________________________________ 
Subfamily Riodininae Grote, 1895 (1827)   —   True Metalmarks

Mesosemia esperanza

Schaus, 1913

Esperanza Eyed-Metalmark

Costa Rica and Panama

Mesosemia coelestis

Godman & Salvin, 1885

Pale-blue Eyed-Metalmark

Costa Rica to Colombia

Mesosemia albipuncta

Schaus, 1913

White-spotted Eyed-Metalmark

Costa Rica and Panama

Mesosemia hesperina hesperina

Butler, 1874

Hesperina Eyed-Metalmark

Nicaragua to Venezuela

Mesosemia zonalis

Godman & Salvin, 1885

Whitened Eyed-Metalmark

Honduras to Colombia

Mesosemia ceropia

H. Druce, 1874

Spying Eyed-Metalmark

Costa Rica to Colombia

Mesosemia hypermegala

Stichel, 1909

Hiding Eyed-Metalmark

Costa Rica to Colombia

Mesosemia telegone telegone

(Boisduval, 1836)

Violet-washed Eyed-Metalmark

SE Mexico to Colombia

Mesosemia lamachus

Hewitson, 1857

Purple-washed Eyed-Metalmark

[= tertica], E & W Mexico to
Colombia

Mesosemia carissima

H. Bates, 1866

Blue-patched Eyed-Metalmark

Nicaragua to Panama

Mesosemia grandis

H. Druce, 1874

Giant Eyed-Metalmark

Costa Rica and Panama

Mesosemia gaudiolum

H. Bates, 1865

Gaudy Eyed-Metalmark

SE Mexico to Costa Rica

Mesosemia gemina

J. De la Maza & R. G. De la
Maza, 1980

Turquoise Eyed-Metalmark

SE Mexico

Mesosemia asa asa

Hewitson, 1869

Deep-blue Eyed-Metalmark

Nicaragua to Panama

Mesosemia asa asopis

Stichel, 1909

Deep-blue Eyed-Metalmark

Panama and Colombia

Mesosemia harveyi

DeVries & J. Hall, 1996

Harvey's Eyed-Metalmark

Costa Rica and Panama

Leucochimona vestalis vestalis

(H. Bates, 1865)

Vestalis Metalmark

SE Mexico and Guatemala

Leucochimona vestalis leucogaea

(Godman & Salvin, 1885)

Vestalis Metalmark

Guatemala to Panama

Leucochimona lepida nivalis

(Godman & Salvin, 1885)

Satyr Metalmark

SE Mexico to Guatemala

Leucochimona lepida lepida

(Godman & Salvin, 1885)

Satyr Metalmark

Nicaragua to Panama

Leucochimona iphias

Stichel, 1909

Iphias Metalmark

Panama

Leucochimona lagora

(Herrich-Schäffer, [1853])

Lagora Metalmark

Honduras to Guianas & Ecuador

Leucochimona molina

(Godman & Salvin, 1885)

Molina Metalmark

Nicaragua to Panama

Perophthalma tullius

(Fabricius, 1787)

Tullius Metalmark

Costa Rica to Brazil

Perophthalma lasus

Westwood, 1851

Lasus Metalmark

SE Mexico to Panama

Hyphilaria thasus

(Stoll, 1780)

Thasus Metalmark

[= subrutila], SE Mexico S
Brazil & Paraguay, Guianas

Hyphilaria parthenis

(Westwood, 1851)

Fire-banded Metalmark

C Panama to Bolivia & S Brazil,
Guianas

Napaea heteroea

(H. Bates, 1867)

Blue-washed Metalmark

Guianas, Venezuela, Colombia to
S Peru, Amazonas

Napaea actoris

(Cramer, 1776)

Actoris Metalmark

far SE Peru, Bolivia, C & S
Amazonia

Napaea beltiana

(H. Bates, 1867)

