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Family PAPILIONIDAE Latreille, [1802] – SWALLOWTAILS

  Subfamily Baroniinae

Baronia  Salvin, 1893  

Baronia brevicornis Salvin, 1893  Baronia  SW Mexico and Chiapas  iN

Baronia brevicornis brevicornis Salvin, 1893  SW Mexico  iN  M T1 T2 S1 S2 S3 L1 L2 I H

Baronia brevicornis rufodiscalis J. Maza & J. White, 1987  Chiapas  iN  M T S L

  Subfamily Parnassiinae

  Tribe Parnassiini

Parnassius  Latreille, 1804  

Parnassius eversmanni Ménétriés, [1850]  Eversmann's Parnassian    iN

Parnassius eversmanni thor H. Edwards, 1881  Alaska, YK, W NT, N BC  M S L I

Parnassius clodius Ménétriés, 1855  Clodius Parnassian    iN

Parnassius clodius strohbeeni Sternitzky, 1945  Santa Cruz Mtns, CA; apparently extinct  M T S1 S2 S3

Parnassius clodius clodius Ménétriés, 1855  N CA; Marin to Trinity cos.  M S I

Parnassius clodius claudianus Stichel, 1907  far N CA (Siskiyous) to SW BC; Cascades westward  M S1 S2 Ab L1 L2 Ha

Parnassius clodius pseudogallatinus Bryk, 1913  Cascades of N WA & S BC  M S

Parnassius clodius incredibilis Bryk, 1935  Mt. St. Elias, Alaska; types possibly mislabeled  M T

Parnassius clodius sol Bryk & Eisner, 1932  CA; Tulare to Modoc & S Siskiyou Cos.; Warners?; usually < 7000'  M S L I H

Parnassius clodius baldur W. H. Edwards, 1877  CA; S Sierra Nevada, usually above 7000'  M S L1 L2 L3

Parnassius clodius S Sierra Nevada segregate ,  Southern Sierra Nevada (Kern-Tulare Cos.), CA  M S

Parnassius clodius altaurus Dyar, 1903  [= shepardi]; SE BC, E WA, ID, W MT, NW WY, E OR  M S L

Parnassius clodius menetriesii H. Edwards, 1877  N UT  M S1 S2 L

Parnassius phoebus (Fabricius, 1793)  Phoebus Parnassian    iN

Parnassius phoebus apricatus Stichel, 1906  S Alaska to NW BC  M T S L

Parnassius phoebus golovinus W. Holland, 1930  Seward Peninsula, Alaska  M T S

Parnassius smintheus E. Doubleday, 1847  Rocky Mountain Parnassian    iN

Parnassius smintheus sternitzkyi McDunnough, 1937  Siskiyou/ S Cascade Mts, NW CA-SW OR  M T S1 S2 L I Ha

Parnassius smintheus olympianna Burdick, 1941  Olympic Mts, NW WA; Vancouver I., SW BC  M T S L

Parnassius smintheus behrii W. H. Edwards, 1870  CA Sierra Nevada; ref.16  iN  M S1 S2 L Ha

Parnassius smintheus yukonensis Eisner, 1969  SW YK, NW BC  M S

Parnassius smintheus smintheus E. Doubleday, 1847  Rocky Mtns., E BC-W ALTA to N MT; ref.151  M T S L

Parnassius smintheus magnus W. G. Wright, 1905  S BC, E WA, E OR, ID, W MT, NW WY  M T S L1 L2 L3 H

Parnassius smintheus maximus Bryk & Eisner, 1937  C & E MT  M S

Parnassius smintheus sayii W. H. Edwards, 1863  [= hermodur]; NE NV, UT, S & E WY, W SD, W NB, CO, N NM  M T S1 S2 L1 L2 L3 I1 I2 H

Parnassius smintheus pseudorotgeri Eisner, 1966  SW CO; San Juan Mtns.  M T S L

  Subfamily Papilioninae

  Tribe Leptocircini

Eurytides  Hübner, [1821]  [includes Mimoides & others]

Eurytides iphitas Hübner, [1821]  TL: [Brazil]  M T

Eurytides dolicaon (Cramer, 1775)    iN

Eurytides dolicaon septentrionalis K. Brown, 1994  central Panama  M T S

Eurytides dolicaon hebreus K. Brown & Lamas, 1994  TL: Colombia  M T

Eurytides dolicaon tromes (Rothschild & Jordan, 1906)  TL: Venezuela  M T

Eurytides dolicaon cauraensis Möhn, 2002  TL: Venezuela

Eurytides dolicaon dolicaon (Cramer, 1775)  TL: East Indies  M T

Eurytides dolicaon deileon (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Eurytides dolicaon deicoon (C. Felder & R. Felder, 1864)  TL: Brazil  M T

Eurytides callias (Rothschild & Jordan, 1906)    iN

Eurytides callias attali Brévignon, 2006  TL: French Guiana

Eurytides callias callias (Rothschild & Jordan, 1906)    iN  M T

Eurytides columbus (Kollar, 1850)    iN  M T

Eurytides serville (Godart, [1824])    iN

Eurytides serville acritus (Rothschild & Jordan, 1906)  TL: Venezuela  M T

Eurytides serville serville (Godart, [1824])  TL: South America  M T L

Eurytides orabilis (A. Butler, 1872)  Thick-edged Kite-Swallowtail    iN

Eurytides orabilis orabilis (A. Butler, 1872)  Costa Rica & Panama  M T S

Eurytides orabilis isocharis (Rothschild & Jordan, 1906)  TL: Colombia  M T

Eurytides salvini (H. Bates, 1864)  Salvin’s Kite-Swallowtail  SE Mexico to Honduras  iN  M T S L

Eurytides bellerophon (Dalman, 1823)    iN  M T

Eurytides telesilaus (C. Felder & R. Felder, 1864)    iN

Eurytides telesilaus dolius (Rothschild & Jordan, 1906)  C Panama to Trinidad and NW Ecuador  M T S L

Eurytides telesilaus telesilaus (C. Felder & R. Felder, 1864)  TL: Colombia; Brazil (Am)  M T

Eurytides telesilaus salobrensis (R.F. d'Almeida, 1941)  TL: Brazil (MT)  M T

Eurytides telesilaus vitellus (Fruhstorfer, 1907)  TL: Brazil (Pr)  M T

Eurytides stenodesmus (Rothschild & Jordan, 1906)    iN  M T

Eurytides earis (Rothschild & Jordan, 1906)    iN  M T

Eurytides orthosilaus (Weymer, 1899)  TL: Paraguay  M T

Eurytides helios (Rothschild & Jordan, 1906)    iN  M T

Eurytides protesilaus (Linnaeus, 1758)    iN

Eurytides protesilaus dariensis (Rothschild & Jordan, 1906)  Costa Rica & Panama to N Venezuela  M T S L I

Eurytides protesilaus archesilaus (C. Felder & R. Felder, 1865)  TL: Colombia  M T S I1 I2

Eurytides protesilaus protesilaus (Linnaeus, 1758)  TL: North America  M T HS

Eurytides protesilaus nigricornis (Staudinger, 1884)  TL: Brazil (SP)  M T

Eurytides penthesilaus (C. Felder & R. Felder, 1865)  Five-striped Kite-Swallowtail  SE Mexico to Guatemala and El Salvador

Eurytides penthesilaus penthesilaus (C. Felder & R. Felder, 1865)  SE Mexico to Guatemala and El Salvador  M T S

Eurytides penthesilaus vilatelai (J. Maza & R.G. Maza, 2022)  TL: Mexico (Oax)

Eurytides macrosilaus (G. Gray, [1853])  Five-striped Kite-Swallowtail    iN

Eurytides macrosilaus macrosilaus (G. Gray, [1853])  Honduras to Costa Rica  M T S L

Eurytides macrosilaus leucones (Rothschild & Jordan, 1906)  TL: Colombia  M T

Eurytides molops (Rothschild & Jordan, 1906)    iN

Eurytides molops molops (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Eurytides molops hetaerius (Rothschild & Jordan, 1906)  TL: Colombia; Guyana; Peru; Surinam; Ecuador; Bolivia  M T

Eurytides molops megalurus (Rothschild & Jordan, 1906)  TL: Brazil (MG)  M T

Eurytides glaucolaus (H. Bates, 1864)  Glaucolaus Swallowtail    iN

Eurytides glaucolaus glaucolaus (H. Bates, 1864)  Panama to C Colombia  M T S

Eurytides glaucolaus melaenus (Rothschild & Jordan, 1906)  TL: Colombia  M T

Eurytides glaucolaus leucas (Rothschild & Jordan, 1906)  TL: Peru  M T

Eurytides leucosilaus (J. Zikán, 1937)  TL: Brazil (Am)  M T

Eurytides aguiari (R.F. d'Almeida, 1937)    iN  M T

Eurytides epidaus (E. Doubleday, 1846)  Mexican Kite-Swallowtail    iN

Eurytides epidaus tepicus (Rothschild & Jordan, 1906)  NW Mexico  M T S L

Eurytides epidaus fenochionis (Salvin & Godman, 1868)  SW Mexico  M T S L I H

Eurytides epidaus epidaus (E. Doubleday, 1846)  E Mexico to NW Costa Rica  M T S1 S2 L I1 I2

Eurytides paleoaztlanica (J. Maza & R.G. Maza, 2022)  TL: Mexico (Jal)

Eurytides tlahuica (RG. Maza & J. Maza, 2022)  TL: Mexico (Mor)

Eurytides agesilaus (Guérin-Méneville & Percheron, 1835)    iN

Eurytides agesilaus fortis (Rothschild & Jordan, 1906)  W Mexico  M T S Hy L

Eurytides agesilaus neosilaus (Hopffer, 1865)  E Mexico to Costa Rica  M T S L

Eurytides agesilaus eimeri (Rothschild & Jordan, 1906)  Panama & W Colombia  M T S L

Eurytides agesilaus agesilaus (Guérin-Méneville & Percheron, 1835)  TL: Colombia  M T

Eurytides agesilaus autosilaus (H. Bates, 1861)  TL: Brazil (Am)  M T

Eurytides agesilaus montanum (Küppers, 1974)  TL: Peru  M T

Eurytides agesilaus viridis (Röber, 1926)  TL: Colombia?

