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FINDING A TAXON BY NAME: Hold down the CONTROL KEY (on PC) or the COMMAND KEY (on Mac), press "F", then type the taxon name in the search box that appears. Use the arrows next to the search box to scroll through found choices. VIEWING THE THUMBNAIL PAGES: Once you've found a taxon, and its name appears in blue letters, click on it to display thumbnails. For the taxa in black letters we do not have images yet. Thumbnail pages exist for Families, Subfamilies, Tribes, Genera, Species and Subspecies. Caution: with slower internet connection, download of family thumbnails could take a very long time to complete, thus it is best to view species thumbnails. |
SEARCH ENGINE: Google SEARCH BOX at the top of a webpage allows you to search the entire website, not only this webpage. Use it when unsure about the family the taxon is from, or name spelling. VIEWING THE SPECIES PAGES: Click on one of the blue letters to the right of the taxon name and distribution: M - Main Species Page, T - Types, S - Specimens, Ab - Aberrations, Hy - hybrids, HS - historic specimens, L - Live Adults, I - Immatures, H - Habitats and Foodplants, G - genitalia. Extensive image collections are split into several pages as denoted by numbers after letters, e.g. S1 or S2. |
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Family NYMPHALIDAE Rafinesque, 1815 – BRUSHFOOTS Subfamily Libytheinae Libytheana Michener, 1943 Libytheana carinenta (Cramer, 1777) American Snout iN Libytheana carinenta bachmanii (Kirtland, 1851) E North America, [rarely Cuba] M T S L Libytheana carinenta larvata (Strecker, [1878]) C & S TX, NE Mexico, strays far N (regularly to to E CO) M T S1 S2 S3 Ma L1 L2 L3 L4 I H Libytheana carinenta mexicana Michener, 1943 W & S Mexico to Panama M T S L Libytheana carinenta carinenta (Cramer, 1777) South America, [Panama?] M S HS L Libytheana fulvescens (Lathy, 1904) Dominican Snout Dominica M T S Libytheana motya (Hübner, [1823]) Cuban Snout Cuba, [S TX as rare stray] iN M S Libytheana terena (Godart, 1819) Antillean Snout Hispaniola, Jamaica, Puerto Rico, [Cuba as stray?] iN M T S Prolibythea Scudder, [1889] a fossil genus Prolibythea vagabunda Scudder, 1889 Fossil from Colorado Subfamily Danainae Tribe Danaini Archaeolycorea Martins, 1989 a fossil genus Archaeolycorea ferreirai Martins, 1989 Fossil from Brazil (SP) Anetia Hübner, [1823] Anetia briarea (Godart, 1819) Many-spotted King iN Anetia briarea briarea (Godart, 1819) Hispaniola M T S L Anetia briarea numidia Hübner, [1823] Cuba & Isla de Juventud M S L Anetia cubana (Salvin, 1869) Cuban King E Cuba M T S L Anetia jaegeri (Ménétriés, 1832) Hispaniolan King Hispaniola, [Jamaica?] iN M S L I Anetia pantheratus (Martyn, 1797) Great King iN Anetia pantheratus pantheratus (Martyn, 1797) Hispaniola M T S L Anetia pantheratus clarescens (A. Hall, 1925) Cuba M T S Anetia thirza (Geyer, [1833]) Cloud-forest King iN Anetia thirza thirza (Geyer, [1833]) C Mexico to El Salvador M T S L Anetia thirza insignis (Salvin, 1869) Costa Rica & Panama M T S L Lycorea E. Doubleday, [1847] Lycorea halia (Hübner, 1816) iN Lycorea halia halia (Hübner, 1816) TL: Surinam M T Lycorea halia atergatis E. Doubleday, [1847] continental; S TX (rare stray) to N South America, [St. Lucia] M T S Ab L Lycorea halia cinnamomea Weymer, 1883 TL: Brazil (Am) M T Lycorea halia cleobaea (Godart, 1819) Hispaniola, [Jamaica, Puerto Rico] M T S L Lycorea halia demeter C. Felder & R. Felder, 1865 Cuba M T S Lycorea halia discreta Haensch, 1909 TL: Brazil ([MG]) M T Lycorea halia fasciata Haensch, 1909 TL: [Ecuador] M T Lycorea halia pales C. Felder & R. Felder, 1862 TL: Brazil (Am) M T S Lycorea halia transiens N. Riley, 1919 TL: Brazil (Am) M T Lycorea halia [n. ssp.#1] Lamas, MS TL: Peru Lycorea halia [n. ssp.#2] Lamas, MS TL: Trinidad Lycorea ilione (Cramer, 1775) iN Lycorea ilione ilione (Cramer, 1775) TL: Brazil Lycorea ilione albescens (Distant, 1876) S Mexico to Panama M S L Lycorea ilione decolorata (Haensch, 1909) TL: Ecuador M T Lycorea ilione lamirus (Latreille, [1817]) TL: ["Peru"] M T Lycorea ilione suffusca Winhard, 2019 ref.360 Lycorea ilione phenarete (E. Doubleday, [1847]) TL: Bolivia M T Lycorea pasinuntia (Stoll, 1780) iN M T Danaus Kluk, 1780 Danaus cleophile (Godart, 1819) Caribbean Queen Jamaica & Hispaniola iN M T S1 S2 Danaus eresimus (Cramer, 1777) Soldier iN Danaus eresimus eresimus (Cramer, 1777) Grenada & Barbados, possibly only as vagrants from S M T S Danaus eresimus dilucida W. Forbes, 1939 TL: Brazil (Pa) M T Danaus eresimus erginus (Godman & Salvin, 1897) TL: Peru M T Danaus eresimus estevana Talbot, 1943 TL: Venezuela M T Danaus eresimus montezuma Talbot, 1943 continental; S TX to Colombia (strays N) M T S1 S2 L1 L2 Danaus eresimus plexaure (Godart, 1819) TL: Brazil Danaus eresimus tethys W. Forbes, [1944] S Florida, Gr. Antilles (except P. Rico), Grand Cayman M T S L Danaus eresimus [n. ssp.] Lamas, MS Peru Danaus erippus (Cramer, 1775) iN M T Danaus gilippus (Cramer, 1775) iN Danaus gilippus thersippus (H. Bates, 1863) continental; SW USA & TX (strays far N) to Colombia M T S1 S2 Ab Hy L1 L2 L3 L4 I H F Danaus gilippus berenice (Cramer, 1779) SE USA, Bahamas, Cuba, Caymans M S L I Danaus gilippus jamaicensis (H. Bates, 1864) Jamaica M T S Danaus gilippus cleothera (Godart, 1819) Hispaniola, Mona, rarely in W Puerto Rico M T S Danaus gilippus S Lesser Antilles segregate , [nr. xanthippus], S Lesser Antilles M L Danaus gilippus hermippus (C. Felder & R. Felder, 1865) TL: Colombia M T Danaus gilippus wheeleri Talbot, 1943 TL: Colombia M T Danaus gilippus candidus A. Clark, 1941 TL: Peru M T Danaus gilippus xanthippus (C. Felder & R. Felder, 1860) TL: Brazil M T Danaus gilippus nivosus (Godman & Salvin, 1897) TL: Peru M T Danaus gilippus gilippus (Cramer, 1775) TL: Brazil (RJ) Danaus gilippus [n. ssp.] Lamas, MS Grenada Danaus plexippus (Linnaeus, 1758) Monarch iN Danaus plexippus plexippus (Linnaeus, 1758) N & C Am., Cuba, N Baham. (to Crooked I.); migratory M T S1 S2 Hy HS L1 L2 L3 L4 L5 I1 I2 H F Ha1 Ha2 Danaus plexippus leucogyne (A. Butler, 1884) Virgin Islands, N Leeward Is.; possibly a form of D. p. megalippe M T S Danaus plexippus megalippe (Hübner, [1826]) E Cuba, S Bahamas, Gr. Antilles, Virgin Is., L. Antilles, Caymans M S L I Danaus plexippus nigrippus (Haensch, 1909) TL: [Ecuador] M T Danaus plexippus portoricensis A. Clark, 1941 Puerto Rico; possibly only a form of D. p. megalippe M T S Danaus plexippus tobagi A. Clark, 1941 TL: Tobago M T Tribe Ithomiini Elzunia Bryk, 1937 Elzunia humboldt (Latreille, [1809]) iN Elzunia humboldt humboldt (Latreille, [1809]) TL: ["Peru"] M S L Elzunia humboldt flavomaculata (Staudinger, 1885) TL: Colombia M T Elzunia humboldt albomaculata (Haensch, 1903) TL: Ecuador M T Elzunia humboldt cassandrina (Srnka, 1885) TL: Ecuador M T L Elzunia humboldt judsoni R. Fox, 1956 TL: Peru M T S Elzunia humboldt microguttata (Röber, 1927) TL: Colombia M T S Elzunia humboldt carlosi Le Crom & Llorente, 2018 TL: Colombia Elzunia humboldt willmotti Le Crom, Andrade & Llorente, 2018 TL: Colombia Elzunia bomplandii (Guérin-Méneville, [1844]) TL: Colombia Elzunia bomplandii bomplandii (Guérin-Méneville, [1844]) TL: Colombia M S L Elzunia bomplandii atahualpa R. Fox, 1956 TL: Peru M T S Elzunia bomplandii descandollesi (Staudinger, 1885) TL: Colombia M T S Elzunia bomplandii tamasea (Hewitson, 1873) TL: Colombia M T Elzunia bomplandii latreillei (Staudinger, 1885) TL: Colombia M T S Elzunia bomplandii golondrina Willmott, Lamas & J. Hall, 2021 TL: Ecuador Elzunia regalis (Stichel, 1903) TL: Colombia Elzunia regalis regalis (Stichel, 1903) TL: Colombia M T Elzunia regalis joiceyi (Kaye, 1918) TL: Colombia M T Elzunia regalis purace Constantino & Salazar, 2007 TL: Colombia Elzunia regalis quinduensis Constantino & Salazar, 2007 TL: Colombia Elzunia pavonii (A. Butler, 1873) iN M T S L1 L2 Tithorea E. Doubleday, 1847 Tithorea harmonia (Cramer, 1777) iN Tithorea harmonia harmonia (Cramer, 1777) TL: Surinam M T S Tithorea harmonia brunnea Haensch, 1905 TL: Peru M T Tithorea harmonia caissara (J. Zikán, 1941) TL: Brazil (ES) M T Tithorea harmonia cuparina H. Bates, 1862 TL: Brazil (Pa) M T Tithorea harmonia dorada K. Brown, 1977 TL: Venezuela M T S Tithorea harmonia egaensis A. Butler, 1873 TL: Brazil (Am) M T Tithorea harmonia flacilla Godman & Salvin, 1898 TL: Colombia M T Tithorea harmonia furia Staudinger, [1884] TL: Venezuela M T Tithorea harmonia furina Godman & Salvin, 1898 TL: Colombia M T S Tithorea harmonia gilberti K. Brown, 1977 TL: Peru M T Tithorea harmonia helicaon Godman & Salvin, 1879 Nicaragua to Panama M T S Tithorea harmonia hermias Godman & Salvin, 1898 TL: Ecuador M T Tithorea harmonia hermina Haensch, 1903 TL: Ecuador M T Tithorea harmonia hippothous Godman & Salvin, 1879 SE Mexico & Guatemala M T S L Tithorea harmonia irene (Drury, 1782) Panama & Colombia M T S Tithorea harmonia lateflava (Haensch, 1909) TL: Bolivia M T Tithorea harmonia manabiana R. Fox, 1956 TL: Ecuador M T Tithorea harmonia martina R. Fox, 1956 TL: Peru M T L Tithorea harmonia megara (Godart, 1819) TL: Antilles M T Tithorea harmonia melanina Haensch, 1905 TL: Peru M T Tithorea harmonia neitha Hopffer, 1874 TL: Peru M T Tithorea harmonia pseudethra A. Butler, 1873 TL: Brazil M T Tithorea harmonia pseudonyma Staudinger, 1894 TL: Bolivia M T Tithorea harmonia salvadoris Staudinger, 1885 S Chiapas to Honduras & El Salvador M T S Tithorea harmonia sulphurata (J. Zikán, 1941) TL: Brazil (Am) M T Tithorea harmonia [n. ssp.#1] Lamas, MS TL: Colombia Tithorea harmonia [n. ssp.#2] K. Brown, MS TL: Venezuela Tithorea harmonia [n. ssp.#3] K. Brown, MS TL: Brazil (Pe) Tithorea harmonia [n. ssp.#4] K. Brown, MS TL: Brazil (Pa) Tithorea harmonia [n. ssp.#5] K. Brown, MS TL: Brazil (Pa) Tithorea harmonia [n. ssp.#6] K. Brown, MS TL: Guyana Tithorea tarricina Hewitson, [1858] iN Tithorea tarricina tarricina Hewitson, [1858] TL: Colombia M T Tithorea tarricina bonita Haensch, 1903 TL: Ecuador M T Tithorea tarricina duenna H. Bates, 1864 SE Mexico & Guatemala M T S L Tithorea tarricina franciscoi K. Brown, 1977 TL: Venezuela M T S Tithorea tarricina hecalesina C. Felder & R. Felder, 1865 TL: Colombia M T Tithorea tarricina lecromi Vitale & Rodríguez, 2004 ref.298 Tithorea tarricina parola Godman & Salvin, 1898 TL: Colombia M T Tithorea tarricina pinthias Godman & Salvin, 1878 Honduras to Panama M T S L Tithorea tarricina rafaeli Neild, 2008 ref.266 Tithorea tarricina tagarma Hewitson, 1874 TL: Bolivia M T L Tithorea pacifica Willmott & Lamas, 2004 TL: Ecuador Tithorea pacifica pacifica Willmott & Lamas, 2004 TL: Ecuador Tithorea pacifica concordia Willmott & Lamas, 2004 E Panama M T S Tithorea pacifica euphonia Willmott & Lamas, 2004 TL: Colombia M T Aeria Hübner, 1816 Aeria elara (Hewitson, 1855) iN Aeria elara elara (Hewitson, 1855) TL: Brazil M T Aeria elara elarina (Oberthür, 1879) TL: Brazil M T S Aeria elara [n. ssp.#1] Lamas, MS TL: Peru Aeria elara [n. ssp.#2] K. Brown, MS TL: Brazil (SP) Aeria elara [n. ssp.#3] K. Brown, MS TL: Colombia Aeria eurimedia (Cramer, 1777) iN Aeria eurimedia Costa Rican segregate , Costa Rica & Panama M L Aeria eurimedia eurimedia (Cramer, 1777) TL: Surinam Aeria eurimedia agna Godman & Salvin, 1879 Panama to NW Venezuela M T S L Aeria eurimedia latistriga M. Hering, 1925 TL: Colombia Aeria eurimedia negricola (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Aeria eurimedia pacifica Godman & Salvin, 1879 SE Mexico to Honduras M T S1 S2 L Aeria eurimedia sisenna Weymer, 1899 TL: Bolivia M T Aeria eurimedia [n. ssp.] Lamas, MS Costa Rica Aeria olena Weymer, 1875 iN Aeria olena olena Weymer, 1875 TL: Brazil M T Aeria olena [n. ssp.#1] K. Brown, MS TL: Brazil (MT) Aeria olena [n. ssp.#2] K. Brown, MS TL: Brazil (Pe) Athesis E. Doubleday, 1847 Athesis acrisione Hewitson, 1869 iN Athesis acrisione acrisione Hewitson, 1869 TL: Ecuador M T S Athesis acrisione deflavata Niepelt, 1928 TL: Colombia M T Athesis clearista E. Doubleday, 1847 iN Athesis clearista clearista E. Doubleday, 1847 TL: Venezuela M T S L Athesis clearista colombiensis Kaye, 1918 TL: Colombia M T Athesis clearista [n. ssp.] Lamas, MS Colombia Athesis vitrala Kaye, 1918 iN M T Eutresis E. Doubleday, 1847 Eutresis dilucida Staudinger, 1885 Smoky Tigerwing Costa Rica & W Panama M T S Eutresis hypereia E. Doubleday, 1847 iN Eutresis hypereia theope Godman & Salvin, 1877 Costa Rica & Panama M T OI S L Eutresis hypereia Panamanian segregate , Panama M S Eutresis hypereia hypereia E. Doubleday, 1847 TL: Venezuela M T S Eutresis hypereia antioquensis Staudinger, 1885 TL: Colombia M T Eutresis hypereia banosana R. Fox, 1956 TL: Ecuador M T Eutresis hypereia hyspa Godman & Salvin, 1879 TL: Ecuador M T Eutresis hypereia imeriensis K. Brown, 1977 TL: Brazil (Am) Eutresis hypereia imitatrix Staudinger, 1876 TL: Peru M T Eutresis hypereia [n. ssp.] Lamas, MS Panama Athyrtis C. Felder & R. Felder, 1862 Athyrtis mechanitis C. Felder & R. Felder, 1862 iN Athyrtis mechanitis mechanitis C. Felder & R. Felder, 1862 TL: Colombia M T Athyrtis mechanitis oberthueri Srnka, 1885 TL: Ecuador M T Athyrtis mechanitis salvini Srnka, 1884 TL: Peru M T Paititia Lamas, 1979 Paititia neglecta Lamas, 1979 TL: Peru M T Olyras E. Doubleday, 1847 Olyras crathis E. Doubleday, 1847 iN Olyras crathis staudingeri Godman & Salvin, 1897 Costa Rica & Panama M T S Olyras crathis Panamanian segregate , Panama M S Olyras crathis crathis E. Doubleday, 1847 TL: Venezuela M T S Olyras crathis browni Neild, 2008 ref.266 Olyras crathis montagui A. Butler, 1870 TL: Colombia M T Olyras crathis sticheli Haensch, 1905 TL: Peru M T Olyras crathis weeksi R. Fox, 1956 TL: Ecuador M T Olyras crathis [n. ssp.#1] Lamas, MS TL: Panama Olyras crathis [n. ssp.#2] Lamas, MS TL: Colombia Olyras insignis Salvin, 1869 Insignis Tigerwing iN Olyras insignis insignis Salvin, 1869 Costa Rica & Panama M T S Olyras insignis praestans Godman & Salvin, 1897 TL: Colombia M T Olyras insignis translucens Hewitson, 1872 TL: Ecuador M T Olyras theon H. Bates, 1866 Rusty Tigerwing SE Mexico, Guatemala iN M T S L Patricia R. Fox, 1940 Patricia demylus (Godman & Salvin, 1879) iN Patricia demylus demylus (Godman & Salvin, 1879) TL: Ecuador M T S Patricia demylus gemellus R. Fox, 1960 TL: Bolivia M T S Patricia demylus nebulosa Lamas, 1979 TL: Peru M T Patricia demylus [n. ssp.] Lamas, MS Peru Patricia dercyllidas (Hewitson, 1864) iN Patricia dercyllidas dercyllidas (Hewitson, 1864) TL: Colombia M T S Patricia dercyllidas hazelea R. Fox, 1956 TL: Ecuador M T S Patricia dercyllidas [n. ssp.#1] Lamas, MS TL: Peru Patricia dercyllidas [n. ssp.#2] Lamas, MS TL: Peru Patricia dercyllidas [n. ssp.#3] Lamas, MS TL: Peru Patricia hewitsonii (Srnka, 1885) iN M T Patricia oligyrtis (Hewitson, 1877) TL: Ecuador Patricia oligyrtis oligyrtis (Hewitson, 1877) TL: Ecuador M T S Patricia oligyrtis [n. ssp.] Lamas, MS Peru Melinaea Hübner, 1816 Melinaea ethra (Godart, 1819) iN M T S Melinaea idae (C. Felder & R. Felder, 1862) Idae Tigerwing iN Melinaea idae idae (C. Felder & R. Felder, 1862) Panama, Colombia M T S L Melinaea idae vespertina R. Fox, 1945 TL: Ecuador M T Melinaea isocomma W. Forbes, 1948 iN Melinaea isocomma isocomma W. Forbes, 1948 TL: Colombia Melinaea isocomma simulator R. Fox, 1960 TL: Ecuador M T Melinaea isocomma [n. ssp.] Lamas, MS Peru Melinaea lilis (E. Doubleday, 1847) iN Melinaea lilis lilis (E. Doubleday, 1847) TL: Venezuela M T Melinaea lilis dodona Hopffer, 1874 TL: Bolivia M T Melinaea lilis ezra R. Fox, 1939 TL: Colombia M T S Melinaea lilis flavicans C. Hoffmann, 1924 W Mexico (NAY to GRO) M T S Melinaea lilis imitata H. Bates, 1864 E & S Mexico to Costa Rica (Atlantic slope) M T S L Melinaea lilis kayei K. Brown, 1977 TL: Guyana M T Melinaea lilis lateapicalis A. Hall, 1935 TL: Venezuela M T S Melinaea lilis messatis (Hewitson, [1856]) Panama, Colombia M T OI Melinaea lilis parallelis A. Butler, 1873 SE Costa Rica & Panama (Atlantic slope) M T S L Melinaea lilis sola Kaye, 1925 TL: Trinidad M T Melinaea scylax Salvin, 1871 Scylax Tigerwing Costa Rica to Colombia (Pacific slope) iN M T S L Melinaea ludovica (Cramer, 1780) iN Melinaea ludovica ludovica (Cramer, 1780) TL: Surinam M T S Melinaea ludovica manuelito Tessmann, 1928 TL: Peru M T Melinaea ludovica paraiya Reakirt, 1866 TL: Brazil (RJ/SC) Melinaea marsaeus (Hewitson, 1860) iN Melinaea marsaeus marsaeus (Hewitson, 1860) TL: Brazil ([Am]) M T S Melinaea marsaeus clara Rosenberg & Talbot, 1914 TL: Peru M T S Melinaea marsaeus macaria Godman & Salvin, 1898 TL: Colombia M T Melinaea marsaeus messenina C. Felder & R. Felder, 1865 TL: Colombia M T S Melinaea marsaeus mothone (Hewitson, 1860) TL: Colombia M T Melinaea marsaeus phasiana A. Butler, 1870 TL: Peru M T Melinaea marsaeus pothete R.F. d'Almeida, 1945 TL: Brazil (Rd) M T S Melinaea marsaeus rileyi R. Fox, 1942 TL: Peru M T Melinaea marsaeus [n. ssp.] Lamas, MS Peru Melinaea menophilus (Hewitson, [1856]) iN Melinaea menophilus menophilus (Hewitson, [1856]) TL: Colombia M T S Melinaea menophilus cocana Haensch, 1903 TL: Ecuador M T Melinaea menophilus ernestoi K. Brown, 1977 TL: Colombia M T Melinaea menophilus hicetas Godman & Salvin, 1879 TL: Peru M T Melinaea menophilus maenius (Hewitson, 1860) TL: Brazil ([Am]) M T Melinaea menophilus mediatrix Weymer, 1891 TL: French Guiana M T S Melinaea menophilus orestes Salvin, 1871 TL: Peru M T S Melinaea menophilus zaneka A. Butler, 1870 TL: Ecuador M T Melinaea menophilus [n. ssp.#1] Lamas, MS TL: Peru Melinaea menophilus [n. ssp.#2] Lamas, MS TL: Peru Melinaea menophilus [n. ssp.#3] Lamas, MS TL: Brazil (Rd) Melinaea menophilus [n. ssp.#4] K. Brown, MS TL: Brazil (Rd) Melinaea mnasias (Hewitson, [1856]) TL: [Brazil (Pa)] Melinaea mnasias mnasias (Hewitson, [1856]) TL: [Brazil (Pa)] M T S Melinaea mnasias abitagua K. Brown, 1977 TL: Ecuador M T Melinaea mnasias comma W. Forbes, 1927 TL: Peru Melinaea mnasias eratosthenes A. Hall, 1935 TL: French Guiana M T S Melinaea mnasias lucifer H. Bates, 1862 TL: Brazil (Am) M T Melinaea mnasias lutzi R. Fox, 1942 TL: Peru M T Melinaea mnasias neblinae K. Brown, 1977 TL: Brazil (Am) M T Melinaea mnasias romualdo R. Fox, 1965 TL: Peru M T Melinaea mnasias rondonia K. Brown, 1977 TL: Brazil (Rd) M T Melinaea mnasias tecta Haensch, 1909 TL: Guyana M T Melinaea mnasias thera C. Felder & R. Felder, 1865 TL: ? M T Melinaea mnasias [n. ssp.#1] Lamas, MS TL: Peru Melinaea mnasias [n. ssp.#2] K. Brown, MS TL: Venezuela Melinaea mnasias [n. ssp.#3] K. Brown, MS TL: Brazil (Pa) Melinaea mneme (Linnaeus, 1763) iN Melinaea mneme mneme (Linnaeus, 1763) TL: China M T Melinaea mneme mauensis Weymer, 1891 TL: Brazil (Am) M T S Melinaea mneme [n. ssp.#1] K. Brown, MS TL: Venezuela Melinaea mneme [n. ssp.#2] K. Brown, MS TL: Brazil (Rd) Melinaea mnemopsis Berg, 1897 TL: Peru Melinaea mnemopsis mnemopsis Berg, 1897 TL: Peru M T Melinaea mnemopsis [n. ssp.] K. Brown, MS Brazil (Rd) Melinaea satevis (E. Doubleday, 1847) iN Melinaea satevis satevis (E. Doubleday, 1847) TL: Bolivia M T Melinaea satevis aurantia W. Forbes, 1942 TL: Venezuela M T S Melinaea satevis crameri Godman & Salvin, 1898 TL: Guyana M T S Melinaea satevis cydon Godman & Salvin, 1879 TL: Brazil (Am); Peru M T Melinaea satevis flavomacula Weymer, 1894 TL: Brazil (Am) M T Melinaea satevis lamasi K. Brown, 1977 TL: Peru M T S Melinaea satevis maelus (Hewitson, 1860) TL: Brazil ([Am]) M T Melinaea satevis maeonis Hewitson, 1869 TL: Ecuador M T Melinaea satevis tarapotensis Haensch, 1909 TL: Peru M T Melinaea satevis zamora Haensch, 1909 TL: Peru M T S Methona E. Doubleday, 1847 Methona confusa A. Butler, 1873 iN Methona confusa Panamanian segregate , Panama M S L Methona confusa confusa A. Butler, 1873 TL: Brazil (Pa) M T S L Methona confusa psamathe Godman & Salvin, 1898 TL: Ecuador M T S Methona confusa [n. ssp.#1] Lamas, MS TL: Panama Methona confusa [n. ssp.#2] Lamas, MS TL: Panama Methona curvifascia Weymer, 1883 iN M T S Methona grandior (W. Forbes, 1944) iN Methona grandior grandior (W. Forbes, 1944) TL: French Guiana Methona grandior incana (W. Forbes, 1944) TL: Ecuador M T Methona grandior batesi Lamas & Willmott, 2021 ref.318 Methona maxima (W. Forbes, 1944) iN Methona maxima maxima (W. Forbes, 1944) TL: Bolivia M T Methona maxima nigerrima (W. Forbes, 1944) TL: Peru Methona megisto C. Felder & R. Felder, 1860 iN M T S Methona singularis (Staudinger, [1884]) iN M T S Methona themisto (Hübner, 1818) iN Methona themisto themisto (Hübner, 1818) TL: Brazil M T S Methona themisto [n. ssp.] Lamas, MS Brazil (SP) Thyridia Hübner, 1816 Thyridia psidii (Linnaeus, 1758) iN Thyridia psidii melantho H. Bates, 1866 S Mexico (Chiapas) to Panama M T S L Thyridia psidii aedesia E. Doubleday, 1847 TL: Venezuela M T S Thyridia psidii psidii (Linnaeus, 1758) TL: America [Guianas] M OD TI IL Txt Thyridia psidii ino C. Felder & R. Felder, 1862 TL: Brazil (Am) M T S Thyridia psidii cetoides (Rosenberg & Talbot, 1914) TL: Peru M T S Thyridia psidii pallida Godman & Salvin, 1898 TL: Brazil (MT) M T S Thyridia psidii hippodamia (Fabricius, 1775) TL: [Brazil] M T Scada W. F. Kirby, 1871 Scada karschina (Herbst, 1792) iN Scada karschina karschina (Herbst, 1792) TL: Brazil (RJ) M T Scada karschina delicata Talbot, 1932 TL: Brazil? M T Scada kusa (Hewitson, 1872) iN M T Scada reckia (Hübner, [1808]) iN Scada reckia reckia (Hübner, [1808]) TL: ? Scada reckia bernhardi Neild, 2008 ref.266 Scada reckia ethica (Hewitson, [1861]) TL: Ecuador M T S Scada reckia huascara R. Fox, 1967 TL: Colombia Scada reckia junina Bryk, 1953 TL: Peru M T Scada reckia labyrintha Lamas, 1985 TL: Peru M T S Scada reckia majuscula Haensch, 1905 TL: Guyana M T Scada reckia ortygia (H. Druce, 1876) TL: Peru M T Scada reckia philemon (C. Felder & R. Felder, 1865) TL: Venezuela? M T Scada reckia theaphia (H. Bates, 1862) TL: Brazil (Pa) M T Scada reckia ruiza Lamas & Willmott, 2021 ref.318 Scada reckia [n. ssp.#1] K. Brown, MS TL: Brazil (Am) Scada reckia [n. ssp.#2] K. Brown, MS TL: Brazil (Rd) Scada reckia [n. ssp.#3] K. Brown, MS TL: Brazil (Pa) Scada zemira (Hewitson, 1856) iN M T L Scada zibia (Hewitson, 1856) iN Scada zibia zibia (Hewitson, 1856) TL: Colombia M T S Scada zibia batesi Haensch, 1903 TL: Ecuador M T S Scada zibia perpuncta Kaye, 1918 TL: Colombia M T Scada zibia quotidiana Haensch, 1909 TL: Ecuador M T Scada zibia xanthina (H. Bates, 1866) Nicaragua to Panama M T S L Scada zibia zeroca R. Fox, 1967 TL: Ecuador M T L Sais Hübner, 1816 Sais browni Takahashi, 1977 TL: Colombia Sais browni browni Takahashi, 1977 TL: Colombia Sais browni demarmelsi Neild, 2008 ref.266 Sais rosalia (Cramer, 1779) iN Sais rosalia rosalia (Cramer, 1779) TL: Surinam M T Sais rosalia badia Haensch, 1905 TL: Upper Amazon M T Sais rosalia brasiliensis Talbot, 1928 TL: Brazil (MT) M T Sais rosalia camariensis Haensch, 1905 TL: Guyana M T S Sais rosalia klagesi Avinoff, 1926 TL: Brazil (Am) M T Sais rosalia mosella (Hewitson, 1867) TL: Venezuela M T Sais rosalia promissa Weymer, 1883 TL: Brazil (Am) M T Sais rosalia rosalinde Weymer, 1890 TL: Brazil (Pa) M T Sais rosalia schatzi J. Zikán, 1941 TL: Brazil (Am) M T S Sais rosalia virchowii Dewitz, 1877 TL: Venezuela M T S Sais rosalia zitella (Hewitson, 1868) TL: Peru M T Sais rosalia [n. ssp.#1] Lamas, MS TL: Colombia Sais rosalia [n. ssp.#2] Lamas, MS TL: Venezuela Sais rosalia [n. ssp.#3] Lamas, MS TL: Guyana Sais rosalia [n. ssp.#4] K. Brown, MS TL: Brazil Sais rosalia [n. ssp.#5] K. Brown, MS TL: Brazil Forbestra R. Fox, 1967 Forbestra equicola (Cramer, 1780) iN Forbestra equicola equicola (Cramer, 1780) TL: Surinam M T Forbestra equicola equicoloides (Godman & Salvin, 1898) TL: Ecuador; Peru; Brazil (Am) M T S Forbestra equicola [n. ssp.] K. Brown, MS Brazil (Rd) Forbestra olivencia (H. Bates, 1862) iN Forbestra olivencia olivencia (H. Bates, 1862) TL: Brazil (Am) M T S Forbestra olivencia aeneola R. Fox, 1967 TL: Peru M T S Forbestra olivencia juntana (Haensch, 1903) TL: Ecuador M T Forbestra olivencia oiticicai (R.F. d'Almeida, 1951) TL: Brazil (Pa) M T S Forbestra olivencia truncata (A. Butler, 1877) TL: Brazil (Am) M T Forbestra olivencia [n. ssp.] Lamas, MS Peru Forbestra olivencia boliviana Winhard, 2020 ref.361 Forbestra proceris (Weymer, 1883) iN M T Mechanitis Fabricius, 1807 Mechanitis lysimnia (Fabricius, 1793) iN Mechanitis lysimnia lysimnia (Fabricius, 1793) TL: ? M T Mechanitis lysimnia bipuncta W. Forbes, 1948 TL: Venezuela Mechanitis lysimnia elisa (Guérin-Méneville, [1844]) TL: Bolivia M T Mechanitis lysimnia labotas Distant, 1876 Costa Rica M T S L Mechanitis lysimnia limnaea W. Forbes, 1930 TL: French Guiana M T Mechanitis lysimnia macrinus Hewitson, 1860 Panama & Colombia M T S L Mechanitis lysimnia menecles Hewitson, 1860 TL: Brazil (Am) M T Mechanitis lysimnia nesaea Hübner, [1820] TL: ? M T Mechanitis lysimnia neukircheni Neild, 2008 TL: Venezuela Mechanitis lysimnia ocona H. Druce, 1876 TL: Peru M T S Mechanitis lysimnia roqueensis Bryk, 1953 TL: Peru M T Mechanitis lysimnia solaria W. Forbes, 1948 TL: Venezuela Mechanitis lysimnia utemaia Reakirt, 1866 E & SW Mexico to Costa Rica M T S L Mechanitis lysimnia [n. ssp.#1] K. Brown, MS TL: Colombia Mechanitis lysimnia [n. ssp.#2] K. Brown, MS TL: Colombia Mechanitis lysimnia [n. ssp.#3] Lamas, MS TL: Peru Mechanitis mazaeus Hewitson, 1860 iN Mechanitis mazaeus mazaeus Hewitson, 1860 TL: Brazil (Am) M T S Mechanitis mazaeus beebei W. Forbes, 1948 TL: Venezuela Mechanitis mazaeus egaensis H. Bates, 1862 TL: Brazil (Am) M T Mechanitis mazaeus fallax A. Butler, 1873 TL: Colombia M T Mechanitis mazaeus lanei R. Fox, 1967 TL: Brazil (Pa) M T Mechanitis mazaeus pannifera A. Butler, 1877 TL: Brazil (Am/Pa) M T Mechanitis mazaeus pothetoides R.F. d'Almeida, 1951 TL: Brazil (Rd) M T S Mechanitis mazaeus visenda A. Butler, 1877 TL: Brazil (Pa) M T S Mechanitis mazaeus [n. ssp.#1] K. Brown, MS TL: Ecuador Mechanitis mazaeus [n. ssp.#2] K. Brown, MS TL: Brazil (Rd) Mechanitis messenoides C. Felder & R. Felder, 1865 iN Mechanitis messenoides messenoides C. Felder & R. Felder, 1865 TL: Colombia M T S Mechanitis messenoides holmgreni Bryk, 1953 TL: Peru M T S Mechanitis messenoides deceptus A. Butler, 1873 TL: Ecuador M T S Mechanitis menapis Hewitson, [1856] iN Mechanitis menapis menapis Hewitson, [1856] TL: Colombia M T S Mechanitis menapis caribensis R. Fox, 1967 TL: Venezuela M T Mechanitis menapis dariensis K. Brown, 1977 Panama M S Mechanitis menapis doryssus H. Bates, 1864 SE Mexico & Guatemala M T S L I Mechanitis menapis mantineus Hewitson, 1869 TL: Ecuador M T Mechanitis menapis occasiva R. Fox, 1967 TL: Colombia M T Mechanitis menapis saturata Godman, 1901 Honduras to Costa Rica M T S L I Mechanitis menapis [n. ssp.] Lamas, MS Colombia Mechanitis polymnia (Linnaeus, 1758) iN Mechanitis polymnia polymnia (Linnaeus, 1758) TL: Surinam M T S HS Mechanitis polymnia angustifascia Talbot, 1928 TL: Brazil (MT) M T Mechanitis polymnia apicenotata J. Zikán, 1941 TL: Brazil (Am) M T Mechanitis polymnia bolivarensis R. Fox, 1967 TL: Venezuela M T S Mechanitis polymnia casabranca Haensch, 1905 TL: Brazil (MG) M T Mechanitis polymnia caucaensis Haensch, 1909 TL: Colombia M T Mechanitis polymnia chimborazona H. Bates, 1864 TL: Ecuador M T Mechanitis polymnia dorissides Staudinger, [1884] TL: Peru M T Mechanitis polymnia eurydice Haensch, 1905 TL: Peru M T Mechanitis polymnia isthmia H. Bates, 1863 Costa Rica & Panama M T S L I Mechanitis polymnia kayei R. Fox, 1967 TL: Trinidad M T Mechanitis polymnia lycidice H. Bates, 1864 SE Mexico to Honduras M T S L I Mechanitis polymnia mauensis W. Forbes, 1948 TL: Brazil (Pa) Mechanitis polymnia proceriformis Bryk, 1953 TL: Peru M T Mechanitis polymnia veritabilis A. Butler, 1873 TL: Colombia; Venezuela M T S Mechanitis polymnia werneri M. Hering, 1925 TL: Colombia M T Mechanitis polymnia [n. ssp.#1] K. Brown, MS TL: Colombia Mechanitis polymnia [n. ssp.#2] Lamas, MS TL: Colombia Mechanitis polymnia [n. ssp.#3] Lamas, MS TL: Peru Mechanitis polymnia [n. ssp.#4] Lamas, MS TL: Peru Mechanitis polymnia [n. ssp.#5] Lamas, MS TL: Ecuador Mechanitis polymnia [n. ssp.#6] Lamas, MS TL: Brazil Epityches R.F. d'Almeida, 1938 Epityches eupompe (Geyer, 1832) S Brazil; ref.16 iN M T S Epityches ferra (Haensch, 1909) Peru; ref.16 M T S Epityches thyridiana (Haensch, 1909) Bolivia; ref.16 M T S Hyalyris Boisduval, 1870 Hyalyris antea (Hewitson, 1869) iN Hyalyris antea antea (Hewitson, 1869) TL: Ecuador M T Hyalyris antea achuar Vitale & Bollino, 2000 TL: Ecuador M T Hyalyris antea amarilla Vitale & Bollino, 2000 TL: Ecuador M T L Hyalyris antea atagalpa (Haensch, 1905) TL: Peru M T S Hyalyris antea cana (Haensch, 1905) TL: Colombia M T Hyalyris antea flebilis R. Fox, 1971 TL: Venezuela M T Hyalyris antea frater (Salvin, 1869) TL: Peru M T S L Hyalyris antea lalimba Lamas & Willmott, 2021 ref.318 Hyalyris antea [n. ssp.#1] Lamas & Willmott, MS TL: Venezuela Hyalyris antea [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyalyris coeno (E. Doubleday, 1847) iN Hyalyris coeno coeno (E. Doubleday, 1847) TL: Venezuela M T Hyalyris coeno acceptabilis (Weeks, 1902) TL: Bolivia M T Hyalyris coeno angustior (Schaus & T. Cockerell, 1923) TL: Colombia M T Hyalyris coeno atrata R. Fox, 1971 TL: Peru M T Hyalyris coeno avinoffi R. Fox, 1971 TL: Venezuela M T Hyalyris coeno florida (Röber, 1930) TL: Colombia M T Hyalyris coeno norella (Hewitson, [1859]) TL: Ecuador M T Hyalyris coeno norellana (Haensch, 1903) TL: Ecuador M T Hyalyris coeno [n. ssp.] Lamas & Willmott, MS Peru Hyalyris excelsa (C. Felder & R. Felder, 1862) iN Hyalyris excelsa decumana (Godman & Salvin, 1879) Costa Rica & Panama M T S Hyalyris excelsa Panamanian s segregate , Panama M S Hyalyris excelsa excelsa (C. Felder & R. Felder, 1862) TL: Colombia M T Hyalyris excelsa robertus (Bargmann, 1929) TL: Colombia Hyalyris excelsa tachira (T. Racheli & L. Racheli, 1998) TL: Venezuela Hyalyris excelsa [n. ssp.#1] Lamas & Willmott, MS TL: Panama Hyalyris excelsa [n. ssp.#2] Lamas & Willmott, MS TL: Panama Hyalyris excelsa [n. ssp.#3] Willmott & Lamas, MS TL: Ecuador Hyalyris fiammetta (Hewitson, 1852) TL: S. Amer. M T S Hyalyris juninensis R. Fox & Real, 1971 TL: Peru Hyalyris juninensis juninensis R. Fox & Real, 1971 TL: Peru M T Hyalyris juninensis [n. ssp.] Lamas, MS Peru Hyalyris latilimbata (Weymer, 1890) iN M T Hyalyris leptalina (C. Felder & R. Felder, 1865) TL: Brazil Hyalyris leptalina leptalina (C. Felder & R. Felder, 1865) TL: Brazil M T S Hyalyris leptalina [n. ssp.] Lamas, MS Brazil Hyalyris mestra (Hopffer, 1874) iN Hyalyris mestra mestra (Hopffer, 1874) TL: Peru M T Hyalyris mestra fassli R. Fox & Real, 1971 TL: Bolivia Hyalyris mestra personata R. Fox, 1971 TL: Ecuador M T Hyalyris mestra [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyalyris mestra [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyalyris ocna (Herrich-Schäffer, 1865) iN Hyalyris ocna ocna (Herrich-Schäffer, 1865) TL: Colombia M T Hyalyris ocna aurea Lamas & Willmott, 2021 ref.318 Hyalyris ocna equatoria Willmott & Lamas, 2021 ref.318 Hyalyris ocna [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyalyris ocna [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyalyris oulita (Hewitson, [1859]) iN Hyalyris oulita oulita (Hewitson, [1859]) TL: [Peru] M T L Hyalyris oulita lurida (A. Butler, 1873) TL: Peru M T S Hyalyris oulita metella (Hopffer, 1874) TL: Peru M T L Hyalyris oulita [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyalyris oulita [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyalyris oulita [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hyalyris oulita [n. ssp.#4] Lamas & Willmott, MS TL: Bolivia Hyalyris yasunina Willmott, Lamas & J. Hall, 2020 ref.317 Hyalyris praxilla (Hewitson, 1870) iN Hyalyris praxilla praxilla (Hewitson, 1870) TL: Ecuador M T Hyalyris praxilla leuca Willmott & Lamas, 2021 ref.318 Hyalyris praxilla abscondita Lamas, Radford & Willmott, 2021 ref.318 Hyalyris schlingeri Real, 1971 TL: Peru M T Hyalyris lactea Willmott, Lamas & J. Hall, 2020 ref.317 Hyalyris lactea lactea Willmott, Lamas & J. Hall, 2020 ref.317 Hyalyris lactea granatina Willmott & Lamas, 2020 ref.317 Hyalyris lactea crocata Willmott & Lamas, 2020 ref.317 Napeogenes H. Bates, 1862 Napeogenes aethra (Hewitson, 1869) TL: Ecuador M T Napeogenes apulia (Hewitson, 1858) iN Napeogenes apulia apulia (Hewitson, 1858) TL: Colombia M T Napeogenes apulia palmasensis R. Fox & Real, 1971 TL: Peru M T Napeogenes apulia woytkowskii R. Fox & Real, 1971 TL: Peru M T Napeogenes apulia ocnita Willmott & Vitale, 2008 ref.357 Napeogenes apulia sangay Vitale & Willmott, 2008 ref.357 Napeogenes apulia [n. ssp.] Lamas, MS Peru Napeogenes benigna Weymer, 1899 TL: Colombia Napeogenes benigna benigna Weymer, 1899 TL: Colombia M T Napeogenes benigna sandra Vitale & Willmott, 2008 ref.357 Napeogenes cranto C. Felder & R. Felder, 1865 iN Napeogenes cranto paedaretus Godman & Salvin, 1878 Costa Rica & Panama M T S Napeogenes cranto reali R. Fox, 1968 Panama M T S Napeogenes cranto Panamanian segregate , Panama M S Napeogenes cranto cranto C. Felder & R. Felder, 1865 TL: Colombia M T Napeogenes cranto marilenae Vitale & Bollino, 2001 TL: Colombia Napeogenes cranto chocoana Vitale & Bollino, 2008 ref.357 M T Napeogenes duessa (Hewitson, [1859]) iN Napeogenes duessa duessa (Hewitson, [1859]) TL: [Peru] M T S Napeogenes duessa anitae R. Fox & Real, 1971 TL: Ecuador Napeogenes duessa cunza Vitale & Willmott, 2008 ref.357 Napeogenes duessa incas Bryk, 1953 TL: Peru M T Napeogenes duessa jamariensis R.F. d'Almeida, 1951 TL: Brazil (Rd) M T Napeogenes duessa mastersi R. Fox & Real, 1971 TL: Bolivia M T Napeogenes duessa orellana Vitale & Willmott, 2008 ref.357 Napeogenes duessa [n. ssp.#1] Lamas, MS TL: Peru Napeogenes duessa [n. ssp.#2] Lamas, MS TL: Peru Napeogenes duessa [n. ssp.#3] Lamas, MS TL: Peru Napeogenes flossina A. Butler, 1873 TL: Colombia Napeogenes flossina flossina A. Butler, 1873 TL: Colombia M T Napeogenes flossina baumanni R. Fox & Real, 1971 TL: Bolivia M T Napeogenes flossina hypsaea Staudinger, 1885 TL: Colombia M T Napeogenes flossina [n. ssp.#1] Lamas, MS TL: Peru Napeogenes flossina [n. ssp.#2] Lamas, MS TL: Peru Napeogenes garwoodae Willmott & J. Hall, 2020 ref.317 iN Napeogenes glycera Godman, 1899 iN Napeogenes glycera glycera Godman, 1899 TL: Ecuador M T Napeogenes glycera ellariformis Strand, 1912 TL: Peru M T Napeogenes glycera eunomia Godman, 1899 TL: Peru M T Napeogenes glycera nausica Weymer, 1899 TL: Ecuador M T Napeogenes glycera olyrina Haensch, 1905 TL: Bolivia M T Napeogenes glycera mirador Lamas, Willmott & Radford, 2021 ref.318 Napeogenes glycera [n. ssp.] Lamas, MS Peru Napeogenes gracilis Haensch, 1905 TL: Bolivia M T Napeogenes harbona (Hewitson, 1869) iN Napeogenes harbona chiguinda Willmott & Vitale, 2008 ref.357 Napeogenes harbona harbona (Hewitson, 1869) TL: Ecuador M T Napeogenes harbona domiduca (Hewitson, 1876) TL: Bolivia M T S Napeogenes harbona forsteri R. Fox & Real, 1971 TL: Peru M T Napeogenes harbona podocarpus Vitale & Willmott, 2008 ref.357 Napeogenes harbona [n. ssp.#1] Lamas, MS TL: Peru Napeogenes harbona [n. ssp.#2] Lamas, MS TL: Peru Napeogenes harbona [n. ssp.#3] Lamas, MS TL: Peru Napeogenes inachia (Hewitson, 1855) iN Napeogenes inachia inachia (Hewitson, 1855) TL: [Brazil (Pa)] M T Napeogenes inachia johnsoni R. Fox & Real, 1971 TL: Colombia M T Napeogenes inachia moles Haensch, 1905 TL: Guyana M T Napeogenes inachia patientia Lamas, 1985 TL: Peru M T L Napeogenes inachia pozziana (Oberthür, 1879) TL: [Ecuador] M T L Napeogenes inachia pyrois H. Bates, 1862 TL: Brazil (Pa) M T Napeogenes inachia sulphurina H. Bates, 1862 TL: Brazil (Ba) M T S Napeogenes inachia [n. ssp.#1] Lamas, MS TL: Peru Napeogenes inachia [n. ssp.#2] Lamas, MS TL: Peru Napeogenes inachia tijatawaman Brévignon, 2007 ref.176 Napeogenes inachia [n. ssp.#3] K. Brown, MS TL: Brazil Napeogenes inachia [n. ssp.#4] Lamas, MS TL: Brazil (Am) Napeogenes inachia [n. ssp.#5] K. Brown, MS TL: Brazil (Am) Napeogenes inachia [n. ssp.#6] K. Brown, MS TL: Brazil (Rd) Napeogenes inachia [n. ssp.#7] K. Brown, MS TL: Brazil Napeogenes inachia [n. ssp.#8] K. Brown, MS TL: Brazil (Pe) Napeogenes inachia [n. ssp.#9] K. Brown, MS TL: Brazil (MG) Napeogenes inachia juanjuiensis R. Fox & Real, 1971 TL: Peru M T Napeogenes inachia [n. ssp.#10] Lamas, MS TL: Peru Napeogenes inachia [n. ssp.#11] Lamas, MS TL: Peru Napeogenes paruensis R.F. d'Almeida, 1958 TL: Brazil (Pa) Napeogenes paruensis paruensis R.F. d'Almeida, 1958 ref.305 M T Napeogenes paruensis tawaman Brévignon, 2007 ref.304,176 Napeogenes larilla (Hewitson, 1877) iN Napeogenes larilla larilla (Hewitson, 1877) TL: Ecuador M T Napeogenes larilla delavillae Neild, 2008 ref.266 Napeogenes larilla [n. ssp.#1] Lamas, MS TL: Colombia Napeogenes larilla pastazana Vitale & Willmott, 2008 ref.357 Napeogenes larilla reventador Willmott & Vitale, 2008 ref.357 Napeogenes larilla [n. ssp.#2] Lamas, MS TL: Peru Napeogenes larina (Hewitson, [1856]) iN M T Napeogenes lycora (Hewitson, 1870) TL: Ecuador Napeogenes lycora lycora (Hewitson, 1870) TL: Ecuador M T Napeogenes lycora attali Vitale & Willmott, 2008 ref.357 Napeogenes peridia (Hewitson, [1854]) iN Napeogenes peridia hemisticta Schaus, 1913 Costa Rica & W Panama M T S L Napeogenes peridia hemimelaena Godman & Salvin, 1877 Panama M T S Napeogenes peridia peridia (Hewitson, [1854]) TL: Colombia M T S Napeogenes peridia hoppi M. Hering, 1925 TL: Colombia M T S Napeogenes peridia decora Godman, 1899 TL: Ecuador M T S Napeogenes peridia willmotti Vitale & Constantino, 2012 TL: Colombia Napeogenes pharo (C. Felder & R. Felder, 1862) iN Napeogenes pharo pharo (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T S Napeogenes pharo astarte R. Fox, 1943 TL: Brazil (Am) M T Napeogenes pharo azeka (Hewitson, 1869) TL: Colombia M T Napeogenes pharo elva Godman, 1899 TL: Colombia M T Napeogenes pharo gryne R.F. d'Almeida, 1951 TL: Brazil (Am) M T Napeogenes pharo lamia (Hewitson, 1869) TL: Ecuador M T Napeogenes pharo osuna (Hewitson, 1876) TL: Bolivia M T Napeogenes pharo pheranthes H. Bates, 1862 TL: Brazil (Am) M T Napeogenes pharo [n. ssp.#1] Lamas, MS TL: Ecuador Napeogenes pharo [n. ssp.#2] Lamas, MS TL: Peru Napeogenes pharo [n. ssp.#3] K. Brown, MS TL: Brazil Napeogenes pharo [n. ssp.#4] Lamas, MS TL: Brazil (Pa) Napeogenes rhezia (Geyer, [1834]) iN Napeogenes rhezia rhezia (Geyer, [1834]) TL: S. Amer. M T Napeogenes rhezia adelphe H. Bates, 1862 TL: Brazil (Pa) M T Napeogenes rhezia adulta Haensch, 1905 TL: Guyana M T Napeogenes rhezia catamelas W. Forbes, 1942 TL: Venezuela M T Napeogenes rhezia cyrianassa (E. Doubleday, [1847]) TL: Brazil (Pa) M T Napeogenes rhezia tunantina H. Bates, 1862 TL: Brazil (Am) M T S Napeogenes rhezia yanetta (Hewitson, 1867) TL: Brazil (ES) M T Napeogenes rhezia [n. ssp.#1] K. Brown, MS TL: Venezuela Napeogenes rhezia itu Brévignon, 2007 ref.176 Napeogenes rhezia [n. ssp.#2] K. Brown, MS TL: Brazil (Pa) Napeogenes rhezia [n. ssp.#3] K. Brown, MS TL: Brazil (Am) Napeogenes rhezia [n. ssp.#4] K. Brown, MS TL: Brazil (Rd) Napeogenes rhezia achaea (Hewitson, 1869) TL: Ecuador M T Napeogenes rhezia curvilutea Kaye, 1918 TL: Peru M T Napeogenes rhezia nicolayi R. Fox & Real, 1971 TL: Colombia M T S Napeogenes rhezia [n. ssp.#5] Lamas, MS TL: Peru Napeogenes rhezia [n. ssp.#6] Lamas, MS TL: Peru Napeogenes sodalis Haensch, 1905 TL: Peru M T Napeogenes stella (Hewitson, [1855]) iN Napeogenes stella Panamanian segregate , Panama M S L Napeogenes stella stella (Hewitson, [1855]) TL: Colombia M T S Napeogenes stella aster Godman, 1899 TL: Ecuador M T Napeogenes stella browni T. Racheli & L. Racheli, 1998 TL: Venezuela Napeogenes stella opacella Kaye, 1918 TL: Colombia M T S Napeogenes stella delgadoi Vitale & Constantino, 2012 ref.297 M T Napeogenes stella caucaensis Constantino & Vitale, 2012 ref.297 M T Napeogenes sulphureophila Bryk, 1937 TL: not specified M T Napeogenes sylphis (Guérin-Méneville, [1844]) iN Napeogenes sylphis sylphis (Guérin-Méneville, [1844]) TL: Bolivia Napeogenes sylphis acreana R.F. d'Almeida, 1958 TL: Brazil (Ac) M T Napeogenes sylphis corena (Hewitson, [1861]) TL: Brazil (Am) M T Napeogenes sylphis ercilla (Hewitson, 1858) TL: Brazil (Am) M T Napeogenes sylphis ithra (Hewitson, 1855) TL: [Brazil (Pa)] M T Napeogenes sylphis potaronus Kaye, 1905 TL: Guyana M T Napeogenes sylphis rindgei R. Fox & Real, 1971 TL: Peru M T Napeogenes sylphis thira (Hewitson, 1874) TL: Peru M T Napeogenes sylphis [n. ssp.#1] Lamas, MS TL: Peru Napeogenes sylphis [n. ssp.#2] Lamas, MS TL: Brazil (Am) Napeogenes sylphis ehu Brévignon, 2007 ref.176 Napeogenes tolosa (Hewitson, 1855) Tolosa Tigerwing iN Napeogenes tolosa tolosa (Hewitson, 1855) SE Mexico, Guatemala, Honduras M T S L Napeogenes tolosa tepehua (R. de la Maza & J. de la Maza, 2018) ref.245 Napeogenes tolosa mombachoensis Brabant & Maes, 1997 Nicaragua M T L Napeogenes tolosa amara Godman, 1899 Nicaragua to Panama M T S L Napeogenes tolosa Panamanian s segregate , Panama M S Napeogenes tolosa chrispina (Hewitson, 1874) TL: Colombia M T Napeogenes tolosa diaphanosa Kaye, 1918 TL: Colombia M T Napeogenes tolosa panamensis Vitale, Constantino & Delgado, 2012 ref.297 M T Napeogenes tolosa [n. ssp.] Lamas, MS Panama Napeogenes verticilla (Hewitson, 1874) TL: Peru M T Napeogenes zurippa (Hewitson, [1876]) iN Napeogenes zurippa zurippa (Hewitson, [1876]) TL: Bolivia M T Napeogenes zurippa deucalion Haensch, 1905 TL: Peru M T S Napeogenes zurippa iquitensis Staudinger, [1884] TL: Peru M T Napeogenes zurippa otaxes Godman, 1899 TL: Peru M T S Napeogenes zurippa pyrrho H. Druce, 1876 TL: Peru M T Napeogenes zurippa [n. ssp.] K. Brown, MS Brazil (Rd) Napeogenes zurippa quadrilis Haensch, 1903 TL: Ecuador M T S Hypothyris Hübner, 1821 Hypothyris anastasia (H. Bates, 1862) iN Hypothyris anastasia anastasia (H. Bates, 1862) TL: Brazil (Am) M T Hypothyris anastasia acreana R.F. d'Almeida, 1958 TL: Brazil (Ac) M T Hypothyris anastasia anastasina (Staudinger, 1885) TL: Peru M T Hypothyris anastasia arpi (R.F. d'Almeida, 1958) TL: Brazil (Pa) M T Hypothyris anastasia bicolora (Haensch, 1903) TL: Ecuador M T Hypothyris anastasia castanea (A. Butler, 1877) TL: Brazil (Am) M T Hypothyris anastasia honesta (Weymer, 1883) TL: Ecuador M T S Hypothyris anastasia niphas R.F. d'Almeida, 1945 TL: Brazil (Rd) M T Hypothyris anastasia porsenna (Srnka, 1885) TL: Peru M T Hypothyris anastasia [n. ssp.#1] Lamas, MS TL: Peru Hypothyris anastasia [n. ssp.#2] Lamas, MS TL: Peru Hypothyris cantobrica (Hewitson, 1876) iN Hypothyris cantobrica cantobrica (Hewitson, 1876) TL: Bolivia M T Hypothyris cantobrica carvalhoi (R.F. d'Almeida, 1954) TL: Brazil (Pa) M T Hypothyris cantobrica nundina (R.F. d'Almeida, 1945) TL: Brazil (Rd) M T Hypothyris cantobrica schunkeae (Lamas, 1979) TL: Peru M T Hypothyris cantobrica zamorita Willmott, Boyer & Vitale, 2021 ref.318 Hypothyris cantobrica [n. ssp.#1] K. Brown, MS TL: Brazil Hypothyris cantobrica [n. ssp.#2] K. Brown, MS TL: Brazil (Pa) Hypothyris cantobrica [n. ssp.#3] K. Brown, MS TL: Brazil (Pa) Hypothyris connexa (A. Hall, 1939) TL: Guyana Hypothyris connexa connexa (A. Hall, 1939) TL: Guyana M T Hypothyris connexa lema K. Brown, 1977 TL: Venezuela M T Hypothyris connexa [n. ssp.] K. Brown, MS Venezuela Hypothyris daphnis R.F. d'Almeida, 1945 iN Hypothyris daphnis daphnis R.F. d'Almeida, 1945 TL: Brazil (Rd) M T S Hypothyris daphnis amapaensis K. Brown, 1980 TL: Brazil (Ap) M T Hypothyris daphnis clenchi K. Brown, 1980 TL: Brazil (Pa) M T Hypothyris daphnis daphnoides R.F. d'Almeida, 1945 TL: Brazil (Pa) M T Hypothyris daphnis madeira K. Brown, 1980 TL: Brazil (Am) M T Hypothyris euclea (Godart, 1819) iN Hypothyris euclea valora (Haensch, 1909) S Mexico to Panama M T S L Hypothyris euclea Costa Rican segregate , Costa Rica M S L Hypothyris euclea euclea (Godart, 1819) TL: Antilles M T S Hypothyris euclea barii (H. Bates, 1862) TL: Brazil (Pa) M T Hypothyris euclea caldasensis R. Fox, 1971 TL: Colombia M T Hypothyris euclea callanga (Haensch, 1905) TL: Peru M T Hypothyris euclea forbesi R. Fox, 1941 TL: Venezuela M T Hypothyris euclea intermedia (A. Butler, 1873) TL: Colombia M T Hypothyris euclea interrupta (J. Zikán, 1941) TL: Brazil (Am) M T Hypothyris euclea laphria (E. Doubleday, 1847) TL: Brazil M S Hypothyris euclea micheneri R. Fox, 1971 TL: Colombia M T Hypothyris euclea nina (Haensch, 1905) TL: Bolivia M T Hypothyris euclea nonia R.F. d'Almeida, 1945 TL: Brazil (Ac) M T Hypothyris euclea peruviana (Staudinger, 1885) TL: Peru M T L Hypothyris euclea philetaera (Hewitson, 1876) TL: ? M T Hypothyris euclea pyrippe (Hopffer, 1874) TL: Peru M T Hypothyris euclea [n. ssp.#1] Lamas, MS TL: Peru Hypothyris euclea [n. ssp.#2] Lamas, MS TL: Peru Hypothyris euclea [n. ssp.#3] Lamas, MS TL: Peru Hypothyris euclea [n. ssp.#4] Lamas, MS TL: Peru Hypothyris euclea [n. ssp.#5] K. Brown, MS TL: Brazil (Rd) Hypothyris fluonia (Hewitson, 1854) iN Hypothyris fluonia fluonia (Hewitson, 1854) TL: Brazil (Am) M T S Hypothyris fluonia baiochii T. Racheli & L. Racheli, 1998 TL: Venezuela Hypothyris fluonia berna (Haensch, 1903) TL: Ecuador M T S L Hypothyris fluonia flavigera (N. Riley, 1919) TL: Brazil (Am) M T Hypothyris fluonia fulvifascia (Talbot, 1932) TL: Peru M T Hypothyris fluonia hygia (Godman, 1899) TL: Surinam M T Hypothyris fluonia iberina R.F. d'Almeida, 1945 TL: Brazil (Rd) M T S Hypothyris fluonia manaos (H. Bates, 1862) TL: Brazil (Am) M T Hypothyris fluonia pardalina (Hopffer, 1874) TL: Peru M T Hypothyris fluonia rowena (Hewitson, [1857]) TL: Colombia M T Hypothyris fluonia seminigra (Rosenberg & Talbot, 1914) TL: Peru M T S Hypothyris fluonia uchiza Lamas, 1979 TL: Peru M T Hypothyris fluonia viola (Haensch, 1905) TL: Bolivia M T Hypothyris fluonia violantilla R.F. d'Almeida, 1952 TL: Brazil (MT) M T Hypothyris fluonia [n. ssp.#1] Lamas, MS TL: Ecuador Hypothyris fluonia [n. ssp.#2] Lamas, MS TL: Peru Hypothyris fluonia [n. ssp.#3] K. Brown, MS TL: Brazil (Am) Hypothyris fluonia [n. ssp.#4] K. Brown, MS TL: Brazil (Pa) Hypothyris fluonia [n. ssp.#5] K. Brown, MS TL: Brazil (Pa) Hypothyris gemella R. Fox, 1971 TL: Guyana Hypothyris gemella gemella R. Fox, 1971 TL: Guyana M T Hypothyris gemella mossi Neild, 2008 ref.266 Hypothyris gemella foxi Neild, 2008 ref.266 Hypothyris gemella batesi Neild, 2008 ref.266 Hypothyris gemella anduzei Neild, 2008 ref.266 Hypothyris gemella pseudoglycon Neild, 2008 ref.266 Hypothyris gemella [n. ssp.] K. Brown, MS Brazil Hypothyris leprieuri (Feisthamel, 1835) iN Hypothyris leprieuri leprieuri (Feisthamel, 1835) TL: French Guiana Hypothyris leprieuri catilla (Hewitson, 1876) TL: Bolivia M T S Hypothyris leprieuri ignorata (Haensch, 1905) TL: Brazil (Pa) M T Hypothyris leprieuri michaelisi (Haensch, 1909) TL: Brazil (Pa) M T Hypothyris leprieuri nemea (Weymer, 1899) TL: Colombia M T Hypothyris leprieuri ninyas R.F. d'Almeida, 1945 TL: Brazil (Rd) M T Hypothyris leprieuri [n. ssp.#1] Lamas, MS TL: Peru Hypothyris leprieuri [n. ssp.#2] K. Brown, MS TL: Brazil Hypothyris leprieuri [n. ssp.#3] K. Brown, MS TL: Brazil (Pa) Hypothyris leprieuri [n. ssp.#4] K. Brown, MS TL: Brazil (Pa) Hypothyris leprieuri [n. ssp.#5] K. Brown, MS TL: Brazil Hypothyris leprieuri [n. ssp.#6] K. Brown, MS TL: Brazil Hypothyris lycaste (Fabricius, 1793) Lycaste Tigerwing iN Hypothyris lycaste dionaea (Hewitson, 1854) SE Mexico & Guatemala M T S L Hypothyris lycaste callispila (H. Bates, 1866) Costa Rica & Panama M T IL Hypothyris lycaste Panamanian segregate , Panama M S Hypothyris lycaste lycaste (Fabricius, 1793) Panama & Colombia M T IL L Hypothyris lycaste antonia (Hewitson, 1869) TL: Ecuador M T Hypothyris lycaste fraterna (Haensch, 1909) TL: Venezuela M T Hypothyris lycaste glabra (Godman, 1899) TL: Colombia M T Hypothyris lycaste limosa R. Fox, 1971 TL: Colombia M T Hypothyris lycaste limpida (Haensch, 1905) TL: Colombia M T Hypothyris lycaste mergelena (Hewitson, [1855]) TL: Colombia M T Hypothyris lycaste [n. ssp.] Lamas, MS Panama Hypothyris mamercus (Hewitson, 1869) iN Hypothyris mamercus mamercus (Hewitson, 1869) TL: Ecuador M T S Hypothyris mamercus maenas (Haensch, 1909) TL: [Peru] M T Hypothyris mamercus poemne R.F. d'Almeida, 1939 TL: Brazil (Pa) M T Hypothyris mamercus polymnides (Haensch, 1905) TL: Colombia M T Hypothyris mamercus [n. ssp.#1] Lamas, MS TL: Peru Hypothyris mamercus [n. ssp.#2] Lamas, MS TL: Peru Hypothyris mamercus [n. ssp.#3] K. Brown, MS TL: Brazil (Rd) Hypothyris mamercus [n. ssp.#4] K. Brown, MS TL: Brazil Hypothyris mamercus [n. ssp.#5] K. Brown, MS TL: Brazil (Pa) Hypothyris mansuetus (Hewitson, 1860) iN Hypothyris mansuetus mansuetus (Hewitson, 1860) TL: [Peru] M T S Hypothyris mansuetus amica (Weymer, 1883) TL: Colombia M T S Hypothyris mansuetus deemae R. Fox, 1943 TL: Peru M T Hypothyris mansuetus klotsi R. Fox, 1941 TL: Peru M T Hypothyris mansuetus meterus (Hewitson, 1860) TL: [Peru] M T Hypothyris mansuetus zephyrus R. Fox, 1945 TL: Ecuador M T Hypothyris mansuetus [n. ssp.] Lamas, MS Peru Hypothyris moebiusi (Haensch, 1903) iN Hypothyris moebiusi moebiusi (Haensch, 1903) TL: Ecuador M T Hypothyris moebiusi unicolora (Tessmann, 1928) TL: Peru M T Hypothyris moebiusi [n. ssp.] Lamas, MS Peru Hypothyris ninonia (Hübner, [1806]) iN Hypothyris ninonia ninonia (Hübner, [1806]) TL: ? Hypothyris ninonia aetha R. Fox & Real, 1971 TL: Bolivia M T Hypothyris ninonia antonina (Staudinger, [1884]) TL: Peru M T Hypothyris ninonia apollinis (Staudinger, [1884]) TL: Peru M T Hypothyris ninonia caquetacola R. Fox, 1971 TL: Colombia M T Hypothyris ninonia colophonia R.F. d'Almeida, 1945 TL: Brazil (Rr) M T Hypothyris ninonia completa (Haensch, 1909) TL: Brazil (Pa) M T Hypothyris ninonia completomaculata (J. Zikán, 1941) TL: Brazil (Am) M T Hypothyris ninonia cornelie (Guérin-Méneville, [1844]) TL: Bolivia M T Hypothyris ninonia daeta (Boisduval, 1836) TL: Brazil M T Hypothyris ninonia daetina (Weymer, 1899) TL: Brazil (Ba) M T Hypothyris ninonia diphes R. Fox, 1971 TL: Colombia M T Hypothyris ninonia fimbria (Hewitson, 1855) TL: Colombia M T Hypothyris ninonia granadensis (Haensch, 1905) TL: Colombia M T Hypothyris ninonia latefasciata (Haensch, 1909) TL: Upper Amazon M T Hypothyris ninonia latipennis (Tessmann, 1928) TL: Peru M T Hypothyris ninonia matouryensis Brévignon, 2007 ref.176 Hypothyris ninonia mutilla (Hewitson, 1867) TL: Guyana M T Hypothyris ninonia mysotis (Haensch, 1909) TL: Venezuela M T Hypothyris ninonia neimyi (N. Riley, 1919) TL: Brazil (Rd) M T Hypothyris ninonia pellucida (Haensch, 1905) TL: Guyana M T Hypothyris ninonia vallina (Haensch, 1909) TL: Venezuela M T Hypothyris ninonia [n. ssp.#1] Lamas, MS TL: Peru Hypothyris ninonia [n. ssp.#2] Lamas, MS TL: Peru Hypothyris ninonia [n. ssp.#3] Lamas, MS TL: Peru Hypothyris ninonia [n. ssp.#4] K. Brown, MS TL: Brazil (Pa) Hypothyris ninonia [n. ssp.#5] K. Brown, MS TL: Brazil (Am) Hypothyris ninonia [n. ssp.#6] K. Brown, MS TL: Brazil (Am) Hypothyris ninonia [n. ssp.#7] K. Brown, MS TL: Brazil (Am) Hypothyris semifulva (Salvin, 1869) iN Hypothyris semifulva semifulva (Salvin, 1869) TL: Ecuador M T Hypothyris semifulva angelina (Haensch, 1905) TL: Peru M T S Hypothyris semifulva dalmeidai R. Fox & Real, 1971 TL: Brazil (Rd) M T Hypothyris semifulva fulminans (A. Butler, 1873) TL: Colombia M T Hypothyris semifulva meteroides R. Fox, 1971 TL: Peru M T Hypothyris semifulva pallisteri R. Fox & Real, 1971 TL: Peru M T S Hypothyris semifulva putumayoensis R. Fox & Real, 1971 TL: Colombia M T Hypothyris semifulva satura (Haensch, 1903) TL: Ecuador M T Hypothyris semifulva soror (Srnka, 1885) TL: Peru M T Hypothyris semifulva virgilini (N. Riley, 1919) TL: Brazil (Ac) M T Hypothyris semifulva [n. ssp.#1] Lamas, MS TL: Ecuador Hypothyris semifulva [n. ssp.#2] Lamas, MS TL: Peru Hypothyris thea (Hewitson, 1852) TL: Brazil (Am) Hypothyris thea thea (Hewitson, 1852) TL: Brazil (Am) M T Hypothyris thea mayi R.F. d'Almeida, 1945 TL: Brazil (Pa) M T S Hypothyris thea theatina (Haensch, 1909) TL: Upper Amazon M T Hypothyris thea [n. ssp.#1] Lamas, MS TL: Peru Hypothyris thea [n. ssp.#2] K. Brown, MS TL: Bolivia Hypothyris vallonia (Hewitson, [1853]) TL: Brazil (Pa) Hypothyris vallonia vallonia (Hewitson, [1853]) TL: Brazil (Pa) M T Hypothyris vallonia glycon (Godman, 1899) TL: Guyana; Surinam M T Hypothyris vallonia [n. ssp.#1] K. Brown, MS TL: Brazil Hypothyris vallonia [n. ssp.#2] K. Brown, MS TL: Brazil (Pa) Hypothyris xanthostola (H. Bates, 1862) TL: Brazil (Pa) Hypothyris xanthostola xanthostola (H. Bates, 1862) TL: Brazil (Pa) M T S Hypothyris xanthostola bellatula (R.F. d'Almeida, 1945) TL: Brazil (Rd) M T Hypothyris xanthostola desmora Bryk, 1937 TL: Brazil (Pa) M T Hypothyris xanthostola ipoti Brévignon, 2007 ref.176 Hypothyris [n. sp.] Lamas, MS Peru Pagyris Boisduval, 1870 Pagyris euryanassa (C. Felder & R. Felder, 1860) ref.16 iN M T Pagyris cymothoe (Hewitson, [1855]) iN Pagyris cymothoe cymothoe (Hewitson, [1855]) TL: Venezuela M T S L Pagyris cymothoe sylvella (Hewitson, 1868) TL: [Ecuador] M T S Pagyris priscilla Lamas, 1986 TL: Bolivia M T Pagyris renelichyi Neild, 2008 Venezuela; ref.266 Pagyris ulla (Hewitson, [1857]) iN Pagyris ulla ulla (Hewitson, [1857]) TL: Colombia M T Pagyris ulla carmilla Lamas, 1986 TL: Peru M T Pagyris ulla dimidiata (Staudinger, 1885) TL: Colombia M T Pagyris ulla leonila Lamas, 1986 TL: Peru M T Pagyris ulla vanilla Lamas, 1986 TL: Ecuador M T Pagyris ulla zorrilla Lamas, 1986 TL: Ecuador M T Ithomia Hübner, 1816 Ithomia agnosia Hewitson, [1855] iN Ithomia agnosia agnosia Hewitson, [1855] TL: Venezuela M T S Ithomia agnosia pellucida Weymer, 1875 TL: Trinidad M T Ithomia agnosia zikani R.F. d'Almeida, 1940 TL: Brazil (MG) M T S Ithomia agnosia zumba Lamas & Willmott, 2021 ref.318 Ithomia amarilla Haensch, 1903 iN M T S L Ithomia arduinna R.F. d'Almeida, 1952 iN Ithomia arduinna arduinna R.F. d'Almeida, 1952 TL: Bolivia M T S L Ithomia arduinna ardua Lamas, Vitale & Boyer, 2021 ref.318 Ithomia arduinna [n. ssp.] Lamas, MS Peru Ithomia avella Hewitson, 1854 iN Ithomia avella avella Hewitson, 1854 TL: Colombia M T Ithomia avella adamsi Neild, 2008 ref.266 Ithomia avella cesleria Hewitson, [1855] TL: Colombia M T Ithomia avella deliciae R. Fox, 1941 TL: Colombia Ithomia avella epona Hewitson, 1869 TL: Ecuador M T S Ithomia avella katherineae R. Fox, 1941 TL: Colombia M T Ithomia avella miraculosa Lamas, 2003 TL: Colombia M T Ithomia avella roberti Neild, 2008 ref.266 Ithomia avella salazari Vitale & Bollino, 2000 TL: Ecuador M T Ithomia celemia Hewitson, [1854] iN Ithomia celemia plaginota A. Butler & H. Druce, 1872 Costa Rica & Panama M T S Ithomia celemia Panamanian segregate , Panama M S Ithomia celemia celemia Hewitson, [1854] TL: Colombia M T S Ithomia celemia lurida Haensch, 1909 TL: Colombia M T Ithomia celemia [n. ssp.] Lamas, MS Panama Ithomia cleora Hewitson, 1855 iN M T S L Ithomia diasia Hewitson, 1854 iN Ithomia diasia diasia Hewitson, 1854 TL: Colombia M T Ithomia diasia galata Hewitson, 1855 Panama to Colombia M T S Ithomia diasia hippocrenis H. Bates, 1866 Nicaragua to Panama M T S L Ithomia diasia leucophaena M. Hering, 1925 TL: Colombia Ithomia diasia morena Haensch, 1903 TL: Ecuador M T S Ithomia diasia ossuna Haensch, 1909 TL: Colombia M T Ithomia drymo Hübner, 1816 iN M T Ithomia eleonora Haensch, 1905 iN M T S Ithomia ellara Hewitson, 1874 iN M T Ithomia heraldica H. Bates, 1866 Heraldica Clearwing iN Ithomia heraldica heraldica H. Bates, 1866 Costa Rica & Panama M T S L Ithomia heraldica Panamanian segregate , Panama M S L Ithomia hyala Hewitson, [1856] iN Ithomia hyala Panamanian segregate , Costa Rica & Panama M S Ithomia hyala hyala Hewitson, [1856] TL: Ecuador M T S Ithomia hyala sincerus Bargmann, 1929 TL: [Colombia] M T Ithomia hyala [n. ssp.] Lamas, MS Panama Ithomia iphianassa E. Doubleday, 1847 iN Ithomia iphianassa iphianassa E. Doubleday, 1847 TL: Venezuela M T S Ithomia iphianassa alienassa Haensch, 1905 TL: Colombia M T S Ithomia iphianassa anaphissa Herrich-Schäffer, 1865 TL: Colombia M T Ithomia iphianassa ethilla Neustetter, 1929 TL: Colombia M T Ithomia iphianassa florencia Winhard, 2020 ref.361 Ithomia iphianassa panamensis H. Bates, 1863 Panama M T OI S L Ithomia iphianassa phanessa Herrich-Schäffer, 1865 TL: Colombia M T S Ithomia iphianassa occidentissa Lamas & Willmott, 2021 ref.318 Ithomia jucunda Godman & Salvin, 1878 Jucunda Clearwing iN Ithomia jucunda jucunda Godman & Salvin, 1878 Panama M T S Ithomia jucunda bolivari Schaus, 1913 Nicaragua to Panama (Atlantic slope) M T S L Ithomia jucunda centromaculata Weymer, 1899 Panama & Colombia M T OI S Ithomia jucunda lamasi Vitale & Bollino, 2000 TL: Ecuador M T Ithomia jucunda [n. ssp.] Lamas, MS Colombia Ithomia lagusa Hewitson, [1856] iN Ithomia lagusa lagusa Hewitson, [1856] TL: Colombia M T Ithomia lagusa adelinda Hewitson, 1868 TL: Ecuador M T Ithomia lagusa linda Hewitson, 1872 TL: Ecuador M T Ithomia lagusa peruana Salvin, 1869 TL: Peru M T S L Ithomia lagusa rachelii Vitale & Bollino, 2000 TL: Ecuador M T Ithomia lagusa theuda Hewitson, 1872 TL: Ecuador M T Ithomia lagusa [n. ssp.#1] Lamas, MS TL: Colombia Ithomia lagusa [n. ssp.#2] Lamas, MS TL: Colombia Ithomia lagusa [n. ssp.#3] Lamas, MS TL: Colombia Ithomia hymettia (Staudinger, 1885) ref.318 Ithomia hymettia hymettia (Staudinger, 1885) ref.318 M T S Ithomia hymettia petitia Lamas & Willmott, 2021 ref.318 Ithomia leila Hewitson, 1852 Leila’s Clearwing SE Mexico & Guatemala iN M T S L Ithomia lichyi R.F. d'Almeida, 1939 iN Ithomia lichyi lichyi R.F. d'Almeida, 1939 TL: Brazil ([RJ]) M T Ithomia lichyi dalmeidai R. Fox, 1941 TL: Peru M T Ithomia lichyi neivai R.F. d'Almeida, 1940 TL: Brazil (Ac) M T S Ithomia lichyi [n. ssp.] Lamas, MS Brazil Ithomia patilla Hewitson, 1852 Patilla Clearwing SE Mexico to Panama iN M T S L Ithomia praeithomia Vitale & Bollino, 2003 TL: Peru Ithomia praeithomia praeithomia Vitale & Bollino, 2003 TL: Peru Ithomia praeithomia [n. ssp.] Lamas, MS Peru Ithomia pseudoagalla Rebel, 1902 iN M T S L Ithomia salapia Hewitson, [1853] iN Ithomia salapia salapia Hewitson, [1853] TL: Ecuador M T S Ithomia salapia aquinia Hopffer, 1874 TL: Bolivia M T L Ithomia salapia ardea Hewitson, 1855 TL: Bolivia M T Ithomia salapia derasa Hewitson, 1855 TL: Nicaragua M T S Ithomia salapia diaphana Bryk, 1937 TL: Peru M T S Ithomia terra Hewitson, [1853] iN Ithomia terra terra Hewitson, [1853] TL: Colombia M T S Ithomia terra jasinskii Vitale & Bollino, 2000 TL: Ecuador M T Ithomia terra oenanthe Weymer, 1899 TL: Colombia M T Ithomia terra terrana Haensch, 1903 TL: Ecuador M T Ithomia terra vulcana Haensch, 1909 Costa Rica & Panama M T S L Ithomia terra [n. ssp.#1] Lamas, MS TL: Colombia Ithomia terra [n. ssp.#2] Lamas, MS TL: Peru Ithomia xenos (H. Bates, 1866) Xenos Clearwing iN Ithomia xenos xenos (H. Bates, 1866) Costa Rica & Panama M T S L Ithomia xenos Panamanian segregate , Panama M S Ithomia xenos [n. ssp.] Lamas, MS Panama Megoleria Constantino, 1999 Megoleria orestilla (Hewitson, 1867) iN Megoleria orestilla orestilla (Hewitson, 1867) TL: Colombia M T Megoleria orestilla polymacula (Rosenberg & Talbot, 1914) TL: Peru M T S Megoleria orestilla polylla Lamas & Willmott, 2008 ref.334 Megoleria susiana (C. Felder & R. Felder, 1862) iN Megoleria susiana susiana (C. Felder & R. Felder, 1862) TL: Colombia M T S L Megoleria susiana sandra Constantino, 1999 TL: Colombia M T Megoleria susiana susanna (Staudinger, [1884]) TL: Colombia M T S Hyposcada Godman & Salvin, 1879 Hyposcada anchiala (Hewitson, 1868) iN Hyposcada anchiala Panamanian segregate , Panama M S Hyposcada anchiala anchiala (Hewitson, 1868) TL: Peru M T Hyposcada anchiala abidula J. Zikán, 1941 TL: Brazil (Am) M T Hyposcada anchiala cynara R.F. d'Almeida, 1945 TL: Brazil (Am) M T Hyposcada anchiala ecuadorina Bryk, 1953 TL: Ecuador M T S Hyposcada anchiala fallax (Staudinger, [1884]) TL: Peru M S Hyposcada anchiala gallardi Brévignon, 1993 TL: French Guiana M T Hyposcada anchiala interrupta Tessmann, 1928 TL: Peru M T S Hyposcada anchiala kezia (Hewitson, 1868) TL: Peru M T S Hyposcada anchiala mendax R. Fox, 1941 TL: Peru M T Hyposcada anchiala rezia Haensch, 1905 TL: Peru M T S Hyposcada anchiala richardsi R. Fox, 1941 TL: Peru M T S Hyposcada anchiala [n. ssp.#1] Lamas & Willmott, MS TL: Panama Hyposcada anchiala [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyposcada attilodes Kaye, 1918 TL: Bolivia M T Hyposcada [n. sp.] Lamas & Willmott, MS not specified Hyposcada [n. sp.] [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyposcada [n. sp.] [n. ssp.#2] Lamas & Willmott, MS TL: Bolivia Hyposcada dujardini Brévignon, 1993 TL: French Guiana Hyposcada dujardini dujardini Brévignon, 1993 TL: French Guiana M T S Hyposcada dujardini humboldti Neild, 2008 ref.266 Hyposcada illinissa (Hewitson, [1852]) iN Hyposcada illinissa illinissa (Hewitson, [1852]) TL: [Brazil (Am)] M T S Hyposcada illinissa abida (Hewitson, 1871) TL: Colombia M T Hyposcada illinissa aesion (Godman & Salvin, 1878) Panama M T S L Hyposcada illinissa brisotis (Haensch, 1909) TL: Bolivia M T S Hyposcada illinissa cynthia R. Fox, 1941 TL: Ecuador M T Hyposcada illinissa dolabella (Hewitson, 1876) TL: Bolivia M T S Hyposcada illinissa ida Haensch, 1903 TL: Ecuador M T L Hyposcada illinissa idina Haensch, 1905 TL: [Peru] M T Hyposcada illinissa ilerdinoides (Staudinger, [1884]) TL: Brazil (Am) M T Hyposcada illinissa margarita R. Fox, 1941 TL: Peru M T Hyposcada illinissa napirida J. Zikán, 1941 TL: Brazil (Am) M T Hyposcada illinissa napoensis Vitale & Bollino, 2001 TL: Ecuador M T Hyposcada illinissa roberto Neild, 2008 ref.266 Hyposcada illinissa sinilia (Herrich-Schäffer, 1865) TL: Colombia M T L Hyposcada illinissa tundayme S. Padrón, 2008 ref.275 Hyposcada illinissa varia Winhard, 2018 ref.359 Hyposcada illinissa mocoa Winhard, 2018 ref.349,359 Hyposcada illinissa morona Lamas & Willmott, 2021 ref.318 Hyposcada illinissa [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#4] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#5] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#6] Lamas & Willmott, MS TL: Peru Hyposcada illinissa [n. ssp.#7] Lamas & Willmott, MS TL: Brazil (Am) Hyposcada schausi R. Fox, 1941 TL: Colombia Hyposcada schausi schausi R. Fox, 1941 TL: Colombia M T Hyposcada schausi lactea Vitale & Bollino, 2001 TL: Ecuador Hyposcada taliata (Hewitson, 1874) TL: Peru Hyposcada taliata taliata (Hewitson, 1874) TL: Peru M T Hyposcada taliata laetitia Lamas & Willmott, 2021 ref.318 Hyposcada virginiana (Hewitson, [1855]) Virginiana Clearwing iN Hyposcada virginiana virginiana (Hewitson, [1855]) SE Mexico to Honduras M T S L Hyposcada virginiana adelphina (H. Bates, 1866) Panama M T TI L Hyposcada virginiana consobrina (Godman & Salvin, 1892) TL: Ecuador M T Hyposcada virginiana evanides Haensch, 1909 Nicaragua to Panama M T S L Hyposcada virginiana neustetteri Bargmann, 1928 TL: Colombia M T Hyposcada zarepha (Hewitson, 1869) iN Hyposcada zarepha zarepha (Hewitson, 1869) TL: French Guiana M T Hyposcada zarepha bonplandi Neild, 2008 ref.266 Hyposcada zarepha pseudegra (J. Zikán, 1941) TL: Brazil (Pa) M T Hyposcada kena (Hewitson, 1872) ref.318 Hyposcada kena kena (Hewitson, 1872) TL: Ecuador M T Hyposcada kena flexibilis (Haensch, 1909) TL: Peru M T S Hyposcada kena luisa (Tessmann, 1928) TL: Peru M T Hyposcada kena virina (Haensch, 1909) TL: Bolivia M T Hyposcada kena unki Lamas, Willmott & Boyer, 2021 ref.318 Hyposcada kena [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hyposcada kena [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hyposcada kena [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hyposcada kena [n. ssp.