombia

Synale cynaxa

(Hewitson, 1867)

Black-veined Ruby-eye

E & S Mexico to Costa Rica

Carystus ploetzi

Mielke & Casagrande, 2002

Ploetz's Ruby-eye

E & S Mexico to central Brazil &
Peru

Carystus phorcus phorcus

(Cramer, 1777)

White-patched Ruby-eye

E & W Mexico to Argentina

Tisias quadrata hermoda

(Hewitson, 1870)

Quadrate Ruby-eye

Panama to Ecuador

Tisias myna

(Mabille, 1889)

Myna Ruby-eye

Costa Rica and Panama

Cobalus calvina

(Hewitson, 1866)

White-rayed Ruby-Eye

TL: Brazil (Pa)

Cobalus virbius virbius

(Cramer, 1777)

White-edged Ruby-eye

Costa Rica to S Brazil &
Paraguay

Cobalus fidicula

(Hewitson, 1877)

White-centered Ruby-eye

S Mexico to Panama

Dubiella fiscella belpa

Evans, 1955

Yellow-striped Ruby-eye

S Mexico to Ecuador, Venezuela,
Guianas

Carystina aurifer

(Godman & Salvin, 1879)

Golden-spotted Ruby-eye

Costa Rica to W Ecuador

Damas clavus

(Herrich-Schäffer, 1869)

Violet-washed Skipper

E & S Mexico to S Brazil &
Bolivia

Damas immaculata

Nicolay, 1973

Immaculate Ruby-eye

Panama

Orphe gerasa

(Hewitson, 1867)

Gerasa Skipper

Nicaragua to Amazon basin &
Guianas

Orphe vatinius

Godman, 1901

Vatinius Skipper

Costa Rica, Colombia, Venezuela,
Guyana, French Guiana, Brazil,
Peru, Ecuador

Carystoides basoches

(Latreille, [1824])

Basoches Ruby-eye

S Mexico to S Brazil & Paraguay

Carystoides lila

Evans, 1955

Lila Ruby-eye

S Mexico to S Ecuador

Carystoides escalantei

H. Freeman, 1969

Escalante's Ruby-eye

E Mexico to Costa Rica

Carystoides abrahami

H. Freeman, 1969

Abraham’s Ruby-eye

E & W Mexico

Carystoides floresi

H. Freeman, 1969

Freeman’s Ruby-eye

E Mexico

Carystoides mexicana

H. Freeman, 1969

Mexican Ruby-eye

E Mexico

Carystoides hondura

Evans, 1955

Honduran Ruby-eye

Honduras to Costa Rica

Carystoides lebbaeus lebbaeus

(Hewitson, 1876)

Lebbaeus Ruby-eye

Costa Rica to NW Ecuador

Carystoides sicania orbius

(Godman, 1901)

Yellow-spotted Ruby-eye

S Mexico to Ecuador & Guianas

Carystoides maroma

(Möschler, 1877)

Maroma Ruby-eye

Panama to Amazon basin

Lychnuchoides saptine

(Godman & Salvin, 1879)

Golden-banded Ruby-eye

S Mexico (OAX, CHIS) to Costa
Rica

Calpodes ethlius

(Stoll, 1782)

Brazilian Skipper

SE USA (strays far N), E & W
Mexico to Argentina, West Indies

Panoquina panoquin

(Scudder, 1863)

Salt Marsh Skipper

coastal salt marshes, New
England to S TX

Panoquina panoquinoides
panoquinoides

(Skinner, 1891)

Obscure Skipper

S FL to S TX, Mexican Gulf Coast
to Costa Rica, most West Indies

Panoquina panoquinoides eugeon

(Godman & Salvin, 1896)

Obscure Skipper

Grenada, Grenadines

Panoquina errans

(Skinner, 1892)

Wandering Skipper

S CA, W Mexico

Panoquina nero

(Fabricius, 1798)

Nero Skipper

Hispaniola and Puerto Rico

Panoquina corrupta

(Herrich-Schäffer, 1865)

Corrupt Skipper

Cuba and Isla de Juventud

Panoquina ocola ocola

(W. H. Edwards, 1863)

Ocola Skipper

Greater Antilles, Texas to
Argentina

Panoquina ocola distipuncta

K. Johnson & Matusik, 1988

Ocola Skipper

SW Dominican Republic

Panoquina hecebolus

(Scudder, 1872)

Hecebolus Skipper

S TX, E & S Mexico to Paraguay

Panoquina lucas

(Fabricius, 1793)

Purple-washed Skipper

[= sylvicola, = woodruffi], Gr.
& Less. Antilles, S TX to
Argentina

Panoquina evansi

(H. Freeman, 1946)

Evans’ Skipper

S TX, E & W Mexico to W Panama

Panoquina fusina sonta

Evans, 1955

Fusina Skipper

Panama & Colombia

Panoquina fusina viola

Evans, 1955

Fusina Skipper

[Honduras?, Colombia?], Bolivia,
SE Brazil, Paraguay

Panoquina pauper pauper

(Mabille, 1878)

Pauper Skipper

SE Mexico to Colombia

Panoquina evadnes

(Stoll, 1781)

Evadnes Skipper

S Mexico to Argentina

Zenis jebus hemizona

(Dyar, 1918)

Purple-stained Skipper

[= janka], E & W Mexico to
Ecuador, Peru, Guianas, Trinidad

Zenis minos

(Latreille, [1824])

Minos Skipper

SE Mexico, Honduras, Costa Rica,
S Brazil

Saliana chiomara

(Hewitson, 1867)

Chiomara Saliana

Panama to Peru & Brazil

Saliana triangularis

(Kaye, 1914)

Triangular Saliana

E & S Mexico to Argentina

Saliana fusta

Evans, 1955

Suffused Saliana

E & W Mexico to Brazil & Bolivia

Saliana placens

(Butler, 1874)

Ochreous Saliana

Costa Rica to Colombia & Ecuador

Saliana hewitsoni

(Riley, 1926)

Green Saliana

SE Mexico to Bolivia

Saliana esperi esperi

Evans, 1955

Perching Saliana

E & W Mexico to S Brazil and
Bolivia

Saliana esperi soroa

D. Smith & L. Hernandez, 1992

Perching Saliana

W Cuba

Saliana antoninus

(Latreille, [1824])

Persistent Saliana

S Mexico to S Brazil

Saliana longirostris

(Sepp, [1840])

Shy Saliana

E & W Mexico to S Brazil

Saliana morsa

Evans, 1955

Morsa Saliana

Taboga Is., Panama, to Guianas &
Amazon basin

Saliana salius

(Cramer, 1775)

Sullied Saliana

S Mexico to S Brazil & Uruguay

Saliana saladin saladin

Evans, 1955

Violet-tipped Saliana

S Mexico to Colombia & Peru

Saliana severus

(Mabille, 1895)

Dark Saliana

S Mexico to Colombia & Ecuador

Thracides phidon

(Cramer, 1779)

Jewel-studded Skipper

E & W Mexico to S Brazil

Thracides thrasea

(Hewitson, 1866)

Thrasea Skipper

S Mexico (OAX), Belize, Guianas,
Amazon basin to Bolivia