Eurytides marcellus (Cramer, 1777)  Zebra Swallowtail  E USA & Canada  iN  M T S1 S2 HS L1 L2 L3 L4 L5 L6 I1 I2 I3 I4 H

Eurytides zonaria (A. Butler, 1869)  Hispaniolan Kite-Swallowtail  Hispaniola  iN  M T S

Eurytides philolaus (Boisduval, 1836)  Dark Kite-Swallowtail    iN

Eurytides philolaus philolaus (Boisduval, 1836)  Texas and Sinaloa S to Costa Rica  M T S1 S2 Hy L1 L2 I

Eurytides philolaus huasteco (RG. Maza & J. Maza, 2022)  TL: Mexico (Tam)  M T

Eurytides philolaus xanticles (H. Bates, 1863)  Panama, N Colombia, N Venezuela  M T S L

Eurytides anaxilaus (C. Felder & R. Felder, 1865)  Small Kite-Swallowtail  Panama, N Colombia, N Venezuela  iN  M T S

Eurytides celadon (Lucas, 1852)  Cuban Kite-Swallowtail  Cuba & Isla de Juventud  iN  M T S

Eurytides marcellinus (E. Doubleday, [1845])  Jamaican Kite-Swallowtail  Jamaica  iN  M T S

Eurytides thyastes (Drury, 1782)    iN

Eurytides thyastes occidentalis R. F. Maza, 1982  SW Mexico  M T

Eurytides thyastes marchandii (Boisduval, 1836)  E Mexico to Honduras  M S

Eurytides thyastes panamensis (Oberthür, 1879)  Nicaragua and Costa Rica to NW Ecuador  M T S L I

Eurytides thyastes thyastinus (Oberthür, 1879)  TL: Ecuador  M T

Eurytides thyastes thyastes (Drury, 1782)  TL: Brazil (RJ)  M T

Eurytides calliste (H. Bates, 1864)  Yellow Kite-Swallowtail    iN

Eurytides calliste calliste (H. Bates, 1864)  SE Mexico to El Salvador and Honduras  M T S

Eurytides calliste olbius (Rothschild & Jordan, 1906)  Costa Rica to W Panama  M T S L

Eurytides dioxippus (Hewitson, [1856])  Thick-bordered Kite-Swallowtail    iN

Eurytides dioxippus lacandones (H. Bates, 1864)  SE Mexico to W Panama  M T S L

Eurytides dioxippus dioxippus (Hewitson, [1856])  C Panama to Colombia  M T S L

Eurytides dioxippus marae T. Racheli, Bollino & Sala, 1990  TL: Venezuela  M T

Eurytides dioxippus diores (Rothschild & Jordan, 1906)  TL: Peru  M T

Eurytides dioxippus flotti Schäffler & Frankenbach, 2011  TL: not specified

Eurytides leucaspis (Godart, 1819)    iN

Eurytides leucaspis lamidis (K. Brown & Lamas, 1994)  TL: Colombia  M T

Eurytides leucaspis leucaspis (Godart, 1819)  TL: Peru?  M T

Eurytides asius (Fabricius, 1781)    iN  M T

Eurytides lysithous (Hübner, [1821])    iN

Eurytides lysithous sebastianus (Oberthür, 1879)  TL: Brazil  M T

Eurytides lysithous harrisianus (Swainson, 1822)  TL: ?  M T

Eurytides lysithous rurik (Eschscholtz, 1821)  TL: Brazil (SC)  M T

Eurytides lysithous lysithous (Hübner, [1821])  TL: ?  M T

Eurytides lysithous eupatorion (Lucas, [1859])  TL: Brazil  M T

Eurytides thymbraeus (Boisduval, 1836)  White-crescent Swallowtail    iN

Eurytides thymbraeus thymbraeus (Boisduval, 1836)  SE Mexico to Honduras  M T S L

Eurytides thymbraeus aconophos (G. Gray, [1853])  W Mexico  M S L I H

Eurytides pausanias (Hewitson, 1852)    iN

Eurytides pausanias prasinus (Rothschild & Jordan, 1906)  Costa Rica & W Panama  M T S L

Eurytides pausanias cleombrotus (Strecker, 1885)  E Panama to N Ecuador  M S L

Eurytides pausanias hermolaus (Guenée, 1872)  TL: Venezuela  M T

Eurytides pausanias tabaquita (Kaye, 1925)  TL: Trinidad  M T

Eurytides pausanias pausanias (Hewitson, 1852)  TL: Surinam; Brazil (Am)  M T S

Eurytides protodamas (Godart, 1819)    iN  M T

Eurytides phaon (Boisduval, 1836)  Variable Swallowtail    iN

Eurytides phaon phaon (Boisduval, 1836)  NE Mexico to Panama  M T S1 S2 L1 L2 I

Eurytides phaon whitei (RG. Maza & J. Maza, 2022)  TL: Mexico (Que)

Eurytides phaon poncei (J. Maza & R.G. Maza, 2022)  TL: Mexico (Oax)

Eurytides phaon therodamas (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Eurytides euryleon (Hewitson, [1856])    iN

Eurytides euryleon clusoculis (A. Butler, 1872)  Costa Rica & Panama  M T S1 S2 I1 I2

Eurytides euryleon euryleon (Hewitson, [1856])  TL: Colombia  M T

Eurytides euryleon pithonius (Rothschild & Jordan, 1906)  TL: Colombia  M T S I

Eurytides euryleon rodriguezi (Le Crom, Constantino & Salazar, 2002)  TL: Colombia  M T

Eurytides euryleon haenschi (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Eurytides euryleon anatmus (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Eurytides ilus (Fabricius, 1793)  Ilus Swallowtail    iN

Eurytides ilus occiduus (Vázquez, 1957)  W Mexico  M T S Ha

Eurytides ilus branchus (E. Doubleday, 1846)  E Mexico to N Costa Rica  M T S1 S2 L I

Eurytides ilus ilus (Fabricius, 1793)  Panama to N Colombia & C Venezuela  M T S L

Eurytides ariarathes (Esper, 1788)  ref.10  iN

Eurytides ariarathes evagorides (K. Brown & Lamas, 1994)  TL: Venezuela  M T

Eurytides ariarathes demerara (K. Brown & Lamas, 1994)  TL: Guyana  M T

Eurytides ariarathes ariarathes (Esper, 1788)  TL: South America  M T OI

Eurytides ariarathes gayi (Lucas, 1852)  TL: Peru  M T

Eurytides ariarathes illuminatus (Niepelt, 1928)  TL: Colombia  M T

Eurytides ariarathes arianus (Staudinger, 1884)  TL: Brazil (Am); Peru  M T

Eurytides ariarathes metagenes (Rothschild & Jordan, 1906)  TL: Brazil (Pa)  M T

Eurytides ariarathes leuctra (Rothschild & Jordan, 1906)  TL: Brazil (Go)  M T

Eurytides xeniades (Hewitson, 1867)    iN

Eurytides xeniades isus (Oberthür, 1879)  TL: Colombia  M T

Eurytides xeniades halex (Rothschild & Jordan, 1906)  TL: Colombia  M T

Eurytides xeniades xeniades (Hewitson, 1867)  TL: Ecuador  M T Ma L

Eurytides xeniades tabaconas (Joicey & Talbot, 1918)  TL: Peru  M T

Eurytides xeniades imaus (Rothschild & Jordan, 1906)  TL: Peru  M T

Eurytides xeniades signatus (Tyler, K. Brown & Wilson, 1994)  TL: [Peru]  M T S

Eurytides xynias (Hewitson, 1875)    iN

Eurytides xynias trapeza (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Eurytides xynias xynias (Hewitson, 1875)  TL: Bolivia  M T

Eurytides microdamas (Burmeister, 1878)    iN  M T

  Tribe Troidini

Battus  Scopoli, 1777  

Battus philenor (Linnaeus, 1771)  Pipevine Swallowtail    iN

Battus philenor philenor (Linnaeus, 1771)  Mexico, S & E USA & Canada  M T S L1 L2 L3 L4 I1 I2 I3 H1 H2 F

Battus philenor hirsuta (Skinner, 1908)  central and northern California  M T S L I H

Battus philenor insularis Vázquez, 1957  Islas Revillagigedo, Colima  M T S

Battus philenor orsua (Godman & Salvin, 1889)  Islas Tres Marias, Nayarit  M T S

Battus philenor acauda (Oberthür, 1879)  Yucatan Peninsula and surrounding areas  M T S L I

Battus philenor frailescana J. Maza & R.G. Maza, 2022  TL: Mexico (Chia)

Battus devilliersii (Godart, 1823)  De Villiers' Swallowtail  Cuba, N & S Andros  iN  M S

Battus zetides (Munroe, 1971)  Zetides Swallowtail  Hispaniola  iN  M S

Battus polydamas (Linnaeus, 1758)  Polydamas Swallowtail    iN

Battus polydamas polydamas (Linnaeus, 1758)  continental  M T S1 S2 HS L1 L2 I1 I2 I3 H

Battus polydamas lucayus (Rothschild & Jordan, 1906)  SE USA & Bahamas  M T S L I

Battus polydamas cubensis (Dufrane, 1946)  Cuba & Isla de Juventud  M T S

Battus polydamas jamaicensis (Rothschild & Jordan, 1906)  Jamaica  M T S

Battus polydamas polycrates (Hopffer, 1865)  Hispaniola  M T S L

Battus polydamas thyamus (Rothschild & Jordan, 1906)  Virgin Islands, Puerto Rico, Culebra  M T S L I

Battus polydamas antiquus (Rothschild & Jordan, 1906)  Antigua; extinct  M S

Battus polydamas christopheranus (A. Hall, 1936)  St. Kitts, St. Christopher, Montserrat, Nevis  M T

Battus polydamas neodamas (Lucas, 1852)  Guadeloupe & Marie-Galante  M T S

Battus polydamas dominicus (Rothschild & Jordan, 1906)  Dominica  M T S

Battus polydamas cebriones (Dalman, 1823)  [= xenodamus]; Martinique  M T S

Battus polydamas lucianus (Rothschild & Jordan, 1906)  St. Lucia  M T S

Battus polydamas vincentius (Rothschild & Jordan, 1906)  St. Vincent (possibly northern Grenadines)  M T S

Battus polydamas grenadensis (A. Hall, 1930)  Grenada, southern Grenadines  M T L

Battus polydamas streckerianus (Honrath, 1884)  TL: Peru  M T

Battus polydamas peruanus (Fuchs, 1954)  TL: Peru

Battus polydamas atahualpa T. Racheli & Pischedda, 1987  TL: Peru

Battus polydamas weyrauchi Lamas, 1998  TL: Peru  M T

Battus polydamas renani Lamas, 1998  TL: Peru  M T

Battus polydamas psittacus (J. Molina, 1782)  TL: Chile  M T

Battus madyes (E. Doubleday, 1846)    iN

Battus madyes adloni (Ehrmann, 1926)  TL: Ecuador  M T

Battus madyes philetas (Hewitson, 1869)  TL: Ecuador  M T

Battus madyes lojaensis Möhn, 1999  TL: Ecuador

Battus madyes magnimacula (Joicey & Talbot, 1925)  TL: Peru  M T

Battus madyes frankenbachi Möhn, 2001  TL: Peru

Battus madyes plinius (Weymer, 1890)  TL: Peru  M T

Battus madyes buechei Lamas, 1998  TL: Peru  M T

Battus madyes callangaensis Möhn, 2001  TL: Peru

Battus madyes chlorodamas (Guenée, 1872)  TL: Peru  M T

Battus madyes montebanus (Dyar, 1913)  TL: Peru  M T

Battus madyes crispatus Lamas, 2001  TL: Peru  M T

Battus madyes madyes (E. Doubleday, 1846)  TL: Bolivia  M T

Battus madyes tucumanus (Rothschild & Jordan, 1906)  TL: Argentina  M T

Battus eracon (Godman & Salvin, 1897)  West-Mexican Swallowtail  W Mexico  iN  M T S