#4] Lamas & Willmott, MS TL: Peru Hyposcada kena [n. ssp.#5] Lamas & Willmott, MS TL: Colombia Hyposcada kena [n. ssp.#6] Lamas & Willmott, MS TL: Colombia Oleria Hübner, 1816 Oleria [n. sp.#1] Willmott & Lamas, MS TL: Guyana Oleria aegle (Fabricius, 1776) iN Oleria aegle aegle (Fabricius, 1776) TL: India Oleria aegle divisa (Aurivillius, 1929) TL: Brazil (Am) M T S Oleria aegle egra (Hewitson, [1852]) TL: [Brazil (Pa)] M T Oleria aegle [n. ssp.] Lamas & Willmott, MS Brazil (Rd) Oleria agarista (C. Felder & R. Felder, 1862) iN Oleria agarista agarista (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Oleria agarista idalie R. Fox, 1941 TL: Peru Oleria agarista [n. ssp.] Willmott & Lamas, MS Colombia Oleria alexina (Hewitson, [1859]) iN M T S Oleria amalda (Hewitson, [1857]) iN Oleria amalda amalda (Hewitson, [1857]) TL: Colombia M T S Oleria amalda amaldina (Haensch, 1909) TL: Colombia M T L Oleria amalda faunula (Bargmann, 1929) TL: Colombia M T Oleria amalda modesta (Haensch, 1903) TL: Ecuador M T S Oleria amalda [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria amalda [n. ssp.#2] Lamas & Willmott, MS TL: Colombia Oleria amalda [n. ssp.#3] Neild, MS TL: Venezuela Oleria antaxis (Haensch, 1909) iN Oleria antaxis antaxis (Haensch, 1909) TL: [Brazil (Pa)] M T S Oleria antaxis machadoi R.F. d'Almeida, 1962 TL: Brazil (Ap) M T Oleria antaxis saulensis Brévignon, 1993 TL: French Guiana M T Oleria aquata (Weymer, 1875) iN M T S Oleria assimilis (Haensch, 1903) iN Oleria assimilis assimilis (Haensch, 1903) TL: Ecuador M T S L Oleria assimilis [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria assimilis [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria assimilis [n. ssp.#3] Lamas & Willmott, MS TL: Peru Oleria assimilis [n. ssp.#4] Lamas & Willmott, MS TL: Peru Oleria astrea (Cramer, 1775) iN Oleria astrea astrea (Cramer, 1775) TL: Brazil M S Oleria astrea burchelli (Sanders, 1904) TL: Brazil (Pa) M T S L Oleria astrea chloris Lamas & Brévignon, 1993 TL: French Guiana M T Oleria astrea [n. ssp.#1] Willmott & Lamas, MS TL: Brazil (RJ) Oleria astrea [n. ssp.#2] Lamas & Willmott, MS TL: Brazil (Rd) Oleria thiemei (Oberthür, 1879) Thieme's Clearwing Panama to Colombia iN M T S Oleria athalina (Staudinger, [1884]) iN Oleria athalina athalina (Staudinger, [1884]) TL: Ecuador or Bolivia M T Oleria athalina banjana (Haensch, 1903) TL: Ecuador M T S Oleria athalina epimakrena (Haensch, 1905) TL: Bolivia M T S Oleria athalina [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria athalina [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria attalia (Hewitson, 1855) iN Oleria attalia attalia (Hewitson, 1855) TL: Bolivia M T S L Oleria attalia tabera (Hewitson, 1869) TL: Ecuador M T Oleria attalia [n. ssp.] Lamas & Willmott, MS Peru Oleria baizana (Haensch, 1903) iN Oleria baizana baizana (Haensch, 1903) TL: Ecuador M T Oleria baizana munaycha T. Racheli & L. Racheli, 2003 ref.286 Oleria bifurcata Willmott & Lamas, 2020 ref.317 Oleria bioculata (Haensch, 1905) iN Oleria bioculata bioculata (Haensch, 1905) TL: Bolivia M T Oleria bioculata tapio Bryk, 1953 TL: Peru M T Oleria boyeri Neild, 2008 ref.266 Oleria chimaera Willmott & Lamas, 2020 ref.317 Oleria cyrene (Latreille, [1809]) iN Oleria cyrene cyrene (Latreille, [1809]) TL: Peru M T Oleria cyrene solida (Weymer, 1883) TL: Ecuador M T Oleria cyrene susianoides Winhard, 2019 ref.351,360 Oleria deronda (Hewitson, 1876) iN Oleria deronda deronda (Hewitson, 1876) TL: Bolivia M T Oleria deronda valida (Haensch, 1905) TL: Peru M T S Oleria deronda [n. ssp.] Lamas & Willmott, MS Peru Oleria derondina (Haensch, 1909) iN Oleria derondina derondina (Haensch, 1909) TL: Bolivia M T S Oleria derondina mandorina Lamas, 2003 TL: Peru M T Oleria derondina [n. ssp.] Lamas & Willmott, MS Peru Oleria didymaea (Hewitson, 1876) iN Oleria didymaea didymaea (Hewitson, 1876) TL: Bolivia M T S L Oleria didymaea denuda (N. Riley, 1919) TL: Brazil (Ac) M T Oleria didymaea ramona (Haensch, 1909) TL: Peru M T Oleria didymaea [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria didymaea [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria didymaea [n. ssp.#3] Lamas & Willmott, MS TL: Peru Oleria didymaea [n. ssp.#4] Lamas & Willmott, MS TL: Brazil (Rd) Oleria enania (Haensch, 1909) TL: Peru Oleria enania enania (Haensch, 1909) TL: Peru M T Oleria enania itacoaiensis R.F. d'Almeida, 1964 TL: Brazil (Am) M T Oleria enania [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria enania [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria estella (Hewitson, 1868) iN Oleria estella estella (Hewitson, 1868) TL: Ecuador M T S Oleria estella subosa (Haensch, 1909) TL: Bolivia M T S Oleria fasciata (Haensch, 1903) iN Oleria fasciata fasciata (Haensch, 1903) TL: Ecuador M T Oleria fasciata melchori Lamas, 2003 TL: Peru M T Oleria fasciata [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria fasciata [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria flora (Cramer, 1779) TL: Surinam Oleria flora flora (Cramer, 1779) TL: Surinam M L Oleria flora [n. ssp.] Willmott & Lamas, MS Brazil (Pa) Oleria fumata (Haensch, 1905) iN Oleria fumata fumata (Haensch, 1905) TL: Colombia M T L Oleria fumata richardina Lamas, 1994 TL: Colombia M T Oleria gunilla (Hewitson, 1858) iN Oleria gunilla gunilla (Hewitson, 1858) TL: [Brazil (Am)] M T Oleria gunilla borilis (Haensch, 1909) TL: Peru M T Oleria gunilla escura (Haensch, 1903) TL: Ecuador M T Oleria gunilla lerdina (Staudinger, 1885) TL: Peru M T Oleria gunilla lota (Hewitson, 1872) TL: Ecuador M T L Oleria gunilla lourdes Freitas & Willmott, 2016 TL: Brazil (MT) Oleria gunilla lubilerda (Haensch, 1905) TL: Colombia M T S Oleria gunilla praemona (Haensch, 1909) TL: [Peru] M T Oleria gunilla serdolis (Haensch, 1909) TL: [Peru] M T S Oleria gunilla [n. ssp.#1] Lamas & Willmott, MS TL: Brazil (Pa) Oleria gunilla [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria ilerdina (Hewitson, 1858) iN Oleria ilerdina ilerdina (Hewitson, 1858) TL: Peru M T Oleria ilerdina hermieri Brévignon, 1993 TL: French Guiana M T Oleria ilerdina lerida (W. F. Kirby, 1878) TL: Ecuador M T Oleria ilerdina priscilla (Hewitson, 1858) TL: [Brazil (Am)] M T Oleria makrena (Hewitson, 1854) iN Oleria makrena Panamanian segregate , Panama M S Oleria makrena makrena (Hewitson, 1854) TL: Venezuela M S Oleria makrena caucana (Staudinger, 1885) TL: Colombia M L Oleria makrena [n. ssp.#1] Willmott & Lamas, MS TL: Panama Oleria makrena [n. ssp.#2] Lamas & Willmott, MS TL: Colombia Oleria makrena [n. ssp.#3] Lamas & Willmott, MS TL: Colombia Oleria onega (Hewitson, [1852]) iN Oleria onega onega (Hewitson, [1852]) TL: [Brazil (Am)] M T S Oleria onega bocca (N. Riley, 1919) TL: Brazil (Am) M T Oleria onega crispinilla (Hopffer, 1874) TL: Peru M S Oleria onega epicharme (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Oleria onega ilerda (Hewitson, [1854]) TL: Colombia Oleria onega janarilla (Hewitson, 1863) TL: Ecuador Oleria onega lentita Lamas, 1985 TL: Peru M T S L Oleria onega perspicua (A. Butler, 1877) TL: Brazil (Am) M T Oleria onega astigara Willmott & Lamas, 2021 ref.318 Oleria onega [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria onega [n. ssp.#2] Willmott & Lamas, MS TL: Colombia Oleria onega [n. ssp.#3] Lamas & Willmott, MS TL: Peru Oleria padilla (Hewitson, 1863) iN Oleria padilla padilla (Hewitson, 1863) TL: Ecuador M T S Oleria padilla cajamarcensis Baumann, 1985 TL: Peru Oleria padilla gorkyi Lamas, 2003 TL: Peru M T Oleria padilla pseudmakrena Baumann, 1985 TL: Bolivia M S Oleria padilla [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria padilla [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria paula Weymer, 1883 Paula’s Clearwing iN Oleria paula paula (Weymer, 1883) E & SW Mexico to Panama M S L Ha Oleria paula Panamanian segregate , Osa Peninsula, Costa Rica, to W Panama, Pacific slope M S Oleria paula [n. ssp.#1] Lamas & Willmott, MS TL: Panama Oleria paula [n. ssp.#2] Lamas & Willmott, MS TL: Colombia Oleria phenomoe (E. Doubleday, [1847]) iN Oleria phenomoe phenomoe (E. Doubleday, [1847]) TL: Venezuela M T S Oleria phenomoe apexatrata Neild, 2008 ref.266 Oleria quadrata (Haensch, 1903) iN Oleria quadrata quadrata (Haensch, 1903) TL: Ecuador M S Oleria quadrata [n. ssp.] Willmott & Lamas, MS Colombia Oleria quintina (C. Felder & R. Felder, 1865) iN M T S L Oleria radina (Haensch, 1909) iN Oleria radina radina (Haensch, 1909) TL: Colombia Oleria radina bonita Willmott & Lamas, 2021 ref.318 Oleria radina [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria radina [n. ssp.#2] Willmott & Lamas, MS TL: Colombia Oleria rubescens (A. Butler & H. Druce, 1872) Rubescens Clearwing Nicaragua to Panama iN M T S Oleria santineza (Haensch, 1903) iN Oleria santineza santineza (Haensch, 1903) TL: Ecuador Oleria santineza reducida Neild, 2008 ref.266 M S Oleria santineza veneae Neild, 2008 ref.266 Oleria santineza [n. ssp.] Willmott & Lamas, MS Colombia Oleria sexmaculata (Haensch, 1903) iN Oleria sexmaculata sexmaculata (Haensch, 1903) TL: Ecuador Oleria sexmaculata lerda (Haensch, 1909) TL: [Peru] Oleria sexmaculata [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria sexmaculata [n. ssp.#2] Willmott & Lamas, MS TL: Colombia Oleria sexmaculata [n. ssp.#3] Lamas & Willmott, MS TL: Peru Oleria similigena R.F. d'Almeida, 1962 TL: Brazil (Ap) Oleria similigena similigena R.F. d'Almeida, 1962 TL: Brazil (Ap) M S Oleria similigena [n. ssp.#1] Willmott & Lamas, MS TL: Guyana Oleria similigena [n. ssp.#2] Willmott & Lamas, MS TL: Brazil (Pa) Oleria synnova (Hewitson, [1859]) TL: [Brazil (Am)] M T Oleria tigilla (Weymer, 1899) iN Oleria tigilla tigilla (Weymer, 1899) TL: Ecuador M S Oleria tigilla raya Lamas & Willmott, 2021 ref.318 Oleria tigilla [n. ssp.#1] Lamas & Willmott, MS TL: Peru Oleria tigilla [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria tremona (Haensch, 1909) iN Oleria tremona tremona (Haensch, 1909) TL: Colombia Oleria tremona ochoai Lamas, 2003 TL: Peru M T Oleria tremona schoenfelderi Baumann, 1985 TL: Bolivia Oleria tremona benigna Lamas & Willmott, 2021 ref.318 Oleria tremona [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Oleria tremona [n. ssp.#2] Lamas & Willmott, MS TL: Peru Oleria vicina (Salvin, 1869) Vicina Clearwing Costa Rica & W Panama iN M T IL L Oleria victorine (Guérin-Méneville, [1844]) iN Oleria victorine victorine (Guérin-Méneville, [1844]) TL: Bolivia M S Oleria victorine graziella (Oberthür, 1879) TL: Venezuela M T S Oleria victorine manora (Schaus, 1902) TL: Brazil (RJ) M S Oleria victorine sarilis (Haensch, 1909) TL: [Peru] M L Oleria victorine vicenteae Neild, 2008 ref.266 Oleria victorine [n. ssp.] Lamas & Willmott, MS Brazil (Rd) Oleria [n. sp.#2] Lamas & Willmott, MS TL: Peru Oleria zea (Hewitson, [1855]) Zea Clearwing iN Oleria zea zea (Hewitson, [1855]) SE Mexico to El Salvador & Honduras M T S L Oleria zea diazi J. Maza & Lamas, 1978 SW Mexico M T S Oleria zelica (Hewitson, 1856) iN Oleria zelica zelica (Hewitson, 1856) TL: Ecuador M T S Oleria zelica daguana (Bargmann, 1928) TL: Colombia M T Oleria zelica pagasa (H. Druce, 1875) Costa Rica & Panama M T S L Ollantaya K. Brown & A. Freitas, 1994 Ollantaya aegineta (Hewitson, 1869) TL: Ecuador Ollantaya aegineta aegineta (Hewitson, 1869) ref.194 M T Ollantaya aegineta cleobulina (Hewitson, 1876) ref.194 M T Ollantaya aegineta inelegans (Hewitson, 1877) ref.194 M T S Ollantaya aegineta [n. ssp.] Lamas & Willmott, MS Peru Ollantaya canilla (Hewitson, 1874) ref.194 iN Ollantaya canilla canilla (Hewitson, 1874) TL: Peru M T Ollantaya canilla [n. ssp.#1] Lamas & Willmott, MS TL: Peru Ollantaya canilla [n. ssp.#2] Willmott & Lamas, MS TL: Bolivia Ollantaya olerioides (R.F. d'Almeida, 1952) TL: Bolivia Ollantaya olerioides olerioides (R.F. d'Almeida, 1952) ref.194 M T Ollantaya olerioides baeza Willmott & Lamas, 2021 ref.318 Ollantaya olerioides [n. ssp.] Lamas & Willmott, MS Peru Ollantaya [n. sp.] Lamas & Willmott, MS Peru Ceratinia Hübner, 1816 Ceratinia cayana (Salvin, 1869) iN Ceratinia cayana cayana (Salvin, 1869) TL: French Guiana M T Ceratinia cayana giparanaensis R.F. d'Almeida, 1964 TL: Brazil (Rd) M T Ceratinia iolaia (Hewitson, [1856]) iN Ceratinia iolaia iolaia (Hewitson, [1856]) TL: Colombia M T Ceratinia iolaia conveniens Bryk, 1937 TL: Colombia M T Ceratinia iolaia rehni R. Fox, 1941 TL: Colombia M T Ceratinia neso (Hübner, [1806]) iN Ceratinia neso neso (Hübner, [1806]) TL: [Brazil] M T Ceratinia neso espriella (Hewitson, 1868) TL: Ecuador M T Ceratinia neso hamlini (Weeks, 1906) TL: Venezuela M T Ceratinia neso immaculata Neild, 2008 ref.266 Ceratinia neso naranja Neild, 2008 ref.266 Ceratinia neso nisea (Godart, 1819) TL: French Guiana; Surinam M T Ceratinia neso niselina (J. Zikán, 1941) TL: Brazil (Am) M T Ceratinia neso peruensis (Haensch, 1905) TL: Peru M T Ceratinia neso tarapotis (Haensch, 1909) TL: [Peru] M T Ceratinia neso [n. ssp.#1] Lamas, MS TL: Colombia Ceratinia neso [n. ssp.#2] Lamas, MS TL: Venezuela Ceratinia neso [n. ssp.#3] Lamas, MS TL: Venezuela Ceratinia neso [n. ssp.#4] Lamas, MS TL: Peru Ceratinia neso [n. ssp.#5] Lamas, MS TL: Bolivia Ceratinia neso [n. ssp.#6] Lamas, MS TL: Brazil (Rd) Ceratinia neso [n. ssp.#7] Lamas, MS TL: Brazil (Pa) Ceratinia tutia (Hewitson, 1852) iN Ceratinia tutia tutia (Hewitson, 1852) TL: Amazon M T L Ceratinia tutia Northern segregate , SE Mexico, Guatemala, Honduras M S Ceratinia tutia Costa Rican segregate , Costa Rica M S L Ceratinia tutia azara (Hewitson, [1854]) TL: Ecuador M S Ceratinia tutia dorilla (H. Bates, 1864) Panama M T S L1 L2 Ceratinia tutia singularis (Rebel, 1902) TL: Ecuador M T Ceratinia tutia tosca (Schaus, 1902) TL: Colombia M T Ceratinia tutia transversa (M. Hering, 1925) TL: Colombia M S Ceratinia tutia [n. ssp.#1] Lamas, MS TL: Mexico Ceratinia tutia [n. ssp.#2] Lamas, MS TL: Costa Rica Ceratinia tutia [n. ssp.#3] Lamas, MS TL: Panama Ceratinia tutia [n. ssp.#4] Lamas, MS TL: Colombia Ceratinia tutia [n. ssp.#5] Lamas, MS TL: Venezuela Ceratinia poecila (H. Bates, 1862) TL: Colombia Ceratinia poecila poecila (H. Bates, 1862) TL: Colombia M T Ceratinia poecila azarina (Weymer, 1899) TL: Ecuador M T Ceratinia poecila callichroma (Staudinger, 1885) TL: Ecuador M T Ceratinia poecila chanchamaya (Haensch, 1905) TL: Peru M T Ceratinia poecila fuscens (Haensch, 1905) TL: Bolivia M T Ceratinia poecila hopfferi (Weymer, 1899) TL: [Peru] M T Ceratinia poecila poeciloides (N. Riley, 1919) TL: Brazil (Am) M T Ceratinia poecila porrecta (Haensch, 1905) TL: Bolivia M T Ceratinia poecila radiosa (Haensch, 1903) TL: Ecuador M T Ceratinia poecila robusta (Haensch, 1905) TL: Bolivia M T Ceratinia poecila selenides (Weymer, 1899) TL: Peru M T Ceratinia poecila [n. ssp.#1] Lamas, MS TL: Peru Ceratinia poecila [n. ssp.#2] Lamas, MS TL: Peru Ceratinia poecila [n. ssp.#3] Lamas, MS TL: Peru Ceratinia poecila [n. ssp.#4] Lamas, MS TL: Peru Callithomia H. Bates, 1862 Callithomia alexirrhoe H. Bates, 1862 iN Callithomia alexirrhoe alexirrhoe H. Bates, 1862 TL: Brazil (Am) M T Callithomia alexirrhoe butes Godman & Salvin, 1898 TL: [Peru] M T S Callithomia alexirrhoe schulzi Haensch, 1905 TL: Brazil (Pa) M T Callithomia alexirrhoe thornax H. Bates, 1862 TL: Brazil (Am) M T Callithomia alexirrhoe zeuxippe H. Bates, 1862 TL: Brazil (Pa) M T Callithomia alexirrhoe [n. ssp.#1] Lamas, MS TL: Colombia Callithomia alexirrhoe [n. ssp.#2] Lamas, MS TL: Brazil (MT) Callithomia hezia (Hewitson, [1854]) Hezia Clearwing iN Callithomia hezia hezia (Hewitson, [1854]) SE Mexico to Honduras M T S L Callithomia hezia beronilla (Hewitson, 1874) TL: Colombia M T Callithomia hezia formosa Winhard, 2019 ref.360 Callithomia hezia hedila Godman & Salvin, 1879 SE Mexico to Guatemala M T S Callithomia hezia nigra Winhard, 2019 ref.360 Callithomia hezia perija Masters, 1973 TL: Venezuela M T Callithomia hezia phagesia (Hewitson, 1869) TL: Ecuador M T Callithomia hezia pulchra Winhard, 2019 ref.360 Callithomia hezia renati Neild, 2008 ref.266 Callithomia hezia smalli R. Fox, 1968 Panama M T Callithomia hezia tridactyla Dewitz, 1877 TL: Colombia M T Callithomia hezia villula (Hewitson, 1858) TL: Colombia M T Callithomia hezia wellingi R. Fox, 1968 SW Mexico (S Oaxaca; Sierra Madre del Sur) M T S Callithomia hezia [n. ssp.#1] Lamas, MS TL: Colombia Callithomia hezia [n. ssp.#2] Lamas, MS TL: Colombia Callithomia hydra C. Felder & R. Felder, 1865 TL: Venezuela Callithomia hydra hydra C. Felder & R. Felder, 1865 TL: Venezuela M T Callithomia hydra megaleas Godman & Salvin, 1898 [Panama], W Costa Rica M T S Callithomia hydra panamensis Godman & Salvin, 1878 Panama M T S1 S2 Callithomia lenea (Cramer, 1779) iN Callithomia lenea lenea (Cramer, 1779) TL: Surinam M T Callithomia lenea agrippina (Hewitson, 1863) Panama to Colombia M T S Callithomia lenea alpho (C. Felder & R. Felder, 1865) TL: Venezuela M T Callithomia lenea epidero (H. Bates, 1862) TL: Brazil ([Pa]) M T Callithomia lenea fumantis (Haensch, 1909) TL: Colombia M T Callithomia lenea inturna (R. Fox, 1941) TL: Peru M T Callithomia lenea methonella (Weymer, 1875) TL: Brazil (SC) M T L Callithomia lenea nikita (Haensch, 1909) TL: Venezuela M T Callithomia lenea obfuscata (A. Butler, 1873) TL: Brazil (Am) M T Callithomia lenea pulcheria (Hewitson, 1870) TL: Ecuador M T Callithomia lenea skinneri Masters, 1973 TL: Venezuela Callithomia lenea siparia (Kaye, 1922) TL: Trinidad M T Callithomia lenea travassosi R.F. d'Almeida, 1958 TL: Brazil (MT) M T Callithomia lenea xantho (C. Felder & R. Felder, 1860) TL: Brazil (Ba) M T Callithomia lenea zelie (Guérin-Méneville, [1844]) TL: Bolivia M T S Callithomia lenea bella Neild, 2008 ref.266 Callithomia lenea [n. ssp.#1] Lamas, MS TL: Venezuela Callithomia lenea [n. ssp.#2] Lamas, MS TL: Venezuela Dircenna E. Doubleday, 1847 Dircenna adina (Hewitson, [1855]) iN Dircenna adina adina (Hewitson, [1855]) TL: Venezuela M T S Dircenna adina columbiana E. Krüger, 1925 TL: Colombia M T Dircenna adina lorica Weymer, 1875 TL: Guyana M T S Dircenna adina marica (C. Felder & R. Felder, 1865) TL: Venezuela M T S Dircenna adina steinheili Staudinger, [1884] TL: Colombia M T S Dircenna adina stevei Neild, 2008 ref.266 Dircenna adina suna Haensch, 1903 TL: Ecuador M T S Dircenna adina willmotti Neild, 2008 ref.266 Dircenna adina xanthophane Hopffer, 1874 TL: Peru M T S Dircenna adina [n. ssp.#1] Lamas, MS TL: Brazil (Rr) Dircenna adina [n. ssp.#2] Lamas, MS TL: Colombia Dircenna adina [n. ssp.#3] Lamas, MS TL: Colombia Dircenna adina [n. ssp.#4] Lamas, MS TL: Peru Dircenna dero (Hübner, 1823) iN Dircenna dero Northern segregate , SE Mexico & Guatemala M S Dircenna dero dero (Hübner, 1823) TL: Surinam M S Dircenna dero caliginosa J. Zikán, 1941 TL: Brazil (Am) M T L Dircenna dero celtina Burmeister, 1878 TL: ? M T Dircenna dero christopheri Neild, 2008 ref.266 Dircenna dero euchytma (C. Felder & R. Felder, 1865) Nicaragua to Colombia & Venezuela M T S L Dircenna dero euteles Erschoff, 1875 TL: French Guiana M S Dircenna dero rhoeo C. Felder & R. Felder, 1860 TL: Brazil ([RJ]) M T S Dircenna dero [n. ssp.#1] Lamas, MS TL: Mexico Dircenna dero [n. ssp.#2] Lamas, MS TL: Peru Dircenna dero [n. ssp.#3] Lamas, MS TL: Brazil (Pa) Dircenna jemina (Geyer, 1837) iN Dircenna jemina Northern segregate , E Mexico to Panama M S Dircenna jemina jemina (Geyer, 1837) TL: Venezuela M T S Dircenna jemina chiriquensis Haensch, 1909 Costa Rica & Panama (Pacific slope) M T S Dircenna jemina reynoldsi Neild, 2008 ref.266 Dircenna jemina varina (Hewitson, 1869) TL: Ecuador M T S Dircenna jemina visina Haensch, 1903 TL: Ecuador M T S Dircenna jemina [n. ssp.#1] Lamas, MS TL: Guatemala Dircenna jemina [n. ssp.#2] Lamas, MS TL: Panama Dircenna jemina [n. ssp.#3] Lamas, MS TL: Colombia Dircenna jemina [n. ssp.#4] Lamas, MS TL: Colombia Dircenna jemina [n. ssp.#5] Lamas, MS TL: Colombia Dircenna jemina [n. ssp.#6] Lamas, MS TL: Colombia Dircenna jemina [n. ssp.#7] Lamas, MS TL: Ecuador Dircenna klugii (Geyer, 1837) Klug’s Clearwing iN Dircenna klugii klugii (Geyer, 1837) S TX (very rare stray), E & W Mexico to Nicaragua M S L Dircenna klugii Southern segregate , Costa Rica & Panama M S L Dircenna klugii [n. ssp.] Lamas, MS Costa Rica Dircenna loreta Haensch, 1903 iN Dircenna loreta loreta Haensch, 1903 TL: Ecuador M T Dircenna loreta acreana R.F. d'Almeida, 1950 TL: Brazil (Ac) M T Dircenna loreta melini Bryk, 1953 TL: Peru M T Dircenna loreta [n. ssp.] Lamas, MS Brazil (Pa) Dircenna olyras (C. Felder & R. Felder, 1865) iN Dircenna olyras Panamanian segregate , Panama M S L Dircenna olyras olyras (C. Felder & R. Felder, 1865) TL: Colombia M T Dircenna olyras bairdii Reakirt, 1868 TL: Colombia M T Dircenna olyras relata A. Butler & H. Druce, 1872 Nicaragua to Panama M T S L Dircenna olyras [n. ssp.#1] Lamas, MS TL: Panama Dircenna olyras [n. ssp.#2] Lamas, MS TL: Ecuador Hyalenna W. Forbes, 1942 Hyalenna paradoxa (Staudinger, [1884]) iN Hyalenna paradoxa paradoxa (Staudinger, [1884]) TL: Colombia M T S Hyalenna paradoxa praestigiosa (Haensch, 1903) TL: Ecuador M T S Hyalenna paradoxa catenata (Kaye, 1918) TL: Peru M T Hyalenna paradoxa incachaca Lamas & Willmott, 2006 ref.332 Hyalenna hugia (Schaus, 1902) TL: Bolivia M T S Hyalenna perasippa (Hewitson, 1877) TL: Ecuador Hyalenna perasippa valencia Willmott & Lamas, 2006 ref.332 M S Hyalenna perasippa ortygiosa Willmott & Lamas, 2006 ref.332 M S Hyalenna perasippa perasippa (Hewitson, 1877) TL: Ecuador M T Hyalenna perasippa solitaria Lamas & Willmott, 2006 ref.332 M T Hyalenna buckleyi Willmott & Lamas, 2006 TL: Ecuador Hyalenna buckleyi buckleyi Willmott & Lamas, 2006 ref.332 M T Hyalenna buckleyi pomacocha Lamas & Willmott, 2006 ref.332 Hyalenna pascua (Schaus, 1902) iN M T S Hyalenna alidella (Hewitson, 1869) TL: Colombia Hyalenna alidella exsulans Lamas & Willmott, 2006 Panama (Darien); ref.332 M T Hyalenna alidella vesca Lamas & Willmott, 2006 ref.332 M T S Hyalenna alidella alidella (Hewitson, 1869) TL: Colombia M T S Hyalenna alidella cinereola Lamas & Willmott, 2006 ref.332 M S Hyalenna alidella scantilla (Hewitson, 1877) TL: Ecuador M T S Hyalenna alidella dirama (Haensch, 1905) TL: Bolivia M T Hyalenna alidella minna (Schaus, 1902) TL: Bolivia M T S Hyalenna sulmona (Hewitson, 1877) iN Hyalenna sulmona tersa Willmott & Lamas, 2006 ref.332 M T Hyalenna sulmona hyalina Lamas & Willmott, 2006 ref.332 M T S Hyalenna sulmona lobusa (Haensch, 1909) TL: Colombia M T Hyalenna sulmona sulmona (Hewitson, 1877) TL: Ecuador M T S Hyalenna sulmona balsamica Lamas & Willmott, 2006 ref.332 M T Episcada Godman & Salvin, 1879 Episcada apuleia (Hewitson, 1868) iN Episcada apuleia apuleia (Hewitson, 1868) TL: Ecuador M T Episcada apuleia santanella (Haensch, 1903) TL: Ecuador M T Episcada apuleia cora Haensch, 1909 TL: Bolivia M T Episcada apuleia [n. ssp.] Lamas & Willmott, MS Peru Episcada clausina (Hewitson, 1876) iN Episcada clausina clausina (Hewitson, 1876) TL: Bolivia M T S Episcada clausina bomboiza Willmott & Lamas, 2021 ref.318 Episcada doto (Hübner, [1806]) TL: Brazil ("RJ") Episcada doto doto (Hübner, [1806]) TL: Brazil ("RJ") M T Episcada doto paquito Neild, 2008 ref.266 Episcada doto [n. ssp.] Lamas & Willmott, MS Peru Episcada hemixanthe (C. Felder & R. Felder, 1865) iN Episcada hymen Haensch, 1905 TL: Bolivia Episcada hymen hymen Haensch, 1905 TL: Bolivia M T Episcada hymen [n. ssp.#1] Lamas & Willmott, MS TL: Peru Episcada hymen [n. ssp.#2] Lamas & Willmott, MS TL: Peru Episcada hymenaea (Prittwitz, 1865) iN Episcada hymenaea hymenaea (Prittwitz, 1865) TL: Brazil (RJ) M T S Episcada hymenaea caucana (K. Brown, 1970) TL: Colombia Episcada hymenaea centralis (K. Brown & O. Mielke, 1970) TL: Brazil (MT) M T Episcada hymenaea guianensis Neild, 2008 ref.266 Episcada hymenaea mirifica Lamas & Willmott, 2021 ref.318 Episcada [n. sp.#1] Lamas & Willmott, MS TL: not specified Episcada [n. sp.#1] [n. ssp.#1] Lamas & Willmott, MS TL: Peru Episcada [n. sp.#1] [n. ssp.#2] Lamas & Willmott, MS TL: Peru Episcada [n. sp.#1] [n. ssp.#3] Lamas & Willmott, MS TL: Peru Episcada [n. sp.#1] [n. ssp.#4] Lamas & Willmott, MS TL: Bolivia Episcada mira (Hewitson, 1877) iN M T Episcada philoclea (Hewitson, [1855]) iN M T S Episcada pichita Lamas & Willmott, 2020 ref.317 iN Episcada polita Weymer, 1899 iN Episcada polita polita Weymer, 1899 TL: Colombia M T Episcada polita cabenis Haensch, 1905 TL: Colombia M T Episcada polita aprica Neild, 2008 ref.266 Episcada polita raymondi Neild, 2008 ref.266 Episcada polita archaia Neild, 2008 ref.266 Episcada polita pichincha Willmott & Lamas, 2021 ref.318 Episcada polita flava Winhard, 2021 ref.318 Episcada [n. sp.#2] Lamas & Willmott, MS TL: Peru Episcada salvinia (H. Bates, 1864) Salvin's Clearwing iN Episcada salvinia salvinia (H. Bates, 1864) E Mexico, Guatemala M T L Episcada salvinia apia (C. Felder & R. Felder, 1865) TL: Colombia M T L Episcada salvinia opleri Lamas, 1978 Costa Rica & Panama M T S L Episcada salvinia portilla J. Maza & Lamas, 1978 W Mexico (NAY to OAX) M T S Episcada striposis Haensch, 1909 iN M T S Episcada sulphurea Haensch, 1905 TL: Bolivia M T S Episcada arcadia Lamas & Willmott, 2021 ref.318 Episcada arcadia arcadia Lamas & Willmott, 2021 ref.318 Episcada arcadia navarro Lamas & Willmott, 2021 ref.318 Episcada sylpha Haensch, 1905 TL: Venezuela M T S Episcada sylvo (Geyer, 1832) ref.319 iN M T S Episcada ticidella (Hewitson, 1869) iN Episcada ticidella ticidella (Hewitson, 1869) TL: Ecuador M T Episcada ticidella [n. ssp.#1] Lamas & Willmott, MS TL: Peru Episcada ticidella [n. ssp.#2] Lamas & Willmott, MS TL: Peru Episcada trapezula Brabant & Bischler, 2003 TL: Peru M S Episcada vitrea R.F. d'Almeida & O. Mielke, 1967 TL: Brazil (RJ) M T Episcada zajciwi R.F. d'Almeida & O. Mielke, 1967 iN Episcada zajciwi zajciwi R.F. d'Almeida & O. Mielke, 1967 TL: Brazil (Ce) M T Episcada zajciwi canaria (K. Brown & R.F. d'Almeida, 1970) TL: Brazil (ES) M T S Episcada zajciwi [n. ssp.] Lamas & Willmott, MS Brazil (Ce) Episcada [n. sp.#3] Lamas & Willmott, MS TL: Peru Haenschia Lamas, 2004 Haenschia derama (Haensch, 1905) iN M T Haenschia sidonia (Haensch, 1905) TL: Peru M T Haenschia [n. sp.#1] Lamas, MS TL: Peru Haenschia [n. sp.#2] Lamas, MS TL: Peru Pteronymia A. Butler & H. Druce, 1872 Pteronymia alcmena (Godman & Salvin, 1877) Alcmena Clearwing iN Pteronymia alcmena alcmena (Godman & Salvin, 1877) S Mexico (Chiapas), Guatemala, Honduras, Nicaragua M T S L Pteronymia alcmena Panamanian segregate , Panama M S Pteronymia alcmena [n. ssp.] Lamas & Willmott, MS Panama Pteronymia aletta (Hewitson, [1855]) iN Pteronymia aletta agalla Godman & Salvin, 1879 Costa Rica & Panama (Pacific slope) M T S Pteronymia aletta Panamanian segregate , Panama M S Pteronymia aletta aletta (Hewitson, [1855]) TL: Venezuela M T S Pteronymia aletta lilla (Hewitson, 1856) TL: Ecuador M T Pteronymia aletta [n. ssp.#1] Lamas & Willmott, MS TL: Panama Pteronymia aletta [n. ssp.#2] Willmott & Lamas, MS TL: Colombia Pteronymia aletta [n. ssp.#3] Willmott & Lamas, MS TL: Colombia Pteronymia alicia Neild, 2008 ref.266 M S Pteronymia alida (Hewitson, 1855) iN Pteronymia alida alida (Hewitson, 1855) TL: ? M T S Pteronymia alida dispaena (Hewitson, 1876) TL: Bolivia M T S Pteronymia alida zabina (Hewitson, [1857]) TL: Colombia M T Pteronymia alida georgei Neild, 2008 ref.266 Pteronymia alida cosanga Willmott & Lamas, 2021 ref.318 Pteronymia alida francisca Willmott & Lamas, 2021 ref.318 Pteronymia alida luctuosa Willmott & Lamas, 2021 ref.318 Pteronymia alida [n. ssp.#1] Lamas & Willmott, MS TL: Colombia Pteronymia alida [n. ssp.#2] Willmott & Lamas, MS TL: Ecuador Pteronymia alida [n. ssp.#3] Lamas & Willmott, MS TL: Peru Pteronymia alida [n. ssp.#4] Lamas & Willmott, MS TL: Peru Pteronymia alida [n. ssp.#5] Lamas & Willmott, MS TL: Peru Pteronymia alina Haensch, 1909 iN Pteronymia alina alina Haensch, 1909 TL: Bolivia M T Pteronymia alina machay T. Racheli & L. Racheli, 2003 ref.286 Pteronymia alina mielkei Lamas, 2003 TL: Peru M T Pteronymia alina sumaq T. Racheli & L. Racheli, 2003 ref.286 Pteronymia alina [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pteronymia alina [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia alina [n. ssp.#3] Lamas & Willmott, MS TL: Peru Pteronymia alina [n. ssp.#4] Lamas & Willmott, MS TL: Peru Pteronymia alissa (Hewitson, 1869) iN Pteronymia alissa alissa (Hewitson, 1869) TL: Ecuador M T L Pteronymia alissa amandes Kaye, 1921 TL: Trinidad M T S Pteronymia alissa [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Pteronymia alissa [n. ssp.#2] Willmott & Lamas, MS TL: Colombia Pteronymia andreas (Weeks, 1901) TL: Bolivia Pteronymia andreas andreas (Weeks, 1901) TL: Bolivia M T L Pteronymia andreas [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia artena (Hewitson, [1855]) Artena Clearwing iN Pteronymia artena praedicta J. Maza & Lamas, 1982 SW Mexico (GRO, S OAX) M T S Pteronymia artena artena (Hewitson, [1855]) E Mexico, Guatemala, Honduras M T Pteronymia artena Costa Rican segregate , Costa Rica M S Pteronymia artena Panamanian segregate , Panama M S L Pteronymia artena beebei R. Fox & J. Fox, 1947 TL: Venezuela M T S Pteronymia artena afrania (Hopffer, 1874) TL: Bolivia M T L Pteronymia artena [n. ssp.#1] Lamas & Willmott, MS TL: Costa Rica Pteronymia artena [n. ssp.#2] Lamas & Willmott, MS TL: Panama Pteronymia artena [n. ssp.#3] Willmott & Lamas, MS TL: Colombia Pteronymia artena [n. ssp.#4] Willmott & Lamas, MS TL: Colombia Pteronymia asopo (C. Felder & R. Felder, 1865) TL: Venezuela M T S Pteronymia calgiria Schaus, 1902 iN M T S Pteronymia mariannae Lamas, Willmott & J. Hall, 2020 ref.317 Pteronymia mariannae mariannae Lamas, Willmott & J. Hall, 2020 ref.317 Pteronymia mariannae chisinche Willmott & Lamas, 2020 ref.317 Pteronymia carlia Schaus, 1902 iN M T Pteronymia cotytto (Guérin-Méneville, [1844]) Cotytto Clearwing iN Pteronymia cotytto cotytto (Guérin-Méneville, [1844]) E & S Mexico to Nicaragua, [Costa Rica?] M T S L Pteronymia cotytto Panamanian segregate , Panama M S Pteronymia cotytto [n. ssp.] Lamas & Willmott, MS Panama Pteronymia cuneata Willmott, Lamas & J. Hall, 2020 ref.317 Pteronymia dispar Haensch, 1905 TL: Colombia M T Pteronymia donella (C. Felder & R. Felder, 1865) iN Pteronymia donella Panamanian segregate , Costa Rica & Panama M S Pteronymia donella donella (C. Felder & R. Felder, 1865) TL: Colombia M T Pteronymia donella donata Haensch, 1909 Costa Rica & Panama (Pacific slope) M T S Pteronymia donella mariagabrielae Neild & De Sousa, 2008 ref.266 Pteronymia donella [n. ssp.] Lamas & Willmott, MS Panama Pteronymia dorothyae Neild, 2008 TL: Venezuela Pteronymia dorothyae dorothyae Neild, 2008 ref.266 Pteronymia dorothyae marjorieae Neild, 2008 ref.266 Pteronymia euritea (Cramer, 1780) iN M S Pteronymia [n. sp.#1] Lamas & Willmott, MS TL: Panama Pteronymia forsteri Baumann, 1985 iN M T S Pteronymia fulvimargo A. Butler & H. Druce, 1872 Fulvimargo Clearwing Costa Rica & W Panama M T S Pteronymia fumida Schaus, 1913 Fumida Clearwing iN Pteronymia fumida fumida Schaus, 1913 Costa Rica & Panama M T S Pteronymia fumida pacifica E. Krüger, 1925 TL: Colombia Pteronymia fumida ortizi T. Racheli & L. Racheli, 2003 ref.286 Pteronymia gertschi R. Fox, 1945 TL: Ecuador Pteronymia gertschi gertschi R. Fox, 1945 TL: Ecuador M T Pteronymia gertschi [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia glauca Haensch, 1903 iN M T Pteronymia granica (Hewitson, 1877) TL: Ecuador Pteronymia granica granica (Hewitson, 1877) TL: Ecuador M T Pteronymia granica [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia [n. sp.#2] Lamas & Willmott, MS TL: Mexico (Gue) Pteronymia hara (Hewitson, 1877) iN Pteronymia hara Costa Rican segregate , Costa Rica M S Pteronymia hara Panamanian segregate , Panama M S Pteronymia hara hara (Hewitson, 1877) TL: Ecuador M T Pteronymia hara brunneata Bryk, 1937 TL: Ecuador M T Pteronymia hara semonis Haensch, 1909 TL: Ecuador M T Pteronymia hara ancha Neild, 2008 ref.266 Pteronymia hara albida Neild, 2008 ref.266 Pteronymia hara estrecha Neild, 2008 ref.266 Pteronymia hara olerina Willmott & Lamas, 2021 ref.318 Pteronymia hara [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia inania Haensch, 1903 TL: Ecuador Pteronymia inania inania Haensch, 1903 TL: Ecuador M T Pteronymia inania evonia Haensch, 1905 TL: Colombia M T S Pteronymia inania bethana Willmott & Lamas, 2021 ref.318 Pteronymia inania [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia latilla (Hewitson, [1855]) iN Pteronymia latilla fulvescens Godman & Salvin, 1879 Costa Rica & W Panama M T S L Pteronymia latilla Panamanian segregate , Panama M S L Pteronymia latilla latilla (Hewitson, [1855]) TL: Venezuela M T S Pteronymia latilla barilla Haensch, 1903 TL: Ecuador M T Pteronymia latilla nigricans Bryk, 1937 TL: Colombia Pteronymia latilla [n. ssp.] Lamas & Willmott, MS Colombia Pteronymia laura (Staudinger, 1885) iN M T Pteronymia lonera (A. Butler & H. Druce, 1872) Lonera Clearwing Costa Rica & W Panama M T S Pteronymia medellina Haensch, 1905 iN Pteronymia medellina medellina Haensch, 1905 TL: Colombia M T S Pteronymia medellina [n. ssp.] Willmott & Lamas, MS Colombia Pteronymia obscuratus (Fabricius, 1793) Obscure Clearwing Panama M T S Pteronymia olimba Haensch, 1905 TL: Peru M T Pteronymia oneida (Hewitson, 1855) iN Pteronymia oneida oneida (Hewitson, 1855) TL: Colombia M T Pteronymia oneida lisae Salazar, 1992 TL: Colombia Pteronymia oneida sebastiani Neild, 2008 ref.266 Pteronymia oneida merida Neild, 2008 ref.266 M S Pteronymia oneida ricardi Neild, 2008 ref.266 Pteronymia oneida [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pteronymia oneida [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia ozia (Hewitson, 1870) iN Pteronymia ozia ozia (Hewitson, 1870) TL: Ecuador M T Pteronymia ozia browni R. Fox, 1945 TL: Ecuador M T Pteronymia ozia tanampaya Haensch, 1909 TL: Bolivia M T S Pteronymia parva (Salvin, 1869) Parva Clearwing [S Mexico?], Costa Rica & Panama iN M T S Pteronymia peteri Neild, 2008 ref.266 Pteronymia picta (Salvin, 1869) iN Pteronymia picta notilla A. Butler & H. Druce, 1872 Costa Rica to Panama M T S Pteronymia picta Panamanian segregate , Panama M S Pteronymia picta picta (Salvin, 1869) TL: Colombia M T Pteronymia picta [n. ssp.#1] Lamas & Willmott, MS TL: Panama Pteronymia picta [n. ssp.#2] Lamas & Willmott, MS TL: Colombia Pteronymia primula (H. Bates, 1862) iN Pteronymia primula primula (H. Bates, 1862) TL: Brazil (Am) M T Pteronymia primula auricula Haensch, 1905 TL: Colombia M T S Pteronymia primula crocea Neild, 2008 ref.266 Pteronymia primula [n. ssp.] Lamas & Willmott, MS Bolivia Pteronymia rufocincta (Salvin, 1869) West-Mexican Clearwing W Mexico (NAY to OAX) iN M T S L Pteronymia sao (Hübner, [1813]) iN Pteronymia sao sao (Hübner, [1813]) TL: [Brazil] Pteronymia sao antisao (H. Bates, 1862) TL: [Brazil (Am)] M T S Pteronymia sao guntheri Lamas, 1985 TL: Peru M T S Pteronymia sao loreto Lamas & Willmott, 2021 ref.318 Pteronymia sao [n. ssp.#1] Lamas & Willmott, MS TL: Brazil (Pa) Pteronymia sao [n. ssp.#2] Lamas & Willmott, MS TL: Bolivia Pteronymia serrata Haensch, 1903 iN Pteronymia serrata serrata Haensch, 1903 TL: Ecuador M T Pteronymia serrata amplificata Kaye, 1918 TL: Ecuador M T Pteronymia serrata triangula Winhard, 2019 ref.360 Pteronymia serrata [n. ssp.] Lamas & Willmott, MS Peru Pteronymia sexpunctata (Bryk, 1953) TL: Peru Pteronymia sexpunctata sexpunctata (Bryk, 1953) TL: Peru M T Pteronymia sexpunctata [n. ssp.#1] Willmott & Lamas, MS TL: Colombia Pteronymia sexpunctata [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia simplex (Salvin, 1869) Simple Clearwing iN Pteronymia simplex simplex (Salvin, 1869) Costa Rica & Panama M T S L Pteronymia simplex fenochioi Lamas, 1978 E Mexico to Nicaragua M T S L Pteronymia simplex timagenes Godman & Salvin, 1889 SW Mexico (GRO, S OAX) M T S Pteronymia tamina Haensch, 1909 iN Pteronymia tamina tamina Haensch, 1909 TL: Peru M T Pteronymia tamina [n. ssp.] Willmott & Lamas, MS Ecuador Pteronymia teresita (Hewitson, 1863) iN M T Pteronymia thabena (Hewitson, 1869) iN Pteronymia thabena thabena (Hewitson, 1869) TL: Ecuador M T Pteronymia thabena denticulata Haensch, 1905 TL: Bolivia M T S Pteronymia thabena [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pteronymia thabena [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia ticida (Hewitson, 1869) iN Pteronymia ticida ticida (Hewitson, 1869) TL: Ecuador M T Pteronymia ticida citrina Neild, 2008 ref.266 Pteronymia ticida yungava Haensch, 1905 TL: Bolivia M T S Pteronymia ticida melanoptera (Hewitson, 1877) TL: Ecuador M T S Pteronymia ticida [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pteronymia ticida [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia ticida [n. ssp.#3] Lamas & Willmott, MS TL: Peru Pteronymia tucuna (H. Bates, 1862) iN M T S Pteronymia veia (Hewitson, [1853]) iN Pteronymia veia veia (Hewitson, [1853]) TL: ? M T S Pteronymia veia brunnea Bryk, 1937 TL: ? M T S Pteronymia veia florea Neild, 2008 ref.266 M L Pteronymia veia linzera (Herrich-Schäffer, 1865) TL: Colombia M T S Pteronymia veia pomacocha T. Racheli & L. Racheli, 2003 TL: Peru Pteronymia veia [n. ssp.] Lamas & Willmott, MS Colombia Pteronymia luisa Willmott, Lamas & J. Hall, 2020 Ecuador; ref.317 Pteronymia vestilla (Hewitson, [1853]) iN Pteronymia vestilla vestilla (Hewitson, [1853]) TL: Brazil (Am) M T Pteronymia vestilla sparsa Haensch, 1903 TL: Ecuador M T Pteronymia vestilla ucaya Haensch, 1909 TL: Peru M T Pteronymia vestilla [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pteronymia vestilla [n. ssp.#2] Lamas & Willmott, MS TL: Peru Pteronymia zerlina (Hewitson, [1856]) iN Pteronymia zerlina zerlina (Hewitson, [1856]) TL: Colombia M T Pteronymia zerlina amarillana Neild, 2008 ref.266 Pteronymia zerlina cristalina Neild, 2008 ref.266 Pteronymia zerlina nubivaga R. Fox & J. Fox, 1947 TL: Venezuela M T S Pteronymia zerlina pronuba (Hewitson, 1870) TL: Ecuador M T Pteronymia zerlina [n. ssp.] Lamas & Willmott, MS Colombia Velamysta Haensch, 1909 Velamysta peninna (Hewitson, 1855) iN Velamysta peninna peninna (Hewitson, 1855) TL: Bolivia M T Velamysta peninna [n. ssp.] Lamas, MS Peru Velamysta phengites R. Fox, 1945 TL: Ecuador Velamysta phengites phengites R. Fox, 1945 TL: Ecuador M T Velamysta phengites perlucens Winhard, 2017 ref.317 Velamysta phengites [n. ssp.#1] Lamas, MS TL: Colombia Velamysta phengites [n. ssp.#2] Lamas, MS TL: Peru Velamysta phengites [n. ssp.#3] Lamas, MS TL: Peru Velamysta pupilla (Hewitson, 1874) iN Velamysta pupilla pupilla (Hewitson, 1874) TL: Bolivia M T Velamysta pupilla anomala (Staudinger, [1884]) TL: Colombia M T Velamysta cruxifera (Hewitson, 1877) TL: Ecuador Velamysta cruxifera cruxifera (Hewitson, 1877) ref.335 M T S Velamysta cruxifera greeneyi Vitale & Bollino, 2003 ref.335 Velamysta desmondi Kell & Willmott, 2020 ref.317 iN Godyris Boisduval, 1870 Godyris crinippa (Hewitson, 1874) iN Godyris crinippa crinippa (Hewitson, 1874) TL: Bolivia M T Godyris crinippa [n. ssp.] Lamas, MS Peru Godyris dircenna (C. Felder & R. Felder, 1865) iN M T Godyris duillia (Hewitson, 1854) iN M T S Godyris hewitsoni (Haensch, 1903) TL: ["Bolivia"] Godyris hewitsoni hewitsoni (Haensch, 1903) TL: ["Bolivia"] M T Godyris hewitsoni [n. ssp.#1] Lamas, MS TL: Colombia Godyris hewitsoni [n. ssp.#2] Lamas, MS TL: Peru Godyris hewitsoni [n. ssp.#3] Lamas, MS TL: Peru Godyris hewitsoni [n. ssp.#4] Lamas, MS TL: Peru Godyris kedema (Hewitson, [1855]) iN Godyris kedema furina (Godman & Salvin, [1879]) Panama M T S L Godyris kedema Panamanian segregate , Panama M S Godyris kedema kedema (Hewitson, [1855]) TL: Venezuela M T S Godyris kedema albinotata (A. Butler, 1873) TL: Colombia M T Godyris kedema [n. ssp.] Lamas, MS Panama Godyris lauta (Haensch, 1910) TL: [Ecuador] Godyris lauta lauta (Haensch, 1910) TL: [Ecuador] M T Godyris lauta [n. ssp.] Lamas, MS Peru Godyris nepos (Weymer, 1875) iN M T Godyris nero (Hewitson, [1855]) Nero Clearwing iN Godyris nero nero (Hewitson, [1855]) E Mexico to Nicaragua M T S L Godyris nero Southern segregate , Costa Rica M S Godyris nero [n. ssp.] Lamas, MS Costa Rica Godyris panthyale (C. Felder & R. Felder, 1862) iN Godyris panthyale panthyale (C. Felder & R. Felder, 1862) TL: Colombia M T S Godyris panthyale quinta (Staudinger, 1885) TL: Colombia M T Godyris panthyale [n. ssp.] Lamas, MS Peru Godyris sappho (Haensch, 1910) TL: Bolivia Godyris sappho sappho (Haensch, 1910) TL: Bolivia M T Godyris sappho tika (T. Racheli & L. Racheli, 2003) TL: Peru Godyris sappho [n. ssp.] Lamas, MS Peru Godyris zavaleta (Hewitson, [1855]) iN Godyris zavaleta sosunga (Reakirt, [1866]) SE Mexico to Nicaragua M S L Godyris zavaleta caesiopicta (Niepelt, 1915) [= sorites], Costa Rica & Panama (Atlantic slope) M T S1 S2 L Godyris zavaleta zygia (Godman & Salvin, 1877) Costa Rica & Panama (Pacific slope) M T OI S Godyris zavaleta Panamanian s segregate , Panama M S L Godyris zavaleta zavaleta (Hewitson, [1855]) TL: [Colombia] M T Godyris zavaleta christiani Neild, 2008 ref.266 Godyris zavaleta eutelina Brévignon, 1993 TL: French Guiana M S Godyris zavaleta gonussa (Hewitson, [1856]) TL: Colombia M T Godyris zavaleta huallaga R. Fox, 1941 TL: Peru M T Godyris zavaleta matronalis (Weymer, 1883) TL: Ecuador M T S Godyris zavaleta petersii (Dewitz, 1877) TL: Colombia M T Godyris zavaleta rosata Vitale & G. Rodriguez, 2004 ref.298 Godyris zavaleta telesilla (Hewitson, 1863) TL: Ecuador M T Godyris zavaleta [n. ssp.#1] Lamas, MS TL: Panama Godyris zavaleta [n. ssp.#2] Lamas, MS TL: Panama Godyris zavaleta [n. ssp.#3] Lamas, MS TL: Peru Godyris zavaleta [n. ssp.#4] Lamas, MS TL: Peru Godyris zavaleta [n. ssp.#5] Lamas, MS TL: Peru Godyris zavaleta [n. ssp.#6] Lamas, MS TL: Peru Godyris zavaleta [n. ssp.#7] Lamas, MS TL: Peru Godyris zavaleta [n. ssp.#8] Lamas, MS TL: Bolivia Godyris zavaleta [n. ssp.#9] Lamas, MS TL: Brazil (Rd) Pachacutia Willmott & Lamas, 2007 Pachacutia baroni Willmott & Lamas, 2007 ref.333 M S Pachacutia mantura (Hewitson, 1876) TL: Bolivia Pachacutia mantura joroni Lamas & Willmott, 2007 ref.333 M T Pachacutia mantura honrathi (Srnka, 1885) TL: Peru M T Pachacutia mantura mantura (Hewitson, 1876) TL: Bolivia M T Pachacutia cleomella (Hewitson, 1874) TL: Bolivia M T Pachacutia germaini Lamas & Willmott, 2007 ref.333 M T S Veladyris R. Fox, 1945 Veladyris electrea (M. Brabant, 2004) TL: Peru Veladyris cytharista (Hewitson, 1874) iN M T S Veladyris pardalis (Salvin, 1869) iN Veladyris pardalis pardalis (Salvin, 1869) TL: Ecuador M T Veladyris pardalis aurea Lamas, 1980 TL: Ecuador M T S L Veladyris pardalis christina Lamas, 1980 TL: Peru M T L Veladyris pardalis totumbra (Kaye, 1919) TL: Peru M T Veladyris pardalis [n. ssp.] Lamas & Vitale, MS Peru Veladyris [n. sp.] Lamas & Vitale, MS Peru Hypoleria Godman & Salvin, 1879 Hypoleria adasa (Hewitson, [1855]) iN Hypoleria adasa adasa (Hewitson, [1855]) TL: Brazil M T Hypoleria adasa parcilimbata R.F. d'Almeida, 1943 TL: Brazil (SP) M T Hypoleria alema (Hewitson, [1857]) iN Hypoleria alema alema (Hewitson, [1857]) TL: Colombia M T Hypoleria alema cajona Haensch, 1905 TL: Peru M T S L Hypoleria alema chrysodonia (H. Bates, 1862) TL: Brazil (Am) M T S L Hypoleria alema consimilis Talbot, 1928 TL: Brazil (MT) M T Hypoleria alema garleppi Haensch, 1905 TL: Peru M T Hypoleria alema ina (Hewitson, [1859]) TL: Ecuador M T Hypoleria alema inconspicua Neild, 2008 ref.266 Hypoleria alema indecora Haensch, 1905 TL: Guyana M T Hypoleria alema karschi Haensch, 1903 TL: Ecuador M T Hypoleria alema niclae Neild, 2008 ref.266 Hypoleria alema oreas Weymer, 1899 TL: Brazil (ES) M T Hypoleria alema pastaza Willmott & Lamas, 2021 ref.318 Hypoleria alema proxima Weymer, 1899 TL: Brazil (SP) M T Hypoleria alema [n. ssp.#1] Lamas, MS TL: Brazil (Am) Hypoleria alema [n. ssp.#2] Lamas, MS TL: Brazil (Rd) Hypoleria alema [n. ssp.#3] Lamas, MS TL: French Guiana Hypoleria asellia (Hopffer, 1874) TL: Peru M T Hypoleria aureliana (H. Bates, 1862) TL: Brazil (Am) M T Hypoleria lavinia (Hewitson, [1855]) iN Hypoleria lavinia lavinia (Hewitson, [1855]) TL: [Ecuador] M T Hypoleria lavinia cassotis (H. Bates, 1864) S Mexico to Honduras M T S L Hypoleria lavinia libera Godman & Salvin, 1879 Panama M T S L Hypoleria lavinia meridana R. Fox, 1948 TL: Venezuela M T S Hypoleria lavinia rhene (Godman & Salvin, 1878) Nicaragua to W Panama M T S1 S2 L Hypoleria lavinia riffarthi Haensch, 1905 TL: Ecuador M T Hypoleria lavinia vanilia (Herrich-Schäffer, 1865) Panama to Colombia M T S L Hypoleria lavinia vaniliana Kaye, 1919 TL: Colombia M T Hypoleria lavinia aquilonalis Winhard, 2019 ref.360 Hypoleria mulviana R.F. d'Almeida, 1958 TL: Brazil (Pa) Hypoleria ocalea (E. Doubleday, 1847) iN Hypoleria ocalea ocalea (E. Doubleday, 1847) TL: Venezuela; "Brazil" M T S Hypoleria ocalea gephira (Hewitson, [1856]) TL: Colombia M T Hypoleria ocalea [n. ssp.#1] Lamas, MS TL: Colombia Hypoleria ocalea [n. ssp.#2] Lamas, MS TL: Colombia Hypoleria sarepta (Hewitson, [1852]) iN Hypoleria sarepta sarepta (Hewitson, [1852]) TL: [Brazil (Am)] M T S Hypoleria sarepta cidonia (Hewitson, [1857]) TL: Colombia M T S Hypoleria sarepta famina Haensch, 1909 TL: Peru M T Hypoleria sarepta goiana R.F. d'Almeida, 1951 TL: Brazil (Go) M T S Hypoleria sarepta olerioides R.F. d'Almeida, 1951 TL: Brazil (Rd) M T Hypoleria sarepta oriana (Hewitson, [1859]) TL: Brazil (Am) M T Hypoleria sarepta orianula Bryk, 1953 TL: Peru Hypoleria sarepta virginia (Hewitson, [1853]) TL: Brazil (Am) M T Hypoleria sarepta vitiosa Lamas, 1985 TL: Peru M T L Hypoleria sarepta [n. ssp.#1] Lamas, MS TL: Peru Hypoleria sarepta [n. ssp.#2] Lamas, MS TL: Peru Hypoleria xenophis Haensch, 1909 TL: Peru M T Brevioleria Lamas, 2004 Brevioleria aelia (Hewitson, 1852) iN Brevioleria aelia aelia (Hewitson, 1852) TL: Brazil (Am) M T Brevioleria aelia brevicula (R.F. d'Almeida, 1951) TL: Peru M T Brevioleria aelia jamariensis (R.F. d'Almeida, 1951) TL: Brazil (Rd) M T L Brevioleria aelia orolina (Hewitson, [1861]) TL: Brazil (Am) M T Brevioleria aelia pachiteae (Tessmann, 1928) TL: Peru M T L Brevioleria aelia plisthenes (R.F. d'Almeida, 1958) TL: Brazil (MG) M T Brevioleria aelia yawara (J. Zikán, 1942) TL: Brazil (Am) M T Brevioleria aelia [n. ssp.] Lamas, MS Colombia Brevioleria arzalia (Hewitson, 1876) iN Brevioleria arzalia arzalia (Hewitson, 1876) TL: Bolivia M T Brevioleria arzalia loronia Lamas & Willmott, 2001 ref.318 Brevioleria coenina (Hewitson, 1869) TL: Ecuador M T Brevioleria seba (Hewitson, 1872) iN Brevioleria seba seba (Hewitson, 1872) TL: ? M T Brevioleria seba emyra (Haensch, 1905) TL: Brazil (MG) M T Brevioleria seba oculata (Haensch, 1903) TL: Ecuador M T Brevioleria seba [n. ssp.#1] Lamas, MS TL: Peru Brevioleria seba [n. ssp.#2] Lamas, MS TL: Peru Brevioleria seba [n. ssp.#3] Lamas, MS TL: Peru Brevioleria [n. sp.] Lamas, MS Brazil (Pa) Mcclungia R. Fox, 1940 Mcclungia cymo (Hübner, [1806]) iN Mcclungia cymo cymo (Hübner, [1806]) TL: Brazil M T Mcclungia cymo arcuata (Tessmann, 1928) TL: Peru M T S Mcclungia cymo collenettei (Talbot, 1928) TL: Brazil (MT) M T Mcclungia cymo fallens (Haensch, 1905) TL: Brazil (MG) M T Mcclungia cymo salonina (Hewitson, 1855) TL: Bolivia M T L Mcclungia cymo subtilis (Haensch, 1903) TL: Ecuador M T L Mcclungia cymo trepotis (Haensch, 1910) TL: Bolivia M T S Mcclungia cymo wana (A. Hall, 1930) TL: Guyana M T S Mcclungia cymo [n. ssp.] Lamas, MS Brazil (Rd) Mcclungia cymo iguazu Winhard, 2020 ref.361 Greta Hemming, 1934 Greta andromica (Hewitson, [1855]) iN Greta andromica lyra (Salvin, 1869) E Mexico to Panama M T S L Greta andromica Panamanian segregate , Panama M S Greta andromica andromica (Hewitson, [1855]) TL: Venezuela M T S L Greta andromica andania (Hopffer, 1874) TL: Peru M T S L Greta andromica dromica (Haensch, 1910) TL: Colombia M T Greta andromica nerina (Haensch, 1905) TL: Colombia M T Greta andromica trifenestra R. Fox, 1941 TL: Trinidad M T Greta andromica [n. ssp.] Lamas & Willmott, MS Peru Greta annette (Guérin-Méneville, [1844]) White-spotted Clearwing iN Greta annette annette (Guérin-Méneville, [1844]) E Mexico, Guatemala, Honduras M T S L Greta annette championi Lamas, 1978 Nicaragua to Panama M T S L Greta annette moschion (Godman, 1901) W Mexico (NAY to S OAX) M T S L Ha Greta clavijoi Neild, 2008 ref.266 M S Greta cubana (Herrich-Schäffer, 1862) Cuban Clearwing Cuba iN M T S I Greta diaphanus (Drury, 1773) iN M T S1 S2 L I Greta morgane (Geyer, 1837) Rusty Clearwing iN Greta morgane morgane (Geyer, 1837) W Mexico (NAY to S OAX) M S L Greta morgane oto (Hewitson, [1855]) E Mexico to Panama M T S L1 L2 Greta [n. sp.] Lamas & Willmott, MS Peru Hypomenitis R. Fox, 1945 Hypomenitis alphesiboea (Hewitson, 1869) ref.184 iN M T S Hypomenitis candida Lamas & Willmott, 2020 ref.317 Hypomenitis [n. sp.#1] Lamas & Willmott, MS TL: Peru Hypomenitis [n. sp.#2] Lamas & Willmott, MS TL: Peru Hypomenitis depauperata (Boisduval, 1870) iN Hypomenitis depauperata depauperata (Boisduval, 1870) ref.184 M T S Hypomenitis depauperata umbrosa (Haensch, 1903) ref.184 M T Hypomenitis dercetis (E. Doubleday, 1847) iN Hypomenitis dercetis dercetis (E. Doubleday, 1847) ref.184 M T S Hypomenitis dercetis [n. ssp.] Lamas & Willmott, MS Peru Hypomenitis enigma (Haensch, 1905) TL: Bolivia Hypomenitis enigma enigma (Haensch, 1905) ref.184 M T S Hypomenitis enigma koechlini (Lamas, 2003) ref.184 M T Hypomenitis enigma pseudortygia (Neild & Willmott, 2008) ref.266 M T Hypomenitis enigma [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hypomenitis enigma [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hypomenitis enigma [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hypomenitis enigma [n. ssp.#4] Lamas & Willmott, MS TL: Peru Hypomenitis jamesiana Willmott, Lamas & J. Hall, 2020 ref.317 Hypomenitis esula (Hewitson, 1855) TL: Colombia Hypomenitis esula esula (Hewitson, 1855) ref.184 M T Hypomenitis esula [n. ssp.] Lamas & Willmott, MS Colombia Hypomenitis [n. sp.#3] Lamas & Willmott, MS TL: Peru Hypomenitis gabiglooris Brabant & Bischler, 2005 Gabi's Clearwing Nicaragua to W Panama; ref.184 iN M S Hypomenitis gardneri (Weeks, 1901) iN Hypomenitis gardneri gardneri (Weeks, 1901) ref.184 M T S Hypomenitis gardneri devriesi (D'Abrera, 1984) ref.184 M T Hypomenitis gardneri [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hypomenitis gardneri [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hypomenitis gardneri [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hypomenitis gardneri [n. ssp.#4] Lamas & Willmott, MS TL: Peru Hypomenitis hermana (Haensch, 1903) iN Hypomenitis hermana hermana (Haensch, 1903) ref.184 M T Hypomenitis hermana joiceyi (Kaye, 1918) ref.184 M T Hypomenitis hermana [n. ssp.] Willmott & Lamas, MS Colombia Hypomenitis hermana serafina Willmott & Lamas, 2021 ref.318 Hypomenitis hermana chamba Willmott & Lamas, 2021 ref.318 Hypomenitis libethris (C. Felder & R. Felder, 1865) iN Hypomenitis libethris libethris (C. Felder & R. Felder, 1865) ref.184 M T S L Hypomenitis libethris [n. ssp.] Willmott & Lamas, MS Colombia Hypomenitis lojana Vitale & Bollino, 2001 ref.184 M T Hypomenitis lydia (Weymer, 1899) [= flava]; ref.184 iN M T Hypomenitis ochretis (Haensch, 1903) iN Hypomenitis ochretis ochretis (Haensch, 1903) ref.184 M T Hypomenitis ochretis [n. ssp.] Willmott & Lamas, MS Colombia Hypomenitis oneidodes (Kaye, 1918) TL: Ecuador Hypomenitis oneidodes oneidodes (Kaye, 1918) ref.184 M T S Hypomenitis oneidodes guarumales Boyer, Petit & Willmott, 2021 ref.318 Hypomenitis oneidodes nicolasi Willmott & Lamas, 2021 ref.318 Hypomenitis ortygia (Weymer, 1890) iN Hypomenitis ortygia ortygia (Weymer, 1890) [= panthyaloides]; ref.184 M T Hypomenitis ortygia pyrczi Vitale & Bollino, 2001 ref.184 Hypomenitis ortygia [n. ssp.#1] Lamas & Willmott, MS TL: Colombia Hypomenitis ortygia [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hypomenitis ortygia [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hypomenitis [n. sp.#4] Willmott & Lamas, MS TL: Colombia Hypomenitis polissena (Hewitson, 1863) iN Hypomenitis polissena polissena (Hewitson, 1863) ref.184 M T Hypomenitis polissena colombiana Vitale & Bollino, 2001 ref.184 Hypomenitis polissena umbrana (Haensch, 1908) Costa Rica & Panama; ref.184 M T S L Hypomenitis polissena bolivariana Neild, 2008 ref.266 Hypomenitis polissena [n. ssp.] Lamas & Willmott, MS Colombia Hypomenitis theudelinda (Hewitson, [1861]) iN Hypomenitis theudelinda theudelinda (Hewitson, [1861]) ref.184 M T Hypomenitis theudelinda bonita Vitale & Bollino, 2001 ref.184 Hypomenitis theudelinda cyrcilla (Hewitson, 1874) ref.184 M T Hypomenitis theudelinda zalmunna (Hewitson, 1869) ref.184 M T L Hypomenitis theudelinda [n. ssp.#1] Lamas & Willmott, MS TL: Peru Hypomenitis theudelinda [n. ssp.#2] Lamas & Willmott, MS TL: Peru Hypomenitis theudelinda [n. ssp.#3] Lamas & Willmott, MS TL: Peru Hypomenitis [n. sp.#5] Lamas & Willmott, MS Undescribed Clearwing (Costa Rica) Costa Rica Heterosais Godman & Salvin, 1880 Heterosais edessa (Hewitson, [1855]) iN M T S Heterosais giulia (Hewitson, [1855]) iN Heterosais giulia cadra (Godman & Salvin, 1878) Nicaragua to Panama M T S Heterosais giulia Panamanian segregate , Panama M S L Heterosais giulia giulia (Hewitson, [1855]) TL: Colombia M T S Heterosais giulia pallidula Haensch, 1903 TL: Ecuador M T Heterosais giulia caucaensis Constantino, Gil & Valencia, 2005 TL: Colombia M T Heterosais giulia [n. ssp.#1] Lamas, MS TL: Colombia Heterosais giulia [n. ssp.#2] Lamas, MS TL: Brazil (Am) Heterosais nephele (H. Bates, 1862) TL: Brazil (Am) Heterosais nephele nephele (H. Bates, 1862) TL: Brazil (Am) M T S Heterosais nephele gedera (Hewitson, 1869) TL: [Colombia] M T Pseudoscada Godman & Salvin, 1879 Pseudoscada acilla (Hewitson, 1867) iN Pseudoscada acilla acilla (Hewitson, 1867) TL: Brazil (MG/[RJ]) M T Pseudoscada acilla quadrifasciata Talbot, 1928 TL: Brazil (MT) M T Pseudoscada acilla [n. ssp.] Lamas & Willmott, MS Brazil (Rd) Pseudoscada erruca (Hewitson, 1855) iN M T S Pseudoscada florula (Hewitson, [1855]) iN Pseudoscada florula florula (Hewitson, [1855]) TL: French Guiana M T Pseudoscada florula aureola (H. Bates, 1862) TL: Brazil (Am) M T S V Pseudoscada florula exornata Haensch, [1908] TL: Guyana M T Pseudoscada florula genetyllis (R.F. d'Almeida, 1922) TL: Brazil (RJ) M T S Pseudoscada florula gracilis Tessmann, 1928 TL: Peru M T S V Pseudoscada florula [n. ssp.#1] Lamas & Willmott, MS TL: Peru Pseudoscada florula [n. ssp.#2] Lamas & Willmott, MS TL: Brazil (Rd) Pseudoscada timna (Hewitson, [1855]) iN Pseudoscada timna timna (Hewitson, [1855]) TL: Venezuela M T S L Pseudoscada timna utilla (Hewitson, 1856) TL: Colombia M T Pseudoscada pusio (Godman & Salvin, 1877) Timna Clearwing Nicaragua to Panama iN M T S L Pseudoscada troetschi (Staudinger, [1884]) iN Pseudoscada troetschi troetschi (Staudinger, [1884]) TL: Colombia M T Pseudoscada troetschi saturata (Staudinger, 1885) Panama and Colombia M T S L Pseudoscada troetschi colombiana Winhard, 2020 ref.237,361 Pseudoscada troetschi litoralis Winhard, 2020 ref.236,361 Subfamily Heliconiinae Tribe Acraeini Actinote Hübner, [1819] Actinote abana (Hewitson, 1868) ref.12 iN Actinote abana abana (Hewitson, 1868) TL: Ecuador M T Actinote abana capnodes Jordan, 1913 TL: Colombia M T Actinote erinome (C. Felder & R. Felder, 1861) ref.12 iN Actinote erinome erinome (C. Felder & R. Felder, 1861) TL: Ecuador M T S Actinote erinome mathani Oberthür, 1917 TL: Peru M T Actinote erinome testacea (Salvin & Godman, 1868) TL: Bolivia M T S Actinote erinome [n. ssp.] Lamas, MS Ecuador Actinote euryleuca Jordan, 1910 ref.12 M T L Actinote hylonome (E. Doubleday, 1844) ref.12 iN Actinote hylonome hylonome (E. Doubleday, 1844) TL: Colombia M T Actinote hylonome basilia (Lamas, 1995) TL: Venezuela Actinote roseus Winhard, 2017 ref.12,Winhard_2017 Actinote intensa Jordan, 1910 ref.12 M T S Actinote johncoulsoni Willmott, Lamas & J. Hall, 2017 ref.12 Actinote alcione (Hewitson, 1852) ref.12 iN Actinote alcione alcione (Hewitson, 1852) TL: Ecuador M T S Actinote alcione corduba (Hewitson, 1874) TL: Bolivia M T S Actinote alcione elatus H. Druce, 1903 TL: Ecuador M T Actinote alcione flaugeri Neild & Romero, 2008 TL: Venezuela Actinote alcione miguelangeli Neild & Romero, 2008 TL: Venezuela Actinote alcione sodalis A. Butler, 1877 TL: Peru M T S L Actinote alcione theophila (Dognin, 1887) TL: Ecuador M T Actinote alcione varians Jordan, 1910 TL: Colombia M T Actinote anaxo (Hopffer, 1874) ref.12 iN Actinote anaxo anaxo (Hopffer, 1874) TL: Peru M T Actinote anaxo comta Jordan, 1910 TL: Peru M T Actinote anaxo [n. ssp.] Lamas, MS Peru Actinote callianthe (C. Felder & R. Felder, 1862) ref.12 iN Actinote callianthe callianthe (C. Felder & R. Felder, 1862) TL: Colombia M T Actinote callianthe alla Jordan, 1910 TL: Venezuela M T Actinote callianthe amoena Jordan, 1913 TL: Colombia M T Actinote callianthe borgesi Neild & Romero, 2008 TL: Venezuela Actinote callianthe rufa Jordan, 1910 TL: Ecuador M T Actinote dicaeus (Latreille, [1817]) ref.12 iN Actinote dicaeus dicaeus (Latreille, [1817]) TL: ["Peru"] M T Actinote dicaeus albofasciata (Hewitson, 1869) TL: Ecuador M T S L Actinote dicaeus amida (Hewitson, 1854) TL: Colombia M T Actinote dicaeus callianira Geyer, 1837 TL: Mexico? M T Actinote dicaeus carica (Weymer, 1890) TL: Peru M T Actinote dicaeus eulalia Oberthür, 1917 TL: Colombia M T Actinote dicaeus euris Jordan, 1910 TL: Peru M T S Actinote dicaeus jucunda Jordan, 1910 TL: Bolivia M T Actinote dicaeus laverna (E. Doubleday, [1847]) TL: Venezuela M T Actinote dicaeus luisi Neild & Romero, 2008 TL: Venezuela Actinote dicaeus ozinta (Schaus, 1902) TL: Venezuela M T Actinote eresia (C. Felder & R. Felder, 1862) ref.12 iN Actinote eresia eresia (C. Felder & R. Felder, 1862) TL: Colombia M T Actinote eresia binghamae Dyar, 1913 TL: Peru M T Actinote eresia eresina (Hopffer, 1874) TL: Bolivia M T Actinote eresia lacrymosa Oberthür, 1917 TL: Colombia M T Actinote eresia leptogramma Jordan, 1913 TL: Ecuador M T Actinote eresia [n. ssp.#1] Lamas, MS TL: Peru Actinote eresia [n. ssp.#2] Lamas, MS TL: Peru Actinote eresia [n. ssp.#3] Lamas, MS TL: Peru Actinote griseata A. Butler, 1873 TL: Peru M T Actinote hilaris Jordan, 1910 ref.12 iN Actinote hilaris hilaris Jordan, 1910 TL: Peru M T S L Actinote hilaris desmiala Jordan, 1913 TL: Ecuador M T S Actinote hilaris [n. ssp.] Lamas, MS Peru Actinote momina Jordan, 1910 ref.12 M T Actinote negra (C. Felder & R. Felder, 1862) ref.12 iN Actinote negra negra (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T L Actinote negra avuncula Oberthür, 1917 TL: Peru M T Actinote negra demonica (Hopffer, 1874) TL: Bolivia M T S L Actinote negra euclia (Dognin, 1887) TL: Ecuador M T S Actinote negra menoetes Oberthür, 1917 TL: Ecuador M T Actinote negra scotosis Jordan, 1910 TL: Ecuador M T Actinote negra sobrina Jordan, 1910 TL: Peru M T Actinote negra [n. ssp.] Lamas, MS Peru Actinote neleus (Latreille, [1813]) ref.12 iN Actinote neleus neleus (Latreille, [1813]) TL: [S. Amer.] M T S Actinote neleus bermudezi Neild & Romero, 2008 TL: Venezuela Actinote ozomene (Godart, 1819) ref.12 iN Actinote ozomene ozomene (Godart, 1819) TL: S. Amer. M T Actinote ozomene calimene Rebel, 1902 TL: [Colombia] M T Actinote ozomene carbonaria (Hopffer, 1874) TL: Bolivia M T Actinote ozomene cleasa (Hewitson, 1868) TL: Ecuador M T Actinote ozomene gabrielae Rebel, 1902 TL: Ecuador M T Actinote ozomene nox (H. Bates, 1864) [= leucomelas], E & S Mexico to Panama M T S1 S2 L I Actinote rubrocellulata Hayward, 1960 ref.12 iN Actinote rubrocellulata rubrocellulata Hayward, 1960 TL: Peru M T S Actinote rubrocellulata [n. ssp.] Lamas, MS Peru Actinote stratonice (Latreille, [1813]) ref.12 iN Actinote stratonice stratonice (Latreille, [1813]) TL: [Colombia] M T L Actinote stratonice acipha (Hewitson, 1861) TL: Colombia M T Actinote stratonice adoxa Jordan, 1910 TL: Colombia M T Actinote stratonice aereta Jordan, 1913 TL: Ecuador M T Actinote stratonice marthae Jordan, 1910 TL: Colombia M T Actinote stratonice meridana Jordan, 1910 TL: Venezuela M T Actinote stratonice oaxaca J. Miller & L. Miller, 1979 S Oaxaca (Sierra Madre del Sur) M T S L Actinote tenebrosa (Hewitson, 1868) ref.12 iN Actinote tenebrosa tenebrosa (Hewitson, 1868) TL: Ecuador M T S Actinote tenebrosa erebia Oberthür, 1917 TL: Peru M T L Actinote tenebrosa [n. ssp.] Lamas, MS Peru Actinote trinacria (C. Felder & R. Felder, 1862) ref.12 iN Actinote trinacria trinacria (C. Felder & R. Felder, 1862) TL: Colombia M T Actinote trinacria chea H. Druce, 1903 TL: Colombia M T Actinote trinacria naura (H. Druce, 1875) TL: Venezuela M T Actinote trinacria santamarta (Winhard, 2017) ref.12,Winhard_2017 Actinote trinacria tenebrarum Oberthür, 1917 TL: Colombia M T Actinote trinacria vivasi Neild & Romero, 2008 TL: Venezuela Actinote [n. sp.#1] Lamas, MS TL: Peru Actinote [n. sp.