Clouded Metalmark

[= malis, = aza], Chiapas to
Bolivia, Amazonas

Napaea rufolimba

J. Hall, 2005

Red-margined Metalmark

SE Mexico to W & C Ecuador

Napaea eucharila

(H. Bates, 1867)

White-stitched Metalmark

[= picina], SE Mexico to
Bolivia, Paraguay, S Brazil,
Guianas, Trinidad

Voltinia danforthi

(A. Warren & Opler, 1999)

Ancient Metalmark

NW Mexico (Sonora to Jalisco)

Voltinia theata

Stichel, 1910

Blue-banded Metalmark

Costa Rica to W Ecuador

Voltinia radiata

(Godman & Salvin, 1886)

White-rayed Metalmark

Costa Rica to Colombia & W
Ecuador

Voltinia umbra

(Boisduval, 1870) (W Mexican
segregate)

Quilted Metalmark

W Mexico to Chiapas

Voltinia umbra

(Boisduval, 1870) (Chiapas
segregate)

Quilted Metalmark

S-central Chiapas (very large
adults)

Voltinia umbra

(Boisduval, 1870) (Typical
segregate)

Quilted Metalmark

E Mexico to Panama

Hermathena oweni

Schaus, 1913

Peppered Metalmark

SE Mexico to Costa Rica

Hermathena eburna

J. Hall & Harvey, 2005

White Metalmark

[= regional records of H.
candidata], Costa Rica to W
Ecuador

Ithomiola floralis celtilla

(Hewitson, 1870)

Clearwing Metalmark

E Panama to Bolivia

Ithomiola calculosa

J. Hall & Harvey, 2005

Calculosa Metalmark

Costa Rica to C Panama

Ithomiola theages tessera

Hall, 2005

White-spotted Metalmark

SE Mexico to Costa Rica

Ithomiola theages theages

(Godman & Salvin, 1878)

White-spotted Metalmark

Costa Rica to NW Ecuador

Ithomiola cribralis

(Stichel, 1915)

Cloud-spotted Metalmark

C Panama to W Ecuador

Eurybia cyclopia cyclopia

Stichel, 1910

Fire-bordered Eurybia

Costa Rica to Ecuador

Eurybia unxia

Godman & Salvin, 1885

Azure-winged Eurybia

[= fulgens], Costa Rica and
Panama

Eurybia lycisca

Westwood, 1851

Blue-winged Eurybia

SE Mexico to Ecuador

Eurybia patrona persona

Staudinger, 1876

Great Eurybia

Honduras to Panama

Eurybia elvina elvina

Stichel, 1910

Blind Eurybia

E & W Mexico to Panama

Lyropteryx lyra cleadas

H. Druce, 1875

Lyra Metalmark

SE Mexico to Panama

Necyria larunda

Godman & Salvin, 1885

Larunda Metalmark

Chiapas & Guatemala

Necyria duellona beltiana

Hewitson, 1870

White-dashed Metalmark

Nicaragua to Panama

Necyria ingaretha

Hewitson, 1872

Orange-spotted Metalmark

Nicaragua & Costa Rica

Cyrenia martia pyrippe

Godman & Salvin, 1878

Red-spotted Metalmark

Costa Rica & Panama

Cyrenia martia martia

Westwood, 1851

Red-spotted Metalmark

Amazonas

Ancyluris inca inca

(Saunders, 1850)

Inca Metalmark

S Mexico to Panama

Ancyluris jurgensenii
jurgensenii

(Saunders, 1850)

Costa-spotted Metalmark

S Mexico to Panama

Rhetus arcius beutelspacheri

Llorente, 1988

Long-tailed Metalmark

W Mexico

Rhetus arcius thia

(Morisse, 1838)

Long-tailed Metalmark

E Mexico

Rhetus arcius castigatus

Stichel, 1909

Long-tailed Metalmark

Guatemala to Costa Rica

Rhetus dysoni caligosus

Stichel, 1929

Dyson's Metalmark

Costa Rica to Colombia

Rhetus periander naevianus

Stichel, 1910

Periander Metalmark

S Mexico to Panama

Chorinea bogota

(Saunders, 1859)