Battus polystictus (A. Butler, 1874)    iN

Battus polystictus galenus (Fruhstorfer, 1907)  TL: Surinam  M T

Battus polystictus polystictus (A. Butler, 1874)  TL: Brazil (RS)  M T

Battus laodamas (C. Felder & R. Felder, 1859)    iN

Battus laodamas iopas (Godman & Salvin, 1897)  W Mexico  M T S L

Battus laodamas copanae (Reakirt, 1863)  E Mexico to N Costa Rica  M T S L

Battus laodamas rhipidius (Rothschild & Jordan, 1906)  Costa Rica to Panama  M T S

Battus laodamas laodamas (C. Felder & R. Felder, 1859)  TL: Colombia  M T

Battus ingenuus (Dyar, 1907)  Dyar's Swallowtail  SE Mexico to E Venezuela, Trinidad, and Ecuador  iN  M T S1 S2 L I1 I2

Battus lycidas (Cramer, 1777)  Cramer's Swallowtail  SE Mexico to N Bolivia and S Pará, Brazil  iN  M T S L

Battus belus (Cramer, 1777)    iN

Battus belus varus (Kollar, 1850)  TL: Colombia  M T

Battus belus belus (Cramer, 1777)  TL: Surinam  M T

Battus belus aureochloris K. Brown, 1994  TL: Brazil (Pa)

Battus belus belemus (H. Bates, 1864)  TL: Brazil (Pa)  M T

Battus belus cochabamba (Weeks, 1901)  TL: Bolivia  M T

Battus crassus (Cramer, 1777)    iN

Battus crassus lepidus (C. Felder & R. Felder, 1861)  Costa Rica to E Venezuela and W Ecuador  M T S L

Battus crassus crassus (Cramer, 1777)  TL: Surinam  M T

Battus crassus hirundo (Röber, 1925)  TL: Bolivia  M T

Battus crassus paraensis K. Brown, 1994  TL: Brazil (Pa)  M T

Euryades  C. Felder & R. Felder, 1864  

Euryades corethrus (Boisduval, 1836)    iN  M T

Euryades duponchelii (Lucas, 1839)    iN  M T

Parides  Hübner, [1819]  

Parides gundlachianus (C. Felder & R. Felder, 1864)  Cuban Cattleheart    iN

Parides gundlachianus alayoi L. Hernández, Alayón & D. Smith, 1995  Pinar del Río, W Cuba  M T S

Parides gundlachianus gundlachianus (C. Felder & R. Felder, 1864)  E Cuba  M T S

Parides ascanius (Cramer, 1775)    iN  M T

Parides agavus (Drury, 1782)    iN  M T

Parides proneus (Hübner, [1831])    iN  M T

Parides phalaecus (Hewitson, 1869)    iN

Parides phalaecus phalaecus (Hewitson, 1869)  TL: Ecuador  M T

Parides phalaecus nieva Möhn, 1999  TL: Peru

Parides bunichus (Hübner, [1821])    iN

Parides bunichus diodorus (Hopffer, 1865)  TL: Brazil  M T

Parides bunichus bunichus (Hübner, [1821])  TL: ?

Parides bunichus almas Casagrande & O. Mielke, 2007  TL: Brazil

Parides bunichus chamissonia (Eschscholtz, 1821)  TL: Brazil  M T

Parides bunichus perrhebus (Boisduval, 1836)  TL: Paraguay; Brazil (RS)  M T

Parides bunichus damocrates (Guenée, 1872)  TL: Argentina

Parides montezuma (Westwood, 1842)  Montezuma’s Cattleheart  NE & NW Mexico to Costa Rica  iN

Parides montezuma montezuma (Westwood, 1842)  NE & NW Mexico to Costa Rica  iN  M S L1 L2 H

Parides montezuma tyleri RG. Maza & J. Maza, 2022  TL: Mexico (SLP)

Parides montezuma tulana (Reakirt, 1863)  TL: Mexico (Chia)

Parides alopius (Godman & Salvin, 1890)  White-dotted Cattleheart  S Arizona (stray), W Mexico  iN  M T S L I

Parides photinus (E. Doubleday, 1844)  Pink-spotted Cattleheart  NE & NW Mexico to Costa Rica  iN  M T S L I H

Parides klagesi (Ehrmann, 1904)  TL: Venezuela  M T S

Parides vercingetorix (Oberthür, 1888)  TL: French Guiana  M T

Parides quadratus (Staudinger, 1890)  TL: Brazil (Am)

Parides quadratus spoliatus (Staudinger, 1898)  TL: Brazil (Am); Peru  M T

Parides quadratus quadratus (Staudinger, 1890)  TL: Brazil (Am)

Parides pizarro (Staudinger, 1884)  TL: Brazil (Am); Peru

Parides pizarro pizarro (Staudinger, 1884)  TL: Brazil (Am); Peru  M T

Parides pizarro kuhlmanni (E. May, 1925)  TL: Brazil (Ac)  M T S

Parides pizarro steinbachi (Rothschild, 1905)  TL: Bolivia  M T S

Parides hahneli (Staudinger, 1882)  TL: Brazil (Am)  M T

Parides mithras (Grose-Smith, 1902)  TL: Guyana

Parides mithras mithras (Grose-Smith, 1902)  TL: Guyana  M T

Parides mithras marajoara K. Brown, 1994  TL: Brazil (Pa)

Parides chabrias (Hewitson, 1852)    iN

Parides chabrias chabrias (Hewitson, 1852)  TL: Brazil (Am)  M T

Parides chabrias ygdrasilla Hemming, 1935  TL: ?  M T

Parides aeneas (Linnaeus, 1758)    iN

Parides aeneas bolivar (Hewitson, 1850)  TL: Brazil (Am)  M T

Parides aeneas huallaga T. Racheli, 1988  TL: Peru  M T

Parides aeneas damis (Rothschild & Jordan, 1906)  TL: Peru  M T

Parides aeneas tucha T. Racheli, 1988  TL: Peru  M T

Parides aeneas lamasi T. Racheli, 1988  TL: Peru  M S

Parides aeneas locris (Rothschild & Jordan, 1906)  TL: Bolivia  M T

Parides aeneas opalinus (A. Butler, 1877)  TL: Brazil (Am)  M T

Parides aeneas gabrieliensis (Bryk, 1953)  TL: Brazil (Am)  M S

Parides aeneas aeneas (Linnaeus, 1758)  TL: Asia  M T HS

Parides aeneas didas K. Brown & Lamas, 1994  TL: Surinam  M T

Parides aeneas lucasi K. Brown & Lamas, 1994  TL: French Guiana  M T

Parides aeneas marcius (Hübner, [1816])  TL: ?  M S

Parides aeneas linoides K. Brown & Lamas, 1994  TL: Brazil (Pa)  M T

Parides tros (Fabricius, 1793)    iN

Parides tros tros (Fabricius, 1793)  TL: Brazil

Parides tros danunciae O. Mielke, Casagrande & C. Mielke, 2000  TL: Brazil (Pr)  M T

Parides orellana (Hewitson, 1852)    iN  M T

Parides burchellanus (Westwood, 1872)    iN

Parides burchellanus burchellanus (Westwood, 1872)  TL: Brazil (MG)  M T

Parides burchellanus jaguarae (Foetterle, 1902)  TL: Brazil (MG; SP)  M T

Parides panthonus (Cramer, 1780)  TL: Surinam

Parides panthonus panthonus (Cramer, 1780)  TL: Surinam  M T

Parides panthonus phylarchus (Hopffer, 1865)  TL: French Guiana  M T

Parides aglaope (G. Gray, [1853])    iN

Parides aglaope aglaope (G. Gray, [1853])  TL: Brazil (Pa)  M T

Parides aglaope lysimachus (Honrath, 1888)  TL: Peru

Parides aglaope ecaudatus (Joicey & Talbot, 1924)  TL: Brazil (MT)  M T

Parides aglaope callicles (H. Bates, 1861)  TL: Bolivia  M T

Parides aglaope castilhoi R.F. d'Almeida, 1967  TL: Brazil (SP)  M T

Parides lysander Cramer, 1775    iN

Parides lysander orinocoensis Constantino, Le Crom & Salazar, [2002]  TL: Colombia  M T

Parides lysander brissonius (Hübner, [1819])  TL: ?  M T S

Parides lysander lysander (Cramer, 1775)  TL: Surinam  M T IL L

Parides lysander parsodes (G. Gray, [1853])  TL: Brazil (Pa)  M T

Parides lysander antalcidas Tyler, K. Brown & Wilson, 1994  TL: Brazil (Pa)  M T

Parides lysander mattogrossensis (Talbot, 1928)  TL: Brazil (MT)  M T

Parides echemon (Hübner, [1813])    iN

Parides echemon empistocles Küppers, 1975  TL: Peru

Parides echemon pisander (C. Felder & R. Felder, 1865)  TL: ?  M T

Parides echemon ergeteles (G. Gray, [1853])  TL: Brazil (Pa)  M T

Parides echemon echemon (Hübner, [1813])  TL: ?  M T

Parides eurimedes (Stoll, 1782)    iN

Parides eurimedes mylotes (H. Bates, 1861)  S Mexico to Costa Rica  M T S1 S2 L I

Parides eurimedes mycale (Godman & Salvin, 1890)  Panama and far N Colombia  M T S L

Parides eurimedes arriphus (Boisduval, 1836)  TL: Colombia  M T S

Parides eurimedes emilius Constantino, 1999  TL: Colombia  M T

Parides eurimedes antheas (Rothschild & Jordan, 1906)  TL: Colombia  M T S

Parides eurimedes eurimedes (Stoll, 1782)  TL: Berbices  M T S

Parides eurimedes agathokles (Kollar, 1850)  TL: [Colombia]  M T

Parides eurimedes timias (G. Gray, [1853])  TL: Ecuador  M T

Parides neophilus (Geyer, 1837)    iN

Parides neophilus parianus (Rothschild & Jordan, 1906)  TL: Trinidad; Venezuela  M T

Parides neophilus neophilus (Geyer, 1837)  TL: Surinam

Parides neophilus olivencius (H. Bates, 1861)  TL: Brazil (Am)  M T

Parides neophilus anaximenes (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T L

Parides neophilus consus (Rothschild & Jordan, 1906)  TL: Bolivia  M T

Parides neophilus ecbolius (Rothschild & Jordan, 1906)  TL: Brazil (Pa)  M T

Parides neophilus eurybates (G. Gray, [1853])  TL: [Brazil]  M T

Parides zacynthus (Fabricius, 1793)    iN

Parides zacynthus polymetus (Godart, 1819)  TL: Brazil; "Peru"  M T

Parides zacynthus zacynthus (Fabricius, 1793)  TL: Brazil  M HS

Parides childrenae (G. Gray, 1832)  Green-celled Cattleheart    iN

Parides childrenae childrenae (G. Gray, 1832)  SE Mexico to Panama  M T S1 S2 L I