#2] Lamas, MS TL: Peru Actinote alalia (C. Felder & R. Felder, 1860) iN M T Actinote alberti Neild & Romero, 2008 TL: Venezuela Actinote alberti alberti Neild & Romero, 2008 ref.266 M T Actinote alberti skinneri Romero & Neild, 2008 ref.266 M T Actinote anteas (E. Doubleday, [1847]) iN Actinote anteas Central American segregate , S Mexico to Panama M S L Actinote anteas anteas (E. Doubleday, [1847]) TL: Venezuela M T Actinote anteas brettia Oberthür, 1917 TL: Colombia M T Actinote anteas byssa Oberthür, 1917 TL: Venezuela M T Actinote anteas cedestis Jordan, 1913 TL: Ecuador M T Actinote anteas crassinia (Hopffer, 1874) TL: Peru M T S Actinote anteas pierrei Neild, 2008 ref.266 Actinote anteas suspecta Jordan, 1913 TL: Ecuador M T Actinote anteas terpsinoe (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Actinote ballettae Neild & Romero, 2008 TL: Venezuela Actinote ballettae ballettae Neild & Romero, 2008 ref.266 M T Actinote ballettae bonyae Neild & Romero, 2008 ref.266 Actinote bonita Penz, 1996 TL: Brazil (MG) M T Actinote brylla Oberthür, 1917 iN M T Actinote canutia (Hopffer, 1874) iN M T Actinote carycina Jordan, 1913 iN M T S Actinote catarina Penz, 1996 iN M T Actinote conspicua Jordan, 1913 iN M T Actinote dalmeidai Francini, 1996 iN M T Actinote discrepans R.F. d'Almeida, 1958 iN M T Actinote eberti Francini, Freitas & Penz, 2004 ref.205 Actinote genitrix R.F. d'Almeida, 1922 iN Actinote genitrix genitrix R.F. d'Almeida, 1922 TL: Brazil ([RJ]) M T Actinote genitrix costae Neild, 2008 ref.266 Actinote guatemalena (H. Bates, 1864) Guatemalan Actinote iN Actinote guatemalena guatemalena (H. Bates, 1864) Chiapas to Costa Rica M T S L Actinote guatemalena guerrerensis J. Maza, 1982 SW Mexico (Sierra Madre del Sur) M T S Actinote guatemalena veraecrucis Jordan, 1913 SE Mexico M T S1 S2 Actinote kennethi Freitas, Willmott & J. Hall, 2009 ref.312 M S Actinote lapitha (Staudinger, 1885) Pale Actinote iN Actinote lapitha lapitha (Staudinger, 1885) C Chiapas to Costa Rica & Panama M T S Actinote lapitha calderoni Schaus, 1920 W Chiapas to El Salvador M T S Actinote latior Jordan, 1913 TL: Colombia M T Actinote mamita (Burmeister, 1861) iN Actinote mamita mamita (Burmeister, 1861) TL: Argentina M T S Actinote mamita elena A. Hall, 1921 TL: Colombia M T Actinote mamita mitama (Schaus, 1902) TL: Brazil (Pr) M T Actinote melampeplos Godman & Salvin, 1881 Bow-winged Actinote S Mexico (S Oaxaca), Ecuador iN Actinote melampeplos melampeplos Godman & Salvin, 1881 TL: Costa Rica M T S L Actinote melampeplos [n. ssp.] Lamas, MS Ecuador Actinote melanisans Oberthür, 1917 iN M T Actinote morio Oberthür, 1917 TL: Brazil (MG) M T Actinote parapheles Jordan, 1913 iN M T Actinote pellenea Hübner, [1821] iN Actinote pellenea pellenea Hübner, [1821] TL: [Brazil] M T Actinote pellenea adriana (Hopffer, 1874) TL: Bolivia M T Actinote pellenea calymma Jordan, 1913 TL: Paraguay M T1 T2 Actinote pellenea crucis Jordan, 1913 TL: Bolivia M T Actinote pellenea epiphaea Jordan, 1913 TL: Peru M T Actinote pellenea equatoria (H. Bates, 1864) TL: Ecuador M T Actinote pellenea fernandezi Neild & Romero, 2008 ref.266 Actinote pellenea hahneli Jordan, 1913 TL: Venezuela M T Actinote pellenea hyalina Jordan, 1913 TL: [Peru] M T Actinote pellenea limbata Jordan, 1913 TL: [Colombia] M T Actinote pellenea mucia (Hopffer, 1874) TL: Peru M T Actinote pellenea perfulva Jordan, 1913 TL: Colombia M T Actinote pellenea subbadia Jordan, 1913 TL: Venezuela M T Actinote pellenea subhyalina (Staudinger, 1885) TL: Peru M T Actinote pellenea trinitatis Jordan, 1913 TL: Trinidad M T Actinote pellenea [n. ssp.#1] Lamas, MS TL: Ecuador Actinote pellenea [n. ssp.#2] Lamas, MS TL: Peru Actinote pellenea [n. ssp.#3] Lamas, MS TL: Peru Actinote pellenea [n. ssp.#4] Lamas, MS TL: Bolivia Actinote pratensis Francini, Freitas & Penz, 2004 ref.205 iN Actinote quadra (Schaus, 1902) iN M T Actinote rhodope R.F. d'Almeida, 1923 iN M T S Actinote romeroi Neild & Costa, 2008 ref.266 M T Actinote rufina Oberthür, 1917 iN M T S Actinote surima (Schaus, 1902) iN Actinote surima surima (Schaus, 1902) TL: Brazil (Pr) M T Actinote surima perisa Jordan, 1913 TL: Argentina M T Actinote thalia (Linnaeus, 1758) iN Actinote thalia thalia (Linnaeus, 1758) TL: Indiis M T S Actinote thalia eupelia Jordan, 1913 TL: Bolivia M T Actinote thalia pyrrha (Fabricius, 1775) TL: Brazil ([RJ]) M T S Actinote thalia sainorum Neild & Romero, 2008 ref.266 M S Actinote zikani R.F. d'Almeida, 1951 TL: Brazil (SP) M T Actinote [n. sp.#3] Lamas, MS TL: Peru Actinote [n. sp.#4] Lamas, MS TL: Peru Actinote [n. sp.#5] Lamas, MS TL: Peru Actinote [n. sp.#6] Lamas, MS TL: Peru Actinote [n. sp.#7] Lamas, MS TL: Peru Actinote [n. sp.#8] Penz, MS TL: Brazil Tribe Heliconiini Dione Hübner, [1819] Dione incarnata N. Riley, 1926 Gulf Fritillary ref.11,88, iN.312743 Dione incarnata incarnata N. Riley, 1926 SW USA, C & S TX (strays far N) through C Amer.; HI; ref.11,88 M T S1 S2 L I H Dione incarnata nigrior (Michener, 1942) SE USA (strays far N); ref.11,88 iN M T S1 S2 Ab HS L1 L2 I H Dione insularis (Maynard, 1889) Gulf Fritillary Bahamas, West Indies except southern Lesser Antilles; ref.88 M T S L Dione vanillae (Linnaeus, 1758) Gulf Fritillary S Lesser Antilles (St. Lucia, Barbados), Trinidad, N S. Amer. ...; ref.10,88 iN M S HS L Dione forbesi (Michener, 1942) ref.88 Dione galapagensis (W. Holland, 1890) ref.88 M T S Dione lucina (C. Felder & R. Felder, 1862) ref.88 M T Dione maculosa Stichel, [1908] ref.88 M T Dione dodona Lamas & Farfán, 2022 Peru; ref.89 M T Dione moneta Hübner, [1825] iN Dione moneta poeyii A. Butler, 1873 E & W Mexico (strays N into USA) to Panama M T S L I H Dione moneta butleri Stichel, [1908] TL: Colombia M T Dione moneta moneta Hübner, [1825] TL: ? Dione glycera (C. Felder & R. Felder, 1861) iN M T S Dione juno (Cramer, 1779) Juno Longwing iN Dione juno huascuma (Reakirt, 1866) E & W Mexico through Central America M S Ab L1 L2 I Dione juno juno (Cramer, 1779) Lesser Antilles from Martinique southward, NE South America M S Dione juno andicola (H. Bates, 1864) TL: Ecuador M T Dione juno miraculosa M. Hering, 1926 TL: Peru M T L Dione juno suffumata K. Brown & O. Mielke, 1972 TL: Paraguay Podotricha Michener, 1942 Podotricha judith (Guérin-Méneville, [1844]) iN Podotricha judith judith (Guérin-Méneville, [1844]) TL: Colombia M T Podotricha judith caucana (N. Riley, 1926) TL: Colombia M T Podotricha judith mellosa (Stichel, 1906) TL: Ecuador M T L Podotricha judith straminea (N. Riley, 1926) TL: Ecuador Podotricha telesiphe (Hewitson, 1867) iN Podotricha telesiphe telesiphe (Hewitson, 1867) TL: Ecuador M T S Podotricha telesiphe tithraustes (Salvin, 1871) TL: Ecuador M T Dryadula Michener, 1942 Dryadula phaetusa (Linnaeus, 1758) Banded Longwing S TX (rare stray), E & S Mexico to Brazil iN M T S HS L1 L2 Dryas Hübner, [1807] Dryas iulia (Fabricius, 1775) Julia iN Dryas iulia largo Clench, 1975 S FL (strays N) M S L I Dryas iulia nudeola (M. Bates, 1934) Cuba M T S L Dryas iulia carteri (N. Riley, 1926) N Bahamas M T S L Dryas iulia zoe L. Miller & Steinhauser, 1992 Grand Cayman & Cayman Brac M T Dryas iulia delila (Fabricius, 1775) Jamaica M S Dryas iulia fucatus (Boddaert, 1783) [= hispaniola], Hispaniola & satellite islands M S1 S2 L Dryas iulia iulia (Fabricius, 1775) Mona, Puerto Rico, Tortola, St. John, St. Thomas, St. Croix M T S L Dryas dominicana (A. Hall, 1917) Julia Guadeloupe & Dominica; ref.88 Dryas dominicana warneri (A. Hall, 1936) St. Kitts, Montserrat, Saba M T S Dryas dominicana dominicana (A. Hall, 1917) Guadeloupe & Dominica M T S Dryas lucia (N. Riley, 1926) Julia St. Lucia; ref.88 Dryas lucia martinica Enrico & Pinchon, 1969 Martinique M T S Dryas lucia lucia (N. Riley, 1926) St. Lucia M T S Dryas alcionea (Cramer, 1779) ref.88 Dryas alcionea framptoni (N. Riley, 1926) St. Vincent, some Grenadines, Grenada, [Barbados?] M T S L Dryas alcionea alcionea (Cramer, 1779) TL: Surinam M T S Dryas alcionea moderata (N. Riley, 1926) S TX (strays far N), E & W Mexico through Central America M S L1 L2 I H Ha Philaethria Billberg, 1820 Philaethria diatonica (Fruhstorfer, 1912) Northern Green Longwing [= Central American "dido" records], S Mexico to Panama (strays) iN M T S L I Philaethria dido (Linnaeus, 1763) iN Philaethria dido panamensis Constantino & Salazar, 2010 TL: Panama M T Philaethria dido chocoensis Constantino, 1999 TL: Colombia M T Philaethria dido dido (Linnaeus, 1763) TL: S. Amer. M S HS Philaethria constantinoi Salazar, 1991 TL: Colombia M T S Philaethria browni Constantino & Salazar, 2010 TL: Venezuela M T Philaethria romeroi Constantino & Salazar, 2010 TL: Venezuela M T Philaethria neildi Constantino & Salazar, 2010 TL: Venezuela Philaethria neildi winhardi Constantino & Salazar, 2010 TL: Colombia M T Philaethria neildi neildi Constantino & Salazar, 2010 TL: Venezuela M T Philaethria neildi tachiraensis Constantino & Salazar, 2010 TL: Venezuela M T Philaethria ostara (Röber, 1906) iN Philaethria ostara araguensis Constantino & Salazar, 2010 TL: Venezuela M T Philaethria ostara meridensis Constantino & Salazar, 2010 TL: Venezuela M T Philaethria ostara ostara (Röber, 1906) TL: Colombia M S Philaethria andrei Brévignon, 2002 iN Philaethria andrei orinocensis Constantino & Salazar, 2010 TL: Venezuela M T Philaethria andrei andrei Brévignon, 2002 TL: French Guiana M T Philaethria pygmalion (Fruhstorfer, 1912) iN Philaethria pygmalion metaensis Constantino & Salazar, 2010 TL: Colombia M T Philaethria pygmalion pygmalion (Fruhstorfer, 1912) TL: Brazil (Pa) M T S Philaethria wernickei (Röber, 1906) iN M S Neruda J. Turner, 1976 Neruda aoede (Hübner, [1813]) iN Neruda aoede aoede (Hübner, [1813]) TL: [Brazil] Neruda aoede aliciae Neukirchen, 2000 TL: Ecuador M T Neruda aoede astydamia (Erichson, [1849]) TL: Guyana M T Neruda aoede auca Neukirchen, 1997 TL: Ecuador M T Neruda aoede ayacuchensis (Neukirchen, 1992) TL: Venezuela M T Neruda aoede bartletti (H. Druce, 1876) TL: Peru M T Neruda aoede centurius (Neukirchen, 1994) TL: French Guiana M T Neruda aoede cupidineus (Stichel, 1906) TL: Peru M T Neruda aoede emmelina (Oberthür, 1902) TL: Guyana M T Neruda aoede eurycleia (K. Brown, 1973) TL: Brazil (MT) M T Neruda aoede faleria (Fruhstorfer, 1910) TL: Brazil (MT) Neruda aoede lucretius (Weymer, 1891) TL: Brazil (Am) M T Neruda aoede manu (Lamas, 1976) TL: Peru Neruda aoede philipi (K. Brown, 1976) TL: Bolivia M T Neruda aoede [n. ssp.] K. Brown, MS Brazil Neruda godmani (Staudinger, 1882) TL: Colombia M T Neruda metharme (Erichson, [1849]) iN Neruda metharme metharme (Erichson, [1849]) TL: Guyana M T S Neruda metharme adelinae (Cast, 2019) ref.182 Neruda metharme makiritare K. Brown & F. Fernández, 1985 TL: Venezuela M T Neruda metharme perseis (Stichel, 1923) TL: Colombia M T Neruda metis Moreira & O. Mielke, 2010 ref.248 Eueides Hübner, 1816 Eueides aliphera (Godart, 1819) iN Eueides aliphera aliphera (Godart, 1819) TL: Brazil M T S Eueides aliphera cyllenella Seitz, 1912 TL: ? M T Eueides aliphera gracilis Stichel, 1903 E & S Mexico to Panama M T S L Ha Eueides emsleyi K. Brown, 1976 TL: Colombia Eueides emsleyi emsleyi K. Brown, 1976 TL: Colombia Eueides emsleyi esmeraldensis Cast, 2013 TL: Ecuador M T Eueides heliconioides C. Felder & R. Felder, 1861 iN Eueides heliconioides heliconioides C. Felder & R. Felder, 1861 TL: Ecuador M T S L Eueides heliconioides eanes Hewitson, 1861 TL: Peru M T Eueides heliconioides koenigi H. Holzinger & R. Holzinger, 1993 TL: Peru Eueides isabella (Stoll, 1781) iN Eueides isabella isabella (Stoll, 1781) TL: Surinam Eueides isabella arquata Stichel, 1903 TL: Colombia M T Eueides isabella cleobaea Geyer, 1832 Cuba M S Eueides isabella dianasa (Hübner, [1806]) TL: Brazil (Ba) M T Eueides isabella dissoluta Stichel, 1903 TL: Peru M T Eueides isabella dynastes C. Felder & R. Felder, 1861 TL: Venezuela; Colombia M T Eueides isabella ecuadorensis Strand, 1912 TL: Ecuador Eueides isabella eva (Fabricius, 1793) [= zoracon], E Mexico (strays far N) through Central America M T S L1 L2 I Eueides isabella hippolinus A. Butler, 1873 TL: Peru M T Eueides isabella huebneri Ménétriés, 1857 TL: Brazil M T Eueides isabella melphis (Godart, 1819) Hispaniola M T S Eueides isabella nigricornis R. G. Maza, 1982 SW Mexico M T Eueides isabella [n. ssp.] K. Brown, MS Brazil (Ce) Eueides lampeto H. Bates, 1862 iN Eueides lampeto lampeto H. Bates, 1862 TL: Brazil (Am) M T Eueides lampeto acacetes Hewitson, 1869 TL: Ecuador M T Eueides lampeto apicalis Röber, 1927 TL: Ecuador M T Eueides lampeto brownsbergensis Gernaat & Beckles, 2010 TL: Suriname Eueides lampeto carbo Seitz, 1913 TL: Ecuador M T Eueides lampeto concisa Lamas, 1985 TL: Peru M T Eueides lampeto nigrofulva Kaye, 1906 TL: Guyana M T Eueides lampeto [n. ssp.#1] K. Brown, MS TL: French Guiana Eueides lampeto [n. ssp.#2] K. Brown, MS TL: Brazil (Am) Eueides libitina Staudinger, 1885 iN Eueides libitina libitina Staudinger, 1885 TL: French Guiana M T Eueides libitina malleti Lamas, 1998 TL: Peru M T Eueides lineata Salvin & Godman, 1868 Thick-edged Longwing S Mexico to Panama iN M T S L Eueides lybia (Fabricius, 1775) iN Eueides lybia lybia (Fabricius, 1775) TL: India Eueides lybia lybioides Staudinger, 1876 Pacific slope of Costa Rica and Panama M T S L Eueides lybia olympia (Fabricius, 1793) Atlantic slope from Nicaragua to Colombia M T S L Eueides lybia orinocensis K. Brown & F. Fernández, 1985 TL: Venezuela M T Eueides lybia otelloi K. Brown & F. Fernández, 1985 TL: Venezuela M T Eueides lybia salcedoi K. Brown & F. Fernández, 1985 TL: Venezuela M T Eueides pavana Ménétriés, 1857 iN M T Eueides procula E. Doubleday, [1847] iN Eueides procula procula E. Doubleday, [1847] TL: Venezuela M T Eueides procula asidia Schaus, 1920 SE Mexico to Nicaragua M T S Ab L Eueides procula browni H. Holzinger & R. Holzinger, 1974 TL: Venezuela Eueides procula edias Hewitson, 1861 TL: Colombia M T Eueides procula eurysaces Hewitson, 1864 TL: Ecuador M T Eueides procula kuenowii Dewitz, 1877 TL: Colombia M T Eueides procula vulgiformis A. Butler & H. Druce, 1872 Costa Rica & Panama M T S Eueides tales (Cramer, 1775) iN Eueides tales tales (Cramer, 1775) TL: Surinam Eueides tales barcellinus J. Zikán, 1937 TL: Brazil (Am) M T Eueides tales calathus Stichel, 1909 TL: Ecuador M T Eueides tales cognata Weymer, 1890 TL: Colombia M T Eueides tales franciscus K. Brown & H. Holzinger, 1973 TL: Venezuela M T Eueides tales llanera Winhard, 2019 ref.360 Eueides tales michaeli J. Zikán, 1937 TL: Peru M T Eueides tales pseudeanes Boullet & Le Cerf, 1910 TL: Venezuela M T Eueides tales pythagoras W. F. Kirby, [1900] TL: Brazil (Pa) M T S Eueides tales surdus Stichel, 1903 TL: Brazil (Pa); Guyana M T Eueides tales tabernula Lamas, 1985 TL: Peru Eueides tales xenophanes C. Felder & R. Felder, 1865 TL: Colombia M T Eueides tales [n. ssp.#1] K. Brown, MS TL: French Guiana Eueides tales [n. ssp.#2] K. Brown, MS TL: Brazil (Pa) Eueides tales guyanensis Lacomme, 2011 TL: not specified Eueides vibilia (Godart, 1819) iN Eueides vibilia vibilia (Godart, 1819) TL: Brazil M T Eueides vibilia mereaui (Hübner, 1823) TL: Brazil M T Eueides vibilia unifasciatus A. Butler, 1873 TL: Brazil (Am) M T Eueides vibilia vialis Stichel, 1903 SE Mexico to Panama M T S L Eueides vibilia vicinalis Stichel, 1903 TL: Ecuador M T Eueides vibilia [n. ssp.] K. Brown, MS Colombia Heliconius Kluk, 1780 Heliconius antiochus (Linnaeus, 1767) iN Heliconius antiochus antiochus (Linnaeus, 1767) TL: Indiis M T Heliconius antiochus aranea (Fabricius, 1793) TL: ? Heliconius antiochus araneides Staudinger, 1897 TL: Venezuela M T Heliconius antiochus salvinii Dewitz, 1877 TL: [Venezuela] M T Heliconius antiochus [n. ssp.] Lamas, MS Peru Heliconius astraea Staudinger, 1897 TL: Brazil (Am) Heliconius astraea astraea Staudinger, 1897 TL: Brazil (Am) M T Heliconius astraea rondonia K. Brown, 1973 TL: Brazil (Rd) M T S Heliconius atthis (E. Doubleday, 1847) iN M T L Heliconius besckei (Ménétriés, 1857) iN M T Heliconius burneyi (Hübner, [1831]) iN Heliconius burneyi burneyi (Hübner, [1831]) TL: Brazil M T Heliconius burneyi ada Neustetter, 1925 TL: Brazil (Rd) M T Heliconius burneyi anjae Neukirchen, 1995 TL: Brazil (Am) M T Heliconius burneyi boliviensis Neukirchen, 1995 TL: Bolivia M T Heliconius burneyi catharinae Staudinger, 1885 TL: [Brazil (Pa)] M T Heliconius burneyi huebneri Staudinger, 1897 TL: [Peru] M T Heliconius burneyi jamesi Neukirchen, 1995 TL: Peru M T Heliconius burneyi koenigi Neukirchen, 1995 TL: Peru M T Heliconius burneyi lindigii C. Felder & R. Felder, 1865 TL: Colombia M T Heliconius burneyi skinneri K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius charithonia (Linnaeus, 1767) Zebra Longwing iN Heliconius charithonia vazquezae W. Comstock & F. Brown, 1950 S TX (strays far N), E & W Mexico through Central America M T S Ab L1 L2 I H Ha Heliconius charithonia tuckeri W. Comstock & F. Brown, 1950 E TX to FL (strays N), Grand Bahama, Great Abaco M T S L I Heliconius charithonia ramsdeni W. Comstock & F. Brown, 1950 Cuba, Andros, [New Providence?] M T S Heliconius charithonia simulator Röber, 1921 Jamaica M S L Heliconius charithonia churchi W. Comstock & F. Brown, 1950 Hispaniola & satellite islands M T S L Heliconius charithonia charithonia (Linnaeus, 1767) Mona, Puerto Rico, Virgin Islands, some Leeward Is. to Montserrat M T S L Heliconius charithonia antiquus Lamas, 1988 St Kitts & Antigua; as an occasional form on other islands M T S Heliconius charithonia bassleri W. Comstock & F. Brown, 1950 TL: Colombia M T Heliconius clysonymus Latreille, [1817] iN Heliconius clysonymus clysonymus Latreille, [1817] TL: [Colombia] M T Heliconius clysonymus hygiana (Hewitson, 1867) TL: Ecuador M T Heliconius clysonymus montanus Salvin, 1871 Costa Rica & Panama M T TI L Heliconius clysonymus tabaconas K. Brown, 1976 TL: Peru M T Heliconius congener Weymer, 1890 iN Heliconius congener congener Weymer, 1890 TL: Ecuador M T L Heliconius congener aquilionaris K. Brown, 1976 TL: Colombia M T Heliconius congener ocannensis Stichel, 1905 TL: Colombia M T Heliconius cydno (E. Doubleday, 1847) iN Heliconius cydno cydno (E. Doubleday, 1847) TL: Colombia M T Heliconius cydno alithea (Hewitson, 1869) TL: Ecuador M T L Heliconius cydno barinasensis Masters, 1973 TL: Venezuela Heliconius cydno chioneus H. Bates, 1864 SE Costa Rica to Colombia M T S L Heliconius cydno cordula Neustetter, 1913 TL: Venezuela M T Heliconius cydno cydnides Staudinger, 1885 TL: Colombia M T Heliconius cydno gadouae K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius cydno galanthus H. Bates, 1864 SE Mexico to E Costa Rica M T S L Heliconius cydno hermogenes (Hewitson, [1858]) TL: [Colombia] M T Heliconius cydno lisethae Neukirchen, 1995 TL: Colombia M T Heliconius cydno wanningeri Neukirchen, 1991 TL: Colombia M T Heliconius cydno weymeri Staudinger, 1897 TL: Colombia M T Ab Heliconius cydno zelinde A. Butler, 1869 TL: W coast of America M T Ab Heliconius pachinus Salvin, 1871 Pachinus Longwing Pacific slope of Costa Rica and Panama iN M T S L I Heliconius demeter Staudinger, 1897 iN Heliconius demeter demeter Staudinger, 1897 TL: Peru M T Heliconius demeter angeli Neukirchen, 1997 TL: Peru Heliconius demeter beebei J. Turner, 1966 TL: Guyana M T Heliconius demeter bouqueti Nöldner, 1901 TL: French Guiana M T Heliconius demeter karinae Neukirchen, 1990 TL: Brazil (Pa) M T Heliconius demeter neildi Neukirchen, 1997 TL: Ecuador M T Heliconius demeter terrasanta K. Brown & Benson, 1975 TL: Brazil (Pa) M T Heliconius demeter titan Neukirchen, 1995 TL: Brazil (Am) M T Heliconius demeter turneri K. Brown & Benson, 1975 TL: Brazil (Am) M T Heliconius demeter zikani K. Brown & Benson, 1975 TL: Brazil (Am) M T Heliconius demeter joroni Lamas & Rosser, 2018 Zoological Journal of the Linnean Society 186(1): 233-249) Heliconius eratosignis (Joicey & Talbot, 1925) iN Heliconius eratosignis eratosignis (Joicey & Talbot, 1925) ref.291 M T Heliconius eratosignis tambopata Lamas, 1985 ref.291 Heliconius eratosignis ucayalensis H. Holzinger & R. Holzinger, 1975 ref.291 Heliconius eratosignis ulysses K. Brown & Benson, 1975 ref.291 M T Heliconius doris (Linnaeus, 1771) iN Heliconius doris viridis Staudinger, 1885 SE Mexico to Panama M T S L Heliconius doris doris (Linnaeus, 1771) TL: Surinam M T Heliconius doris dives (Oberthür, 1920) TL: Colombia M T Heliconius doris obscurus Weymer, 1891 TL: Colombia M T Heliconius doris [n. ssp.] Neukirchen, MS Trinidad Heliconius egeria (Cramer, 1775) TL: Surinam Heliconius egeria egeria (Cramer, 1775) TL: Surinam M T Heliconius egeria christiani Neukirchen, 1997 TL: Brazil (Pa) M T Heliconius egeria egerides Staudinger, 1897 TL: Brazil (Am) M T Heliconius egeria homogena Bryk, 1953 TL: Brazil (Am) M T Heliconius egeria hyas Weymer, 1883 TL: [Brazil (Am)] M T Heliconius egeria keithbrowni Neukirchen, 1992 TL: Venezuela M T Heliconius egeria mariasibyllae Neukirchen, 1991 TL: Brazil (Pa) M T Heliconius eleuchia (Hewitson, [1854]) White-edged Longwing iN Heliconius eleuchia eleuchia (Hewitson, [1854]) Costa Rica to Colombia M T L1 L2 Ha Heliconius eleuchia eleusinus Staudinger, 1885 TL: Colombia M T Heliconius eleuchia primularis A. Butler, 1869 TL: Ecuador M T L Heliconius elevatus Nöldner, 1901 iN Heliconius elevatus elevatus Nöldner, 1901 TL: Amazonas M T Heliconius elevatus bari (Oberthür, 1902) TL: French Guiana M T Heliconius elevatus lapis Lamas, 1976 TL: Peru Heliconius elevatus perchlora Joicey & Kaye, 1917 TL: Bolivia M T Heliconius elevatus pseudocupidineus Neustetter, 1931 TL: Peru M T Heliconius elevatus roraima J. Turner, 1966 TL: Guyana M T Heliconius elevatus schmassmanni Joicey & Talbot, 1925 TL: Brazil (MT) M T Heliconius elevatus sonjae Neukirchen, 1997 TL: Brazil (Pa) M T Heliconius elevatus taracuanus Bryk, 1953 TL: Brazil (Am) Heliconius elevatus tumatumari Kaye, 1906 TL: Guyana M T Heliconius elevatus willmotti Neukirchen, 1997 TL: Ecuador M T Heliconius elevatus zoelleri Neukirchen, 1990 TL: Venezuela M T Heliconius erato (Linnaeus, 1758) iN Heliconius erato erato (Linnaeus, 1758) TL: Surinam M T1 T2 HS Heliconius erato adana J. Turner, 1967 TL: [Trinidad] M T Heliconius erato amalfreda Riffarth, 1901 TL: Brazil (Pa) M T Heliconius erato amazona Staudinger, 1897 TL: Brazil (Pa) M T Heliconius erato amphitrite Riffarth, 1901 TL: Peru M T Heliconius erato chestertonii (Hewitson, 1872) TL: Colombia M T Heliconius erato colombina Staudinger, 1897 TL: Colombia M T Heliconius erato cruentus Lamas, 1998 W Mexico M T S L Heliconius erato cyrbia (Godart, 1819) TL: America M T Heliconius erato demophoon Ménétriés, 1855 Nicaragua M T OI S Heliconius erato dignus Stichel, 1923 TL: Colombia M T Heliconius erato emma Riffarth, 1901 TL: Peru; "Ecuador" M T Heliconius erato estrella H. Bates, 1862 TL: Brazil (Pa) M T Heliconius erato etylus Salvin, 1871 TL: Ecuador M T Heliconius erato favorinus Hopffer, 1874 TL: Bolivia M T Heliconius erato guarica Reakirt, 1868 TL: Colombia M T Heliconius erato hydara (Hewitson, 1867) Panama to Brazil M T1 T2 L Heliconius erato lativitta A. Butler, 1877 TL: Brazil (Am); Ecuador M T Heliconius erato lichyi K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius erato luscombei Lamas, 1976 TL: Peru Heliconius erato magnifica Riffarth, 1900 TL: Guyana Heliconius erato microclea Kaye, 1907 TL: Peru M T Heliconius erato notabilis Salvin & Godman, 1868 TL: Ecuador M T Heliconius erato petiverana (E. Doubleday, 1847) S TX, E Mexico to Panama M T S Ab L Heliconius erato phyllis (Fabricius, 1775) TL: Brazil M T Heliconius erato reductimacula Bryk, 1953 TL: Brazil (Am) Heliconius erato tobagoensis Barcant, 1982 TL: Tobago M T Heliconius erato venus Staudinger, 1882 SE Panama (Darien), W Colombia (Choco) M T L Heliconius erato venustus Salvin, 1871 TL: Bolivia M T Heliconius ethilla (Godart, 1819) iN Heliconius ethilla ethilla (Godart, 1819) TL: Antilles M T Heliconius ethilla adela Neustetter, 1912 TL: Peru M T Heliconius ethilla aerotome C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Heliconius ethilla cephallenia C. Felder & R. Felder, 1865 TL: Surinam M T Heliconius ethilla chapadensis K. Brown, 1973 TL: Brazil (MT) M T Heliconius ethilla claudia Godman & Salvin, 1881 Panama M T S Heliconius ethilla eucoma (Hübner, [1831]) TL: Brazil Heliconius ethilla flavofasciatus Weymer, 1894 TL: Brazil (Pa) M T Heliconius ethilla flavomaculatus Weymer, 1894 TL: Brazil (Pe) M T Heliconius ethilla hyalina Neustetter, 1928 TL: Brazil (Rr) M T Heliconius ethilla jaruensis K. Brown, 1976 TL: Brazil (Rd) M T Heliconius ethilla latona Neukirchen, 1998 TL: Colombia M T Heliconius ethilla mentor Weymer, 1883 TL: Colombia M T Heliconius ethilla metalilis A. Butler, 1873 TL: Venezuela M T Heliconius ethilla michaelianius Lamas, 1988 TL: Peru M T Heliconius ethilla narcaea (Godart, 1819) TL: Antilles M T Heliconius ethilla nebulosa Kaye, 1916 TL: Peru M T Heliconius ethilla neukircheni Lamas, 1998 TL: Peru M T Heliconius ethilla numismaticus Weymer, 1894 TL: Brazil (Pa) M T Heliconius ethilla penthesilea Neukirchen, 1994 TL: Brazil (Am) M T Heliconius ethilla polychrous C. Felder & R. Felder, 1865 TL: Brazil ([SP]) M T Heliconius ethilla semiflavidus Weymer, 1894 TL: Colombia M T Heliconius ethilla thielei Riffarth, 1900 TL: French Guiana M T Heliconius ethilla tyndarus Weymer, 1897 TL: Bolivia M T Heliconius ethilla yuruani K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius hecale (Fabricius, 1776) iN Heliconius hecale hecale (Fabricius, 1776) TL: Surinam M T Heliconius hecale anderida (Hewitson, [1853]) TL: [Venezuela] M T Heliconius hecale annetta Riffarth, 1900 TL: Colombia M T Heliconius hecale australis K. Brown, 1976 TL: Ecuador M T Heliconius hecale barcanti K. Brown, 1976 TL: Venezuela M T Heliconius hecale clearei A. Hall, 1930 TL: Guyana M T Heliconius hecale discomaculatus Weymer, 1891 Honduras M T Heliconius hecale ennius Weymer, 1891 TL: Brazil (Am) Heliconius hecale felix Weymer, 1894 TL: Peru M T Heliconius hecale fornarina (Hewitson, 1854) C & SW Chiapas to Guatemala M T S Heliconius hecale holcophorus Staudinger, 1897 TL: Colombia M T Heliconius hecale humboldti Neustetter, 1928 TL: Peru M T Heliconius hecale ithaca C. Felder & R. Felder, 1862 TL: Colombia M T Heliconius hecale latus Riffarth, 1900 TL: Brazil (Pa) M T Heliconius hecale melicerta H. Bates, 1866 Panama & Colombia M T S L Heliconius hecale metellus Weymer, 1894 TL: Brazil (Pa) M T Heliconius hecale naxos Neukirchen, 1998 TL: Brazil (Pa) Heliconius hecale nigrofasciatus Weymer, 1894 TL: Brazil (Am) M T S Heliconius hecale novatus H. Bates, 1867 TL: Brazil (Pa/Ma) M T Heliconius hecale paraensis Riffarth, 1900 TL: Brazil (Pa) M T Heliconius hecale paulus Neukirchen, 1998 TL: Brazil (Pa) Heliconius hecale quitalena (Hewitson, [1853]) TL: Ecuador M T Heliconius hecale rosalesi K. Brown & F. Fernández, 1976 TL: Venezuela M T Heliconius hecale shanki Lamas & K. Brown, 1976 TL: Peru Heliconius hecale sisyphus Salvin, 1871 TL: Peru M T Heliconius hecale sulphureus Weymer, 1894 TL: Brazil (Am) M T Heliconius hecale vetustus A. Butler, 1873 TL: Guyana M T Heliconius hecale zeus Neukirchen, 1995 TL: Bolivia M T Heliconius hecale zuleika (Hewitson, 1854) SE Mexico to Panama M T S1 S2 S3 L I H Heliconius hecalesia (Hewitson, [1854]) iN Heliconius hecalesia hecalesia (Hewitson, [1854]) TL: Colombia Heliconius hecalesia eximius Stichel, 1923 TL: Colombia Heliconius hecalesia formosus H. Bates, 1866 Costa Rica & Panama M T S L Heliconius hecalesia gynaesia (Hewitson, 1875) TL: ? M T Heliconius hecalesia longarena (Hewitson, 1875) TL: Colombia M T Heliconius hecalesia octavia H. Bates, 1866 SE Mexico to Nicaragua M T S L Heliconius hecalesia romeroi K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius hecuba (Hewitson, [1858]) iN Heliconius hecuba hecuba (Hewitson, [1858]) TL: Colombia M T Heliconius hecuba bonplandi Neukirchen, 1991 TL: Ecuador M T Heliconius hecuba cassandra C. Felder & R. Felder, 1862 TL: Colombia M T Heliconius hecuba choarina (Hewitson, 1872) TL: Ecuador M T Heliconius hecuba creusa H. Holzinger & R. Holzinger, 1989 TL: Colombia M T Heliconius hecuba crispus Staudinger, 1885 TL: Colombia Heliconius hecuba flava K. Brown, 1979 TL: Ecuador M T Heliconius hecuba lamasi Neukirchen, 1991 TL: Ecuador Heliconius hecuba salazari Neukirchen, 1993 TL: Colombia M T Heliconius hecuba tolima Fassl, 1912 TL: Colombia M T Heliconius hecuba walteri Salazar, 1998 TL: Colombia M T Heliconius hermathena (Hewitson, [1854]) iN Heliconius hermathena hermathena (Hewitson, [1854]) TL: [Brazil] M T Heliconius hermathena duckei K. Brown & Benson, 1977 TL: Brazil (Pa) M T Heliconius hermathena renatae K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius hermathena sabinae Neukirchen, 1992 TL: Brazil (Am) M T Heliconius hermathena sheppardi K. Brown & Benson, 1977 TL: Brazil (Am) M T Heliconius hermathena vereatta Stichel, 1912 TL: Brazil (Pa) M T Heliconius heurippa (Hewitson, [1854]) iN M S Heliconius hierax (Hewitson, 1869) iN M T Heliconius himera (Hewitson, 1867) iN M T Heliconius hortense (Guérin-Méneville, [1844]) Mexican Longwing E & W Mexico to El Salvador & Honduras iN M S Ab L Heliconius ismenius Latreille, [1817] iN Heliconius ismenius ismenius Latreille, [1817] TL: [S. Amer.] M T Heliconius ismenius boulleti Neustetter, 1928 TL: French Guiana M T Heliconius ismenius clarescens A. Butler, 1875 Costa Rica & Panama M T S L Heliconius ismenius fasciatus Godman & Salvin, 1877 Panama M T TI Heliconius ismenius metaphorus Weymer, 1883 TL: Ecuador M T Heliconius ismenius occidentalis Neustetter, 1928 TL: Colombia M T Heliconius ismenius telchinia (E. Doubleday, 1847) E Mexico to Panama M T S L Heliconius ismenius tilletti K. Brown & F. Fernández, 1976 TL: Venezuela M T Heliconius lalitae Brévignon, 1996 TL: French Guiana M T Heliconius leucadia H. Bates, 1862 iN Heliconius leucadia leucadia H. Bates, 1862 TL: Brazil (Am) M T Heliconius leucadia andromeda Neukirchen, 1996 TL: Ecuador M T Heliconius leucadia birgitae Neukirchen, 1996 TL: Peru M T Heliconius leucadia pseudorhea Staudinger, 1897 TL: Brazil (Am) M T S Heliconius luciana Lichy, 1960 TL: Venezuela Heliconius luciana luciana Lichy, 1970 TL: Venezuela M T Heliconius luciana watunna Lichy, 1960 TL: Venezuela Heliconius melpomene (Linnaeus, 1758) iN Heliconius melpomene melpomene (Linnaeus, 1758) TL: America M T1 T2 T3 Heliconius melpomene aglaope C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Heliconius melpomene amandus Grose-Smith & W. F. Kirby, 1892 TL: Bolivia M T Heliconius melpomene amaryllis C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Heliconius melpomene anduzei K. Brown & F. Fernández, 1985 TL: Venezuela M T Heliconius melpomene bellula J. Turner, 1971 TL: Colombia M T Heliconius melpomene burchelli Poulton, 1910 TL: Brazil (Go) M T Heliconius melpomene cythera (Hewitson, 1869) TL: Ecuador M T Heliconius melpomene ecuadorensis Emsley, 1964 TL: Ecuador M T Heliconius melpomene euryades Riffarth, 1900 TL: Peru M T Heliconius melpomene flagrans Stichel, 1919 TL: Trinidad M T Heliconius melpomene intersectus Neustetter, 1928 TL: Brazil ([Pa]) M T Heliconius melpomene madeira N. Riley, 1919 TL: Brazil (Am) M T Heliconius melpomene malleti Lamas, 1988 TL: Ecuador M T Heliconius melpomene martinae Cast & Le Crom, 2012 TL: Colombia M T Heliconius melpomene meriana J. Turner, 1967 TL: Surinam M T Heliconius melpomene michellae Neukirchen, 1997 TL: Peru M T Heliconius melpomene nanna Stichel, 1899 TL: Brazil (ES/MG) M T Heliconius melpomene penelope Staudinger, 1894 TL: Bolivia M T Heliconius melpomene plesseni Riffarth, 1907 TL: Ecuador M T Heliconius melpomene pyrforus Kaye, 1907 TL: Guyana M T Heliconius melpomene rosina (Boisduval, 1870) Costa Rica & Panama M T S L Heliconius melpomene schunkei Lamas, 1976 TL: Peru M T L Heliconius melpomene tessa Barcant, 1982 TL: Tobago M T Heliconius melpomene thelxiope (Hübner, [1806]) TL: [Brazil] Heliconius melpomene thelxiopeia Staudinger, 1897 TL: French Guiana M T Heliconius melpomene vicina Ménétriés, 1857 TL: Brazil M T Heliconius melpomene vulcanus A. Butler, 1865 Panama & Colombia M T L Heliconius melpomene xenoclea (Hewitson, [1853]) TL: ? M T Heliconius nattereri C. Felder & R. Felder, 1865 TL: Brazil ([Ba]) M T Heliconius numata (Cramer, 1780) iN Heliconius numata numata (Cramer, 1780) TL: Surinam M T Heliconius numata arcuella H. Druce, 1874 TL: Peru M T Heliconius numata aristiona (Hewitson, [1853]) TL: Colombia M T Heliconius numata aulicus Weymer, 1883 TL: [Venezuela] M T Heliconius numata aurora H. Bates, 1862 TL: Brazil (Am) M T Heliconius numata bicoloratus A. Butler, 1873 TL: Peru M T L Heliconius numata ethra (Hübner, [1831]) TL: Brazil M T Heliconius numata euphone C. Felder & R. Felder, 1862 TL: Colombia M T Heliconius numata geminatus Weymer, 1894 TL: Brazil (Am) M T Heliconius numata holzingeri F. Fernández & K. Brown, 1976 TL: Venezuela Heliconius numata ignotus Joicey & Kaye, 1917 TL: Peru M T Heliconius numata illustris Weymer, 1894 TL: Peru M T Heliconius numata jiparanaensis Neustetter, 1931 TL: Brazil (Rd) M T S Heliconius numata lenaeus Weymer, 1891 TL: Colombia Heliconius numata lyrcaeus Weymer, 1891 TL: Peru? M T Heliconius numata mavors Weymer, 1894 TL: [Brazil (Am)] M T Heliconius numata messene C. Felder & R. Felder, 1862 TL: Colombia M T Heliconius numata mirus Weymer, 1894 TL: Bolivia M T Heliconius numata nubifer A. Butler, 1875 TL: Brazil (Am) M T Heliconius numata peeblesi Joicey & Talbot, 1925 TL: Venezuela M T Heliconius numata pratti Joicey & Kaye, 1917 TL: Peru M T Heliconius numata robigus Weymer, 1875 TL: Venezuela M T Heliconius numata silvana (Stoll, 1781) TL: Surinam M T Heliconius numata sourensis K. Brown, 1976 TL: Brazil (Pa) M T Heliconius numata superioris A. Butler, 1875 TL: Brazil (Am/Pa) M T Heliconius numata talboti Joicey & Kaye, 1917 TL: Peru M T Heliconius numata tarapotensis Riffarth, 1901 TL: Peru M T Heliconius numata zobrysi Fruhstorfer, 1910 TL: Brazil (MT) M T Heliconius pardalinus H. Bates, 1862 iN Heliconius pardalinus pardalinus H. Bates, 1862 TL: Brazil (Am) M T Heliconius pardalinus ariadne Neukirchen, 1995 TL: Bolivia M T Heliconius pardalinus butleri K. Brown, 1976 TL: Peru M T Heliconius pardalinus dilatus Weymer, 1894 TL: Peru M T Heliconius pardalinus gloria Winhard, 2019 ref.360 Heliconius pardalinus julia Neukirchen, 2000 TL: Ecuador Heliconius pardalinus lucescens Weymer, 1894 TL: Brazil (Pa) M T Heliconius pardalinus maeon Weymer, 1891 TL: ? M T L Heliconius pardalinus orteguaza K. Brown, 1976 TL: Colombia M T Heliconius pardalinus radiosus A. Butler, 1873 TL: Brazil (Pa) M T Heliconius pardalinus sergestus Weymer, 1894 TL: Peru M T Heliconius pardalinus tithoreides Staudinger, 1900 TL: Peru M T Heliconius peruvianus (C. Felder & R. Felder, 1859) iN M S Heliconius ricini (Linnaeus, 1758) iN Heliconius ricini ricini (Linnaeus, 1758) TL: America M HS Heliconius ricini insulanus (Stichel, 1909) TL: Trinidad M T Heliconius sapho (Drury, 1782) Sapho Longwing iN Heliconius sapho sapho (Drury, 1782) Panama M OI IL L Heliconius sapho candidus K. Brown, 1976 TL: Ecuador M T Heliconius sapho chocoensis K. Brown & Benson, 1975 TL: Colombia M T Heliconius sapho leuce (E. Doubleday, 1847) SE Mexico to Costa Rica; Atlantic slope M S L I Heliconius hewitsoni (Hewitson, 1875) Hewitson's Longwing Costa Rica & Panama; Pacific slope iN M T S L Heliconius sara (Fabricius, 1793) iN Heliconius sara sara (Fabricius, 1793) TL: Guyana M T Heliconius sara apseudes (Hübner, [1813]) TL: Brazil M S Heliconius sara brevimaculata Riffarth, 1901 TL: Colombia M T Heliconius sara elektra Neukirchen, 1998 TL: Colombia M T Heliconius sara fulgidus Stichel, 1906 Guatemala to Panama; Atlantic slope M T L Heliconius sara magdalena H. Bates, 1864 Panama to Colombia & Venezuela M T S L Heliconius sara sprucei H. Bates, 1864 TL: Ecuador M T L Heliconius sara theudela (Hewitson, 1874) Costa Rica & Panama; Pacific slope M T S L I H Heliconius sara veraepacis H. Bates, 1864 SE Mexico to Costa Rica (Atlantic Slope) M T S L Heliconius sara williami Neukirchen, 1994 TL: Trinidad M T Heliconius telesiphe (E. Doubleday, 1847) iN Heliconius telesiphe telesiphe (E. Doubleday, 1847) TL: Bolivia M T L Heliconius telesiphe cretacea Neustetter, 1916 TL: French Guiana M T Heliconius telesiphe sotericus Salvin, 1871 TL: Ecuador M T Heliconius timareta (Hewitson, 1867) iN Heliconius timareta timareta (Hewitson, 1867) TL: Ecuador M T Heliconius timareta timoratus Lamas, 1998 TL: Peru M T Heliconius tristero A. V.Z. Brower, 1996 TL: Colombia M T Heliconius wallacei Reakirt, 1866 iN Heliconius wallacei wallacei Reakirt, 1866 TL: [Brazil] M T Heliconius wallacei araguaia K. Brown, 1976 TL: Brazil (Go) M T Heliconius wallacei colon Weymer, 1891 TL: [Brazil (Am)] M T Heliconius wallacei flavescens Weymer, 1891 TL: Ecuador M T L Heliconius wallacei kayei Neustetter, 1929 TL: Trinidad M T Heliconius wallacei mimulinus A. Butler, 1873 TL: Colombia M T Heliconius xanthocles H. Bates, 1862 iN Heliconius xanthocles xanthocles H. Bates, 1862 TL: Guyana; "French Guiana" M T Heliconius xanthocles buechei Neukirchen, 1992 TL: Venezuela M T Heliconius xanthocles cleoxanthe H. Holzinger & R. Holzinger, 1972 TL: Venezuela Heliconius xanthocles donatia Fruhstorfer, 1910 TL: Brazil (MT) M T Heliconius xanthocles explicata K. Brown, 1976 TL: Colombia M T Heliconius xanthocles hippocrene H. Holzinger & K. Brown, 1982 TL: Bolivia Heliconius xanthocles melete C. Felder & R. Felder, 1865 TL: Colombia M T Heliconius xanthocles melior Staudinger, 1897 TL: Peru M T Heliconius xanthocles melittus Staudinger, 1897 TL: Peru M T Heliconius xanthocles napoensis H. Holzinger & K. Brown, 1982 TL: Ecuador M S Heliconius xanthocles paraplesius H. Bates, 1867 TL: Brazil (Ma) M T Heliconius xanthocles quindecim Lamas, 1976 TL: Peru Heliconius xanthocles rindgei H. Holzinger & K. Brown, 1982 TL: Colombia M T Heliconius xanthocles similatus J. Zikán, 1937 TL: Brazil (Am) M T Heliconius xanthocles vala Staudinger, 1885 TL: French Guiana M T Heliconius xanthocles zamora H. Holzinger & K. Brown, 1982 TL: Ecuador Tribe Argynnini Euptoieta E. Doubleday, 1848 Euptoieta bogotana Staudinger, 1885 iN M T Euptoieta claudia (Cramer, 1775) Variegated Fritillary S N Am. (strays far N) to Panama, Bahamas, Gr. Antilles, Barbuda iN Euptoieta claudia claudia (Cramer, 1775) TL: Jamaica M S1 S2 HS L1 L2 L3 L4 L5 L6 I1 I2 I3 H F Euptoieta claudia daunius (Herbst, 1798) TL: East Indies M T Euptoieta hegesia (Cramer, 1779) Mexican Fritillary iN Euptoieta hegesia hegesia (Cramer, 1779) Bahamas, Caymans, Jamaica, Cuba, Hispaniola, Mona M T S Ab L Euptoieta hegesia meridiania Stichel, 1938 [= hoffmanni], S TX, W MX (strays N), E Mexico to Argentina M T S L1 L2 I1 I2 Euptoieta hegesia watsoni W. Comstock, 1944 Puerto Rico M T S L Euptoieta hortensia (Blanchard, 1852) iN M T Euptoieta perdistincta A. Hall, 1930 TL: Peru M T Euptoieta poasina Schaus, 1913 TL: Costa Rica M T Euptoieta sunides (Hewitson, 1877) TL: Ecuador M T Euptoieta thekla A. Hall, 1919 iN M T Euptoieta [n. sp.] Lamas, MS Peru Yramea Reuss, 1920 Yramea cytheris (Drury, 1773) iN Yramea cytheris cytheris (Drury, 1773) TL: Falkland Is. M T Yramea cytheris siga (Geyer, 1832) TL: Java M T Yramea inca (Staudinger, 1894) iN Yramea inca inca (Staudinger, 1894) TL: Bolivia M T Yramea inca cora (Lehmann, 1913) TL: Peru M T Yramea inca [n. ssp.] Lamas, MS Argentina Yramea lathonioides (Blanchard, 1852) iN M T Yramea modesta (Blanchard, 1852) iN M T Yramea sobrina (Weymer, 1890) iN M T Yramea [n. sp.] Lamas & Grados, MS Peru Boloria Moore, 1900 Boloria alaskensis (W. Holland, 1900) Mountain Fritillary iN Boloria alaskensis alaskensis (W. Holland, 1900) Nunavut W to AK, S to C BC M S L Ha Boloria alaskensis halli Klots, 1940 Wind River Mountains, NW WY M S1 S2 L Ha Boloria eunomia (Esper, 1800) Bog Fritillary iN Boloria eunomia triclaris (Hübner, [1821]) Labrador & vic. M S Boloria eunomia dawsoni (W. Barnes & McDunnough, 1916) N WI, MI, MN, S boreal Canada to Yukon in taiga habitats M T S1 S2 Boloria eunomia nichollae (W. Barnes & Benjamin, 1926) Rocky Mountains of C BC & ALTA M S Boloria eunomia denali (Klots, 1940) AK, N montane YK M S L Boloria eunomia ursadentis Ferris & Groothuis, 1971 Beartooth Plateau of N WY and S MT M S L Ha Boloria eunomia laddi (Klots, 1940) Snowy Range of Albany and Carbon Cos., WY, similar in Big Horn Mts of N-central WY M S1 S2 Boloria eunomia caelestis (Hemming, 1933) north and central Colorado Rocky Mountains M S L Ha Boloria myrina (Cramer, 1777) Silver-bordered Fritillary [former selene]; ref.16 Boloria myrina myrina (Cramer, 1777) TL: USA (NY) M S L Ha Boloria myrina nebraskensis (W. Holland, 1928) TL: USA (Ne) M S Boloria myrina sabulocollis Kohler, 1977 TL: USA (Ne) M S Boloria myrina tollandensis (W. Barnes & Benjamin, 1925) TL: USA (Co) M S1 S2 L I Ha Boloria myrina Pacific Northwestern s segregate , E OR, E WA, S ID, NW WY M S1 S2 L Boloria myrina albequina (W. Holland, 1928) TL: not specified M S Boloria myrina atrocostalis (Huard, 1927) TL: not specified M S Ab L Boloria myrina terraenovae (W. Holland, 1928) TL: not specified M S Boloria natazhati (A. Gibson, 1920) Cryptic Fritillary iN Boloria natazhati natazhati (A. Gibson, 1920) Wrangell & St. Elias Mts., Montana Mt., YK M T Boloria natazhati nabokovi D. Stallings & Turner, 1947 N BC, probably S YK, White Mt., MacKenzie Mts., Bernard H., NWT M S Ha Boloria natazhati bankslandia J. Shepard, 1998 Victoria Island, NWT (and adjacent mainland) M S Boloria freija (Thunberg, 1791) Freija Fritillary iN Boloria freija freija (Thunberg, 1791) TL: Sweden M T S1 S2 L Ha Boloria freija tarquinius (J. Curtis, 1835) E Canadian arctic tundra M S Boloria freija browni (Higgins, 1953) TL: USA (Co) M S L Ha Boloria chariclea (Schneider, 1794) Purplish Fritillary iN Boloria chariclea montinus (Scudder, 1863) TL: not specified M S L Ha Boloria chariclea boisduvalii (Duponchel, [1832]) SE Canadian arctic (occurrs between B. c. arctica & B. c. grandis) M S1 S2 Boloria chariclea arctica (Zetterstedt, [1839]) NE Canadian arctic to N YK M S Boloria chariclea grandis (W. Barnes & McDunnough, 1916) taiga from Labrador to YK & BC, [to MT?] M S1 S2 L Ha Boloria chariclea butleri (W. H. Edwards, 1883) montane AK & YK to N BC M S1 S2 L Ha Boloria chariclea rainieri (W. Barnes & McDunnough, 1913) TL: not specified M T S L Boloria chariclea ingens (W. Barnes & McDunnough, 1918) TL: USA (Wy) M S L Ha Boloria chariclea helena (W. H. Edwards, 1871) TL: USA (Co) M T S1 S2 Ab L1 L2 L3 Ha Boloria kriemhild (Strecker, 1879) Relict Fritillary iN M S L1 L2 Ha Boloria epithore (W. H. Edwards, 1864) Pacific Fritillary iN Boloria epithore sigridae (J. Shepard, 2001) TL: not specified M T Boloria epithore chermocki E. Perkins & S. Perkins, 1966 TL: USA (Or) M T S1 S2 L1 L2 Ha Boloria epithore NE Oregon segregate , NE Oregon and surrounding areas M S Ab L Boloria epithore epithore (W. H. Edwards, 1864) TL: USA (Ca) M S L H Boloria epithore sierra E. Perkins, 1973 TL: USA (Ca) M T S L Boloria improba (A. Butler, 1877) Dingy Fritillary iN Boloria improba improba (A. Butler, 1877) Scandanavia, Nunavut, NWT M S Boloria improba youngi (W. Holland, 1900) AK, YK M S L Ha Boloria improba nunatak Scott, 1998 Rocky Mountains of Alberta & British Columbia M T S Boloria improba harryi (Ferris, 1984) NW Wyoming M T S1 S2 S3 Boloria improba acrocnema Gall & Sperling, 1980 SW Colorado Rocky Mountains M T S L Boloria bellona (Fabricius, 1775) Meadow Fritillary iN Boloria bellona bellona (Fabricius, 1775) TL: not specified M S L1 L2 Boloria bellona toddi (W. Holland, 1928) TL: not specified M S1 S2 L Boloria bellona jenistai D. Stallings & Turner, 1947 boreal habitats from YK to Labrador M T S Boloria bellona S Rocky Mountain segregate , Wyoming, NW Colorado M S Boloria frigga (Thunberg, 1791) Frigga Fritillary iN Boloria frigga saga (Staudinger, 1861) TL: Canada M S1 S2 L Ha Boloria frigga gibsoni (W. Barnes & Benjamin, 1926) arctic; AK to Baffin Island M S L I Ha Boloria frigga sagata (W. Barnes & Benjamin, 1923) TL: USA (Co) M S1 S2 L Ha Boloria alberta (W. H. Edwards, 1890) Alberta Fritillary iN M S Ha Boloria polaris (Boisduval, [1828]) Polaris Fritillary iN Boloria polaris polaris (Boisduval, [1828]) TL: Norway M S L Boloria polaris stellata Masters, 1972 vic. Churchill, Manitoba M S Boloria astarte (E. Doubleday, [1847]) Astarte Fritillary iN Boloria astarte tschukotkensis (Wyatt, 1961) Seward Peninsula, AK Boloria astarte alaskia Grishin, 2024 Seward Peninsula, AK; ref.154 M T S Boloria astarte distincta (A. Gibson, 1920) most of AK, YK, NW BC, W NWT M S Ha Boloria astarte astarte (E. Doubleday, 1847) far N WA, MT, Rocky Mountains of BC-Alberta M S Argynnis Fabricius, 1807 Argynnis diana (Cramer, 1777) Diana Fritillary iN Argynnis diana arkansana Pavulaan, 2021 TL: USA (Ar) Argynnis diana diana (Cramer, 1777) TL: USA (Va) M S Ab L1 L2 I F Ha Argynnis cybele (Fabricius, 1775) Great Spangled Fritillary ref.16 iN Argynnis cybele cybele (Fabricius, 1775) TL: USA (NY) M S L1 L2 L3 L4 LAb Ha Argynnis cybele novascotiae McDunnough, 1935 TL: not specified M S Argynnis cybele krautwurmi W. Holland, 1931 TL: USA (Mi) M S L Argynnis cybele pseudocarpenteri F. Chermock & R. Chermock, 1940 TL: not specified M T S Argynnis cybele carpenterii W. H. Edwards, 1876 TL: USA (NM) M T Argynnis cybele neomexicana Grishin, 2022 [former carpenterii] SW CO & NC NM; ref.16 M S Argynnis cybele charlottii W. Barnes, 1897 TL: USA (Co) M S L I Ha Argynnis leto Behr, 1862 Great Spangled Fritillary ref.16 Argynnis leto letona (dos Passos & Grey, 1945) TL: USA (Ut) M S I Ha Argynnis leto leto Behr, 1862 TL: USA (Ca) M T S L1 L2 Ha Argynnis leto pugetensis (F. Chermock & Frechin, 1947) TL: USA (Wa) M T1 T2 S L1 L2 Ha Argynnis leto eileenae (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S I Argynnis aphrodite (Fabricius, 1787) Aphrodite Fritillary iN Argynnis aphrodite cullasaja (Gatrelle, 1998) TL: USA (NC) M T Argynnis aphrodite aphrodite (Fabricius, 1787) TL: USA (NY) M S L Argynnis aphrodite alcestis W. H. Edwards, [1877] TL: USA (Il) M T S Ab L Argynnis aphrodite ethne Hemming, 1933 TL: not specified M T S1 S2 Ab L1 L2 L3 L4 Ha Argynnis aphrodite W Colorado segregate (, ) W Colorado M S Argynnis aphrodite byblis (W. Barnes & Benjamin, 1926) TL: USA (Az) M S L Argynnis aphrodite columbia H. Edwards, 1877 TL: USA (Ak) M T S Argynnis aphrodite whitehousei Gunder, 1932 TL: not specified M T S Argynnis aphrodite manitoba F. Chermock & R. Chermock, 1940 TL: not specified M T1 T2 Argynnis aphrodite winni Gunder, 1932 TL: not specified M S Argynnis idalia (Drury, 1773) Regal Fritillary iN Argynnis idalia idalia (Drury, 1773) TL: USA (NY) M T S L I Argynnis idalia occidentalis (B. Williams, 2002) TL: USA (Il) M S L1 L2 L3 L4 Ha Argynnis nokomis W. H. Edwards, 1862 Nokomis Fritillary iN Argynnis nokomis coerulescens W. Holland, 1900 NW Mexico, extirpated from SE AZ M T S L I Argynnis nokomis nitocris W. H. Edwards, 1874 TL: USA (Az) M T S L1 L2 H Argynnis nokomis nokomis W. H. Edwards, 1862 TL: USA (Co) M T S Argynnis nokomis tularosa R. Holland, 2010 TL: USA (NM) M T OD Argynnis nokomis apacheana Skinner, 1918 TL: USA (Ca) M T S1 S2 L1 L2 I1 I2 Ha Argynnis nokomis carsonensis (Austin, 1998) TL: USA (Nv) M S L Ha Argynnis nokomis wenona (dos Passos & Grey, 1945) NE Mexico M T Argynnis nokomis melaena (Mooser & García, 1980) C Mexico (Aguascalientes) M S Argynnis edwardsii Reakirt, 1866 Edwards' Fritillary iN M T S1 S2 Ab L1 L2 Ha Argynnis coronis Behr, 1864 Coronis Fritillary iN Argynnis coronis hennei Gunder, 1934 TL: USA (Ca) M T S Argynnis coronis coronis Behr, 1864 TL: not specified M T S L1 L2 Argynnis coronis simaetha (dos Passos & Grey, 1945) TL: USA (Wa) M T S L Ha Argynnis coronis snyderi Skinner, 1897 TL: USA (Ut) M T S1 S2 L Argynnis coronis halcyone W. H. Edwards, [1869] TL: USA (Co) M T S1 S2 S3 L1 L2 Ha Argynnis coronis carolae dos Passos & Grey, 1942 USA (Nv); ref.11, iN.1267545 M T S L Ha Argynnis coronis semiramis W. H. Edwards, 1886 TL: USA (Ca) M T S L Argynnis zerene Boisduval, 1852 Zerene Fritillary iN Argynnis zerene sitka (Hammond, Harry & McCorkle, 2001) TL: USA (Ak) M T Argynnis zerene bremnerii W. H. Edwards, 1872 TL: USA (Wa) M T S Argynnis zerene Willamette Valley, W OR segregate (, ) apparently extinct M S Argynnis zerene hippolyta W. H. Edwards, 1879 TL: USA (Or) M T S L1 L2 L3 I Ha Argynnis zerene gloriosa (Moeck, 1957) TL: USA (Or) M S Argynnis zerene S OR coastal segregate (, ) M S Argynnis zerene behrensii W. H. Edwards, 1869 TL: USA (Ca) M T S I Argynnis zerene sonomensis (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S Argynnis zerene puntareyes (J. Emmel & T. Emmel, 1998) TL: USA (Ca) M T S1 S2 L Argynnis zerene myrtleae (dos Passos & Grey, 1945) TL: USA (Ca) M T Argynnis zerene monticola Behr, 1863 TL: USA (Ca) M T S Argynnis zerene malcolmi J. A. Comstock, 1920 TL: USA (Ca) M T S L Argynnis zerene zerene Boisduval, 1852 TL: USA (Ca) M T S L Ha Argynnis zerene E-central OR Cascades segregate (, ) M L Ha Argynnis zerene gunderi J. A. Comstock, 1925 TL: USA (Ca) M T S1 S2 L1 L2 Ha Argynnis zerene picta McDunnough, 1924 TL: not specified M T S L Argynnis zerene platina Skinner, 1897 TL: USA (Ut) M S L Argynnis zerene sinope (dos Passos & Grey, 1945) TL: USA (Co) M S1 S2 S3 L1 L2 Ha Argynnis zerene atossa W. H. Edwards, 1890 USA (Ca); ref.11 M T S1 S2 S3 Argynnis callippe Boisduval, 1852 Callippe Fritillary iN Argynnis callippe adiasteoides (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T1 T2 Argynnis callippe liliana H. Edwards, 1877 TL: USA (Ca) M S L Argynnis callippe callippe Boisduval, 1852 TL: USA (Ca) M T S Argynnis callippe comstocki Gunder, 1925 TL: USA (Ca) M T S L1 L2 H Argynnis callippe macaria W. H. Edwards, 1877 TL: USA (Ca) M T S L Argynnis callippe laurina W. G. Wright, 1905 TL: USA (Ca) M T S Ab L Argynnis callippe rupestris Behr, 1863 TL: USA (Ca) M T S Argynnis callippe inornata W. H. Edwards, 1872 TL: USA (Ca) M S Argynnis callippe juba Boisduval, 1869 TL: USA (Ca) M T S Argynnis callippe hagemani (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T1 T2 S Argynnis callippe shasta (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S L Argynnis callippe elaine (dos Passos & Grey, 1945) TL: USA (Or) M T S L1 L2 Ha Argynnis callippe N Siskiyous & W Cascades, low elevations segregate (, ) N Siskiyous, W Cascades, Oregon, below 3500' M L Argynnis callippe W OR Cascades, high elevations segregate (, ) W OR Cascades, above 3500' M S Argynnis callippe semivirida McDunnough, 1924 TL: not specified M S1 S2 L1 L2 Ha Argynnis callippe chilcotinensis (C. Guppy & J. Shepard, 2001) TL: not specified M S Argynnis callippe nevadensis W. H. Edwards, 1870 TL: USA (Nv) M S L Argynnis callippe harmonia (dos Passos & Grey, 1945) TL: USA (Nv) M S L Argynnis callippe calgariana McDunnough, 1924 TL: not specified M S Argynnis callippe gallatini McDunnough, 1929 TL: not specified M S1 S2 Ab L1 L2 Ha Argynnis callippe meadii W. H. Edwards, 1872 TL: USA (Co) M T S1 S2 L1 L2 Ha Argynnis egleis Behr, 1862 Great Basin Fritillary iN Argynnis egleis tehachapina J. A. Comstock, 1920 TL: USA (Ca) M T S L Argynnis egleis egleis Behr, 1862 TL: USA (Ca) M T S L1 L2 L3 Argynnis egleis Warner Mts. segregate (, ) Warner Mts., Lake Co. OR / Modoc Co., CA M S L Argynnis egleis oweni W. H. Edwards, 1892 TL: USA (Ca) M T S L Argynnis egleis mattooni (J. Emmel & T. Emmel, 1998) TL: USA (Ca) M T S L Ha Argynnis egleis yolaboli (J. Emmel & T. Emmel, 1998) TL: USA (Ca) M T Argynnis egleis moecki (Hammond & Dornfeld, 1983) TL: USA (Or) M S L Ha Argynnis egleis toiyabe (Howe, 1975) TL: USA (Nv) M T S Argynnis egleis reidi (Austin, 1998) TL: USA (Nv) M T1 T2 L Ha Argynnis egleis linda dos Passos & Grey, 1942 TL: USA (Id) M S Argynnis egleis Stansbury Mts. segregate (, ) Stansbury Mts., Tooele Co., Utah M S Argynnis egleis utahensis Skinner, 1919 TL: USA (Ut) M S Argynnis egleis secreta (dos Passos & Grey, 1945) TL: USA (Co) M S1 S2 L Ha Argynnis egleis macdunnoughi Gunder, 1932 TL: not specified M S Argynnis egleis morrellensis Kohler, 2020 TL: USA (Mt) Argynnis egleis kutoyisiks Kohler, 2020 TL: USA (Mt) Argynnis egleis albrighti Gunder, 1932 TL: not specified M S Argynnis adiaste W. H. Edwards, 1864 Unsilvered Fritillary iN Argynnis adiaste adiaste W. H. Edwards, 1864 TL: USA (Ca) M T S L Argynnis adiaste clemencei J. A. Comstock, 1925 TL: USA (Ca) M T S L Argynnis atlantis W. H. Edwards, 1862 Atlantis Fritillary iN Argynnis atlantis atlantis W. H. Edwards, 1862 TL: USA (NY) M T S L Argynnis atlantis canadensis dos Passos, 1935 TL: not specified M T S Argynnis atlantis brittanae Pavulaan, 2022 TL: USA (WV) Argynnis atlantis hollandi F. Chermock & R. Chermock, 1940 TL: not specified M T1 T2 S Argynnis atlantis pahasapa (Spomer, Scott & Kondla, 1998) TL: USA (SD) M T Argynnis atlantis sorocko (Scott, Kondla & Spomer, 1998) TL: USA (Co) M T S L Argynnis irene Boisduval, 1869 Northwestern Fritillary ref.11 Argynnis irene irene Boisduval, 1869 TL: USA (Ca) M T S L Argynnis irene hanseni (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S Argynnis irene cottlei J. A. Comstock, 1925 TL: USA (Ca) M S Argynnis irene dodgei Gunder, 1931 TL: USA (Or) M T S L1 L2 Ha Argynnis nausicaa W. H. Edwards, 1874 Northwestern Fritillary ref.11 Argynnis nausicaa elko (Austin, 1984) TL: USA (Nv) M T S L Ha Argynnis nausicaa greyi (Moeck, 1950) TL: USA (Nv) M T S Ha Argynnis nausicaa viola (dos Passos & Grey, 1945) TL: USA (Id) M T S Argynnis nausicaa tetonia (dos Passos & Grey, 1945) TL: USA (Wy) M T S1 S2 L Argynnis nausicaa chitone W. H. Edwards, 1879 TL: USA (Ut) M T S L Argynnis nausicaa schellbachi (Garth, 1949) TL: USA (Az) M T S L Argynnis nausicaa electa W. H. Edwards, 1878 TL: USA (Co) M T S L Ha Argynnis nausicaa dorothea (Moeck, 1947) TL: USA (NM) M T S L Argynnis nausicaa capitanensis (R. Holland, 1988) TL: USA (NM) M T S Argynnis nausicaa nausicaa W. H. Edwards, 1874 TL: USA (Az) M T S L1 L2 L3 L4 L5 L6 Argynnis hesperis W. H. Edwards, 1864 Northwestern Fritillary iN Argynnis hesperis hutchinsi (dos Passos & Grey, 1947) TL: not specified M S L Argynnis hesperis brico (Kondla, Scott & Spomer, 1998) TL: not specified M T S Argynnis hesperis beani (W. Barnes & Benjamin, 1926) TL: not specified M T S Argynnis hesperis dennisi (dos Passos & Grey, 1945) TL: not specified M T S1 S2 Argynnis hesperis lurana (dos Passos & Grey, 1945) TL: USA (SD) M T S1 S2 L Ha Argynnis hesperis hesperis W. H. Edwards, 1864 TL: USA (Co) M T S1 S2 S3 S4 Ab L Ha Argynnis hesperis ratonensis (Scott, 1981) TL: USA (NM) M T S Argynnis hydaspe Boisduval, 1869 Hydaspe Fritillary iN Argynnis hydaspe viridicornis J. A. Comstock, 1925 TL: not specified M T S L1 L2 Argynnis hydaspe hydaspe Boisduval, 1869 TL: USA (Ca) M T S L Argynnis hydaspe tehama (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S I Argynnis hydaspe davisi (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S Argynnis hydaspe rhodope W. H. Edwards, 1874 TL: Canada (BC) M T S1 S2 L1 L2 Ha Argynnis hydaspe NW OR & SW WA Coast Range segregate (, ) M S L Ha Argynnis hydaspe minor (dos Passos & Grey, 1947) TL: not specified M T S Argynnis mormonia Boisduval, 1869 Mormon Fritillary ref.16 iN Argynnis mormonia mormonia Boisduval, 1869 TL: USA (Nv) M T S L Ha Argynnis mormonia obsidiana (J. Emmel, T. Emmel & Mattoon, 1998) TL: USA (Ca) M T S L Ha Argynnis bischoffii W. H. Edwards, 1870 Mormon Fritillary ref.16 Argynnis bischoffii bischoffii W. H. Edwards, 1870 TL: USA (Ak) M S L Argynnis bischoffii opis W. H. Edwards, 1874 TL: not specified M S Argynnis bischoffii washingtonia W. Barnes & McDunnough, 1913 TL: not specified M S L Argynnis bischoffii erinna W. H. Edwards, 1883 TL: USA (Wa) M T S L Ha Argynnis bischoffii kimimela Marrone, Spomer & Scott, 2008 Black Hills of SD & WY M S L Ha Argynnis bischoffii eurynome W. H. Edwards, 1872 TL: USA (Co) M T S Ab L1 L2 L3 Ha Argynnis bischoffii Warner Mts. segregate (, ) M S Argynnis bischoffii artonis W. H. Edwards, 1881 TL: USA (Nv) M T S L Ha Argynnis bischoffii luski W. Barnes & McDunnough, 1913 TL: USA (Az) M S L1 L2 H Subfamily Limenitidinae Tribe Limenitidini Limenitis Fabricius, 1807 Limenitis arthemis (Drury, 1773) White Admiral iN Limenitis arthemis rubrofasciata (W. Barnes & McDunnough, 1916) TL: not specified M S1 S2 S3 Limenitis arthemis arthemis (Drury, 1773) TL: USA (NY) M S1 S2 L1 L2 Limenitis arthemis astyanax (Fabricius, 1775) TL: not specified M S1 S2 S3 S4 S5 Hy HS L1 L2 L3 L4 L5 I H F Limenitis arthemis arizonensis W. H. Edwards, 1882 TL: USA (Az) M T S1 S2 L I H Limenitis weidemeyerii W. H. Edwards, 1861 Weidemeyer’s Admiral iN Limenitis weidemeyerii weidemeyerii W. H. Edwards, 1861 TL: USA (Co) M S1 S2 S3 L Ha Limenitis weidemeyerii oberfoelli F. Brown, 1960 TL: USA (ND) M S1 S2 L Ha Limenitis weidemeyerii latifascia S. Perkins & E. Perkins, 1967 TL: USA (Id) M S1 S2 Hy L Limenitis weidemeyerii nevadae (W. Barnes & Benjamin, 1924) TL: USA (Nv) M S1 S2 Limenitis weidemeyerii angustifascia (W. Barnes & McDunnough, 1912) TL: USA (Az) M S1 S2 L1 L2 I Ha Limenitis weidemeyerii siennafascia Austin & Mullins, 1984 TL: USA (Az) M T S L Limenitis weidemeyerii Coahuila segregate , N Coahuila, Mexico M S L Ha Limenitis lorquini Boisduval, 1852 Lorquin's Admiral iN Limenitis lorquini powelli Austin & J. Emmel, 1998 TL: not specified M T S1 S2 L I Limenitis lorquini pallidafacies Austin & J. Emmel, 1998 TL: USA (Nv) M T S L Ha Limenitis lorquini lorquini Boisduval, 1852 TL: USA (Ca) M T S1 S2 S3 Hy L1 L2 L3 Ha Limenitis lorquini ilgae C. Guppy, 2001 TL: not specified M S L Ha Limenitis lorquini burrisonii Maynard, 1891 TL: not specified M S Hy L Ha Limenitis archippus (Cramer, 1775) Viceroy iN Limenitis archippus floridensis Strecker, 1878 vagrant in Cuba M T S1 S2 L H Limenitis archippus watsoni (dos Passos, 1938) TL: USA (La) M S1 S2 L Limenitis archippus archippus (Cramer, 1775) TL: USA (NY) M S1 S2 S3 S4 S5 Hy L1 L2 I Limenitis archippus idaho Austin, 1998 TL: USA (Nv) M T S1 S2 Ha Limenitis archippus lahontani Herlan, 1971 TL: USA (Nv) M T S1 S2 Limenitis archippus obsoleta W. H. Edwards, 1882 NW Mexico M T S1 S2 L I Limenitis archippus hoffmanni R. Chermock, 1947 E Mexico M T S1 S2 L H Limenitis archippus Baja California Sur segregate , Baja California Sur - only 1 specimen known M S Adelpha Hübner, [1819] Adelpha alala (Hewitson, 1847) iN Adelpha alala alala (Hewitson, 1847) TL: Venezuela M T Adelpha alala completa Fruhstorfer, 1907 TL: Colombia M T Adelpha alala negra (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T S L Adelpha aricia (Hewitson, 1847) iN Adelpha aricia aricia (Hewitson, 1847) TL: Bolivia M T Adelpha aricia portunus A. Hall, 1938 TL: Peru M T Adelpha aricia serenita Fruhstorfer, 1915 TL: Peru M T Adelpha aricia [n. ssp.] Lamas, MS Peru Adelpha corcyra (Hewitson, 1847) iN Adelpha corcyra corcyra (Hewitson, 1847) TL: Colombia M T Adelpha corcyra aretina Fruhstorfer, 1907 TL: Ecuador M T Adelpha corcyra collina (Hewitson, 1847) TL: Ecuador M T Adelpha corcyra dognini Willmott, 2003 TL: Ecuador M T S Adelpha corcyra salazari Willmott, 2003 TL: Colombia M T Adelpha pithys (H. Bates, 1864) Pithy Sister SW & S Mexico to El Salvador, [Honduras?] iN M T S L Adelpha donysa (Hewitson, 1847) Donysa Sister iN Adelpha donysa SW Mexican segregate , SW Mexico (MICH to OAX) M S Adelpha donysa donysa (Hewitson, 1847) SE Mexico to Guatemala M T S Adelpha donysa albifilum Steinhauser, 1974 El Salvador, Honduras M T Adelpha tracta (A. Butler, 1872) Tracta Sister Nicaragua to W Panama iN M T S1 S2 L I Adelpha bredowii Geyer, 1837 Mexican Sister SW & S-central Mexico, local in NW Mexico; cloud forest iN M T S L Adelpha californica (A. Butler, 1865) California Sister Baja California (N) to S WA, Cascade-Sierra Nevada, westward iN M T S L1 L2 I Ha Adelpha eulalia (E. Doubleday, [1848]) Arizona Sister far SE CA, S NV, UT, AZ, NM, S CO, TX (strays N), Mx to Guat. iN M T S1 S2 L1 L2 L3 I1 I2 I3 I4 H Adelpha diocles Godman & Salvin, 1878 Short-tailed Sister iN Adelpha diocles diocles Godman & Salvin, 1878 E Costa Rica & W Panama (Chiriqui massif) M T S Adelpha diocles creton Godman, 1901 E-central & S Mexico to Guatemala & El Salvador M T S L Adelpha herbita Weymer, 1907 TL: Brazil (SC) Adelpha herbita herbita Weymer, 1907 TL: Brazil (SC) Adelpha herbita perdita Willmott & Freitas, 2016 ref.313 Adelpha zea (Hewitson, 1850) iN M T Adelpha nea (Hewitson, 1847) Nea Sister iN Adelpha nea nea (Hewitson, 1847) Costa Rica to Venezuela, to S Peru, Guianas, Amazonian Brazil M T S Adelpha nea sentia Godman & Salvin, 1884 SE Mexico (OAX, VER), Belize M T S Adelpha paroeca (H. Bates, 1864) Paroeca Sister iN Adelpha paroeca paroeca (H. Bates, 1864) [= emathia], E & W Mexico to W Panama M T S L Adelpha paroeca pseudodonysa Salazar, 2000 TL: Colombia M T Adelpha paraena (H. Bates, 1865) iN Adelpha paraena paraena (H. Bates, 1865) TL: Brazil (Pa) M T Adelpha paraena lecromi Willmott, 2003 C Panama to N Colombia M S Adelpha paraena massilia (C. Felder & R. Felder, 1867) E & W Mexico to C Panama M T S L I Ha Adelpha paraena reyi Neild, 1996 TL: Venezuela M T Adelpha radiata Fruhstorfer, 1915 iN Adelpha radiata radiata Fruhstorfer, 1915 TL: Brazil (SC) M T Adelpha radiata aiellae Willmott & J. Hall, 1999 NW Costa Rica to NW Ecuador M T S I Adelpha radiata explicator Willmott & J. Hall, 1999 TL: Ecuador Adelpha radiata gilletella Brévignon, 1995 TL: French Guiana Adelpha radiata myrlea Fruhstorfer, 1915 TL: Brazil (ES) M T Adelpha radiata romeroi Willmott & Neild, 2003 TL: Venezuela Adelpha serpa (Boisduval, 1836) iN Adelpha serpa serpa (Boisduval, 1836) TL: Brazil M T Adelpha serpa celerio (H. Bates, 1864) E & W Mexico to W Colombia, NW Venezuela, M T S1 S2 L I1 I2 I3 Adelpha serpa diadochus Fruhstorfer, 1915 TL: Peru M T S Adelpha serpa duiliae Fruhstorfer, 1913 TL: Ecuador M T Adelpha seriphia (C. Felder & R. Felder, 1867) iN Adelpha seriphia seriphia (C. Felder & R. Felder, 1867) TL: Venezuela M T Adelpha seriphia aquillia Fruhstorfer, 1915 TL: Colombia M T Adelpha seriphia naryce Fruhstorfer, 1915 TL: Peru M T Adelpha seriphia barcanti Willmott, 2003 TL: Trinidad M T Adelpha seriphia egregia Röber, 1927 TL: Colombia M T Adelpha seriphia pione Godman & Salvin, 1884 TL: Panama M T Adelpha seriphia therasia Fruhstorfer, 1915 TL: Bolivia M T Adelpha hyas (Doyère, [1840]) iN Adelpha hyas hyas (Doyère, [1840]) TL: ? M S Adelpha hyas hewitsoni Willmott & J. Hall, 1999 TL: Ecuador Adelpha hyas viracocha A. Hall, 1938 TL: Peru M T Adelpha godmani Fruhstorfer, 1913 Godman's Sister Chiapas to W Ecuador iN M T S L Adelpha margarita Willmott & J. Hall, 2013 iN Adelpha margarita margarita Willmott & J. Hall, 2013 ref.321 Adelpha margarita garleppi Willmott, 2013 ref.321 Adelpha abia (Hewitson, 1850) iN M T Adelpha abyla (Hewitson, 1850) Jamaican Sister Jamaica iN M T S Adelpha adstricta Fruhstorfer, 1915 iN Adelpha amazona Austin & Jasinski, 1999 iN Adelpha argentea Willmott & J. Hall, 1995 iN M T Adelpha atlantica Willmott, 2003 TL: Brazil (RJ) M T Adelpha attica (C. Felder & R. Felder, 1867) iN Adelpha attica attica (C. Felder & R. Felder, 1867) TL: Colombia M T S Adelpha attica hemileuca Willmott & J. Hall, 1999 NW Ecuador, W Colombia, 1 Panama specimen nr. nominate form M T Adelpha barnesia Schaus, 1902 iN Adelpha barnesia barnesia Schaus, 1902 TL: Bolivia M T Adelpha barnesia leucas Fruhstorfer, 1915 E & W Mexico to NW Venezuela & W Ecuador M T S I Adelpha barnesia trinita Kaye, 1914 TL: Trinidad M T Adelpha basiloides (H. Bates, 1865) Spot-celled Sister S TX, E & W Mexico to N Venezuela & W Ecuador iN M T S1 S2 L I Adelpha boeotia (C. Felder & R. Felder, 1867) Oberthur's Sister iN Adelpha boeotia boeotia (C. Felder & R. Felder, 1867) E Panama to Bolivia, SW Brazil, Guianas M T S L Adelpha boeotia fidicula Fruhstorfer, 1915 TL: Brazil (ES) M T Adelpha boeotia oberthurii (Boisduval, 1870) [= jacquelinae], SE Mexico to W Panama M T L Adelpha boreas (A. Butler, 1866) Boreas Sister iN Adelpha boreas boreas (A. Butler, 1866) E Costa Rica to Venezuela & Bolivia M T S L Adelpha boreas kayei A. Hall, 1939 TL: Guyana M T Adelpha calliphane