Northern Chorinea

Guatemala to Colombia

Ithomeis eulema imitatrix

Godman & Salvin, 1878

Northern Mimic-metalmark

Costa Rica & Panama

Isapis agyrtus hera

Godman & Salvin, 1886

Yellow-based Metalmark

S Mexico to Panama

Brachyglenis esthema esthema

C. Felder & R. Felder, 1862

White-rayed Metalmark

Costa Rica & Colombia

Brachyglenis dinora

(H. Bates, 1866)

Dinora Metalmark

Nicaragua to Colombia

Brachyglenis dodone

(Godman & Salvin, 1886)

Orange-bellied Metalmark

S Mexico to Panama

Brachyglenis sp.



Mystery Metalmark

[nr. dodone], Costa Rica &
Panama

Notheme erota diadema

Stichel, 1910

Tawny Metalmark

S Mexico to Bolivia

Monethe albertus rudolphus

Godman & Salvin, 1885

Yellow-patched Metalmark

Honduras to Colombia

Chalodeta chaonitis

(Hewitson, 1866)

Orange-stitched Metalmark

Mexico to Brazil, Trinidad

Chalodeta lypera

(H. Bates, 1868)

Green-stitched Metalmark

Guatemala to Brazil

Metacharis victrix

(Hewitson, 1870)

Victrix Metalmark

Nicaragua to W Ecuador

Metacharis umbrata

Stichel, 1929

Shadowed Metalmark

[DeVries 1997: 163 "umbrata";
pl. 25 "onorata"], Nicaragua to
W Ecuador

Metacharis syloes

Hewitson, 1877

Confused Metalmark

N Venezuela to Bolivia

Metacharis smalli

Hall, 2005

Small's Metalmark

Panama

Cariomothis poeciloptera
poeciloptera

(Godman & Salvin, 1878)

Fiery Metalmark

Costa Rica & Panama

Cariomothis poeciloptera
melusina

(Staudinger, 1888)

Fiery Metalmark

Panama & Colombia

Pheles melanchroia

(C. Felder & R. Felder, 1865)

Melancholy Metalmark

Mexico & Guatemala

Pheles strigosa strigosa

(Staudinger, 1876)

Strigosa Metalmark

S Mexico to Colombia

Pheles incerta

Staudinger, [1887]

Incerta Metalmark

Panama to Peru

Pheles eulesca

(Dyar, 1909)

Dyar's Metalmark

S Mexico to Panama

Pheles ochracea

(Stichel, 1910)

Ochracea Metalmark

Panama to Venezuela

Pheles bicolor

(Godman & Salvin, 1886)

Yellowed Metalmark

Panama, [Costa Rica?]

Syrmatia nyx

(Hübner, [1817])

Spotted Tailwing

Costa Rica to Brazil, Trinidad

Syrmatia aethiops

Staudinger, 1888

White-tipped Tailwing

Costa Rica to Brazil

Chamaelimnas cydonia

Stichel, 1910

Cydonia Metalmark

[= villagomes], Panama, [Costa
Rica?]

Detritivora nicolayi

(J. Hall & Harvey, 2001)

Nicolay's Metalmark

Costa Rica & Panama

Detritivora smalli

(J. Hall & Harvey, 2001)

Small's Metalmark

E Panama to W Ecuador

Detritivora hermodora

(C. Felder & R. Felder, 1861)

Hermodora Metalmark

Costa Rica to Venezuela

Detritivora gynaea

(Godart, [1824])

Gynaea Metalmark

Costa Rica to Brazil

Detritivora barnesi

(J. Hall & Harvey, 2001)

Barnes' Metalmark

SE Mexico to W Ecuador

Charis anius

(Cramer, 1776)

Anius Metalmark

Honduras to Brazil

Calephelis borealis

(Grote & Robinson, 1866)