Parides childrenae latifasciata (E. Krüger, 1925)  TL: Colombia  M T S

Parides childrenae esmeraldas Winhard, 2019  ref.ref.Willmott_Lamas_2020s,360

Parides childrenae unimacula (Joicey & Talbot, 1922)  TL: Ecuador  M T S L

Parides sesostris (Cramer, 1779)    iN

Parides sesostris zestos (G. Gray, [1853])  SE Mexico to Costa Rica  M T S1 S2 L I1 I2

Parides sesostris tarquinius (Boisduval, 1836)  Panama, transandean Colombia, N Venezuela  M T S L

Parides sesostris sesostris (Cramer, 1779)  TL: Surinam  M T S

Parides sesostris trinitensis K. Brown, 1994  TL: Trinidad  M T

Parides sesostris zischkai (Forster, 1955)  TL: Bolivia  M T

Parides panares (G. Gray, [1853])  Wedge-spotted Cattleheart    iN

Parides panares panares (G. Gray, [1853])  Oaxaca  M T1 T2 S L

Parides panares lycimenes (Boisduval, 1870)  Chiapas to Panama  M T S L I1 I2

Parides panares erythrus (Rothschild & Jordan, 1906)  TL: Colombia  M T

Parides panares paralius (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Parides panares rachelii K. Brown, 1994  TL: Venezuela  M T

Parides panares tachira T. Racheli, 1991  TL: Venezuela  M T

Parides vertumnus (Cramer, 1779)    iN

Parides vertumnus vertumnus (Cramer, 1779)  TL: Surinam  M T

Parides vertumnus pseudobogotanus (R. Krüger, 1926)  TL: French Guiana  M T

Parides vertumnus pyrophanus (J. Zikán, 1937)  TL: Brazil (Am)  M T

Parides vertumnus bogotanus (C. Felder & R. Felder, 1864)  TL: Colombia  M T

Parides vertumnus diceros (G. Gray, [1853])  TL: Brazil (Pa)  M T

Parides vertumnus cutora (G. Gray, [1853])  TL: Brazil (Am)  M T

Parides vertumnus astorius (J. Zikán, 1940)  TL: Brazil (Am)  M T

Parides vertumnus autumnus (Staudinger, 1898)  TL: Peru  M T

Parides vertumnus yuracares (Rothschild & Jordan, 1906)  TL: Bolivia  M T

Parides cutorina (Staudinger, 1898)  TL: Brazil (Am); Peru  M T

Parides phosphorus (H. Bates, 1861)    iN

Parides phosphorus gratianus (Hewitson, 1861)  TL: Colombia  M T

Parides phosphorus laurae Bollino & Costa, 2004  TL: Venezuela  M T

Parides phosphorus phosphorus (H. Bates, 1861)  TL: Guyana  M T S

Parides phosphorus elena L. Racheli & T. Racheli, 2021  TL: Peru

Parides phosphorus ernesto L. Racheli & T. Racheli, 2021  TL: Peru

Parides phosphorus zopyron Lamas, 1998  TL: Peru  M T

Parides phosphorus vavi T. Racheli, 1992  TL: Peru

Parides phosphorus mattia L. Racheli & T. Racheli, 2021  TL: Brazil

Parides anchises (Linnaeus, 1758)    iN

Parides anchises marthilia R. G. Maza, 1999  Chiapas  M T

Parides anchises lagartero RG. Maza & J. Maza, 2019  TL: Mexico

Parides anchises farfan K. Brown, 1994  Panama (Pacific Coast)  M T S

Parides anchises alyattes (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Parides anchises serapis (Boisduval, 1836)  TL: Colombia  M T

Parides anchises osyris (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Parides anchises maurocosta Bollino, L. Racheli & T. Racheli, 2021  TL: Venezuela

Parides anchises cymochles (E. Doubleday, 1844)  TL: Trinidad  M T

Parides anchises orinocensis (Rousseau-Decelle, 1960)  TL: Venezuela  M T

Parides anchises anchises (Linnaeus, 1758)  TL: America  M T HS

Parides anchises bukuti Brévignon, 1998  TL: French Guiana

Parides anchises thelios (G. Gray, [1853])  TL: Brazil (Pa)  M T

Parides anchises humaita D'Abrera, 1981  TL: Brazil (Am)  M T

Parides anchises drucei (A. Butler, 1874)  TL: Ecuador  M T

Parides anchises nielseni Bollino & Salazar, 2001  TL: Colombia  M T

Parides anchises koenigi T. Racheli, 1989  TL: Peru  M T

Parides anchises marinae T. Racheli, 1987  TL: Peru  M T

Parides anchises etias (Rothschild & Jordan, 1906)  TL: Bolivia  M T

Parides anchises orbignyanus (Lucas, 1852)  TL: Argentina  M T

Parides anchises foetterlei (Rothschild & Jordan, 1906)  TL: Brazil (SP)  M T

Parides anchises nephalion (Godart, 1819)  TL: Brazil  M T

Parides anchises stilbon (Kollar, 1839)  TL: Brazil  M T

Parides iphidamas (Fabricius, 1793)  Iphidamas Cattleheart    iN

Parides iphidamas kaneek RG. Maza & J. Maza, 2019  TL: Mexico

Parides iphidamas tacanaensis RG. Maza & J. Maza, 2019  TL: Mexico

Parides iphidamas trinitaria RG. Maza & J. Maza, 2019  TL: Mexico

Parides iphidamas iphidamas (Fabricius, 1793)  SE Mexico to Costa Rica  M T S1 S2 L I

Parides iphidamas Atlantic Coast, Panama segregate ,  Atlantic Coast, Panama  M S

Parides iphidamas elatos (Rothschild & Jordan, 1906)  TL: Colombia  M T

Parides iphidamas gorgonae Vélez & Salazar, 1991  TL: Colombia

Parides iphidamas phalias (Rothschild & Jordan, 1906)  TL: Colombia  M T

Parides iphidamas teneates (Rothschild & Jordan, 1906)  TL: Venezuela  M T

Parides iphidamas calogyna (Rothschild & Jordan, 1906)  TL: Ecuador; Colombia  M T

Parides iphidamas ayabacensis (Joicey & Talbot, 1918)  TL: Peru  M T

Parides iphidamas angustus Winhard, 2017  ref.234,358

Parides erithalion (Boisduval, 1836)    iN

Parides erithalion trichopus (Rothschild & Jordan, 1906)  W Mexico  M T S L I Ha

Parides erithalion polyzelus (C. Felder & R. Felder, 1865)  E Mexico to Costa Rica  M T S L I H

Parides erithalion izapa RG. Maza & J. Maza, 2022  TL: Mexico (Chia)

Parides erithalion sadyattes (H. Druce, 1874)  Nicaragua and N Costa Rica  M T S

Parides erithalion smalli K. Brown, 1994  S Costa Rica & Panama  M T1 T2 L

Parides erithalion erithalion (Boisduval, 1836)  TL: Colombia; "Jamaica"  M T

Parides erithalion cauca (Oberthür, 1879)  TL: Colombia  M S I

Parides erithalion chocoensis Constantino, Le Crom & Salazar, 2002  TL: Colombia  M T

Parides erithalion palmasensis K. Brown, 1994  TL: Ecuador  M T

Parides erithalion zeuxis (Lucas, 1852)  TL: Venezuela  M T

Parides erithalion inini Brévignon, 1998  TL: French Guiana

Parides erithalion keithi T. Racheli, 1991  TL: Venezuela  M S

Parides erithalion browni Le Crom, Constantino & Salazar, 2002  TL: Colombia

Parides erithalion kruegeri (Niepelt, 1927)  TL: Colombia  M T

Parides erithalion guillerminae Pischedda & T. Racheli, 1986  TL: Ecuador  M T

Parides erithalion lacydes (Hewitson, 1869)  TL: Ecuador  M T

Parides erithalion chinchipensis (Joicey & Talbot, 1918)  TL: Peru  M T

Parides erithalion xanthias (Rothschild & Jordan, 1906)  TL: Peru  M T

Parides erithalion callegarii T. Racheli & L. Racheli, 1996  TL: Peru

Parides erithalion racheliorum Lamas, 1998  TL: Peru  M T

Parides erithalion blanca T. Racheli & Möhn, 2001  TL: Peru

Parides erithalion yaminahua Pischedda & T. Racheli, 1987  TL: Peru

Parides erithalion erlaces (G. Gray, [1853])  TL: Bolivia  M T L

Parides donaji R. de la Maza & J. de la Maza, 2020  ref.246

  Tribe Papilionini, Subtribe Papilionina

Heraclides  Hübner, [1819]  

Heraclides hyppason (Cramer, 1775)    iN  M T

Heraclides chiansiades (Westwood, 1872)    iN

Heraclides chiansiades dospassosi (Rütimeyer, 1969)  TL: Colombia  M T

Heraclides chiansiades chiansiades (Westwood, 1872)  TL: Ecuador  M T

Heraclides chiansiades maroni (Moreau, 1923)  TL: French Guiana  M T

Heraclides chiansiades maripasoula Brévignon, 2006  TL: French Guiana

Heraclides chiansiades mossi K. Brown, 1994  TL: Brazil (Pa)  M T

Heraclides isidorus (E. Doubleday, 1846)  Ruby-patched Swallowtail    iN

Heraclides isidorus rhodostictus (A. Butler & H. Druce, 1874)  Costa Rica & W Panama  M T S L

Heraclides isidorus pacificus (Rothschild & Jordan, 1906)  E Panama to Ecuador  M T S

Heraclides isidorus nymphius (Rothschild & Jordan, 1906)  TL: Colombia  M T

Heraclides isidorus brises (Rothschild & Jordan, 1906)  TL: Colombia  M T

Heraclides isidorus autana T. Racheli & L. Racheli, 1995  TL: Venezuela  M S

Heraclides isidorus flavescens (Oberthür, 1879)  TL: Colombia  M T

Heraclides isidorus isidorus (E. Doubleday, 1846)  TL: Bolivia  M T

Heraclides isidorus tingo T. Racheli & L. Racheli, 1995  TL: Peru

Heraclides anchisiades (Esper, 1788)  Ruby-spotted Swallowtail    iN

Heraclides anchisiades idaeus (Fabricius, 1793)  Texas to Colombia & NW Venezuela  M T S L I1 I2 I3

Heraclides anchisiades lamasi K. Brown, 1994  TL: Ecuador  M T

Heraclides anchisiades philastrius (Fruhstorfer, 1915)  TL: Trinidad  M T

Heraclides anchisiades anchisiades (Esper, 1788)  TL: Surinam  M T OI

Heraclides anchisiades capys (Hübner, [1809])  TL: ?