Fruhstorfer, 1915 iN M T Adelpha capucinus (Walch, 1775) iN Adelpha capucinus capucinus (Walch, 1775) TL: East Indies M T S Adelpha capucinus gutierrezi Neild, 1996 TL: Venezuela M T Adelpha capucinus velia (C. Felder & R. Felder, 1867) TL: Colombia M T Adelpha cocala (Cramer, 1779) iN Adelpha cocala cocala (Cramer, 1779) TL: Surinam M T S Adelpha cocala caninia Fruhstorfer, 1915 TL: Brazil (SC/SP) M T Adelpha cocala didia Fruhstorfer, 1915 TL: Brazil (ES) M T Adelpha cocala lorzae (Boisduval, 1870) SE Mexico (OAX, CHIS) to NW Venezuela & W Ecuador M T S1 S2 I Adelpha cocala orellanae Neild, 1996 TL: Venezuela M T Adelpha coryneta (Hewitson, 1874) iN M T Adelpha cytherea (Linnaeus, 1758) iN Adelpha cytherea cytherea (Linnaeus, 1758) TL: Indiis M T S Adelpha cytherea aea (C. Felder & R. Felder, 1867) TL: Brazil? M T Adelpha cytherea daguana Fruhstorfer, 1913 E Costa Rica (as intergrades with marcia) to NW Venez. & W Ecuad. M T S L Adelpha cytherea insularis Fruhstorfer, 1913 TL: Trinidad M T Adelpha cytherea marcia Fruhstorfer, 1913 S Mexico through E Costa Rica M T S1 S2 L I Adelpha cytherea nahua Grose-Smith, 1898 TL: Venezuela M T Adelpha cytherea olbia (C. Felder & R. Felder, 1867) TL: Colombia M T Adelpha delinita Fruhstorfer, 1913 iN Adelpha delinita delinita Fruhstorfer, 1913 TL: Colombia M T S L Adelpha delinita utina A. Hall, 1938 [= escalantei], SE Mexico to W Panama M T S Adelpha demialba (A. Butler, 1872) White-spotted Sister Costa Rica to E Panama; cloud forest iN M T S L I1 I2 Adelpha diazi Beutelspacher, 1975 Diaz's Sister E Mexico iN M T S Adelpha epione (Godart, [1824]) iN Adelpha epione epione (Godart, [1824]) TL: Brazil Adelpha epione agilla Fruhstorfer, 1907 TL: Colombia M T S Adelpha epizygis Fruhstorfer, 1915 iN Adelpha epizygis epizygis Fruhstorfer, 1915 TL: Brazil (SP) M T Adelpha epizygis epona Fruhstorfer, 1915 TL: [Brazil (Ba)] M T Adelpha erotia (Hewitson, 1847) Erotia Sister iN Adelpha erotia erotia (Hewitson, 1847) SE Mexico (Chiapas) to Bolivia, Amazonian Brazil & Guianas M T S L Adelpha erotia caphira (Hewitson, 1869) TL: Venezuela M T Adelpha erymanthis Godman & Salvin, 1884 Erymanthis Sister iN Adelpha erymanthis erymanthis Godman & Salvin, 1884 S Mexico (Chiapas) to C Panama M T S I Adelpha erymanthis esperanza Balcázar & Willmott, 2003 Sierra de Juárez, Oaxaca, Mexico M S HS Adelpha erymanthis fortunata Willmott, 2003 TL: Ecuador Adelpha ethelda (Hewitson, 1867) iN Adelpha ethelda Northern segregate , SE Mexico M S Adelpha ethelda ethelda (Hewitson, 1867) TL: Ecuador M T Adelpha ethelda eponina Staudinger, 1886 TL: Colombia M T Adelpha ethelda galbao Brévignon, 1995 TL: French Guiana Adelpha ethelda sophax Godman & Salvin, 1878 E Costa Rica to C Panama; cloud forest M T L Adelpha ethelda zalmona (Hewitson, 1871) TL: Colombia M T Adelpha ethelda [n. ssp.] Willmott, MS Mexico Adelpha fabricia Fruhstorfer, 1913 iN M T Adelpha falcipennis Fruhstorfer, 1915 iN M T S Adelpha felderi (Boisduval, 1870) Felder's Sister E Mexico to Honduras iN M T S L Adelpha fessonia (Hewitson, 1847) Band-celled Sister iN Adelpha fessonia fessonia (Hewitson, 1847) S TX, E & W Mexico to W Panama M T S1 S2 L1 L2 I Ha Adelpha fessonia cestus (Hewitson, 1847) TL: Venezuela M T Adelpha fessonia ernestoi Willmott, 2003 TL: Colombia M T Adelpha lapitha A. Hall, 1929 Hispaniolan Sister Hispaniola M S Adelpha gavina Fruhstorfer, 1915 iN M T Adelpha gelania (Godart, [1824]) Antillean Sister iN Adelpha gelania gelania (Godart, [1824]) Hispaniola M S Adelpha gelania arecosa (Hewitson, 1847) Puerto Rico M T S L Adelpha heraclea (C. Felder & R. Felder, 1867) Heraclea Sister iN Adelpha heraclea heraclea (C. Felder & R. Felder, 1867) Guatemala to SE Peru, Brazil M T L Adelpha heraclea antonii Neild, 1996 TL: Venezuela Adelpha heraclea makkeda (Hewitson, 1871) TL: Brazil (Pa) M T Adelpha hesterbergi Willmott & J. Hall, 1999 Hesterberg's Sister Costa Rica, NW Ecuador, [Panama?, W Colombia?] iN M T Adelpha iphicleola (H. Bates, 1864) Iphicleola Sister iN Adelpha iphicleola iphicleola (H. Bates, 1864) [= massilides], E & W Mexico to E Panama M T S1 S2 L I Adelpha iphicleola gortyna Fruhstorfer, 1915 Costa Rica to W Panama, possibly only as intergrades with above M T S L Adelpha iphicleola iphimedia Fruhstorfer, 1915 Cuba & Isla de Juventud M T S Adelpha iphicleola leucates Fruhstorfer, 1915 TL: Brazil (Ba) M T Adelpha iphicleola thessalita Willmott & J. Hall, 1999 TL: Ecuador Adelpha iphiclus (Linnaeus, 1758) Iphiclus Sister iN Adelpha iphiclus iphiclus (Linnaeus, 1758) W & S Mexico to Bolivia, Guianas, Trinidad M T S1 S2 HS L I Adelpha iphiclus ephesa (Ménétriés, 1857) TL: Brazil M T Adelpha iphiclus estrecha Willmott & J. Hall, 1999 TL: Ecuador M T S Adelpha irmina (E. Doubleday, [1848]) iN Adelpha irmina irmina (E. Doubleday, [1848]) TL: Venezuela M T Adelpha irmina tumida (A. Butler, 1873) TL: Peru; Colombia M T S L Adelpha irmina arcana Benmesbah & Costa, 2019 ref.192 Adelpha irmina [n. ssp.] Lamas, MS Peru Adelpha jordani Fruhstorfer, 1913 iN M T S Adelpha justina (C. Felder & R. Felder, 1861) iN Adelpha justina justina (C. Felder & R. Felder, 1861) TL: Colombia M T S Adelpha justina inesae Orellana, 1996 TL: Venezuela M T Adelpha justina justinella Fruhstorfer, 1907 TL: Venezuela M T Adelpha justina maira Orellana, 1996 TL: Venezuela M T Adelpha justina pichincha Willmott & J. Hall, 2013 TL: Ecuador Adelpha justina valentina Fruhstorfer, 1915 TL: Colombia; Bolivia M T S Adelpha lamasi Willmott & J. Hall, 1999 iN M T Adelpha leuceria (H. Druce, 1874) iN M T S L Adelpha juanna Grose-Smith, 1898 TL: Colombia M T Adelpha leucerioides Beutelspacher, 1975 Tawny-striped Sister E Mexico to Guatemala iN Adelpha leucerioides SW Mexican segregate , SW Mexico M S Adelpha leucerioides leucerioides Beutelspacher, 1975 TL: Mexico (Ver) M T S Adelpha leucophthalma (Latreille, [1809]) Leucophthalma Sister iN Adelpha leucophthalma leucophthalma (Latreille, [1809]) Nicaragua to Colombia M T S1 S2 L I Adelpha leucophthalma irminella Strand, 1918 TL: Ecuador M T Adelpha leucophthalma smalli Willmott, 2003 Cerro Montuoso, Azuero Peninsula, Herrera, Panama M S Adelpha levona Steinhauser & L. Miller, 1977 iN M T Adelpha melanthe (H. Bates, 1864) Rayed Sister E & S Mexico to C Panama iN M T S1 S2 L I1 I2 Adelpha lycorias (Godart, [1824]) iN Adelpha lycorias melanippe Godman & Salvin, 1884 TL: Colombia M T Adelpha lycorias spruceana (H. Bates, 1864) TL: Ecuador M T Adelpha lycorias wallisii (Dewitz, 1877) TL: Colombia M T Adelpha lycorias lara (Hewitson, 1850) TL: Venezuela M T S L Adelpha lycorias lycorias (Godart, [1824]) TL: Brazil M T Adelpha malea (C. Felder & R. Felder, 1861) iN Adelpha malea malea (C. Felder & R. Felder, 1861) TL: Venezuela M T Adelpha malea aethalia (C. Felder & R. Felder, 1867) TL: Colombia M T S Adelpha malea fundania Fruhstorfer, 1915 E Mexico to NW Venezuela M T S I Adelpha malea goyama Schaus, 1902 TL: Brazil (Pr) M T Adelpha malea ixia (C. Felder & R. Felder, 1867) TL: Venezuela M T Adelpha melona (Hewitson, 1847) iN Adelpha melona melona (Hewitson, 1847) TL: Colombia M T Adelpha melona deborah Weeks, 1901 TL: Colombia M T Adelpha melona leucocoma Fruhstorfer, 1915 TL: Upper Amazon M T S Adelpha melona neildi Willmott, 2003 C Panama M L Adelpha melona pseudarete Fruhstorfer, 1915 TL: ? M T Adelpha mesentina (Cramer, 1777) iN M T Adelpha messana (C. Felder & R. Felder, 1867) Messana Sister iN Adelpha messana messana (C. Felder & R. Felder, 1867) Guatemala to Colombia, NW Ecuador, NW Venezuela M T S L Adelpha messana delphicola Fruhstorfer, 1910 TL: [Peru] M T Adelpha messana seminivea Freitas, Willmott & Woodbury, 2019 ref.218 Adelpha milleri Beutelspacher, 1976 Miller’s Sister SE Mexico & Belize M T S Adelpha mythra (Godart, [1824]) iN M T Adelpha naxia (C. Felder & R. Felder, 1867) Naxia Sister iN Adelpha naxia naxia (C. Felder & R. Felder, 1867) W & S Mexico to Bolivia, Brazilian Amazon, Guianas M T S L I Ha Adelpha naxia hieronica Fruhstorfer, 1913 TL: Trinidad M T Adelpha naxia oteroi Neild, 1996 TL: Venezuela M T Adelpha olynthia (C. Felder & R. Felder, 1867) iN M T S Adelpha phylaca (H. Bates, 1866) Phylaca Sister iN Adelpha phylaca phylaca (H. Bates, 1866) E & W Mexico to W Honduras M T S1 S2 L Adelpha phylaca joffrei Neild, 1996 TL: Venezuela M T Adelpha phylaca pseudaethalia A. Hall, 1938 S Mexico (Chiapas), to NW Venezuela & W Ecuador M T S I1 I2 Adelpha plesaure Hübner, 1823 iN Adelpha plesaure plesaure Hübner, 1823 TL: Brazil M T Adelpha plesaure phliassa (Godart, [1824]) TL: French Guiana; Brazil M T S Adelpha plesaure pseudomalea A. Hall, 1938 TL: Venezuela M T Adelpha plesaure symona Kaye, 1925 TL: Trinidad M T Adelpha pollina Fruhstorfer, 1915 iN M T S Adelpha poltius A. Hall, 1938 iN M T Adelpha rothschildi Fruhstorfer, 1913 iN M T Adelpha salmoneus (A. Butler, 1866) iN Adelpha salmoneus salmoneus (A. Butler, 1866) TL: Venezuela M T Adelpha salmoneus colada (C. Felder & R. Felder, 1867) TL: Colombia M T Adelpha salmoneus emilia Fruhstorfer, 1908 TL: Colombia M T Adelpha salmoneus salmonides A. Hall, 1938 SE Mexico to C Panama M T S1 S2 L I Adelpha salus A. Hall, 1935 iN Adelpha salus Northern segregate , SE Mexico to W Panama M S Adelpha salus salus A. Hall, 1935 TL: Colombia M T L Adelpha salus emmeli Willmott & J. Hall, 1999 TL: Ecuador Adelpha salus [n. ssp.] Willmott, MS Mexico Adelpha saundersii (Hewitson, 1867) iN Adelpha saundersii saundersii (Hewitson, 1867) TL: Ecuador M T S L Adelpha saundersii frontina A. Hall, 1935 TL: Colombia M T Adelpha saundersii helepecki Weeks, 1901 TL: Bolivia M T S Adelpha shuara Willmott & J. Hall, 1995 iN M T S Adelpha sichaeus (A. Butler, 1866) iN M T S L Adelpha stilesiana DeVries & I. Chacón, 1982 Talamanca Sister Talamanca massif, Costa Rica & W Panama, also Vol. Chiriqui M T S Adelpha syma (Godart, [1824]) iN M T Adelpha thesprotia (C. Felder & R. Felder, 1867) iN M T Adelpha thessalia (C. Felder & R. Felder, 1867) iN Adelpha thessalia thessalia (C. Felder & R. Felder, 1867) TL: Brazil (Am) M T S Adelpha thessalia cesilas Fruhstorfer, 1915 TL: Bolivia M T Adelpha thessalia indefecta Fruhstorfer, 1913 TL: Paraguay M T Adelpha thoasa (Hewitson, 1850) iN Adelpha thoasa thoasa (Hewitson, 1850) TL: [Brazil (Pa)] M T Adelpha thoasa calliphiclea (A. Butler, [1870]) TL: Surinam M T Adelpha thoasa gerona (Hewitson, 1867) TL: Brazil (MG) M T Adelpha thoasa manilia Fruhstorfer, 1915 TL: Bolivia M T Adelpha viola Fruhstorfer, 1913 TL: Brazil (SP) Adelpha viola viola Fruhstorfer, 1913 TL: Brazil (SP) M T Adelpha viola pseudococala A. Hall, 1933 TL: Brazil (Pa); Guyana; Peru M T Adelpha ximena (C. Felder & R. Felder, 1862) iN Adelpha ximena ximena (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Adelpha ximena mossi A. Hall, 1933 TL: Brazil (Pa) M T Adelpha zina (Hewitson, 1867) Zina Sister iN Adelpha zina zina (Hewitson, 1867) E Panama to SW Ecuador & Venezuela M T S L Adelpha zina desousae Neild & Orellana, 1996 TL: Venezuela Adelpha zina enope A. Hall, 1938 TL: Colombia M T Adelpha zina inachia Staudinger, 1886 TL: Colombia M T Adelpha zina irma Fruhstorfer, 1907 TL: Peru M T S L Adelpha zina lacina (A. Butler, 1872) Nicaragua to W Panama M T OI S1 S2 L I Adelpha zina pyrczi Neild & Orellana, 1996 TL: Venezuela M T Subfamily Apaturinae Asterocampa Röber, 1916 Asterocampa celtis (Boisduval & Le Conte, [1835]) Hackberry Emperor iN Asterocampa celtis celtis (Boisduval & Le Conte, [1835]) E North America (except as others) M T S Ab L1 L2 L3 L4 Asterocampa celtis reinthali Friedlander, 1987 SE USA coastal plain (SC to FL, W to E TX) M T S1 S2 L Asterocampa celtis antonia (W. H. Edwards, [1878]) [= jeffermont] SW USA (SE CA to TX, N to SE CO), N & C Mexico M T S1 S2 L1 L2 I H Asterocampa leilia (W. H. Edwards, 1874) Empress leilia S & C AZ, SW NM, S & W TX, E & NW Mexico, Baja California iN M T S L1 L2 L3 I H Asterocampa clyton (Boisduval & Le Conte, [1835]) Tawny Emperor iN Asterocampa clyton clyton (Boisduval & Le Conte, [1835]) E North America (except as others) M T S1 S2 Ab L1 L2 L3 Asterocampa clyton flora (W. H. Edwards, 1876) SE USA coastal plain (SC to FL, W to E TX) M T S1 S2 L I Asterocampa clyton texana (Skinner, 1911) S AZ, S NM, W & C TX, NW Mexico (SON, CHIH) M T S L I Asterocampa clyton louisa D. Stallings & J. Turner, 1947 S TX, NE Mexico (to SLP) M T S L1 L2 I Asterocampa idyja (Geyer, [1828]) Dusky Emperor iN Asterocampa idyja idyja (Geyer, [1828]) Cuba & Isla de Juventud, Hispaniola, Puerto Rico M T S1 S2 Asterocampa idyja argus (H. Bates, 1864) SE AZ (as vagrant), E & W Mexico to Costa Rica M T S L1 L2 I Doxocopa Hübner, [1819] Doxocopa agathina (Cramer, 1777) iN Doxocopa agathina agathina (Cramer, 1777) TL: Surinam M T Doxocopa agathina vacuna (Godart, [1824]) TL: Brazil M T Doxocopa callianira (Ménétriés, 1855) Nicaraguan Emperor S Mexico to Costa Rica iN M T S L Doxocopa clothilda (C. Felder & R. Felder, 1867) Clothilda Emperor Costa Rica to Colombia iN M T S L Doxocopa cyane (Latreille, [1813]) iN Doxocopa cyane cyane (Latreille, [1813]) TL: [Peru] M T S L Doxocopa cyane burmeisteri (Godman & Salvin, 1884) TL: Argentina M T S Doxocopa cyane mexicana Bryk, 1953 E & S Mexico to Colombia M T S Doxocopa cyane vespertina Lamas, 1999 TL: Peru M T Doxocopa elis (C. Felder & R. Felder, 1861) iN M T S Doxocopa excelsa (Gillott, 1927) Broad-banded Emperor Nicaragua & Costa Rica M T S Doxocopa kallina (Staudinger, 1886) iN M T Doxocopa laure (Drury, 1773) Silver Emperor ref.133 iN Doxocopa laure laure (Drury, 1773) continental; MX to Panama M T S Doxocopa laure laura (Hübner, [1823]) Jamaica M T S Doxocopa druryi (Hübner, [1825]) Silver Emperor Cuba; ref.133 Doxocopa druryi druryi (Hübner, [1825]) Cuba M S Doxocopa druryi acca (C. Felder & R. Felder, 1867) continental; S TX, SE AZ (uncommon) to Panama M T S L1 L2 L3 I Doxocopa druryi mima (Fruhstorfer, 1907) TL: Colombia M T Doxocopa druryi griseldis (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Doxocopa druryi laurona (Schaus, 1902) TL: Brazil (RJ) M T Doxocopa laurentia (Godart, [1824]) iN Doxocopa laurentia laurentia (Godart, [1824]) TL: Brazil M T Ab Doxocopa laurentia cherubina (C. Felder & R. Felder, 1867) S Mexico to Colombia M T S L Doxocopa laurentia thalysia (Fruhstorfer, 1907) TL: Ecuador M T Doxocopa lavinia (A. Butler, 1866) iN Doxocopa lavinia ducaudi Brévignon, 2008 TL: French Guiana Doxocopa lavinia lavinia (A. Butler, 1866) iN M T Doxocopa linda (C. Felder & R. Felder, 1862) iN Doxocopa linda linda (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Doxocopa linda carwa Lamas, 1999 TL: Peru M T Doxocopa linda godmani (Dannatt, 1904) TL: Venezuela Doxocopa linda mileta (Boisduval, 1870) TL: Brazil M T Doxocopa linda plesaurina (A. Butler & H. Druce, 1872) Costa Rica & Panama M T S Doxocopa pavon (Latreille, [1809]) iN Doxocopa pavon pavon (Latreille, [1809]) TL: Ecuador M T Doxocopa pavon theodora (Lucas, 1857) S TX, E & W Mexico to Colombia M T OI S L1 L2 Doxocopa thoe (Godart, [1824]) Hispaniolan Emperor Hispaniola iN M T S Doxocopa zalmunna (A. Butler, 1869) TL: Brazil M T Doxocopa zunilda (Godart, [1824]) iN Doxocopa zunilda zunilda (Godart, [1824]) TL: Brazil M T Doxocopa zunilda felderi (Godman & Salvin, 1884) Costa Rica to Colombia M T S L Doxocopa zunilda floris (Fruhstorfer, 1907) TL: Peru M T Doxocopa wilmattae (T. Cockerell, 1907) Fossil from Colorado Subfamily Biblidinae Tribe Biblidini Biblis Fabricius, 1807 Biblis aganisa Boisduval, 1836 Red Rim SW USA & S TX (strays N), E & W Mexico at least to Costa Rica M T S1 S2 L I Biblis hyperia (Cramer, 1779) Red Rim iN Biblis hyperia hyperia (Cramer, 1779) Hispaniola, Mona, Puerto Rico, Culebra, Virgin Is. to St. Lucia M T S L Biblis hyperia laticlavia (Thieme, 1904) TL: Ecuador M T Biblis hyperia rosanti Brévignon, 2006 TL: French Guiana Biblis hyperia nectanabis (Fruhstorfer, 1909) TL: Brazil (RS) M T Biblis hyperia pacifica (A. Hall, 1928) TL: Ecuador M T Biblis hyperia [n. ssp.] Lamas, MS Peru Vila W. F. Kirby, 1871 Vila azeca (E. Doubleday, [1848]) iN Vila azeca azeca (E. Doubleday, [1848]) TL: Bolivia M T Vila azeca cacica Staudinger, 1886 TL: Ecuador M T Vila azeca mariana (H. Bates, 1865) TL: Brazil (Am) M T Vila azeca semistalachtis A. Hall, 1928 TL: Colombia M T Vila azeca stalachtoides (H. Bates, 1867) TL: Brazil (Ma/Pa) M T Vila emilia (Cramer, 1779) iN Vila emilia emilia (Cramer, 1779) TL: Surinam Vila emilia caecilia (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T S Vila emilia carmenae Le Crom, 1990 TL: Colombia M T Vila emilia matteii Neild, 1996 TL: Venezuela M T Vila emilia sinefascia A. Hall, 1935 TL: [Peru; Brazil (Am)] M T Vila eueidiformis Joicey & Talbot, 1918 iN M T Mestra Hübner, [1825] Mestra amymone (Ménétriés, 1857) Northern Mestra S TX (strays far N), E & W Mexico to Costa Rica iN M S Ma L1 L2 I1 I2 I3 H Mestra dorcas (Fabricius, 1775) Jamaican Mestra Jamaica iN M T S1 S2 L Mestra hersilia (Fabricius, 1776) South American Mestra iN Mestra hersilia hersilia (Fabricius, 1776) N S America, Trinidad, Tobago, Barbados, Grenada to Dominica M T S L Mestra hersilia latimargo (A. Hall, 1929) TL: Ecuador M T Mestra hersilia semifulva (C. Felder & R. Felder, 1867) TL: Colombia M T Mestra hersilia apicalis (Staudinger, 1886) TL: Brazil (SP) M T Mestra hersilia hypermestra Hübner, [1825] TL: ? M T Archimestra Munroe, 1949 Archimestra teleboas (Ménétriés, 1832) Archimestra Hispaniola iN M S L Tribe Epicaliini Catonephele Hübner, [1819] ref.198 Catonephele acontius (Linnaeus, 1771) iN Catonephele acontius acontius (Linnaeus, 1771) TL: China M T S Catonephele acontius caeruleus Jenkins, 1985 TL: Bolivia Catonephele antinoe (Godart, [1824]) iN M T Catonephele chromis E. Doubleday, [1848] iN Catonephele chromis chromis (E. Doubleday, [1848]) TL: Honduras M T S L Catonephele chromis godmani Stichel, 1901 Honduras to Panama M T S Catonephele cortesi R. G. Maza, 1982 West-Mexican Banner W Mexico iN M T S L Ha Catonephele mexicana Jenkins & R. G. Maza, 1985 East-Mexican Banner E Mexico to Costa Rica iN M S1 S2 L I Catonephele numilia (Cramer, 1775) iN Catonephele numilia numilia (Cramer, 1775) TL: Surinam Catonephele numilia esite (R. Felder, 1869) E Mexico to W Ecuador, Venezuela & Trinidad M T S1 S2 L1 L2 I1 I2 Catonephele numilia immaculata Jenkins, 1985 SW Mexico M T S L Ha Catonephele numilia neogermanica Stichel, 1899 TL: Paraguay M T Catonephele numilia penthia (Hewitson, 1852) TL: Brazil (RJ) M T S Catonephele nyctimus (Westwood, 1850) iN M T Catonephele orites Stichel, 1899 Broken Banner Costa Rica to NW Ecuador iN M T S L Catonephele sabrina (Hewitson, 1852) iN M T Catonephele salacia (Hewitson, 1852) iN M T Catonephele salambria (C. Felder & R. Felder, 1861) iN M T Nessaea Hübner, [1819] ref.198 Nessaea aglaura (E. Doubleday, [1848]) Northern Nessaea iN Nessaea aglaura aglaura (E. Doubleday, [1848]) S Mexico to W Ecuador M T S L Nessaea aglaura regina (Salvin, 1869) TL: Venezuela M T Nessaea aglaura thalia Bargmann, 1928 TL: [Colombia] M T S Nessaea batesii (C. Felder & R. Felder, 1860) iN Nessaea batesii batesii (C. Felder & R. Felder, 1860) TL: [Brazil (Pa)] M T Nessaea batesii magniplaga Röber, 1928 TL: Bolivia Nessaea hewitsonii (C. Felder & R. Felder, 1859) iN Nessaea hewitsonii hewitsonii (C. Felder & R. Felder, 1859) TL: [Brazil (Am)] M T S Nessaea hewitsonii boliviensis Jenkins, 1989 TL: Bolivia M T S Nessaea obrinus (Linnaeus, 1758) iN Nessaea obrinus obrinus (Linnaeus, 1758) TL: India M S Nessaea obrinus faventia Fruhstorfer, 1910 TL: Brazil (MT) M T Nessaea obrinus lesoudieri Le Moult, 1933 TL: Brazil (Am) M T Myscelia E. Doubleday, [1845] ref.198 Myscelia aracynthia (Dalman, 1823) Hispaniolan Bluewing Hispaniola iN M T S Myscelia capenas (Hewitson, [1857]) iN Myscelia capenas capenas (Hewitson, [1857]) TL: Brazil (Am) M T Myscelia capenas octomaculata (A. Butler, 1873) TL: Peru M T S Myscelia cyananthe C. Felder & R. Felder, 1867 Blackened Bluewing iN Myscelia cyananthe cyananthe C. Felder & R. Felder, 1867 E & S Mexico M T S Ab L Myscelia cyananthe diaziana R. G. Maza & J. Maza, 1985 Chiapas M T Myscelia cyananthe skinneri Mengel, 1894 NW Mexico, stray to S AZ, S CA, S NM M T S L Myscelia cyananthe streckeri Skinner, 1889 Baja California M T S L Myscelia cyaniris E. Doubleday, [1848] Whitened Bluewing iN Myscelia cyaniris cyaniris E. Doubleday, [1848] E Mexico to Panama M T S L Myscelia cyaniris albescens A. Hall, 1935 TL: Venezuela M T Myscelia cyaniris alvaradia R. G. Maza & Díaz, 1982 W Mexico M T S Ha Myscelia cyaniris millerorum Jenkins, 1984 TL: Ecuador Myscelia ethusa (Doyère, [1840]) Mexican Bluewing iN Myscelia ethusa ethusa (Doyère, [1840]) S TX, E & W Mexico M T S L1 L2 I H Myscelia ethusa chiapensis Jenkins, 1984 Chiapas M T Myscelia ethusa cyanecula C. Felder & R. Felder, 1867 TL: Mexico M T S Myscelia ethusa pattenia A. Butler & H. Druce, 1872 Guatemala to Costa Rica M T S L Myscelia hypatia Strecker, 1900 TL: [Brazil (Pa)] Myscelia leucocyana C. Felder & R. Felder, 1861 Pale Bluewing iN Myscelia leucocyana leucocyana C. Felder & R. Felder, 1861 Panama to Venezuela M T S L Myscelia leucocyana smalli Jenkins, 1984 Nicaragua & Costa Rica M T L Myscelia milloi Oberthür, 1916 iN M S Myscelia orsis (Drury, 1782) iN M S Tribe Cybdelini Cybdelis Boisduval, 1836 ref.198 Cybdelis boliviana Salvin, 1869 iN Cybdelis boliviana boliviana Salvin, 1869 TL: Bolivia M T Cybdelis boliviana [n. ssp.] Lamas, MS Peru Cybdelis mnasylus E. Doubleday, [1848] iN M T Cybdelis petronita Burmeister, 1861 ref.198 M T Cybdelis thrasylla C. Felder & R. Felder, 1859 ref.198 M T Cybdelis phaesyla (Hübner, [1831]) iN M S Sea sophronia (Godart, [1824]) TL: Brazil Sea sophronia sophronia (Godart, [1824]) iN M S Sea sophronia jenkinsi (F.M.S. Dias, R.R. Siewert & Freitas, 2019) ref.198 Tribe Eunicini Eunica alcmena (E. Doubleday, [1847]) Dark Purplewing iN Eunica alcmena alcmena (E. Doubleday, [1847]) S Mexico to Colombia & W Venezuela M T S L Eunica alcmena flora C. Felder & R. Felder, 1862 TL: Brazil (Am) M T S Eunica alpais (Godart, [1824]) iN Eunica alpais alpais (Godart, [1824]) TL: Brazil M T S Eunica alpais excelsa Godman & Salvin, 1877 S Mexico (Oaxaca, Chiapas) to E Panama M T S L Eunica araucana C. Felder & R. Felder, 1862 Araucana Purplewing W Panama, Colombia, W Venezuela, Ecuador iN M T S Eunica amelia (Cramer, 1777) iN Eunica amelia amelia (Cramer, 1777) TL: Surinam Eunica amelia erroneata Oberthür, 1916 TL: Peru; Brazil (Am) M T S Eunica amycla (Godart, [1824]) iN M T Eunica anna (Cramer, 1780) iN M T S Eunica bechina (Hewitson, 1852) iN Eunica bechina bechina (Hewitson, 1852) TL: Brazil (Am) M T Eunica bechina magnipunctata Talbot, 1928 TL: Brazil (MT) M T Eunica brunnea Salvin, 1869 TL: Peru M T Eunica caelina (Godart, [1824]) iN Eunica caelina caelina (Godart, [1824]) TL: Brazil M T S Eunica caelina agustina R. G. Maza, 1982 W Mexico M T S Eunica caelina alycia Fruhstorfer, 1909 TL: Upper Amazon M T S Eunica caelina augusta H. Bates, 1866 SE Mexico to W Panama M T S Eunica caelina olympias C. Felder & R. Felder, 1862 E Panama to Peru & Venezuela M T S Eunica caralis (Hewitson, [1857]) Caralis Purplewing iN Eunica caralis caralis (Hewitson, [1857]) Honduras, Panama, Colombia, Venezuela M T S Eunica caralis ariba Fruhstorfer, 1908 TL: Brazil? M S Eunica carias (Hewitson, [1857]) Carias Purplewing iN Eunica carias carias (Hewitson, [1857]) Costa Rica to Colombia M T S L Eunica carias cabira C. Felder & R. Felder, 1861 TL: Venezuela M S Eunica carias tenebrosa Salvin, 1869 TL: Peru M T S Eunica chlororhoa Salvin, 1869 iN Eunica chlororhoa chlororhoa Salvin, 1869 TL: Peru M T Eunica chlororhoa mira Godman & Salvin, 1877 Honduras to Colombia M T S L Eunica clytia (Hewitson, 1852) Clytia Purplewing W Panama to Peru & C Brazil iN M T S L Eunica concordia (Hewitson, 1852) iN M T S Eunica cuvierii (Godart, 1819) iN M S Eunica eburnea Fruhstorfer, 1907 iN M T Eunica eurota (Cramer, 1775) iN Eunica eurota eurota (Cramer, 1775) TL: West Indies M T Eunica eurota dolores (Prittwitz, 1871) TL: [Brazil (RJ)] M T Eunica evelide H. Bates, 1864 iN Eunica evelide evelide H. Bates, 1864 TL: [Colombia] M T S Eunica evelide elegans Salvin, 1869 TL: Bolivia; Peru M T S Eunica heraclitus Gundlach, 1881 Cuban Purplewing Cuba; apparently extinct M IL Eunica incognita Jenkins, 1990 iN M T Eunica ingens Seitz, 1915 iN M T Eunica interphasis Jenkins, 1990 TL: Brazil (Rd) M T Eunica macris (Godart, [1824]) iN Eunica macris macris (Godart, [1824]) TL: Brazil M T Eunica macris [n. ssp.] Lamas, MS Peru Eunica maja (Fabricius, 1775) iN Eunica maja maja (Fabricius, 1775) TL: Brazil M T S Eunica maja noerina A. Hall, 1935 TL: [Brazil (Am)] M T S Eunica malvina H. Bates, 1864 iN Eunica malvina malvina H. Bates, 1864 TL: [Brazil (Pa)] M T Eunica malvina albida Jenkins, 1990 SE Mexico to Panama M T S Eunica malvina almae Vargas, Llorente & Luis, 1996 W Mexico M T Eunica margarita (Godart, [1824]) iN M T Eunica marsolia (Godart, [1824]) iN Eunica marsolia marsolia (Godart, [1824]) TL: Brazil M S Eunica marsolia fasula Fruhstorfer, 1909 TL: [Peru] M T S Eunica marsolia paraensis Jenkins, 1990 TL: Brazil (Pa) M T Eunica monima (Stoll, 1782) Dingy Purplewing SE AZ & S TX (strays), E & W Mexico to Venez.; Gr. Antilles, S FL iN M T S L1 L2 Eunica mygdonia (Godart, [1824]) iN Eunica mygdonia mygdonia (Godart, [1824]) TL: Brazil M T S Eunica mygdonia omoa A. Hall, 1919 S Mexico to Panama M T S L Eunica norica (Hewitson, 1852) Midnight Purplewing iN Eunica norica norica (Hewitson, 1852) Costa Rica to Colombia, Venezuela & W Ecuador M T S L Eunica norica occia Fruhstorfer, 1909 TL: Peru M T S Eunica orphise (Cramer, 1775) iN M T S Eunica phasis C. Felder & R. Felder, 1862 iN M T Eunica pomona (C. Felder & R. Felder, 1867) iN Eunica pomona pomona (C. Felder & R. Felder, 1867) TL: Colombia M T Eunica pomona amata H. Druce, 1874 S Mexico (S Chiapas) to E Panama M T TI S L Eunica pusilla H. Bates, 1864 Pusilla Purplewing Costa Rica to Bolivia & SE Brazil iN M T S L Eunica sophonisba (Cramer, 1780) iN Eunica sophonisba sophonisba (Cramer, 1780) TL: Surinam M T S Eunica sophonisba agele Seitz, 1915 TL: Brazil (Am); Peru; Colombia M T Eunica sydonia (Godart, [1824]) iN Eunica sydonia sydonia (Godart, [1824]) TL: Brazil M T S Eunica sydonia caresa (Hewitson, [1857]) SE Mexico to E Panama M T S L Eunica tatila (Herrich-Schäffer, [1855]) Florida Purplewing iN Eunica tatila tatila (Herrich-Schäffer, [1855]) E & W Mexico (strays far N from E MX) to Costa Rica M T S L Eunica tatila bellaria Fruhstorfer, 1908 Panama to S Brazil & Argentina M T S Eunica tatila tatilina Fruhstorfer, 1908 TL: Upper Amazon? M T Eunica tatila tatilista Kaye, 1926 S FL, Bahamas, Greater Antilles, Mona M S L H Eunica veronica H. Bates, 1864 iN M T Eunica viola H. Bates, 1864 iN M T S Eunica violetta Staudinger, [1885] TL: Peru M T Eunica volumna (Godart, [1824]) iN Eunica volumna volumna (Godart, [1824]) TL: Brazil M T S Eunica volumna celma (Hewitson, 1852) TL: [Brazil (Am)] M T S Eunica volumna venusia (C. Felder & R. Felder, 1867) SE Mexico (Chiapas) to Colombia M T S L Tribe Ageroniini Batesia C. Felder & R. Felder, 1862 Batesia hypochlora C. Felder & R. Felder, 1862 iN Batesia hypochlora hypochlora C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Batesia hypochlora hypoxantha Salvin & Godman, 1868 TL: Peru M T S Ectima E. Doubleday, [1848] Ectima erycinoides C. Felder & R. Felder, 1867 iN Ectima erycinoides Northern segregate , SE Mexico to Panama M S L Ectima erycinoides erycinoides C. Felder & R. Felder, 1867 TL: Colombia M T Ectima erycinoides ecuadorensis Jenkins, 1985 TL: Ecuador Ectima erycinoides [n. ssp.] Lamas, MS Mexico (Chia) Ectima iona E. Doubleday, [1848] iN M T Ectima lirides Staudinger, [1885] iN M T S Ectima thecla (Fabricius, 1796) iN Ectima thecla thecla (Fabricius, 1796) TL: Indiis M T Ectima thecla lirina C. Felder & R. Felder, 1867 TL: Brazil? M T Ectima thecla peruviana Bryk, 1953 TL: Peru Hamadryas Hübner, [1806] Hamadryas albicornis (Staudinger, [1885]) TL: Peru M T Hamadryas alicia (H. Bates, 1865) TL: Brazil (Am) M T Hamadryas amphichloe (Boisduval, 1870) iN Hamadryas amphichloe amphichloe (Boisduval, 1870) TL: Ecuador M T Hamadryas amphichloe lamasi Jenkins, 1983 TL: Peru M T Hamadryas amphinome (Linnaeus, 1767) iN Hamadryas amphinome amphinome (Linnaeus, 1767) TL: S. Amer. M T Hamadryas amphinome fumosa (Fruhstorfer, 1915) TL: Colombia M T Hamadryas amphinome mazai Jenkins, 1983 W Mexico M S L Ha Hamadryas amphinome mexicana (Lucas, 1853) E Mexico to Colombia, Cuba M T S L Hamadryas arete (E. Doubleday, 1847) iN M T Hamadryas arinome (Lucas, 1853) iN Hamadryas arinome arinome (Lucas, 1853) TL: French Guiana M T Hamadryas arinome arienis (Godman & Salvin, 1883) Costa Rica to Bolivia M T S L Hamadryas arinome obnubila (Fruhstorfer, 1916) TL: Brazil (ES) M T Hamadryas atlantis (H. Bates, 1864) Black-patched Cracker iN Hamadryas atlantis atlantis (H. Bates, 1864) S Mexico (Chiapas), Guatemala, Honduras M T S Hamadryas atlantis lelaps (Godman & Salvin, 1883) W Mexico M T S L Hamadryas belladonna (H. Bates, 1865) iN M T Hamadryas chloe (Stoll, 1787) iN Hamadryas chloe chloe (Stoll, 1787) TL: Surinam M T Hamadryas chloe daphnis (Staudinger, 1886) TL: Peru M T Hamadryas chloe obidona (Fruhstorfer, 1914) TL: Brazil (Pa) M T Hamadryas chloe rhea (Fruhstorfer, 1907) TL: Brazil (ES/MG) M T Hamadryas epinome (C. Felder & R. Felder, 1867) iN M T Hamadryas februa (Hübner, [1823]) iN Hamadryas februa NW Mexican segregate , NW Mexico M S L I Hamadryas februa ferentina (Godart, [1824]) S TX, E & S Mexico to Colombia, Trinidad, Guianas & Amazon M T S L1 L2 I Hamadryas februa ferox (Staudinger, 1886) S FL (rare stray), Caribbean isls., Pan to Venez; ref.135,136 M T S1 S2 L Hamadryas februa februa (Hübner, [1823]) TL: ? M T Hamadryas feronia (Linnaeus, 1758) iN Hamadryas feronia feronia (Linnaeus, 1758) TL: India M T HS Hamadryas feronia farinulenta (Fruhstorfer, 1916) S TX (rare stray), E & W Mexico to W Peru, Venezuela, Trinidad M T S L Hamadryas fornax (Hübner, [1823]) iN Hamadryas fornax fornax (Hübner, [1823]) TL: ? Hamadryas fornax fornacalia (Fruhstorfer, 1907) E Mexico to Colombia & Venezuela M T S L Hamadryas glauconome (H. Bates, 1864) Glaucous Cracker iN Hamadryas glauconome glauconome (H. Bates, 1864) S TX (rare stray), E & S Mexico to Costa Rica (Panama?) M T S L Hamadryas glauconome grisea Jenkins, 1983 NW Mexico M T S L Hamadryas guatemalena (H. Bates, 1864) Guatamalan Cracker iN Hamadryas guatemalena guatemalena (H. Bates, 1864) SE Mexico (much of Chiapas, Yucatan Peninsula) to Costa Rica M T S L Hamadryas guatemalena marmarice (Fruhstorfer, 1916) S TX, most of Mexico M T S L I Hamadryas iphthime (H. Bates, 1864) Brownish Cracker iN Hamadryas iphthime joannae Jenkins, 1983 S TX (rare stray), E & S Mexico to Costa Rica M S1 S2 L1 L2 Hamadryas iphthime iphthime (H. Bates, 1864) Costa Rica, Panama to Guyana, Colombia, Peru, Bolivia, S Brazil M T L Hamadryas iphthime joannae Jenkins, 1983 S TX (rare stray), E & S Mexico to Costa Rica M S1 S2 L1 L2 Hamadryas iphthime griffoni Brévignon, 2006 TL: French Guiana Hamadryas julitta (Fruhstorfer, 1914) Yucatan Cracker [= honorina], Yucatan Peninsula iN M T S L Hamadryas laodamia (Cramer, 1777) iN Hamadryas laodamia laodamia (Cramer, 1777) TL: Surinam M T Hamadryas laodamia saurites (Fruhstorfer, 1916) SE Mexico to Colombia, N Venezuela, Trinidad M T S L I Hamadryas rosandra (Fruhstorfer, 1916) TL: Brazil (Am) M T Hamadryas velutina (H. Bates, 1865) iN Hamadryas velutina velutina (H. Bates, 1865) TL: [Brazil (Am)] M T Hamadryas velutina browni Jenkins, 1983 TL: Brazil (MT) M T Panacea Godman & Salvin, 1883 Panacea procilla (Hewitson, [1854]) iN Panacea procilla procilla (Hewitson, [1854]) TL: Colombia M T S Panacea procilla bleuzeni Plantrou & Attal, 1986 TL: French Guiana M T Panacea procilla divalis (H. Bates, 1868) TL: Peru M T S Panacea procilla lysimache Godman & Salvin, 1883 Costa Rica & Panama M T Panacea procilla mamorensis A. Hall, 1917 TL: Brazil (Rd) M T Panacea prola (E. Doubleday, [1848]) iN Panacea prola prola (E. Doubleday, [1848]) TL: Colombia M T S Panacea prola amazonica Fruhstorfer, 1915 TL: [Peru] M T S Panacea regina (H. Bates, 1864) iN Panacea regina regina (H. Bates, 1864) TL: Brazil (Am) M T S Panacea regina chalcothea (H. Bates, 1868) TL: Colombia; [Peru] M T S Tribe Epiphilini Asterope Hübner, [1819] Asterope batesii (Hewitson, 1850) TL: Brazil (Am) M T Asterope buckleyi (Hewitson, 1869) TL: Ecuador M T Asterope degandii (Hewitson, 1858) Red-based Banner iN Asterope degandii degandii (Hewitson, 1858) Peru M T OI S Asterope degandii adamsi (Lathy, 1903) TL: Peru M T Asterope degandii allyni Jenkins, 1987 TL: Peru M T Asterope degandii bartletti (Godman & Salvin, 1878) TL: Peru M T Asterope degandii caelestis Attal & Hiltbrand, 2006 ref.160 M T Asterope degandii orientalis Attal, 1999 TL: Brazil (Pa) M T Asterope degandii mocoensis Attal, Pacl & Hiltbrand, 2020 ref.161 Asterope leprieuri (Feisthamel, 1835) iN Asterope leprieuri leprieuri (Feisthamel, 1835) TL: [Brazil (Pa)] M T Asterope leprieuri depuiseti (C. Felder & R. Felder, 1861) TL: Ecuador M T S Asterope leprieuri optima (A. Butler, 1869) TL: Peru M T Asterope leprieuri philotima (Rebel, 1912) TL: Peru M T S Asterope markii (Hewitson, 1857) iN Asterope markii markii (Hewitson, 1857) TL: [Brazil (Am)] M T S Asterope markii ackeryi Jenkins, 1987 TL: Brazil (Pa) M T Asterope markii boyi (Röber, 1924) TL: Brazil ([Pa]) M T Asterope markii davisii (A. Butler, 1877) TL: Peru M T Asterope markii gallardi Neukirchen, 1996 TL: French Guiana Asterope markii hewitsoni (Staudinger, 1886) TL: Colombia M T Asterope markii werneri Neukirchen, 1995 TL: Brazil (Am) M T Asterope sapphira (Hübner, [1816]) TL: Brazil M T Asterope whitelyi (Salvin, 1869) TL: Peru Asterope whitelyi whitelyi (Salvin, 1869) TL: Peru M T Asterope whitelyi cyaneogyna Attal & Hiltbrand, 2006 ref.160 M T Asterope whitelyi glennae Attal & Hiltbrand, 2006 ref.160 M T Asterope whitelyi srnkai (Honrath, 1884) TL: Peru M T Bolboneura Godman & Salvin, 1877 Bolboneura sylphis (H. Bates, 1864) Variable Banner iN Bolboneura sylphis sylphis (H. Bates, 1864) C & W Chiapas,Guatemala, Honduras M T S L Bolboneura sylphis beatrix R. G. Maza, 1985 SW Mexico (MICH, GRO, MOR, SW PUE, C & S OAX) M T L Bolboneura sylphis lacandona R. G. Maza & J. Maza, 1985 E Chiapas M T Bolboneura sylphis veracruzana (Draudt, 1931) E Mexico M T S L Epiphile E. Doubleday, [1845] Epiphile adrasta Hewitson, 1861 Common Banner iN Epiphile adrasta adrasta Hewitson, 1861 S TX (as vagrant), E Mexico to W Panama M T S L Epiphile adrasta bandusia Fruhstorfer, 1912 C Panama to W Ecuador M T S L Epiphile adrasta calixto Attal & Crosson, 2003 ref.159 Epiphile adrasta escalantei Descimon & Mast, 1979 W Mexico (SIN to S OAX) M S L Epiphile boliviana Röber, 1914 iN Epiphile boliviana boliviana Röber, 1914 TL: Bolivia M T Epiphile boliviana grosi Attal & Crosson, 2003 ref.159 M T Epiphile boliviana lamasi Neild, 1996 TL: Venezuela M T Epiphile chrysites (Latreille, [1809]) Chrysites Banner iN Epiphile chrysites chrysites (Latreille, [1809]) [W Panama?], Venezuela to Bolivia M T S L Epiphile chrysites [n. ssp.] Lamas, MS Bolivia M T Epiphile dilecta Röber, 1914 iN Epiphile dilecta dilecta Röber, 1914 TL: Bolivia M T Epiphile dilecta jenkinsi Lamas, 1995 TL: Peru Epiphile dilecta venezuelensis Jenkins, 1986 TL: Venezuela M T Epiphile epicaste Hewitson, 1857 iN Epiphile epicaste epicaste Hewitson, 1857 TL: Colombia M T Epiphile epicaste [n. ssp.] Lamas, MS Peru Epiphile epimenes Hewitson, 1857 iN Epiphile epimenes epimenes Hewitson, 1857 TL: Colombia M T Epiphile epimenes electra Staudinger, [1885] TL: Venezuela M T Epiphile epimenes kalbreyeri Fassl, 1912 TL: Colombia M T Epiphile eriopis Hewitson, 1857 iN Epiphile eriopis eriopis Hewitson, 1857 TL: Colombia M T Epiphile eriopis devriesi Jenkins, 1986 Nicaragua to Colombia M T S Epiphile grandis A. Butler, 1872 Grand Banner Costa Rica & Panama M T OI S Epiphile hermosa J. Maza & Díaz, 1978 Beautiful Banner SE Mexico, Guatemala iN M T S L Epiphile hubneri Hewitson, 1861 iN Epiphile iblis C. Felder & R. Felder, 1861 iN Epiphile iblis iblis C. Felder & R. Felder, 1861 TL: Venezuela; Colombia M T Epiphile iblis plusios Godman & Salvin, 1883 Costa Rica & Panama M T S L Epiphile iblis plutonia H. Bates, 1864 SE Mexico & Guatemala M T S Epiphile iblis pyrczi Attal, 1999 TL: Colombia M T Epiphile imperator Attal, 2005 TL: Ecuador Epiphile imperator imperator Attal, 2005 ref.157 M T Epiphile imperator duponti Attal, 2005 ref.157 M T Epiphile imperator willmotti Attal, 2005 ref.157 Epiphile lampethusa E. Doubleday, [1848] iN Epiphile lampethusa lampethusa E. Doubleday, [1848] TL: Bolivia M T Epiphile lampethusa zipa Mengel, 1899 TL: not specified M T Epiphile latifasciata Röber, 1914 iN Epiphile latifasciata latifasciata Röber, 1914 TL: Bolivia M T Epiphile latifasciata obscurior Attal, 1997 TL: Ecuador M T Epiphile orea (Hübner, [1823]) iN Epiphile orea orea (Hübner, [1823]) TL: ? Epiphile orea distalis Attal, 1997 TL: Ecuador M T Epiphile orea helios Attal & Crosson, 2003 ref.159 Epiphile orea negrina C. Felder & R. Felder, 1862 TL: Brazil (Am) M T S Epiphile neildi Attal, 2005 iN Epiphile neildi neildi Attal, 2005 ref.157 M T Epiphile neildi bakuru Attal & G. Rodríguez, 2006 ref.160 M T Lucinia Hübner, [1823] Lucinia sida Hübner, [1823] Caribbean Banner iN Lucinia sida sida Hübner, [1823] Cuba M T S Lucinia sida albomaculata Rindge, 1955 Bahamas M T S Lucinia sida torrebia (Ménétriés, 1832) Hispaniola M S Lucinia cadma (Drury, 1773) Jamaican Banner Jamaica M T S Nica Hübner, [1826] Nica flavilla (Godart, [1824]) iN Nica flavilla bachiana (R. G. Maza & J. Maza, 1985) W Mexico M T S L Nica flavilla E Mexican segregate , E Mexico M S L Nica flavilla canthara (E. Doubleday, 1849) Nicaragua to Venezuela (at least) M T S L1 L2 Nica flavilla flavilla (Godart, [1824]) TL: Brazil M T Nica flavilla lucillae Neild, 1996 TL: Venezuela M T Nica flavilla sylvestris H. Bates, 1864 TL: Brazil (Am) M T Nica flavilla [n. ssp.] Lamas, MS Mexico Peria W. F. Kirby, 1871 Peria lamis (Cramer, 1779) iN M S Pyrrhogyra Hübner, [1819] Pyrrhogyra amphiro H. Bates, 1865 iN Pyrrhogyra amphiro amphiro H. Bates, 1865 TL: Brazil (Am) M T S Pyrrhogyra amphiro agilis Brévignon, 1995 TL: French Guiana Pyrrhogyra amphiro juani Staudinger, 1886 TL: Colombia M T Pyrrhogyra amphiro typhoeus C. Felder & R. Felder, 1867 TL: Colombia M T Pyrrhogyra crameri Aurivillius, 1882 iN Pyrrhogyra crameri Northern segregate , Nicaragua to Panama M L Pyrrhogyra crameri nautaca Fruhstorfer, 1908 TL: Colombia; "Upper Amazon" M T Pyrrhogyra crameri undine Bargmann, 1929 TL: Colombia M T Pyrrhogyra crameri hagnodorus Fruhstorfer, 1908 TL: Peru M T Pyrrhogyra crameri crameri Aurivillius, 1882 TL: Surinam M T Pyrrhogyra crameri [n. ssp.] Lamas, MS Panama Pyrrhogyra edocla (E. Doubleday, [1848]) Green-spotted Banner iN Pyrrhogyra edocla edocla (E. Doubleday, [1848]) E Mexico to Colombia & Venezuela M T S L Pyrrhogyra edocla cuparina H. Bates, 1865 TL: Brazil (Pa) M T Pyrrhogyra edocla lysanias C. Felder & R. Felder, 1862 TL: Brazil (Am) M T S Pyrrhogyra edocla paradisea R. G. Maza & J. Maza, 1985 W Mexico M T S Pyrrhogyra edocla perscita Brévignon, 1996 TL: French Guiana Pyrrhogyra neaerea (Linnaeus, 1758) iN Pyrrhogyra neaerea neaerea (Linnaeus, 1758) TL: Indiis M T HS Pyrrhogyra neaerea arge Gosse, 1880 TL: Paraguay M T L Pyrrhogyra neaerea argina Fruhstorfer, 1908 TL: Upper Amazon M T Pyrrhogyra neaerea hypsenor Godman & Salvin, 1884 E & W Mexico to Panama M T S L Pyrrhogyra neaerea kheili Fruhstorfer, 1908 TL: Colombia M T Pyrrhogyra neaerea ophni A. Butler, 1870 TL: Brazil (MG) M T Pyrrhogyra neaerea susarion Fruhstorfer, 1912 TL: Brazil (Pe) M T Pyrrhogyra otolais H. Bates, 1864 Double-banded Banner iN Pyrrhogyra otolais otolais H. Bates, 1864 E & W Mexico to Panama M T S L Pyrrhogyra otolais nasica Staudinger, 1886 TL: Colombia M T Pyrrhogyra otolais olivenca Fruhstorfer, 1908 TL: Brazil (Am) M T S Pyrrhogyra otolais placita Brévignon, 1996 TL: French Guiana Pyrrhogyra otolais seitzi Fruhstorfer, 1908 TL: Bolivia M T Pyrrhogyra stratonicus Fruhstorfer, 1908 iN Pyrrhogyra stratonicus stratonicus Fruhstorfer, 1908 TL: Upper Amazon M T S Pyrrhogyra stratonicus matthias Brévignon, 1995 TL: French Guiana Temenis Hübner, [1819] Temenis huebneri Fruhstorfer, 1907 iN Temenis huebneri huebneri Fruhstorfer, 1907 TL: Brazil (MG) M T Temenis huebneri korallion Fruhstorfer, 1912 TL: Brazil (SP) M T Temenis huebneri pedania Fruhstorfer, 1912 TL: Brazil (Pe) M T Temenis laothoe (Cramer, 1777) iN Temenis laothoe quilapayunia R. G. Maza & Turrent, 1985 W Mexico M T S L Ha Temenis laothoe E Mexican segregate , S TX (as vagrant), E Mexico M S L Temenis laothoe hondurensis Fruhstorfer, 1907 S Mexico (Chiapas) to Colombia M T S L I Temenis laothoe laothoe (Cramer, 1777) TL: Surinam M T Temenis laothoe bahiana Fruhstorfer, 1907 TL: Brazil (Ba) M T Temenis laothoe columbiana Fruhstorfer, 1907 TL: Colombia M T Temenis laothoe meridionalis Ebert, 1965 TL: Brazil (SP) M T S Temenis laothoe santina Fruhstorfer, 1907 TL: Brazil (ES); Paraguay M T Callicorina Smart, 1975 Callicorina pulchra (Hewitson, 1861) ref.12 iN Callicorina pulchra Northern segregate , Costa Rica & Panama Callicorina pulchra pulchra (Hewitson, 1861) TL: Colombia M T Callicorina pulchra amazonica Fruhstorfer, 1907 TL: [Brazil (Am)] Callicorina pulchra pallidior (Oberthür, 1901) TL: Peru; Bolivia; Brazil (Pa) M T Callicorina pulchra [n. ssp.] Lamas, MS Panama Tribe Eubagini Dynamine Hübner, [1819] Dynamine aerata (A. Butler, 1877) iN Dynamine aerata aerata (A. Butler, 1877) TL: Brazil (Am) M T S Dynamine aerata [n. ssp.#1] Lamas, MS TL: Venezuela Dynamine aerata [n. ssp.#2] Lamas, MS TL: Paraguay Dynamine agacles (Dalman, 1823) iN Dynamine agacles agacles (Dalman, 1823) TL: Brazil M T Dynamine agacles core Röber, 1915 Costa Rica to Colombia M T S L Dynamine agatha (Oberthür, 1916) TL: Brazil (Am) M T Dynamine amplias (Hewitson, 1859) iN M T Dynamine anubis (Hewitson, 1859) iN Dynamine anubis anubis (Hewitson, 1859) TL: Brazil (Am) M T Dynamine anubis pieridoides (C. Felder & R. Felder, 1867) TL: Colombia M T Dynamine arene Hübner, [1823] iN M S Dynamine artemisia (Fabricius, 1793) iN Dynamine artemisia Northern segregate , E & S Mexico to Panama M S Ab L Dynamine artemisia artemisia (Fabricius, 1793) TL: America M S Dynamine artemisia ackeryi Neild, 1996 TL: Venezuela M T Dynamine artemisia glauce (H. Bates, 1865) TL: Brazil (Am); "Guatemala" M T S Dynamine artemisia [n. ssp.#1] Lamas, MS TL: Mexico Dynamine artemisia [n. ssp.#2] Lamas, MS TL: Venezuela Dynamine ate (Godman & Salvin, 1883) Little Sailor SE Mexico (SE Chiapas), Guatemala, Honduras, Costa Rica iN M T S Dynamine athemon (Linnaeus, 1758) iN Dynamine athemon athemon (Linnaeus, 1758) TL: America Dynamine athemon amazonica Röber, 1915 TL: Brazil (Am) M S Dynamine athemon athemaena (Hübner, [1824]) TL: Brazil Dynamine athemon barreiroi A. Fernández, 1928 TL: Peru M T L Dynamine athemon isolda A. Hall, 1919 TL: Colombia M T Dynamine athemon maeon (E. Doubleday, 1849) Panama & Colombia M T OI L Dynamine athemon niveata (A. Butler, 1877) TL: Brazil (Am) M T Dynamine chiquita Willmott & J. Hall, 2010 ref.320 Dynamine chryseis (H. Bates, 1865) Thick-rimmed Sailor Nicaragua to Amazon Basin iN M T S L Dynamine coenus (Fabricius, 1793) iN Dynamine coenus coenus (Fabricius, 1793) TL: Indiis M T Dynamine coenus albidula Weeks, 1901 TL: Bolivia M T Dynamine coenus leucothea (H. Bates, 1865) TL: Brazil (Am) M T Dynamine colombiana Talbot, 1932 Colombian Sailor Nicaragua to Colombia & NW Venezuela iN M T L Dynamine dyonis Geyer, 1837 Blue-eyed Sailor S TX, E & W Mexico to Panama iN M S L Dynamine erchia (Hewitson, 1852) iN M T Dynamine gisella (Hewitson, 1857) iN M T Dynamine haenschi A. Hall, 1917 iN M T Dynamine hecuba (Schaus, 1913) Costa Rican Sailor Costa Rica M T S Dynamine ines (Godart, [1824]) iN Dynamine ines ines (Godart, [1824]) TL: Brazil M T Dynamine ines geta (Godman & Salvin, 1878) TL: Bolivia M S Dynamine ines [n. ssp.] Lamas, MS Colombia Dynamine intermedia Talbot, 1932 TL: Peru M T Dynamine laugieri (Oberthür, 1916) TL: French Guiana M T Dynamine meridionalis Röber, 1915 iN Dynamine myrrhina (E. Doubleday, 1849) iN M T Dynamine myrson (E. Doubleday, 1849) iN Dynamine myrson myrson (E. Doubleday, 1849) TL: Brazil (Pa) M T S Dynamine myrson manriquei Neild, 1996 TL: Peru M T Dynamine neoris (Hewitson, 1859) TL: [Brazil (Am)] M T Dynamine onias (Hewitson, 1857) iN M T Dynamine paulina (H. Bates, 1865) iN Dynamine paulina paulina (H. Bates, 1865) TL: Brazil (Am) M T Dynamine paulina thalassina (Boisduval, 1870) Guatemala to Colombia M T S L Dynamine pebana Staudinger, [1885] iN Dynamine pebana pebana Staudinger, [1885] TL: Peru M T Dynamine pebana elisa Brévignon, 1995 TL: French Guiana Dynamine perpetua (H. Bates, 1865) TL: Brazil (Am) M T Dynamine persis (Hewitson, 1859) TL: Brazil (Am) M T Dynamine postverta (Cramer, 1779) iN Dynamine postverta postverta (Cramer, 1779) TL: Surinam M S Dynamine postverta mexicana R.F. d'Almeida, 1952 [= mylitta], S TX (rare stray), E & W Mexico, C America, Cuba M T S L1 L2 Dynamine racidula (Hewitson, 1852) iN Dynamine racidula racidula (Hewitson, 1852) TL: Brazil (Am) M T S L Dynamine racidula trembathi Neild, 1996 TL: Venezuela M T Dynamine sara (H. Bates, 1865) iN M T S Dynamine serina (Fabricius, 1775) Caribbean Sailor iN Dynamine serina serina (Fabricius, 1775) [= egaea], Jamaica M T S Dynamine serina calais M. Bates, 1936 Cuba M T IL Dynamine serina zetes (Ménétriés, 1832) Hispaniola M S Dynamine setabis (E. Doubleday, 1849) iN Dynamine setabis setabis (E. Doubleday, 1849) TL: Venezuela; Colombia M T Dynamine setabis agnes Röber, 1915 TL: Peru M T S Dynamine sosthenes (Hewitson, 1869) Thin-rimmed Sailor iN Dynamine sosthenes sosthenes (Hewitson, 1869) Nicaragua & Costa Rica M T S Dynamine sosthenes smerdis Tessmann, 1928 TL: Peru M T S Dynamine sosthenes venezolensis Tessmann, 1928 TL: Venezuela M T Dynamine theseus (C. Felder & R. Felder, 1861) White Sailor S Mexico to Colombia & Venezuela iN M T S L Dynamine tithia (Hübner, 1823) iN Dynamine tithia tithia (Hübner, [1823]) TL: Brazil M T Dynamine tithia dentilinea Talbot, 1932 TL: Ecuador M T Dynamine tithia salpensa (C. Felder & R. Felder, 1862) Andes and upper Amazon basin, Colombia to Bolivia (hypothetical for Central America) M T S Dynamine tithia [n. ssp.] Lamas, MS Colombia Dynamine vicaria (H. Bates, 1865) iN Dynamine vicaria vicaria (H. Bates, 1865) TL: Brazil (Am) M T Dynamine vicaria gillotti A. Hall, 1930 Costa Rica M T S Dynamine vicaria hoppi M. Hering, 1926 TL: Colombia M T Dynamine zenobia (H. Bates, 1865) TL: Brazil (Am) Dynamine zenobia zenobia (H. Bates, 1865) TL: Brazil (Am) M T S Dynamine zenobia ampliata J. Zikán, 1937 TL: Colombia M T Dynamine [n. sp.] Lamas, MS Peru Tribe Callicorini Haematera E. Doubleday, 1849 Haematera pyrame (Hübner, [1819]) iN Haematera pyrame Northern segregate , Nicaragua to Colombia M S L Haematera pyrame pyrame (Hübner, [1819]) TL: French Guiana M T Haematera pyrame rubra Kaye, 1904 TL: Trinidad Haematera pyrame thysbe E. Doubleday, 1849 TL: Venezuela; Colombia M T Haematera pyrame [n. ssp.#1] Lamas, MS TL: Panama Haematera pyrame [n. ssp.#2] Lamas, MS TL: Peru Haematera pyrame [n. ssp.#3] Lamas, MS TL: Peru Antigonis C. Felder, 1861 Antigonis pharsalia (Hewitson, 1852) TL: [Brazil (Am)] Antigonis pharsalia pharsalia (Hewitson, 1852) TL: [Brazil (Am)] M T Antigonis pharsalia felderi H. Bates, 1864 TL: Brazil (Am) M T Callicore Hübner, [1819] Callicore astarte (Cramer, 1779) iN Callicore astarte astarte (Cramer, 1779) TL: Surinam M T Callicore astarte antillena (Kaye, 1914) TL: Trinidad; "St. Lucia" Callicore astarte astartoides (Dillon, 1948) TL: Bolivia M T Callicore astarte casta (Salvin, 1869) Veracruz & Oaxaca, Mexico M T S Callicore astarte codomannus (Fabricius, 1781) TL: Brazil M T Callicore astarte lilliputa (Dillon, 1948) TL: Colombia M T Callicore astarte otheres (Fruhstorfer, 1916) TL: Colombia Callicore astarte panamaensis Kariya, 2006 TL: Panama Callicore astarte patelina (Hewitson, 1853) Chiapas to Costa Rica M T S L Callicore astarte selima (Guenée, 1872) TL: Brazil (MG) M T Callicore astarte staudingeri (Dillon, 1948) Panama (Darien), Colombia (Antioquia), Venezuela (Aragua, Trujillo, Perija) M T S Callicore astarte stratiotes (C. Felder & R. Felder, 1861) TL: Ecuador M T S Callicore astarte [n. ssp.] Lamas, MS Brazil Callicore ines (Hopp, 1922) iN M T Callicore cynosura (E. Doubleday, [1847]) iN Callicore cynosura cynosura (E. Doubleday, [1847]) TL: Bolivia M T S Callicore cynosura amazona (H. Bates, 1864) TL: Brazil (Pa) M T Callicore cynosura fulva (Dillon, 1948) TL: Brazil (Am) M T Callicore excelsior (Hewitson, [1858]) iN Callicore excelsior excelsior (Hewitson, [1858]) TL: Brazil (Am) M T Callicore excelsior arirambae (Ducke, 1913) TL: Brazil (Pa) M T Callicore excelsior elatior (Oberthür, 1916) TL: Ecuador Callicore excelsior excelsissima (Staudinger, [1885]) TL: Brazil (Am) M T Callicore excelsior inferior (A. Butler, 1877) TL: Peru M T Callicore excelsior marisolae Neukirchen, 1995 TL: Brazil (Am) M T Callicore excelsior mauensis (Fassl, 1922) TL: Brazil (Am) Callicore excelsior michaeli (Staudinger, 1890) TL: Brazil (Am) M T Callicore excelsior micheneri (Dillon, 1948) TL: Colombia M T Callicore excelsior ockendeni (Oberthür, 1916) TL: Peru M T Callicore excelsior pastazza (Staudinger, 1886) TL: Peru M T Callicore excelsior splendida (Weymer, 1890) TL: Peru M T Callicore sorana (Godart, [1824]) iN Callicore sorana sorana (Godart, [1824]) TL: Brazil M T L Callicore sorana horstii (Mengel, 1916) TL: Bolivia M T Callicore cyclops (Staudinger, 1891) iN M T Callicore pitheas (Latreille, [1813]) Two-eyed Eighty-eight W Chiapas to Venezuela & W Ecuador iN M T S1 S2 L I1 I2 Callicore texa (Hewitson, [1855]) iN Callicore texa texa (Hewitson, [1855]) TL: Colombia M T Callicore texa aretas (Hewitson, 1857) TL: Venezuela M T Callicore texa bari (Oberthür, 1916) TL: French Guiana; "Peru" M T Callicore texa boyeri Attal, 1999 TL: Ecuador M T Callicore texa fassli (Röber, 1915) TL: Bolivia M T Callicore texa heroica (Fruhstorfer, 1916) Veracruz, N Oaxaca M T S Callicore texa kayei (A. Hall, 1917) TL: Trinidad M T Callicore texa loxicha R. G. Maza & J. Maza, 1983 S Oaxaca (Sierra Madre del Sur) M T S L Callicore texa maimuna (Hewitson, 1858) TL: [Peru] M T Callicore texa maximilla (Fruhstorfer, 1916) TL: Brazil (ES) M T Callicore texa sigillata (Kotzsch, 1939) TL: Peru M T Callicore texa skinneri Neild, 1996 TL: Venezuela Callicore texa strympli (Fassl, 1922) TL: Brazil (Pa) Callicore texa tacana R. G. Maza & J. Maza, 1983 W Chiapas to Guatemala M T S Callicore texa titania (Salvin, 1869) far SE Mexico (E Chiapas) to Colombia M T S L I Callicore hydaspes (Drury, 1782) iN M T Callicore lyca (E. Doubleday, [1847]) Six-spotted Eighty-eight iN Callicore lyca lyca (E. Doubleday, [1847]) S Mexico to Honduras M T S Callicore lyca aegina (C. Felder & R. Felder, 1861) TL: Ecuador M T S L Callicore lyca aerias (Godman & Salvin, 1883) Nicaragua to Panama M T S1 S2 L I1 I2 Callicore lyca bella (Röber, 1924) TL: Colombia M T Callicore lyca exultans (Fruhstorfer, 1916) TL: Bolivia M T Callicore lyca mena (Staudinger, 1886) TL: Peru M T Callicore lyca mionina (Hewitson, 1855) TL: Colombia M T Callicore lyca odilia (Oberthür, 1916) TL: Colombia M T Callicore lyca salamis (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Callicore lyca sticheli (Dillon, 1948) TL: Honduras M T Callicore brome (Doyère, [1840]) iN Callicore brome brome (Doyère, [1840]) TL: ? M T Callicore brome angusta Attal, 2000 Costa Rica M T L I Callicore brome danieli Attal & Crosson, 2003 Panama (Darien); ref.159 M T L Callicore maronensis (Oberthür, 1916) iN M T Callicore hesperis (Guérin-Méneville, [1844]) iN M T Callicore atacama (Hewitson, 1852) iN Callicore atacama folschveilleri (Attal, 2008) Costa Rica & W Panama M T Callicore atacama manova (Fruhstorfer, 1916) TL: Colombia M T Callicore atacama faustina (H. Bates, 1866) Costa Rica (Atlantic slope Talamancas) & Panama M T Callicore atacama atacama (Hewitson, 1852) TL: Ecuador M T Callicore felderi (Hewitson, 1864) iN Callicore felderi felderi (Hewitson, 1864) TL: [Peru] M T Callicore felderi cajetani (Guenée, 1872) TL: Peru M T Catagramma Boisduval, 1836 Catagramma eunomia Hewitson, 1853 iN Catagramma eunomia eunomia Hewitson, 1853 ref.213 M T Catagramma eunomia alani (Attal & Crosson, 2003) ref.213 M T Catagramma eunomia atahualpa (Attal, 2006) ref.213 Catagramma eunomia ferrerorum Attal, 2000 ref.213 Catagramma eunomia incarnata Röber, 1915 ref.213 M T Catagramma eunomia valeriae (Neild, 1996) ref.213 M T Catagramma hydarnis (Godart, [1824]) ref.213 iN M T Catagramma hystaspes (Fabricius, 1781) iN Catagramma hystaspes hystaspes (Fabricius, 1781) ref.213 M T Catagramma hystaspes discrepans Stichel, 1936 ref.213 M T Catagramma hystaspes zelphanta Hewitson, 1858 ref.213 M T S Catagramma pygas (Godart, [1824]) iN Catagramma pygas pygas (Godart, [1824]) ref.213 M T Catagramma pygas aphidna Hewitson, 1869 ref.213 M T Catagramma pygas concolor Talbot, 1928 ref.213 M T Catagramma pygas cyllene E. Doubleday, [1847] ref.213 M T S Catagramma pygas eucale Fruhstorfer, 1916 ref.213 M T Catagramma pygas lalannensis Brévignon, 1995 ref.213 Catagramma pygas rondoni Ribeiro, 1931 ref.213 M T Catagramma pygas thamyras Ménétriés, 1857 ref.213 M T Catagramma tolima Hewitson, 1852 iN Catagramma tolima tolima Hewitson, 1852 ref.213 M T Catagramma tolima bugaba Staudinger, 1876 W Costa Rica & Panama; ref.213 M T S1 S2 L I Catagramma tolima chimana Oberthür, 1916 ref.213 M T Catagramma tolima aurantiaca Oberthür, 1916 TL: Colombia M T Catagramma tolima denina Hewitson, 1858 ref.213 M T Catagramma tolima guatemalena H. Bates, 1866 Guatemala; ref.213 M T TI L Catagramma tolima pacifica H. Bates, 1866 C & S Chiapas to W Guatemala; ref.213 M T S Catagramma tolima peralta Dillon, 1948 Honduras to E Costa Rica & Panama; ref.213 M T S1 S2 L I Catagramma tolima tehuana (R. G. Maza & J. Maza, 1983) Veracruz, Tabasco, Oaxaca, N Chiapas (vic. Palenque); ref.213 M T S L Catagramma pyracmon (Godart, [1824]) iN Catagramma pyracmon pyracmon (Godart, [1824]) ref.213 M T Catagramma pyracmon isaura J. Zikán, 1937 ref.213 M T Catagramma pyracmon peristera Hewitson, 1853 ref.213 M T Catagramma pyracmon piraya (Dillon, 1948) ref.213 M T Catagramma pyracmon pujoli Oberthür, 1916 ref.213 M T Archaeogramma Attal & Viloria, 2014 Archaeogramma claritae Costa, 2014 ref.188 Diaethria Billberg, 1820 Diaethria anna (Guérin-Méneville, [1844]) Anna’s Eighty-eight iN Diaethria anna anna (Guérin-Méneville, [1844]) E Mexico to Costa Rica M T S L1 L2 I Diaethria anna mixteca J. Maza, 1977 SW Mexico M T S Ab L Diaethria anna salvadorensis (Franz & Schröder, 1954) SW Chiapas to El Salvador M T S Diaethria astala (Guérin-Méneville, [1844]) Astala Eighty-eight iN Diaethria astala astala (Guérin-Méneville, [1844]) E Mexico to Colombia M T S L1 L2 Diaethria astala asteroide R. G. Maza & R. F. Maza, 1985 SW Mexico M T Diaethria asteria (Godman & Salvin, 1894) West-Mexican Eighty-eight W Mexico iN M T S Diaethria bacchis (E. Doubleday, 1849) White-patched Eighty-eight E & W Mexico & Guatemala iN M T S L Diaethria candrena (Godart, [1824]) iN Diaethria candrena candrena (Godart, [1824]) TL: Brazil M T Diaethria candrena longfieldae (Talbot, 1928) TL: Brazil (MT) M T Diaethria ceryx (Hewitson, 1864) iN M T Diaethria clymena (Cramer, 1775) iN Diaethria clymena clymena (Cramer, 1775) TL: Guyana M T Diaethria clymena aurelia (Guenée, 1872) TL: ? M T Diaethria clymena beleses (Godman & Salvin, 1889) TL: Panama M T Diaethria clymena bourcieri (Guenée, 1872) TL: Ecuador M T Diaethria clymena colombiana (Viette, 1958) TL: Colombia M T Diaethria clymena consobrina (Guérin-Méneville, [1844]) TL: Colombia M T Diaethria clymena dodone (Guenée, 1872) TL: Colombia M T Diaethria clymena janeira (C. Felder, 1862) TL: Brazil (RJ) M T Diaethria clymena juani Neild, 1996 TL: Venezuela M T Diaethria clymena marchalii (Guérin-Méneville, [1844]) Nicaragua to Brazil M T S L Diaethria clymena meridionalis (H. Bates, 1864) TL: Brazil (RS) M T Diaethria clymena peruviana (Guenée, 1872) TL: Peru M T Diaethria clymena seropina (Röber, 1924) TL: Brazil (Pa) Diaethria eluina (Hewitson, [1855]) iN Diaethria eluina eluina (Hewitson, [1855]) TL: [Brazil (MG)] M T Diaethria eluina lidwina (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Diaethria eluina splendidus (Oberthür, 1916) TL: Brazil (Go) M T Diaethria euclides (Latreille, [1809]) iN Diaethria euclides euclides (Latreille, [1809]) TL: Peru Diaethria euclides artemis (Röber, 1915) TL: Colombia M T Diaethria euclides gueneei (Röber, 1915) TL: Ecuador M T Diaethria euclides metiscus (E. Doubleday, 1849) TL: Venezuela M T Diaethria euclides phlogea (Salvin & Godman, 1868) TL: Colombia M T Diaethria euclides phlogeides (Röber, 1915) TL: Colombia M T Diaethria gabaza (Hewitson, [1855]) iN Diaethria gabaza gabaza (Hewitson, [1855]) TL: Colombia M T Diaethria gabaza eupepla (Salvin & Godman, 1868) Guatemala to Colombia M T TI Diaethria gabaza gabazina (Oberthür, 1916) TL: Colombia M T Diaethria kolyma (Hewitson, 1852) iN Diaethria kolyma kolyma (Hewitson, 1852) TL: Ecuador M T Diaethria kolyma pasithea (Hewitson, 1864) TL: Ecuador Diaethria neglecta (Salvin, 1869) iN Diaethria neglecta neglecta (Salvin, 1869) TL: Bolivia; Peru; Ecuador; Colombia; "Guatemala" M T S Diaethria neglecta merida (Honrath, 1884) TL: Venezuela M T Diaethria nystographa (Guenée, 1872) iN Diaethria nystographa nystographa (Guenée, 1872) TL: Ecuador M T Diaethria nystographa aliciae Neild, 1996 TL: Venezuela M T Diaethria nystographa charis (Oberthür, 1916) TL: Colombia M T Diaethria nystographa panthalis (Honrath, 1884) TL: Venezuela M T Diaethria nystographa perezi Lamas, 1995 TL: Peru Diaethria pandama (E. Doubleday, [1848]) Orange-striped Eighty-eight E & W Mexico to Panama iN M T S L Perisama E. Doubleday, 1849 Perisama barnesi Schaus, 1913 Costa Rican Eighty-eight Costa Rica (Carrillo Belt, Atlantic slope, to Talamancas); ref.12,213 M T S Perisama vaninka (Hewitson, [1855]) ref.12 iN Perisama vaninka vaninka (Hewitson, [1855]) ref.12,213 Perisama vaninka delafuentei (Neild, 1996) ref.12 Perisama vaninka doris (C. Felder & R. Felder, 1861) ref.12,213 M S Perisama vaninka gonalia Fruhstorfer, 1916 ref.12,213 Perisama vaninka volara (Hewitson, 1868) ref.12,213 Perisama vaninka [n. ssp.] Lamas, MS Colombia Perisama cardases (Hewitson, 1869) ref.12,375,213 iN M T Perisama campaspe (Hewitson, 1869) ref.12,375,213 iN M T Perisama diotima (Hewitson, 1852) ref.12,375,213 iN M T S Perisama cecidas (Hewitson, 1869) ref.12,375,213 iN M T Perisama alicia (Hewitson, 1868) iN M T S Perisama antioquia Attal & Crosson, 1996 iN M T Perisama bomplandii (Guérin-Méneville, [1844]) iN Perisama bomplandii bomplandii (Guérin-Méneville, [1844]) TL: Colombia M T S Perisama bomplandii albipennis A. Butler, 1873 TL: Peru M T Perisama bomplandii equatorialis (Guenée, 1872) TL: Ecuador M T Perisama bomplandii parabomplandii Dognin, 1899 TL: Colombia M T Perisama bomplandii reyi Attal & Crosson, 2003 ref.159 Perisama bomplandii velasteguii Lamas, 1995 TL: Ecuador M T Perisama cabirnia (Hewitson, 1874) iN M T Perisama calamis (Hewitson, 1869) iN M T S Perisama camelita (Hewitson, [1876]) iN M T S Perisama canoma H. Druce, 1874 iN Perisama canoma canoma H. Druce, 1874 TL: Peru M T S Perisama canoma cotyora (Hewitson, 1874) TL: Bolivia M T Perisama canoma cyaneis Attal & Crosson, 1997 TL: Ecuador M T Perisama canoma lamasi Descimon & Mast, 1995 TL: Peru M S Perisama canoma magaly Attal & Crosson, 1999 TL: Ecuador M T Perisama canoma thryoessa Fruhstorfer, 1916 TL: Peru M T Perisama clisithera (Hewitson, 1874) iN M T S Perisama cloelia (Hewitson, 1868) TL: Ecuador M T Perisama comnena (Hewitson, 1868) iN Perisama comnena comnena (Hewitson, 1868) TL: Bolivia M T S Perisama comnena attali Neild, 1996 TL: Venezuela M T Perisama comnena bertha Oberthür, 1916 TL: Colombia M T Perisama comnena bolivar Attal & Crosson, 1996 TL: Venezuela M T Perisama comnena comnenoides Oberthür, 1916 TL: Peru M T Perisama comnena divergens A. Butler, 1873 TL: Ecuador M T Perisama comnena ismaeli Attal & Crosson, 1996 TL: Ecuador Perisama dorbignyi (Guérin-Méneville, [1844]) iN Perisama dorbignyi dorbignyi (Guérin-Méneville, [1844]) TL: Colombia M T S Perisama dorbignyi aldasi Attal & Crosson, 1996 TL: Ecuador M T Perisama dorbignyi aristoteles Oberthür, 1916 TL: Ecuador M T Perisama dorbignyi jurinei (Guenée, 1872) TL: Peru M T S Perisama dorbignyi klugi Descimon & Mast, 1995 TL: Peru M S Perisama dorbignyi mola Dognin, 1899 TL: Ecuador M T Perisama dorbignyi paula Oberthür, 1916 TL: Colombia M T Perisama euriclea (E. Doubleday, [1847]) TL: Venezuela M T Perisama gisco Godman & Salvin, 1880 iN M T L Perisama goeringi H. Druce, 1875 TL: Venezuela M T Perisama hazarma (Hewitson, 1877) TL: Ecuador M T S Perisama humboldtii (Guérin-Méneville, [1844]) iN Perisama humboldtii humboldtii (Guérin-Méneville, [1844]) TL: Colombia M T Perisama humboldtii intermedia Oberthür, 1916 TL: Ecuador M T Perisama humboldtii tringa (Guenée, 1872) TL: Peru M T S Perisama humboldtii tristis Oberthür, 1916 TL: Venezuela M T Perisama humboldtii [n. ssp.] Lamas, MS Colombia Perisama lanice (Hewitson, 1868) TL: Ecuador Perisama lanice lanice (Hewitson, 1868) TL: Ecuador M T Perisama lanice latimargo Oberthür, 1916 TL: Peru M T Perisama lanice picteti (Guenée, 1872) TL: Peru M T Perisama lebasii (Guérin-Méneville, [1844]) iN Perisama lebasii lebasii (Guérin-Méneville, [1844]) TL: Colombia M T Perisama lebasii ambatensis Oberthür, 1916 TL: Ecuador M T Perisama lebasii astuta Dognin, 1899 TL: Colombia M T Perisama lebasii hilara Salvin, 1869 TL: Peru M T S Perisama lebasii koenigi Descimon & Mast, 1995 TL: Peru M T Perisama lebasii selva Attal & Crosson, 1997 TL: Ecuador M T Perisama morona (Hewitson, 1868) iN Perisama morona morona (Hewitson, 1868) TL: Bolivia M T Perisama morona moronina Röber, 1915 TL: Peru M T Perisama nevada Attal & Crosson, 1996 iN M T Ma Perisama oppelii (Latreille, [1809]) iN Perisama oppelii oppelii (Latreille, [1809]) TL: Peru M T S Perisama oppelii amalia Oberthür, 1916 TL: Colombia Perisama oppelii angustifasciata Röber, 1924 TL: Colombia Perisama oppelii bleuzeni Attal & Crosson, 1996 TL: Venezuela M T Perisama oppelii cristal Attal & Crosson, 1996 TL: Ecuador M T S Perisama oppelii eminens Oberthür, 1881 TL: Peru M T Perisama oppelii erebina Oberthür, 1916 TL: Colombia M T Perisama oppelii iphigenia Oberthür, 1916 TL: Bolivia M T Perisama oppelii viridinota A. Butler, 1873 TL: Peru M T Perisama oppelii xanthica (Hewitson, 1868) TL: Peru M T Perisama ouma Dognin, 1891 iN M T S Perisama paralicia Fruhstorfer, 1916 iN Perisama paralicia paralicia Fruhstorfer, 1916 TL: Peru M T S Perisama paralicia bonita Attal & Crosson, 1997 TL: Ecuador M T S Perisama patara (Hewitson, [1855]) TL: Colombia Perisama patara patara (Hewitson, [1855]) TL: Colombia Perisama patara arletta Descimon & Mast, 1995 TL: Peru M S Perisama patara gallardi Attal & Crosson, 1996 TL: Colombia M T Perisama philinus E. Doubleday, 1849 iN Perisama philinus philinus E. Doubleday, 1849 TL: Bolivia M T S Perisama philinus descimoni Mast, 1995 TL: Peru M S Perisama philinus nyctimene (Hewitson, 1868) TL: Ecuador M T S Perisama philinus saussurei (Guenée, 1872) TL: Brazil Perisama satanas Attal & Crosson, 1996 TL: Colombia M T Perisama tristrigosa A. Butler, 1873 iN M T S Perisama tryphena (Hewitson, [1857]) iN Perisama tryphena tryphena (Hewitson, [1857]) TL: Colombia M T S Perisama tryphena affinis Crosson & Attal, 1995 TL: Peru M T Perisama tryphena asormani Attal & Crosson, 1996 TL: Venezuela M T Perisama tryphena cauca Attal & Crosson, 1996 TL: Colombia M T Perisama tryphena fernandezi Attal & Crosson, 1996 TL: Venezuela Perisama tryphena pallida Attal & Crosson, 1996 TL: Ecuador M T Perisama tryphena pastaza Attal & Crosson, 1996 TL: Ecuador Perisama typhania Oberthür, 1916 iN M T Perisama vitringa (Hewitson, 1858) iN Perisama vitringa vitringa (Hewitson, 1858) TL: [Peru] M T Perisama vitringa huallaga Attal & Crosson, 1996 TL: Peru M T S Perisama vitringa ochreipennis A. Butler, 1873 TL: Bolivia M T S Perisama yeba (Hewitson, [1857]) iN Perisama yeba yeba (Hewitson, [1857]) TL: Colombia M T Perisama yeba emma Oberthür, 1916 TL: Venezuela M T Perisama yeba guerini C. Felder & R. Felder, 1867 TL: Colombia M T Perisama yeba johannes Oberthür, 1916 TL: Ecuador M T Perisama yeba malvina Kretzschmar, [1894] TL: Colombia M T Perisama yeba nimbala Attal & Crosson, 1996 TL: Ecuador M T Perisama [n. sp.] Lamas, MS Peru Subfamily Cyrestinae Tribe Cyrestini Marpesia Hübner, 1818 Marpesia berania (Hewitson, 1852) iN Marpesia berania berania (Hewitson, 1852) TL: Ecuador; Brazil (Am) M T S Marpesia berania fruhstorferi (Seitz, 1914) Honduras to Panama M T S L Marpesia chiron (Fabricius, 1775) Many-banded Daggerwing S TX (strays N), E & W Mexico to Argentina, Greater Antilles iN Marpesia chiron chiron (Fabricius, 1775) TL: [Haiti] M T S HS L1 L2 Marpesia chiron chironides (Staudinger, 1886) TL: Cuba M T Marpesia chiron marius (Cramer, 1779) TL: Surinam Marpesia corinna (Latreille, [1813]) iN M T S Marpesia corita (Westwood, 1850) Orange-banded Daggerwing iN Marpesia corita corita (Westwood, 1850) E & S Mexico, Guatemala, Honduras M T S L Ha Marpesia corita phiale (Godman & Salvin, 1878) SW Mexico (S Chiapas) & W Guatemala M T S L Marpesia crethon (Fabricius, 1776) iN M T S Marpesia egina (H. Bates, 1865) iN M T Marpesia eleuchea Hübner, 1818 Caribbean Daggerwing iN Marpesia eleuchea eleuchea Hübner, 1818 Cuba & Isla de Juventud, rare stray to S FL Keys M S Marpesia eleuchea bahamensis Munroe, 1971 Bahamas M T S Marpesia eleuchea dospassosi Munroe, 1971 Hispaniola M T S Marpesia eleuchea pellenis (Godart, [1824]) Jamaica M S L Marpesia furcula (Fabricius, 1793) iN Marpesia furcula Northern segregate , [= "iole" records for Central America], Nicaragua to Panama M L Marpesia furcula furcula (Fabricius, 1793) TL: Jamaica; India Marpesia furcula violetta (A. Hall, 1929) TL: Ecuador M T Marpesia furcula oechalia (Westwood, 1850) TL: Bolivia M T Marpesia furcula [n. ssp.] Lamas, MS Panama Marpesia harmonia (Klug, 1836) Pale Daggerwing SE Mexico & Guatemala iN M S L Marpesia livius (W. F. Kirby, 1871) iN Marpesia livius livius (W. F. Kirby, 1871) TL: Ecuador M T S Marpesia livius alcibiades (Staudinger, 1876) Costa Rica & Panama M T S Marpesia marcella (C. Felder & R. Felder, 1861) iN Marpesia marcella marcella (C. Felder & R. Felder, 1861) TL: Colombia; "Ecuador" M T S Marpesia marcella valetta (A. Butler & H. Druce, 1872) Guatemala to Costa Rica M T S Marpesia marcella [n. ssp.] Lamas, MS Panama Marpesia merops (Doyère, [1840]) Spot-banded Daggerwing Guatemala to Brazil iN M T S L1 L2 Marpesia orsilochus (Fabricius, 1776) iN M S Marpesia petreus (Cramer, 1776) Ruddy Daggerwing iN Marpesia petreus Northern segregate , S FL, S TX, E & W Mexico (strays far N) to W Ecuador M S1 S2 L1 L2 L3 I Marpesia petreus petreus (Cramer, 1776) Guianas, Amazonian Colombia to Bolivia M T S L Marpesia petreus damicorum Brévignon, 2001 Mona, Puerto Rico, Lesser Antilles (St. Kitts, Guadeloupe, etc.) M S L Marpesia petreus rheophila Lamas, 1995 TL: Peru Marpesia petreus [n. ssp.] Lamas, MS Mexico Marpesia themistocles (Fabricius, 1793) iN Marpesia themistocles themistocles (Fabricius, 1793) TL: ? Marpesia themistocles norica (Hewitson, 1852) TL: [Brazil (Am)] M T S Marpesia tutelina (Hewitson, 1852) iN M T Marpesia zerynthia Hübner, [1823] iN Marpesia zerynthia zerynthia Hübner, [1823] TL: ? M T Marpesia zerynthia dentigera (Fruhstorfer, 1907) [= coresia], E & S Mexico to Ecuador, one-time stray to S TX M T S L1 L2 Marpesia pantepuiana Costa, Attal & Viloria, 2016 ref.189 Subfamily Nymphalinae Tribe Coeini Pycina E. Doubleday, [1849] Pycina zamba E. Doubleday, [1849] iN Pycina zamba zamba E. Doubleday, [1849] TL: Venezuela M T S Pycina zamba zelys Godman & Salvin, 1884 S Mexico to Panama M T S Historis Hübner, [1819] Historis acheronta (Fabricius, 1775) Tailed Cecropian iN Historis acheronta acheronta (Fabricius, 1775) continental; S TX & AZ (as stray), E & W Mexico to Brazil M T S L1 L2 I Historis acheronta cadmus (Cramer, 1775) Jamaica M OI Historis acheronta semele (M. Bates, 1939) Cuba and Hispaniola M S Historis odius (Fabricius, 1775) Orion Cecropian iN Historis odius odius (Fabricius, 1775) Greater Antilles M T S L Historis odius caloucaera Brévignon, 2003 Windward and Leeward Islands M OI Historis odius dious Lamas, 1995 continental; S TX (as stray), E & W Mexico to Argentina M T S L Baeotus Hemming, 1939 Baeotus aeilus (Stoll, 1780) iN M T S Baeotus beotus (E. Doubleday, [1849]) Graphic Beauty SE Mexico to Amazon Basin iN M T S1 S2 L I Baeotus deucalion (C. Felder & R. Felder, 1860) iN M T S Baeotus japetus (Staudinger, [1885]) iN M T S Tribe Nymphalini Colobura Billberg, 1820 Colobura annulata Willmott, Constantino & J. Hall, 2001 New Beauty S Mexico to W Ecuador, Venezuela to Bolivia & Guianas, Trinidad iN M T S L I Colobura dirce (Linnaeus, 1758) Dirce Beauty iN Colobura dirce dirce (Linnaeus, 1758) E & W Mexico to Argentina M T S HS L I Colobura dirce wolcotti W. Comstock, 1942 Greater Antilles (Jamaica, Cuba, Hispaniola, Puerto Rico) M T S L Tigridia Hübner, [1819] Tigridia acesta (Linnaeus, 1758) iN Tigridia acesta Northern segregate , SE Mexico to Panama M S L I Tigridia acesta acesta (Linnaeus, 1758) TL: Indiis Tigridia acesta columbina (Neustetter, 1929) TL: Colombia M T Tigridia acesta fulvescens (A. Butler, 1873) TL: Peru M T Tigridia acesta latifascia (A. Butler, 1873) TL: Colombia M T Tigridia acesta ochracea (Bryk, 1953) TL: Peru Tigridia acesta tapajona (A. Butler, 1873) TL: Brazil (Pa) M T Tigridia acesta [n. ssp.