Northern Metalmark

E OK, MO, IN to VA, local in New
England

Calephelis muticum

McAlpine, 1937

Swamp Metalmark

N AR, MO, KY?, WO to MI, OH

Calephelis virginiensis

(Guérin-Méneville, [1832])

Little Metalmark

E TX to FL, N to VA

Calephelis laverna laverna

(Godman & Salvin, 1886)

Laverna Calephelis

S Mexico to Brazil

Calephelis laverna parva

Austin, 1993

Laverna Calephelis

Osa Peninsula, SW Costa Rica

Calephelis argyrodines

(H. Bates, 1866)

Argyrodines Calephelis

S Mexico to Costa Rica

Calephelis nemesis australis

(W. H. Edwards, 1877)

Fatal Calephelis

S TX, NE Mexico

Calephelis nemesis nemesis

(W. H. Edwards, 1871)

Fatal Calephelis

SE AZ, S NM, NW Mexico

Calephelis nemesis dammersi

McAlpine, 1971

Fatal Calephelis

[= californica], S CA, Baja
California

Calephelis bajaensis

McAlpine, 1971

Baja Calephelis

Baja California Sur

Calephelis exiguus

Austin, 1993

Exigua Calephelis

Costa Rica

Calephelis costaricicola

Strand, 1916

Costa Rican Calephelis

S Mexico to Panama

Calephelis perditalis perditalis

W. Barnes & McDunnough, 1918

Rounded Calephelis

S TX to SE Mexico

Calephelis perditalis donahuei

McAlpine, 1971

Rounded Calephelis

vic. Jacala, Hidalgo, Mexico

Calephelis mexicana

McAlpine, 1971

Mexican Calephelis

E & W Mexico

Calephelis sixola

McAlpine, 1971

Greater Calephelis

Costa Rica

Calephelis wrighti

W. Holland, 1930

Wright’s Calephelis

Baja California, lower Colorado
River drainage

Calephelis fulmen

Stichel, 1910

Fulmen Calephelis

E & W Mexico to Panama

Calephelis rawsoni

McAlpine, 1939

Rawson’s Calephelis

W & S TX, NE Mexico

Calephelis freemani

McAlpine, 1971

Freeman's Calephelis

Davis Mts., Jeff Davis Co., TX

Calephelis arizonensis

McAlpine, 1971

Arizona Calephelis

SE AZ, SW NM, NW Mexico

Calephelis sinaloensis
sinaloensis

McAlpine, 1971

Sinaloan Calephelis

NW Mexico (S Sinaloa)

Calephelis sinaloensis nuevoleon

McAlpine, 1971

Sinaloan Calephelis

NE & N-central Mexico (Nuevo
Leon to Durango)

Calephelis dreisbachi

McAlpine, 1971

Dreisbach's Calephelis

NW Mexico (Sinaloa-Nayarit), [SE
Arizona?]

Calephelis stallingsi

McAlpine, 1971

Stallings' Calephelis

E Mexico to Guatemala

Calephelis matheri

McAlpine, 1971

Mather’s Calephelis

W-central Mexico

Calephelis huasteca

McAlpine, 1971

Huastecan Calephelis

E Mexico

Calephelis montezuma

McAlpine, 1971

Montezuma’s Calephelis

E & W Mexico

Calephelis acapulcoensis

McAlpine, 1971

Acapulco Calephelis

SW Mexico

Calephelis azteca

McAlpine, 1971

Aztec Calephelis

E Mexico

Calephelis yucatana

McAlpine, 1971

Yucatan Calephelis

Yucatan Peninsula and vicinity

Calephelis maya

McAlpine, 1971

Mayan Calephelis

Yucatan Peninsula and vicinity

Calephelis yautepequensis

R. G. De la Maza & Turrent, 1977

Morelos Calephelis

S-central Mexico

Calephelis wellingi wellingi

McAlpine, 1971

Welling's Calephelis

SE Mexico to Belize

Calephelis wellingi baleuensis

McAlpine, 1971

Welling's Calephelis

Alta Verapaz, Guatemala

Calephelis sacapulas

McAlpine, 1971

Sacapulas Calephelis

Guatemala, [S Mexico?]