Heraclides rogeri (Boisduval, 1836)  Pink-spotted Swallowtail  Yucatan Peninsula and surrounding areas  iN  M S L

Heraclides pharnaces (E. Doubleday, 1846)  Pink-spotted Swallowtail  S Texas, most of Mexico, to NW Guatemala  M T S L I

Heraclides polycharmus (Godman & Salvin, 1890)  TL: Mexico (Sin)

Heraclides polycharmus polycharmus (Godman & Salvin, 1890)  TL: Mexico (Sin)  M T

Heraclides polycharmus intiana (J. Maza & R.G. Maza, 2022)  TL: Mexico (Chia)

Heraclides epenetus (Hewitson, 1861)    iN  M T

Heraclides erostratus (Westwood, 1847)  Erostratus Swallowtail    iN

Heraclides erostratus erostratinus (Vázquez, 1947)  E Mexico  M T S L

Heraclides erostratus vazquezae (Beutelspacher, 1986)  SW Mexico  M T S L Ha

Heraclides erostratus erostratus (Westwood, 1847)  Chiapas, Guatemala, El Salvador  M T S

Heraclides oxynius (Geyer, [1827])  Oxynius Swallowtail  Cuba  M T S

Heraclides pelaus (Fabricius, 1775)  Caribbean Swallowtail    iN

Heraclides pelaus atkinsi (M. Bates, 1935)  Cuba  M T S

Heraclides pelaus pelaus (Fabricius, 1775)  Jamaica  M T S

Heraclides pelaus imerius (Godart, 1819)  Hispaniola  M T S

Heraclides pelaus puertoricensis Möhn, 1999  Puerto Rico  M S L

Heraclides garleppi (Staudinger, 1892)    iN

Heraclides garleppi interruptus (Staudinger, 1892)  TL: Brazil (Am); Peru  M T

Heraclides garleppi lecerfi K. Brown & Lamas, 1994  TL: French Guiana  M T

Heraclides garleppi garleppi (Staudinger, 1892)  TL: Bolivia  M T

Heraclides torquatus (Cramer, 1777)  Torquatus Swallowtail    iN

Heraclides torquatus mazai (Beutelspacher, 1977)  W Mexico to W El Salvador  M T S

Heraclides torquatus atsukoae (Okano, 1985)  southern Veracruz to E El Salvador  M S L

Heraclides torquatus tolus (Godman & Salvin, 1890)  NE Mexico to C Veracruz  M T S L1 L2

Heraclides torquatus tolmides (Godman & Salvin, 1890)  Guatemala to Panama  M T S1 S2 I1 I2

Heraclides torquatus jeani K. Brown & Lamas, 1994  TL: Colombia  M T

Heraclides torquatus leptalea (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Heraclides torquatus torquatus (Cramer, 1777)  TL: Surinam  M T

Heraclides torquatus polybius (Swainson, 1823)  TL: Brazil (MG)  M T TI

Heraclides hectorides (Esper, 1794)    iN  M T

Heraclides himeros (Hopffer, 1865)    iN

Heraclides himeros baia (Rothschild & Jordan, 1906)  TL: Brazil (Ba)  M T

Heraclides himeros himeros (Hopffer, 1865)  TL: Brazil  M T

Heraclides lamarchei (Staudinger, 1892)    iN  M T

Heraclides androgeus (Cramer, 1775)  Androgeus Swallowtail    iN

Heraclides androgeus epidaurus (Godman & Salvin, 1890)  Greater Antilles (minus Jamaica), E Mexico to W Ecuador  M T S1 S2 L I1 I2

Heraclides androgeus reyesorum Vargas, Llorente & Luis, 2013  W Mexico  M S L

Heraclides androgeus androgeus (Cramer, 1775)  TL: Surinam  M T

Heraclides androgeus iouanalao Brévignon, Deknuydt, Romé & Ravat, 2021  TL: Martinique

Heraclides androgeus laodocus (Fabricius, 1793)  TL: Brazil  M T

Heraclides thersites (Fabricius, 1775)  False Androgeus Swallowtail  Jamaica  iN  M T S L

Heraclides astyalus (Godart, 1819)    iN

Heraclides astyalus hippomedon (C. Felder & R. Felder, 1859)  Panama, Colombia, E to Guianas & S to mid-Amazon  M T S L

Heraclides astyalus phanias (Rothschild & Jordan, 1906)  TL: Ecuador  M T

Heraclides astyalus astyalus (Godart, 1819)  TL: Brazil  M T

Heraclides anchicayaensis Constantino, Le Crom & Salazar, 2002    iN  M T

Heraclides pallas (G. Gray, [1853])  Broad-banded Swallowtail  S Texas to N Costa Rica  iN

Heraclides pallas pallas (G. Gray, [1853])  S Texas, E Mexico to N Costa Rica  M T S1 S2 Ab L I

Heraclides pallas mixteca (RG. Maza & J. Maza, 2022)  TL: Mexico (Oax)

Heraclides pallas bajaensis (J. Brown & Faulkner, 1992)  W Mexico  M T S L I Ha

Heraclides ornythion (Boisduval, 1836)  Ornythion Swallowtail  S Texas to Honduras  iN

Heraclides ornythion ornythion (Boisduval, 1836)  S Texas, E Mexico to Honduras  M TI S L1 L2 I1 I2 I3 H

Heraclides ornythion W Mexican segregate ,  S Arizona, W Mexico  M S Ha

Heraclides ponceana (Schaus, 1911)  Schaus' Swallowtail  S Florida & northern Bahamas; ref.11  iN

Heraclides ponceana latefasciatus Grishin, 2020  [former temenes] Cuba; ref.11  M S L

Heraclides ponceana ponceana (Schaus, 1911)  S Florida & northern Bahamas; ref.11  M T S L I H

Heraclides ponceana majasi L. Miller, 1987  Crooked & Acklin islands, central Bahamas  M T S

Heraclides ponceana bjorndalae (Clench, 1979)  Mayaguana, Turks & Caicos & Inagua islands; ref.11  M T S

Heraclides aristodemus (Esper, 1794)  Aristodemus Swallowtail  [= temenes, Cuba?], Hispaniola, Mona, Puerto Rico; ref.11  iN  M T S L

Heraclides thoas (Linnaeus, 1771)  Thoas Swallowtail    iN

Heraclides thoas autocles (Rothschild & Jordan, 1906)  Texas to Panama  M T S1 S2 L I1 I2

Heraclides thoas nealces (Rothschild & Jordan, 1906)  Panama to Venezuela, Colombia, W Ecuador, far NW Peru  M T S L I

Heraclides thoas thoas (Linnaeus, 1771)  TL: Surinam  M OD TI HS Txt

Heraclides thoas cinyras (Ménétriés, 1857)  TL: Brazil (Ba)  M T

Heraclides thoas brasiliensis (Rothschild & Jordan, 1906)  TL: Brazil (RJ)  M T

Heraclides thoas thoantiades (Burmeister, 1878)  TL: Argentina  M T

Heraclides oviedo (Gundlach, 1866)  Cuban King Swallowtail  Cuba  iN  M T S

Heraclides homothoas (Rothschild & Jordan, 1906)  False Giant Swallowtail  Costa Rica, Panama, N Colombia, N Venezuela, Trinidad  iN  M T S I

Heraclides rumiko Shiraiwa & Grishin, 2014  Western Giant Swallowtail  SW US: CA to TX, S to Panama  iN  M T1 T2 T3 T4 T5 S1 S2 S3 S4 L1 L2 I H

Heraclides cresphontes (Cramer, 1777)  Eastern Giant Swallowtail  E US: Can. to TX, introd. to Cuba  iN  M T OD TI S1 S2 S3 S4 S5 S6 S7 S8 HS L1 L2 I Txt

Heraclides melonius (Rothschild & Jordan, 1906)  Jamaican Giant Swallowtail  Jamaica  iN  M T S L

Heraclides paeon (Boisduval, 1836)  Straight-banded Swallowtail    iN

Heraclides paeon thrason (C. Felder & R. Felder, 1865)  Yucatan Peninsula to N Venezuela and W Peru  M T S L I

Heraclides paeon paeon (Boisduval, 1836)  TL: Chile  M T S

Heraclides paeon escomeli (T. Cockerell, 1927)  TL: Peru  M T OD

Heraclides aristor (Godart, 1819)  Scarce Hispaniolan Swallowtail  Hispaniola  M S

Heraclides caiguanabus (Poey, [1852])  Poey's Swallowtail  Cuba  iN  M T S

Heraclides andraemon Hübner, [1823]  Bahamian Swallowtail    iN

Heraclides andraemon bonhotei (Sharpe, 1900)  S Florida & Bahamas  M T S

Heraclides andraemon andraemon Hübner, [1823]  Cuba, Little Cayman & Cayman Brac, Jamaica  M OI S

Heraclides andraemon tailori (Rothschild & Jordan, 1906)  Grand Cayman  M T S

Heraclides machaonides (Esper, 1796)  Machaonides Swallowtail  Hispaniola  iN  M T OI S L

Heraclides peleides (Esper, 1793)  Mysterious Swallowtail  TL: Indien?  M OI

Pterourus  Scopoli, 1777  

Pterourus esperanza (Beutelspacher, 1975)  Esperanza Swallowtail  N Oaxaca  M T S L

Pterourus palamedes (Drury, 1773)  Palamedes Swallowtail    iN

Pterourus palamedes palamedes (Drury, 1773)  SE USA  M T S HS L1 L2 I H

Pterourus palamedes leontis (Rothschild & Jordan, 1906)  NE Mexico  M T S L I Ha

Pterourus troilus (Linnaeus, 1758)  Spicebush Swallowtail    iN

Pterourus troilus troilus (Linnaeus, 1758)  E USA & SE Canada  M T S HS L1 L2 L3 L4 I H

Pterourus troilus fakahatcheensis Gatrelle, 2000  S Florida  M T S

Pterourus pilumnus (Boisduval, 1836)  Three-tailed Tiger Swallowtail  S USA (as stray), Mexico to Guatemala  iN  M T S L1 L2

Pterourus multicaudata (W. F. Kirby, 1884)  Two-tailed Tiger Swallowtail (Xochiquetzal)    iN

Pterourus multicaudata pusillus (Austin & J. Emmel, 1998)  NW USA to Baja California  M T S L I1 I2 Ha

Pterourus multicaudata multicaudata (W. F. Kirby, 1884)  SW USA to central Mexico  M T S L1 L2 I1 I2 H

Pterourus multicaudata grandiosus (Austin & J. Emmel, 1998)  central Mexico to Guatemala  M T S L I Ha

Pterourus eurymedon (Lucas, 1852)  Pale Tiger Swallowtail  W USA & Canada, Baja California Norte  iN  M T S L1 L2 L3 I H

Pterourus rutulus (Lucas, 1852)  Western Tiger Swallowtail  [= arizonensis]; W USA & Canada, Baja California N  iN  M T S L1 L2 I Ha

Pterourus canadensis (Rothschild & Jordan, 1906)  Canadian Tiger Swallowtail  Alaska to northern New England  iN  M T S L1 L2 F Ha

Pterourus appalachiensis Pavulaan & D. Wright, 2002  Appalachian Tiger Swallowtail  Appalachian Mtns., E USA  iN  M T S L1 L2

Pterourus glaucus (Linnaeus, 1758)  Eastern Tiger Swallowtail    iN

Pterourus glaucus glaucus (Linnaeus, 1758)  E USA, far NE Tamaulipas  M T1 T2 S1 S2 S3 Ab L1 L2 L3 L4 L5 L6 L7 I F