#1] Lamas, MS TL: Mexico (Chia) Tigridia acesta [n. ssp.#2] Lamas, MS TL: Venezuela Tigridia acesta [n. ssp.#3] Lamas, MS TL: Bolivia Smyrna Hübner, [1823] Smyrna blomfildia (Fabricius, 1781) iN Smyrna blomfildia blomfildia (Fabricius, 1781) TL: Brazil M T Smyrna blomfildia datis Fruhstorfer, 1908 S TX, E & W Mexico to Panama M T S L I Smyrna karwinskii Geyer, [1833] Karwinski’s Beauty E & W Mexico to Nicaragua iN M S L Hypanartia Hübner, [1821] Hypanartia bella (Fabricius, 1793) iN M T Hypanartia celestia Lamas, Willmott & J. Hall, 2001 TL: Peru M T S Hypanartia charon (Hewitson, 1878) TL: Ecuador M T L Hypanartia christophori Jasinski, 1998 iN M S Hypanartia cinderella Lamas, Willmott & J. Hall, 2001 iN M T L Hypanartia dione (Latreille, [1813]) iN Hypanartia dione dione (Latreille, [1813]) TL: [Colombia] M T S Hypanartia dione arcaei (Salvin, 1871) Costa Rica & Panama M T S L Hypanartia dione disjuncta Willmott, J. Hall & Lamas, 2001 E & W Mexico, Guatemala, Honduras M T S L Hypanartia fassli Willmott, J. Hall & Lamas, 2001 TL: Colombia M T Hypanartia godmanii (H. Bates, 1864) Godman's Mapwing E & W Mexico to Colombia iN M T S L Hypanartia kefersteini (E. Doubleday, [1847]) iN M T Hypanartia lethe (Fabricius, 1793) Orange Mapwing S TX (rare stray), E & S Mexico to Brazil iN Hypanartia lethe lethe (Fabricius, 1793) TL: Indiis M S L Hypanartia lethe rosamariae Neild, 2008 ref.266 Hypanartia lindigii (C. Felder & R. Felder, 1862) iN M T L Hypanartia paullus (Fabricius, 1793) Antillean Mapwing Greater Antilles (Jamaica, Cuba, Hispaniola, Puerto Rico) iN M T S L Hypanartia splendida Rothschild, 1903 TL: Peru M T Hypanartia trimaculata Willmott, J. Hall & Lamas, 2001 iN Hypanartia trimaculata trimaculata Willmott, J. Hall & Lamas, 2001 TL: Colombia M T Hypanartia trimaculata autumna Willmott, J. Hall & Lamas, 2001 [Mexican "kefersteini" records], E & S Mexico to Panama M T S L Hypanartia trimaculata fountaineae Neild, 2008 ref.266 Vanessa Fabricius, 1807 Vanessa altissima (Rosenberg & Talbot, 1914) iN M T S Vanessa annabella (W. D. Field, 1971) West Coast Lady iN M T S Ab L1 L2 I Ha Vanessa atalanta (Linnaeus, 1758) Red Admiral iN Vanessa atalanta rubria (Fruhstorfer, 1909) Gr. Antilles (not PR), N Bahamas (strays), N Am. to Guat., Venez. M T S Ab L1 L2 L3 I H Vanessa braziliensis (Moore, 1883) iN M T Vanessa cardui (Linnaeus, 1758) Painted Lady nearly cosmopolitan as vagrant; uncommon or vagrant in W. Indies iN M T S L1 L2 L3 I Vanessa carye (Hübner, [1812]) iN M T Vanessa myrinna (E. Doubleday, 1849) iN M T HS Vanessa tameamea Escholtz, 1821 Kameamea Endemic to Hawaii iN M S L Ha Vanessa terpsichore (Philippi, 1859) iN M S L Vanessa virginiensis (Drury, 1773) American Lady N Bahamas, Gr. Antilles, Hawaii, Canary Is.; N. Am. to Colombia iN M T S Ab L1 L2 L3 L4 I H Vanessa amerindica J. Miller & F. Brown, 1989 a fossil from Colorado Aglais Dalman, 1816 Aglais io (Linnaeus, 1758) TL: Sweden Aglais milberti (Godart, 1819) Milbert's Tortoiseshell iN Aglais milberti viola dos Passos, 1938 TL: Canada (NL) M S Aglais milberti milberti (Godart, 1819) TL: USA (Pa) M S1 S2 L Aglais milberti subpallida (T. Cockerell, 1889) TL: USA (Co) M S1 S2 L I Ha Aglais milberti pullum (Austin, 1998) TL: USA (Az) M T S L Nymphalis Kluk, 1780 Nymphalis l-album (Esper, 1781) Compton Tortoiseshell iN Nymphalis l-album j-album (Boisduval & Le Conte, [1835]) TL: USA (In) M S L Nymphalis l-album watsoni (G. Hall, 1924) TL: Canada (BC) M S Nymphalis californica (Boisduval, 1852) California Tortoiseshell iN M T S1 S2 L I Ha1 Ha2 Nymphalis antiopa (Linnaeus, 1758) Mourning Cloak iN Nymphalis antiopa antiopa (Linnaeus, 1758) TL: Sweden; America M T S1 S2 S3 Ab L1 L2 L3 I H Txt Nymphalis cyanomelas (E. Doubleday, [1848]) Mexican Tortoiseshell S Mexico to El Salvador M T S Polygonia Hübner, [1819] Polygonia interrogationis (Fabricius, 1798) Question Mark iN M T S1 S2 L1 L2 L3 L4 I1 I2 H F Polygonia comma (T. Harris, 1841) Comma Anglewing iN M S1 S2 L1 L2 Polygonia g-argenteum (E. Doubleday, 1848) Mexican Anglewing Mexico iN M T S L Polygonia satyrus (W. H. Edwards, 1869) Satyr Anglewing iN Polygonia satyrus satyrus (W. H. Edwards, 1869) TL: USA (Co) M T S1 S2 S3 L1 L2 I H Polygonia satyrus neomarsyas dos Passos, 1969 [= transcanada] M T S1 S2 L Ha Polygonia progne (Cramer, 1775) Gray Anglewing iN M T S1 S2 L Polygonia oreas (W. H. Edwards, 1869) Oreas Anglewing iN Polygonia oreas oreas (W. H. Edwards, 1869) TL: USA (Ca) M S1 S2 Polygonia oreas silenus (W. H. Edwards, 1870) TL: USA (Or) M S L Ha Polygonia oreas threatfuli C. Guppy & J. Shepard, 2001 TL: Canada (BC) M S1 S2 Polygonia oreas satellow Scott, 2006 TL: USA (Co) M S Polygonia oreas nigrozephyrus Scott, 1984 TL: USA (Co) M S Polygonia gracilis (Grote & Robinson, 1867) Hoary Anglewing iN Polygonia gracilis gracilis (Grote & Robinson, 1867) TL: USA (NH) M S1 S2 L Polygonia gracilis zephyrus (W. H. Edwards, 1870) TL: USA (Nv) M T S1 S2 L1 L2 L3 L4 I1 I2 H Ha Polygonia haroldii (Dewitz, 1877) Spotless Anglewing Mexico iN M T S1 S2 L Polygonia faunus (W. H. Edwards, 1862) Green Anglewing iN Polygonia faunus smythi A. Clark, 1937 TL: USA (Va) M S1 S2 Polygonia faunus faunus (W. H. Edwards, 1862) TL: USA (NY) M S1 S2 L Polygonia faunus arcticus Leussler, 1935 [= cenveray] M T S1 S2 L Polygonia faunus rusticus (W. H. Edwards, 1874) TL: USA (Ca) M S L Polygonia faunus fulvescens J. Emmel, T. Emmel & Mattoon, 1998 TL: USA (Ca) M T S1 S2 Polygonia faunus hylas (W. H. Edwards, 1872) TL: USA (Co) M S1 S2 L Jupitellia , [] a fossil genus Jupitellia charon (Scudder, 1889) Fossil from Colorado Tribe Victorinini Anartia Hübner, [1819] Anartia amathea (Linnaeus, 1758) Red Peacock iN Anartia amathea amathea (Linnaeus, 1758) Panama to Argentina; Grenada, Barbados, Antigua M S Hy HS L Anartia amathea roeselia (Eschscholtz, 1821) TL: Brazil M T Anartia amathea sticheli Fruhstorfer, 1907 TL: Bolivia M T Anartia amathea [n. ssp.] Lamas, MS Lesser Antilles Anartia chrysopelea Hübner, [1831] Cuban Peacock Cuba & Isla de Juventud, rare stray to S FL iN M T S Anartia fatima (Fabricius, 1793) Banded Peacock iN Anartia fatima fatima (Fabricius, 1793) S TX, E & S Mexico to far NW Colombia M T S Hy L1 L2 I H Anartia fatima colima Lamas, 1995 NW Mexico M T S L Anartia jatrophae (Linnaeus, 1763) White Peacock iN Anartia jatrophae jatrophae (Linnaeus, 1763) South America, Lesser Antilles N to Barbuda & St. Kitts M T S1 S2 HS L Anartia jatrophae guantanamo Munroe, 1942 S FL (strays N), Bahamas, Cuba, Turks & Caicos M T S L1 L2 Anartia jatrophae intermedia Munroe, 1942 St. Croix M T Anartia jatrophae jamaicensis Möschler, 1886 Jamaica, Grand Cayman, [Cayman Brac?] M S1 S2 L Anartia jatrophae luteipicta Fruhstorfer, 1907 S TX (strays far N), E & W Mexico through Central America M T S Ab Ma L1 L2 L3 L4 I H Anartia jatrophae saturata Staudinger, [1885] Hispaniola, Great Inagua M T S Anartia jatrophae semifusca Munroe, 1942 Puerto Rico, Mona, E to Virgin Islands M T S L Anartia lytrea (Godart, 1819) Hispaniolan Peacock Hispaniola (Dominican Republic); rare stray to Cuba iN M T S Metamorpha Hübner, [1819] Metamorpha elissa Hübner, [1819] Metamorpha iN Metamorpha elissa elissa Hübner, [1819] Panama to Guianas and Peru M S L Metamorpha elissa alberti Neild, 2008 ref.266 Metamorpha elissa pulsitia R. Fox & A. Forbes, 1971 TL: Bolivia M T Napeocles H. Bates, 1864 Napeocles jucunda (Hübner, [1808]) iN Napeocles jucunda jucunda (Hübner, [1808]) TL: ? M T Napeocles jucunda caesari Neild, 2008 ref.266 Siproeta Hübner, [1823] Siproeta epaphus (Latreille, [1813]) Rusty-tipped Page iN Siproeta epaphus epaphus (Latreille, [1813]) S TX, S NM (rare strays), E & W Mexico to Peru M T S L1 L2 I H Siproeta epaphus gadoui Masters, 1967 TL: Venezuela M T Siproeta epaphus trayja Hübner, [1823] TL: Brazil M L Siproeta stelenes (Linnaeus, 1758) Malachite iN Siproeta stelenes stelenes (Linnaeus, 1758) Jamaica, Hispaniola, Puerto Rico, St. Kitts, St. Croix M S1 S2 HS L Siproeta stelenes biplagiata (Fruhstorfer, 1907) S TX (strays N), E & W Mexico to Amazon, S FL, Cuba, Caymans M T S L1 L2 I H Siproeta stelenes meridionalis (Fruhstorfer, 1909) TL: Brazil (RS) M T S L Siproeta stelenes sophene (Fruhstorfer, 1907) TL: Ecuador M T Siproeta superba (H. Bates, 1864) Broad-banded Page iN Siproeta superba superba (H. Bates, 1864) S Mexico, Guatemala, Honduras M T S L Siproeta superba euoe R. Fox & A. Forbes, 1971 Costa Rica M T S Tribe Junoniini Hypolimnas Hübner, [1819] Hypolimnas misippus (Linnaeus, 1764) Mimic NE S. Am, Gr. & Lesser Antilles, N continental and Yucatan coasts (as st ... iN M T S HS Junonia Hübner, [1819] Junonia coenia Hübner, [1822] Northern Buckeye E USA (strays N), Bermuda, Cuba & Isla de Juventud iN M T S1 S2 Ab L1 L2 L3 L4 L5 L6 L7 I H Junonia grisea Austin & J. Emmel, 1998 Northern Buckeye W USA (strays N & NE), Baja California to SW Oregon to W TX iN M T S L1 L2 Ha Junonia nigrosuffusa W. Barnes & McDunnough, 1916 Dark Buckeye SW USA & W TX to SW Mexico (strays N) iN M T S L Junonia stemosa Grishin, 2020 Twintip buckeye C & S TX (strays N); ref.6 iN M S1 S2 L1 L2 Junonia zonalis C. Felder & R. Felder, 1867 Tropical Buckeye iN Junonia zonalis zonalis C. Felder & R. Felder, 1867 S FL, Bahamas, Cuba, Cayman Is., Jamaica, Hispaniola; C. Am.? M S IL L Junonia zonalis michaelisi Fruhstorfer, 1907 Puerto Rico M T L Junonia zonalis swifti Brévignon, 2004 Guadeloupe and nearby islands; possibly other I. in Less. Antilles M S Junonia pacoma Grishin, 2020 Pacific mangrove buckeye Son to Jal & BCS, coastal; ref.6,16 iN M S L Junonia neildi Brévignon, 2004 Mangrove Buckeye Guadeloupe; possibly elsewhere in Lesser Antilles iN Junonia neildi varia Grishin, 2020 Gulf Coast mangroves, S TX to Belize (at least), ? along Pacific; ref.6 M S L Junonia neildi Caribbean segregate , S FL, Bahamas, Greater Antilles, Cayman Islands M S1 S2 L Junonia neildi neildi Brévignon, 2004 Guadeloupe; possibly elsewhere in Lesser Antilles M S Junonia litoralis Brévignon, 2009 Litoral Buckeye French Guiana & Trinidad iN Junonia wahlbergi Brévignon, 2008 Wahlberg's Buckeye French Guiana & Trinidad Junonia evarete (Cramer, 1779) Tropical Buckeye iN Junonia evarete evarete (Cramer, 1779) NE South America (at least) M T OI L Junonia evarete flirtea (Fabricius, 1793) TL: India Junonia evarete fuscescens (A. Butler, 1901) TL: Ecuador M L Junonia evarete huacapistana W. Forbes, [1929] TL: Peru Junonia evarete lima W. Forbes, [1929] TL: Peru Junonia evarete oscura Neild, 2008 ref.266 M T Junonia evarete occidentalis C. Felder & R. Felder, 1862 TL: Brazil (Am) Junonia evarete [n. ssp.] Lamas, MS Mexico (Oax) Junonia divaricata C. Felder & R. Felder, 1867 iN Junonia genoveva (Cramer, 1780) Mangrove Buckeye iN Junonia genoveva genoveva (Cramer, 1780) NE South America, possibly Central America M T OI Junonia genoveva constricta C. Felder & R. Felder, 1867 TL: Colombia M T L Junonia genoveva hilaris C. Felder & R. Felder, 1867 TL: Paraguay Junonia genoveva incarnata C. Felder & R. Felder, 1867 TL: Colombia M T Junonia genoveva infuscata C. Felder & R. Felder, 1867 TL: Brazil (Ba) M T Junonia genoveva vivida W. Forbes, [1929] TL: Guyana; Surinam M T Junonia genoveva [n. ssp.#1] Lamas, MS TL: Mexico (Oax) Junonia genoveva [n. ssp.#2] Lamas, MS TL: Peru Junonia vestina C. Felder & R. Felder, 1867 iN Junonia vestina vestina C. Felder & R. Felder, 1867 TL: ? Junonia vestina livia Fruhstorfer, 1912 TL: Bolivia M T Tribe Melitaeini Euphydryas Scudder, 1872 Euphydryas gillettii (W. Barnes, 1897) Gillett's Checkerspot iN M S1 S2 L I Ha Euphydryas editha (Boisduval, 1852) Edith's Checkerspot iN Euphydryas editha taylori (W. H. Edwards, 1888) TL: Canada (BC) M S1 S2 S3 S4 Ab L Euphydryas editha editha (Boisduval, 1852) TL: USA (Ca) M T S1 S2 L Euphydryas editha insularis T. Emmel & J. Emmel, 1974 TL: USA (Ca) M T Euphydryas editha augustina (W. G. Wright, 1905) TL: USA (Ca) M T S1 S2 L I Euphydryas editha quino (Behr, 1863) TL: USA (Ca) M T S1 S2 L I H Ha Euphydryas editha ehrlichi Baughman & Murphy, 1998 TL: USA (Ca) M S Euphydryas editha nubigena (Behr, 1863) TL: USA (Ca) M T S L Ha Euphydryas editha tahoensis Austin & Murphy, 1998 TL: USA (Nv) M S L Euphydryas editha aurilacus Gunder, 1928 TL: USA (Ca) M S L Euphydryas editha lawrencei Gunder, 1931 TL: USA (Or) M T S1 S2 L Ha Euphydryas editha colonia (W. G. Wright, 1905) TL: USA (Or) M T S L Euphydryas editha rubicunda (H. Edwards, 1881) TL: USA (Ca) M S1 S2 L H Euphydryas editha baroni (W. H. Edwards, 1879) TL: USA (Ca) M S Ab L I Euphydryas editha karinae Baughman & Murphy, 1998 TL: USA (Ca) M S Euphydryas editha luestherae Murphy & P. Ehrlich, 1980 TL: USA (Ca) M T S I Euphydryas editha monoensis Gunder, 1928 TL: USA (Ca) M S Ab Euphydryas editha edithana (Strand, 1915) TL: USA (Ca) M T S L Ha Euphydryas editha C Oregon segregate , Pine Mtn. area, Deschutes Co., OR M S L Ha Euphydryas editha Ochoco Mts. segregate , vic. Big Summit Prairie, Ochoco Mts., OR M S Euphydryas editha NE Oregon segregate , Blue Mountains, NE Oregon M S L Euphydryas editha bingi Baughman & Murphy, 1998 S end Warner Mts., CA, high elevation M S L Euphydryas editha mattooni Austin & Murphy, 1998 TL: USA (Ca) M T1 T2 T3 Euphydryas editha owyheensis Austin & Murphy, 1998 TL: USA (Nv) M T1 T2 Ha Euphydryas editha lehmani Gunder, 1929 TL: USA (Nv) M T S Ha Euphydryas editha gunnisonensis F. Brown, [1971] TL: USA (Co) M S1 S2 Euphydryas editha alebarki Ferris, [1971] TL: USA (Wy) M T S1 S2 S3 L Ha Euphydryas editha montanus McDunnough, 1928 TL: USA (Wy) M S L Euphydryas editha hutchinsi McDunnough, 1928 TL: USA (Mt) M S Euphydryas editha beani (Skinner, 1897) TL: Canada (Ab) M S Euphydryas chalcedona (E. Doubleday, [1847]) Chalcedon Checkerspot iN Euphydryas chalcedona klotsi dos Passos, 1938 TL: USA (Az) M S L1 L2 I Euphydryas chalcedona kingstonensis T. Emmel & J. Emmel, 1973 TL: USA (Ca) M T S1 S2 L Euphydryas chalcedona corralensis T. Emmel & J. Emmel, 1973 TL: USA (Ca) M T S1 S2 L H Ha Euphydryas chalcedona hennei Scott, 1981 TL: USA (Ca) M T S1 S2 L1 L2 L3 L4 I H Ha Euphydryas chalcedona sierra (W. G. Wright, 1905) TL: USA (Ca) M T S1 S2 L H Euphydryas chalcedona olancha (W. G. Wright, 1905) TL: USA (Ca) M T S Ha Euphydryas chalcedona macglashanii (Rivers, 1888) TL: USA (Ca) M S1 S2 Euphydryas chalcedona dwinellei (H. Edwards, 1881) TL: USA (Ca) M S1 S2 S3 Ab Euphydryas chalcedona trinitina Baughman & Murphy, 1998 TL: USA (Ca) M L I Ha Euphydryas chalcedona chalcedona (E. Doubleday, [1847]) [= sinecat] M T1 T2 S1 S2 S3 Ab L1 L2 L3 L4 L5 I Ha Euphydryas colon (W. H. Edwards, 1881) Snowberry Checkerspot iN Euphydryas colon colon (W. H. Edwards, 1881) TL: USA (Wa) M S Ab L1 L2 Ha Euphydryas colon svihlae Pelham, 2008 TL: USA (Wa) M S Euphydryas colon E-central OR Cascades segregate , M L Ha Euphydryas colon SE Cascades, Warner Mts. segregate , [nr. wallacensis] M S1 S2 L1 L2 L3 Ha Euphydryas colon wallacensis Gunder, 1928 TL: USA (Id) M T S1 S2 S3 Ab L Ha Euphydryas colon nevadensis Bauer, 1975 TL: USA (Nv) M S Ha Euphydryas colon central WA Cascades segregate , [often called paradoxa in literature] C WA Cascades from C Yakima to far S Chelan Co. M L1 L2 L3 I Ha Euphydryas colon paradoxa McDunnough, 1927 TL: Canada (BC) M S Euphydryas anicia (E. Doubleday, [1847]) Anicia Checkerspot iN Euphydryas anicia helvia (Scudder, 1869) TL: USA (Ak) M S1 S2 Euphydryas anicia anicia (E. Doubleday, [1847]) TL: Canada (Ab) M S1 S2 Euphydryas anicia howlandi D. Stallings & Turner, 1947 TL: USA (Mt) M T S1 S2 Euphydryas anicia hopfingeri Gunder, 1934 TL: USA (Wa) M S Euphydryas anicia N WA Cascades segregate , M Ab L Ha Euphydryas anicia NE OR, high elevations segregate , [nr. howlandi] M S Euphydryas anicia bakeri D. Stallings & Turner, 1945 TL: USA (Or) M T S1 S2 Euphydryas anicia effi D. Stallings & Turner, 1945 TL: USA (Id) M T S1 S2 Euphydryas anicia windi Gunder, 1932 TL: USA (Wy) M S Ab L Ha Euphydryas anicia maria (Skinner, 1899) TL: USA (Ut) M S1 S2 Euphydryas anicia brucei (W. H. Edwards, 1888) TL: USA (Co) M T S L1 L2 L3 Euphydryas anicia eurytion (Mead, 1875) TL: USA (Co) M T S Ab L1 L2 H Euphydryas anicia wecoeut M. Fisher, Spomer & Scott, 2006 TL: USA (Co) M T S Euphydryas anicia alena W. Barnes & Benjamin, 1926 TL: USA (Ut) M S L Euphydryas anicia magdalena W. Barnes & McDunnough, 1918 TL: USA (Az) M S L1 L2 L3 H Euphydryas anicia hermosa (W. G. Wright, 1905) TL: USA (Az) M T S L1 L2 I Euphydryas anicia chuskae (Ferris & R. Holland, 1980) TL: USA (NM) M T S Euphydryas anicia carmentis W. Barnes & Benjamin, 1926 TL: USA (Co) M S1 S2 Euphydryas anicia cloudcrofti (Ferris & R. Holland, 1980) TL: USA (NM) M T S L Euphydryas anicia capella (W. Barnes, 1897) TL: USA (Co) M S1 S2 S3 L Euphydryas anicia bernadetta Leussler, 1920 [= rorina] M T S L1 L2 L3 L4 Ha Euphydryas anicia veazieae Fender & Jewett, 1953 TL: USA (Or) M S1 S2 Ab L Ha Euphydryas anicia macyi Fender & Jewett, 1953 TL: USA (Or) M S1 S2 Euphydryas anicia wheeleri (H. Edwards, 1881) TL: USA (Nv) M S L Ha Euphydryas anicia morandi (Gunder, 1928) TL: USA (Nv) M T S Euphydryas anicia variicolor Baughman & Murphy, 1998 TL: USA (Ca) M S L Euphydryas phaeton (Drury, 1773) Baltimore Checkerspot iN Euphydryas phaeton borealis F. Chermock & R. Chermock, 1940 TL: USA (Me) Euphydryas phaeton phaeton (Drury, 1773) TL: USA (NY) M T S Ab L1 L2 I H Euphydryas phaeton clarki Pavulaan, 2021 TL: USA (Md) M T Euphydryas phaeton ozarkae Masters, 1968 TL: USA (Mo) M S1 S2 Ab L Poladryas Bauer, 1961 Poladryas arachne (W. H. Edwards, 1869) Arachne Checkerspot iN Poladryas arachne arachne (W. H. Edwards, 1869) TL: USA (Co) M T S1 S2 S3 L Ha Poladryas arachne gilensis (W. Holland, 1930) TL: USA (Az) M T S Poladryas arachne expedita Austin, 1998 TL: USA (Nv) M T I Poladryas arachne nympha (W. H. Edwards, 1884) TL: USA (Az) M T S Ab L1 L2 Poladryas arachne monache (J. A. Comstock, 1918) TL: USA (Ca) M T S Poladryas minuta (W. H. Edwards, 1861) Dotted Checkerspot iN Poladryas minuta minuta (W. H. Edwards, 1861) [= simador]; far SE CO, E NM, N & W TX, NE Mexico M T S I Poladryas minuta NW Mexican segregate , NW Mexico (SON, CHIH, DGO) M S Chlosyne A. Butler, 1870 Chlosyne gaudialis (H. Bates, 1864) Gaudy Checkerspot iN Chlosyne gaudialis gaudialis (H. Bates, 1864) SE Mexico to Panama M T S L Chlosyne gaudialis wellingi L. Miller & Rotger, 1979 S Oaxaca (Sierra Madre del Sur) M S Chlosyne narva (Fabricius, 1793) Narva Checkerspot iN Chlosyne narva narva (Fabricius, 1793) Nicaragua to Colombia and Venezuela M S Chlosyne narva bonpland (Latreille, [1809]) Costa Rica to W Ecuador M T S1 S2 L Chlosyne janais (Drury, 1782) Crimson-patch Checkerspot iN Chlosyne janais janais (Drury, 1782) C & S TX, T & S Mexico to Colombia M S L1 L2 L3 L4 I H Chlosyne janais gloriosa Bauer, 1960 W Mexico M T S1 S2 L Chlosyne janais marianna Röber, [1914] S-C Mexico (Balsas Basin) M T S L Chlosyne ehrenbergii (Geyer, [1833]) White-rayed Checkerspot Mexico iN M S L Chlosyne definita (E. Aaron, [1885]) Definite Checkerspot iN Chlosyne definita definita (E. Aaron, [1885]) S NM, S & W TX, NE Mexico M T S L I H Chlosyne definita dolosa Grishin, 2023 NW Mexico; ref.151 Chlosyne anastasia (Hemming, 1934) Definite Checkerspot C & NW Mexico; ref.151 M T S L Chlosyne eumeda (Godman & Salvin, 1894) Eumeda Checkerspot W Mexico (Nayarit to Guerrero) iN Chlosyne eumeda NW Mexican segregate , Sonora & Sinaloa, NW Mexico M S L I Chlosyne eumeda eumeda (Godman & Salvin, 1894) TL: Mexico (Gue) M T S L Chlosyne marina (Geyer, 1837) Marina Checkerspot Sonora, Chihuahua iN Chlosyne marina marina (Geyer, 1837) TL: Mexico M T S Chlosyne marina SW Mexico segregate , Guerrero and Oaxaca M S L Chlosyne marina Chiapas segregate , Chiapas M S Chlosyne melitaeoides (C. Felder & R. Felder, 1867) Melitaeoides Checkerspot S TX (as vagrant), E Mexico iN M T S L Chlosyne erodyle (H. Bates, 1864) Erodyle Checkerspot iN Chlosyne erodyle Oaxaca segregate , Oaxaca M S L Chlosyne erodyle erodyle (H. Bates, 1864) Chiapas to NW Costa Rica M T S L Chlosyne poecile (C. Felder & R. Felder, 1867) Poecile Checkerspot SW Costa Rica to Venezuela iN Chlosyne poecile poecile (C. Felder & R. Felder, 1867) TL: Colombia M T S L Chlosyne poecile rubrigutta Röber, [1914] TL: Colombia M T Chlosyne poecile rodriguezi Neild, 2008 TL: Venezuela Chlosyne poecile intermedia Neild, 2008 TL: Venezuela Chlosyne poecile murphyi Neild, 2008 TL: Venezuela Chlosyne pardelina Grishin, 2023 Banded Checkerspot [infrasub. of Higgins 1960] S TX, NE Mexico; ref.151 M T S L H Chlosyne endeis (Godman & Salvin, 1894) Banded Checkerspot W Mexico iN M T S Chlosyne melanarge (H. Bates, 1864) Cream-banded Checkerspot E Mexico to Costa Rica iN M T S L Chlosyne hippodrome (Geyer, 1837) Simple Checkerspot iN Chlosyne hippodrome hippodrome (Geyer, 1837) E & W Mexico to Colombia M S L1 L2 Chlosyne hippodrome fabricii Higgins, 1960 TL: Indiis Chlosyne rosita A. Hall, 1924 Rosita Checkerspot iN Chlosyne rosita rosita A. Hall, 1924 Chiapas, Guatemala, El Salvador M T S1 S2 S3 S4 Chlosyne rosita browni Bauer, 1961 S TX (as stray), E Mexico M T S L I Chlosyne rosita mazarum L. Miller & Rotger, 1979 TL: Mexico (Mor) Chlosyne rosita montana A. Hall, 1924 C and NW Mexico, SE AZ (as stray) M T S Ab L Chlosyne rosita riobalsensis Bauer, 1961 SW Mexico (Rio Balsas basin) [mazarum = form of riobalsensis] M T S1 S2 L Ha Chlosyne perlula (C. Felder & R. Felder, 1861) Perula Checkerspot Venezuela & Colombia iN M T S Chlosyne ezra (Hewitson, 1864) Ezra Checkerspot Costa Rica & Panama iN M T S L Chlosyne theona (Ménétriés, 1855) Theona Checkerspot iN Chlosyne theona costaricensis (Austin & M. Smith, 1998) NW Costa Rica, SW Nicaragua M S L Chlosyne theona theona (Ménétriés, 1855) Chiapas to Nicaragua M S L Chlosyne theona minimus (Austin & M. Smith, 1998) Yucatan Peninsula to Honduras M S L Chlosyne theona mullinsi (Austin & M. Smith, 1998) SW & C Mexico M S L I Chlosyne theona brocki (Austin & M. Smith, 1998) NW Mexico (Sonora to N Nayarit) M S L I Chlosyne theona thekla (W. H. Edwards, 1870) far NW Mexico (N Sonora, SE AZ, SW NM M T S L1 L2 I H Chlosyne theona hondana (Weymer, 1890) TL: Colombia M T Chlosyne bollii (W. H. Edwards, [1878]) Theona Checkerspot C Texas, NE Mexico; ref.151 M T S Ab L1 L2 L3 I H Chlosyne chinatiensis (Tinkham, 1944) Theona Checkerspot W TX, N-C Mexico (Dur); ref.11,151 iN M S I Chlosyne cyneas (Godman & Salvin, 1878) iN M T S1 S2 Hy L1 L2 L3 I H Chlosyne cynisca (Godman & Salvin, 1882) Oaxacan Checkerspot Oaxaca, Puebla iN M T S L Chlosyne fulvia (W. H. Edwards, 1879) Fulvia Checkerspot W NE, W KS, E CO, E NM, W TX, far NE Mexico; ref.11 iN M T S1 S2 L F Chlosyne coronado (M. Smith & Brock, 1988) Fulvia Checkerspot AZ, SW NM, far NW Mexico; ref.11 iN Chlosyne coronado pariaensis (M. Smith & Brock, 1988) S-central UT, N-central AZ; ref.11 M T S L I Ha Chlosyne coronado coronado (M. Smith & Brock, 1988) AZ, SW NM, far NW Mexico; ref.11 M T S L1 L2 L3 I H Chlosyne leanira (C. Felder & R. Felder, 1860) Leanira Checkerspot iN Chlosyne leanira oregonensis Bauer, 1975 TL: USA (Or) M S L Ha Chlosyne leanira leanira (C. Felder & R. Felder, 1860) TL: USA (Ca) M T S L I Chlosyne leanira obsoleta (H. Edwards, 1877) TL: USA (Ca) M T S Chlosyne leanira nebularum (Austin & M. Smith, 1998) TL: USA (Ca) M T S L Chlosyne leanira elegans (Priestaf & J. Emmel, 1998) TL: USA (Ca) M T S L1 I F Ha Chlosyne leanira wrightii (W. H. Edwards, 1886) TL: USA (Ca) M T S L1 L2 I H Chlosyne leanira alma (Strecker, [1878]) TL: USA (Az) M T S L1 L2 I1 I2 H Ha Chlosyne leanira basinensis (Austin & M. Smith, 1998) TL: USA (Ca) M T S Ha Chlosyne leanira flavodorsalis (Austin & M. Smith, 1998) TL: USA (Ut) M T S Chlosyne leanira austrima (Austin & M. Smith, 1998) TL: Mexico (BCN) M T S Chlosyne nycteis (E. Doubleday, [1847]) Silvery Checkerspot iN Chlosyne nycteis pastoron Pelham, 2008 [= obsoleta] M T L Chlosyne nycteis nycteis (E. Doubleday, [1847]) TL: USA M T OI S1 S2 L1 L2 L3 L4 L5 L6 L7 I F Chlosyne nycteis reversa (F. Chermock & R. Chermock, 1940) TL: Canada (Mt) M S Chlosyne nycteis drusius (W. H. Edwards, 1884) TL: USA (Co) M S1 S2 L1 L2 Ha Chlosyne gorgone (Hübner, [1810]) Gorgone Checkerspot iN Chlosyne gorgone gorgone (Hübner, [1810]) TL: ? M T S Chlosyne gorgone carlota (Reakirt, 1866) TL: USA (Mo) M T S Ab HS L1 L2 L3 I F Chlosyne californica (W. G. Wright, 1905) California Patch iN M T S L1 L2 I H Chlosyne lacinia (Geyer, 1837) Bordered Patch iN Chlosyne lacinia lacinia (Geyer, 1837) C Mexico to Panama M T S L1 L2 Chlosyne lacinia adjutrix Scudder, 1875 NE Mexico, S & C Texas (strays N) M T S Ab L1 L2 L3 L4 L5 I1 I2 H Chlosyne lacinia crocale (W. H. Edwards, 1874) NW Mexico, SW USA (strays N) M T S L1 L2 I Chlosyne lacinia saundersi (E. Doubleday, [1847]) TL: Venezuela M T L Chlosyne kendallorum Opler, 1999 Kendall's Checkerspot NE Mexico iN M T S Chlosyne harrisii (Scudder, 1863) Harris's Checkerspot iN Chlosyne harrisii harrisii (Scudder, 1863) TL: USA (Me/NH/Ma) M S L I Ha Chlosyne harrisii albimontana (Avinoff, 1930) TL: USA (NH) M S Chlosyne harrisii liggetti (Avinoff, 1930) TL: USA (Pa) M S Chlosyne harrisii hanhami (Fletcher, 1903) TL: USA (Mn) M S Chlosyne hoffmanni (Behr, 1863) Hoffmann's Checkerspot iN Chlosyne hoffmanni hoffmanni (Behr, 1863) TL: USA (Ca) M T S L1 L2 Chlosyne hoffmanni segregata (W. Barnes & McDunnough, 1918) TL: USA (Or) M T S L Ha Chlosyne hoffmanni manchada Bauer, 1960 TL: USA (Wa) M T L Ha Chlosyne whitneyi (Behr, 1863) Sierra Nevada Checkerspot iN M T S L Chlosyne acastus (W. H. Edwards, 1874) Sagebrush Checkerspot iN Chlosyne acastus sabina (W. G. Wright, 1905) TL: USA (Az) M T S L I Chlosyne acastus neumoegeni (Skinner, 1895) TL: USA (Ut) M T S L1 L2 L3 I H Ha Chlosyne acastus acastus (W. H. Edwards, 1874) TL: USA (Ut) M S I Chlosyne acastus arkanyon M. Fisher & Scott, 2006 TL: USA (Co) M T S1 S2 L1 L2 Ha Chlosyne acastus vallismortis (J. Johnson, 1938) TL: USA (Ca) M T S Chlosyne acastus robusta Austin, 1998 TL: USA (Nv) M T S Chlosyne acastus waucoba J. Emmel, T. Emmel & Mattoon, 1998 TL: USA (Ca) M T Chlosyne acastus far N Great Basin segregate , [nr. acastus] M S Chlosyne acastus dorothyi Bauer, 1975 TL: USA (Or) M S Chlosyne gabbii (Behr, 1863) Gabb’s Checkerspot iN Chlosyne gabbii gabbii (Behr, 1863) TL: USA (Ca) M T S L1 L2 I Ha Chlosyne gabbii atrifasciata Hawks & J. Emmel, 1998 TL: USA (Ca) Chlosyne palla (Boisduval, 1852) Northern Checkerspot iN Chlosyne palla sterope (W. H. Edwards, 1870) NW US & SW Can; ref.16 M T L Ha Chlosyne palla E Columbia Gorge segregate , M S L Ha Chlosyne palla E-central OR Cascades segregate , M L Ha Chlosyne palla eremita (W. G. Wright, 1905) TL: USA (Ca) M T S L Ha Chlosyne palla palla (Boisduval, 1852) TL: USA (Ca) M S L Ha Chlosyne palla altasierra J. Emmel, T. Emmel & Mattoon, 1998 TL: USA (Ca) M T L Chlosyne palla pola (Boisduval, 1869) ref.20 M T Chlosyne palla australomontana J. Emmel, T. Emmel & Mattoon, 1998 TL: USA (Ca) M T S L1 L2 L3 Chlosyne flavula (W. Barnes & McDunnough, 1918) Northern Checkerspot ref.20 Chlosyne flavula blackmorei Pelham, 2008 TL: not specified M S L I Ha Chlosyne flavula flavula (W. Barnes & McDunnough, 1918) TL: USA (Co) M S1 S2 L1 L2 L3 L4 Ha Chlosyne flavula calydon (W. Holland, 1931) TL: USA (Co) M S Chlosyne damoetas (Skinner, 1902) Rockslide Checkerspot iN Chlosyne damoetas damoetas (Skinner, 1902) TL: USA (Co) M S1 S2 L1 L2 I Ha Chlosyne damoetas windriver Scott, 1998 TL: USA (Wy) M S L Chlosyne damoetas altalus Scott, 1998 TL: not specified M T Microtia H. Bates, 1864 Microtia elvira Grishin, 2023 Elfoid S AZ & W Mexico; ref.146 M T S L I Microtia elva H. Bates, 1864 Elf ref.146 iN Microtia elva elva H. Bates, 1864 S TX (as stray), E Mexico to Costa Rica M T S1 S2 L Microtia elva horni Rebel, 1906 S Mexico & Guatemala M T S L Microtia elva bogotana Grishin, 2024 E Colombia; ref.154 M T Microtia dymas (W. H. Edwards, 1877) Tiny Checkerspot ref.12 iN Microtia dymas dymas (W. H. Edwards, 1877) C TX to NE Mexico M T S L1 L2 I1 I2 I3 Microtia dymas chara (W. H. Edwards, [1884]) S AZ, S NM, W TX, NW Mexico, Baja California Sur M T S L1 L2 I H Microtia dymas imperialis Bauer, 1959 S CA, Baja California Norte M T S L Ha Microtia perse (W. H. Edwards, 1882) Arizona Checkerspot S AZ, SW NM, N Sonora, NW Chihuahua; ref.12 iN M T S L I H Microtia elada (Hewitson, 1868) Elada Checkerspot ref.12 iN Microtia elada ulrica (W. H. Edwards, 1877) C & W TX, NE Mexico M T S L1 L2 L3 I H Microtia elada elada (Hewitson, 1868) C & S Mexico M T S L Microtia elada hepburni (Godman, 1901) NW Mexico M T S L Microtia anomalus (Godman & Salvin, 1897) Anomalus Checkerspot W Mexico (Nay to Mich); ref.12 iN Microtia anomalus anomalus (Godman & Salvin, 1897) TL: Mexico (Col) M T S Microtia coracara (Dyar, 1912) Coracara Checkerspot SW Mexico (Mor, Gue, Oax); ref.12 iN M T S L Gnathotriche C. Felder & R. Felder, 1862 Gnathotriche exclamationis (Kollar, 1850) iN Gnathotriche exclamationis exclamationis (Kollar, 1850) TL: Venezuela M T Gnathotriche exclamationis elaea (Hewitson, 1869) TL: Ecuador M T Gnathotriche exclamationis sodalis Staudinger, 1885 TL: Colombia M T Gnathotriche mundina (H. Druce, 1876) iN Gnathotriche mundina mundina (H. Druce, 1876) TL: Peru M T Gnathotriche mundina erebia (Röber, [1914]) TL: Peru M T Gnathotriche mundina fallax (Staudinger, 1885) TL: Peru M T Gnathotriche mundina hopfferi (Röber, [1914]) TL: [Bolivia] M T Gnathotriche mundina rosina (Dognin, 1888) TL: Ecuador M T Gnathotriche mundina steinii (Dewitz, 1877) TL: Colombia M T Higginsius Hemming, 1964 Higginsius fasciata (Hopffer, 1874) iN M T Higginsius miriam (Dognin, 1888) TL: Ecuador M T Antillea Higgins, 1959 Antillea pelops (Drury, 1773) Antillean Crescent iN Antillea pelops pelops (Drury, 1773) Hispaniola, Puerto Rico, St. Kitts M T S L Antillea pelops anocaona (Herrich-Schäffer, 1865) Cuba M T S Antillea pelops pygmaea (Godart, 1819) Jamaica M T S Antillea proclea (E. Doubleday, [1847]) Jamaican Crescent Jamaica iN M T S L Atlantea Higgins, 1959 Atlantea cryptadia Sommer & A. Schwartz, 1980 Hispaniolan Checkerspot Hispaniola M T S Atlantea pantoni (Kaye, 1906) Jamaican Checkerspot Jamaica iN M T S L Atlantea perezi (Herrich-Schäffer, 1862) Cuban Checkerspot NE Cuba iN M T S L Atlantea tulita (Dewitz, 1877) Puerto Rican Checkerspot Puerto Rico iN M S L I Phystis Higgins, 1981 Phystis simois (Hewitson, 1864) ref.12 iN M T Phystis pratti (A. Hall, 1935) ref.12 Phystis pratti pratti (A. Hall, 1935) TL: Peru M T Phystis pratti chinchipensis (Hayward, 1964) TL: Peru M T Phystis variegata (Röber, 1913) ref.12 M T Phystis fontus (A. Hall, 1928) ref.12 Phystis fontus fontus (A. Hall, 1928) TL: Guyana M T Phystis fontus schmitzorum (Austin & O. Mielke, 1993) TL: Brazil (Rd) M T Mazia Higgins, 1981 Mazia amazonica (H. Bates, 1864) iN Mazia amazonica amazonica (H. Bates, 1864) TL: Brazil (Pa) M T S Mazia amazonica cocha Lamas, 1995 TL: Peru Mazia amazonica tambopata Lamas, 1995 TL: Peru Tisona Higgins, 1981 Tisona saladillensis (Giacomelli, 1911) TL: Argentina Tisona saladillensis saladillensis (Giacomelli, 1911) TL: Argentina M T Tisona saladillensis clarior Higgins, 1981 TL: Bolivia M T Ortilia Higgins, 1981 Ortilia gentina Higgins, 1981 iN M T Ortilia liriope (Cramer, 1775) iN M S Notilia Grishin, 2021 Notilia dicoma (Hewitson, 1864) ref.12 iN M T Notilia orthia (Hewitson, 1864) ref.12 iN M T Notilia orticas (Schaus, 1902) ref.12 iN Notilia orticas orticas (Schaus, 1902) TL: Brazil (Pr) M T L Notilia orticas zamora (A. Hall, 1917) TL: Venezuela M T Notilia polinella (A. Hall, 1928) ref.12 M T Notilia sejona (Schaus, 1902) ref.12 iN M T Notilia velica (Hewitson, 1864) ref.12 iN Notilia velica velica (Hewitson, 1864) TL: ? M T Notilia velica durnfordi (Godman & Salvin, 1878) TL: Argentina M T Levinata Grishin, 2021 Levinata levina (Hewitson, 1872) ref.12 iN M T Anthanassa Scudder, 1875 Anthanassa acesas (Hewitson, 1864) iN Anthanassa acesas acesas (Hewitson, 1864) TL: Venezuela M T Anthanassa acesas castianira (Godman & Salvin, 1880) TL: Colombia M T Anthanassa annulata Higgins, 1981 iN M T Anthanassa ardys (Hewitson, 1864) Ardys Crescent iN Anthanassa ardys ardys (Hewitson, 1864) E & W Mexico through Oaxaca M T S L Anthanassa ardys [n. ssp.] Lamas, MS Nicaragua to Colombia M S L Anthanassa subota (Godman & Salvin, 1878) Cryptic Crescent Chiapas & Guatemala iN Anthanassa subota [n. ssp.] Lamas, MS W Mexico (Nayarit to Oaxaca) M S Anthanassa subota subota (Godman & Salvin, 1878) TL: Guatemala M T S Anthanassa argentea (Godman & Salvin, 1882) Chestnut Crescent S TX (as stray), E & S Mexico, Guatemala, Nicaragua iN M T S L Anthanassa atronia (H. Bates, 1866) Brown Crescent E & S Mexico to Panama iN M T S L Anthanassa crithona (Salvin, 1871) Orange-banded Crescent Nicaragua to Panama iN M T S L Anthanassa dracaena (C. Felder & R. Felder, 1867) iN Anthanassa dracaena dracaena (C. Felder & R. Felder, 1867) TL: Colombia M T Anthanassa dracaena phlegias (Godman, 1901) S Mexico to Costa Rica M T S L Anthanassa drusilla (C. Felder & R. Felder, 1861) Orange-patched Crescent iN Anthanassa drusilla drusilla (C. Felder & R. Felder, 1861) Panama, Colombia, Venezuela, Ecuador M T S L Anthanassa drusilla alceta (Hewitson, 1869) TL: Ecuador M T L Anthanassa drusilla halli Neild, 2008 ref.266 Anthanassa drusilla forbesi Neild, 2008 ref.266 Anthanassa drusilla higginsi Neild, 2008 ref.266 Anthanassa drusilla lelex (H. Bates, 1864) S TX (rare stray), E & S Mexico to Panama M T S L Anthanassa drusilla verena (Hewitson, 1864) TL: Bolivia M T Anthanassa drymaea (Godman & Salvin, 1878) Weak-banded Crescent S Mexico (Chiapas), Guatemala, Honduras iN M T S L Anthanassa frisia (Poey, 1832) Cuban Crescent S FL (strays N), N Bahamas, Cuba, Hispaniola, Jamaica iN M T S L Anthanassa hermas (Hewitson, 1864) iN Anthanassa hermas dubia (A. Hall, 1929) TL: Venezuela M T Anthanassa hermas hermas (Hewitson, 1864) TL: [Brazil (Pe)] M T Anthanassa hermas taeniata (Röber, 1913) TL: Peru M T L Anthanassa tulcis (H. Bates, 1864) Pale-banded Crescent SE AZ (as stray), S TX, E & W Mexico to Panama iN M T S L1 L2 L3 I Anthanassa nebulosa (Godman & Salvin, 1878) Nebulosa Crescent iN Anthanassa nebulosa nebulosa (Godman & Salvin, 1878) Chiapas & Guatemala M T S Anthanassa nebulosa alexon (Godman & Salvin, 1889) [= subconcolor], E & W Mexico M T S L Anthanassa otanes (Hewitson, 1864) Blackened Crescent iN Anthanassa otanes otanes (Hewitson, 1864) [= sopolis], Chiapas, W Guatemala to Costa Rica; also W Mexico? M T S L Anthanassa otanes cyno (Godman & Salvin, 1889) E Mexico M T S Anthanassa otanes fulviplaga (A. Butler, 1872) Costa Rica & Panama M T S L Anthanassa otanes oaxaca Beutelspacher, 1990 vic. Sierra de Juarez, Oaxaca, Mexico M T S L Anthanassa ptolyca (H. Bates, 1864) Darkened Crescent iN Anthanassa ptolyca E Mexican segregate , S TX (rare stray), E Mexico M S L Anthanassa ptolyca ptolyca (H. Bates, 1864) Chiapas to Venezuela M T S L Anthanassa ptolyca amator (A. Hall, 1929) W Mexico M T S L Anthanassa sitalces (Godman & Salvin, 1882) Montane Crescent iN Anthanassa sitalces sitalces (Godman & Salvin, 1882) Chiapas & Guatemala M T S Anthanassa sitalces cortes (A. Hall, 1917) W Mexico to Oaxaca M T S L Anthanassa sosis (Godman & Salvin, 1878) Sosis Crescent [= dora], Costa Rica to Colombia iN M T S L Anthanassa texana (W. H. Edwards, 1863) Texan Crescent AZ to TX (strays far N), Baja, E & W Mexico to Guatemala iN M T S Ab L I H Anthanassa seminole (Skinner, 1911) Texan Crescent SC to FL, E to LA & far E TX; ref.146 M S L H Castilia Higgins, 1981 Castilia angusta (Hewitson, 1868) iN M T Castilia castilla (C. Felder & R. Felder, 1862) iN Castilia castilla castilla (C. Felder & R. Felder, 1862) TL: Colombia M T S Castilia castilla occidentalis (Fassl, 1912) TL: Colombia Castilia chiapaensis (Beutelspacher, 1990) Chiapas Crescent S Mexico (Chiapas) iN M T S Castilia chinantlensis (R. F. Maza, 1978) Chinantlan Crescent SE Mexico (Sierra de Juarez, Oaxaca) iN M T S Castilia eranites (Hewitson, 1857) Mimic Crescent S Mexico to Colombia & Venezuela iN M T S L Castilia fulgora (Godman & Salvin, 1878) Fulgora Crescent Costa Rica (Atlantic slope, Cordillera Central) & W Panama iN M T TI L Castilia griseobasalis (Röber, 1913) Gray-based Crescent E & W Mexico to Costa Rica iN M T S L Castilia guaya (A. Hall, 1929) TL: Ecuador M T Castilia myia (Hewitson, [1864]) Mayan Crescent E & W Mexico to Panama iN M T S L Castilia neria (Hewitson, 1869) iN M T Castilia nortbrundii (Weeks, 1901) TL: Bolivia M T Castilia ofella (Hewitson, [1864]) White-dotted Crescent S Mexico, Guatemala to Colombia, Venezuela, Trinidad iN M S L1 L2 Castilia perilla (Hewitson, 1852) iN M T L Telenassa Higgins, 1981 Telenassa abas (Hewitson, 1864) iN M T Telenassa berenice (C. Felder & R. Felder, 1862) iN Telenassa berenice berenice (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T S Telenassa berenice drusinilla (Röber, 1913) TL: Argentina M T Telenassa delphia (C. Felder & R. Felder, 1861) iN Telenassa delphia delphia (C. Felder & R. Felder, 1861) TL: Colombia M T Telenassa delphia alani Neild & Orellana, 2008 ref.266 Telenassa delphia austini Neild, 2008 ref.266 Telenassa delphia catenaria (Godman & Salvin, 1880) TL: Colombia M T Telenassa delphia elaphina (Röber, 1913) TL: Bolivia M T Telenassa delphia flavocincta (Dognin, 1887) TL: Ecuador M T Telenassa delphia gaujoni (Dognin, 1887) TL: Ecuador M T Telenassa delphia nana (H. Druce, 1874) TL: Peru M T Telenassa delphia nussia (H. Druce, 1876) TL: Peru M T Telenassa delphia trimaculata (Hewitson, 1869) TL: Ecuador M T L Telenassa jana (C. Felder & R. Felder, 1867) iN M T Telenassa notus (A. Hall, 1917) TL: Peru M T Telenassa sepultus (A. Hall, 1928) TL: Peru M T Telenassa teletusa (Godart, [1824]) iN Telenassa teletusa teletusa (Godart, [1824]) TL: Brazil M T Telenassa teletusa burchelli (Moulton, 1909) TL: Brazil (Go) M T Ithra Grishin, 2021 Ithra ithra (W. F. Kirby, 1900) ref.12 iN M T Dagon Higgins, 1981 Dagon catula (Hopffer, 1874) iN M T Dagon morena (Röber, 1913) iN M T Dagon pusilla (Salvin, 1869) iN M T Eresia Boisduval, 1836 Eresia actinote Salvin, 1869 iN Eresia actinote actinote Salvin, 1869 TL: Peru M T Eresia actinote limbata (Röber, 1913) TL: Bolivia M T Eresia actinote [n. ssp.] Lamas, MS Peru Eresia carme E. Doubleday, [1847] iN Eresia carme carme E. Doubleday, [1847] TL: Venezuela M T Eresia carme mejiai Neild & De Sousa, 2008 ref.266 Eresia carme judithae Neild, 2008 ref.266 Eresia casiphia Hewitson, 1869 iN M T Eresia clio (Linnaeus, 1758) Clio Crescent iN Eresia clio clio (Linnaeus, 1758) [= clara], S Mexico to Brazil & Bolivia M T S L Eresia clio carmenae Neild, 2008 ref.266 Eresia clio reducta (A. Hall, 1929) TL: Ecuador M T Eresia datis Hewitson, [1864] iN Eresia datis datis Hewitson, [1864] TL: [Peru] M T Eresia datis corybassa Hewitson, 1874 TL: Bolivia M T Eresia datis fassli (Röber, 1913) TL: Colombia M T Eresia datis fenestrata (Röber, 1913) TL: Peru M T Eresia datis magniplaga (Röber, 1927) TL: Colombia M T Eresia datis manto (A. Hall, 1930) TL: Colombia M T Eresia datis margaretha Hewitson, 1872 TL: Colombia M T Eresia datis moesta Salvin & Godman, 1868 TL: Ecuador M T L Eresia datis phaedima Salvin & Godman, 1868 TL: Peru M T Eresia datis [n. ssp.] Lamas, MS Colombia Eresia emerantia Hewitson, 1857 iN Eresia emerantia emerantia Hewitson, 1857 TL: Colombia M T Eresia emerantia coela H. Druce, 1874 Costa Rica & NE Panama M T S L Eresia emerantia sestia Hewitson, 1869 TL: Ecuador M T Eresia erysice (Geyer, 1832) TL: S. Amer. Eresia erysice erysice (Geyer, 1832) TL: S. Amer. Eresia erysice etesiae (A. Hall, 1928) TL: French Guiana M T Eresia estebana (A. Hall, 1929) TL: Venezuela Eresia estebana estebana (A. Hall, 1929) TL: Venezuela M T Eresia estebana camachoi Neild, 2008 ref.266 Eresia eunice (Hübner, [1807]) iN Eresia eunice eunice (Hübner, [1807]) TL: [Brazil] M T Eresia eunice drypetis Godman & Salvin, 1878 Panama, Colombia M T OI L Eresia eunice esora Hewitson, 1857 TL: Brazil M T Eresia eunice gudruna Röber, 1913 TL: [Brazil (Am)] M T Eresia eunice klagesii (Weeks, 1906) TL: Venezuela M T Eresia eunice mechanitis Godman & Salvin, 1878 Nicaragua to Panama M T L Eresia eunice polymnia (Röber, 1913) TL: Colombia Eresia eunice wallacei Neild, 2008 ref.266 Eresia eunice [n. ssp.] Lamas, MS Peru Eresia ithomioides Hewitson, [1864] iN Eresia ithomioides ithomioides Hewitson, [1864] TL: Colombia M T Eresia ithomioides alsina Hewitson, 1869 Nicaragua, Costa Rica, W Panama M T S L Eresia ithomioides anomala Higgins, 1981 TL: Colombia M T Eresia ithomioides cissia (A. Hall, 1928) TL: Colombia M T Eresia ithomioides eutropia Hewitson, 1874 [= melania], Costa Rica, Panama, W Colombia M T S L Eresia ithomioides mimas (Staudinger, 1885) TL: Colombia M T Eresia ithomioides poecilina H. Bates, 1866 Costa Rica & Panama M T S Eresia ithomioides quintilla Hewitson, 1872 TL: Ecuador M T Eresia ithomioides [n. ssp.] Lamas, MS Colombia Eresia lansdorfi (Godart, 1819) iN M T Eresia letitia Hewitson, 1869 iN Eresia letitia letitia Hewitson, 1869 TL: Ecuador M T Eresia letitia ocellata (Röber, 1913) TL: Peru M T Eresia nauplius (Linnaeus, 1758) iN Eresia nauplius nauplius (Linnaeus, 1758) TL: Indiis Eresia nauplius extensa (A. Hall, 1929) TL: Brazil (MT) M T Eresia nauplius plagiata (Röber, 1913) TL: Peru M T Eresia olivencia H. Bates, 1864 TL: Brazil (Am) M T Eresia pelonia Hewitson, 1852 iN Eresia pelonia pelonia Hewitson, 1852 TL: Ecuador M T L Eresia pelonia callonia (Staudinger, 1885) TL: Peru M T Eresia pelonia callonioides Strand, 1912 TL: Peru M T Eresia pelonia ithomiola Salvin, 1869 TL: Peru M T L Eresia perna Hewitson, 1852 iN Eresia perna perna Hewitson, 1852 TL: Brazil (RJ) M T Eresia perna aveyrona H. Bates, 1864 TL: Brazil (Pa) M T Eresia perna mylitta Hewitson, 1869 TL: Ecuador M T Eresia perna [n. ssp.] Lamas, MS Brazil Eresia phillyra Hewitson, 1852 Longwing Crescent iN Eresia phillyra phillyra Hewitson, 1852 E & S Mexico, Guatemala M T S L Eresia phillyra nigripennis Salvin, 1869 Costa Rica, [Panama?] M T OI Eresia polina Hewitson, 1852 iN Eresia polina polina Hewitson, 1852 TL: Ecuador M T Eresia polina joaquini Neild, 2008 ref.266 Eresia polina laias Godman & Salvin, 1879 TL: Colombia M T Eresia polina lycus (A. Hall, 1928) TL: Colombia M T Eresia polina oblita (Staudinger, 1885) TL: Venezuela M T Eresia sticta Schaus, 1913 Costa Rican Crescent Costa Rica (Carrillo belt, Atlantic slope), [Colombia?] iN M T S L Janatella Higgins, 1981 Janatella fellula (Schaus, 1902) iN M T Janatella hera (Cramer, 1779) iN M T Janatella leucodesma (C. Felder & R. Felder, 1861) Whitened Crescent Nicaragua to Venezuela & Trinidad iN M T S L Tegosa Higgins, 1981 Tegosa anieta (Hewitson, 1864) Black-bordered Crescent iN Tegosa anieta anieta (Hewitson, 1864) Nicaragua to Venezuela & Trinidad M T S L Tegosa anieta cluvia (Godman & Salvin, 1882) Chiapas & Guatemala M T S L Tegosa anieta lirina (Röber, 1913) TL: Bolivia M T Tegosa anieta luka Higgins, 1981 S TX (rare stray), E Mexico to Honduras M T S L Tegosa claudina (Eschscholtz, 1821) Claudina Crescent SE Mexico to Argentina iN M T OI S L Tegosa etia (Hewitson, 1868) iN M T Tegosa flavida (Hewitson, 1868) iN M T Tegosa fragilis (H. Bates, 1864) iN M T Tegosa guatemalena (H. Bates, 1864) Guatemalan Crescent E & W Mexico, Guatemala, Belize iN M T S L Tegosa infrequens Higgins, 1981 iN M T Tegosa nazaria (C. Felder & R. Felder, 1867) iN M T Tegosa nigrella (H. Bates, 1866) Nigrella Crescent iN Tegosa nigrella nigrella (H. Bates, 1866) S Mexico (Chiapas), Guatemala, Honduras M T S L Tegosa nigrella niveonotis (A. Butler & H. Druce, 1872) Costa Rica (Atlantic slope) M T S L Tegosa orobia (Hewitson, 1864) iN Tegosa orobia orobia (Hewitson, 1864) TL: ? M T Tegosa orobia ursula (Staudinger, 1894) TL: Bolivia M T Tegosa pastazena (H. Bates, 1864) iN M T Tegosa selene (Röber, 1913) iN M T Tegosa serpia Higgins, 1981 iN M T Tegosa tissoides (A. Hall, 1928) iN M T Phyciodes Hübner, [1819] Phyciodes graphica (R. Felder, 1869) Vesta Crescent [= vesta] iN Phyciodes graphica graphica (R. Felder, 1869) TL: Mexico (Oax) M T S Ma L1 L2 L3 I H Phyciodes graphica vesta (W. H. Edwards, 1869) TL: USA (Tx) M T Phyciodes pallescens (R. Felder, 1869) Mexican Crescent E & W Mexico, SE AZ as rare stray iN M T S L Phyciodes picta (W. H. Edwards, 1865) Painted Crescent iN Phyciodes picta picta (W. H. Edwards, 1865) TL: USA (Nb) M S L1 L2 Phyciodes picta canace W. H. Edwards, 1871 TL: USA (Az) M T S L1 L2 Phyciodes orseis W. H. Edwards, 1871 California Crescent iN Phyciodes orseis orseis W. H. Edwards, 1871 TL: USA (Ca) M S1 S2 L I Ha Phyciodes orseis Warner Mts. segregate , Warner Mts. (OR/CA) W to SE Siskiyou Co., CA M S Phyciodes orseis herlani Bauer, 1975 TL: USA (Nv) M T S L Phyciodes pallida (W. H. Edwards, 1864) Pale Crescent iN Phyciodes pallida pallida (W. H. Edwards, 1864) TL: USA (Co) M S L Ha Phyciodes pallida barnesi Skinner, 1897 TL: USA (Co) M S1 S2 Phyciodes mylitta (W. H. Edwards, 1861) Mylitta Crescent iN Phyciodes mylitta mylitta (W. H. Edwards, 1861) BC to MT, S to S CA M T S1 S2 L1 L2 Ha Phyciodes mylitta arizonensis Bauer, 1975 AZ, NM, W TX, N to E CO, far NW Mexico M S Ab L1 L2 I Phyciodes mylitta mexicana A. Hall, 1928 TL: Mexico (Ver) M T S L Ha Phyciodes mylitta thebais Godman & Salvin, 1878 [= arida] M T S L Phyciodes phaon (W. H. Edwards, 1864) Phaon Crescent iN Phyciodes phaon phaon (W. H. Edwards, 1864) E & C Mexico, TX to FL & VA (strays far N), Cuba, Grand Cayman; ref.16 M T S Ab L1 L2 L3 I F Phyciodes phaon maya A. Hall, 1928 Chiapas & Guatemala M T Phyciodes jalapeno Scott, 1998 Phaon Crescent NW Mexico & Baja (N), S CA, S AZ; ref.16 M T S L Ha Phyciodes orantain Scott, 1998 Pearl Crescent ref.16 M T S1 S2 S3 Ha Phyciodes tharos (Drury, 1773) Pearl Crescent iN Phyciodes tharos tharos (Drury, 1773) TL: [USA (NY)] M T S1 S2 S3 Ab L1 L2 L3 L4 L5 L6 L7 L8 Ha Phyciodes tharos distincta Bauer, 1975 CA, AZ Phyciodes tharos riocolorado Scott, 1992 TL: USA (Ut) M T S Phyciodes incognitus Gatrelle, 2004 Incognito Crescent ref.16 M T L Phyciodes diminutor Scott, 1998 Diminutor Crescent ref.16 M T L Phyciodes cocyta (Cramer, 1777) Northern Crescent iN Phyciodes cocyta cocyta (Cramer, 1777) TL: not specified M S L Ha Phyciodes cocyta selenis (W. Kirby, 1837) TL: not specified M T S1 S2 L1 L2 Ha Phyciodes cocyta pascoensis W. G. Wright, 1905 TL: USA (Wa) M T S Phyciodes cocyta arenacolor Austin, 1998 TL: USA (Nv) M T S Phyciodes cocyta apache Scott & Opler, 2017 TL: USA (NM) Phyciodes anasazi Scott, 1994 Anasaz Crescent ref.16 Phyciodes anasazi anasazi Scott, 1994 TL: USA (Co) M T S Phyciodes anasazi apsaalooke Scott, 1994 ref.16 M S Phyciodes batesii (Reakirt, [1866]) Tawny Crescent iN Phyciodes batesii lakota Scott, 1994 TL: USA (Ne) M T S1 S2 L Ha Phyciodes batesii saskatchewan Scott, 2006 TL: not specified M T Phyciodes batesii batesii (Reakirt, [1866]) TL: USA (Va) M T OD S Phyciodes batesii maconensis Gatrelle, 1998 TL: USA (NC) M T L1 L2 Phyciodes pulchella (Boisduval, 1852) Field Crescent iN Phyciodes pulchella tutchone Scott, 1994 TL: not specified M T S L Phyciodes pulchella owimba Scott, 1998 TL: not specified M S Ab L1 L2 Ha Phyciodes pulchella Siskiyou segregate , Siskiyou Mts., SW OR, NW CA M L Ha Phyciodes pulchella Willamette Valley segregate , Willamette Valley, W Oregon M S L Phyciodes pulchella pulchella (Boisduval, 1852) TL: USA (Ca) M T S L1 L2 L3 H F Ha Phyciodes pulchella deltarufa Scott, 1998 TL: USA (Ca) M T Phyciodes pulchella S CA Sierra Nevada, low elevations segregate , M S L1 L2 Phyciodes pulchella montana (Behr, 1863) TL: USA (Ca) M T S1 S2 L Ha Phyciodes pulchella OR Cascades, high elevations segregate , [nr. montana] M S L Ha Phyciodes pulchella vallis Austin, 1998 TL: USA (Nv) M S Phyciodes pulchella inornatus Austin, 1998 TL: USA (Nv) M S L1 L2 Phyciodes pulchella shoshoni Scott, 1994 TL: USA (Nv) M S L Ha Phyciodes pulchella camillus W. H. Edwards, 1871 TL: USA (Co) M T S1 S2 L1 L2 L3 Ha Tribe Nymphalinae-incertae-sedis Prodryas , [] a fossil genus Prodryas persephone Scudder, 1878 Fossil from Colorado Subfamily Charaxinae Tribe Anaeini Coenophlebia C. Felder & R. Felder, 1862 Coenophlebia archidona (Hewitson, 1860) iN M T Consul Hübner, [1807] Consul electra (Westwood, 1850) Pearly Leafwing iN Consul electra electra (Westwood, 1850) E Mexico to Panama M T S Ab L Consul electra adustus Lamas, 2003 W Mexico (NAY to MICH) M T S Consul excellens (H. Bates, 1864) Black-veined Leafwing iN Consul excellens excellens (H. Bates, 1864) E Chiapas to Guatemala M T S L Consul excellens genini (Le Cerf, 1922) S Mexico (Oaxaca, W Chiapas) to Guatemala M T S L Consul fabius (Cramer, 1776) iN Consul fabius fabius (Cramer, 1776) TL: Surinam M T Consul fabius albinotatus (A. Butler, 1874) TL: Colombia M T Consul fabius bogotanus (A. Butler, 1874) TL: Colombia M T Consul fabius castaneus (A. Butler, 1874) TL: Brazil (Am) M T Consul fabius cecrops (E. Doubleday, [1849]) E & W Mexico to Bolivia M T S Ab L1 L2 Consul fabius diffusus (A. Butler, 1875) TL: Ecuador M T Consul fabius divisus (A. Butler, 1874) TL: Peru M T Consul fabius drurii (A. Butler, 1874) TL: Brazil M T Consul fabius fassli (Röber, 1916) TL: Colombia M T Consul fabius fulvus (A. Butler, 1875) TL: Peru M T Consul fabius ochraceus (A. Butler, 1874) TL: French Guiana M T Consul fabius quadridentatus (A. Butler, 1874) TL: Bolivia M T Consul fabius semifulvus (A. Butler, 1875) TL: Ecuador M T Consul fabius superba (Niepelt, 1923) TL: Colombia M T Consul fabius [n. ssp.] Lamas, MS Ecuador Consul panariste (Hewitson, 1856) iN Consul panariste panariste (Hewitson, 1856) TL: Colombia M T S Consul panariste jansoni (Salvin, 1871) Guatemala to Panama M T S IL Consul panariste ludmilla (Fassl, 1912) TL: Colombia M T Consul panariste pandrosa (Niepelt, 1927) TL: Colombia M T Hypna Hübner, [1819] Hypna clytemnestra (Cramer, 1777) iN Hypna clytemnestra clytemnestra (Cramer, 1777) TL: Surinam M T Hypna clytemnestra corumbaensis Talbot, 1928 TL: Brazil (MT) M T Hypna clytemnestra forbesi Godman & Salvin, 1884 TL: Brazil (Pe) M T Hypna clytemnestra huebneri A. Butler, 1866 TL: Brazil M T Hypna clytemnestra iphigenia Herrich-Schäffer, 1862 Cuba M T S Hypna clytemnestra mexicana A. Hall, 1917 W Mexico, Guatemala M T S L Ha Hypna clytemnestra negra C. Felder & R. Felder, 1862 Nicaragua to Bolivia M T S L Hypna clytemnestra rufescens A. Butler, 1866 TL: Venezuela M T Polygrapha Staudinger, 1887 Polygrapha cyanea (Salvin & Godman, 1868) iN Polygrapha cyanea cyanea (Salvin & Godman, 1868) TL: Ecuador M T S Polygrapha cyanea silvaorum Constantino & Salazar, 1998 TL: Colombia M T S Polygrapha suprema (Schaus, 1920) iN M T S Siderone Hübner, [1823] Siderone galanthis (Cramer, 1775) Red-striped Leafwing iN Siderone galanthis Mexican segregate , E & W Mexico to Central America M S L Siderone galanthis nemesis (Illiger, 1801) Cuba & Isla de Juventud, Hispaniola, Puerto Rico M T S Siderone galanthis galanthis (Cramer, 1775) Central America to Amazon Basin M T S L Siderone galanthis thebais C. Felder & R. Felder, 1862 TL: Colombia M T Siderone galanthis [n. ssp.] Lamas, MS Mexico Siderone syntyche Hewitson, [1854] Red-patched Leafwing iN Siderone syntyche syntyche Hewitson, [1854] S Mexico through Central America M T S Siderone syntyche angustifascia A. Hall, 1917 TL: Ecuador M T Siderone syntyche mars H. Bates, 1860 TL: Brazil (Am) M T Phantos Dias, [2018] Phantos callidryas (R. Felder, 1869) Pale Leafwing E & W Mexico to Panama iN M T S1 S2 L Phantos opalina (Godman & Salvin, 1884) ref.197 M T Zaretis Hübner, [1819] Zaretis delassisei X. Choimet, 2009 ref.185 Zaretis ellops (Ménétriés, 1855) Seasonal Leafwing [= anzuletta], E & W Mexico to Colombia & NW Venezuela iN M T S L Zaretis isidora (Cramer, 1779) Isidora Leafwing [S Mexico] to Ecuador & Argentina iN M T S L Zaretis strigosus (Gmelin, [1790]) ref.197 iN M T Zaretis falcis Dias, Casagrande & O. Mielke, 2012 ref.196 Zaretis crawfordhilli Dias, 2018 ref.197 iN Zaretis elianahenrichae Dias, 2018 ref.197 iN Zaretis hurin Dias, 2018 ref.197 iN Zaretis mirandahenrichae Dias, 2018 ref.197 iN Zaretis itys (Cramer, 1777) Itys Leafwing S Mexico (SE Chiapas) to Amazon Basin and Bolivia iN M T S Zaretis itylus (Westwood, 1850) ref.196 M T Zaretis syene (Hewitson, 1856) iN M T S Zaretis pythagoras Willmott & J. Hall, 2004 iN M S Anaea Hübner, [1819] Anaea aidea (Guérin-Méneville, [1844]) Tropical Leafwing S TX (strays far N), E & W Mexico to Costa Rica iN M T S1 S2 S3 L1 L2 I H Anaea andria Scudder, 1875 Goatweed Leafwing SE AZ, far N Mexico, to E CO, E to IL, N FL, strays N iN M T S L1 L2 L3 I1 I2 H Anaea troglodyta (Fabricius, 1775) Florida Leafwing iN Anaea troglodyta floridalis F. Johnson & W. Comstock, 1941 S FL M T S L Anaea troglodyta cubana (H. Druce, 1905) Cuba & Isla de Juventud, Grand Cayman M T S Anaea troglodyta portia (Fabricius, 1775) Jamaica M T S1 S2 L Anaea troglodyta troglodyta (Fabricius, 1775) Hispaniola M T S L Anaea troglodyta borinquenalis F. Johnson & W. Comstock, 1941 Mona Island & Puerto Rico, [Culebra] M T S L Anaea troglodyta astina (Fabricius, 1793) Virgin Islands (St. Thomas, St. Croix, Tortola) M T S Anaea troglodyta minor A. Hall, 1936 N Lesser Antilles (St. Bartholomew & St. Kitts to Guadeloupe) M T S Fountainea Rydon, 1971 Fountainea centaurus (C. Felder & R. Felder, 1867) iN M T S Fountainea eurypyle (C. Felder & R. Felder, 1862) iN Fountainea eurypyle eurypyle (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Fountainea eurypyle confusa (A. Hall, 1929) E Mexico to Panama M T S L I Fountainea eurypyle glanzi (Rotger, Escalante & Coronado, 1965) W Mexico (SIN to S OAX) M T S Ha Fountainea glycerium (E. Doubleday, [1849]) Angled Leafwing iN Fountainea glycerium glycerium (E. Doubleday, [1849]) E & W Mexico to Panama M T S L1 L2 Fountainea glycerium comstocki (Witt, 1972) TL: Colombia M T Fountainea glycerium cratais (Hewitson, 1874) TL: Bolivia M T Fountainea glycerium venezuelana (F. Johnson & W. Comstock, 1941) TL: Venezuela M T Fountainea glycerium yucatanum (Witt, 1980) Yucatan Peninsula M OI Fountainea johnsoni (Avinoff & Shoumatoff, 1941) Jamaican Leafwing Jamaica, Hispaniola M T S Fountainea halice (Godart, [1824]) iN Fountainea halice halice (Godart, [1824]) TL: Brazil Fountainea halice chrysophana (H. Bates, 1866) Costa Rica & Panama M T L Fountainea halice evelina (W. Comstock, 1961) TL: Bolivia M T Fountainea halice fumata (A. Hall, 1935) TL: Venezuela M T Fountainea halice martinezi (J. Maza & Díaz, 1978) SE Mexico (VER, N OAX) M T S Fountainea halice maya (Witt, 1980) Yucatan Peninsula M T Fountainea halice moretta (H. Druce, 1877) TL: Brazil (Pe) M T Fountainea halice tehuana (A. Hall, 1917) W Mexico (JAL to S CHIS) M T S Fountainea nessus (Latreille, [1813]) Cindy's Leafwing iN M T S L Fountainea rayoensis (J. Maza & Díaz, 1978) Rayo Leafwing SW Mexico (NAY to OAX) iN M T S Fountainea nobilis (H. Bates, 1864) Noble Leafwing iN Fountainea nobilis nobilis (H. Bates, 1864) S Mexico (SW Chiapas) & Guatemala M T S Fountainea nobilis caudata (Röber, 1916) TL: Peru M T S Fountainea nobilis pacifica (Vélez & Salazar, 1991) TL: Colombia M T Fountainea nobilis peralta (A. Hall, 1929) Costa Rica & Panama M T S L Fountainea nobilis romeroi (Descimon, 1988) TL: Venezuela Fountainea nobilis titan (C. Felder & R. Felder, 1867) TL: Colombia M T Fountainea ryphea (Cramer, 1775) Ryphea Leafwing iN Fountainea ryphea ryphea (Cramer, 1775) E Mexico to Amazon Basin M T S L Fountainea ryphea ecuadoralis (F. Johnson & W. Comstock, 1941) TL: Ecuador M T Fountainea ryphea phidile (Geyer, 1837) TL: ? M T Fountainea sosippus (Hopffer, 1874) iN M T Fountainea tyrianthina (Salvin & Godman, 1868) ref.295 iN M T S Memphis Hübner, [1819] Memphis acaudata (Röber, 1916) iN M T Memphis acidalia (Hübner, [1819]) iN Memphis acidalia acidalia (Hübner, [1819]) TL: Surinam M T Memphis acidalia memphis (C. Felder & R. Felder, 1867) TL: Colombia M T S Memphis acidalia victoria (H. Druce, 1877) TL: Brazil (RJ) M T Memphis