Calephelis clenchi

McAlpine, 1971

Clench's Calephelis

Guatemala to El Salvador

Calephelis browni

McAlpine, 1971

Brown's Calephelis

S Mexico to Panama

Calephelis sodalis

Austin, 1993

Sodalis Calephelis

Costa Rica, [Panama?]

Calephelis schausi

McAlpine, 1971

Schaus' Calephelis

Honduras to Panama

Calephelis guatemala

McAlpine, 1971

Guatemalan Calephelis

Guatemala

Calephelis tikal

Austin, 1993

Tikal Calephelis

SE Mexico (Yucatan Peninsula) to
Guatemala

Calephelis inca

McAlpine, 1971

Inca Calephelis

Costa Rica to Brazil & Trinidad

Calephelis tapuyo

McAlpine, 1971

Tapuyo Calephelis

possibly Costa Rica

Calephelis candiope

(H. Druce, 1904)

Candiope Calephelis

Costa Rica to Colombia

Calephelis iris

(Staudinger, 1876)

Iris Calephelis

Guatemala to Colombia

Calephelis velutina

(Godman & Salvin, 1878)

Dark Calephelis

S Mexico to Colombia

Parcella amarynthina

(C. Felder & R. Felder, 1865)

Orange-banded Metalmark

Panama to Argentina

Caria ino ino

Godman & Salvin, 1886

Red-bordered Metalmark

W Mexico

Caria ino melicerta

Schaus, 1890

Red-bordered Metalmark

E Mexico, S TX

Caria domitianus vejento

Clench, 1967

Small-patched Metalmark

SE Mexico to Panama

Caria rhacotis

(Godman & Salvin, 1878)

Honduran Metalmark

S Mexico to Peru

Caria stillaticia

Dyar, 1912

Gold-headed Metalmark

[= rabatta], SW Mexico

Caria melino

Dyar, 1912

Melino Metalmark

SE Mexico

Caria mantinea lampeto

Godman & Salvin, 1886

Lampeto Metalmark

S Mexico to Bolivia

Baeotis zonata zonata

R. Felder, 1869

Bumblebee Metalmark

E & W Mexico to Venezuela

Baeotis nesaea

Godman & Salvin, 1889

Nesaea Metalmark

Costa Rica to Peru

Baeotis barce barce

Hewitson, 1875

Barce Metalmark

S Mexico to Honduras

Baeotis sulphurea

(R. Felder, 1869)

Sulphur Metalmark

E Mexico to Guatemala

Baeotis macularia

(Boisduval, 1870)

Sulphur Metalmark

S Mexico (Chiapas) to Colombia

Lasaia meris

(Stoll, 1781)

Variegated Lasaia

S Mexico to Brazil

Lasaia sula sula

Staudinger, 1888

Blue Lasaia

W Mexico to Costa Rica

Lasaia sula peninsularis

Clench, 1972

Blue Lasaia

S TX, E Mexico

Lasaia agesilas callaina

Clench, 1972

Shining-blue Lasaia

E & W Mexico to N Costa Rica

Lasaia agesilas agesilas

(Latreille, [1809])

Shining-blue Lasaia

S Costa Rica to Bolivia & Brazil

Lasaia sessilis

Schaus, 1890

Gray Lasaia

E & S Mexico to Costa Rica

Lasaia pseudomeris

Clench, 1972

Clench's Lasaia

Costa Rica to Venezuela & Peru

Lasaia maria maria

Clench, 1972

Blue-gray Lasaia

W Mexico to Guatemala, Honduras

Lasaia maria anna

Clench, 1972

Blue-gray Lasaia

E Mexico

Lasaia oileus

Godman, 1903

Oily Lasaia

Belize to Paraguay, French
Guiana, Trinidad

Exoplisia hypochalybe
hypochalybe

(C. Felder & R. Felder, 1861)