Pterourus glaucus maynardi (Gauthier, 1984)  [= australis]; C & S Florida  M S L I

Pterourus alexiares (Hopffer, 1865)  Mexican Tiger Swallowtail    iN

Pterourus alexiares garcia (Rothschild & Jordan, 1906)  NE Mexico, W Texas  M S L

Pterourus alexiares alexiares (Hopffer, 1865)  east-central Mexico  M T S

Pterourus hellanichus (Hewitson, 1868)    iN  M T

Pterourus scamander (Boisduval, 1836)    iN

Pterourus scamander joergenseni (Röber, 1925)  TL: Argentina  M T

Pterourus scamander scamander (Boisduval, 1836)  TL: Brazil  M T

Pterourus scamander grayi (Boisduval, 1836)  TL: Brazil ([RS])  M T

Pterourus homerus (Fabricius, 1793)  Homerus Swallowtail  Jamaica  iN  M S IL L

Pterourus garamas (Geyer, [1829])  Magnificent Swallowtail    iN

Pterourus garamas garamas (Geyer, [1829])  C and W Mexico  M T S Hy L I

Pterourus garamas tapatio (J. Maza & R.G. Maza, 2022)  TL: Mexico (Jal)

Pterourus garamas NW Mexican segregate ,  nr. garamas  M S L

Pterourus garamas abderus (Hopffer, 1856)  E Mexico  M T S L

Pterourus garamas baroni (Rothschild & Jordan, 1906)  SW Mexico  M T S

Pterourus garamas electryon (H. Bates, 1864)  Chiapas to El Salvador & Honduras  M T S

Pterourus garamas syedra (Godman & Salvin, 1878)  Costa Rica & W Panama  M T S L I

Pterourus garamas faisthuberi (Schäffler & Frankenbach, 2011)  TL: not specified

Pterourus garamas moehni (Schäffler & Frankenbach, 2011)  TL: not specified

Pterourus victorinus (E. Doubleday, 1844)  Victorine Swallowtail  E Mexico to Nicaragua

Pterourus victorinus victorinus (E. Doubleday, 1844)  E Mexico to Nicaragua  M T S L I Ha

Pterourus victorinus morelius (Rothschild & Jordan, 1906)  W Mexico  M T S Hy L

Pterourus victorinus NW Mexican segregate ,  NW Mexico  M S

Pterourus coroebus (C. Felder & R. Felder, 1861)    iN

Pterourus coroebus vulneratus (A. Butler, 1872)  N Costa Rica  M T S

Pterourus coroebus laetitia (A. Butler, 1872)  S Costa Rica & W Panama  M T1 T2 S1 S2 I1 I2

Pterourus coroebus syndemis Tyler, K. Brown & Wilson, 1994  TL: Colombia  M T S I

Pterourus coroebus coroebus (C. Felder & R. Felder, 1861)  TL: Venezuela  M T S

Pterourus lemoulti (Rousseau-Decelle, 1933)  TL: Venezuela  M T

Pterourus menatius (Hübner, [1819])    iN

Pterourus menatius menatius (Hübner, [1819])  TL: Surinam  M T Ma L

Pterourus menatius ctesiades (Rothschild & Jordan, 1906)  TL: Peru  M T

Pterourus menatius coelebs (Rothschild & Jordan, 1906)  TL: Peru  M T

Pterourus menatius elvirae Séraphin, 2015  TL: Colombia

Pterourus menatius levis Winhard, 2021  TL: Colombia

Pterourus menatius timana Winhard, 2021  TL: Colombia

Pterourus menatius karljordani Neild, Costa, Jost & Grishin, 2022  TL: Venezuela

Pterourus menatius eurotas (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Pterourus menatius lenaeus (E. Doubleday, 1846)  TL: Bolivia  M T

Pterourus cleotas (G. Gray, 1832)    iN  M T

Pterourus birchallii (Hewitson, 1863)  Green-washed Swallowtail    iN

Pterourus birchallii bryki (Strand, 1930)  Honduras to central Panama  M T S I1 I2

Pterourus birchallii birchallii (Hewitson, 1863)  SE Panama to C & W Colombia  M T S

Pterourus xanthopleura (Salvin & Godman, 1868)    iN

Pterourus xanthopleura ballofi Brévignon, 2006  TL: French Guiana

Pterourus xanthopleura xanthopleura (Salvin & Godman, 1868)    iN  M T

Pterourus warscewiczii (Hopffer, 1865)    iN

Pterourus warscewiczii jelskii (Oberthür, 1881)  TL: Peru  M T S

Pterourus warscewiczii mercedes (Rothschild & Jordan, 1906)  TL: Peru  M T

Pterourus warscewiczii warscewiczii (Hopffer, 1865)  TL: Bolivia  M T

Pterourus cacicus (Lucas, 1852)    iN

Pterourus cacicus cacicus (Lucas, 1852)  TL: Colombia  M T

Pterourus cacicus nesrinae (Koçak, 1983)  TL: Colombia  M T

Pterourus cacicus upanensis (Talbot, 1929)  TL: Ecuador  M T

Pterourus cacicus mendozaensis (Bollino & Sala, 1994)  TL: Peru  M T S

Pterourus cacicus inca (Rothschild & Jordan, 1906)  TL: Peru  M T

Pterourus euterpinus (Salvin & Godman, 1868)    iN  M T

Pterourus bachus (C. Felder & R. Felder, 1865)    iN

Pterourus bachus bachus (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Pterourus bachus chrysomelus (Rothschild & Jordan, 1906)  TL: Peru; Bolivia  M T S

Pterourus neyi (Niepelt, 1909)  TL: Ecuador

Pterourus neyi neyi (Niepelt, 1909)  TL: Ecuador  M T

Pterourus neyi josianeae Möhn, 2001  TL: Peru  M T

Pterourus neyi bedoci (Le Cerf, 1925)  TL: French Guiana

Pterourus zagreus (E. Doubleday, 1847)    iN

Pterourus zagreus zagreus (E. Doubleday, 1847)  TL: Venezuela  M T

Pterourus zagreus shuar L. Racheli, Bollino & T. Racheli, 2012  TL: Ecuador

Pterourus zagreus batesi T. Racheli & L. Racheli, 1996  TL: Peru

Pterourus zagreus chrysoxanthus (Fruhstorfer, 1915)  TL: Peru  M T

Pterourus zagreus nigroapicalis Bollino & Sala, 1998  TL: Peru  M T

Pterourus zagreus baueri Möhn, 2001  TL: Brazil (Am)  M T

Pterourus ascolius (C. Felder & R. Felder, 1864)  Orange Mimic-Swallowtail    iN

Pterourus ascolius zalates (Godman & Salvin, 1890)  Costa Rica to C Panama  M T S

Pterourus ascolius ascolius (C. Felder & R. Felder, 1864)  N Colombia (S. Santa Marta to Magdalena V.)  M T S L

Pterourus ascolius daguanus (Rothschild & Jordan, 1906)  W Colombia (Choco)  M T S

Pterourus ascolius rosenbergi (H. Druce, 1903)  TL: Ecuador  M T

Pterourus ascolius sarapiqui L. Racheli & T. Racheli, 2011  TL: not specified

Papilio  Linnaeus, 1758  

Papilio demoleus Linnaeus, 1758  Lime Swallowtail    iN

Papilio demoleus malayanus Wallace, 1865  Introduced to Greater Antilles, orginally Sumatra and the Malaysian peninsula  M S L

Papilio demoleus demoleus Linnaeus, 1758  Asian, shown here for comparison  M S HS

Papilio xuthus Linnaeus, 1767  Asian Swallowtail  Introduced to Hawaii  iN

Papilio xuthus xuthus Linnaeus, 1767  TL: [China]  M T S L I

Papilio indra Reakirt, 1866  Indra Swallowtail    iN

Papilio indra shastensis J. Emmel & T. Emmel, 1998  NW CA, SW OR  M T S L

Papilio indra indra Reakirt, 1866  most of range; NE ORE-WA, ID, MT, N UT, WY, W SD, CO  M T S L I Ha

Papilio indra N & C Sierra Nevada segregate ,  [ "i. indra"]; N & C Sierra Nevada, CA  M S

Papilio indra phyllisae J. Emmel, 1982  S Sierra Nevada, CA  M T S L

Papilio indra pergamus H. Edwards, 1874  Los Angeles, W San Berdo., Riverside, San Diego cos., CA  M T S L H

Papilio indra fordi J. A. Comstock & Martin, 1956  Calico S to Granite Mts.; Sheephole, Coxcomb, Eagle Mts., CA  M T S

Papilio indra parvindra J. Emmel, T. Emmel & Griffin, 2008  [= pygmaeus], Dead Mtns., San Bernardino Co., CA  M T S

Papilio indra martini J. Emmel & T. Emmel, 1966  Providence Mtns., San Bernardino Co., CA  M T S

Papilio indra panamintensis J. Emmel, 1982  Panamint Mts., Death Valley region, CA  M T S

Papilio indra nevadensis T. Emmel & J. Emmel, 1971  C & N NV  M T S L Ha

Papilio indra calcicola J. Emmel & Griffin, 1998  Muddy Mtns., Clark Co., NV, and vicinity, to E Death Valley area  M T S1 S2 I Ha

Papilio indra minori Cross, 1937  W CO, E UT, NW NM, NE AZ  M S L I Ha

Papilio indra kaibabensis Bauer, 1955  N AZ, S UT  M T S L

Papilio machaon Linnaeus, 1758  Old World Swallowtail    iN

Papilio machaon aliaska Scudder, 1869  Alaska, NW Canada  M S L I Ha

Papilio machaon pikei Sperling, 1987  Peace River region, ALTA-BC  M S

Papilio machaon hudsonianus A. Clark, 1932  NE Canada (and far NE USA)  iN  M T S

Papilio bairdii W. H. Edwards, [1866]  'Baird's' Old World Swallowtail  ref.138

Papilio bairdii dodi McDunnough, 1939  northern Great Plains, Alberta to NE Colorado  iN  M T S

Papilio bairdii oregonia W. H. Edwards, 1876  Columbia Basin, NW USA  iN  M T S Hy L

Papilio bairdii bairdii W. H. Edwards, [1866]  southern Rocky Mountains to SE California  iN  M T S1 S2 L I H

Papilio brevicauda Saunders, 1868  Short-tailed Swallowtail    iN

Papilio brevicauda brevicauda Saunders, 1868  S Laborador, Newfoundland, Anticosti Island  M S I

Papilio brevicauda bretonensis McDunnough, 1939  Maritime Provinces  M T S I

Papilio brevicauda gaspeensis McDunnough, 1934  Gaspe Peninsula, Quebec  M S

Papilio joanae J. Heitzman, 1973  Ozark Swallowtail  Ozark region, central USA  M T S

Papilio polyxenes Fabricius, 1775  Black Swallowtail    iN

Papilio polyxenes polyxenes Fabricius, 1775  Cuba  M T S

Papilio polyxenes asterius Stoll, 1782  Canada, E USA, S to El Salvador  M T S1 S2 HS L1 L2 L3 L4 L5 L6 I1 I2 I3 H F