alberta (H. Druce, 1876) iN M T S Memphis ambrosia (H. Druce, 1874) Ambrosia Leafwing TL: Panama Memphis ambrosia ambrosia (H. Druce, 1874) Costa Rica & Panama M T S L Memphis ambrosia phoebe (H. Druce, 1877) TL: Bolivia M T Memphis anassa (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Memphis anna (Staudinger, 1897) TL: Brazil (Am); Peru Memphis anna anna (Staudinger, 1897) TL: Brazil (Am); Peru M T Memphis anna elina (Staudinger, 1897) TL: Colombia M T S Memphis appias (Hübner, [1825]) iN M S Memphis arginussa (Geyer, 1832) iN Memphis arginussa arginussa (Geyer, 1832) TL: Brazil M T S Memphis eubaena (Boisduval, 1870) Mottled Leafwing SE Mexico to Panama; ref.ref.Dias_et_el_2021 M T S Memphis onophis (C. Felder & R. Felder, 1861) ref.199 M T S Memphis artacaena (Hewitson, 1869) White-patched Leafwing SE Mexico to Colombia iN M T S L I Memphis aulica (Röber, 1916) Aulica Leafwing Costa Rica & Panama M T S Memphis aureola (H. Bates, 1866) Aureola Leafwing E & S Mexico to Colombia iN Memphis aureola aureola (H. Bates, 1866) E & S Mexico to Colombia M T S L Memphis aureola pueblaensis Dottax & Salazar, 2011 Colombia Memphis basilia (Stoll, 1780) iN Memphis basilia basilia (Stoll, 1780) TL: Surinam M T Memphis basilia bella (W. Comstock, 1961) TL: Colombia Memphis basilia drucei (Staudinger, 1887) TL: [Peru] M T Memphis beatrix (H. Druce, 1874) Beatrix Leafwing Costa Rica & Panama iN M T S Memphis boliviana (H. Druce, 1877) TL: Bolivia M T Memphis catinka (H. Druce, 1877) TL: ? M T Memphis cerealia (H. Druce, 1877) TL: Peru M T Memphis cleomestra (Hewitson, 1869) iN Memphis cleomestra cleomestra (Hewitson, 1869) TL: Colombia M T Memphis cleomestra ada (A. Butler, 1875) Costa Rica & Panama M T S Memphis cluvia (Hopffer, 1874) TL: Bolivia M T Memphis dia (Godman & Salvin, 1884) Dia Leafwing iN Memphis dia dia (Godman & Salvin, 1884) SE Mexico to Panama M T S Memphis dia divina (Röber, 1916) TL: Bolivia M T Memphis falcata (Hopffer, 1874) TL: Peru M S Memphis forreri (Godman & Salvin, 1884) Forrer's Leafwing S TX (rare vagrant), E & W Mexico to Costa Rica iN M T S L Memphis glauce (C. Felder & R. Felder, 1862) iN Memphis glauce glauce (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T S Memphis glauce centralis (Röber, 1916) Honduras to Panama M T S Memphis glauce cicla (Möschler, 1877) TL: Surinam M T Memphis glauce felderi (Röber, 1916) TL: Ecuador M T Memphis grandis (H. Druce, 1877) TL: ? M T S Memphis hedemanni (R. Felder, 1869) Hedemann's Leafwing SE Mexico, Guatemala M T S Memphis herbacea (A. Butler & H. Druce, 1872) Herbacea Leafwing SE Mexico to Costa Rica iN M T S Memphis hirta (Weymer, 1907) iN M T Memphis iphis (Latreille, [1813]) TL: ? M T S Memphis juliani Constantino, 1999 TL: Colombia M T Memphis laertes (Cramer, 1775) iN M T S Memphis laura (H. Druce, 1877) iN Memphis laura laura (H. Druce, 1877) TL: Panama M T Memphis laura balboa (A. Hall, 1927) Costa Rica & Panama M T S Memphis laura caucana (Joicey & Talbot, 1922) TL: Colombia M T S Memphis laura rosae (Fassl, 1909) TL: Colombia M T S Memphis lemnos (H. Druce, 1877) TL: Peru M T L Memphis leonida (Stoll, 1782) iN Memphis leonida leonida (Stoll, 1782) TL: Surinam M T S L Memphis leonida editha (W. Comstock, 1961) TL: Brazil (SC) M T S Memphis lineata (Salvin, 1869) iN M T Memphis lorna (H. Druce, 1877) iN M T Memphis lyceus (H. Druce, 1877) Lyceus Leafwing Costa Rica to Bolivia iN M T S Memphis maria Pyrcz & Neild, 1996 TL: Venezuela M T Memphis moeris (C. Felder & R. Felder, 1867) iN M T Memphis montesino Pyrcz, 1995 TL: Venezuela M T Memphis mora (H. Druce, 1874) iN Memphis mora mora (H. Druce, 1874) TL: Colombia M T Memphis mora annetta (W. Comstock, 1961) TL: Ecuador M T Memphis mora montana (Röber, 1916) TL: Peru M T S L Memphis mora orthesia (Godman & Salvin, 1884) SE Mexico to Panama M T S Memphis moruus (Fabricius, 1775) iN Memphis moruus moruus (Fabricius, 1775) TL: India M T Memphis moruus boisduvali (W. Comstock, 1961) E Mexico to Panama M T S L Memphis moruus leonila (W. Comstock, 1961) TL: Ecuador M T S Memphis moruus morpheus (Staudinger, [1886]) TL: Brazil (Am); Peru M T S L Memphis moruus phila (H. Druce, 1877) TL: Colombia M T Memphis moruus stheno (Prittwitz, 1865) TL: Brazil (RJ) M T Memphis neidhoeferi (Rotger, Escalante & Coronado, 1965) Wavy-edged Leafwing SE Mexico to Costa Rica M T S Memphis nenia (H. Druce, 1877) TL: Brazil (SP) Memphis nenia nenia (H. Druce, 1877) TL: Brazil (SP) M T Memphis nenia threnodion (Bargmann, 1929) TL: Colombia M T Memphis oenomais (Boisduval, 1870) Edge Leafwing E Mexico through Central America iN M T S L Memphis offa (H. Druce, 1877) iN Memphis offa offa (H. Druce, 1877) TL: Ecuador M T S Memphis offa gudrun (Niepelt, 1924) TL: Colombia M T Memphis otrere (Hübner, [1825]) iN M T S Memphis paulus Orellana & Costa, 2014 ref.188 Memphis pasibula (E. Doubleday, [1849]) iN M T S Memphis perenna (Godman & Salvin, 1884) Perenna Leafwing iN Memphis perenna perenna (Godman & Salvin, 1884) E & SW Mexico & Guatemala M T S L Memphis perenna austrina (W. Comstock, 1961) TL: Colombia M S Memphis lankesteri (A. Hall, 1935) Lankester's Leafwing Costa Rica; ref.199 M T S Memphis phantes (Hopffer, 1874) iN Memphis phantes phantes (Hopffer, 1874) TL: Peru M T Memphis phantes vicinia (Staudinger, 1887) TL: Peru M T S Memphis philumena (E. Doubleday, [1849]) iN Memphis philumena philumena (E. Doubleday, [1849]) TL: Bolivia M T Memphis philumena chaeronea (C. Felder & R. Felder, 1861) TL: Colombia M T Memphis philumena corita (Fruhstorfer, 1916) TL: Brazil (ES) M T S Memphis philumena indigotica (Salvin, 1869) Costa Rica & Panama M T S Memphis philumena xenica (H. Bates, 1864) E Mexico & Guatemala M T S Memphis pithyusa (R. Felder, 1869) Pale-spotted Leafwing S TX, E & W Mexico to Bolivia; ref.199 iN M T S L Memphis polycarmes (Fabricius, 1775) iN M S Memphis polyxo (H. Druce, 1874) iN M T Memphis praxias (Hopffer, 1874) iN Memphis praxias praxias (Hopffer, 1874) TL: Peru Memphis praxias oblita (A. Hall, 1929) TL: Ecuador M T Memphis schausiana (Godman & Salvin, 1894) Great Leafwing C Veracruz M T S Memphis proserpina (Salvin, 1869) Proserpina Leafwing SE Mexico to Costa Rica iN M T S Memphis elara (Godman & Salvin, 1897) Elara Leafwing Costa Rica & W Panama M T S Memphis kingi (L. Miller & Nicolay, 1971) King's Leafwing Panama (Cerro Campana, Chiriqui, Canal Zone) M T Memphis pseudiphis (Staudinger, 1887) iN M T S Memphis salinasi Pyrcz, 1993 TL: Venezuela M T Memphis smalli T.J. Riley & F.M.S. Dias, 2021 ref.199 Memphis verticordia (Hübner, [1831]) Hispaniolan Leafwing Hispaniola iN Memphis verticordia verticordia (Hübner, [1831]) TL: Cuba M T S Memphis echemus (E. Doubleday, [1849]) Cuban Leafwing iN Memphis echemus echemus (E. Doubleday, [1849]) Cuba & Isla de Juventud; one-time stray to Texas M T S Memphis echemus bahamae (Witt, 1972) Bahamas: New Providence & Andros M T S Memphis echemus danieliana (Witt, 1972) Cayman Islands M T S Memphis intermedia (Witt, 1972) Bahaman Leafwing iN Memphis intermedia intermedia (Witt, 1972) Islands of Great Inagua Bank & Turks, Caicos M T S Memphis intermedia venus L. Miller, Simon & Harvey, 1992 Islands of Crooked - Acklins Bank M T1 T2 Memphis intermedia mayaguanae L. Miller, Simon & Harvey, 1992 Mayaguana Bank M T1 T2 Memphis dominicana (Godman & Salvin, 1884) Dominican Leafwing iN Memphis dominicana dominicana (Godman & Salvin, 1884) Dominica M T S Memphis dominicana luciana (A. Hall, 1929) St. Lucia M T S Memphis viloriae Pyrcz & Neild, 1996 TL: Venezuela M T Memphis wellingi L. Miller & J. Miller, 1976 Welling's Leafwing SW Mexico (S OAX, S CHIS) M T S Memphis xenippa (A. Hall, 1935) TL: Colombia M T Memphis xenocles (Westwood, 1850) iN Memphis xenocles xenocles (Westwood, 1850) TL: Bolivia M T S Memphis xenocles carolina (W. Comstock, 1961) SE Mexico to Panama M S L Memphis xenocles fisilis (A. Hall, 1935) TL: Colombia M T Memphis xenocles marginalis (A. Hall, 1935) TL: Brazil (RJ/MG/SP) M T S Memphis xenocrates (Westwood, 1850) iN Memphis xenocrates xenocrates (Westwood, 1850) ref.295 M T Memphis xenocrates ernestoi (Salazar & Constantino, 2001) ref.295 M S Memphis xenocrates punctimarginale (Kaye, 1918) ref.295 M T Tribe Anaemorphini Anaeomorpha Rothschild, 1894 Anaeomorpha splendida Rothschild, 1894 TL: Peru Anaeomorpha splendida splendida Rothschild, 1894 TL: Peru M T S Anaeomorpha splendida esmeralda Attal & M. Büche, 2008 ref.158 Anaeomorpha mirifica M. Simon & Willmott, 2017 ref.314 Tribe Preponini Archaeoprepona Fruhstorfer, 1915 Archaeoprepona chromus (Guérin-Méneville, [1844]) ref.274 iN Archaeoprepona chromus chromus (Guérin-Méneville, [1844]) TL: Colombia M T Archaeoprepona chromus macaraoensis (Schäffler & Frankenbach, 2011) TL: not specified Archaeoprepona chromus marankiariensis (Schäffler & Frankenbach, 2011) TL: not specified Archaeoprepona priene (Hewitson, 1859) ref.274 M T Archaeoprepona licomedes (Cramer, 1777) iN Archaeoprepona licomedes licomedes (Cramer, 1777) TL: Surinam M T Archaeoprepona licomedes pacifica Constantino & Salazar, 1998 TL: Colombia M T Archaeoprepona chalciope (Hübner, [1823]) iN M T Archaeoprepona amphimachus (Fabricius, 1775) iN Archaeoprepona amphimachus amphimachus (Fabricius, 1775) TL: Indiis M T Archaeoprepona amphimachus amphiktion (Fruhstorfer, 1916) E Mexico to Colombia M T S1 S2 L I1 I2 I3 Archaeoprepona amphimachus baroni J. Maza, 1982 SW Mexico (GRO, S OAX) M T S Archaeoprepona amphimachus pseudomeander (Fruhstorfer, 1906) TL: Brazil (RS) M T Archaeoprepona amphimachus symaithus (Fruhstorfer, 1916) TL: Ecuador M T Archaeoprepona amphimachus costaricana Schäffler & Frankenbach, 2011 TL: not specified Archaeoprepona amphimachus esmeraldica Schäffler & Frankenbach, 2011 TL: not specified Archaeoprepona amphimachus araguaensis Schäffler & Frankenbach, 2011 TL: not specified Archaeoprepona meander (Cramer, 1775) iN Archaeoprepona meander meander (Cramer, 1775) TL: Surinam Archaeoprepona meander castorina (E. May, 1932) TL: Brazil (RJ) M T Archaeoprepona meander megabates (Fruhstorfer, 1916) Panama to Bolivia M T S L Archaeoprepona meander phoebus (Boisduval, 1870) SE Mexico to Honduras M T S Archaeoprepona camilla (Godman & Salvin, 1884) Pale Prepona iN Archaeoprepona camilla camilla (Godman & Salvin, 1884) Nicaragua to Panama M T S1 S2 L I1 I2 Archaeoprepona camilla metabus (Fruhstorfer, 1916) TL: Colombia M T Archaeoprepona demophon (Linnaeus, 1758) iN Archaeoprepona demophon demophon (Linnaeus, 1758) TL: Indiis M HS Archaeoprepona demophon centralis (Fruhstorfer, 1905) S TX (rare stray), E Mexico through Panama M T S L I1 I2 Archaeoprepona demophon muson (Fruhstorfer, 1905) TL: Colombia; Ecuador; Bolivia M T Archaeoprepona demophon occidentalis (Stoffel & Descimon, 1974) W Mexico (SIN to S OAX) M T S L Archaeoprepona demophon thalpius (Hübner, [1814]) TL: ? M T Archaeoprepona demophoon (Hübner, [1814]) iN Archaeoprepona demophoon demophoon (Hübner, [1814]) TL: ? M T Archaeoprepona demophoon andicola (Fruhstorfer, 1904) TL: Ecuador M T Archaeoprepona demophoon antimache (Hübner, [1819]) TL: Indien Archaeoprepona demophoon crassina (Fruhstorfer, 1904) Cuba M T S Archaeoprepona demophoon gulina (Fruhstorfer, 1904) E Mexico to Amazon basin M T S1 S2 L I1 I2 Archaeoprepona demophoon insulicola (Fruhstorfer, 1897) Hispaniola M T S L Archaeoprepona demophoon ramosorum K. Johnson & Descimon, 1989 Puerto Rico M T S L I Archaeoprepona demophoon mexicana Llorente, Descimon & K. Johnson, 1993 W Mexico (NAY to S OAX) M T S Archaeoprepona demophoon dominici Schäffler & Frankenbach, 2011 TL: not specified Archaeoprepona demophoon niklasi Schäffler & Frankenbach, 2011 TL: not specified Archaeoprepona phaedra (Godman & Salvin, 1884) Falcate Prepona iN Archaeoprepona phaedra SW Mexican segregate , SW Mexico (GRO, S OAX) M S Archaeoprepona phaedra aelia (Godman & Salvin, 1889) SE Mexico possibly to Nicaragua M T S L Archaeoprepona phaedra phaedra (Godman & Salvin, 1884) Costa Rica & Panama M T Mesoprepona Bonfantti, Casagrande & O. Mielke, 2017 Mesoprepona pheridamas (Cramer, 1777) ref.273 iN M T Prepona Boisduval, 1836 Prepona dexamenus Hopffer, 1874 Least Prepona iN Prepona dexamenus dexamenus Hopffer, 1874 Panama to Amazon Basin M T S Prepona dexamenus leuctra Fruhstorfer, 1904 TL: Brazil (ES) M T Prepona dexamenus medinai Beutelspacher, 1981 SE Mexico to Honduras M T S Prepona laertes (Hübner, [1811]) iN Prepona laertes laertes (Hübner, [1811]) TL: ? M T Prepona laertes demodice (Godart, [1824]) TL: French Guiana; Brazil M T1 T2 T3 T4 Prepona laertes louisa A. Butler, 1870 TL: Cuba M T Prepona laertes octavia Fruhstorfer, 1905 E & W Mexico through Central America, recorded introduction to USA: Florida M T1 T2 S1 S2 L Prepona werneri M. Hering & Hopp, 1925 iN M S Prepona pylene Hewitson, [1854] iN Prepona pylene pylene Hewitson, [1854] TL: Brazil (RJ) M T Prepona eugenes H. Bates, 1865 TL: Brazil (Pa) Prepona eugenes eugenes H. Bates, 1865 TL: Brazil (Pa) M T S Prepona eugenes bahiana Fruhstorfer, 1897 TL: Brazil (Ba) M T Prepona eugenes laertides Staudinger, 1898 TL: Bolivia M T Prepona gnorima H. Bates, 1865 Mottled Prepona Honduras to Colombia iN Prepona gnorima gnorima H. Bates, 1865 Honduras to Colombia M T S Prepona gnorima jordani Fruhstorfer, 1905 TL: Ecuador M T S Prepona gnorima philetas Fruhstorfer, 1904 SE Mexico to Honduras M T S Prepona deiphile (Godart, [1824]) iN Prepona deiphile Nayarit segregate , Sierra de San Juan, Nayarit, Mexico M S L Prepona deiphile lambertoana Llorente, Luis & F. González, 1992 Michoacán, Estado de Mexico M T1 T2 S Prepona deiphile brooksiana Godman & Salvin, 1889 E Mexico (SLP to N OAX) M T TI S L I Prepona deiphile escalantiana Stoffel & Mast, 1973 SE Mexico (vic. Los Tuxtlas, Veracruz) M S1 S2 Prepona deiphile NW Chiapas segregate , NW Chiapas M S Prepona deiphile diaziana L. Miller & J. Miller, 1976 E Chiapas M T S Prepona deiphile salvadora Llorente, Luis & F. González, 1992 El Salvador M T S Prepona deiphile lygia Fruhstorfer, 1904 Costa Rica & Panama M T S Prepona deiphile photidia Fruhstorfer, 1912 TL: Colombia Prepona deiphile sphacteria Fruhstorfer, 1916 TL: Peru Prepona deiphile neoterpe Honrath, 1884 TL: Peru M T Prepona deiphile deiphile (Godart, [1824]) TL: Brazil Prepona deiphile xenagoras Hewitson, 1875 TL: Bolivia M T Prepona ibarra Beutelspacher, 1982 Guerrero, SW Mexico M T S Prepona silvana Turrent & García, 2019 Turrent & García (2019, Revista de la Sociedad Mexicana de Lepidopterolog ... Prepona praeneste Hewitson, 1859 iN Prepona praeneste praeneste Hewitson, 1859 TL: Colombia M T Prepona praeneste abrupta Biedermann, 1936 TL: Peru M T Prepona praeneste buckleyana Hewitson, 1876 TL: Bolivia M T Prepona praeneste choroniensis Lichy, 1975 TL: Venezuela M T Prepona praeneste confusa Niepelt, 1913 TL: Peru M T Prepona praeneste isabelae Orellana, 2000 TL: Venezuela Prepona praeneste virago Biedermann, 1938 TL: Colombia M T S Prepona aedon (Hewitson, 1848) iN Prepona aedon aedon (Hewitson, 1848) ref.274 M T Prepona aedon pepitoensis (Michael, 1930) ref.274 M T Prepona aedon rodriguezi (Schaus, 1918) S Mexico (SE Chiapas) to Costa Rica; ref.274 M T S L Prepona amydon (Hewitson, [1854]) iN Prepona amydon amydon (Hewitson, [1854]) ref.274 M T S Prepona amydon amydonius (Staudinger, [1885]) ref.274 M T Prepona amydon aristoxenus (Niepelt, 1913) ref.274 M T Prepona amydon aurantiaca (Fruhstorfer, 1897) ref.274 M T Prepona amydon bogotana (Fruhstorfer, 1895) ref.274 M T Prepona amydon boliviensis (Fruhstorfer, 1895) ref.274 M T Prepona amydon excelsior (Lathy, 1924) ref.274 M T Prepona amydon ferdinandi (Fruhstorfer, 1895) ref.274 M T Prepona amydon frontina (Fruhstorfer, 1895) ref.274 M T Prepona amydon lacandona (R. G. Maza & J. Maza, 1999) SE Mexico (SE Chiapas) to Honduras; ref.274 M T S Prepona amydon oaxacata (Kruck, 1931) S Mexico (N OAX) to Guatemala; ref.274 M T S L Prepona amydon phalcidon (Hewitson, 1855) ref.274 M T1 T2 T3 T4 T5 T6 Prepona amydon philatelica (DeVries, 1980) Nicaragua to Panama; ref.274 M T S L I1 I2 Prepona amydon rubella (Michael, 1930) ref.274 M T Prepona amydon smalli (L. Miller & Nicolay, 1971) Panama (vic. Canal Zone); ref.274 M T S Prepona amydon uniformis (Michael, 1929) ref.274 M T Prepona amydon zenodorus (Hewitson, 1870) ref.274 M T Prepona amydon [n. ssp.] (Neild, MS) Venezuela Prepona hewitsonius (H. Bates, 1860) iN Prepona hewitsonius hewitsonius (H. Bates, 1860) ref.274 M T Prepona hewitsonius beata (Staudinger, [1885]) ref.274 M T Prepona hewitsonius beatifica (Hewitson, 1869) ref.274 M T Prepona hewitsonius stuarti (Godman & Salvin, 1882) ref.274 M T S Prepona claudina (Godart, [1824]) iN Prepona claudina claudina (Godart, [1824]) ref.274 M T1 T2 T3 T4 T5 S Prepona claudina annetta (G. Gray, 1832) ref.274 M T Prepona claudina croesus (Staudinger, 1896) ref.274 M T Prepona claudina delavillae (Neild, 1996) ref.274 Prepona claudina godmani (Fruhstorfer, 1895) ref.274 M T Prepona claudina lugens (Staudinger, 1886) ref.274 M T S Prepona claudina patriciae (Attal, 2000) ref.274 Prepona claudina sardanapalus (H. Bates, 1860) ref.274 M T1 T2 Prepona narcissus (Staudinger, [1885]) iN Prepona narcissus narcissus (Staudinger, [1885]) ref.274 M T Prepona narcissus stoffeli (Mast & Descimon, 1972) ref.274 Prepona narcissus tapajonus (Fassl, 1921) ref.274 M T1 T2 Subfamily Satyrinae Tribe Morphini Antirrhea Hübner, [1822] Antirrhea adoptiva (Weymer, 1909) iN Antirrhea adoptiva adoptiva (Weymer, 1909) TL: Colombia M T Antirrhea adoptiva porphyrosticta (Watkins, 1928) TL: Ecuador M T Antirrhea adoptiva [n. ssp.#1] Orellana, MS TL: Venezuela Antirrhea adoptiva [n. ssp.#2] Lamas, MS TL: Peru Antirrhea archaea Hübner, [1822] iN Antirrhea geryon (C. Felder & R. Felder, 1862) iN Antirrhea geryon geryon (C. Felder & R. Felder, 1862) TL: Colombia M T Antirrhea geryon geryonides (Weymer, 1909) TL: Ecuador M T Antirrhea geryon weymeri Salazar, Constantino & López, 1998 TL: Colombia M T S Antirrhea hela (C. Felder & R. Felder, 1862) iN M T Antirrhea ornata (A. Butler, 1870) iN M T Antirrhea phasiana (A. Butler, 1870) iN M T S Antirrhea philoctetes (Linnaeus, 1758) iN Antirrhea philoctetes philoctetes (Linnaeus, 1758) TL: Indiis M T HS Antirrhea philoctetes murena (Staudinger, [1886]) TL: Brazil (Am) M T S Antirrhea philaretes (C. Felder & R. Felder, 1862) iN Antirrhea philaretes casta (H. Bates, 1865) SE Mexico, Guatemala M T S Antirrhea philaretes tomasia (A. Butler, 1875) Costa Rica & Panama (Pacific slope) M T S L Antirrhea philaretes philaretes (C. Felder & R. Felder, 1862) TL: Colombia M T Antirrhea philaretes lindigii (C. Felder & R. Felder, 1862) [= miltiades], Guatemala to Amazon Basin; Atlantic slope M T S1 S2 L I1 I2 Antirrhea philaretes intermedia Salazar, Constantino & López, 1998 TL: Colombia M T Antirrhea philaretes avernus (Hopffer, 1874) TL: Peru M T Antirrhea philaretes theodori (Fruhstorfer, 1907) TL: Brazil (Am) M T Antirrhea philaretes lumejia Orellana, 2004 TL: Venezuela Antirrhea philaretes piaroa Orellana & Neild, 2008 ref.266 Antirrhea ulei Strand, 1912 iN Antirrhea pterocopha (Salvin & Godman, 1868) Brilliant Antirrhea iN Antirrhea pterocopha pterocopha (Salvin & Godman, 1868) Costa Rica & W Panama (Atlantic slope; Carrillo Belt) M T S Antirrhea pterocopha chocoensis Salazar, 1997 TL: Colombia M T S Antirrhea taygetina (A. Butler, 1868) iN Antirrhea taygetina taygetina (A. Butler, 1868) TL: Brazil (Am) M T S Antirrhea taygetina rodwayi (A. Hall, 1939) TL: Guyana M T Antirrhea taygetina [n. ssp.] Vane-Wright, MS Ecuador M S Antirrhea undulata (M. Hering & Hopp, 1925) iN M T Antirrhea watkinsi Rosenberg & Talbot, 1914 iN Antirrhea watkinsi watkinsi Rosenberg & Talbot, 1914 TL: Peru M T S Antirrhea watkinsi callephebus (Tessmann, 1928) TL: Peru M T Caerois Hübner, [1819] Caerois chorinaeus (Fabricius, 1775) iN Caerois chorinaeus chorinaeus (Fabricius, 1775) TL: Surinam M T HS Caerois chorinaeus protonoe Fruhstorfer, 1912 TL: Peru M T Caerois chorinaeus rufomarginata Lathy, 1918 TL: Peru M T Caerois chorinaeus [n. ssp.] Lamas, MS Peru Caerois gerdrudtus (Fabricius, 1793) False Antirrhea Costa Rica to W Ecuador iN M S L I1 I2 Morpho Fabricius, 1807 Morpho absoloni E. May, 1924 TL: Brazil (Ac) M T Morpho achilles (Linnaeus, 1758) iN Morpho achilles achilles (Linnaeus, 1758) TL: America M T S HS Morpho achilles agamedes Fruhstorfer, 1912 TL: Peru M T Morpho achilles fischeri Weber, 1962 TL: Peru M T Morpho achilles glaisi Le Moult & Réal, 1962 TL: Venezuela M T Morpho achilles guaraunos Le Moult, 1925 TL: Venezuela M T Morpho achilles patroclus C. Felder & R. Felder, 1861 TL: Colombia M T Morpho achilles phokylides Fruhstorfer, 1912 TL: Bolivia M T Morpho achilles vitrea A. Butler, 1866 TL: Bolivia M T Morpho aega (Hübner, [1822]) iN Morpho aega aega (Hübner, [1822]) TL: ? M T Ab Morpho aega amargosensis Otero & Soares, 1996 TL: Brazil (Ba) M T Morpho amphitryon Staudinger, 1887 iN Morpho amphitryon amphitryon Staudinger, 1887 TL: Peru M T Morpho amphitryon cinereus Duchêne, 1985 TL: Peru M T Morpho amphitryon susarion Fruhstorfer, 1913 TL: Bolivia M T Morpho anaxibia (Esper, [1801]) iN M T Morpho athena Otero, 1966 iN M T Morpho aurora Westwood, 1851 iN Morpho aurora aurora Westwood, 1851 TL: Bolivia M T Morpho aurora aureola Fruhstorfer, 1913 TL: Peru Morpho cisseis C. Felder & R. Felder, 1860 iN Morpho cisseis cisseis C. Felder & R. Felder, 1860 TL: Brazil (Am) M T Morpho cisseis gahua Blandin, 1988 TL: Peru M T Morpho cisseis phanodemus Hewitson, 1869 TL: Ecuador M T Morpho cypris Westwood, 1851 iN Morpho cypris cypris Westwood, 1851 TL: Colombia M T Morpho cypris aphrodite Le Moult & Réal, 1962 Honduras, Nicaragua M T S Morpho cypris bugaba Staudinger, 1887 Costa Rica & Panama M T S Morpho cypris chrysonicus Fruhstorfer, 1913 TL: Venezuela M T Morpho cypris limonensis Schäffler, Frankenbach & Dufek, 2011 TL: not specified Morpho cypris ceibaensis Schäffler, Frankenbach & Dufek, 2011 TL: not specified Morpho deidamia (Hübner, [1819]) iN Morpho deidamia deidamia (Hübner, [1819]) TL: Surinam M T HS Morpho deidamia diffusa Le Moult & Réal, 1962 TL: Brazil (Ma) M T Morpho deidamia diomedes Weber, 1944 TL: [Peru] Morpho deidamia electra Röber, 1903 TL: Bolivia M T Morpho deidamia granadensis C. Felder & R. Felder, 1867 Panama to Colombia M T S L Morpho deidamia guaraura Le Cerf, 1925 TL: Venezuela M T Morpho deidamia hermione Röber, 1903 TL: Colombia M T Morpho deidamia jacki Neild, 2008 ref.266 Morpho deidamia lycanor Fruhstorfer, 1907 TL: Ecuador M T Morpho deidamia neoptolemus Wood, 1863 TL: [Brazil (Am)] M T Morpho deidamia polybaptus A. Butler, 1875 Nicaragua to Panama; Atlantic slope M T S Morpho deidamia pyrrhus Staudinger, 1887 TL: Peru M T Morpho deidamia steinbachi Le Moult & Réal, 1962 TL: Bolivia M T Morpho epistrophus (Fabricius, 1796) iN Morpho epistrophus epistrophus (Fabricius, 1796) TL: Brazil (RJ) M T Morpho epistrophus argentinus Fruhstorfer, 1907 TL: Argentina M T Morpho epistrophus catenaria Perry, 1811 TL: Brazil M T Morpho epistrophus titei Le Moult & Réal, 1962 TL: Paraguay M T Morpho eugenia Deyrolle, 1860 iN M T Morpho hecuba (Linnaeus, 1771) iN Morpho hecuba hecuba (Linnaeus, 1771) TL: French Guiana M S Morpho hecuba obidonus Fruhstorfer, 1905 TL: Brazil (Pa) M T Morpho hecuba polyidos Fruhstorfer, 1912 TL: Venezuela M T Morpho hecuba werneri Hopp, 1921 TL: Colombia M T Morpho helenor (Cramer, 1776) iN Morpho helenor helenor (Cramer, 1776) TL: Surinam M T Morpho helenor achillaena (Hübner, [1823]) TL: ? M T1 T2 Morpho helenor achillides C. Felder & R. Felder, 1867 TL: Brazil M T1 T2 Morpho helenor anakreon Fruhstorfer, 1910 TL: Brazil (Pe) M T Morpho helenor charapensis Le Moult & Réal, 1962 TL: Peru M T Morpho helenor coelestis A. Butler, 1866 TL: Brazil M T Morpho helenor cortone Fruhstorfer, 1913 TL: Colombia M T Morpho helenor corydon Guenée, 1859 TL: "Colombia"; Venezuela M T Morpho helenor guerrerensis Le Moult & Réal, 1962 SW Mexico (MICH to S OAX) M T S L Morpho helenor insularis Fruhstorfer, 1912 TL: Trinidad M T Morpho helenor leontius C. Felder & R. Felder, 1867 TL: Colombia M T Morpho helenor macrophthalmus Fruhstorfer, 1913 TL: Colombia M T Morpho helenor maculata Röber, 1903 TL: Ecuador M T Morpho helenor marajoensis Le Moult & Réal, 1962 TL: Brazil (Pa) M T Morpho helenor marinita A. Butler, 1872 SW Costa Rica to Panama (Pacific slope) M T S L I Morpho helenor montezuma Guenée, 1859 E Mexico to Honduras M T S L Morpho helenor narcissus Staudinger, 1887 Nicaragua to Panama M T S L I Morpho helenor octavia H. Bates, 1864 SW Mexico (Chiapas) to El Salvador M T S L Morpho helenor packeri Neild, 2008 ref.266 Morpho helenor papirius Hopffer, 1874 TL: Peru M T Morpho helenor peleides Kollar, 1850 Panama to Colombia M T S L Morpho helenor peleus Röber, 1903 TL: Venezuela M T Morpho helenor pindarus Fruhstorfer, 1910 TL: Brazil (MT) M T Morpho helenor popilius Hopffer, 1874 TL: "Bolivia"; "Venezuela"; [Colombia] M T Morpho helenor rugitaeniatus Fruhstorfer, 1907 TL: Ecuador M T Morpho helenor telamon Röber, 1903 TL: Colombia M T Morpho helenor theodorus Fruhstorfer, 1907 TL: Brazil (Am) M T Morpho helenor tucupita Le Moult, 1925 TL: Venezuela M T Morpho helenor ululina Le Moult & Réal, 1962 TL: Venezuela M T Morpho helenor violaceus Fruhstorfer, 1912 TL: Brazil (SC) M T Morpho helenor zonaras Fruhstorfer, 1912 Panama M T Morpho helenor prometa Gareca & Blandin, 2011 TL: not specified Morpho helenor coibaensis Jongerling, 2011 TL: not specified Morpho helenor caldarai Schäffler & Frankenbach, 2011 TL: not specified Morpho hercules (Dalman, 1823) iN M T Morpho lympharis A. Butler, 1873 iN Morpho lympharis lympharis A. Butler, 1873 TL: Peru M T Morpho lympharis descimokoenigi Blandin, 1993 TL: Peru M T Morpho lympharis eros Staudinger, 1892 TL: Bolivia M T Morpho lympharis stoffeli Le Moult & Réal, 1962 TL: Peru M T Morpho marcus (Schaller, 1785) iN Morpho marcus marcus (Schaller, 1785) TL: ? M T Morpho marcus intermedia Kaye, 1917 TL: Peru M T Morpho marcus major Lathy, 1905 TL: Peru M T Morpho menelaus (Linnaeus, 1758) iN Morpho menelaus menelaus (Linnaeus, 1758) TL: [Surinam] M T HS Morpho menelaus alexandrovna H. Druce, 1874 TL: Peru M T Morpho menelaus argentiferus Fruhstorfer, 1913 TL: Peru M T Morpho menelaus assarpai Röber, 1903 TL: Peru M T Morpho menelaus coeruleus (Perry, 1810) TL: ? M T Morpho menelaus didius Hopffer, 1874 TL: Peru M T1 T2 Morpho menelaus eberti Weber, 1963 TL: Brazil (Pe) M T Morpho menelaus godartii Guérin-Méneville, [1844] TL: Bolivia M T Morpho menelaus titogilberti Blandin & Gareca, 2011 TL: not specified Morpho menelaus julanthiscus Fruhstorfer, 1907 TL: Ecuador M T Morpho menelaus kesselringi H. Fischer, 1962 TL: Brazil (Pa) M T Morpho menelaus laurellae Neild, 2008 ref.266 Morpho menelaus neildi Blandin, 2008 ref.266 Morpho menelaus occidentalis C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Morpho menelaus orinocensis Le Moult, 1925 TL: Venezuela M T Morpho menelaus terrestris A. Butler, 1866 TL: Brazil (Am) M T Morpho menelaus verae Weber, 1951 TL: Brazil (Pa) M T Morpho menelaus zischkai H. Fischer, 1962 TL: Bolivia M T Morpho amathonte Deyrolle, 1860 Amathonte Morpho Nicaragua to Panama; ref.168 M T S L Morpho niepelti Röber, 1927 iN M T Morpho polyphemus Westwood, [1850] White Morpho iN Morpho polyphemus polyphemus Westwood, [1850] SE AZ (rare stray), W Mexico to El Salvador M T S1 S2 L I Morpho polyphemus catalina Corea & I. Chacón, 1984 Honduras to NW Costa Rica M T S I Morpho polyphemus luna A. Butler, 1869 SE Mexico to Guatemala M T S L Morpho portis (Hübner, [1821]) iN Morpho portis portis (Hübner, [1821]) TL: ? M T Morpho portis thamyris C. Felder & R. Felder, 1867 TL: Brazil M T Morpho rhetenor (Cramer, 1775) iN Morpho rhetenor rhetenor (Cramer, 1775) TL: Surinam M T S Morpho rhetenor cacica Staudinger, 1876 TL: Peru M T Morpho rhetenor columbianus E. Krüger, 1925 TL: Colombia M T1 T2 Morpho rhetenor equatenor Le Moult & Réal, 1962 TL: Ecuador M T Morpho rhetenor hightoni Neild, 2008 ref.266 Morpho rhetenor helena Staudinger, 1890 TL: Peru M T Morpho rhetenor subtusmurina Le Moult & Réal, 1962 TL: Brazil (MT) M T Morpho rhodopteron Godman & Salvin, 1880 iN M T Morpho richardus Fruhstorfer, 1898 TL: Brazil (MG) M T Morpho sulkowskyi Kollar, 1850 iN Morpho sulkowskyi sulkowskyi Kollar, 1850 TL: Colombia M T Morpho sulkowskyi hoppiana Niepelt, 1923 TL: Colombia M T Morpho sulkowskyi selenaris Le Moult & Réal, 1962 TL: Peru M T Morpho sulkowskyi sirene Niepelt, 1911 TL: Ecuador M T Morpho sulkowskyi [n. ssp.#1] Lamas, MS TL: Peru M T Morpho sulkowskyi [n. ssp.#2] Lamas, MS TL: Peru Morpho telemachus (Linnaeus, 1758) iN Morpho telemachus telemachus (Linnaeus, 1758) TL: [Surinam] M T Morpho telemachus iphiclus C. Felder & R. Felder, 1862 TL: Colombia M T Morpho telemachus lilianae Le Moult, 1927 TL: Venezuela M T Morpho telemachus [n. ssp.] Ebert, MS Brazil (Pe) Morpho justitiae Salvin & Godman, 1868 Stub-tailed Morpho SE Mexico (E OAX, far SE VER, N & C CHIS) to Guatemala & Honduras; ref.144 Morpho justitiae schweizeri (R. F. Maza, 1987) SE Mexico; vic. Los Tuxtlas, SE Veracruz; ref.144 M T S Morpho justitiae oaxacensis Le Moult & Réal, 1962 SE Mexico (N Oaxaca); ref.144 M T S Morpho justitiae neoescalantei Schäffler, 2009 ref.144 Morpho justitiae mercedes J. Maza, 2023 ref.144 Morpho justitiae justitiae Salvin & Godman, 1868 SE Mexico (E OAX, far SE VER, N & C CHIS) to Guatemala & Honduras; ref.144 M T S L Morpho malintzin R. G. Maza & J. Maza, 2023 ref.144 Morpho aquarius A. Butler, 1872 ref.144 Morpho aquarius nebulosus R. G. Maza, 2023 ref.144 Morpho aquarius monica J. Maza, 2023 ref.144 Morpho aquarius aquarius A. Butler, 1872 ref.144 M T Morpho theseus Deyrolle, 1860 iN Morpho theseus theseus Deyrolle, 1860 TL: Colombia M T Morpho theseus claritae Neild, 2008 ref.266 Morpho theseus heraldica Niepelt, 1927 Costa Rica & Panama M T S Morpho theseus juturna A. Butler, 1870 TL: Colombia M T Morpho theseus pacificus E. Krüger, 1925 TL: Colombia M T Morpho theseus perlmani Neild, 2001 TL: Venezuela M T Morpho theseus triangulifera Le Moult & Réal, 1962 TL: Ecuador M T Morpho theseus yaritanus Fruhstorfer, 1913 TL: Venezuela M T Morpho theseus [n. ssp.] Llorente & Luis, MS Mexico Morpho theseus chirripoensis Schäffler, Frankenbach & Dufek, 2011 TL: not specified Morpho uraneis H. Bates, 1865 iN M T Morpho zephyritis A. Butler, 1873 TL: Peru M T Tribe Brassolini Bia Hübner, [1819] Bia actorion (Linnaeus, 1763) iN Bia actorion actorion (Linnaeus, 1763) TL: Indiis M HS Bia actorion decaerulea Weymer, 1911 TL: [Brazil (Am)] M T Bia actorion rebeli Bryk, 1953 TL: Peru M T S Bia peruana Röber, 1904 TL: [Peru] M T Blepolenis Röber, 1906 Blepolenis bassus (C. Felder & R. Felder, 1867) iN M T Blepolenis batea (Hübner, [1821]) iN Blepolenis batea batea (Hübner, [1821]) TL: [Brazil] M T Blepolenis batea didymaon (C. Felder & R. Felder, 1867) TL: Brazil M T Blepolenis batea praegrandis (Fruhstorfer, 1907) TL: Paraguay M T Blepolenis catharinae (Stichel, 1902) iN M T Brassolis Fabricius, 1807 Brassolis astyra Godart, [1824] iN Brassolis astyra astyra Godart, [1824] TL: Brazil M T Brassolis astyra philocala Stichel, 1904 TL: Brazil (RS) M T Brassolis haenschi Stichel, 1902 iN Brassolis haenschi haenschi Stichel, 1902 TL: Ecuador Brassolis haenschi maritimus Stichel, 1903 TL: Venezuela M T Brassolis haenschi rufescens Rothschild, 1916 TL: Peru M T Brassolis isthmia H. Bates, 1864 Small-spotted Owl-Butterfly Guatemala to Colombia; Atlantic slope iN M T S Brassolis granadensis Stichel, 1902 Colombian Owl-Butterfly iN Brassolis granadensis granadensis Stichel, 1902 NW Costa Rica to W Ecuador; Pacific slope M T S I Brassolis granadensis boliviana Rothschild, 1916 TL: Bolivia M T Brassolis sophorae (Linnaeus, 1758) iN Brassolis sophorae sophorae (Linnaeus, 1758) TL: America M T HS Brassolis sophorae ardens Stichel, 1903 TL: Peru M T Brassolis sophorae dinizi R.F. d'Almeida, 1956 TL: Brazil (Pb) M T Brassolis sophorae laurentii Stichel, 1925 TL: Brazil (MG) M T Brassolis sophorae luridus Stichel, 1902 TL: Colombia M T Brassolis sophorae philomela Stichel, 1925 TL: Ecuador M T Brassolis sophorae vulpeculus Stichel, 1902 TL: Paraguay M T Caligo Hübner, [1819] Caligo arisbe Hübner, [1822] iN Caligo arisbe arisbe Hübner, [1822] TL: ? M T Caligo arisbe fulgens Rothschild, 1916 TL: Brazil (RJ/SP) M T Caligo atreus (Kollar, 1850) iN Caligo atreus atreus (Kollar, 1850) TL: Colombia M T Caligo atreus agesilaus H. Druce, 1902 TL: Ecuador M T Caligo atreus ajax (E. Doubleday, [1849]) TL: Venezuela; "Ecuador" M T Caligo atreus dentina H. Druce, 1874 TL: Peru M T Caligo atreus dionysos Fruhstorfer, 1912 [Chiapas?], Costa Rica & Panama M T S1 S2 L I1 I2 Caligo bellerophon Stichel, 1903 iN M T Caligo beltrao (Illiger, 1801) iN M S Caligo brasiliensis (C. Felder, 1862) iN Caligo brasiliensis brasiliensis (C. Felder, 1862) TL: Brazil (RJ) M T Caligo brasiliensis caesius Stichel, 1903 TL: Venezuela Caligo brasiliensis galba (Deyrolle, 1874) TL: Colombia M T Caligo brasiliensis minor Kaye, 1904 TL: Trinidad M T Caligo brasiliensis morpheus Stichel, 1903 TL: Ecuador; Colombia M T Caligo brasiliensis sulanus Fruhstorfer, 1904 SE Mexico to Panama M T S1 S2 L I1 I2 Caligo euphorbus (C. Felder & R. Felder, 1862) iN Caligo euphorbus euphorbus (C. Felder & R. Felder, 1862) TL: Brazil (Am) M T Caligo euphorbus menoetius Staudinger, 1887 TL: Brazil (Am) M T Caligo eurilochus (Cramer, 1775) iN Caligo eurilochus eurilochus (Cramer, 1775) TL: Surinam M T Caligo eurilochus delectans Joicey & Kaye, 1917 TL: Venezuela M T Caligo eurilochus livius Staudinger, [1886] TL: "Panama"; Peru; Brazil (Pa) M T Caligo eurilochus mattogrossensis Ribeiro, 1931 TL: Brazil (Rd) M T Caligo eurilochus pallidus Fruhstorfer, 1912 TL: Bolivia M T Caligo idomeneus (Linnaeus, 1758) iN Caligo idomeneus idomeneus (Linnaeus, 1758) TL: S. Amer. M T HS Caligo idomeneus ariphron Fruhstorfer, 1910 TL: Brazil (ES) M T Caligo idomeneus idomenides Fruhstorfer, 1903 TL: Peru M T Caligo idomeneus rhoetus Staudinger, [1886] TL: Brazil (Pa/Am) M T Caligo illioneus (Cramer, 1775) iN Caligo illioneus illioneus (Cramer, 1775) TL: Surinam M T Caligo illioneus oberon A. Butler, 1870 Costa Rica & Panama M T S1 S2 L I Caligo illioneus pampeiro Fruhstorfer, 1904 TL: Paraguay M T Caligo illioneus pheidriades Fruhstorfer, 1912 TL: Bolivia M T Caligo illioneus praxsiodus Fruhstorfer, 1912 TL: Peru M T Caligo martia (Godart, [1824]) iN Caligo oberthurii (Deyrolle, 1872) iN Caligo oberthurii oberthurii (Deyrolle, 1872) TL: Colombia M T Caligo oberthurii phokilides Fruhstorfer, 1912 TL: Peru Caligo oedipus Stichel, 1903 Fruhstorfer's Owl-Butterfly iN Caligo oedipus oedipus Stichel, 1903 Panama to Colombia M T S1 S2 L I Caligo oedipus fruhstorferi Stichel, 1904 SE Mexico to Honduras M T S L Caligo oileus C. Felder & R. Felder, 1861 iN Caligo oileus oileus C. Felder & R. Felder, 1861 TL: Venezuela; Colombia M T Caligo oileus scamander (Boisduval, 1870) Guatemala to Panama M T S Caligo oileus umbratilis Stichel, 1903 TL: Peru M T Caligo placidianus Staudinger, 1887 iN M T S Caligo prometheus (Kollar, 1850) iN Caligo prometheus prometheus (Kollar, 1850) TL: Colombia M T Caligo prometheus atlas Röber, 1904 TL: Ecuador M T Caligo superbus Staudinger, 1887 iN Caligo superbus superbus Staudinger, 1887 TL: Peru M T S Caligo superbus agamemnon Weymer, 1890 TL: Ecuador M T Caligo suzanna (Deyrolle, 1872) iN M T Caligo telamonius (C. Felder & R. Felder, 1862) iN Caligo telamonius telamonius (C. Felder & R. Felder, 1862) TL: Colombia M T Caligo telamonius memnon (C. Felder & R. Felder, 1867) E & SW Mexico to Nicaragua M T S L I Caligo telamonius menus Fruhstorfer, 1903 Costa Rica to Colombia M T S1 S2 L I1 I2 Caligo telamonius modestus Niepelt, 1934 TL: Surinam Caligo teucer (Linnaeus, 1758) iN Caligo teucer teucer (Linnaeus, 1758) TL: America M T HS Caligo teucer ecuadora Joicey & Kaye, 1917 TL: Ecuador M T L Caligo teucer insulanus Stichel, 1904 TL: Trinidad M T Caligo teucer japetus Stichel, 1903 TL: Paraguay M T Caligo teucer nubilus Fruhstorfer, 1907 TL: Guyana M T Caligo teucer obidonus Fruhstorfer, 1904 TL: Brazil (Pa) M T Caligo teucer phorkys Fruhstorfer, 1912 TL: Bolivia M T Caligo teucer semicaerulea Joicey & Kaye, 1917 TL: Peru M T Caligo uranus Herrich-Schäffer, 1850 Gold-edged Owl-Butterfly E Mexico to Honduras iN M T S L Caligo zeuxippus H. Druce, 1902 iN Caligo zeuxippus zeuxippus H. Druce, 1902 TL: Ecuador M T Caligo zeuxippus obscurus Niepelt, 1934 TL: Colombia M T Caligopsis Seydel, 1924 Caligopsis seleucida (Hewitson, 1877) iN Caligopsis seleucida seleucida (Hewitson, 1877) TL: Bolivia M T Caligopsis seleucida juruana (Fruhstorfer, 1912) TL: Brazil (Ac) M T Caligopsis [n. sp.] Lamas, MS Peru M T Catoblepia Stichel, 1901 Catoblepia amphirhoe (Hübner, [1825]) iN M T Catoblepia berecynthia (Cramer, 1777) iN Catoblepia berecynthia berecynthia (Cramer, 1777) TL: Surinam M T S Catoblepia berecynthia adjecta Stichel, 1906 TL: Bolivia M T S Catoblepia berecynthia berecynthina (Hopffer, 1874) TL: Peru M T Catoblepia berecynthia halli Bristow, 1981 TL: Guyana M T Catoblepia berecynthia luxuriosus Stichel, 1902 TL: Colombia M T Catoblepia berecynthia meridaensis Bristow, 1981 TL: Venezuela Catoblepia berecynthia midas Stichel, 1908 TL: Ecuador M T Catoblepia berecynthia unditaenia Fruhstorfer, 1907 TL: Brazil (RJ) M T Catoblepia generosa Stichel, 1902 TL: Ecuador M T Catoblepia orgetorix (Hewitson, 1870) Orange-rimmed Owl-Butterfly iN Catoblepia orgetorix orgetorix (Hewitson, 1870) Honduras & Nicaragua M T TI Catoblepia orgetorix championi Bristow, 1981 Nicaragua to Panama; Atlantic slope M T S1 S2 L I1 I2 Catoblepia orgetorix magnalis Stichel, 1902 TL: Ecuador M T Catoblepia orgetorix rothschildi Casagrande & Lamas, 2004 TL: Colombia M T Catoblepia soranus (Westwood, 1851) iN M T Catoblepia versitincta Stichel, 1901 iN Catoblepia versitincta versitincta Stichel, 1901 TL: Surinam; French Guiana M T Catoblepia versitincta mossi Bristow, 1981 TL: Brazil (Pa) M T Catoblepia xanthicles (Godman & Salvin, 1881) Xanthicles Owl-Butterfly iN Catoblepia xanthicles xanthicles (Godman & Salvin, 1881) Costa Rica & Panama M T S Catoblepia xanthicles belisar Stichel, 1904 TL: Bolivia M T Catoblepia xanthicles cyparissa Fruhstorfer, 1907 TL: Colombia M T Catoblepia xanthicles mercedensis Bristow, 1981 TL: Peru M T Catoblepia xanthicles occidentalis Bristow, 1981 TL: Ecuador M T Catoblepia xanthicles orientalis Bristow, 1981 TL: Peru M T Catoblepia xanthicles paraensis Bristow, 1981 TL: Brazil (Pa) M T Catoblepia xanthicles sosigenes (Fruhstorfer, 1913) TL: French Guiana M T Catoblepia xanthus (Linnaeus, 1758) iN Catoblepia xanthus xanthus (Linnaeus, 1758) TL: ? Catoblepia xanthus rivalis Niepelt, 1911 TL: Ecuador M T Dasyophthalma Westwood, 1851 Dasyophthalma creusa (Hübner, [1821]) iN Dasyophthalma creusa creusa (Hübner, [1821]) TL: Brazil M S Dasyophthalma creusa baronesa Stichel, 1904 TL: Brazil (ES) M T S Dasyophthalma geraensis Rebel, 1922 iN M T Dasyophthalma rusina (Godart, [1824]) iN Dasyophthalma rusina rusina (Godart, [1824]) TL: Brazil M T Dasyophthalma rusina delanira Hewitson, 1862 TL: Brazil M T Dasyophthalma rusina principesa Stichel, 1904 TL: Brazil (ES) M T Dasyophthalma vertebralis A. Butler, 1869 TL: Brazil ("Pa") M T S Dynastor E. Doubleday, [1849] Dynastor darius (Fabricius, 1775) iN Dynastor darius darius (Fabricius, 1775) TL: Brazil ([RJ]) M T Dynastor darius anaxarete (Cramer, 1776) TL: Surinam M T Dynastor darius faenius Fruhstorfer, 1912 TL: Brazil (RS) M T Dynastor darius ictericus Stichel, 1904 TL: Paraguay M T Dynastor darius stygianus A. Butler, 1872 S Mexico to W Ecuador M T S L Dynastor macrosiris (Westwood, 1851) iN Dynastor macrosiris macrosiris (Westwood, 1851) TL: French Guiana M T Dynastor macrosiris hannibal Oberthür, 1881 TL: Colombia M T Dynastor macrosiris pharnaces Stichel, 1908 TL: Bolivia Dynastor macrosiris strix (H. Bates, 1864) E Mexico, Guatemala, Costa Rica M T S L Dynastor napoleon E. Doubleday, [1849] iN M T Eryphanis Boisduval, 1870 Eryphanis aesacus (Herrich-Schäffer, 1850) Double-spotted Owl-Butterfly E & S Mexico to Honduras & Nicaragua iN M T S L Eryphanis bubocula (A. Butler, 1872) Bubocula Owl-Butterfly Costa Rica to Colombia and Amazon Basin iN M T S Eryphanis automedon (Cramer, 1775) iN Eryphanis automedon automedon (Cramer, 1775) TL: Surinam M T Eryphanis automedon amphimedon (C. Felder & R. Felder, 1867) TL: Brazil M T Eryphanis automedon cheiremon Fruhstorfer, 1912 TL: Bolivia M T Eryphanis lycomedon (C. Felder & R. Felder, 1862) Split-spotted Owl-Butterfly Guatemala, Costa Rica, Colombia, Ecuador, Bolivia, southern Brazil iN M T S L Eryphanis gerhardi (Weeks, 1902) iN M T Eryphanis reevesii (E. Doubleday, [1849]) iN Eryphanis reevesii reevesii (E. Doubleday, [1849]) TL: [Brazil] M T Eryphanis reevesii pusillus Stichel, 1904 TL: Paraguay M T Eryphanis zolvizora (Hewitson, 1877) iN Eryphanis zolvizora isabelae Neild & De Sousa, 2014 Cordillera de la Costa in N Venezuela Eryphanis zolvizora reyi Bristow, Neild, De Sousa & Huertas, 2014 Cordillera de Mérida in NW Venezuela & N Colombia Eryphanis zolvizora opimus Staudinger, 1887 E slopes of W Andes in Colombia Eryphanis zolvizora casagrande Bristow, 2014 SW Colombia & NW Ecuador Eryphanis zolvizora greeneyi Penz & DeVries, 2008 E Ecuador Eryphanis zolvizora chachapoya Blandin, 2014 N & central Peru M T Eryphanis zolvizora inca Blandin, 2014 central & S Peru Eryphanis zolvizora zolvizora (Hewitson, 1877) Bolivia M T Mielkella Casagrande, 1982 Mielkella singularis (Weymer, 1907) Singular Owl-Butterfly S Mexico (Chiapas), Guatemala iN M T S Opoptera Aurivillius, 1882 Opoptera staudingeri (Godman & Salvin, 1894) Staudinger's Owl-Butterfly iN Opoptera staudingeri staudingeri (Godman & Salvin, 1894) Costa Rica & W Panama M T TI L Opoptera staudingeri mexicana J. Maza & R. G. Maza, 1989 S Mexico (Chiapas), Guatemala M T Opoptera aorsa (Godart, [1824]) iN Opoptera aorsa aorsa (Godart, [1824]) TL: Brazil M T Opoptera aorsa colombiana (Rothschild, 1916) TL: Colombia M T Opoptera aorsa fuscata Stichel, 1908 TL: Brazil (Am) M T Opoptera aorsa hilaris Stichel, 1901 TL: Ecuador Opoptera arsippe (Hopffer, 1874) iN Opoptera arsippe arsippe (Hopffer, 1874) TL: Peru Opoptera arsippe bracteolata Stichel, 1901 TL: Bolivia M T Opoptera arsippe [n. ssp.] Lamas, MS Peru Opoptera fruhstorferi (Röber, 1896) iN M T S Opoptera sulcius (Staudinger, 1887) iN M T Opoptera syme (Hübner, [1821]) iN M T Opoptera [n. sp.] Casagrande, MS Peru M T Opsiphanes E. Doubleday, [1849] Opsiphanes bogotanus Distant, 1875 iN Opsiphanes bogotanus bogotanus Distant, 1875 TL: Colombia M T Opsiphanes bogotanus alajuela Bristow, 1991 Costa Rica; cloud forests 700-1600m M T S Opsiphanes bogotanus blandini Bristow, 1991 TL: Ecuador M T Opsiphanes bogotanus phrataphernes Fruhstorfer, 1912 TL: Colombia M T Opsiphanes blythekitzmillerae Austin & A. Warren, 2007 Minerva's Owl-Butterfly NW Mexico iN M T S L Ha Opsiphanes boisduvallii E. Doubleday, [1849] Orange Owl-Butterfly E & W Mexico (include. BCS) to Nicaragua iN M T S L I Opsiphanes camena Staudinger, [1886] iN M T S Opsiphanes cassiae (Linnaeus, 1758) iN Opsiphanes cassiae cassiae (Linnaeus, 1758) TL: America M T HS Opsiphanes cassiae castaneus Stichel, 1904 Honduras to Colombia M T S L Opsiphanes cassiae crameri C. Felder & R. Felder, 1862 TL: Brazil (Ba) M T Opsiphanes cassiae kleisthenes Fruhstorfer, 1912 TL: Colombia M T Opsiphanes cassiae mexicana Bristow, 1991 SE Mexico, Guatemala M T S L Opsiphanes cassiae rubigatus Stichel, 1904 TL: Ecuador M T S Opsiphanes cassiae strophios Fruhstorfer, 1907 TL: Bolivia M T Opsiphanes cassina C. Felder & R. Felder, 1862 iN Opsiphanes cassina cassina C. Felder & R. Felder, 1862 TL: Brazil (Am) M T Opsiphanes cassina barkeri Bristow, 1991 TL: Ecuador M T Opsiphanes cassina caliensis Bristow, 1991 TL: Colombia M T Opsiphanes cassina chiriquensis Stichel, 1902 [= aiellae], Nicaragua through Panama M T S L Opsiphanes cassina fabricii (Boisduval, 1870) E & W Mexico to Honduras & El Salvador M T S L Opsiphanes cassina merianae Stichel, 1902 TL: Surinam M T Opsiphanes cassina milesi Bristow, 1991 TL: Brazil (Pa) M T Opsiphanes cassina notanda Stichel, 1904 TL: Peru M T Opsiphanes cassina numatius Fruhstorfer, 1912 TL: Colombia M T Opsiphanes invirae (Hübner, [1808]) iN Opsiphanes invirae invirae (Hübner, [1808]) TL: ? Opsiphanes invirae aequatorialis Stichel, 1902 TL: Ecuador M T Opsiphanes invirae agasthenes Fruhstorfer, 1907 TL: Peru M T Opsiphanes invirae amplificatus Stichel, 1904 TL: Paraguay M T Opsiphanes invirae camposi Bristow, 1991 TL: Ecuador M T Opsiphanes invirae cuspidatus Stichel, 1904 Nicaragua to W Panama M T S Opsiphanes invirae intermedius Stichel, 1902 TL: Brazil (Am) M T Opsiphanes invirae pseudophilon Fruhstorfer, 1907 TL: Brazil (ES) M T Opsiphanes invirae remoliatus Fruhstorfer, 1907 TL: Brazil (SC?) M T Opsiphanes invirae relucens Fruhstorfer, 1907 S Mexico (Chiapas) to Honduras M T S Opsiphanes invirae roraimaensis Bristow, 1991 TL: Guyana M T Opsiphanes invirae sieberti Bristow, 1991 TL: French Guiana M T Opsiphanes invirae sticheli Röber, 1906 E Panama (CZ) to W Venezuela M S Opsiphanes jacobsorum I. Chacón & Montero, 2012 ref.183 Opsiphanes mutatus Stichel, 1902 iN Opsiphanes mutatus mutatus Stichel, 1902 TL: Ecuador M T Opsiphanes mutatus farrago Stichel, 1903 TL: Peru M T Opsiphanes mutatus parodizi Bristow, 1991 TL: Bolivia M T S Opsiphanes tiberius Attal & Blandin, 2020 ref.193 Opsiphanes quiteria (Stoll, 1780) iN Opsiphanes quiteria quiteria (Stoll, 1780) TL: Surinam M T Opsiphanes quiteria angostura Bristow, 1979 TL: Ecuador M T Opsiphanes quiteria badius Stichel, 1902 Panama (CZ) to Colombia & Venezuela M T S L Opsiphanes quiteria bolivianus Stichel, 1902 TL: Bolivia M T Opsiphanes quiteria cardenasi Bristow, 1991 TL: Brazil (Pa) M T Opsiphanes quiteria cauca Röber, 1906 TL: Colombia M T Opsiphanes quiteria erebus Röber, 1927 TL: Peru Opsiphanes quiteria meridionalis Staudinger, 1887 TL: Brazil ([SC]) M T Opsiphanes quiteria phylas Fruhstorfer, 1912 TL: Colombia M T Opsiphanes quiteria quaestor Stichel, 1902 TL: Ecuador M T Opsiphanes quiteria quirinalis Staudinger, 1887 TL: Peru M T Opsiphanes quiteria quirinus Godman & Salvin, 1881 SE Mexico to Panama (Atlantic slope) M T S L Opsiphanes quiteria talamancensis Bristow, 1991 Costa Rica to W Panama M T S Opsiphanes sallei E. Doubleday, [1849] iN Opsiphanes sallei sallei E. Doubleday, [1849] TL: Venezuela M T Opsiphanes sallei colombiana Bristow, 1991 TL: Colombia M T S Opsiphanes sallei kennerleyi Bristow, 1991 TL: Peru M T S Opsiphanes sallei nicandrus Fruhstorfer, 1912 TL: Bolivia M T Opsiphanes tamarindi C. Felder & R. Felder, 1861 Narrow-banded Owl-Butterfly iN Opsiphanes tamarindi tamarindi C. Felder & R. Felder, 1861 E & W Mexico to Colombia M T S L I Opsiphanes tamarindi corrosus Stichel, 1904 TL: Ecuador M T S Opsiphanes tamarindi incolumis Stichel, 1904 TL: Peru M T Opsiphanes tamarindi latifascia Rothschild, 1916 TL: Peru M T Opsiphanes tamarindi xiphos Fruhstorfer, 1907 TL: Venezuela M T Opsiphanes zelotes Hewitson, 1873 iN Opsiphanes zelotes zelotes Hewitson, 1873 TL: Colombia M T Opsiphanes zelotes zelus Stichel, 1908 Panama M S Orobrassolis Casagrande, 1982 Orobrassolis ornamentalis (Stichel, 1906) iN M T Orobrassolis latusoris Penz & Simosen, 2011 TL: not specified Penetes E. Doubleday, [1849] Penetes pamphanis E. Doubleday, [1849] iN M T Selenophanes Staudinger, 1887 Selenophanes cassiope (Cramer, 1775) iN Selenophanes cassiope cassiope (Cramer, 1775) TL: Surinam; Guyana Selenophanes cassiope amplior Stichel, 1902 TL: Colombia M T Selenophanes cassiope andromeda Stichel, 1901 TL: Bolivia M T Selenophanes cassiope cassiopeia (Staudinger, [1886]) TL: Brazil (Am) M T Selenophanes cassiope guarany Casagrande, 1992 TL: Brazil (SP) M T Selenophanes cassiope haraposa Bristow, 1982 TL: Brazil (Pa) M T S Selenophanes cassiope mapiriensis Bristow, 1982 TL: Bolivia M T Selenophanes cassiope perenensis Bristow, 1982 TL: Peru M T Selenophanes cassiope placentia (Fruhstorfer, 1912) TL: Bolivia Selenophanes cassiope theognis Fruhstorfer, 1910 TL: Brazil (MT) M T Selenophanes josephus (Godman & Salvin, 1881) Josephus Owl-Butterfly iN Selenophanes josephus josephus (Godman & Salvin, 1881) Guatemala M T OI Selenophanes josephus excisus (Rothschild, 1916) TL: Colombia M T Selenophanes josephus excultus Stichel, 1902 E Panama to Colombia M T S Selenophanes supremus Stichel, 1901 TL: Ecuador Selenophanes supremus supremus Stichel, 1901 TL: Ecuador M T Selenophanes supremus ditatus Thieme, 1902 TL: Peru M S Aponarope Casagrande, 1982 Aponarope sutor (Stichel, 1916) TL: Brazil (MT) M T S Narope E. Doubleday, [1849] Narope albopunctum Stichel, 1904 TL: Peru M T Narope anartes Hewitson, 1874 iN M T S Narope cauca Casagrande, 2002 iN M T Narope cyllabarus Westwood, 1851 iN M T S Narope cyllarus Westwood, 1851 iN M T Narope cyllastros E. Doubleday, [1849] iN M T Narope cyllene C. Felder & R. Felder, 1859 iN M T Narope denticulatus Talbot, 1928 iN M T Narope guilhermei Casagrande, 1989 TL: Brazil (SC) M T S Narope marmorata Schaus, 1902 iN M T S Narope minor Casagrande, 2002 Small Owl-Butterfly S Mexico (VER, OAX), Guatemala M T OI G Narope nesope Hewitson, 1869 TL: Ecuador M T S Narope obidos Casagrande, 2002 TL: Brazil (Pa) M T S Narope panniculus Stichel, 1904 iN M T S Narope syllabus Staudinger, 1887 iN Narope syllabus syllabus Staudinger, 1887 TL: Peru M T S Narope syllabus stygius Staudinger, 1887 TL: Brazil (Am) M T S Narope testacea Godman & Salvin, 1878 Brown Owl-Butterfly S Mexico to Panama iN M T S Narope ybyra Casagrande, 2002 TL: Bolivia M T Tribe Elymniini Neorinella Martins, Kucera-Santos, Vieira & Fragoso, 1993 a fossil genus Neorinella garciae Martins, Kucera-Santos, Vieira & Fragoso, 1993 Fossil from Brazil (SP) Tribe Haeterini Cithaerias Hübner, [1819] Cithaerias andromeda (Fabricius, 1775) iN Cithaerias andromeda andromeda (Fabricius, 1775) TL: India M T Cithaerias andromeda azurina (J. Zikán, 1942) TL: Brazil (Am) M T Cithaerias andromeda bandusia Staudinger, [1887] TL: Brazil (Am) M T Cithaerias andromeda esmeralda (E. Doubleday, 1845) TL: Brazil (Pa) M T Cithaerias pyropina (Salvin & Godman, 1868) iN Cithaerias pyropina pyropina (Salvin & Godman, 1868) TL: Peru M T S Cithaerias pyropina julia Willmott, 2015 ref.301 Cithaerias pyropina songoana (Langer, 1944) TL: Bolivia M T S Cithaerias pireta (Stoll, 1780) Rusted Clearwing-Satyr iN Cithaerias pireta pireta (Stoll, 1780) [= menander], Guatemala to Colombia M T S L Cithaerias pireta aurora (C. Felder & R. Felder, 1862) TL: [Peru] M T Cithaerias pireta magdalenensis Constantino, 1995 TL: Colombia M T Cithaerias aurorina (Weymer, 1910) iN Cithaerias aurorina aurorina (Weymer, 1910) TL: Upper Amazon M T S Cithaerias aurorina tambopata Lamas, 1998 TL: Peru M T S Cithaerias cliftoni Constantino, 1995 iN M T Cithaerias pyritosa (J. Zikán, 1942) iN M T Dulcedo R.F. d'Almeida, 1951 Dulcedo polita (Hewitson, 1869) Uncolored Clearwing-Satyr Nicaragua to Colombia iN M T S L Haetera Fabricius, 1807 Haetera macleannania (H. Bates, 1865) Reddish Clearwing-Satyr Costa Rica to W Ecuador iN M T TI S IL L Haetera piera (Linnaeus, 1758) iN Haetera piera sanguinolenta Constantino & Salazar, 2007 TL: Colombia M T Haetera piera piera (Linnaeus, 1758) TL: Indiis M HS Haetera piera diaphana Lucas, 1857 TL: Cuba M T Haetera piera negra C. Felder & R. Felder, 1862 TL: [Peru] M T Haetera piera pakitza Lamas, 1998 TL: Peru M T Haetera piera unocellata Weymer, 1910 TL: Bolivia M T Pierella Westwood, 1851 Pierella amalia Weymer, 1885 iN M T Pierella astyoche (Erichson, [1849]) iN Pierella astyoche astyoche (Erichson, [1849]) TL: Guyana M T Pierella astyoche bernhardina Bryk, 1953 TL: Brazil (Am) M T Pierella astyoche stollei Ribeiro, 1931 TL: Brazil (Rd) M T Pierella helvina (Hewitson, 1859) iN Pierella helvina helvina (Hewitson, 1859) TL: Colombia M T S Pierella helvina hymettia Staudinger, [1886] TL: Colombia Pierella helvina incanescens Godman & Salvin, 1877 Guatemala to Panama M T S L Pierella helvina ocreata Salvin & Godman, 1868 Panama to NW Colombia M T S L Pierella helvina pacifica Niepelt, 1924 TL: Colombia M T Pierella hortona (Hewitson, 1854) iN Pierella hortona hortona (Hewitson, 1854) TL: Brazil (Am) M T Pierella hortona albofasciata Rosenberg & Talbot, 1914 TL: Peru M T Pierella hortona [n. ssp.] Lamas, MS Peru Pierella hyalinus (Gmelin, [1790]) iN Pierella hyalinus hyalinus (Gmelin, [1790]) TL: Surinam M T Pierella hyalinus extincta Weymer, 1910 TL: Brazil (Am) M T S Pierella hyalinus schmidti Constantino, 1995 TL: Colombia M T Pierella hyalinus velezi Constantino, 1995 TL: Colombia M T Pierella hyceta (Hewitson, 1859) iN Pierella hyceta hyceta (Hewitson, 1859) TL: [Peru] M T Pierella hyceta ceryce (Hewitson, 1874) TL: Bolivia M T Pierella hyceta latona (C. Felder & R. Felder, 1867) TL: Colombia M T Pierella hyceta [n. ssp.#1] Lamas, MS TL: Peru Pierella hyceta [n. ssp.#2] Lamas, MS TL: Peru Pierella lamia (Sulzer, 1776) iN Pierella lamia lamia (Sulzer, 1776) TL: America M T Pierella lamia boliviana F. Brown, 1948 TL: Bolivia M T Pierella lamia chalybaea Godman, 1905 TL: Brazil (MT) M T S Pierella lena (Linnaeus, 1767) iN Pierella lena lena (Linnaeus, 1767) TL: S. Amer. M T Pierella lena brasiliensis (C. Felder & R. Felder, 1862) TL: [Peru] M T S Pierella lena salma Constantino, 2007 TL: Colombia M T Pierella lucia Weymer, 1885 iN M T Pierella luna (Fabricius, 1793) Moon Satyr iN Pierella luna luna (Fabricius, 1793) Nicaragua to Colombia M T S L Pierella luna lesbia Staudinger, 1887 TL: Colombia M S Pierella luna pallida (Salvin & Godman, 1868) Honduras & Nicaragua M T IL L Pierella luna rubecula Salvin & Godman, 1868 E & S Mexico to Honduras M T S L Pierella nereis (Drury, 1782) iN Pseudohaetera F. Brown, [1943] Pseudohaetera hypaesia (Hewitson, 1854) iN M T Pseudohaetera mimica (Rosenberg & Talbot, 1914) iN M T L Tribe Melanitini Manataria W. F. Kirby, [1902] Manataria maculata (Hopffer, 1874) White-spotted Satyr E & W Mexico to Colombia iN Manataria maculata maculata (Hopffer, 1874) E & W Mexico to Colombia M S HS L I Ha Manataria maculata daguana (Neustetter, 1929) TL: Colombia M T Manataria hercyna (Hübner, [1821]) iN Manataria hercyna hyrnethia Fruhstorfer, 1912 TL: Bolivia; Peru M T Manataria hercyna distincta (Lathy, 1918) TL: French Guiana M T Manataria hercyna hercyna (Hübner, [1821]) TL: [Brazil] M OI Tribe Satyrini, Subtribe Lethina Lethe Hübner, [1819] Lethe anthedon (A. Clark, 1936) Northern Pearly-eye iN Lethe anthedon borealis (A. Clark, 1936) TL: not specified M T S L Lethe anthedon anthedon (A. Clark, 1936) TL: USA (NY) M T S L1 L2 L3 L4 Lethe creola (Skinner, 1897) Creole Pearly-eye iN M T S L1 L2 Lethe portlandia (Fabricius, 1781) Southern Pearly-eye iN Lethe portlandia missarkae J. Heitzman & dos Passos, 1974 TL: USA (Ar) M S L I H Lethe portlandia portlandia (Fabricius, 1781) TL: USA (Ga) M S L Lethe portlandia floralae J. Heitzman & dos Passos, 1974 TL: USA (Fl) M T S L Lethe eurydice (Linnaeus, 1763) Eyed Brown iN Lethe eurydice fumosus (Leussler, 1916) TL: USA (Ne) M T S Lethe eurydice eurydice (Linnaeus, 1763) TL: USA (Pa) M T S L Lethe appalachia R. Chermock, 1947 Appalachian Brown iN Lethe appalachia leeuwi Gatrelle & Arbogast, 1974 TL: USA (Mi) M T L Lethe appalachia appalachia R. Chermock, 1947 TL: USA (NC) M S L1 L2 Ha Tribe Satyrini, Subtribe Calistina Calisto Hübner, [1823] Calisto herophile Hübner, [1823] Cuban Calisto iN Calisto herophile herophile Hübner, [1823] much of Cuba and Isla de Juventud; lowlands M S IL L I Calisto herophile parsonsi Clench, 1943 Cuba: Sierra de Escambray, 500-1000m M T IL Calisto bruneri Michener, 1949 Bruner's Calisto Nipe-Sagua-Baracoa Mts., E Cuba iN M T IL I Calisto smintheus M. Bates, 1935 Smintheus Calisto [= biocellatus, brocheri], Sierra de Maestra, E Cuba, 1000-2500m iN M T IL L I Calisto muripetens M. Bates, 1939 Muripetens Calisto C Cuba; ca. 1000m M T S I Calisto bradleyi Munroe, 1951 Bradley's Calisto W Cuba; 500m M S Calisto brochei Torre, 1973 Broche's Calisto Nipe-Sagua-Baracoa Mts., E Cuba iN M I Calisto israeli Torre, 1973 Israeli Calisto Nipe-Sagua-Baracoa Mts., E Cuba Calisto occulta Núñez, 2012 Cryptic Calisto Nipe-Sagua-Baracoa Mts., E Cuba M I Calisto torrei Núñez, 2013 Guamuhaya Mts., Cuba M T L I Calisto aquilum Núñez, 2013 Guamuhaya Mts., Cuba iN Calisto aquilum aquilum Núñez, 2013 Guamuhaya Mts., Cuba M T L Calisto aquilum occidentalis Núñez, 2013 Guaniguanico Mts., Cuba Calisto dissimulatum Núñez, 2013 Sierra Cristal, Nipe-Sagua-Baracoa & middle of Sierra Maestra, Cuba M T L I Calisto sibylla M. Bates, 1934 Woodland Calisto Bahamas: New Providence, N & S Andros M T S Calisto apollinis M. Bates, 1934 Cuban Calisto Bahamas M T S L Calisto zangis (Fabricius, 1775) Jamaican Calisto Jamaica iN M T S1 S2 L Calisto nubila Lathy, 1899 Puerto Rican Calisto Puerto Rico iN M T S1 S2 L I Calisto anegadensis D. Smith, L. Miller & McKenzie, 1991 Anegada Calisto Anegada (British Virgin Islands) M T Calisto pulchella Lathy, 1899 Sugar cane Calisto iN Calisto pulchella pulchella Lathy, 1899 Hispaniola (widespread, except in Cordillera Central; see below) M T S I Calisto pulchella darlingtoni Clench, 1943 Cordillera Central, W Dominican Republic; montane M T S Calisto raburni Gali, 1985 Raburn's Calisto NE of El Aguacate, Independencia, Dominican Republic M T S Calisto arcas M. Bates, 1939 Tawny-washed Calisto Cordillera Central, Pico Duarte massif, Dominican Republic M T S Calisto chrysaoros M. Bates, 1935 White-banded Calisto Massif de la Hotte, Haiti; Sierra de Baoruco, Dom. Rep. iN M T S L Calisto galii A. Schwartz, 1983 Gali's Calisto TL: Dominican Rep. Calisto galii galii A. Schwartz, 1983 Cordillera Central, W Dominican Republic M T S L Calisto galii choneupsilon A. Schwartz, 1985 Sierra de Neiba, Dominican Republic M T Calisto clenchi A. Schwartz & Gali, 1984 Clench's Calisto Massif de la Selle (Haiti-Dominican Republic) M T S Calisto wetherbeei A. Schwartz & F. González, 1988 Wetherbee's Calisto Cordillera Central, W Dominican Republic M T Calisto neochma A. Schwartz, 1991 Neochma Calisto Dominican Republic Calisto clydoniata A. Schwartz & Gali, 1984 Yellow-stained Calisto Sierra de Neiba, Dominican Republic M T Calisto archebates (Ménétriés, 1832) Yellow-banded Calisto Massif de la Selle (Haiti-D.R.), W Sierra de Baoruco, Dom. Rep. iN M S L I Calisto eleleus M. Bates, 1935 Eleleus Calisto Hispaniola; most abundant in Sierra de Baoruco iN M T S Calisto loxias M. Bates, 1935 Dark Calisto Massif de la Hotte, Haiti; nr. Las Abejas, SW Dom. Republic M T S Calisto woodsi K. Johnson & Hedges, 1998 Woods' Calisto Massif de la Hotte, Haiti M OI Calisto tasajera F. González, A. Schwartz & Wetherbee, 1991 Tasajera Calisto Cordillera Central, Dominican Republic M S L I Calisto schwartzi Gali, 1985 Schwartz's Calisto nr. Aceitillar, Sierra de Baoruco, Pedernales, SW Dom. Republic M T S Calisto lyceius M. Bates, 1935 Saona Calisto Isla Saona & Isla Catalina, Dominican Republic iN M T S Calisto franciscoi Gali, 1985 Francisco's Calisto SW Dom. Republic (Valle de Neiba & Barahona Pen. to Azua) iN M T S Calisto crypta Gali, 1985 Cryptic Calisto vic. Monte Cristi, Dominican Republic iN M T S L I Ha Calisto confusa Lathy, 1899 Confused Calisto Hispaniola (widespread) iN M T S L I Calisto debarriera Clench, 1943 Debarriera Calisto [= gonzalezi, montana, neiba] Haiti, Dom. Rep. (Sierra de Nieba) iN M T1 T2 S Calisto hysius (Godart, [1824]) Godart's Calisto iN Calisto hysius hysius (Godart, [1824]) Hispaniola (S palaeoisland) M T S L I Calisto hysius aleucosticha Correa & A. Schwartz, 1986 N Cordillera Central, Dominican Republic M T Calisto batesi Michener, 1943 Bates' Calisto Hispaniola (most common in N Dominican Republic) iN M T S L Calisto pauli K. Johnson & Hedges, 1998 Paul's Calisto Massif de la Hotte, Haiti M OI Calisto obscura Michener, 1943 Obscure Calisto Hispaniola (widespread) iN M T S L I Calisto ainigma K. Johnson, Quinter & Matusik, 1987 Enigmatic Calisto Cordillera Central, Dominican Republic, possilby an aberration of C. obscura M T Calisto grannus M. Bates, 1939 Grannus Calisto iN Calisto grannus dilemma F. González, 1987 nr. Constanza, La Vega, Dominican Republic M T S L I Calisto grannus dystacta F. González, 1987 Loma Leonor, Santiago Rodriguez, Dominican Republic M T Calisto grannus micheneri Clench, 1944 Cordillera Central, Dominican Republic iN M T S Calisto grannus micrommata A. Schwartz & Gali, 1984 S range Sierra de Neiba, SW Dominican Republic, 1600-1900m M T Calisto grannus grannus M. Bates, 1939 Cordillera Central, Dominican Republic, 1160-2300m M T S L I Calisto grannus amazona F. González, 1987 vic. Pico Duarte, Cordillera Central, Dom. Rep., 1200-1900m M T S Calisto grannus phoinix F. González, 1987 La Palma, E Cordillera Central, La Vega, Dominican Republic M T S Calisto grannus sommeri A. Schwartz & Gali, 1984 E Massif de la Selle, Haiti, to W Sierra de Baoruco, Dom. Rep. M T S Calisto thomasi K. Johnson & Hedges, 1998 Thomas' Calisto Massif de la Hotte, Haiti M OI Calisto tragius M. Bates, 1935 Tragic Calisto Massif de la Selle (Haiti); W Sierra de Baoruco, Dom. Republic M T S Calisto hendersoni Gali, 1985 Henderson's Calisto nr. El Limon, Independencia, Dominican Republic M T S Tribe Satyrini, Subtribe Gyrocheilina Gyrocheilus A. Butler, 1867 Gyrocheilus patrobas (Hewitson, 1862) Red-bordered Satyr iN Gyrocheilus patrobas tritonia (W. H. Edwards, 1874) C & SE AZ, SW NM, far W TX, NW Mexico M T S L I Gyrocheilus patrobas patrobas (Hewitson, 1862) C & S Mexico M T S L Gyrocheilus [n. sp.] N/A, N/A Rust-bordered Satyr NE Mexico M S Tribe Satyrini, Subtribe Pronophilina Pronophila E. Doubleday, [1849] Pronophila unifasciata Lathy, 1906 iN Pronophila unifasciata unifasciata Lathy, 1906 TL: Ecuador M T S Pronophila unifasciata deverra Thieme, 1907 TL: Ecuador Pronophila unifasciata brennus Thieme, 1907 TL: Colombia M T Pronophila unifasciata bogotensis Jurriaanse, 1926 TL: Colombia M T S Pronophila unifasciata australis Hayward, 1949 TL: Argentina M T Pronophila unifasciata donachui Adams & Bernard, 1977 TL: Colombia M T Pronophila unifasciata nanegalito Pyrcz, 2000 TL: Ecuador Pronophila unifasciata deceptoria Pyrcz, 2005 TL: not specified Pronophila unifasciata eugeni Pyrcz, 2005 TL: Colombia Pronophila unifasciata kennethi Pyrcz, 2005 TL: Ecuador M S Pronophila unifasciata antioquiana Pyrcz, 2005 TL: Peru Pronophila unifasciata [n. ssp.] Pyrcz, MS Bolivia Pronophila juliani Adams & Bernard, 1977 iN M T Pronophila timanthes Salvin, 1871 Timanthes Satyr Costa Rica to W Ecuador iN M T OI S L Pronophila obscura A. Butler, 1868 iN Pronophila obscura obscura A. Butler, 1868 TL: Venezuela M T Pronophila obscura cuchillaensis Viloria & Camacho, 1999 TL: Venezuela Pronophila epidipnis Thieme, 1907 iN Pronophila epidipnis epidipnis Thieme, 1907 TL: Venezuela M T Pronophila epidipnis orchewitsoni Adams & Bernard, 1979 TL: Colombia M T S Pronophila epidipnis lathyi Pyrcz, 2000 TL: Ecuador Pronophila epidipnis orcidipnis Pyrcz & Viloria, 2007 ref.285 Pronophila epidipnis perplexa Pyrcz, 2004 TL: Peru Pronophila epidipnis cana Pyrcz, MS TL: Peru Pronophila epidipnis [n. ssp.