Many-banded Metalmark

S Mexico to Colombia

Exoplisia cadmeis

(Hewitson, 1866)

Cadmeis Metalmark

Honduras to Peru

Exoplisia azuleja

Callaghan, Llorente & Luis, 2007

Occidental Metalmark

W Mexico

Melanis pixe pixe

(Boisduval, 1836)

Pixie

[= sexpunctata], S TX, E & W
Mexico to Guatemala

Melanis pixe sanguinea

(Stichel, 1910)

Pixie

Honduras to Colombia

Melanis electron melantho

(Ménétriés, 1855)

Electron Metalmark

Belize & Guatemala to Panama

Melanis gynaeceas

(Godman & Salvin, 1889)

Forgotten Metalmark

E & S Mexico (TAMPS, PUE, VER,
GRO)

Melanis cephise

(Ménétriés, 1855)

White-rayed Metalmark

W Mexico to Costa Rica

Melanis acroleuca huasteca

J. White & A. White, 1989

White-tipped Metalmark

E Mexico

Melanis acroleuca acroleuca

(R. Felder, 1869)

White-tipped Metalmark

W Mexico

Mesene phareus

(Cramer, 1777)

Cell-barred Metalmark

S Mexico to Brazil & Peru

Mesene mygdon

Schaus, 1913

Red Metalmark

Costa Rica to Brazil

Mesene croceella

H. Bates, 1865

Guatemalan Metalmark

E & S Mexico to Costa Rica,
[Panama?]

Mesene margaretta jimena

Callaghan, Llorente & Luis, 2011

Zebra-tipped Metalmark

SW Mexico

Mesene margaretta

(A. White, 1843) (El Salvador
segregate)

Zebra-tipped Metalmark

El Salvador

Mesene margaretta margaretta

(A. White, 1843)

Zebra-tipped Metalmark

E Mexico to Colombia

Mesene silaris

Godman & Salvin, 1878

Yellow Metalmark

S Mexico to Peru, Venezuela,
Guianas

Mesene leucopus

Godman & Salvin, 1886

White-legged Metalmark

S Mexico & Guatemala

Mesenopsis bryaxis

(Hewitson, 1870)

Yellow-centered Metalmark

Nicaragua to Colombia

Mesenopsis melanochlora

(Godman & Salvin, 1878)

Orange-striped Metalmark

Nicaragua to Bolivia

Xenandra helius helius

(Cramer, 1779)

Red-striped Metalmark

Costa Rica to Colombia

Xenandra nigrivenata

Schaus, 1913

Black-veined Metalmark

Costa Rica

Xenandra desora

Schaus, 1928

Red-patched Metalmark

Costa Rica & Panama

Esthemopsis clonia

C. Felder & R. Felder, 1865

Clonia Metalmark

S Mexico to Colombia

Esthemopsis alicia alicia

(H. Bates, 1865)

Alicia's Metalmark

S Mexico & Guatemala

Esthemopsis colaxes colaxes

Hewitson, 1870

Cloaxes Metalmark

Costa Rica to W Ecuador

Esthemopsis pherephatte
pherephatte

(Godart, [1824])

Godart's Metalmark

[= caeruleata], S Mexico to
Panama

Chimastrum argenteum argenteum

(H. Bates, 1866)

Silvery Metalmark

S Mexico to Panama

Symmachia probetor championi

Godman & Salvin, 1886

Red-costa Metalmark

S Mexico & Guatemala

Symmachia probetor probetor

(Stoll, 1782)

Red-costa Metalmark

Nicaragua to Colombia

Symmachia rubina rubina

H. Bates, 1866

Rubina Metalmark

S Mexico to Colombia

Symmachia leena leena

Hewitson, 1870

Rusted Metalmark

Nicaragua to Panama

Symmachia accusatrix

Westwood, 1851

Accused Metalmark

Mexico to Colombia, Ecuador,
Brazil

Symmachia threissa threissa

Hewitson, 1870

White-notched Metalmark

Nicaragua to Colombia