Papilio polyxenes rudkini J. A. Comstock, 1935  [former coloro] far SW USA; ref.81  M T1 T2 S1 S2 L I Ha

Papilio polyxenes stabilis Rothschild & Jordan, 1906  Costa Rica & W Panama  M T S1 S2 L I

Papilio polyxenes americus Kollar, 1850  TL: Colombia  M T S I

Papilio polyxenes costarum Orellana, 2009  TL: Venezuela  M T S

Papilio polyxenes sadalus Lucas, 1852  TL: Ecuador  M T

Papilio polyxenes gerardi Bollino & Vitale, 2002  TL: Peru

Papilio zelicaon Lucas, 1852  Anise Swallowtail  [= coloro] W USA & Canada, Baja California Norte  iN  M T1 T2 S1 S2 Hy L1 L2 L3 I Ha

  Subfamily Praepapilioninae

Praepapilio  , []  a fossil genus

Praepapilio colorado Durden & Rose, 1978  Fossil from Colorado

Praepapilio gracilis Durden & Rose, 1978  Fossil from Colorado

Family PIERIDAE Swainson, 1820 – WHITES and YELLOWS

  Subfamily Coliadinae

  Tribe Euremini

Nathalis  Boisduval, 1836  

Nathalis iole Boisduval, 1836  Dainty Sulphur  S USA (strays N to Canada) to Honduras, Greater Antilles  iN

Nathalis iole iole Boisduval, 1836  TL: Mexico  M T S L1 L2 L3 L4 I1 I2 H F

Nathalis iole natalia Pulido, Andrade & Llorente, 2010  TL: Colombia  M S

Nathalis plauta E. Doubleday, 1847    iN  M T S

Kricogonia  Reakirt, 1863  

Kricogonia lyside (Godart, 1819)  Lyside Sulphur  SE AZ & S TX (strays N) to Venezuela; S FL, Gr. Antilles  iN  M T S1 S2 S3 L1 L2 I H

Kricogonia cabrerai Ramsden, 1920  Cuban Sulphur  Cuba  iN  M T S

Prestonia  Schaus, 1920  

Prestonia clarki Schaus, 1920  West-Mexican Sulphur  W Mexico (Sinaloa to Oaxaca); ref.12  iN  M T S L

Eurema  Hübner, [1819]  

Eurema priddyi (Lathy, 1898)  Priddy's Yellow  Hispaniola  iN  M T S

Eurema lucina (Poey, [1852])  Smudged Yellow  [= forbesi?], Cuba and Isla de Juventud  iN  M T

Eurema daira (Godart, 1819)  Barred Yellow    iN

Eurema daira palmira (Poey, [1852])  Greater & Lesser Antilles, irregularly S FL (as vagrant?)  M T1 T2 S

Eurema daira daira (Godart, 1819)  SE USA, Bahamas, Cuba as irregular vagrant  M T S1 S2 L

Eurema daira sidonia (R. Felder, 1869)  SE AZ & SW NM (as vagrant), W Mexico  M T S1 S2 L I

Eurema daira eugenia (Wallengren, 1860)  S TX (strays N), E Mexico to Panama  M T S1 S2 S3 S4 L

Eurema daira lydia (C. Felder & R. Felder, 1861)  SE Panama, NW South America  M T S

Eurema daira macheti Brévignon, 1996  TL: French Guiana

Eurema ella (Röber, 1909)  Ecuador; ref.154  M T

Eurema elathea (Cramer, 1777)  Banded Yellow    iN

Eurema elathea elathea (Cramer, 1777)  Greater & Lesser Antilles, Bahamas, Caymans  M T S1 S2 L

Eurema elathea vitellina (C. Felder & R. Felder, 1861)  Nicaragua to Colombia & Venezuela  M T S1 S2 L

Eurema elathea lamasi Brévignon, 1993  TL: French Guiana

Eurema elathea [n. ssp.#1] Lamas, MS  TL: Peru

Eurema elathea [n. ssp.#2] Lamas, MS  TL: Venezuela

Eurema flavescens (Chavannes, 1850)  S S. Am.; ref.154

Eurema flavescens flavescens (Chavannes, 1850)  TL: Brazil (SP)  M T

Eurema flavescens obsoleta (Jörgensen, 1932)  TL: Paraguay  M T L

Eurema nigrocincta Dognin, 1889  TL: Ecuador

Eurema nigrocincta nigrocincta Dognin, 1889  TL: Ecuador  M T

Eurema nigrocincta hockingi Lamas, 1981  TL: Peru  M T

Eurema millerorum Llorente & Luis, 1987  Millers' Yellow  SE Mexico to Costa Rica; ref.154  M T S

Eurema agave (Cramer, 1775)  Agave Yellow    iN

Eurema agave agave (Cramer, 1775)  Panama, South America  M T S L

Eurema agave pallida (Chavannes, 1850)  TL: Brazil (SP)  M S

Eurema phiale (Cramer, 1775)    iN

Eurema phiale columbia (C. Felder & R. Felder, 1861)  TL: Venezuela; Colombia  M T

Eurema phiale phiale (Cramer, 1775)  TL: Surinam  M T

Eurema phiale paula (Röber, 1909)  TL: Brazil (SP)  M T

Pyrisitia  A. Butler, 1870  ref.146

Pyrisitia proterpia (Fabricius, 1775)  Tailed Orange  SE AZ, SW NM, S TX (strays N) to Peru, most Greater Antilles  iN  M T S1 S2 S3 L1 L2 I H

Pyrisitia westwoodii (Boisduval, 1836)  Westwood's Yellow  SE AZ (as stray) & E Mexico to Panama; ref.12

Pyrisitia westwoodii westwoodii (Boisduval, 1836)  SE AZ (as stray) & E Mexico to Panama; ref.12  M T S1 S2 L

Pyrisitia westwoodii gabriela Le Crom & Llorente, 2004  [far SE Panama?], Colombia  M T

Pyrisitia dina (Poey, 1832)  Dina Yellow    iN

Pyrisitia dina helios (M. Bates, 1934)  S FL, Bahamas  M T S1 S2 L I H

Pyrisitia dina dina (Poey, 1832)  Cuba  M T S

Pyrisitia parvumbra (Kaye, 1925)  Dina Yellow  Jamaica  M T S L

Pyrisitia mayobanex (M. Bates, 1939)  Dina Yellow  Hispaniola; ref.146  M T S

Pyrisitia memulus (A. Butler, 1871)  Leuce Yellow  Hispaniola; ref.146  M T S1 S2 L

Pyrisitia leuce (Boisduval, 1836)    iN

Pyrisitia leuce athalia (C. Felder & R. Felder, 1865)  Panama, Colombia, Venezuela, Trinidad  M T S1 S2

Pyrisitia leuce antillarum (A. Hall, 1936)  P. Rico, St. Kitts, St. Martin, Guadeloupe, Dominica, St. Lucia  M T S1 S2 L

Pyrisitia leuce saulbatado Brévignon, 2006  TL: French Guiana

Pyrisitia leuce flavilla (H. Bates, 1861)  TL: [Brazil (Am)]  M T

Pyrisitia leuce circumcincta (H. Bates, 1861)  TL: Brazil (Pa)  M T

Pyrisitia leuce leuce (Boisduval, 1836)  TL: Brazil (RS)  M T

Pyrisitia larae (Herrich-Schäffer, 1862)  Confusing Yellow  Cuba, Andros, Great Abaco  iN  M T S

Pyrisitia venusta (Boisduval, 1836)  Pale Yellow    iN

Pyrisitia venusta emanona (Dillon, 1947)  Lesser Antilles (Grenada to Montserrat)  M T S

Pyrisitia venusta venusta (Boisduval, 1836)  Panama, northern South America  M T S L

Pyrisitia venusta aequatorialis (C. Felder & R. Felder, 1861)  TL: Ecuador  M T

Pyrisitia chamberlaini (A. Butler, 1898)  Chamberlain's Yellow  TL: Bahamas

Pyrisitia chamberlaini chamberlaini (A. Butler, 1898)  Great Bahama Bank  M T S

Pyrisitia chamberlaini clenchi (L. Miller, Simon & Harvey, 1992)  Crooked & Acklins Islands  M T

Pyrisitia chamberlaini inaguae (Munroe, 1950)  Great & Little Inagua, far E Cuba  M T S

Pyrisitia chamberlaini mariguanae (M. Bates, 1934)  Mayaguana I. & Caicos Islands  M T S

Pyrisitia nise (Cramer, 1775)  Mimosa Yellow    iN

Pyrisitia nise nelphe (R. Felder, 1869)  S AZ (as stray) & S TX (strays N) to Panama  M T S1 S2 L I1 I2 I3 Ha

Pyrisitia nise nise (Cramer, 1775)  S FL, Grand Bahama, New Providence I., Cuba, Jamaica  M T S

Pyrisitia nise stygma (Boisduval, 1836)  TL: Peru  M T

Pyrisitia nise floscula (Weeks, 1901)  TL: Bolivia  M T

Pyrisitia nise tenella (Boisduval, 1836)  TL: Brazil  M T

Pyrisitia nise [n. ssp.] Lamas, MS  Peru

Pyrisitia lisa (Boisduval & Le Conte, [1830])  Little Yellow    iN

Pyrisitia lisa lisa (Boisduval & Le Conte, [1830])  SE USA (C TX to VA, strays N), Bermuda, N Bahamas  M T S1 S2 Ma L1 L2

Pyrisitia lisa centralis (Herrich-Schäffer, 1865)  SE AZ (as stray), S TX to Panama  M S1 S2 L1 L2 I

Pyrisitia lisa euterpe (Ménétriés, 1832)  Greater and Lesser Antilles to Barbados  M T S1 S2 L

Pyrisitia euterpiformis (Munroe, 1947)  Hispaniolan Yellow  Hispaniola  iN

Pyrisitia euterpiformis euterpiformis (Munroe, 1947)  Hispaniola  iN  M T S

Pyrisitia euterpiformis turlandi W. Turner, 2016  TL: Jamaica

Pyrisitia amelia (Poey, [1852])  Cuban Yellow  Cuba & Isla de Juventud  M T S

Pyrisitia portoricensis (Dewitz, 1877)  Puerto Rican Yellow  Puerto Rico  iN  M T S L

Pyrisitia pyro (Godart, 1819)  Pyro Orange  Hispaniola  iN  M T S1 S2

Pyrisitia messalina (Fabricius, 1787)  Whitish Yellow  Bahamas, Cuba, I. de Juventud, Jamaica, Caymans, [S FL?]  iN  M T S1 S2

Pyrisitia lirina (H. Bates, 1861)    iN  M T

Abaeis  Hübner, [1819]  ref.146

Abaeis brephos (Hübner, [1809])    iN  M T

Abaeis elvina (Godart, 1819)    iN  M T

Abaeis albula (Cramer, 1775)  White Yellow  ref.10  iN

Abaeis albula celata (R. Felder, 1869)  E & W Mexico to Honduras  M T S L

Abaeis albula ceciliana (F.G. Haghenbeck, J.J. García, E.L. Haghenbeck & Turrent, 2021)  TL: Mexico