#1] Pyrcz, MS TL: Colombia Pronophila epidipnis [n. ssp.#2] Pyrcz, MS TL: Bolivia Pronophila tremocrata Pyrcz, 2004 iN M S Pronophila cordillera Westwood, 1851 iN Pronophila cordillera cordillera Westwood, 1851 TL: [Bolivia] M T Pronophila cordillera variabilis A. Butler, 1873 TL: Peru M T S Pronophila cordillera colocasia Thieme, 1907 TL: Peru M T Pronophila cordillera [n. ssp.] Pyrcz & Boyer, MS Peru Pronophila orcus (Latreille, [1813]) iN Pronophila orcus orcus (Latreille, [1813]) TL: ["Peru"] M T S Pronophila orcus locuples Weymer, 1912 TL: Bolivia M S Pronophila orcus nepete Thieme, 1907 TL: Peru M T Pronophila thelebe E. Doubleday, [1849] iN M T Pronophila isobelae Pyrcz, 2000 TL: Ecuador Pronophila isobelae isobelae Pyrcz, 2000 TL: Ecuador Pronophila isobelae benevola Pyrcz, 2000 TL: Ecuador Pronophila attali Pyrcz, 2000 TL: Peru Pronophila rosenbergi Lathy, 1906 iN Pronophila rosenbergi rosenbergi Lathy, 1906 TL: Ecuador M T Pronophila rosenbergi puyango Pyrcz, 2000 TL: Ecuador Pronophila bernardi Pyrcz, 2004 iN Pronophila intercidona Thieme, 1907 iN Pronophila intercidona intercidona Thieme, 1907 TL: Ecuador M T Pronophila intercidona thelebina Thieme, 1907 TL: Bolivia M T Pronophila intercidona [n. ssp.] Pyrcz & Prieto, MS Colombia Pronophila margarita Pyrcz, 2000 TL: Ecuador Corades E. Doubleday, [1849] Corades enyo Hewitson, [1849] iN Corades enyo enyo Hewitson, [1849] TL: Venezuela M T Corades enyo almo Thieme, 1907 TL: Colombia M T L Corades enyo [n. ssp.#1] Pyrcz, MS TL: Ecuador Corades enyo [n. ssp.#2] Pyrcz, MS TL: Bolivia Corades enyo [n. ssp.#3] Pyrcz, MS TL: Ecuador Corades pax Watkins, 1939 TL: Venezuela M T Corades chelonis Hewitson, 1863 iN Corades chelonis chelonis Hewitson, 1863 TL: Colombia M T Corades chelonis rubeta Thieme, 1907 TL: Venezuela Corades chelonis rectilinea Pyrcz & Viloria, 2005 TL: Colombia Corades lactefusa Thieme, 1907 iN M S Corades iduna Hewitson, 1850 iN Corades iduna iduna Hewitson, 1850 TL: Bolivia M T Corades iduna procellaria Thieme, 1907 TL: Argentina Corades iduna marginalis A. Butler, 1873 TL: Peru M T S Corades iduna peruviana A. Butler, 1873 TL: Peru M T S Corades iduna gradosi Lamas, 2003 TL: Peru Corades iduna [n. ssp.] Pyrcz, MS Peru Corades pannonia Hewitson, 1850 iN Corades pannonia pannonia Hewitson, 1850 TL: Venezuela M T Corades pannonia ploas Thieme, 1907 TL: Venezuela; Colombia M T Corades pannonia condorita Lamas, [1997] TL: Peru Corades pannonia albomaculata Staudinger, [1887] TL: Peru M T S Corades pannonia [n. ssp.] Pyrcz, MS Ecuador Corades argentata A. Butler, 1868 iN M T S L Corades sareba Hewitson, 1850 iN Corades sareba sareba Hewitson, 1850 TL: Bolivia M T Corades sareba fusciplaga A. Butler, 1873 TL: Peru M T Corades cybele A. Butler, 1866 iN Corades cybele cybele A. Butler, 1866 TL: Colombia M T Corades cybele semiplena Thieme, 1907 TL: Peru M T Corades cybele [n. ssp.#1] Pyrcz, MS TL: Colombia Corades cybele [n. ssp.#2] Pyrcz, MS TL: Peru Corades cybele [n. ssp.#3] Pyrcz, MS TL: Peru Corades medeba Hewitson, 1850 iN Corades medeba medeba Hewitson, 1850 TL: Bolivia M T S L Corades medeba columbina Staudinger, 1894 TL: Colombia M T L Corades medeba pittieri Pyrcz, 2010 TL: Venezuela Corades melania Staudinger, 1894 iN Corades melania melania Staudinger, 1894 TL: Bolivia M T Corades melania carlos Pyrcz, 2006 TL: Venezuela Corades cistene Hewitson, 1863 iN Corades cistene cistene Hewitson, 1863 TL: [Bolivia] M T Corades cistene generosa Thieme, 1907 TL: Peru; Bolivia M T L Corades cistene [n. ssp.] Pyrcz, MS Ecuador Corades dymantis Thieme, 1907 iN Corades dymantis dymantis Thieme, 1907 TL: Colombia M T Corades dymantis [n. ssp.#1] Pyrcz, MS TL: Colombia Corades dymantis [n. ssp.#2] Pyrcz, MS TL: Ecuador Corades dymantis [n. ssp.#3] Pyrcz, MS TL: Ecuador Corades chirone Hewitson, 1863 iN M T Corades tripunctata Weymer, 1890 TL: Peru Corades tripunctata tripunctata Weymer, 1890 TL: Peru M T Corades tripunctata necrufa Pyrcz, 2004 TL: Peru Corades tripunctata malambo Pyrcz, 2006 TL: Venezuela Corades ulema Hewitson, 1850 iN Corades ulema ulema Hewitson, 1850 TL: Bolivia M T S L Corades ulema mirianae Lamas, [1997] TL: Peru Corades anfortas (Thieme, 1907) iN Corades anfortas anfortas (Thieme, 1907) TL: Peru M T Corades anfortas celestini Pyrcz, 2005 TL: Venezuela Corades tricordatus Hewitson, 1874 TL: Bolivia M T Corades callipolis Hewitson, 1874 TL: Bolivia M T Corades liliaceus Pyrcz, MS TL: Ecuador Corades [n. sp.#1] Pyrcz, MS TL: Peru Corades [n. sp.#2] Lamas, MS TL: Peru Corades [n. sp.#3] Pyrcz, MS TL: not specified Corades [n. sp.#3] [n. ssp.#1] Pyrcz, MS TL: Venezuela Corades [n. sp.#3] [n. ssp.#2] Pyrcz, MS TL: Venezuela Corades [n. sp.#4] Pyrcz, MS TL: Bolivia Daedalma Hewitson, 1858 Daedalma dinias Hewitson, 1858 iN Daedalma dinias dinias Hewitson, 1858 TL: Colombia M T Daedalma dinias salmoni Pyrcz, 2011 ref.281 Daedalma dinias emma Pyrcz & Greeney, 2011 ref.281 M L Daedalma dinias leticia Pyrcz, 2011 ref.281 Daedalma dinias romeliana Pyrcz & Prieto, 2011 ref.281 Daedalma dinias granadillas Pyrcz & Willmott, 2011 ref.281 Daedalma dinias radfordi Pyrcz & Willmott, 2011 TL: not specified Daedalma boliviana Staudinger, 1897 iN Daedalma boliviana boliviana Staudinger, 1897 TL: Bolivia; Peru M T Daedalma boliviana peruviana Pyrcz, 2004 TL: Peru Daedalma fraudata Pyrcz, 2004 TL: Peru M S Daedalma rubroreducta Pyrcz & Willmott, 2011 ref.281 Daedalma drusilla Hewitson, 1858 iN Daedalma drusilla drusilla Hewitson, 1858 TL: Colombia M T Daedalma drusilla tinta Pyrcz, 2011 ref.281 Daedalma drusilla krugeriana Pyrcz, 2011 ref.281 Daedalma inconspicua A. Butler, 1866 iN Daedalma inconspicua inconspicua A. Butler, 1866 TL: Ecuador M T Daedalma inconspicua blancae Pyrcz, 2011 ref.281 Daedalma inconspicua cuencana Pyrcz, 2011 ref.281 Daedalma inconspicua onorei Pyrcz, 2011 ref.281 Daedalma inconspicua orientalis Pyrcz, 2011 ref.281 Daedalma inconspicua variegata Pyrcz, 2011 ref.281 Daedalma inconspicua palacio Dognin, 1891 TL: Ecuador M T Daedalma inconspicua petitia Pyrcz, 2011 ref.281 Daedalma inconspicua tapichalaca Pyrcz & Willmott, 2011 ref.281 Daedalma vertex Pyrcz, 2004 TL: Peru Daedalma vertex vertex Pyrcz, 2004 TL: Peru M T Daedalma vertex milleri Pyrcz, 2011 ref.281 M T Daedalma parvomaculata E. Krüger, 1924 TL: Colombia M T S Daedalma adamsi D'Abrera, 1988 TL: Peru M T Daedalma eliza Pyrcz & Willmott, 2011 ref.281 iN Daedalma dognini Pyrcz, 2011 TL: Peru Daedalma dognini dognini Pyrcz, 2011 ref.281 Daedalma dognini mirianae Pyrcz, 2011 ref.281 M T Thiemeia Weymer, 1912 Thiemeia phoronea (E. Doubleday, [1849]) iN Thiemeia phoronea phoronea (E. Doubleday, [1849]) TL: [Venezuela] M T Thiemeia phoronea obscurata E. Krüger, 1924 TL: Colombia Thiemeia phoronea ortruda (Thieme, 1907) TL: Bolivia M T Junea Hemming, 1964 Junea dorinda (C. Felder & R. Felder, 1862) iN Junea dorinda dorinda (C. Felder & R. Felder, 1862) TL: Colombia M T Junea dorinda quasinegra Pyrcz, 2004 TL: Peru Junea dorinda whitelyi (H. Druce, 1876) TL: Peru M T S Junea dorinda [n. ssp.#1] Pyrcz, MS TL: Ecuador Junea dorinda [n. ssp.#2] Pyrcz, MS TL: Peru Junea doraete (Hewitson, 1858) iN Junea doraete doraete (Hewitson, 1858) TL: Colombia M T Junea doraete antissa (Thieme, 1907) TL: Peru Junea doraete gideon (Thieme, 1907) TL: Peru M T S Junea doraete enipeus (Thieme, 1907) TL: Bolivia M T Junea doraete [n. ssp.#1] Pyrcz & Clavijo, MS TL: Colombia Junea doraete [n. ssp.#2] Pyrcz & Clavijo, MS TL: Colombia Junea doraete [n. ssp.#3] Pyrcz, MS TL: Ecuador Junea doraete [n. ssp.#4] Pyrcz, MS TL: Ecuador Oxeoschistus A. Butler, 1867 Oxeoschistus protogenia (Hewitson, 1862) iN M T Oxeoschistus pronax (Hewitson, 1859) iN M T S Oxeoschistus puerta (Westwood, 1851) iN Oxeoschistus puerta submaculatus A. Butler & H. Druce, 1874 Costa Rica & Panama M T S L Oxeoschistus puerta pervius Thieme, 1907 TL: Colombia M T Oxeoschistus puerta puerta (Westwood, 1851) TL: "Colombia"; Venezuela M T Oxeoschistus puerta magna Pyrcz & Viloria, 2007 TL: Venezuela Oxeoschistus simplex A. Butler, 1868 iN Oxeoschistus simplex simplex A. Butler, 1868 TL: Colombia M T Oxeoschistus simplex fuscus R. Krüger, 1929 TL: Colombia M T Oxeoschistus simplex isolda Thieme, 1907 TL: Ecuador M T Oxeoschistus simplex triplex Pyrcz & Rodriguez, MS TL: Venezuela Oxeoschistus simplex duplex Godman, 1905 TL: Bolivia M T Oxeoschistus simplex garleppi Lamas, 2003 TL: Peru Oxeoschistus iphigenia Pyrcz, 2004 iN M T Oxeoschistus euriphyle A. Butler, 1872 Dot-banded Satyr Costa Rica & Panama iN M T S L Oxeoschistus hilara (H. Bates, 1865) Dot-banded Satyr iN Oxeoschistus hilara hilara (H. Bates, 1865) S Mexico (Chiapas) & Guatemala M T S L Oxeoschistus hilara [n. ssp.#1] Pyrcz, MS TL: Mexico Oxeoschistus hilara [n. ssp.#2] Pyrcz, MS TL: Honduras Oxeoschistus hilara [n. ssp.#3] Pyrcz & Marabuto, MS TL: Honduras Oxeoschistus cothonides Grose-Smith, 1896 Rust-patched Satyr Costa Rica & Panama iN M T S Oxeoschistus cothon Salvin, 1871 Lemon-patched Satyr Costa Rica & W Panama iN M T S L Oxeoschistus tauropolis (Westwood, [1850]) Yellow-patched Satyr iN Oxeoschistus tauropolis tauropolis (Westwood, [1850]) E Mexico to Panama M T S L Oxeoschistus tauropolis W Mexican segregate , SW Mexico M S Oxeoschistus tauropolis [n. ssp.] Pyrcz, MS Mexico Oxeoschistus leucospilos Staudinger, 1876 iN Oxeoschistus leucospilos leucospilos Staudinger, 1876 TL: Peru M T S Oxeoschistus leucospilos pugil (Thieme, 1907) TL: Bolivia M T Oxeoschistus romeo Pyrcz & Fratello, 2005 TL: not specified Oxeoschistus [n. sp.] Pyrcz, MS Guatemala Mygona Thieme, 1907 Mygona irmina (E. Doubleday, [1849]) iN Mygona irmina irmina (E. Doubleday, [1849]) TL: [Venezuela] M T Mygona irmina [n. ssp.#1] Pyrcz, MS TL: Colombia Mygona irmina [n. ssp.#2] Pyrcz, MS TL: Ecuador Mygona prochyta (Hewitson, 1862) iN M T Mygona chyprota (Grose-Smith, 1900) iN M T Mygona thammi (Staudinger, 1876) iN M T Mygona poeania (Hewitson, 1870) TL: Ecuador Mygona poeania poeania (Hewitson, 1870) TL: Ecuador M T Mygona poeania magalyae Pyrcz, 2004 TL: Peru Proboscis Thieme, 1907 Proboscis propylea (Hewitson, 1857) TL: Colombia M T Proboscis pomarancia Pyrcz, 2004 TL: Peru M T S Apexacuta Pyrcz, 2004 Apexacuta orsedice (Hewitson, 1878) iN Apexacuta orsedice orsedice (Hewitson, 1878) TL: Ecuador M T Apexacuta orsedice colombiana (E. Krüger, 1924) TL: Colombia M T S Apexacuta orsedice violacea (Weymer, 1912) TL: Ecuador M T Apexacuta orsedice [n. ssp.] Pyrcz & Clavijo, MS Colombia Apexacuta improvisa Pyrcz, 2004 TL: Peru Apexacuta improvisa improvisa Pyrcz, 2004 TL: Peru M L Apexacuta improvisa [n. ssp.] Pyrcz & Boyer, MS Peru Apexacuta superior Pyrcz, 2004 TL: Peru Apexacuta superior superior Pyrcz, 2004 TL: Peru M T Apexacuta superior [n. ssp.] Pyrcz & Boyer, MS Peru Apexacuta astoreth (Thieme, 1907) iN Apexacuta astoreth astoreth (Thieme, 1907) TL: Bolivia M T Apexacuta astoreth [n. ssp.] Pyrcz & Boyer, MS Peru Apexacuta [n. sp.] Pyrcz & Boyer, MS Peru Lasiophila C. Felder & R. Felder, 1859 Lasiophila cirta C. Felder & R. Felder, 1859 iN Lasiophila cirta cirta C. Felder & R. Felder, 1859 TL: Peru M T Lasiophila cirta atropurpurea Pyrcz, 2004 TL: Peru Lasiophila cirta [n. ssp.#1] Pyrcz, MS TL: Peru Lasiophila cirta [n. ssp.#2] Pyrcz, MS TL: Peru Lasiophila lucuma Pyrcz, 2011 TL: not specified Lasiophila lucuma lucuma Pyrcz, 2011 TL: not specified Lasiophila lucuma [n. ssp.] Pyrcz, MS Peru Lasiophila piscina Thieme, 1903 iN Lasiophila piscina piscina Thieme, 1903 TL: [Peru] M T S Lasiophila piscina [n. ssp.] Pyrcz, MS Peru Lasiophila luna Pyrcz, 2006 iN M S Lasiophila hewitsonia A. Butler, 1868 TL: Bolivia Lasiophila hewitsonia hewitsonia A. Butler, 1868 TL: Bolivia M T Lasiophila hewitsonia neda Thieme, 1907 TL: Peru M T Lasiophila zapatoza (Westwood, 1851) iN Lasiophila zapatoza zapatoza (Westwood, 1851) TL: "Bolivia"; Venezuela M T Lasiophila zapatoza meridae Adams & Bernard, 1981 TL: Venezuela M T Lasiophila zapatoza elusiva Pyrcz & Viloria, 2007 ref.285 Lasiophila zapatoza manaurera Adams & Bernard, 1979 TL: Colombia M T Lasiophila zapatoza sombra Thieme, 1907 TL: Colombia Lasiophila zapatoza [n. ssp.] Pyrcz, MS Venezuela Lasiophila semipartita Weymer, 1912 TL: Colombia Lasiophila prosymna (Hewitson, 1857) iN Lasiophila prosymna prosymna (Hewitson, 1857) TL: Colombia M T Lasiophila prosymna dirempta Thieme, 1907 TL: Ecuador M T Lasiophila parthyene (Hewitson, 1872) iN M T Lasiophila ciris Thieme, 1907 iN Lasiophila ciris ciris Thieme, 1907 TL: Ecuador M S Lasiophila ciris [n. ssp.] Pyrcz, MS Ecuador Lasiophila circe C. Felder & R. Felder, 1859 iN Lasiophila circe circe C. Felder & R. Felder, 1859 TL: Venezuela M T S Lasiophila circe farallonensis Pyrcz & Rodriguez, 2008 TL: not specified Lasiophila circe cnephas Thieme, 1907 TL: Colombia Lasiophila circe arithmetica Pyrcz & Viloria, 2007 ref.285 Lasiophila circe zarathustra Thieme, 1907 TL: Ecuador Lasiophila circe [n. ssp.#1] Pyrcz, MS TL: Colombia Lasiophila circe [n. ssp.#2] Pyrcz, MS TL: Colombia Lasiophila persepolis (Hewitson, 1872) iN Lasiophila persepolis persepolis (Hewitson, 1872) TL: Ecuador M T Lasiophila persepolis [n. ssp.] Pyrcz, MS Ecuador Lasiophila palades (Hewitson, 1872) iN Lasiophila palades palades (Hewitson, 1872) TL: Ecuador M T Lasiophila palades limes Pyrcz, 1999 TL: Ecuador Lasiophila phalaesia (Hewitson, 1868) iN Lasiophila phalaesia phalaesia (Hewitson, 1868) TL: Ecuador M T Lasiophila phalaesia alce Pyrcz, 1999 TL: Ecuador Lasiophila phalaesia confusa Staudinger, 1897 TL: [Ecuador] M T Lasiophila phalaesia kroli Pyrcz, 1999 TL: Ecuador Lasiophila regia Staudinger, 1897 TL: Bolivia M T Lasiophila orbifera A. Butler, 1868 iN Lasiophila orbifera orbifera A. Butler, 1868 TL: "Bolivia"; "Colombia"; Peru M T S Lasiophila orbifera munda Thieme, 1907 TL: Peru Lasiophila orbifera intercepta Thieme, 1907 TL: Ecuador Lasiophila orbifera pura Thieme, 1907 TL: Argentina Lasiophila alkaios Tessmann, 1928 TL: Peru Lasiophila alkaios alkaios Tessmann, 1928 TL: Peru M T Lasiophila alkaios gita Smart, 1975 TL: Peru Lasiophila alkaios [n. ssp.] Pyrcz, MS Peru Lasiophila [n. sp.] Pyrcz & Gareca, MS Bolivia Drucina A. Butler, 1872 Drucina championi Godman & Salvin, 1881 Blue-spotted Satyr iN Drucina championi championi Godman & Salvin, 1881 S Mexico (Chiapas) & Guatemala M T S IL L Drucina championi [n. ssp.] Pyrcz, MS Mexico Drucina leonata A. Butler, 1872 Leonata Satyr Costa Rica & Panama iN M T S L Druphila Pyrcz, 2004 Druphila venerata (A. Butler, 1873) iN Druphila venerata venerata (A. Butler, 1873) TL: Peru M T Druphila venerata [n. ssp.] Pyrcz, MS Peru Druphila [n. sp.] Pyrcz, MS Peru Foetterleia Viloria, 2004 Foetterleia schreineri (Foetterle, 1902) iN M T Pseudomaniola Röber, 1889 Pseudomaniola phaselis (Hewitson, 1862) iN Pseudomaniola phaselis phaselis (Hewitson, 1862) TL: Venezuela M T S Pseudomaniola phaselis pholoe (Staudinger, 1887) TL: Colombia M T S Pseudomaniola phaselis rogersi (Godman & Salvin, 1878) Costa Rica & W Panama M T S IL L Pseudomaniola phaselis macasica (Strand, 1912) TL: Ecuador M T Pseudomaniola phaselis argyritis (Thieme, 1907) TL: Bolivia M T Pseudomaniola gigas (Godman & Salvin, 1877) Orange-bordered Satyr S Mexico & Guatemala iN M T S Pseudomaniola loxo (Dognin, 1891) iN Pseudomaniola loxo loxo (Dognin, 1891) TL: Colombia M T Pseudomaniola loxo [n. ssp.] Pyrcz, MS Ecuador Pseudomaniola ilsa (Thieme, 1907) TL: Colombia M T Pseudomaniola mirabilis (A. Butler, 1873) iN Pseudomaniola mirabilis mirabilis (A. Butler, 1873) TL: Peru M T Pseudomaniola mirabilis extrema Pyrcz, 2004 TL: Peru Pseudomaniola gerlinda (Thieme, 1907) iN M T S Pseudomaniola mena (Grose-Smith, 1900) TL: Bolivia M T Pseudomaniola clethra (Thieme, 1907) TL: Peru M T Pseudomaniola helche (Thieme, 1907) TL: Bolivia M T Pseudomaniola asuba (Thieme, 1907) TL: Bolivia M S Pseudomaniola [n. sp.] Lamas, MS Peru Arhuaco Adams & Bernard, 1977 Arhuaco ica Adams & Bernard, 1977 TL: Colombia M T Arhuaco dryadina (Schaus, 1913) Dryadina Satyr Costa Rica & W Panama M T Cheimas Thieme, 1907 Cheimas opalinus (Staudinger, 1897) TL: Venezuela Cheimas opalinus opalinus (Staudinger, 1897) TL: Venezuela M T Cheimas opalinus cristalinus Pyrcz & Boyer, 2018 ref.283 Cheimas opalinus dominici Pyrcz & Boyer, 2018 ref.283 Cheimas opalinus iosephi Pyrcz & Boyer, 2018 ref.283 Cheimas opalinus rosalinus Pyrcz, 2018 ref.283 Steroma Westwood, [1850] Steroma bega Westwood, [1850] iN Steroma bega bega Westwood, [1850] TL: Venezuela M T Steroma bega andensis C. Felder & R. Felder, 1867 TL: Colombia; "Ecuador" M T S L Steroma modesta Weymer, 1912 iN M T S Steroma polyxo (Godman & Salvin, 1880) iN M T Steroma superba A. Butler, 1868 iN Steroma superba superba A. Butler, 1868 TL: Bolivia M T S Steroma superba [n. ssp.] Pyrcz, MS Peru Steremnia Thieme, 1905 Steremnia lucillae Pyrcz, 2004 TL: not specified M S Steremnia monachella (Weymer, 1911) iN Steremnia monachella monachella (Weymer, 1911) TL: Peru M T S Steremnia monachella adamsiella Pyrcz & Boyer, 2011 TL: Colombia Steremnia monachella [n. ssp.#1] Pyrcz, MS TL: Peru Steremnia monachella [n. ssp.#2] Pyrcz, MS TL: Ecuador Steremnia pronophila (C. Felder & R. Felder, 1867) iN Steremnia pronophila pronophila (C. Felder & R. Felder, 1867) TL: Colombia; "Ecuador" M T Steremnia pronophila fluminea Pyrcz & Viloria, 2007 ref.285 Steremnia selva Adams, 1986 TL: Colombia Steremnia selva selva Adams, 1986 TL: Colombia M T Steremnia selva [n. ssp.] Pyrcz, MS Colombia Steremnia umbracina (A. Butler, 1873) iN Steremnia umbracina umbracina (A. Butler, 1873) TL: Peru M T Steremnia umbracina misella Thieme, 1905 TL: Peru Steremnia agraulis (Weymer, 1912) TL: Bolivia Steremnia agraulis agraulis (Weymer, 1912) TL: Bolivia M T Steremnia agraulis agraulina Pyrcz, MS TL: Peru Steremnia rugilas Thieme, 1905 TL: [Peru] M T Steremnia [n. sp.#1] Pyrcz, MS TL: Ecuador Steremnia [n. sp.#2] Pyrcz, MS TL: not specified Steremnia [n. sp.#2] [n. ssp.#1] Pyrcz, MS TL: Ecuador Steremnia [n. sp.#2] [n. ssp.#2] Pyrcz, MS TL: Ecuador Steremnia [n. sp.#3] Pyrcz, MS TL: Ecuador Eteona E. Doubleday, 1848 Eteona tisiphone (Boisduval, 1836) iN M T Eretris Thieme, 1905 Eretris oculata (C. Felder & R. Felder, 1867) iN M T Eretris suzannae DeVries, 1980 Suzanne's Satyr Costa Rica & W Panama iN M T S Eretris lecromi Pyrcz, 1999 iN M S Eretris depresissima Pyrcz, 1999 TL: Colombia Eretris depresissima depresissima Pyrcz, 1999 TL: Colombia Eretris depresissima niambii Pyrcz, 1999 TL: Colombia Eretris agata Pyrcz & Fratello, 2005 TL: not specified Eretris encycla (C. Felder & R. Felder, 1867) TL: Venezuela Eretris encycla encycla (C. Felder & R. Felder, 1867) TL: Venezuela M T Eretris encycla [n. ssp.#1] Pyrcz, MS TL: Venezuela Eretris encycla [n. ssp.#2] Pyrcz, MS TL: Colombia Eretris encycla [n. ssp.#3] Pyrcz & Willmott, MS TL: Ecuador Eretris neocycla Pyrcz & Viloria, 2009 TL: not specified Eretris calisto (C. Felder & R. Felder, 1867) iN Eretris calisto calisto (C. Felder & R. Felder, 1867) TL: Colombia M T Eretris calisto kogui Adams & Bernard, 1977 TL: Colombia M T Eretris calisto [n. ssp.] Pyrcz, MS Colombia Eretris cuaoensis Viloria & Attal, 2019 ref.192 Eretris ocellifera (C. Felder & R. Felder, 1867) iN Eretris ocellifera ocellifera (C. Felder & R. Felder, 1867) TL: Colombia M T Eretris ocellifera [n. ssp.#1] Pyrcz & Willmott, MS TL: Colombia Eretris ocellifera [n. ssp.#2] Pyrcz & Willmott, MS TL: Colombia Eretris hulda (A. Butler & H. Druce, 1872) Hulda Satyr Costa Rica & Panama iN M T S IL Eretris maria (Schaus, 1920) Maria’s Satyr S Mexico & Guatemala iN M T S Eretris centralis E. Krüger, 1924 TL: Colombia M T S Eretris porphyria (C. Felder & R. Felder, 1867) iN Eretris porphyria porphyria (C. Felder & R. Felder, 1867) TL: Venezuela M T Eretris porphyria decorata (C. Felder & R. Felder, 1867) TL: Colombia M T Eretris porphyria catargyrea (Staudinger, 1887) TL: Colombia M T Eretris porphyria perija Adams & Bernard, 1979 TL: Colombia M T S Eretris porphyria recta Pyrcz & Viloria, 2007 ref.285 Eretris porphyria pseudoperija Pyrcz, 1999 TL: Colombia Eretris porphyria transmaraniona Pyrcz, 2004 TL: Peru Eretris porphyria [n. ssp.#1] Pyrcz, MS TL: Venezuela Eretris porphyria [n. ssp.#2] Pyrcz, MS TL: Venezuela Eretris porphyria [n. ssp.#3] Pyrcz, MS TL: Colombia Eretris porphyria [n. ssp.#4] Pyrcz, MS TL: Colombia Eretris apuleja (C. Felder & R. Felder, 1867) iN Eretris apuleja apuleja (C. Felder & R. Felder, 1867) TL: Venezuela M T Eretris apuleja subrufescens (Grose-Smith & W. F. Kirby, 1895) TL: Costa Rica M T Eretris apuleja bogotana E. Krüger, 1924 TL: [Colombia] M T S Eretris apuleja ochrea Thieme, 1905 TL: Ecuador M T Eretris apuleja altamira Adams & Bernard, 1979 TL: Colombia M T Eretris apuleja remotissima Pyrcz & Viloria, 2007 ref.285 Eretris apuleja socorrensis Pyrcz & Rodriguez, 2008 TL: not specified Eretris apuleja [n. ssp.#1] Pyrcz & Willmott, MS TL: Ecuador Eretris apuleja [n. ssp.#2] Pyrcz & Willmott, MS TL: Ecuador Eretris apuleja [n. ssp.#3] Pyrcz & Willmott, MS TL: Ecuador Eretris apuleja [n. ssp.#4] Pyrcz, MS TL: Colombia Eretris apuleja [n. ssp.#5] Pyrcz, MS TL: Colombia Eretris microrufescens Pyrcz & Wojtusiak, 2010 TL: not specified Eretris suprarufescens Pyrcz & Wojtusiak, 2010 TL: not specified Eretris apuleina (Pyrcz, 2004) iN Eretris apuleina apuleina (Pyrcz, 2004) TL: Ecuador M S Eretris apuleina [n. ssp.] Pyrcz, MS Peru Eretris truncatina (Pyrcz, 2004) TL: Peru Eretris truncatina truncatina (Pyrcz, 2004) TL: Peru M T Eretris truncatina [n. ssp.#1] Pyrcz, MS TL: Peru Eretris truncatina [n. ssp.#2] Pyrcz, MS TL: Peru Eretris subpunctata (Grose-Smith & W. F. Kirby, 1895) iN Eretris subpunctata subpunctata (Grose-Smith & W. F. Kirby, 1895) TL: Bolivia M T S L Eretris subpunctata [n. ssp.#1] Pyrcz, MS TL: Peru Eretris subpunctata [n. ssp.#2] Pyrcz, MS TL: Peru Eretris mendoza (Pyrcz, 2004) iN Eretris mendoza mendoza (Pyrcz, 2004) TL: Peru M T Eretris mendoza [n. ssp.#1] Pyrcz, MS TL: Peru Eretris mendoza [n. ssp.#2] Pyrcz, MS TL: Peru Eretris julieta Pyrcz & Gareca, 2010 TL: not specified Eretris julieta julieta Pyrcz & Gareca, 2010 TL: not specified Eretris julieta [n. ssp.] Pyrcz, MS Bolivia Eretris [n. sp.#1] Pyrcz, MS TL: Bolivia Eretris [n. sp.#2] Pyrcz, MS TL: Ecuador Eretris [n. sp.#3] Pyrcz, MS TL: not specified Eretris [n. sp.#3] [n. ssp.#1] Pyrcz, MS TL: Peru Eretris [n. sp.#3] [n. ssp.#2] Pyrcz, MS TL: Bolivia Eretris [n. sp.#3] [n. ssp.#3] Pyrcz, MS TL: Peru Eretris [n. sp.#4] Pyrcz, MS TL: not specified Eretris [n. sp.#4] [n. ssp.#1] Pyrcz, MS TL: Peru Eretris [n. sp.#4] [n. ssp.#2] Pyrcz, MS TL: Peru Eretris [n. sp.#4] [n. ssp.#3] Pyrcz, MS TL: Peru Eretris [n. sp.#4] [n. ssp.#4] Pyrcz, MS TL: Peru Eretris [n. sp.#4] [n. ssp.#5] Pyrcz, MS TL: Bolivia Eretris [n. sp.#5] Pyrcz, MS TL: Bolivia Eretris [n. sp.#6] Pyrcz, MS TL: not specified Eretris [n. sp.#6] [n. ssp.#1] Pyrcz, MS TL: Peru Eretris [n. sp.#6] [n. ssp.#2] Pyrcz, MS TL: Peru Eretris [n. sp.#6] [n. ssp.#3] Pyrcz, MS TL: Peru Eretris [n. sp.#6] [n. ssp.#4] Pyrcz, MS TL: Bolivia Manerebia Staudinger, 1897 Manerebia leaena (Hewitson, 1861) iN Manerebia leaena leaena (Hewitson, 1861) TL: [Ecuador] M T Manerebia leaena lanassa (C. Felder & R. Felder, 1867) TL: Colombia M T Manerebia leaena gonzalezi Pyrcz & Viloria, 2006 ref.280 Manerebia inderena (Adams, 1986) iN Manerebia inderena inderena (Adams, 1986) TL: Colombia M T S Manerebia inderena antioquiana Pyrcz & Willmott, 2006 ref.280 Manerebia inderena fina Pyrcz & Willmott, 2006 ref.280 Manerebia inderena similis Pyrcz & Willmott, 2006 ref.280 Manerebia inderena clara Pyrcz & Willmott, 2006 ref.280 M L Manerebia inderena leaeniva Pyrcz & Willmott, 2006 ref.280 Manerebia inderena mirena Pyrcz & Willmott, 2006 ref.280 Manerebia inderena [n. ssp.] Pyrcz, MS Ecuador Manerebia prattorum Pyrcz & Willmott, 2006 ref.280 Manerebia prattorum prattorum Pyrcz & Willmott, 2006 TL: Peru M S Manerebia prattorum udimaense Pyrcz, MS TL: Peru Manerebia golondrina Pyrcz & Willmott, 2006 ref.280 Manerebia trimaculata (Hewitson, 1870) TL: Ecuador Manerebia trimaculata trimaculata (Hewitson, 1870) TL: Ecuador M T Manerebia trimaculata [n. ssp.] Pyrcz, MS Ecuador Manerebia undulata Pyrcz & J. Hall, 2006 TL: Ecuador Manerebia undulata undulata Pyrcz & J. Hall, 2006 ref.280 Manerebia undulata milaena Pyrcz & Willmott, 2006 ref.280 Manerebia interrupta (F. Brown, 1944) TL: Ecuador M T Manerebia rufanalis Pyrcz & J. Hall, 2006 TL: Ecuador Manerebia rufanalis rufanalis Pyrcz & J. Hall, 2006 ref.280 Manerebia rufanalis fernandina Pyrcz & Willmott, 2006 ref.280 Manerebia quinterae (Adams & Bernard, 1979) TL: Venezuela M T S Manerebia navarrae (Adams & Bernard, 1979) TL: Colombia M T S Manerebia satura (Weymer, 1911) TL: Peru; "Colombia" Manerebia satura satura (Weymer, 1911) TL: Peru; "Colombia" Manerebia satura keithi (Dyar, 1913) TL: Peru M T Manerebia satura pauperata Pyrcz & Willmott, 2006 ref.280 Manerebia satura lamasi Pyrcz & Willmott, 2006 ref.280 Manerebia germaniae Pyrcz & J. Hall, 2006 TL: Ecuador Manerebia germaniae germaniae Pyrcz & J. Hall, 2006 ref.280 Manerebia germaniae vitalei Pyrcz & Willmott, 2006 ref.280 Manerebia mammuthus Pyrcz & Willmott, 2006 ref.280 Manerebia franciscae (Adams & Bernard, 1981) TL: Venezuela Manerebia franciscae franciscae (Adams & Bernard, 1981) TL: Venezuela M T Manerebia franciscae rodriguezi Pyrcz & Willmott, 2006 ref.280 Manerebia nevadensis E. Krüger, 1925 iN M T S Manerebia mycalesoides (C. Felder & R. Felder, 1867) TL: Colombia M T Manerebia magnifica Pyrcz & Willmott, 2006 ref.280 M T Manerebia levana (Godman, 1905) TL: Colombia Manerebia levana levana (Godman, 1905) TL: Colombia M T Manerebia levana pervaga Pyrcz & Viloria, 2007 TL: Venezuela Manerebia apiculata (C. Felder & R. Felder, 1867) TL: Colombia M T Manerebia ignilineata (Dognin, 1896) TL: Ecuador Manerebia ignilineata ignilineata (Dognin, 1896) TL: Ecuador M T Manerebia ignilineata neglecta (F. Brown, 1944) TL: Ecuador M T Manerebia ignilineata jalca Pyrcz, 2004 TL: Peru Manerebia seducta Pyrcz & Willmott, 2006 ref.280 M T Manerebia diffusa Pyrcz, 2004 TL: Peru M T Manerebia zoippus (H. Druce, 1876) TL: Peru M T Manerebia benigni Pyrcz, 2004 iN Manerebia benigni benigni Pyrcz, 2004 TL: Peru M T Manerebia benigni tessmanni Pyrcz, 2004 TL: Peru M T Manerebia trirufa Pyrcz, 2004 TL: Peru Manerebia trirufa trirufa Pyrcz, 2004 TL: Peru M T Manerebia trirufa [n. ssp.] Pyrcz, MS Peru Manerebia haywardi Pyrcz, 2004 TL: Peru Manerebia haywardi haywardi Pyrcz, 2004 TL: Peru M S Manerebia haywardi [n. ssp.] Pyrcz, MS Peru Manerebia rubescens (A. Butler, 1873) iN M T Manerebia cyclopella Staudinger, 1897 iN M T S Manerebia cyclops Staudinger, 1897 TL: Bolivia M T Manerebia cyclopina Staudinger, 1897 iN Manerebia cyclopina cyclopina Staudinger, 1897 TL: Peru M T Manerebia cyclopina staudingeri Forster, 1964 TL: Bolivia M T S Manerebia monops Hayward, 1949 TL: Argentina M T Manerebia insulsa (Hewitson, 1874) TL: Bolivia M T Manerebia typhlops Staudinger, 1897 TL: Bolivia M T Manerebia pluviosa Pyrcz & Viloria, 2007 ref.280 Manerebia reducta (F. Brown, 1944) TL: Peru M T Manerebia [n. sp.#1] Pyrcz, MS TL: not specified Manerebia [n. sp.#1] [n. ssp.#1] Pyrcz, MS TL: Peru Manerebia [n. sp.#1] [n. ssp.#2] Pyrcz, MS TL: Peru Manerebia [n. sp.#2] Pyrcz, MS TL: Peru Manerebia [n. sp.#3] Pyrcz, MS TL: Peru Manerebia [n. sp.#4] Pyrcz, MS TL: not specified Manerebia [n. sp.#4] [n. ssp.#1] Pyrcz, MS TL: Peru Manerebia [n. sp.#4] [n. ssp.#2] Pyrcz, MS TL: Peru Manerebia [n. sp.#5] Pyrcz, MS TL: Peru Manerebia [n. sp.#6] Pyrcz, MS TL: not specified Manerebia [n. sp.#6] [n. ssp.#1] Pyrcz, MS TL: Peru Manerebia [n. sp.#6] [n. ssp.#2] Pyrcz, MS TL: Bolivia Manerebia [n. sp.#7] Pyrcz, MS TL: Argentina Diaphanos Adams & Bernard, 1981 Diaphanos huberi Adams & Bernard, 1981 TL: Venezuela Diaphanos huberi huberi Adams & Bernard, 1981 TL: Venezuela M T Diaphanos huberi [n. ssp.] Pyrcz & Viloria, MS Venezuela Diaphanos curvignathos Viloria, 1994 TL: Venezuela Diaphanos curvignathos curvignathos Viloria, 1994 TL: Venezuela Diaphanos curvignathos [n. ssp.] Pyrcz & Viloria, MS Venezuela Diaphanos fuscus Viloria, 1994 TL: Venezuela Diaphanos [n. sp.] Viloria, MS Venezuela Idioneurula Strand, 1932 Idioneurula erebioides (C. Felder & R. Felder, 1867) iN Idioneurula erebioides erebioides (C. Felder & R. Felder, 1867) TL: Colombia M T Idioneurula erebioides socorroi Villalobos-Moreno & Salazar, 2013 TL: not specified Idioneurula donegani (Huertas & Arias, 2008) iN Idioneurula eremita Pyrcz & Viloria, 2007 ref.285 Idioneurula jacquelinae (Pyrcz, 1995) TL: Venezuela M T Lymanopoda Westwood, 1851 Lymanopoda panacea (Hewitson, 1869) iN Lymanopoda panacea panacea (Hewitson, 1869) TL: Ecuador M T Lymanopoda panacea gortyna (Weymer, 1890) TL: Colombia M T Lymanopoda panacea ocellifera A. Butler, 1873 TL: Peru M T S Lymanopoda panacea venusia Hopffer, 1874 TL: Peru Lymanopoda affineola Weymer, 1911 iN M T Lymanopoda albomaculata Hewitson, 1861 iN Lymanopoda albomaculata albomaculata Hewitson, 1861 TL: [Bolivia] M T S Lymanopoda albomaculata [n. ssp.] Pyrcz, MS Peru Lymanopoda albocincta Hewitson, 1861 iN Lymanopoda albocincta albocincta Hewitson, 1861 TL: Colombia M T Lymanopoda albocincta issacha A. Butler, 1870 TL: Cuba M T Lymanopoda albocincta intermedia Pyrcz, 2004 TL: Peru Lymanopoda albocincta [n. ssp.#1] Pyrcz, MS TL: Ecuador Lymanopoda albocincta [n. ssp.#2] Pyrcz, MS TL: Peru Lymanopoda florenciaensis Salazar, Henao & Vargas, 2004 iN Lymanopoda apulia Hopffer, 1874 iN M S Lymanopoda caucana Weymer, 1911 iN Lymanopoda caucana caucana Weymer, 1911 TL: Colombia Lymanopoda caucana orientalis Viloria & Camacho, 1999 TL: Venezuela Lymanopoda caeruleata Godman & Salvin, 1880 iN M T Lymanopoda euopis Godman & Salvin, 1878 Costa Rican Satyr Costa Rica iN M T TI S L Lymanopoda cinna Godman & Salvin, 1889 Blue-stained Satyr S Mexico (Chiapas) & Guatemala M T S L Lymanopoda acraeida A. Butler, 1868 iN Lymanopoda acraeida acraeida A. Butler, 1868 TL: Bolivia M T Lymanopoda acraeida zigomala (Hewitson, 1877) TL: Ecuador M T Lymanopoda acraeida chavezi Pyrcz, 2004 TL: Peru Lymanopoda acraeida malia Godman, 1905 TL: Peru M T S Lymanopoda acraeida [n. ssp.] Pyrcz, MS Peru Lymanopoda venosa A. Butler, 1868 iN M T S Lymanopoda obsoleta (Westwood, 1851) iN M T Lymanopoda altis Weymer, 1890 iN M T Lymanopoda maletera Adams & Bernard, 1979 TL: Venezuela M T Lymanopoda dietzi Adams & Bernard, 1981 TL: Venezuela Lymanopoda dietzi dietzi Adams & Bernard, 1981 TL: Venezuela M T S Lymanopoda dietzi neildi Pyrcz, 2010 TL: not specified Lymanopoda dietzi vetula Viloria & Pyrcz, 2010 TL: not specified Lymanopoda dietzi oliva Pyrcz & Boyer, 2010 TL: not specified Lymanopoda dietzi rosanna Viloria & Pyrcz, 2010 TL: Peru Lymanopoda dietzi argentata Pyrcz, 2010 TL: not specified Lymanopoda dietzi [n. ssp.] Pyrcz, MS Venezuela Lymanopoda confusa F. Brown, 1943 iN M T Lymanopoda lecromi Pyrcz & Viloria, 2007 ref.285 Lymanopoda confusiana Pyrcz & Boyer, 2010 TL: not specified Lymanopoda magna Pyrcz, 2004 TL: Peru Lymanopoda caudalis Rosenberg & Talbot, 1914 TL: Peru Lymanopoda caudalis caudalis Rosenberg & Talbot, 1914 TL: Peru M T Lymanopoda caudalis raymondae Pyrcz & Boyer, 2010 TL: Venezuela Lymanopoda balintiana Pyrcz & Boyer, 2010 TL: not specified Lymanopoda inaudita Pyrcz, 2010 iN Lymanopoda necalba Pyrcz & Boyer, 2010 TL: not specified Lymanopoda marianna Staudinger, 1897 TL: Venezuela Lymanopoda marianna marianna Staudinger, 1897 TL: Venezuela M T Lymanopoda marianna veronicae Pyrcz, 2010 TL: not specified Lymanopoda marianna giselae Pyrcz, 2010 TL: not specified Lymanopoda marianna ioanae Pyrcz, 2010 TL: not specified Lymanopoda marianna jacquelinae Pyrcz, 2010 TL: not specified Lymanopoda marianna valentinae Pyrcz, 2010 TL: not specified Lymanopoda labda Hewitson, 1861 iN Lymanopoda labda labda Hewitson, 1861 TL: Colombia M T Lymanopoda labda moritzi Pyrcz & Viloria, 2007 TL: Venezuela Lymanopoda labda lebbaea C. Felder & R. Felder, 1867 TL: Colombia M T Lymanopoda labda bilinskii Pyrcz, 2011 TL: not specified Lymanopoda altaselva Adams & Bernard, 1979 TL: Venezuela M T Lymanopoda nadia Pyrcz, 1999 iN M T Lymanopoda ionius Westwood, 1851 iN Lymanopoda ionius ionius Westwood, 1851 TL: Colombia M T Lymanopoda ionius excisa Weymer, 1911 TL: Colombia Lymanopoda ionius browni Pyrcz, 1999 TL: Ecuador Lymanopoda ionius gargantua Pyrcz & Rodriguez, 2008 TL: not specified Lymanopoda ionius [n. ssp.#1] Pyrcz, MS TL: Colombia Lymanopoda ionius [n. ssp.#2] Pyrcz, MS TL: Colombia Lymanopoda lactea Hewitson, 1861 TL: Colombia M T S Lymanopoda schmidti Adams, 1986 TL: Colombia M T S Lymanopoda labineta Hewitson, 1870 iN Lymanopoda labineta labineta Hewitson, 1870 TL: Ecuador M T Lymanopoda labineta piniasi Pyrcz, 1999 TL: Ecuador Lymanopoda labineta decorata Seydel, 1924 TL: Colombia M T Lymanopoda labineta wojtusiaki Pyrcz, 2011 TL: Ecuador Lymanopoda pieridina Röber, 1927 iN Lymanopoda pieridina pieridina Röber, 1927 TL: Colombia M T Lymanopoda pieridina albicosta Pyrcz, 1999 TL: Colombia Lymanopoda paisa Pyrcz & Rodriguez, 2006 TL: not specified Lymanopoda nivea Staudinger, 1887 iN Lymanopoda nivea nivea Staudinger, 1887 TL: Ecuador M T Lymanopoda nivea sororcula Thieme, 1904 TL: Ecuador Lymanopoda nivea bingo Pyrcz, 1999 TL: Ecuador Lymanopoda nivea bonita Pyrcz, 1999 TL: Ecuador Lymanopoda samius Westwood, 1851 iN Lymanopoda samius samius Westwood, 1851 TL: Colombia M T Lymanopoda samius lineana Pyrcz & Viloria, 2007 ref.285 Lymanopoda ferruginosa A. Butler, 1868 iN Lymanopoda ferruginosa ferruginosa A. Butler, 1868 TL: Bolivia M T S Lymanopoda ferruginosa [n. ssp.] Pyrcz, MS Peru Lymanopoda prusia (Heimlich, 1973) iN M S Lymanopoda rana Weymer, 1911 iN Lymanopoda rana rana Weymer, 1911 TL: Peru M S Lymanopoda rana marrana Pyrcz & Boyer, 2011 TL: not specified Lymanopoda hyagnis Weymer, 1911 iN M T S Lymanopoda translucida Weymer, 1912 TL: Bolivia Lymanopoda translucida translucida Weymer, 1912 TL: Bolivia Lymanopoda translucida umbratilis Rosenberg & Talbot, 1914 TL: Peru M T Lymanopoda translucida leucotecta Rosenberg & Talbot, 1914 TL: Peru M T Lymanopoda hockingiana Pyrcz, 2011 TL: not specified Lymanopoda araneola Pyrcz, 2004 TL: Peru M S Lymanopoda galactea Staudinger, 1897 TL: Bolivia M T Lymanopoda shefteli Dyar, 1913 iN Lymanopoda shefteli shefteli Dyar, 1913 TL: Peru M T Lymanopoda shefteli siviae Fuchs, 1954 TL: Peru Lymanopoda shefteli broweri Pyrcz & Boyer, 2011 TL: not specified Lymanopoda mariposa Pyrcz & Boyer, 2011 iN Lymanopoda ingasayana Pyrcz, 2004 TL: Peru Lymanopoda ingasayana ingasayana Pyrcz, 2004 TL: Peru Lymanopoda ingasayana [n. ssp.] Pyrcz, MS Peru Lymanopoda dyari Pyrcz, 2004 TL: Peru Lymanopoda inde Pyrcz, 2004 TL: Peru Lymanopoda inde inde Pyrcz, 2004 TL: Peru M S Lymanopoda inde [n. ssp.] Pyrcz, MS Peru Lymanopoda eubagioides A. Butler, 1873 iN M T S Lymanopoda hazelana F. Brown, 1943 TL: Ecuador Lymanopoda hazelana hazelana F. Brown, 1943 TL: Ecuador M T Lymanopoda hazelana summa Pyrcz, 1999 TL: Ecuador Lymanopoda hazelana viridia Pyrcz, 2011 TL: Ecuador Lymanopoda melia Weymer, 1911 iN Lymanopoda melia melia Weymer, 1911 TL: Colombia M T Lymanopoda melia alba E. Krüger, 1924 TL: Colombia M T S Lymanopoda melia [n. ssp.#1] Pyrcz, MS TL: Colombia Lymanopoda melia [n. ssp.#2] Pyrcz, MS TL: Ecuador Lymanopoda pascua Pyrcz, 2005 TL: not specified Lymanopoda tolima Weymer, 1911 iN Lymanopoda huilana Weymer, 1890 iN Lymanopoda huilana huilana Weymer, 1890 TL: Colombia M T Lymanopoda huilana cajamarca Adams, 1986 TL: Colombia M T S Lymanopoda huilana dominicae Pyrcz, MS TL: not specified Lymanopoda huilana [n. ssp.#1] Pyrcz, MS TL: Ecuador Lymanopoda huilana [n. ssp.#2] Pyrcz, MS TL: Ecuador Lymanopoda huilana [n. ssp.#3] Pyrcz, MS TL: Ecuador Lymanopoda huilana [n. ssp.#4] Pyrcz, MS TL: Colombia Lymanopoda caracara Pyrcz, Willmott & J. Hall, 1999 TL: Ecuador Lymanopoda caracara caracara Pyrcz, Willmott & J. Hall, 1999 TL: Ecuador Lymanopoda caracara ichu Pyrcz, Willmott & J. Hall, 1999 TL: Ecuador M S Lymanopoda caracara [n. ssp.] Pyrcz, Willmott, MS Ecuador Lymanopoda viventieni (Apolinar, 1924) TL: Colombia M S Lymanopoda mirabilis (Staudinger, 1897) TL: Colombia M T Lymanopoda paramera Adams & Bernard, 1979 TL: Colombia M T Lymanopoda nevada E. Krüger, 1924 iN Lymanopoda nevada nevada E. Krüger, 1924 TL: Colombia M T S Lymanopoda nevada nevadensis (A. Schultze, 1931) TL: Colombia Lymanopoda melendeza Adams, 1986 TL: Colombia M T S Lymanopoda [n. sp.#1] Pyrcz, MS TL: Ecuador Lymanopoda [n. sp.#2] Pyrcz, MS TL: Ecuador Lymanopoda [n. sp.#3] Huertas, MS TL: Colombia Lymanopoda [n. sp.#4] Pyrcz & Clavijo, MS TL: Colombia Lymanopoda [n. sp.#5] Pyrcz, MS TL: Peru Ianussiusa Pyrcz & Viloria, 2004 Ianussiusa maso (Godman, 1905) iN Ianussiusa maso maso (Godman, 1905) TL: Colombia M T Ianussiusa maso santa Pyrcz & Viloria, 2007 ref.285 Ianussiusa maso polyommatus (Röber, 1927) TL: Colombia M T Ianussiusa maso [n. ssp.#1] Pyrcz, MS TL: Ecuador Ianussiusa maso [n. ssp.#2] Pyrcz, MS TL: Colombia Redonda Adams & Bernard, 1981 Redonda empetrus (Thieme, 1905) TL: Venezuela Redonda empetrus empetrus (Thieme, 1905) TL: Venezuela Redonda empetrus bolivari Adams & Bernard, 1981 TL: Venezuela M T Redonda empetrus [n. ssp.] Pyrcz & Viloria, MS Venezuela Redonda bordoni Viloria & Pyrcz, 2003 iN Redonda [n. sp.#1] Viloria & Pyrcz, MS TL: not specified Redonda [n. sp.#1] [n. ssp.#1] Pyrcz & Viloria, MS TL: Venezuela Redonda [n. sp.#1] [n. ssp.#2] Pyrcz & Viloria, MS TL: Venezuela Redonda [n. sp.#1] [n. ssp.#3] Pyrcz & Viloria, MS TL: Venezuela Redonda [n. sp.#2] Viloria & Pyrcz, MS TL: Venezuela Redonda [n. sp.#3] Pyrcz, MS TL: Venezuela Dangond Adams & Bernard, 1979 Dangond dangondi Adams & Bernard, 1979 iN M T Altopedaliodes Forster, 1964 Altopedaliodes cocytia (C. Felder & R. Felder, 1867) iN Altopedaliodes cocytia cocytia (C. Felder & R. Felder, 1867) TL: Colombia M T S Altopedaliodes cocytia tamaensis Viloria & Pyrcz, 2007 TL: Colombia Altopedaliodes nebris (Thieme, 1905) iN M T Altopedaliodes reissi (Weymer, 1890) TL: Colombia Altopedaliodes reissi reissi (Weymer, 1890) TL: Colombia M T Altopedaliodes reissi flavomaculata (E. Krüger, 1924) TL: Colombia M T S Altopedaliodes reissi salazari Le Crom, 1994 TL: Colombia M T Altopedaliodes reissi [n. ssp.] Pyrcz & Willmott, MS Ecuador Altopedaliodes kruegeri Pyrcz, 1999 TL: Colombia Altopedaliodes kruegeri kruegeri Pyrcz, 1999 TL: Colombia M T Altopedaliodes kruegeri belmira Pyrcz & Rodriguez, 2004 TL: not specified Altopedaliodes scydmaena (Hayward, 1968) TL: Ecuador M T S Altopedaliodes tena (Hewitson, 1869) iN Altopedaliodes tena tena (Hewitson, 1869) TL: Ecuador M T Altopedaliodes tena [n. ssp.] Pyrcz, MS Ecuador Altopedaliodes nucea Pyrcz & Viloria, 1999 TL: Ecuador Altopedaliodes pasicles (Hewitson, 1872) TL: Ecuador M T Altopedaliodes perita (Hewitson, 1868) TL: Ecuador Altopedaliodes perita perita (Hewitson, 1868) TL: Ecuador M T Altopedaliodes perita sorda Pyrcz, 2004 TL: Ecuador Altopedaliodes zsolti Pyrcz & Viloria, 1999 TL: Ecuador Altopedaliodes zsolti zsolti Pyrcz & Viloria, 1999 TL: Ecuador M T Altopedaliodes zsolti citra Pyrcz, 2004 TL: Ecuador Altopedaliodes kurti Pyrcz & Viloria, 1999 TL: Ecuador M T Altopedaliodes [n. sp.#1] Pyrcz, MS TL: Colombia Altopedaliodes [n. sp.#2] Pyrcz & Willmott, MS TL: Ecuador Punapedaliodes Forster, 1964 Punapedaliodes flavopunctata (Staudinger, 1894) iN Punapedaliodes flavopunctata flavopunctata (Staudinger, 1894) TL: Bolivia M T Punapedaliodes flavopunctata minima Pyrcz, 2004 TL: Peru Punapedaliodes flavopunctata [n. ssp.] Pyrcz, MS Ecuador Antopedaliodes Forster, 1964 Antopedaliodes albopunctata (Weymer, 1890) iN M T S Antopedaliodes antonia (Staudinger, 1897) iN Antopedaliodes antonia antonia (Staudinger, 1897) TL: Bolivia M T Antopedaliodes antonia anina (Staudinger, 1897) TL: Bolivia M T Antopedaliodes antonia quincedis (Thieme, 1905) TL: Peru M T Antopedaliodes antonia [n. ssp.] Lamas, Viloria & Pyrcz, MS Peru Corderopedaliodes Forster, 1964 Corderopedaliodes corderoi (Dognin, 1893) iN Corderopedaliodes corderoi corderoi (Dognin, 1893) TL: Ecuador M T S Corderopedaliodes corderoi exornata Pyrcz, 2004 TL: Peru Corderopedaliodes buda Pyrcz, 2013 TL: not specified Corderopedaliodes luzangelae Pyrcz & Warren-Gash, 2013 TL: not specified Corderopedaliodes pandates (Hewitson, 1874) iN Corderopedaliodes pandates pandates (Hewitson, 1874) TL: Bolivia M T S Corderopedaliodes pandates olivieri Pyrcz, 2013 TL: not specified Panyapedaliodes Forster, 1964 Panyapedaliodes drymaea (Hewitson, 1858) iN Panyapedaliodes drymaea drymaea (Hewitson, 1858) TL: [Ecuador] M T S Panyapedaliodes drymaea atropulla Pyrcz, 2005 TL: not specified Panyapedaliodes jephtha (Thieme, 1905) iN M T Panyapedaliodes mara (Thieme, 1905) iN Panyapedaliodes monticola (Tessmann, 1928) iN M T Panyapedaliodes muscosa (Thieme, 1905) iN Panyapedaliodes muscosa muscosa (Thieme, 1905) TL: Bolivia M T Panyapedaliodes muscosa sumaco Pyrcz & Rodriguez, 2005 TL: Ecuador Panyapedaliodes muscosa tambito Pyrcz & Rodriguez, 2005 TL: not specified Panyapedaliodes cocorna Pyrcz & Rodriguez, MS TL: Peru Panyapedaliodes panyasis (Hewitson, 1862) iN M T S Panyapedaliodes stellata Pyrcz, 2004 TL: Peru Panyapedaliodes phila (Hewitson, 1862) iN Panyapedaliodes phila phila (Hewitson, 1862) TL: Colombia M T Panyapedaliodes phila certa Pyrcz, 2004 TL: Peru Panyapedaliodes phila combeima (E. Krüger, 1924) TL: Colombia M T S Panyapedaliodes phila philaenis (Thieme, 1905) TL: Ecuador Panyapedaliodes phila pilla Lamas, 2003 TL: Peru Panyapedaliodes phila [n. ssp.] Pyrcz, MS Ecuador Panyapedaliodes puma (Thieme, 1905) iN Panyapedaliodes puma puma (Thieme, 1905) TL: Bolivia M T Panyapedaliodes puma ernsti Pyrcz, 2006 TL: not specified Panyapedaliodes rahab (Thieme, 1905) TL: Peru M T S Panyapedaliodes silpa (Thieme, 1905) iN Panyapedaliodes silpa silpa (Thieme, 1905) TL: Bolivia; Peru M T Panyapedaliodes silpa tomentosa (Weymer, 1912) TL: Colombia M S Panyapedaliodes traceyannae Pyrcz & Viloria, 1999 iN Panyapedaliodes [n. sp.#1] Pyrcz, MS TL: Peru Panyapedaliodes [n. sp.#2] Viloria & Pyrcz, MS TL: Peru Panyapedaliodes [n. sp.#3] Pyrcz, MS TL: Peru Panyapedaliodes [n. sp.#4] Lamas & Pyrcz, MS TL: Peru Panyapedaliodes [n. sp.#5] Pyrcz & Clavijo, MS TL: Colombia Paramo Adams & Bernard, 1977 Paramo oculata (E. Krüger, 1924) iN M T S Parapedaliodes Forster, 1964 Parapedaliodes parepa (Hewitson, 1862) iN Parapedaliodes parepa parepa (Hewitson, 1862) TL: Ecuador M T Parapedaliodes parepa milvia (Thieme, 1905) TL: Peru M T Parapedaliodes parepa [n. ssp.#1] Pyrcz & Viloria, MS TL: Peru Parapedaliodes parepa [n. ssp.#2] Pyrcz & Viloria, MS TL: Ecuador Pherepedaliodes Forster, 1964 Pherepedaliodes naevia (Thieme, 1905) iN Pherepedaliodes naevia naevia (Thieme, 1905) TL: Ecuador Pherepedaliodes naevia [n. ssp.] Pyrcz, MS Peru Pherepedaliodes nubilia Pyrcz & Viloria, 1999 TL: Ecuador Pherepedaliodes angela Pyrcz & Rodriguez, 2005 TL: not specified Pherepedaliodes pheretiades (Grose-Smith & W. F. Kirby, 1894) iN M T Pherepedaliodes [n. sp.#1] Pyrcz, MS TL: Colombia Pherepedaliodes [n. sp.#2] Pyrcz, MS TL: Bolivia Pherepedaliodes [n. sp.#3] Pyrcz, MS TL: Peru Physcopedaliodes Forster, 1964 Physcopedaliodes physcoa (Hewitson, 1862) iN Physcopedaliodes physcoa physcoa (Hewitson, 1862) TL: Bolivia M T Physcopedaliodes physcoa marulla (Thieme, 1905) TL: Peru M T Physcopedaliodes physcoa micromaculata Forster, 1964 TL: Bolivia M T Praepedaliodes Forster, 1964 Praepedaliodes amussis (Thieme, 1905) TL: Colombia Praepedaliodes granulata (A. Butler, 1868) TL: Colombia M T Praepedaliodes phanias (Hewitson, 1862) iN M T Praepedaliodes exul (Thieme, 1905) TL: Brazil (RJ) M T Praepedaliodes duartei Dias, Dolibaina & Pyrcz, 2018 ref.282 Praepedaliodes francinii Freitas & Pyrcz, 2018 ref.282 iN Praepedaliodes landryi Pyrcz & Freitas, 2018 ref.282 iN Praepedaliodes pawlaki Pyrcz & Boyer, 2018 ref.282 Praepedaliodes sequeirae Pyrcz, Dias & Dolbaina, 2018 ref.282 iN Praepedaliodes zaccae Dolibaina, Dias & Pyrcz, 2018 ref.282 Praepronophila Forster, 1964 Praepronophila emma (Staudinger, 1897) TL: Bolivia M T Praepronophila perperna (Hewitson, 1862) iN Praepronophila perperna Panamanian segregate , Costa Rica & Panama M S L Praepronophila perperna perperna (Hewitson, 1862) TL: [S. Amer.] M T S Praepronophila perperna smalli Pyrcz & Viloria, 2007 TL: Panama Praepronophila perperna rodriguezi Pyrcz & Viloria, 2007 TL: Peru Praepronophila petronius (Grose-Smith, 1900) Toothed Satyr iN Praepronophila petronius kerrianna (L. Miller, 1986) Costa Rica & Panama M T S L Praepronophila petronius petronius (Grose-Smith, 1900) Panama & Colombia M T S Ma L Steromapedaliodes Forster, 1964 Steromapedaliodes albonotata (Godman, 1905) iN Steromapedaliodes albonotata albonotata (Godman, 1905) TL: Venezuela M T Steromapedaliodes albonotata privigna (Thieme, 1905) TL: Venezuela Steromapedaliodes albonotata sanchezi Viloria & Pyrcz, 2001 TL: Venezuela Steromapedaliodes albonotata schuberti Viloria & Pyrcz, 2001 TL: Venezuela M T Steromapedaliodes albonotata [n. ssp.#1] Pyrcz, MS TL: Venezuela Steromapedaliodes albonotata [n. ssp.#2] Pyrcz, MS TL: Venezuela Steromapedaliodes albonotata [n. ssp.#3] Pyrcz, MS TL: Venezuela Steromapedaliodes albonotata [n. ssp.#4] Pyrcz, MS TL: Venezuela Steromapedaliodes albarregas (Adams & Bernard, 1981) TL: not specified Steromapedaliodes albarregas albarregas (Adams & Bernard, 1981) TL: not specified M T Steromapedaliodes albarregas [n. ssp.#1] Pyrcz, MS TL: Venezuela Steromapedaliodes albarregas [n. ssp.#2] Pyrcz, MS TL: Venezuela Steromapedaliodes [n. sp.] Pyrcz, MS Venezuela Neopedaliodes Viloria, L. Miller & J. Miller, 2004 Neopedaliodes chingazaensis (Torres & Le Crom, 1997) TL: Colombia Neopedaliodes zipa (Adams, 1986) TL: Colombia M T Neopedaliodes nora (Adams, 1986) iN Neopedaliodes nora nora (Adams, 1986) TL: Colombia M T Neopedaliodes nora stepheni Viloria, L. Miller & J. Miller, 2008 TL: Colombia Neopedaliodes margaretha (Adams, 1986) TL: Colombia Neopedaliodes margaretha margaretha (Adams, 1986) TL: Colombia M T Neopedaliodes margaretha allyni Viloria, L. Miller & J. Miller, 2008 TL: Colombia Neopedaliodes philotera (Hewitson, 1868) TL: ? M T Neopedaliodes lefebvreae Viloria, L. Miller & J. Miller, 2008 TL: Ecuador Neopedaliodes parrhoebia (Hewitson, 1872) iN Neopedaliodes parrhoebia parrhoebia (Hewitson, 1872) TL: Ecuador M T Neopedaliodes parrhoebia phintia (Hewitson, 1877) TL: Ecuador M T Neopedaliodes entella (Thieme, 1905) TL: Ecuador Neopedaliodes juba (Staudinger, 1887) iN Neopedaliodes juba juba (Staudinger, 1887) TL: Ecuador M T Neopedaliodes juba jubilosa Viloria, L. Miller & J. Miller, 2008 TL: Ecuador Neopedaliodes michaeli Viloria, L. Miller & J. Miller, 2008 TL: Ecuador Neopedaliodes michaeli michaeli Viloria, L. Miller & J. Miller, 2008 TL: Ecuador Neopedaliodes michaeli innupta Pyrcz, 2008 TL: Ecuador Neopedaliodes michaeli citrica Viloria, L. Miller & J. Miller, 2008 TL: Ecuador Neopedaliodes proculeja (Thieme, 1905) TL: Peru Neopedaliodes proculeja malaga Pyrcz, 2008 TL: not specified Neopedaliodes proculeja proculeja (Thieme, 1905) TL: Peru M T Neopedaliodes phoenicusa (Hewitson, 1868) TL: Ecuador Neopedaliodes phoenicusa phoenicusa (Hewitson, 1868) TL: Ecuador M T Neopedaliodes phoenicusa [n. ssp.] Pyrcz, MS Ecuador Neopedaliodes yeyo Pyrcz & Rodriguez, 2008 TL: not specified Neopedaliodes [n. sp.] Pyrcz & Prieto, MS Colombia Pedaliodes A. Butler, 1867 Pedaliodes acjanaco Pyrcz, Viloria & Lamas, 2010 TL: Peru M S Pedaliodes adamsi D'Abrera, 1988 TL: Ecuador M T Pedaliodes albicilia Pyrcz, 2004 TL: Peru Pedaliodes albutia Thieme, 1905 TL: Peru M T Pedaliodes [n. sp.#1] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#2] Lamas & Viloria, MS TL: Peru Pedaliodes alusana (Hewitson, 1869) iN M T Pedaliodes amafania Thieme, 1905 iN M T Pedaliodes anchiphilonis Hayward, 1968 TL: Ecuador M T Pedaliodes antigua Adams & Bernard, 1981 iN M T Pedaliodes antulla Thieme, 1905 iN M T Pedaliodes arnotti Adams, 1986 TL: Colombia M T Pedaliodes arturi Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes asconia Thieme, 1905 iN M T Pedaliodes auraria Thieme, 1905 iN M T Pedaliodes aureola Pyrcz, 2004 TL: Peru M S Pedaliodes auristriga Thieme, 1905 TL: Peru M S Pedaliodes baccara Thieme, 1905 iN Pedaliodes baccara baccara Thieme, 1905 TL: Colombia M T Pedaliodes baccara allopatra Viloria & Pyrcz, 2007 ref.285 Pedaliodes balnearia Pyrcz & Viloria, 1999 TL: Ecuador M S Pedaliodes bernardi Adams, 1986 TL: Colombia M T Pedaliodes [n. sp.#3] Viloria, MS TL: Colombia Pedaliodes boettgeri Pyrcz, 2004 iN Pedaliodes caeca Pyrcz & Viloria, 1999 TL: Colombia Pedaliodes canela Pyrcz & Viloria, 1999 TL: Colombia Pedaliodes [n. sp.#4] Lamas & Viloria, MS TL: Peru Pedaliodes cebolleta Adams & Bernard, 1977 TL: Colombia M T Pedaliodes cesarense Adams & Bernard, 1979 iN M T Pedaliodes chaconi Viloria, 1998 TL: Venezuela M T Pedaliodes chrysotaenia (Hopffer, 1874) TL: Peru M T Pedaliodes circumducta Thieme, 1905 Circumducta Satyr SE Mexico, Guatemala iN M T L Pedaliodes cledonia Thieme, 1905 iN Pedaliodes cledonia cledonia Thieme, 1905 TL: Bolivia M T Pedaliodes cledonia molesta Pyrcz, 2004 TL: Peru M S Pedaliodes coca Staudinger, 1894 iN M T Pedaliodes [n. sp.#5] Viloria, MS TL: Colombia Pedaliodes costipunctata Weymer, 1912 TL: Colombia Pedaliodes cremera Godman & Salvin, 1878 Cremera Satyr Costa Rica & Panama iN M T OI S L Pedaliodes croizatorum Viloria & Camacho, 1999 TL: Venezuela Pedaliodes daulis Thieme, 1905 TL: Peru M T Pedaliodes dejecta (H. Bates, 1865) Dejected Satyr iN Pedaliodes dejecta SW Mexican segregate , SW Mexico Pedaliodes dejecta dejecta (H. Bates, 1865) SE Mexico to Colombia M T S L Pedaliodes dejecta [n. ssp.] L. Miller & Llorente, MS Mexico Pedaliodes demathani Pyrcz, 2004 iN Pedaliodes demathani demathani Pyrcz, 2004 TL: Peru M S Pedaliodes demathani pisca Lamas & Viloria, 2008 TL: Peru M S Pedaliodes demathani kuni Pyrcz & Boyer, 2008 TL: Peru M S Pedaliodes demathani chaneli Pyrcz & Boyer, 2008 TL: Peru Pedaliodes demathani benedeki Pyrcz, 2008 TL: Peru Pedaliodes demathani alfa Pyrcz & Boyer, 2008 TL: Peru Pedaliodes demathani ockendeni Lamas & Viloria, 2008 TL: Peru Pedaliodes demathani kalinowskii Pyrcz & Boyer, 2008 TL: Peru Pedaliodes demathani horsti Pyrcz & Viloria, 2008 TL: Peru Pedaliodes demathani nussi Pyrcz, 2008 TL: Bolivia Pedaliodes demarmelsi Viloria, 1995 TL: Venezuela M T Pedaliodes dracula Pyrcz & Viloria, 1999 iN M S Pedaliodes empusa (C. Felder & R. Felder, 1867) TL: Colombia Pedaliodes empusa empusa (C. Felder & R. Felder, 1867) TL: Colombia M T S Pedaliodes empusa medusa Pyrcz & Viloria, 2007 ref.285 Pedaliodes ereiba (C. Felder & R. Felder, 1867) iN M T Pedaliodes erschoffi Pyrcz, 2004 TL: Peru M S Pedaliodes exanima (Erschoff, 1875) TL: Peru M S Pedaliodes fassli Weymer, 1912 TL: Colombia M T Pedaliodes ferratilis A. Butler, 1873 iN M T S Pedaliodes fuscata (C. Felder & R. Felder, 1867) iN M T Pedaliodes [n. sp.#6] Viloria, MS TL: Colombia Pedaliodes [n. sp.#7] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#8] Lamas & Viloria, MS TL: Peru Pedaliodes guicana Adams, 1986 TL: Colombia M T Pedaliodes gustavi Viloria, L. Miller & J. Miller, 2003 TL: Ecuador Pedaliodes hardyi Adams, 1986 iN M T Pedaliodes [n. sp.#9] Viloria & Pyrcz, MS TL: Venezuela Pedaliodes hebena Pyrcz & Viloria, 1999 iN Pedaliodes hewitsoni Staudinger, 1897 iN Pedaliodes hewitsoni hewitsoni Staudinger, 1897 TL: Bolivia M T Pedaliodes hewitsoni primera Weeks, 1901 TL: Bolivia M T S Pedaliodes [n. sp.#10] Lamas & Viloria, MS TL: Peru Pedaliodes hopfferi Staudinger, 1887 iN M T S Pedaliodes [n. sp.#11] Lamas & Viloria, MS TL: Peru Pedaliodes japhleta A. Butler, 1870 TL: [Venezuela] M T Pedaliodes jelskii Pyrcz, 2004 TL: Peru Pedaliodes [n. sp.#12] Viloria, MS TL: Peru Pedaliodes leucocheilus Godman & Salvin, 1880 iN M T Pedaliodes luperca Thieme, 1905 TL: Ecuador Pedaliodes [n. sp.#13] Lamas & Viloria, MS TL: Peru Pedaliodes manis (C. Felder & R. Felder, 1867) Manis Satyr Costa Rica to Venezuela iN M T S L Pedaliodes manneja Thieme, 1905 TL: Colombia; Venezuela M T Pedaliodes maruda Pyrcz, 2004 TL: Peru M S Pedaliodes [n. sp.#14] Viloria, MS TL: Ecuador Pedaliodes melaleuca Weymer, 1890 iN Pedaliodes melaleuca melaleuca Weymer, 1890 TL: Colombia; Ecuador M T Pedaliodes melaleuca [n. ssp.] Viloria, MS Ecuador Pedaliodes [n. sp.#15] Pyrcz & Viloria, MS TL: Ecuador Pedaliodes [n. sp.#16] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#17] Lamas & Viloria, MS TL: Peru Pedaliodes montagna Adams & Bernard, 1981 iN M T S Pedaliodes morenoi Dognin, 1887 iN Pedaliodes morenoi morenoi Dognin, 1887 TL: Ecuador M T Pedaliodes morenoi pilaloensis Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes napaea (H. Bates, 1865) Napaea Satyr S Mexico & Guatemala iN M T S L Pedaliodes [n. sp.#18] L. Miller & Llorente, MS TL: Mexico Pedaliodes negreti Pyrcz, 1999 TL: Colombia Pedaliodes niveonota A. Butler, 1873 TL: Peru M T Pedaliodes obscura Grose-Smith & W. F. Kirby, 1894 TL: Bolivia M T Pedaliodes obstructa Pyrcz & Viloria, 1999 TL: Colombia M T Pedaliodes occulta Pyrcz & Viloria, 1999 TL: Colombia Pedaliodes ochrotaenia (C. Felder & R. Felder, 1867) TL: Colombia M T Pedaliodes ornata Grose-Smith & W. F. Kirby, 1895 iN M T Pedaliodes pacifica E. Krüger, 1924 iN Pedaliodes pactyes (Hewitson, 1874) TL: Bolivia M T Pedaliodes paeonides (Hewitson, 1874) TL: Ecuador M T S Pedaliodes palaepolis (Hewitson, 1878) iN Pedaliodes palaepolis palaepolis (Hewitson, 1878) TL: Bolivia M T S Pedaliodes palaepolis [n. ssp.#1] Lamas & Viloria, MS TL: Peru Pedaliodes palaepolis [n. ssp.#2] Lamas & Viloria, MS TL: Peru Pedaliodes palaepolis [n. ssp.#3] Viloria, MS TL: Ecuador Pedaliodes pallantis (Hewitson, 1862) TL: Colombia M T Pedaliodes palpita Adams, 1986 iN Pedaliodes palpita palpita Adams, 1986 TL: Colombia M T Pedaliodes palpita bedonica Pyrcz, 1999 TL: Colombia Pedaliodes pammenes (Hewitson, 1874) TL: Bolivia M T Pedaliodes paneis (Hewitson, 1862) TL: [Peru] M T Pedaliodes panthides (Hewitson, 1874) TL: Bolivia M T Pedaliodes parakana Viloria & Costa, 2019 ref.192 Pedaliodes parma Thieme, 1905 iN M T Pedaliodes parranda Adams, 1986 TL: Colombia M T Pedaliodes patizathes (Hewitson, 1874) TL: [Bolivia] M T Pedaliodes pausia (Hewitson, 1862) iN Pedaliodes pausia pausia (Hewitson, 1862) TL: Bolivia M T S Pedaliodes pausia uncus Thieme, 1905 TL: Argentina M T S Pedaliodes pedacia (Hewitson, 1868) TL: Ecuador M T Pedaliodes pelinaea (Hewitson, 1878) iN M T Pedaliodes pelinna (Hewitson, 1870) iN M T Pedaliodes perisades (Hewitson, 1874) TL: Bolivia M T Pedaliodes peruda (Hewitson, 1862) iN M T S Pedaliodes peruviana A. Butler, 1873 TL: Peru M T Pedaliodes petri Pyrcz & Viloria, 1999 iN Pedaliodes petri petri Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes petri maasseni Pyrcz, 2004 TL: Peru Pedaliodes peucestas (Hewitson, 1862) iN Pedaliodes peucestas peucestas (Hewitson, 1862) TL: Colombia M T Pedaliodes peucestas restricta Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes peucestas [n. ssp.] Viloria, MS Ecuador Pedaliodes phaea (Hewitson, 1862) iN M T Pedaliodes phaedra (Hewitson, 1870) iN Pedaliodes phaedra phaedra (Hewitson, 1870) TL: Ecuador M T Pedaliodes phaedra niphoessa Thieme, 1905 TL: Colombia M S Pedaliodes phaedra [n. ssp.#1] Viloria, MS TL: Ecuador Pedaliodes phaedra [n. ssp.#2] Viloria, MS TL: Ecuador Pedaliodes phaedra [n. ssp.#3] Viloria, MS TL: Ecuador Pedaliodes phaedra [n. ssp.#4] Viloria, MS TL: Ecuador Pedaliodes phaedra [n. ssp.#5] Viloria, MS TL: Ecuador Pedaliodes phaedra [n. ssp.#6] Viloria, MS TL: Ecuador Pedaliodes phaeina Staudinger, 1897 iN M T Pedaliodes [n. sp.#19] Lamas & Viloria, MS TL: Peru Pedaliodes phanoclea (Hewitson, 1877) TL: Ecuador M T Pedaliodes phazania Grose-Smith, 1900 iN Pedaliodes phazania phazania Grose-Smith, 1900 TL: Ecuador M T Pedaliodes phazania takahashii Adams & Bernard, 1977 TL: Colombia M T Pedaliodes pheres Thieme, 1905 iN M S Pedaliodes pheretias (Hewitson, 1872) TL: Ecuador M T Pedaliodes philonis (Hewitson, 1872) TL: Ecuador M T Pedaliodes phoenissa (Hewitson, 1862) iN M T S Pedaliodes phoenix Lamas, 1999 TL: Peru M T S Pedaliodes phrasa Grose-Smith & W. F. Kirby, 1894 TL: Bolivia M T Pedaliodes phrasicla (Hewitson, 1874) iN Pedaliodes phrasicla phrasicla (Hewitson, 1874) TL: Ecuador M T Pedaliodes phrasicla galaxias Weymer, 1912 TL: Peru; Bolivia M T S Pedaliodes phrasicla immaculata Pyrcz, 1999 TL: Colombia Pedaliodes phrasicla rufa Pyrcz, 2004 TL: Peru Pedaliodes phrasiclea Grose-Smith, 1900 iN M T Pedaliodes phrasis Grose-Smith, 1900 TL: Bolivia M T S Pedaliodes phthiotis (Hewitson, 1874) TL: Ecuador M T Pedaliodes piletha (Hewitson, 1862) TL: Venezuela M T Pedaliodes pimienta Adams, 1986 iN M T Pedaliodes piscolabis Pyrcz, 1999 TL: Colombia Pedaliodes pisonia (Hewitson, 1862) iN M T Pedaliodes plotina (Hewitson, 1862) iN Pedaliodes plotina plotina (Hewitson, 1862) TL: Venezuela M T Pedaliodes plotina lucipara Weymer, 1912 TL: Colombia Pedaliodes plotina pharnaspes (Hewitson, 1874) TL: Venezuela M T Pedaliodes plotina rapha A. Butler, 1870 TL: [Venezuela] M T S Pedaliodes plotina ruricola Pyrcz & Viloria, 2007 ref.285 Pedaliodes plotina [n. ssp.#1] Viloria, MS TL: Venezuela Pedaliodes plotina [n. ssp.#2] Pyrcz, MS TL: Venezuela Pedaliodes poema Pyrcz & Viloria, 1999 TL: Colombia Pedaliodes poesia (Hewitson, 1862) iN M T S Pedaliodes polla Thieme, 1905 iN M T S Pedaliodes pollonia Adams, 1986 iN M T S Pedaliodes polusca (Hewitson, 1862) iN M T S Pedaliodes [n. sp.#20] Lamas & Viloria, MS TL: Peru Pedaliodes pomponia (Hewitson, 1869) iN M T Pedaliodes porcia (Hewitson, 1869) iN Pedaliodes porcia porcia (Hewitson, 1869) TL: Ecuador M T Pedaliodes porcia pallantias (Hewitson, 1874) TL: Ecuador M T Pedaliodes porcia [n. ssp.#1] Viloria, MS TL: Colombia Pedaliodes porcia [n. ssp.#2] Viloria, MS TL: Colombia Pedaliodes porcia [n. ssp.#3] Viloria, MS TL: Colombia Pedaliodes porima Grose-Smith & W. F. Kirby, 1894 iN M T Pedaliodes porina (Hewitson, 1862) iN Pedaliodes porina porina (Hewitson, 1862) TL: ? M T S Pedaliodes porina [n. ssp.] Viloria & Pyrcz, MS Venezuela Pedaliodes praemontagna Pyrcz & Viloria, 2007 ref.285 Pedaliodes praxia (Hewitson, 1877) TL: Ecuador Pedaliodes praxia praxia (Hewitson, 1877) TL: Ecuador M T Pedaliodes praxia [n. ssp.] Viloria, MS Ecuador Pedaliodes praxithea (Hewitson, 1870) iN M T S Pedaliodes proerna (Hewitson, 1862) TL: Colombia Pedaliodes proerna proerna (Hewitson, 1862) TL: Colombia M T Pedaliodes proerna fumaria Thieme, 1905 TL: Venezuela M T Pedaliodes prosa Staudinger, 1894 TL: Bolivia M T Pedaliodes prytanis (Hewitson, 1862) iN M T Pedaliodes puracana E. Krüger, 1924 TL: Colombia M T Pedaliodes pylas (Hewitson, 1862) iN M T Pedaliodes [n. sp.#21] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#22] Lamas & Viloria, MS TL: Peru Pedaliodes ralphi Adams, 1986 TL: Colombia M T Pedaliodes reyi Pyrcz & Viloria, 2007 ref.285 iN Pedaliodes roraimae Strand, 1912 iN M T Pedaliodes rudnyi Pyrcz, 1999 TL: Colombia Pedaliodes rumba Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes [n. sp.#23] Lamas & Viloria, MS TL: Peru Pedaliodes simmias Thieme, 1905 TL: Bolivia M T S Pedaliodes simpla Thieme, 1905 iN Pedaliodes simplissima Pyrcz, 2004 iN M S Pedaliodes socorrae Adams, 1986 TL: Colombia M T Pedaliodes sonata Pyrcz & Viloria, 1999 iN Pedaliodes sophismata Pyrcz, 2004 TL: Peru M S Pedaliodes spina Weymer, 1912 iN Pedaliodes [n. sp.#24] Lamas & Viloria, MS TL: Peru Pedaliodes stuebeli Pyrcz, 2004 TL: Peru M S Pedaliodes suspiro Adams & Bernard, 1979 iN M T Pedaliodes syleus Thieme, 1905 TL: Peru Pedaliodes symmachus Godman & Salvin, 1880 iN M T Pedaliodes sztolcmani Pyrcz, 2004 TL: Peru Pedaliodes sztolcmani sztolcmani Pyrcz, 2004 TL: Peru Pedaliodes sztolcmani gilvaecosta Pyrcz, 2004 TL: Peru Pedaliodes tabaconas Pyrcz & Viloria, 1999 TL: Ecuador Pedaliodes terramaris Viloria & Pyrcz, 2000 TL: Venezuela M T Pedaliodes thiemei Staudinger, 1897 iN M T Pedaliodes transmontana Pyrcz & Viloria, 1999 iN Pedaliodes triaria Godman & Salvin, 1878 Ochre-banded Satyr Costa Rica & Panama iN M T OI S Pedaliodes tucca Thieme, 1905 iN M S Pedaliodes tyro Thieme, 1905 TL: Peru M T Pedaliodes tyrrheoides Adams & Bernard, 1979 iN M T Pedaliodes tyrrheus Godman & Salvin, 1880 iN Pedaliodes tyrrheus tyrrheus Godman & Salvin, 1880 TL: Colombia M T Pedaliodes tyrrheus tairona Adams & Bernard, 1977 TL: Colombia M T Pedaliodes uaniuna Pyrcz & Viloria, 2004 TL: Peru M S Pedaliodes uniformis Weymer, 1912 TL: Bolivia M T Pedaliodes uniplaga Weymer, 1912 TL: not specified M T S Pedaliodes valencia Pyrcz & Viloria, 2007 ref.285 Pedaliodes vallenata Adams & Bernard, 1979 TL: Venezuela M T S Pedaliodes wilhelmi Pyrcz, 1999 TL: Colombia Pedaliodes woytkowskii Pyrcz, 2004 TL: Peru M S Pedaliodes xanthosphenisca Hayward, 1968 TL: Ecuador M T Pedaliodes yutajeana Viloria & Pyrcz, 2000 TL: Venezuela M T Pedaliodes zuleta Adams & Bernard, 1979 iN M T S Pedaliodes zingara Heredia & Viloria , 2004 ref.200 iN Pedaliodes haydoni Pyrcz & Viloria, 2013 TL: Colombia Pedaliodes adrianae Pyrcz & Prieto, 2013 TL: Colombia Pedaliodes rodriguezi Pyrcz & Andrade, 2013 iN Pedaliodes philinae Pyrcz & Prieto, 2013 TL: Colombia Pedaliodes [n. sp.#25] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#26] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#27] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#28] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#29] Viloria, MS TL: Colombia Pedaliodes [n. sp.#30] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#31] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#32] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#33] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#34] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#35] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#36] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#37] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#38] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#39] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#40] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#41] Viloria, MS TL: Bolivia Pedaliodes [n. sp.#42] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#43] Viloria, MS TL: Brazil (SC) Pedaliodes [n. sp.#44] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#45] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#46] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#47] Viloria, MS TL: Bolivia Pedaliodes [n. sp.#48] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#49] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#50] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#51] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#52] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#53] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#54] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#55] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#56] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#57] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#58] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#59] Viloria, MS TL: Colombia Pedaliodes [n. sp.#60] Viloria, MS TL: ? Pedaliodes [n. sp.#61] Viloria, MS TL: Bolivia Pedaliodes [n. sp.#62] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#63] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#64] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#65] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#66] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#67] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#68] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#69] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#70] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#71] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#72] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#73] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#74] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#75] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#76] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#77] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#78] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#79] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#80] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#81] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#82] Viloria, MS TL: Colombia Pedaliodes [n. sp.#83] Lamas & Viloria, MS TL: Peru Pedaliodes [n. sp.#84] Pyrcz, MS TL: Ecuador Pedaliodes [n. sp.#85] Viloria, MS TL: Bolivia Pedaliodes [n. sp.#86] Pyrcz, MS TL: Ecuador Pedaliodes [n. sp.#87] Viloria, MS TL: Ecuador Pedaliodes [n. sp.#88] Viloria, MS TL: Bolivia Pedaliodes [n. sp.#89] L. Miller & Llorente, MS TL: Mexico (Gue) Pedaliodes [n. sp.#90] Lamas, MS TL: Peru Pedaliodes [n. sp.#91] Lamas, MS TL: Peru Pedaliodes [n. sp.#92] Lamas, MS TL: Peru Protopedaliodes Viloria & Pyrcz, 1994 Protopedaliodes kukenani Viloria & Pyrcz, 1994 TL: Venezuela M T Protopedaliodes ridouti Viloria & Pyrcz, 2000 TL: Venezuela M T Protopedaliodes profauna Viloria & Pyrcz, 2000 TL: Venezuela M T Protopedaliodes arekuna Pyrcz & Stachowicz, 2021 TL: Venezuela Argyrophorus Blanchard, 1852 Argyrophorus argenteus Blanchard, 1852 iN M T Argyrophorus blanchardi Pyrcz & Wojtusiak, MS TL: Peru Argyrophorus blanchardi blanchardi Pyrcz & Wojtusiak, MS TL: Peru Argyrophorus blanchardi celendeni Pyrcz & Wojtusiak, MS TL: Peru Argyrophorus blanchardi [n. ssp.] Lamas, MS Peru Argyrophorus chiliensis (Guérin-Méneville, [1830]) iN Argyrophorus chiliensis chiliensis (Guérin-Méneville, [1830]) TL: Chile M T Argyrophorus chiliensis magallanicus (Herrera, 1965) TL: Chile M T Argyrophorus chiliensis elwesi (Bryk, 1944) TL: Argentina Argyrophorus chiliensis [n. ssp.] Pyrcz, MS Argentina Argyrophorus tandilensis (Köhler, 1935) iN M T Argyrophorus antarcticus (Mabille, 1885) TL: [Argentina] M T Argyrophorus gustavi (Staudinger, 1898) iN Argyrophorus gustavi gustavi (Staudinger, 1898) TL: Bolivia M T Argyrophorus gustavi penai (Hayward, 1967) TL: Chile M T Argyrophorus monticolens (A. Butler, 1881) iN Argyrophorus monticolens monticolens (A. Butler, 1881) TL: Chile M T Argyrophorus monticolens [n. ssp.] Pyrcz, MS Argentina Argyrophorus poaoeneis (Heimlich, 1959) TL: Chile M T Argyrophorus angusta Weymer, 1911 TL: Bolivia Argyrophorus angusta angusta Weymer, 1911 TL: Bolivia M S Argyrophorus angusta picota Fuchs, 1954 TL: Peru Argyrophorus lamna Thieme, 1904 iN Argyrophorus lamna lamna Thieme, 1904 TL: Bolivia M S Argyrophorus lamna [n. ssp.#1] Pyrcz, MS TL: Peru M T Argyrophorus lamna [n. ssp.#2] Pyrcz, MS TL: Peru Argyrophorus lamna [n. ssp.#3] Pyrcz & Cerdeńa, MS TL: Peru Argyrophorus williamsianus A. Butler, 1868 TL: [Chile] M T Auca Hayward, 1953 Auca barrosi (C. Silva, 1917) iN Auca barrosi barrosi (C. Silva, 1917) TL: Chile M T Auca barrosi [n. ssp.] Pyrcz, MS Chile Auca coctei (Guérin-Méneville, [1838]) iN Auca coctei coctei (Guérin-Méneville, [1838]) TL: Chile M T S Auca coctei [n. ssp.] Pyrcz, MS Chile Auca pales (Philippi, 1859) TL: Chile Cosmosatyrus C. Felder & R. Felder, 1867 Cosmosatyrus leptoneuroides C. Felder & R. Felder, 1867 iN Cosmosatyrus leptoneuroides leptoneuroides C. Felder & R. Felder, 1867 TL: Chile M T Cosmosatyrus leptoneuroides plumbeola (A. Butler, 1868) TL: Chile M T Cosmosatyrus leptoneuroides [n. ssp.] Pyrcz, MS Argentina Elina Blanchard, 1852 Elina montrolii (Feisthamel, 1839) iN M T S Elina vanessoides Blanchard, 1852 iN M T S Faunula C. Felder & R. Felder, 1867 Faunula leucoglene C. Felder & R. Felder, 1867 iN M T Faunula eleates (Weymer, 1890) iN M T Faunula patagonica (Mabille, 1885) iN M T Faunula dubii Pyrcz, 2012 iN Faunula euripides (Weymer, 1890) TL: Bolivia Faunula euripides euripides (Weymer, 1890) TL: Bolivia M T Faunula euripides salomonis (Köhler, 1939) TL: Argentina M T Faunula euripides [n. ssp.] Pyrcz & Cerdeńa, MS Peru Haywardella Herrera, 1966 Haywardella edmondsii (A. Butler, 1881) iN M T Homoeonympha C. Felder & R. Felder, 1867 Homoeonympha boisduvalii (Blanchard, 1852) iN Homoeonympha boisduvalii boisduvalii (Blanchard, 1852) TL: Chile M T S Homoeonympha boisduvalii pusilla C. Felder & R. Felder, 1867 TL: Chile M T Homoeonympha vesagus (E. Doubleday, [1849]) TL: USA? M T Nelia Hayward, 1953 Nelia calvertii (Elwes, 1903) iN M T Nelia nemyroides (Blanchard, 1852) iN M T Nelia [n. sp.] Pyrcz, MS Chile Neomaenas Wallengren, 1858 Neomaenas coenonymphina A. Butler, 1881 iN M T Neomaenas edmondsii (A. Butler, 1881) TL: Chile M T Neomaenas fractifascia A. Butler, 1881 iN M T Neomaenas inornata (Elwes, 1903) iN M T Neomaenas poliozona (C. Felder & R. Felder, 1867) iN M T Neomaenas [n. sp.] Pyrcz, MS Chile Neomaenas servilia Wallengren, 1858 iN M T Neomaenas wallengrenii A. Butler, 1881 iN M T Neomaenas ambiorix Wallengren, 1858 iN Neomaenas ambiorix ambiorix Wallengren, 1858 TL: Chile M T Neomaenas ambiorix [n. ssp.#1] Pyrcz, MS TL: Chile Neomaenas ambiorix [n. ssp.#2] Pyrcz, MS TL: Chile Neomaenas simplex (A. Butler, 1881) TL: Chile M T Neomaenas humilis (C. Felder & R. Felder, 1867) iN Neomaenas humilis humilis (C. Felder & R. Felder, 1867) TL: Chile M T Neomaenas humilis [n. ssp.] Pyrcz, MS Chile Neomaenas schajovskoii (Hayward, 1954) iN M T Pampasatyrus Hayward, 1953 Pampasatyrus glaucope (C. Felder & R. Felder, 1867) iN Pampasatyrus glaucope glaucope (C. Felder & R. Felder, 1867) TL: Brazil M T Pampasatyrus glaucope boenninghauseni (Foetterle, 1902) TL: Brazil (RJ) M T Pampasatyrus gyrtone (Berg, 1877) iN M T Pampasatyrus quies (Berg, 1877) iN M T Pampasatyrus reticulata (Weymer, 1907) iN Pampasatyrus yacantoensis (Köhler, 1939) iN M T Pampasatyrus nilesi (Weeks, 1902) iN Pampasatyrus nilesi nilesi (Weeks, 1902) TL: Bolivia M T Pampasatyrus nilesi johanna (Weymer, 1911) TL: Argentina Pampasatyrus nilesi [n. ssp.] Pyrcz, MS Peru Pampasatyrus [n. sp.#1] Zacca & Pyrcz, MS TL: Brazil Pampasatyrus [n. sp.#2] Zacca & Pyrcz, MS TL: Argentina Quilaphoetosus Herrera, 1966 Quilaphoetosus janirioides (Blanchard, 1852) iN M T Quilaphoetosus monachus (Blanchard, 1852) iN Quilaphoetosus monachus monachus (Blanchard, 1852) TL: Chile M T Quilaphoetosus monachus valdiviae (C. Felder & R. Felder, 1867) TL: Chile M T Spinantenna Hayward, 1953 Spinantenna tristis (Guérin-Méneville, [1830]) iN M T Tetraphlebia C. Felder & R. Felder, 1867 Tetraphlebia germainii C. Felder & R. Felder, 1867 iN Tetraphlebia germainii germainii C. Felder & R. Felder, 1867 TL: Chile M T Tetraphlebia germainii argentina Hayward, 1962 TL: Argentina M T Chillanella Herrera, 1966 Chillanella stelligera (A. Butler, 1881) iN M T Tribe Satyrini, Subtribe Euptychiina Archeuptychia Forster, 1964 Archeuptychia cluena (Drury, 1782) iN M S Caeruleuptychia Forster, 1964 Caeruleuptychia aegrota (A. Butler, 1867) iN M T S Caeruleuptychia aetherialis (A. Butler, 1877) ref.258 iN M T Caeruleuptychia helios (Weymer, 1911) iN M T Caeruleuptychia trembathi Willmott, Nakahara, J. Hall & Neild, 2017 ref.258 Caeruleuptychia pilata (A. Butler, 1867) iN M T Caeruleuptychia scripta Nakahara, Zacca & Huertas, 2017 ref.260 Caeruleuptychia maryzenderae Lamas & Nakahara, 2017 ref.258 Caeruleuptychia coelestis (A. Butler, 1867) iN M T Caeruleuptychia coelica (Hewitson, 1869) iN M T Caeruleuptychia divina (Weymer, 1911) iN M T Caeruleuptychia cyanites (A. Butler, 1871) iN M T S Caeruleuptychia glauca (Weymer, 1911) iN M T Caeruleuptychia mare (A. Butler, 1869) TL: Brazil (Pa) M T Caeruleuptychia urania (A. Butler, 1867) iN M T S Caeruleuptychia ziza (A. Butler, 1869) TL: Peru M T Caeruleuptychia lobelia (A. Butler, 1870) iN M T S Caeruleuptychia probata (Weymer, 1911) iN M T Caeruleuptychia francisca (A. Butler, 1870) iN M T S Caeruleuptychia scopulata (Godman, 1905) iN M T S Caeruleuptychia saul Brévignon & M. Benmesbah, 2011 ref.179 Caeruleuptychia tenera (Weymer, 1911) iN M T S Caeruleuptychia twalela Brévignon, 2005 ref.174 iN Caeruleuptychia romani (Aurivillius, 1929) ref.201 iN M T Caeruleuptychia penicillata (Godman, 1905) TL: Brazil (Pa) M T S Caeruleuptychia helena (Anken, 1994) iN M OI OD G Caeruleuptychia umbrosa (A. Butler, 1870) iN M T S Caeruleuptychia [n. sp.#1] Lamas, MS TL: Peru Caeruleuptychia [n. sp.#2] Lamas, MS TL: Peru Caeruleuptychia [n. sp.#3] Lamas, MS TL: Peru Caeruleuptychia [n. sp.#4] Lamas, MS TL: Peru Caeruleuptychia [n. sp.#5] Lamas, MS TL: Peru Caeruleuptychia [n. sp.#6] Lamas, MS TL: Peru Capronnieria Forster, 1964 Capronnieria galesus (Godart, [1824]) iN M T S Capronnieria narapa (Schaus, 1902) ref.201 iN M T S Carminda Ebert & Dias, 1998 Carminda griseldis (Weymer, 1911) ref.195 iN M S Carminda paeon (Godart, [1824]) ref.195 iN M T Carminda surpresa Barbosa, Aguiar, Rosa, Zacca & A. Freitas, 2020 iN Carminda umuarama Ebert & Dias, 1997 ref.195 iN M T Cepheuptychia Forster, 1964 Cepheuptychia angelica (A. Butler, 1874) TL: Brazil (RJ) M T Cepheuptychia cephus (Fabricius, 1775) iN M T S Cepheuptychia glaucina (H. Bates, 1864) Dirty-blue Satyr SE Mexico, Guatemala iN M T S L Cepheuptychia [n. sp.] Lamas, MS Peru Chloreuptychia Forster, 1964 Chloreuptychia agatha (A. Butler, 1867) iN M T S Chloreuptychia amethysta Brévignon & M. Benmesbah, 2011 ref.179 iN Chloreuptychia chlorimene (Hübner, [1819]) iN M T S Chloreuptychia gordonsmalli Thurman, MacDonald & Nakahara, 2021 TL: Panama Chloreuptychia [n. sp.#1] Lamas, MS TL: Peru Chloreuptychia [n. sp.#2] Lamas, MS TL: Peru Lazulina Willmott, Nakahara & Espeland, 2023 Lazulina catharina (Staudinger, [1886]) ref.201 iN M T Lazulina hewitsonii (A. Butler, 1867) ref.201 iN M T Lazulina tolumnia (Cramer, 1777) ref.201 iN M S Lazulina [n. sp.] Lamas, MS Peru Amiga Nakahara, Willmott & Espeland, 2019 Amiga arnaca (Fabricius, 1776) Blue-smudged Satyr iN Amiga arnaca arnaca (Fabricius, 1776) Venezuela to Bolivia, Brazil, Guianas; ref.264 M T S L Amiga arnaca adela Nakahara & Espeland, 2019 ref.264 Amiga arnaca indianacristoi Nakahara & Marín, 2019 ref.201,309,272,264 Amiga sericeella (H. Bates, 1864) Blue-topped Satyr S Mexico-Honduras; ref.264 M T S Cissia E. Doubleday, 1848 Cissia penelope (Fabricius, 1775) iN M T S Cissia pompilia (C. Felder & R. Felder, 1867) Plain Satyr [= usitata], E & S Mexico to Colombia iN M T S L1 L2 Cissia proba (Weymer, 1911) iN M T Cissia cleophes (Godman & Salvin, 1889) Cleophes Satyr S Mexico; ref.11 iN M T S L Ha Cissia eous (A. Butler, 1867) ref.367 iN M T Cissia phronius (Godart, [1824]) ref.367 iN Cissia rubricata (W. H. Edwards, 1871) Red Satyr iN Cissia rubricata rubricata (W. H. Edwards, 1871) far NE Mexico; ref.367 M T S Cissia rubricata smithorum (Wind, 1946) N-central Mexico; ref.367 M T S L Cissia cheneyorum (R. Chermock, 1949) Red Satyr NW Mexico; ref.16 M S L I Cissia anabelae (L. Miller, 1976) Red Satyr Chiapas; ref.16 M T Cissia wahala Grishin, 2022 Mexico (Oax, Pue); ref.16 Cissia pseudocleophes (L. Miller, 1976) Red Satyr W Mexico; ref.16 M T S L Cissia joyceae Singer, DeVries & P. Ehrlich, 1983 Joyce's Satyr Costa Rica M T Vareuptychia Forster, 1964 Vareuptychia similis (A. Butler, 1867) One-pupil Satyr W Mexico to Colombia; ref.370 iN M T S L Vareuptychia themis (A. Butler, 1867) Two-pupil Satyr [= undina], W & SE Mexico to Nicaragua iN M T S1 S2 L Ha Vanima Zacca, Casagrande & O. Mielke, 2020 Vanima labe (A. Butler, 1870) Labe Satyr S Mexico to W Ecuador; ref.370 iN M T S L Vanima lesbia (Staudinger, [1886]) ref.370 M T Vanima palladia (A. Butler, 1867) Butler’s Satyr S Mexico to Brazil, Trinidad; ref.370 iN M T S L Cyllopsis R. Felder, 1869 Cyllopsis gemma (Hübner, 1808) Eastern Gemmed-Satyr iN Cyllopsis gemma gemma (Hübner, 1808) S IL to E TX, E to WV, S to FL M S L1 L2 L3 Cyllopsis gemma freemani (D. Stallings & J. Turner, 1947) S TX to Guatemala & Belize M T1 T2 S L Cyllopsis pertepida (Dyar, 1912) Canyonland Gemmed-Satyr iN Cyllopsis pertepida dorothea (Nabokov, 1943) SE NV, S & C UT, S & C CO, far W OK, N & C NM, N & C AZ M S L Cyllopsis pertepida maniola (Nabokov, 1942) SE AZ, SW NM, far NW Mexico (N SON, CHIH) M T S L Cyllopsis pertepida avicula (Nabokov, 1942) W TX, NE Mexico M T S I Ha Cyllopsis pertepida intermedia L. Miller, 1974 NW Mexico (W-central Chihuahua) M T S Cyllopsis pertepida pertepida (Dyar, 1912) W to S-central Mexico M T S L Cyllopsis pyracmon (A. Butler, 1867) Nabokov’s Satyr iN Cyllopsis pyracmon pyracmon (A. Butler, 1867) C & S Mexico to Guatemala M T S1 S2 L Cyllopsis pyracmon henshawi (W. H. Edwards, 1876) SE AZ, SW NM, NW Mexico M T S L I Cyllopsis brocki Grishin, 2023 ref.146 Cyllopsis argentella (A. Butler & H. Druce, 1872) Ochre-patched Gemmed-Satyr Costa Rica & W Panama iN M T S L Cyllopsis caballeroi Beutelspacher, 1982 Cowboy Gemmed-Satyr W Mexico (Nayarit, Jalisco) iN M T S Cyllopsis clinas (Godman & Salvin, 1889) Falcate Gemmed-Satyr SW Mexico (Guerrero) M T S Cyllopsis diazi L. Miller, 1974 Diaz's Gemmed-Satyr W Mexico, [Guatemala?] iN M T L Ha Cyllopsis dospassosi L. Miller, 1974 dos Passos' Gemmed-Satyr NE Mexico iN M T L Cyllopsis emilia I. Chacón & Nishida, 2002 White Gemmed-Satyr Costa Rica (Cerro de la Muerte, Villa Mills) M S Cyllopsis guatemalena L. Miller, 1974 Guatemalan Gemmed-Satyr S Mexico (Chiapas) to Guatemala iN M T S L Cyllopsis hedemanni R. Felder, 1869 Stub-tailed Gemmed-Satyr iN Cyllopsis hedemanni hedemanni R. Felder, 1869 [= vetones], E & S Mexico to W Panama M T S L Cyllopsis hedemanni tamaulipensis L. Miller, 1974 NE Mexico M T S L Cyllopsis hedemanni vetones (Godman & Salvin, 1878) TL: Costa Rica M T Cyllopsis hilaria (Godman, 1901) Two-toned Gemmed-Satyr E Mexico to W Panama iN M T S L Cyllopsis jacquelineae L. Miller, 1974 Jacqueline's Gemmed-Satyr SW Mexico (Guerrero, Oaxaca) iN M T L Cyllopsis nayarit (R. Chermock, 1947) Nayarit Gemmed-Satyr E & W Mexico iN M S L Cyllopsis nelsoni (Godman & Salvin, 1881) Yellow-striped Gemmed-Satyr Guatemala (Cerro de Zunil) iN M T TI L Cyllopsis pallens L. Miller, 1974 Pallid Gemmed-Satyr W Mexico to Guatemala iN M T S L Cyllopsis parvimaculata L. Miller, 1974 Weak-marked Gemmed-Satyr S-central Mexico (TL: Tenancingo, Mexico State) M T Cyllopsis pephredo (Godman, 1901) Big-spiked Gemmed-Satyr SE Mexico to C Panama iN M T S Cyllopsis perplexa L. Miller, 1974 Perplexing Gemmed-Satyr S & SW Mexico iN M T L Cyllopsis philodice (Godman & Salvin, 1878) Pale-banded Gemmed-Satyr Costa Rica & W Panama iN M T S L Cyllopsis pseudopephredo (R. Chermock, 1947) Chermock's Gemmed-Satyr S-central Mexico (Distrito Federal) M T Cyllopsis rogersi (Godman & Salvin, 1878) Costa Rican Gemmed-Satyr Costa Rica & Panama iN M T S L Cyllopsis schausi L. Miller, 1974 Schaus’ Gemmed-Satyr W Mexico to Guatemala M T S Cyllopsis steinhauserorum L. Miller, 1974 Steinhausers' Gemmed-Satyr W Mexico to El Salvador iN M T Cyllopsis suivalenoides L. Miller, 1974 Big-eyed Gemmed-Satyr W Mexico to Nicaragua iN M T S L Cyllopsis suivalens (Dyar, 1914) Dyar’s Gemmed-Satyr iN Cyllopsis suivalens suivalens (Dyar, 1914) E Mexico (Puebla, Veracruz) M T S L Cyllopsis suivalens escalantei L. Miller, 1974 S Mexico to Guatemala M T S L Cyllopsis tomemmeli Warren & Nakahara, 2018 ref.299 Cyllopsis whiteorum L. Miller & J. Maza, 1984 Dark Gemmed-Satyr S Mexico (N Oaxaca) iN M T L Cyllopsis wellingi L. Miller, 1978 Belize Gemmed-Satyr endemic to the Maya Mountains of Belize M T S Cyllopsis windi L. Miller, 1974 Wind's Gemmed-Satyr [Possibly a form of C. nayarit], E & W Mexico iN M T S L Erichthodes Forster, 1964 Erichthodes antonina (C. Felder & R. Felder, 1867) iN M T S Erichthodes jovita (C. Felder & R. Felder, 1867) iN M T Erichthodes julia (Weymer, 1911) iN M T Erichthodes eremita Lamas, Willmott & Radford, 2018 ref.315 Stephenympha Viloria, 2022 Stephenympha arius (Weymer, 1911) TL: Bolivia M T Stephenympha pauliana Viloria, 2022 TL: Venezuela Stephenympha eriphule (A. Butler, 1867) iN M T Atlanteuptychia Freitas, Barbosa & O. Mielke, 2013 Atlanteuptychia ernestina (Weymer, 1911) ref.212 iN M T Euptychia Hübner, 1818 Euptychia alacristata Neild, Nakahara & Fratello, 2014 Amazon basin & S of the Guianas; ref.267 Euptychia aquila Fratello, Nakahara & Brévignon, 2015 ref.206 Euptychia atlantica Nakahara & Freitas, 2017 ref.259 Euptychia attenboroughi Neild, Nakahara, Fratello & Le Crom, 2015 ref.268 Euptychia audacia Brévignon, Fratello & Nakahara, 2015 ref.206 Euptychia boulleti (Le Cerf, 1919) ref.210 M T Euptychia cesarense Pulido, Andrade, C. Peña & Lamas, 2011 TL: Colombia Euptychia cesarense cesarense Pulido, Andrade, C. Peña & Lamas, 2011 ref.279 Euptychia cesarense obtusa Nakahara, 2015 ref.252 Euptychia cesarense viloriai Andrade, Pulido, C. Peña & Lamas, 2011 ref.279 M S Euptychia efraini Ríos, 2019 TL: Colombia Euptychia enyita Nakahara, Lamas & Willmott, 2015 ref.252 iN Euptychia enyo A. Butler, 1867 iN M T Euptychia favonius Nakahara, G. Vega & Willmott, 2016 ref.257 Euptychia fernandae Nakahara & Willmott, 2015 ref.252 Euptychia fetna A. Butler, 1870 Orange-patched Satyr W & S Mexico to Guatemala iN M T S L Euptychia granatina Nakahara, LeCrom & J. Hall, 2015 ref.252 iN Euptychia hannemanni Forster, 1964 iN M T Euptychia insolata A. Butler & H. Druce, 1872 Insolata Satyr Costa Rica & Panama iN M T S L Euptychia jesia A. Butler, 1869 Jesia Satyr S Mexico to Colombia iN M T S L Euptychia juanjoi LeCrom, Nakahara & Lamas, 2015 ref.251 Euptychia lacandona A. Warren & Nakahara, 2015 ref.250 Euptychia marceli Brévignon, 2005 ref.174 iN Euptychia marceli marceli Brévignon, 2005 ref.174 Euptychia marceli divisa M. Benmesbah, Costa, Attal & Viloria, 2021 TL: Venezuela Euptychia meta Weymer, 1911 iN M T Euptychia mollina (Hübner, [1813]) Mollina Satyr E Mexico to Colombia & Venezuela iN Euptychia mollina mollina (Hübner, [1813]) E Mexico to Colombia & Venezuela M T S L Euptychia mollina suzannae Brévignon, 2005 ref.174 Euptychia mollis Staudinger, 1876 Mollis Satyr [Belize?] to Panama iN M T S Euptychia neblina A. Warren & Nakahara, 2015 ref.250 Euptychia neildi Brévignon, 2005 ref.174 Euptychia padroni Nakahara, Lamas & Willmott, 2015 ref.252 M S Euptychia pegasus Nakahara & J. Hall, 2015 ref.252 iN Euptychia picea A. Butler, 1867 iN M T S Euptychia pillaca Nakahara & Willmott, 2015 ref.252 Euptychia roraima Nakahara, Fratello & Harvey, 2014 Guyana; ref.249 Euptychia rubrofasciata L. Miller & J. Miller, 1988 Red-webbed Satyr NW Mexico iN M T S L Euptychia rufocincta Weymer, 1911 iN M T S Euptychia sarah M. Benmesbah & Viloria, 2019 TL: Venezuela Euptychia sophiae Zacca, Nakahara, Dolibaina & F. Dias, 2015 ref.268 Euptychia truncata Nakahara & J. Hall, 2015 ref.252 Euptychia westwoodi A. Butler, 1867 Westwood’s Satyr S Mexico to Colombia & Venezuela iN Euptychia westwoodi westwoodi A. Butler, 1867 S Mexico to Colombia & Venezuela M T S L Euptychia westwoodi muli Brévignon, 2015 ref.174 Euptychia woroina Viloria & Benmesbah, 2020 ref.193 Euptychia [n. sp.#1] Lamas, MS TL: Peru Euptychia [n. sp.#2] Lamas, MS TL: Peru Euptychia [n. sp.#3] Lamas, MS TL: Peru Euptychia [n. sp.#4] Lamas, MS TL: Peru Euptychia [n. sp.#5] Lamas, MS TL: Peru Euptychia [n. sp.#6] Lamas, MS TL: Peru Euptychia [n. sp.#7] Lamas, MS TL: Peru Euptychia [n. sp.#8] Lamas, MS TL: Peru Euptychia [n. sp.#9] Lamas, MS TL: Peru Euptychia [n. sp.#10] Lamas, MS TL: Peru Euptychia [n. sp.#11] Brévignon, MS TL: French Guiana Euptychia [n. sp.#12] Brévignon, MS TL: French Guiana Euptychia [n. sp.#13] Lamas, MS TL: Peru Euptychia [n. sp.#14] Neild, MS TL: Venezuela Euptychia [n. sp.#15] Neild, MS TL: Venezuela Euptychoides Forster, 1964 Euptychoides albofasciata (Hewitson, 1869) iN M T Euptychoides inani (Staudinger, [1886]) ref.201 iN M T Euptychoides laccine (C. Felder & R. Felder, 1867) iN M T S Euptychoides pseudosaturnus Forster, 1964 iN M T Cisandina Nakahara & Espeland, 2022 Cisandina castanya Lamas & Nakahara, 2022 TL: Peru Cisandina esmeralda Nakahara & E. Barbosa, 2022 TL: Brazil Cisandina fida (Weymer, 1911) ref.265 M T Cisandina lea (Cramer, 1777) iN Cisandina philippa (A. Butler, 1867) TL: Brazil (Am) M T S Cisandina sanmarcos (Nakahara & Lamas, 2018) ref.265 Cisandina trinitensis (Brévignon & M. Benmesbah, 2011) ref.179 Optimandes Marín, Nakahara & Willmott, 2019 Optimandes eugenia (C. Felder & R. Felder, 1867) iN Optimandes eugenia eugenia (C. Felder & R. Felder, 1867) ref.316 M T Optimandes eugenia transversa (Weymer, 1911) ref.316 M T Optimandes mocha Willmott, J. Hall & Lamas, 2019 ref.316 iN Graphita Nakahara, Marín & Barbosa, 2016 Graphita griphe (C. Felder & R. Felder, 1867) ref.254 iN M T L Godartiana Forster, 1964 Godartiana astronesthes Lamas & Nakahara, 2018 ref.261 iN Godartiana byses (Godart, [1824]) iN M T Godartiana amadoi M. Paluch, Zacca & Freitas, 2017 ref.364 Godartiana muscosa (A. Butler, 1870) iN M T Godartiana luederwaldti (Spitz, 1931) ref.365 iN M T Godartiana armilla (A. Butler, 1867) ref.365 iN M T Godartiana [n. sp.] Lamas, MS Peru Hermeuptychia Forster, 1964 Hermeuptychia sosybius (Fabricius, 1793) Carolina Satyr C TX & E OK, E to FL, N to S OH & VA iN M T OD TI S1 S2 S3 S4 IL Ma L1 L2 L3 L4 I G Txt Hermeuptychia hermybius Grishin, 2014 South Texas Satyr S TX & NE Mexico; ref.201,187 iN M T1 T2 T3 OD S IL L I G Hermeuptychia occidentalis Grishin, 2021 West Mexico Satyr SW Mexico (Jal, Mor, Gue) Hermeuptychia intricata Grishin, 2014 Intricate Satyr SE US coastal plains to (possibly) Costa Rica; ref.201,187 iN M T OD S IL Ma L Ha G Hermeuptychia clara Nakahara, D. Tan, Lamas & Willmott, 2016 ref.201,256 iN Hermeuptychia cucullina (Weymer, 1911) Cucullina Satyr iN M T OD Hermeuptychia gisella (Hayward, 1957) Gisella Satyr differences (genitalia, mitDNA) from H. cucullina support species status ... iN M T OD S G Hermeuptychia atalanta (A. Butler, 1867) Atalanta Satyr iN M T OI OD S G Hermeuptychia lupita (Reakirt, [1867]) Lupita’s Satyr SE Mexico iN Hermeuptychia hermes (Fabricius, 1775) Hermes Satyr probably in South America only iN M T OD S Hermeuptychia canthe (Hübner, [1811]) iN M T OD Hermeuptychia sinuosa Grishin, 2021 Wavy Stayr MX to Guatemala; ref.20 M T Hermeuptychia maimoune (A. Butler, 1870) Maimoune Satyr ref.201 iN M T OD S Hermeuptychia pimpla (C. Felder & R. Felder, 1862) Pimpla Satyr ref.201 iN M T OD S Hermeuptychia harmonia (A. Butler, 1867) Harmonia Satyr [= calixta], Costa Rica to W Ecuador iN M T OD S L Hermeuptychia fallax (C. Felder & R. Felder, 1862) Fallax Satyr iN Hermeuptychia fallax fallax (C. Felder & R. Felder, 1862) TL: [Peru] M T OD S Hermeuptychia fallax marinha Anken, 1994 TL: Brazil (RJ) M T OI OD G Hermeuptychia undulata (A. Butler, 1867) ref.371 iN M T OD S Hermeuptychia [n. sp.#1] Lamas, MS TL: Colombia Hermeuptychia [n. sp.#2] Lamas, MS TL: Peru Hermeuptychia [n. sp.#3] Lamas, MS TL: Peru Hermeuptychia [n. sp.#4] Lamas, MS TL: Peru Huberonympha Viloria & Costa, 2016 Huberonympha neildi Viloria, Costa, Fratello & Nakahara, 2016 TL: Venezuela Inbio Nakahara & Espeland, 2015 Inbio hilara (C. Felder & R. Felder, 1867) Tawny-cornered Satyr S Mexico to Colombia; ref.253 iN M T S L Magneuptychia Forster, 1964 Magneuptychia libye (Linnaeus, 1767) Blue-gray Satyr SE Mexico to Amazon Basin iN M T S L Magneuptychia lethra (Möschler, 1883) iN M T S Modica Zacca, Casagrande & Willmott, [2023] Modica fugitiva (Lamas, [1997]) iN M S L Modica kamel (Benmesbah & Zacca, 2018) ref.166 Modica maripa (Brévignon, 2005) TL: French Guiana Modica myncea (Cramer, 1780) ref.201 iN M T S Modica confusa (Staudinger, 1887) Confused Satyr SE Mexico to Brazil; ref.201 iN M T S L Deltaya Willmott, Nakahara & Espeland, [2023] Deltaya andrei (Zacca, Casagrande & O. Mielke, 2017) ref.201,366 iN Deltaya ocypete (Fabricius, 1776) ref.201 iN M T Deltaya pallema (Schaus, 1902) ref.201 iN M T S Deltaya louisammour (Benmesbah & Zacca, 2018) ref.201,166 Deltaya opima (Weymer, 1911) ref.201 iN M T Occulta Nakahara & Willmott, [2023] Occulta ocnus (A. Butler, 1867) ref.201 iN M T S Xenovena Marín & Nakahara, [2023] Xenovena murrayae (Brévignon, 2005) ref.174 iN Trico Nakahara & Espeland, [2023] Trico tricolor (Hewitson, 1850) iN Trico tricolor tricolor (Hewitson, 1850) TL: [Brazil (Pa)] M T S Trico tricolor fulgora (A. Butler, 1869) TL: Peru M T Scriptor Nakahara & Espeland, 2020 Scriptor sphenophorus Lamas & Nakahara, 2020 iN Modestia Viloria & M. Benmesbah, 2021 ref.167 Modestia agnata (Schaus, 1913) Agnata Satyr Costa Rica (Atlantic slope); ref.201 iN M T S Modestia analis (Godman, 1905) ref.201 iN M T S Modestia drymo (Schaus, 1913) Drymo Satyr Costa Rica (Atlantic slope); ref.201 M T S Modestia gomezi (Singer, DeVries & P. Ehrlich, 1983) Gomez' Satyr Costa Rica; ref.167 iN M T S L Modestia harpyia (C. Felder & R. Felder, 1867) ref.201 iN Modestia harpyia harpyia (C. Felder & R. Felder, 1867) TL: Surinam M T Modestia harpyia batesii (A. Butler, 1867) TL: Brazil (Pa) M T S Modestia modesta (A. Butler, 1867) ref.167 iN M T S Modestia remypignoux M. Benmesbah & Viloria, 2021 ref.167 Modestia remypignoux remypignoux M. Benmesbah & Viloria, 2021 ref.167 Modestia remypignoux shueyi M. Benmesbah & Viloria, 2021 ref.167 Modestia sylvina (C. Felder & R. Felder, 1867) ref.201 iN M T Colombeia Viloria, Andrade & Le Crom, 2019 Colombeia hotchkissi (Dyar, 1913) ref.201 M T Colombeia mycalesis (Röber, 1927) ref.156 iN M T Colombeia nossis (Hewitson, 1862) ref.201 iN M T Omacha Andrade, Viloria, Henao & Le Crom, 2019 Omacha pax (Huertas, Lamas, G. Fagua & Willmott, 2016) ref.156 iN Megeuptychia Forster, 1964 Megeuptychia antonoe (Cramer, 1775) Cramer's Satyr S Mexico to Amazon Basin iN M T S L Megeuptychia monopunctata Willmott & J. Hall, 1995 iN M T S Megeuptychia souadae (M. Benmesbah, 2015) ref.165 iN Megisto Hübner, [1819] Megisto cymela (Cramer, 1777) Little Wood-Satyr iN Megisto cymela cymela (Cramer, 1777) TL: South Africa M T OI S L1 L2 L3 L4 L5 Ha G Megisto cymela viola (Maynard, 1891) TL: USA (Fl) M TI S L Llorenteana Viloria & Luis-Martínez, [2019] Llorenteana pellonia (Godman, 1901) Plateau Satyr NW Mexico; ref.296 M T S Neonympha Hübner, 1818 Neonympha areolatus (J. E. Smith, 1797) Georgia Satyr iN Neonympha areolatus septentrionalis W. Davis, 1924 TL: USA (NJ) M S Neonympha areolatus areolatus (J. E. Smith, 1797) [= helicta]; ref.134 M T S L1 L2 Ha Neonympha areolatus dadeensis Gatrelle, 1999 TL: USA (Fl) M T Neonympha mitchellii French, 1889 Mitchell's Satyr iN Neonympha mitchellii mitchellii French, 1889 TL: USA (Mi) M S1 S2 L Neonympha mitchellii francisci Parshall & Kral, 1989 TL: USA (NC) M T S L Ha Moneuptychia Forster, 1964 Moneuptychia castrensis (Schaus, 1902) iN M T Moneuptychia giffordi Freitas, Emery & O. Mielke, 2010 ref.209 iN Moneuptychia itapeva Freitas, 2007 ref.208 iN Moneuptychia melchiades (A. Butler, 1877) TL: Argentina M T Moneuptychia montana Freitas, 2015 ref.214 Moneuptychia pervagata Freitas, R. Siewert & O. Mielke, 2015 ref.214 iN Moneuptychia romanina (Bryk, 1953) iN M T Moneuptychia soter (A. Butler, 1877) iN M T Moneuptychia vitellina Freitas & E. Barbosa, 2015 ref.214 iN Moneuptychia viviana (Romieux, 1927) ref.164 M T Moneuptychia wahlbergi Freitas, E. Barbosa, R. Siewert & O. Mielke, 2015 ref.214 iN Paramacera A. Butler, 1868 Paramacera allyni L. Miller, 1972 Arizona Pine-Satyr SE Arizona, far NW Mexico; species status supported by differences in and ...; ref.16 M T S IL L I G Paramacera xicaque (Reakirt, [1867]) Mexican Pine-Satyr E & central Mexico iN M T S IL L G Paramacera rubrosuffusa L. Miller, 1972 Mexican Pine-Satyr S Oaxaca (Sierra Madre del Sur); ref.16 M T S L Paramacera copiosa L. Miller, 1972 Guerrero Pine-Satyr N Oaxaca, Guerrero M T S IL G Paramacera chinanteca L. Miller, 1972 Oaxacan Pine-Satyr N Oaxaca iN M T S IL L G Pareuptychia Forster, 1964 Pareuptychia binocula (A. Butler, 1869) iN M T S Pareuptychia difficilis Forster, 1964 TL: Bolivia M T Pareuptychia hervei Brévignon, 2005 ref.174 Pareuptychia hesionides Forster, 1964 iN Pareuptychia hesionides hesionides Forster, 1964 TL: Bolivia M T S Pareuptychia hesionides deviae Brévignon, 2005 ref.174 Pareuptychia lydia (Cramer, 1777) iN M T Pareuptychia metaleuca (Boisduval, 1870) White-banded Satyr E & S Mexico to Panama iN Pareuptychia metaleuca metaleuca (Boisduval, 1870) TL: Guatemala M T S L1 L2 Pareuptychia metaleuca [n. ssp.#1] Lamas, MS TL: Colombia Pareuptychia metaleuca [n. ssp.#2] Lamas, MS TL: Peru Pareuptychia metaleuca [n. ssp.#3] Lamas, MS TL: Peru Pareuptychia metaleuca [n. ssp.#4] Lamas, MS TL: French Guiana Pareuptychia milleri Nakahara, M. Marín & Neild, 2016 ref.255 Pareuptychia ocirrhoe (Fabricius, 1776) iN Pareuptychia ocirrhoe Northern segregate , [= hesione], E & S Mexico to Ecuador M S L1 L2 Pareuptychia ocirrhoe ocirrhoe (Fabricius, 1776) TL: Surinam M T Pareuptychia ocirrhoe interjecta (R.F. d'Almeida, 1952) TL: Brazil (RJ) M T Pareuptychia ocirrhoe [n. ssp.#1] Lamas, MS TL: Mexico Pareuptychia ocirrhoe [n. ssp.#2] Lamas, MS TL: Peru Pareuptychia ocirrhoe [n. ssp.#3] Lamas, MS TL: Trinidad Pareuptychia summandosa (Gosse, 1880) iN M T Pareuptychia [n. sp.] Lamas, MS Peru Paryphthimoides Forster, 1964 Paryphthimoides poltys (Prittwitz, 1865) iN Paryphthimoides poltys poltys (Prittwitz, 1865) TL: Brazil (RJ) M T OD Paryphthimoides poltys numilia (C. Felder & R. Felder, 1867) TL: Colombia M T Paryphthimoides vestigiata (A. Butler, 1867) iN M T Paryphthimoides fridae Zacca, Casagrande, Huertas & Nakahara, 2020 ref.369 Paryphthimoides jorupe Zacca, Casagrande, Checa & Willmott, 2020 ref.369 Paryphthimoides terrestris (A. Butler, 1867) Cryptic Satyr iN Paryphthimoides terrestris terrestris (A. Butler, 1867) E Mexico to Brazil M T S L Paryphthimoides terrestris araguaianus Zacca, Casagrande & O. Mielke, 2020 ref.369 Paryphthimoides terrestris grevei Zacca, Casagrande & O. Mielke, 2020 ref.369 Paryphthimoides terrestris muyrakytan Zacca, Casagrande & O. Mielke, 2020 ref.369 Paryphthimoides touloulou (M. Benmesbah, 2015) ref.369 Paryphthimoides brixius (Godart, [1824]) iN Paryphthimoides brixius brixius (Godart, [1824]) ref.369 M S Paryphthimoides brixius madre Zacca, O. Mielke, Casagrande & Lamas, 2020 ref.369 Paryphthimoides brixius rouranensis Brévignon, 2020 ref.369 Paryphthimoides brixiola (A. Butler, 1867) TL: Brazil (Pa) M T Paryphthimoides flavofascia (Zacca & R. Siewert, 2014) ref.369 iN Paryphthimoides pseudoconfusa (Singer, DeVries & P. Ehrlich, 1983) Gold-stained Satyr E Mexico to Panama; ref.201 iN M T S L Paryphthimoides sheba (Brévignon & M. Benmesbah, 2011) ref.201,179 Emeryus Zacca, Casagrande & O. Mielke, 2020 Emeryus argulus (Godart, [1824]) iN Emeryus argulus argulus (Godart, [1824]) ref.368 M T Emeryus argulus magnum Zacca, Casagrande & O. Mielke, 2019 ref.368 Emeryus difficilis (Forster, 1964) ref.368 iN M T Emeryus numeria (C. Felder & R. Felder, 1867) ref.368 iN M T Pharneuptychia Forster, 1964 Pharneuptychia haywardiana Lamas, 2010 TL: Bolivia M T Pharneuptychia estoraquensis Henao & Meneses, [2018] TL: Colombia Pharneuptychia innocentia (C. Felder & R. Felder, 1867) iN M T Pharneuptychia phares (Godart, [1824]) iN M T Pharneuptychia pharnabazos (Bryk, 1953) iN M T Pharneuptychia pharnaces (Weymer, 1911) TL: Brazil Pindis R. Felder, 1869 Pindis squamistriga R. Felder, 1869 Variable Satyr E & W Mexico to Guatemala iN M T S L I Pseudeuptychia Forster, 1964 Pseudeuptychia hemileuca (Staudinger, [1886]) TL: Colombia M T Pseudeuptychia languida (A. Butler, 1871) iN Pseudeuptychia languida languida (A. Butler, 1871) TL: Colombia M T Pseudeuptychia languida austrina Nakahara & Lamas, 2018 ref.261 Pseudeuptychia cuzquenya Nakahara & Lamas, 2018 ref.261 Pseudeuptychia callichloris (A. Butler, 1867) ref.201 iN M T Pseudeuptychia herseis (Godart, [1824]) ref.201 iN M T Pseudeuptychia rectilinea (Brévignon, Rosant, Lamas & Willmott, 2019) ref.201,180 iN Pseudeuptychia marica (Weymer, 1911) ref.201 iN M T S Satyrotaygetis Forster, 1964 Satyrotaygetis satyrina (H. Bates, 1865) Wide-bordered Satyr [= gigas (Butler)], S Mexico to Panama; ref.201 iN M T S L1 L2 Satyrotaygetis iris (C. Felder & R. Felder, 1867) ref.201 iN M T S Satyrotaygetis tiessa (Hewitson, 1869) Tiessa Satyr Nicaragua to W Ecuador iN M T S L Splendeuptychia Forster, 1964 Splendeuptychia ambra (Weymer, [1911]) iN Splendeuptychia ashna (Hewitson, 1869) iN M T Splendeuptychia purusana (Aurivillius, 1929) iN M T Splendeuptychia tupinamba A. Freitas, Huertas & Rosa, 2021 ref.201 iN Splendeuptychia gera (Hewitson, 1850) ref.201 iN Splendeuptychia gera gera (Hewitson, 1850) TL: Brazil M T Splendeuptychia gera nortia (Hewitson, 1862) TL: Brazil M T S Splendeuptychia nobilis (Weymer, 1911) ref.201 Splendeuptychia moderata (Weymer, 1911) ref.201 iN M T Splendeuptychia metagera (A. Butler, 1867) ref.201 M T Splendeuptychia [n. sp.#1] Lamas, MS TL: Peru Splendeuptychia [n. sp.#2] Lamas, MS TL: Peru Splendeuptychia [n. sp.#3] Lamas, MS TL: Colombia Argentaria Huertas & Willmott, 2023 Argentaria argyropsacas (Bryk, 1953) ref.201 iN M T Argentaria clementia (A. Butler, 1877) ref.201 iN M T S Argentaria clorimena (Stoll, 1790) ref.201 iN M T Argentaria cosmophila (Hübner, 1823) ref.201 M T Argentaria hygina (A. Butler, 1877) ref.201 iN M T Argentaria itonis (Hewitson, 1862) ref.201 iN M T Argentaria jadea (Brévignon & M. Benmesbah, 2011) ref.201,179 iN Argentaria kendalli (L. Miller, 1976) Kendall's Satyr NE Mexico; ref.201 iN M T S L Argentaria libitina (A. Butler, 1870) ref.201 iN M T Argentaria pagyris (Godart, [1824]) ref.201 iN M T Argentaria quadrina (A. Butler, 1869) ref.201 iN M T Argentaria salvini (A. Butler, 1867) Salvin's Satyr Panama; ref.201 iN M T S Argentaria telesphora (A. Butler, 1877) ref.201 M T Argentaria zischkai (Forster, 1964) ref.201 M T Argentaria [n. sp.#1] Lamas, MS TL: Peru Argentaria [n. sp.#2] Lamas, MS TL: Peru Argentaria [n. sp.#3] Lamas, MS TL: Peru Argentaria [n. sp.#4] Lamas, MS TL: Peru Argentaria [n. sp.#5] Lamas, MS TL: Peru Argentaria [n. sp.#6] Lamas, MS TL: Peru Argentaria [n. sp.#7] Lamas, MS TL: Peru Argentaria [n. sp.#8] Lamas, MS TL: ? Argentaria [n. sp.#9] Lamas, MS TL: Brazil (SP) Argentaria [n. sp.#10] Lamas, MS TL: Brazil (MG) Argentaria [n. sp.#11] Lamas, MS TL: Peru Argentaria [n. sp.#12] Lamas, MS TL: Peru Argentaria [n. sp.#13] Lamas, MS TL: Peru Argentaria [n. sp.#14] Lamas, MS TL: Venezuela Argentaria [n. sp.#15] Lamas, MS TL: Peru Argentaria [n. sp.#16] Lamas, MS TL: Brazil Argentaria [n. sp.#17] Lamas, MS TL: Brazil (ES) Argentaria [n. sp.#18] Lamas, MS TL: Peru Saurona Huertas & Willmott, [2023] Saurona aurigera (Weymer, [1911]) ref.201 iN M T Saurona triangula (Aurivillius, 1929) ref.201 iN M T Saurona [n. sp.] Lamas, MS Peru Nhambikuara Freitas, Barbosa & Zacca, 2018 Nhambikuara mima (A. Butler, 1867) ref.216 iN M T Nhambikuara cerradensis Freitas, Barbosa & Zacca, 2018 ref.216 iN Nhambikuara ackeryi (Huertas, C. Rios & LeCrom, 2009) ref.201 iN Nhambikuara doxes (Godart, [1824]) ref.201 iN M T Nhambikuara furina (Hewitson, 1862) ref.201 iN M T Nhambikuara junonia (A. Butler, 1867) ref.201 iN M T Nhambikuara latia (A. Butler, 1867) ref.201 M T Nhambikuara mercedes (Huertas, 2011) ref.201,226 Nhambikuara toynei (Willmott & J. Hall, 1995) ref.201 iN M T Stegosatyrus Zacca, O. Mielke & Pyrcz, 2013 Stegosatyrus hemiclara Pyrcz, Boyer & Zacca, 2013 ref.363 Stegosatyrus imbrialis (Weeks, 1901) ref.363 iN M T Stegosatyrus ocelloides (Schaus, 1902) ref.363 iN M T Stegosatyrus periphas (Godart, [1824]) ref.363 iN M T Stevenaria Viloria, Costa, Neild & Nakahara, 2016 Stevenaria divergens (A. Butler, 1867) TL: Brazil (Am) M T Stevenaria nakaharai Viloria, Costa, Fratello & Neild, 2016 ref.189 Stevenaria yutajensis Viloria & Costa, 2016 ref.189 Taydebis A. Freitas, 2003 Taydebis clarissa (A. Freitas & O. Mielke, 2011) ref.371 iN Taydebis guria Zacca, Casagrande & O. Mielke, 2021 TL: Brazil Taydebis melobosis (Capronnier, 1874) ref.371 M T Sepona Freitas & E. Barbosa, 2016 Sepona punctata (Weymer, 1911) ref.215 iN M T Harjesia Forster, 1964 Harjesia argentata Nakahara, Zacca & Lamas, 2018 ref.261 Harjesia blanda (Möschler, 1877) iN M T S Harjesia obscura (A. Butler, 1867) iN M T S L Harjesia [n. sp.#1] Lamas, MS TL: Peru Harjesia [n. sp.#2] Lamas, MS TL: Peru Harjesia [n. sp.#3] Lamas, MS TL: Peru Harjesia [n. sp.#4] Lamas, MS TL: Peru Parataygetis Forster, 1964 Parataygetis albinotata (A. Butler, 1867) iN M T S Parataygetis lineata (Godman & Salvin, 1880) White-lined Satyr Costa Rica to Colombia iN M T S L Posttaygetis Forster, 1964 Posttaygetis penelea (Cramer, 1777) Yellow-spiked Satyr Costa Rica to Brazil iN M T S L Forsterinaria R. Gray, 1973 Forsterinaria itatiaia C. Peña & Lamas, 2005 ref.276 iN Forsterinaria quantius (Godart, [1824]) iN M T Forsterinaria necys (Godart, [1824]) iN M T Forsterinaria pronophila (A. Butler, 1867) ref.243 iN M T Forsterinaria neonympha (C. Felder & R. Felder, 1867) White-dotted Satyr [= umbracea; = polyphemus; = cyclops], S Mexico to W Ecuador iN M T S L Forsterinaria stella (Hayward, 1957) iN M T S L Forsterinaria pyrczi C. Peña & Lamas, 2005 ref.276 iN Forsterinaria proxima (Hayward, 1957) iN M T S Forsterinaria pseudinornata (Forster, 1964) TL: Ecuador M T Forsterinaria punctata C. Peña & Lamas, 2005 ref.276 Forsterinaria anophthalma (C. Felder & R. Felder, 1867) iN M T Forsterinaria enjuerma C. Peña & Lamas, 2005 ref.276 iN Forsterinaria pichita C. Peña & Lamas, 2005 ref.276 iN M S Forsterinaria guaniloi C. Peña & Lamas, 2005 iN Forsterinaria pilosa C. Peña & Lamas, 2005 ref.276 Forsterinaria falcata C. Peña & Lamas, 2005 ref.276 Forsterinaria rotunda C. Peña & Lamas, 2005 ref.276 Forsterinaria emo A. Zubek, T. Pyrcz & Boyer, 2014 ref.374 Forsterinaria hannieri A. Zubek, T. Pyrcz & Boyer, 2011 ref.376 Forsterinaria difficilis (Forster, 1964) TL: Bolivia M T Forsterinaria rustica (A. Butler, 1868) iN Forsterinaria rustica rustica (A. Butler, 1868) TL: Bolivia M T S Forsterinaria rustica villarresi (Dognin, 1887) TL: Ecuador M T S Forsterinaria rustica glendita C. Peña & Lamas, 2005 TL: Peru Forsterinaria antje C. Peña & Lamas, 2005 ref.276 M S Forsterinaria inornata (C. Felder & R. Felder, 1867) iN M T S Forsterinaria pallida C. Peña & Lamas, 2005 iN Forsterinaria pallida aurita C. Peña & Lamas, 2005 TL: Ecuador M L Forsterinaria pallida coipa C. Peña & Lamas, 2005 TL: Peru M S Forsterinaria pallida pallida C. Peña & Lamas, 2005 TL: Peru Forsterinaria boliviana (Godman, 1905) iN M T S Forsterinaria chaniorum Viloria & Costa, 2019 TL: Venezuela Orotaygetis Nakahara & Zacca, 2018 Orotaygetis surui Nakahara, Zacca & Lamas, 2018 ref.261 Pseudodebis Forster, 1964 Pseudodebis celia (Cramer, 1779) Celia Satyr Nicaragua to Brazil & Bolivia; ref.243 iN M T S L Pseudodebis darrenthroopi Nakahara & Willmott, 2022 TL: Costa Rica Pseudodebis nakamurai Nakahara & Willmott, 2021 TL: Ecuador Pseudodebis pieti Nakahara & Willmott, 2021 TL: Ecuador Pseudodebis tigrillo Nakahara & Willmott, 2022 TL: Ecuador Pseudodebis puritana (Weeks, 1902) iN Pseudodebis puritana puritana (Weeks, 1902) ref.243 M T S Pseudodebis puritana [n. ssp.] L. Miller, MS Brazil (Pr) M T Pseudodebis marpessa (Hewitson, 1862) iN M T S Pseudodebis dubiosa Forster, 1964 iN M T Pseudodebis euptychidia (A. Butler, 1868) iN M T Pseudodebis valentina (Cramer, 1779) iN M T S Pseudodebis vrazi (Kheil, 1896) ref.261 M T Pseudodebis zimri (A. Butler, 1869) Butler's Satyr SE Mexico to Colombia iN M T S L Taguaiba A. Freitas, Zacca & R. Siewert, [2023] Taguaiba rectifascia (Weymer, 1907) ref.201 iN M T Taguaiba ypthima (Hübner, [1821]) ref.201 iN M T Taguaiba drogoni (Siewert, Zacca, Dias & Freitas, 2013) Drogon's Satyr ref.201 Taguaiba servius (Weymer, 1910) ref.201 Taguaiba fulginia (R.F. d'Almeida, 1922) ref.201 M T Taygetina Forster, 1964 Taygetina banghaasi (Weymer, 1910) Bang-Haas' Satyr Costa Rica & Bolivia iN M T S L Taygetina accacioi Nakahara & Freitas, 2019 ref.263 Taygetina brocki Lamas & Nakahara, 2019 ref.263 Taygetina oreba (A. Butler, 1870) Oreba Satyr Costa Rica to Peru; ref.243 iN M T S Taygetina gulnare (A. Butler, 1870) iN M T S L Taygetina weymeri (Draudt, 1912) Mexican Satyr W Mexico; ref.243 iN M T S L Taygetina kerea (A. Butler, 1869) Kerea Satyr E & W Mexico to Paraguay & Argentina; ref.243 iN M T S L Taygetina peribaea (Godman & Salvin, 1880) ref.243 M T S Taygetis Hübner, [1819] Taygetis acuta Weymer, 1910 iN M T Taygetis angulosa Weymer, 1907 iN M T S Taygetis asterie Weymer, 1910 iN Taygetis chiquitana Forster, 1964 iN M T Taygetis chrysogone E. Doubleday, [1849] iN M T S Taygetis cleopatra C. Felder & R. Felder, 1867 iN M T S Taygetis echo (Cramer, 1775) iN Taygetis echo echo (Cramer, 1775) TL: Surinam M T Taygetis echo koepckei Forster, 1964 TL: Peru M T Taygetis echo velutina Staudinger, [1887] TL: Venezuela M T Taygetis echo [n. ssp.] L. Miller, MS Trinidad Taygetis elegia Weymer, 1910 iN M T Taygetis [n. sp.#1] L. Miller, MS TL: Peru Taygetis godmani Weymer, 1910 iN M T Taygetis inambari L. Miller & Lamas, 1999 iN M T Taygetis inconspicua Draudt, 1931 Inconspicuous Satyr SE Mexico, Guatemala iN M T Taygetis kharisma Brévignon & M. Benmesbah, 2011 ref.179 Taygetis laches Fabricius, 1793 iN Taygetis laches laches (Fabricius, 1793) TL: Guyana M T Taygetis laches marginata Staudinger, [1887] TL: Brazil (SP) M T Taygetis laches [n. ssp.] L. Miller, MS Peru Taygetis [n. sp.#2] L. Miller, MS TL: Peru Taygetis larua C. Felder & R. Felder, 1867 Laura's Satyr Panama to Bolivia iN M T S L Taygetis leuctra A. Butler, 1870 Leuctra Satyr Chiapas to Panama (at least) iN M T S L Taygetis mermeria (Cramer, 1776) iN Taygetis mermeria mermeria (Cramer, 1776) TL: Surinam M T S L Taygetis mermeria excavata A. Butler, 1868 E Mexico to Panama M T S L Taygetis mermeria griseomarginata L. Miller, 1978 W Mexico M T S Taygetis nympha A. Butler, 1868 iN M T Taygetis oyapock Brévignon, 2007 ref.175 iN Taygetis rufomarginata Staudinger, 1888 Rufous-margined Satyr [S Mexico?], Honduras, Costa Rica & Panama iN Taygetis rufomarginata rufomarginata Staudinger, 1888 [S Mexico?], Honduras, Costa Rica & Panama M T S L Taygetis rufomarginata tapulunmi Brévignon, 2005 TL: French Guiana Taygetis salvini Staudinger, [1887] iN M T Taygetis sosis Hopffer, 1874 Sosis Satyr [= xenana godmani], [S Mexico] to Brazil & Peru iN M S L Taygetis sylvia H. Bates, 1866 Sylvia's Satyr Panama to Bolivia iN M T S L Taygetis thamyra (Cramer, 1779) Thamyra Satyr E Mexico to S Brazil iN M T S L Taygetis tripunctata Weymer, 1907 iN M T S Taygetis uncinata Weymer, 1907 Hook-lined Satyr W & S Mexico, Guatemala iN M T S L Taygetis uzza A. Butler, 1869 Uzza Satyr [= salvini], [S Mexico?], Guatemala to Colombia iN M T OI S L Taygetis virgilia (Cramer, 1776) Stub-tailed Satyr E & W Mexico to Brazil iN M T S L Taygetis zippora A. Butler, 1869 TL: French Guiana M T Yphthimoides Forster, 1964 Yphthimoides angularis (A. Butler, 1867) iN M T Yphthimoides bella E. Barbosa & Freitas, 2015 ref.162 Yphthimoides blanquita E. Barbosa, M. Marín & Freitas, 2016 ref.163 iN Yphthimoides borasta (Schaus, 1902) iN M T S Yphthimoides celmis (Godart, [1824]) iN Yphthimoides acmenis (Hübner, 1823) iN M T Yphthimoides cipoensis Freitas, 2004 ref.207 iN Yphthimoides gabriela E. Barbosa, Freitas & M. Paluch, 2015 ref.162 Yphthimoides iserhardi Freitas & E. Barbosa, 2015 ref.162 Yphthimoides kinyoni Lamas & Nakahara, 2021 TL: Peru Yphthimoides leguialimai (Dyar, 1913) iN M T Yphthimoides manasses (C. Felder & R. Felder, 1867) TL: Brazil (Ba) M T Yphthimoides nareta E. Barbosa & Freitas, 2016 ref.163 Yphthimoides ochracea (A. Butler, 1867) iN M T L Yphthimoides ordinaria Freitas, L. Kaminski & O. Mielke, 2012 ref.211 iN Yphthimoides pacta (Weymer, 1911) iN M T Yphthimoides patricia (Hayward, 1957) iN Yphthimoides renata (Stoll, 1780) Renata Satyr S Mexico to W Ecuador iN M T S L1 L2 Yphthimoides saltuensis (Hayward, 1962) TL: ? M T Yphthimoides straminea (A. Butler, 1867) iN M T Yphthimoides yphthima (C. Felder & R. Felder, 1867) iN M T Yphthimoides inornata (Hayward, 1962) ref.277 iN M T Yphthimoides [n. sp.#1] Lamas, MS TL: Peru Yphthimoides [n. sp.#2] Lamas, MS TL: Peru Malaveria Viloria & M. Benmesbah, 2021 ref.167 Malaveria affinis (A. Butler, 1867) argyrospila A.G. Butler 1867 0 Euptychia Brazil (Am) S 3550 156 ...; ref.167 iN M T Malaveria alcinoe (C. Felder & R. Felder, 1867) Simple Satyr S Mexico to Bolivia; ref.167 iN M T S L Malaveria argyrospila (A. Butler, 1867) ref.201 M T Malaveria ballofi M. Benmesbah & Viloria, 2021 ref.167 Malaveria bottoi M. Benmesbah & Viloria, 2021 ref.167 Malaveria duponti M. Benmesbah & Murienne, 2021 ref.167 Malaveria erigone (A. Butler, 1867) ref.201 M T Malaveria grimon (Godart, [1824]) ref.201 iN Malaveria guenzeli (Anken, 1994) ref.201 Malaveria iseai (Viloria, [2022]) ref.201 Malaveria maepius (Godart, [1824]) iN M T Malaveria mimas (Godman, 1905) ref.167 iN M T S Malaveria mimula (Hayward, 1954) iN M T Malaveria mythra (Weymer, 1911) iN M T Malaveria nebulosa (A. Butler, 1867) ref.167 iN M T Malaveria neomaenas (Hayward, 1967) ref.167 M T Malaveria rodriguezi M. Benmesbah & Viloria, 2021 ref.167 iN Koutalina Viloria & M. Benmesbah, 2021 ref.167 Koutalina pamela (Hayward, 1957) ref.167 iN Zischkaia Forster, 1964 Zischkaia amalda (Weymer, 1911) TL: Bolivia M T Zischkaia arctoa Nakahara, 2019 ref.262 Zischkaia chullachaki Nakahara & Zacca, 2019 ref.262 iN Zischkaia baku Zacca, Dolibaina & Dias, 2019 ref.262 iN Zischkaia arenisca Nakahara, Willmott & J. Hall, 2019 ref.262 iN Zischkaia argyrosflecha Nakahara, L. Miller & Huertas, 2019 ref.262 Zischkaia pacarus (Godart, [1824]) iN M T Zischkaia abanico Nakahara & Petit, 2019 ref.262 Zischkaia saundersii (A. Butler, 1867) iN M T Zischkaia josti Nakahara & Kleckner, 2019 ref.262 Zischkaia mielkeorum Dolibaina, Dias & Zacca, 2019 ref.262 Zischkaia warreni Dias, Zacca & Dolibaina, 2019 ref.262 iN Amphidecta A. Butler, 1867 Amphidecta calliomma (C. Felder & R. Felder, 1862) Calliomma Satyr Colombia, Panama iN M T S L Amphidecta clio (Weymer, 1911) ref.201 iN M T Amphidecta pignerator A. Butler, 1867 Stub-tailed Satyr iN Amphidecta pignerator pignerator A. Butler, 1867 SE Mexico (CHIS) to Amazon Basin M T S L Amphidecta pignerator simplicia Weymer, 1910 TL: Brazil (MG) M S Amphidecta reynoldsi Sharpe, 1890 iN Amphidecta reynoldsi reynoldsi Sharpe, 1890 TL: Brazil (Go) M T S Amphidecta reynoldsi [n. ssp.#1] Lamas, MS TL: Brazil (MG/SC) M T Amphidecta reynoldsi [n. ssp.#2] Lamas, MS TL: Colombia Genera-unknown1 , [] not assigned to a genus yet Genera-unknown1 insignis (A. Butler, 1867) TL: [Brazil (Am)] M T Genera-unknown1 ordinata (Weymer, 1911) iN M T Tribe Satyrini, Subtribe Coenonymphina Oressinoma E. Doubleday, [1849] Oressinoma sorata Salvin & Godman, 1868 iN Oressinoma sorata sorata Salvin & Godman, 1868 TL: Bolivia M T S Oressinoma sorata [n. ssp.] Lamas, MS Peru Oressinoma sorina Florczyk, Fåhraeus & Pyrcz, 2021 TL: Peru Oressinoma typhla E. Doubleday, [1849] iN Oressinoma typhla Northern segregate , Costa Rica & Panama M S L Oressinoma typhla typhla E. Doubleday, [1849] TL: Colombia M T Oressinoma typhla boliviana Forster, 1964 TL: Bolivia M T Oressinoma typhla [n. ssp.#1] Lamas, MS TL: Costa Rica Oressinoma typhla [n. ssp.#2] Lamas, MS TL: Peru Oressinoma typhla [n. ssp.#3] Lamas, MS TL: Peru Coenonympha Hübner, [1819] Coenonympha haydenii (W. H. Edwards, 1872) Hayden's Ringlet iN M S L1 L2 Ha Coenonympha tullia (Müller, 1764) Large Heath ref.11 iN Coenonympha tullia mixturata Alpheraky, 1897 this name may only apply to Siberian populations: existence of mixturata in Ameri ... M S Coenonympha tullia yukonensis W. Holland, 1900 USA (Ak) & Can (YT); ref.11 M T S1 S2 L Coenonympha tullia kodiak (W. H. Edwards, 1869) USA (Ak); ref.11 M S1 S2 IL Coenonympha california Westwood, 1851 Common Ringlet ref.11 iN Coenonympha california nipisiquit McDunnough, 1939 salt marshes in NB & QC (Canada). Layberry et al. (1998), Webster (1999), and Sei ... M T S Coenonympha california mcisaaci dos Passos, 1935 TL: Canada (NL) M T S Coenonympha california inornata W. H. Edwards, 1861 TL: Canada (MB) M T S L Ha Coenonympha california benjamini McDunnough, 1928 TL: Canada (AB) M S L Ha Coenonympha california mackenziei D. Davenport, 1936 TL: Canada (NT) M T S Coenonympha california insulana McDunnough, 1928 TL: Canada (BC) M S L Coenonympha california yontocket Porter & Mattoon, 1989 TL: USA (Ca) M T1 T2 S Coenonympha california eryngii H. Edwards, 1877 TL: USA (Ca) M T S L Ha Coenonympha california california Westwood, 1851 TL: USA (Ca) M T S L1 L2 I Ha Coenonympha california columbiana McDunnough, 1928 TL: Canada (BC) M S L Coenonympha california eunomia (Dornfeld, 1967) TL: USA (Or) M T S L Ha Coenonympha california ampelos W. H. Edwards, 1871 TL: USA (Ca/Or) M T S1 S2 L I Ha Coenonympha california mono Burdick, 1942 TL: USA (Ca) M S L Coenonympha california elko W. H. Edwards, 1881 TL: USA (Nv) M S1 S2 Coenonympha california ochracea W. H. Edwards, 1861 TL: USA ([Co]) M T S L Ha Coenonympha california pseudobrenda Austin & R. Gray, 1998 TL: USA (Nv) M S1 S2 L Coenonympha california furcae W. Barnes & Benjamin, 1926 TL: USA (Az) M T S Coenonympha california subfusca W. Barnes & Benjamin, 1926 TL: USA (Az) M T S L Ha Tribe Satyrini, Subtribe Erebiina Cercyonis Scudder, 1875 Cercyonis pegala (Fabricius, 1775) Common Wood-Nymph iN Cercyonis pegala pegala (Fabricius, 1775) TL: [USA (SC)] M T S L1 L2 Cercyonis pegala alope (Fabricius, 1793) (= maritima, = ochracea, = agawamensis); ref.141 M S L Cercyonis pegala texana (W. H. Edwards, 1880) TL: USA (Tx) M T S L I Cercyonis pegala Coahuila, low-elevation segregate , M S L Ha Cercyonis pegala nephele (W. Kirby, 1837) (= olympus, = borealis); ref.141 M S1 S2 S3 S4 L1 L2 I Ha Cercyonis pegala ino G. Hall, 1924 TL: Canada (Ab) M S Cercyonis pegala incana (W. H. Edwards, 1880) TL: USA (Wa) M S Cercyonis pegala boopis (Behr, 1864) TL: USA (Ca) M T S L Cercyonis pegala ariane (Boisduval, 1852) TL: USA (Ca) M S L1 L2 Ha Cercyonis pegala australosierra J. Emmel, T. Emmel & K. Davenport, 2008 TL: USA (Ca) M T I Cercyonis pegala Little Deschutes River segregate , Little Deschutes River & upper Deschutes R., Deschutes Co., OR M S1 S2 L Ha Cercyonis pegala Silver Lake segregate , vic. Silver Lake, Lake Co., OR M S L1 L2 L3 Ha Cercyonis pegala Crump Lake segregate , vic. Crump Lake, Lake Co., OR M S1 S2 Cercyonis pegala Ravendale segregate , M S Cercyonis pegala gabbii (W. H. Edwards, 1870) TL: USA (Or) M S L Cercyonis pegala carsonensis Austin, 1992 TL: USA (Nv) M T1 T2 Cercyonis pegala walkerensis Austin, 1992 TL: USA (Ca) M T1 T2 S Cercyonis pegala wheeleri (W. H. Edwards, 1873) TL: USA (Ca) M T Cercyonis pegala paucilineatus Austin, 1992 TL: USA (Nv) M T1 T2 Cercyonis pegala pluvialis Austin, 1992 TL: USA (Nv) M T L Cercyonis pegala paludum Austin, 1992 TL: USA (Nv) M S L I Ha Cercyonis pegala stephensi (W. G. Wright, 1905) TL: USA (Ca) M T S1 S2 L Ha Cercyonis pegala utahensis Austin, 1992 TL: USA (Ut) M T1 T2 S1 S2 Cercyonis meadii (W. H. Edwards, 1872) Mead’s Wood-Nymph iN Cercyonis meadii meadii (W. H. Edwards, 1872) TL: USA (Co) M T S L1 L2 Ha Cercyonis meadii alamosa T. Emmel & J. Emmel, 1969 TL: USA (Co) M S L1 L2 I H Ha Cercyonis meadii melania (Wind, 1946) Requires confirmation in NE Chihuahua M T S L Cercyonis meadii mexicana (R. Chermock, 1949) SE AZ, Sonora, W Chihuahua, etc.; ref.20 M T S Cercyonis sthenele (Boisduval, 1852) Great Basin Wood-Nymph iN Cercyonis sthenele masoni Cross, 1937 TL: USA (Co) M S L Cercyonis sthenele damei W. Barnes & Benjamin, 1926 Grand Canyon area; ref.20 Cercyonis sthenele paulus (W. H. Edwards, 1879) TL: USA (Nv) M S L1 L2 Cercyonis sthenele W OR Cascades segregate , M S Cercyonis sthenele sineocellata Austin & J. Emmel, 1998 TL: USA (Or) M T S L1 L2 Ha Cercyonis sthenele behrii (F. Grinnell, 1905) TL: USA (Ca) M S L Cercyonis sthenele sthenele (Boisduval, 1852) TL: USA (Ca) M T1 T2 TI S IL Cercyonis hypoleuca Hawks & J. Emmel, 1998 Santa Cruz Is., CA, USA; ref.20 Cercyonis silvestris (W. H. Edwards, 1861) Great Basin Wood-Nymph LT not coonspecific with sthenele; ref.16 Cercyonis silvestris incognita J. Emmel, T. Emmel & Mattoon, 2012 N CA; ref.16 iN M S L Cercyonis silvestris silvestris (W. H. Edwards, 1861) ref.16 M T S1 S2 L Ha Cercyonis oetus (Boisduval, 1869) Small Wood-Nymph iN Cercyonis oetus charon (W. H. Edwards, 1872) TL: USA (Co) M T S L1 L2 L3 L4 Ha Cercyonis oetus phocus (W. H. Edwards, 1874) TL: Canada (BC) M T S Cercyonis oetus oetus (Boisduval, 1869) TL: USA (Ca) M T S L Cercyonis oetus pallescens T. Emmel & J. Emmel, 1971 TL: USA (Nv) M S Ha Cercyonis oetus alkalorum Austin, 1998 TL: USA (Nv) M T1 T2 Erebia Dalman, 1816 Erebia magdalena Strecker, 1880 Magdalena Alpine iN Erebia magdalena hilchie Kemal & Koçak, 2007 N Rocky Mountains; W-central ALTA; E-central BC M T S Erebia magdalena magdalena Strecker, 1880 S Rocky Mountains; S-central MT, NW WY, N UT, CO, N-central NM M T S L I Ha Erebia mackinleyensis Gunder, 1932 Mt. McKinley Alpine N, C & E AK, W YT, N BC iN M S L Ha Erebia fasciata A. Butler, 1868 Banded Alpine iN Erebia fasciata avinoffi W. Holland, 1930 NW AK & E Russia M S Erebia fasciata fasciata A. Butler, 1868 Nunavut to C AK M S L Ha G Erebia discoidalis (W. Kirby, 1837) Red-disked Alpine Canada & NC USA (N MT, N ND, N MN, N WI ) iN M S G Erebia rossii (J. Curtis, 1835) Ross's Alpine iN Erebia rossii kuskoquima W. Holland, 1931 coastal SW AK (Yukon & Kuskoquim River drainages) M S Erebia rossii gabrieli dos Passos, 1949 vic. Denali National Park, AK M S L Ha Erebia rossii rossii (J. Curtis, 1835) most of range M S G Erebia rossii ornata Leussler, 1935 vic. Churchill, Manitoba M T S Erebia disa (Thunberg, 1791) Disa Alpine iN Erebia disa steckeri W. Holland, 1930 N AK to W YK; tundra and forest habitats M S L G Erebia disa subarctica McDunnough, 1937 Richardson Mts. & MacKenzie Delta E to Nunavut M S Erebia mancinus E. Doubleday, [1849] Taiga Alpine taiga; Labrador to Alaska; S in Mts. To Banff, ALTA iN M S Erebia lafontainei Troubridge & Philip, 1983 Lafontaine's Alpine tundra AK, E YT & NW NT iN M OI S IL L Ha G Txt Erebia youngi W. Holland, 1900 Young's Alpine iN Erebia youngi rileyi dos Passos, 1947 mostly tundra, W AK coast & vic. Denali National Park, AK M S IL G Txt Erebia youngi youngi W. Holland, 1900 mostly tundra in C AK to W YK M S IL L Ha G Txt Erebia youngi herscheli Leussler, 1935 mostly tundra, coastal N AK & N YK M T S IL G Txt Erebia occulta Roos & Kimmich, 1983 Eskimo Alpine iN Erebia occulta occulta Roos & Kimmich, 1983 small scree areas in AK & NW YT M OI S IL L G Txt Erebia pawloskii Ménétriés, 1859 Theano Alpine iN Erebia pawloskii alaskensis W. Holland, 1900 AK, YK, N BC M S L Ha Erebia pawloskii canadensis B. Warren, 1931 vic. Churchill, Manitoba M S Erebia pawloskii ethela W. H. Edwards, 1891 higher elevations in SC MT, NW WY & NC CO M S L Ha G Erebia pawloskii demmia B. Warren, 1936 San Juan Range in SW CO M S L Erebia epipsodea A. Butler, 1868 Common Alpine iN Erebia epipsodea remingtoni P. Ehrlich, 1952 SE AK, YT & NW BC M S L Erebia epipsodea sineocellata Skinner, 1889 NE BC, AB (except extreme SW), S SK, SW MB, N ID & N MT M S G Erebia epipsodea epipsodea A. Butler, 1868 S BC, SW AB, NW USA M S L1 L2 Ha G Erebia epipsodea brucei Elwes, 1889 moist meadows in S Rocky Mountains; SE WY, E UT, CO & N NM M T S L Ha Erebia vidleri Elwes, 1898 Vidler's Alpine W BC & NW WA iN M OD S L Ha G Erebia callias W. H. Edwards, 1871 Colorado Alpine iN Erebia callias callias W. H. Edwards, 1871 alpine meadows of SE MT, W WY, NE UT & C CO M S L1 L2 Ha Tribe Satyrini, Subtribe Satyrina Oeneis Hübner, [1819] Oeneis ridingsii (W. H. Edwards, 1865) Ridings' Satyr ref.10 iN Oeneis ridingsii minimus (Austin, 1986) S ALTA E to SW Manitoba, N MT, W ND M T S Oeneis ridingsii wyomingo (Scott, 1998) N CO, WY, MT, UT; flies in late summer M S1 S2 Oeneis ridingsii ridingsii (W. H. Edwards, 1865) C MT to N MN, W NE M S1 S2 L1 L2 Ha Oeneis ridingsii curicata (M. Fisher, Scott & Garhart, 2008) upper Gunnison River Valley, Gunnison Co., CO M T L Ha Oeneis ridingsii coloalbiterra (Garhart & M. Fisher, 2008) Roan Cliffs, Garfield Co., CO M T Oeneis ridingsii neomexicanus (Austin, 1986) S-central AZ & NM M T S L Ha Oeneis ridingsii stretchii (W. H. Edwards, 1870) W WY to W CO, W to SE OR, NV, N AZ M S L Ha Oeneis ridingsii pallidus (Austin, 1986) Sierra Nevada (CA-NV), N to C Oregon M S L1 L2 H Oeneis carmen (A. Warren, Austin, Llorente, Luis & Vargas, 2008) Joboni Satyr northern Coah to southern NL; ref.108,10 iN M T S L H Oeneis pseudochazaroides (Durden, 2008) TL: Mexico Oeneis jutta (Hübner, [1806]) Jutta Arctic iN Oeneis jutta terraenovae dos Passos, 1935 Newfoundland M T S Oeneis jutta balder (Guérin-Méneville, [1832]) SE Canada & NE US W to SE Manitoba M T OI S1 S2 Oeneis jutta ascerta Masters & Sorensen, 1968 N WI, MN, MI, adjacent parts of Ontario M T S L Oeneis jutta ridingiana F. Chermock & R. Chermock, 1940 SW Manitoba & SASK M S Oeneis jutta harperi dos Passos, [1977] N Manitoba & E NWT M T S Oeneis jutta leussleri Bryant, 1935 W NWT M T S Oeneis jutta alaskensis W. Holland, 1900 AK, E to YK & N BC M S1 S2 L I Ha Oeneis jutta chermocki Wyatt, 1965 W ALTA, S BC M S Oeneis jutta reducta McDunnough, 1929 S Rocky Mountains (far E ID, W MT, WY, NE UT, NW CO) M S L I1 I2 Ha Oeneis melissa (Fabricius, 1775) Melissa Arctic iN Oeneis melissa semidea (Say, 1828) White Mountains, New Hampshire M T S Oeneis melissa melissa (Fabricius, 1775) coastal Newfoundland & Labrador M T S Oeneis melissa semplei W. Holland, 1930 interior Labrador, N Quebec, margins of Hudson Bay M S Oeneis melissa assimilis A. Butler, 1868 Nunavut & NWT M S Oeneis melissa gibsoni W. Holland, 1930 W & N YK, AK M S L Oeneis melissa atlinensis C. Guppy & J. Shepard, 2001 NW BC, SW YK M S Oeneis melissa beanii Elwes & J. Edwards, 1893 S & C BC, SW & W-central ALTA, to N-central WA, MT, WY & NE UT M S L Ha Oeneis melissa lucilla W. Barnes & McDunnough, 1918 S Rocky Mountains (CO, N NM) M S L1 L2 Ha Oeneis polixenes (Fabricius, 1775) Polixenes Arctic iN Oeneis polixenes katahdin (Newcomb, 1901) Mt. Katahdin, Maine M T S Oeneis polixenes polixenes (Fabricius, 1775) E Canadian arctic; Quebec to Labrador M S Oeneis polixenes subhyalina (J. Curtis, 1835) arctic islands and coasts; Nunavut & NWT M S Oeneis polixenes beringianus Kurentsov, 1970 Churchill, Manitoba, W to AK, S to S ALTA; E Russia M T S1 S2 L Oeneis polixenes woodi Troubridge & Parshall, 1988 arctic coast of AK & YK M S Oeneis polixenes yukonensis A. Gibson, 1920 SW YK M S Oeneis polixenes brucei (W. H. Edwards, 1891) S Rocky Mountains (NW WY, C & N CO, N NM) M S L Ha Oeneis philipi Troubridge, 1988 Philip's Arctic C AK, YK, N BC iN M S Oeneis alpina Kurentsov, 1970 Sentinel Arctic iN Oeneis alpina excubitor Troubridge, Philip, Scott & J. Shepard, 1982 far NW NWT, YK, N AK M S G Oeneis bore (Esper, 1789) White-veined Arctic iN Oeneis bore taygete Geyer, [1830] Labrador & N Quebec M T S Oeneis bore gaspeensis dos Passos, 1949 montane Gaspé Peninsula, Quebec M S Oeneis bore hanburyi Watkins, 1928 N-central Canadian arctic, from NE Manitoba to YK & E-central AK M S L Ha Oeneis bore mckinleyensis dos Passos, 1949 vic. Denali National Park, AK M S L Ha Oeneis bore fordi dos Passos, 1949 W AK, Russian Far East M S Oeneis bore edwardsi dos Passos, 1949 S Rocky Mountains (S-central MT, NW WY, NE UT, SW CO) M S L Ha Oeneis alberta Elwes, 1893 Alberta Arctic iN Oeneis alberta alberta Elwes, 1893 Canada, N-central US M T S Oeneis alberta ojibwe Scott & Kondla, 2006 SE Manitoba M S Oeneis alberta oslari Skinner, 1911 CO M S1 S2 L Ha Oeneis alberta capulinensis F. Brown, 1970 NE NM & SE CO M S1 S2 Oeneis alberta daura (Strecker, 1894) AZ: Graham & Coconino Cos. M S L1 L2 Ha Oeneis tanana A. Warren & Nakahara, 2016 iN M S Oeneis chryxus (E. Doubleday, [1849]) Chryxus Arctic iN Oeneis chryxus strigulosa McDunnough, 1934 N WI, N MI, S & E Ontario, SW Quebec M S L Oeneis chryxus calais (Scudder, 1865) W Ontario, W to SASK M S Oeneis chryxus caryi Dyar, 1904 boreal ALTA, BC, NWT, YK, far E AK M T S Oeneis chryxus altacordillera Scott, 2006 high Canadian Zone in Rocky Mts. of Colorado M S1 S2 L Ha Oeneis chryxus chryxus (E. Doubleday, [1849]) Rocky Mountains from ALTA to NM M OI S1 S2 Ab HS L Ha Oeneis chryxus socorro R. Holland, 2010 NM: Socorro Co., Mt. Withington M OI Oeneis chryxus valerata Burdick, 1958 Olympic Mts., NW WA & Vancouver I. M T S L Ha Oeneis chryxus stanislaus Hovanitz, 1937 Sierra Nevada, CA-NV M T S L1 L2 Ha Oeneis chryxus ivallda (Mead, 1878) Sierra Nevada, CA M S L Oeneis nevadensis (C. Felder & R. Felder, 1867) Great Arctic iN Oeneis nevadensis gigas A. Butler, 1868 BC to NW OR (coastal) M S L Oeneis nevadensis nevadensis (C. Felder & R. Felder, 1867) BC to N CA (inland) M T S L I Ha Oeneis nevadensis iduna (W. H. Edwards, 1874) CA: Mendocino-Sonoma Cos. M S Oeneis macounii (W. H. Edwards, 1885) Macoun's Arctic iN M S Oeneis uhleri (Reakirt, 1866) Uhler's Arctic iN Oeneis uhleri cairnesi A. Gibson, 1920 NE AK, YK, far NW NWT M T OI OD S L Ha G Oeneis uhleri nahanni Dyar, 1904 Mackenzie Mountains, NWT M T Oeneis uhleri varuna (W. H. Edwards, 1882) S-central Canada and N-central US M S L G Oeneis uhleri reinthali F. Brown, 1953 CO: W slopes M T S L1 L2 Ha G Oeneis uhleri uhleri (Reakirt, 1866) CO: E slopes, E WY M S1 S2 L Ha G Oeneis uhleri SW Utah segregate , Tushar Mts. of Beaver-Piute Cos., Utah M S1 S2 Subfamily Nymphalidae-incertae-sedis Apanthesis , [] a fossil genus Apanthesis leuce Scudder, 1889 Fossil from Colorado Lithodryas , [] a fossil genus Lithodryas styx (Scudder, 1889) Fossil from Colorado Nymphalites , [] a fossil genus Nymphalites obscurum Scudder, 1889 Fossil from Colorado Nymphalites scudderi Beutenmüller & T. Cockerell, 1908 Fossil from Colorado |
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