Abaeis albula marginella (C. Felder & R. Felder, 1861)  Nicaragua to Ecuador  M T S1 S2 L I

Abaeis albula albula (Cramer, 1775)  NE South America to Argentina, Trinidad, St. Vincent  M T S

Abaeis albula totora (Lamas, 1981)  TL: Peru  M T

Abaeis albula espinosae (A. Fernández, 1928)  TL: Peru  M T

Abaeis albula sinoe (Godart, 1819)  TL: Brazil  M T

Abaeis paulina (H. Bates, 1861)  ref.10  iN  M T

Abaeis xantochlora (Kollar, 1850)  Tropical Yellow  ref.10  iN

Abaeis xantochlora xantochlora (Kollar, 1850)  SE Mexico to Colombia & Venezuela  M T S1 S2 L I

Abaeis xantochlora magna (Winhard & Le Crom, 2004)  TL: Colombia

Abaeis xantochlora edentata (Le Crom & Constantino, 2004)  TL: Colombia

Abaeis xantochlora ectriva (A. Butler, 1873)  TL: Ecuador  M T

Abaeis xantochlora pomponia (Hopffer, 1874)  TL: Peru  M T S

Abaeis fabiola (C. Felder & R. Felder, 1861)  ref.10  M T

Abaeis tupuntenem (Lichy, 1976)  ref.10  iN

Abaeis tupuntenem tupuntenem (Lichy, 1976)  TL: Venezuela  M T

Abaeis tupuntenem [n. ssp.] (Lamas, MS)  Venezuela

Abaeis salome (C. Felder & R. Felder, 1861)  ref.10  iN

Abaeis salome jamapa (Reakirt, 1866)  S NM & S TX (as strays) to Panama  M S1 S2 L

Abaeis salome gaugamela (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Abaeis salome santamarta (Le Crom, Winhard & Salazar, 2004)  TL: Colombia

Abaeis salome neglecta (Le Crom & G. Rodríguez, 2004)  TL: Colombia  M T

Abaeis salome limoneus (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Abaeis salome xystra (R.F. d'Almeida, 1936)  TL: Ecuador  M T

Abaeis salome salome (C. Felder & R. Felder, 1861)  TL: Ecuador  M T

Abaeis salome [n. ssp.] (Lamas, MS)  Peru

Abaeis mexicana (Boisduval, 1836)  Mexican Yellow  ref.10  iN

Abaeis mexicana mexicana (Boisduval, 1836)  SW USA (strays widely N) to Panama  M T S L1 L2 I H

Abaeis mexicana bogotana (C. Felder & R. Felder, 1861)  Costa Rica, Panama, Colombia  M T S

Abaeis mexicana citrella (Winhard, Le Crom & Constantino, 2004)  TL: Colombia  M T

Abaeis mexicana tayrona (Winhard, Le Crom & Salazar, 2004)  TL: Colombia

Abaeis boisduvaliana (C. Felder & R. Felder, 1865)  Boisduval’s Yellow  SE AZ, SW NM, S TX to W CR, Cuba, Jamaica; ref.10  iN  M T S1 S2 L1 L2 I H

Abaeis angulata (Wallengren, 1860)  ref.10,Zhang23d  M T

Abaeis gratiosa (E. Doubleday, 1847)  Disjunct Yellow  W Costa Rica to Venezuela, St. Lucia & the Grenadines; ref.10,Zhang23d

Abaeis gratiosa gratiosa (E. Doubleday, 1847)  W Costa Rica to Venezuela, St. Lucia & the Grenadines; ref.10  M T S1 S2

Abaeis gratiosa ruizana (Lamas, 1981)  TL: Peru  M T

Abaeis arbela (Geyer, 1832)  ref.10  iN

Abaeis arbela deflorata (Kollar, 1850)  TL: Colombia  M T

Abaeis arbela gracilis (Avinoff, 1926)  TL: Brazil (Ba)  M T S

Abaeis arbela graduata (A. Butler, 1873)  TL: Brazil (Pa)  M T

Abaeis arbela arbela (Geyer, 1832)  TL: Java  M T S

Abaeis adamsi (Lathy, 1898)  Contrasting Yellow  Jamaica; ref.10  M T S

Abaeis nicippe (Cramer, 1779)  Sleepy Orange  S USA (strays N) to Costa Rica, Gr. Antilles, Caymans, Bahamas  iN  M T S1 S2 Ab L1 L2 L3 L4 I H

Abaeis nicippiformis (Munroe, 1947)  Fast Orange  Hispaniola  iN  M T S

Teriocolias  Röber, 1909  ref.146

Teriocolias deva (E. Doubleday, 1847)  ref.12  iN

Teriocolias deva deva (E. Doubleday, 1847)  TL: French Guiana  M T

Teriocolias deva chilensis (Blanchard, 1852)  TL: Chile  M T

Teriocolias doris (Röber, 1909)  TL: Bolivia  M T

Teriocolias zelia (Lucas, 1852)    iN

Teriocolias zelia zelia (Lucas, 1852)  TL: [Bolivia]  M T

Teriocolias zelia andina W. Forbes, 1928  TL: Peru  M T

Teriocolias zelia mathani (W. D. Field, 1950)  TL: Peru  M T

Teriocolias zelia pacis Röber, 1909  TL: Peru  M T

Teriocolias reticulata (A. Butler, 1871)  Ecuador; ref.12  iN  M T S

  Tribe Coliadini

Colias  Fabricius, 1807  

Colias vitabunda Hovanitz, 1943  Lively Clouded Sulphur  AK & NW Can  iN  M S L I

Colias philodice Godart, 1819  Clouded Sulphur    iN

Colias philodice philodice Godart, 1819    iN  M T S1 S2 S3 Ab L

Colias philodice guatemalena Röber, 1909  Chiapas & Guatemala  iN  M T S

Colias eriphyle W. H. Edwards, 1876  Clouded Sulphur  SW Can & W US  iN  M T S1 S2 L I Ha

Colias eurytheme Boisduval, 1852  Orange Sulphur    iN  M T S1 S2 S3 Ab L1 L2 L3 L4 L5 I Ha

Colias occidentalis Scudder, 1862  Western Sulphur    iN

Colias occidentalis chrysomelas H. Edwards, 1877  TL: USA (Ca)  M S

Colias occidentalis primordialis Hammond & McCorkle, 2017  TL: USA, OR

Colias occidentalis SW Oregon segregate ,  S Cascades, far NE Siskiyous, Jackson -Josephine Co., OR  M S L Ha

Colias occidentalis occidentalis Scudder, 1862  [= nortepacifica]  M S L Ha

Colias occidentalis sullivani Hammond & McCorkle, 2003  TL: USA (Or)  M T1 T2 T3 T4 T5 T6 T7 T8 S

Colias occidentalis resplendens Hammond & McCorkle, 2017  TL: USA, ID

Colias occidentalis NE Oregon mountains segregate ,   M S L Ha

Colias occidentalis lower Snake River drainage segregate ,   M S1 S2 S3 S4 S5 S6 S7

Colias occidentalis pseudochristina Ferris, 1989  TL: USA (Ut)  M T S L I

Colias christina W. H. Edwards, 1863  Christina's Sulphur    iN

Colias christina kluanensis Ferris, 1981  TL: Canada (Yt)  M T S L I

Colias christina christina W. H. Edwards, 1863  TL: Canada (NT)  M S1 S2 S3

Colias christina sacajawea Kohler, 2006  TL: USA (Mt)  M S

Colias christina pseudocolumbiensis C. Guppy & J. Shepard, 2001  TL: Canada (BC)  M S1 S2 S3

Colias christina columbiensis Ferris, 1973  TL: Canada (BC)  M S

Colias christina krauthii Klots, 1935  TL: USA (SD)  M S L I Ha

Colias alexandra W. H. Edwards, 1863  Queen Alexandra's Sulphur    iN

Colias alexandra altamont Kondla & Kohler, 2006  TL: USA (Mt)  M S

Colias alexandra astraea W. H. Edwards, 1872  TL: USA (Wy)  M S Ab L Ha

Colias alexandra alexandra W. H. Edwards, 1863  TL: USA (Co)  M S Ab L I Ha

Colias alexandra altiplano M. Fisher & Scott, 2006  TL: USA (Co)  M S

Colias alexandra apache Ferris, 1988  TL: USA (Az)  M S L

Colias alexandra edwardsii W. H. Edwards, 1870  TL: USA (Nv)  M S1 S2 S3 L I1 I2 Ha

Colias harfordii H. Edwards, 1877  Harford’s Sulphur    iN  M T S1 S2 L I1 I2 Ha

Colias meadii W. H. Edwards, 1871  Mead's Sulphur  ref.11  iN

Colias meadii meadii W. H. Edwards, 1871  TL: USA (Co)  M S L1 L2 L3 I Ha

Colias meadii lemhiensis N. Curtis & Ferris, 1985  TL: USA (Id)  M T

Colias elis Strecker, 1885  Mead's Sulphur  ref.11  iN  M S I

Colias johanseni Troubridge & Philip, 1990  Johansen's Sulphur  Bernard Harbour, District of Mackenzie, Nunavut  M T1 T2

Colias rankinensis Verhulst, 2009  Rankin Inlet Sulphur  Canada: Nunavut & Manitoba  M OI

Colias hecla Lefèbvre, 1836  Hecla Sulphur    iN

Colias hecla hecla Lefèbvre, 1836  TL: Greenland  M S L Ha

Colias hecla hela Strecker, 1880  TL: Canada (Nu)  M S1 S2

Colias hecla zamolodchikovi Churkin & Grieshuber, 2001  Seward Peninsula, AK, possibly E through Yukon & NWT  M S

Colias boothii J. Curtis, 1835  Booth's Sulphur  C Canadian Arctic

Colias boothii canadensis Ferris, 1982    iN  M S Ha

Colias boothii kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001  [olga is preoccupied] Seward Peninsula, AK  M T

Colias boothii thula Hovanitz, 1955  N slope Brooks Range, AK  M S1 S2 Ha

Colias boothii boothii J. Curtis, 1835  C Canadian Arctic  M S1 S2 Ab

Colias nastes Boisduval, 1832  Labrador Sulphur    iN

Colias nastes nastes Boisduval, 1832  Labrador  M S

Colias nastes moina Strecker, 1880  C Canadian Arctic  M S1 S2

Colias nastes aliaska Bang-Haas, 1927  most of Alaska (except Seward Peninsula) and Yukon; blending with moina to E  M S1 S2 S3 L I Ha

Colias nastes dezhnevi Korshunov, 1995  Seward Peninsula, AK  M S

Colias nastes streckeri Grum-Grshimailo, 1895  N WA, N MT, ALTA, N BC  M S1 S2

Colias scudderii Reakirt, 1865  Scudder's Sulphur    iN  M T S1 S2 Ab L Ha

Colias gigantea Strecker, 1900  Giant Sulphur    iN

Colias gigantea inupiat Harry, 2007  Alaskan North Slope, Dalton Highway  M T1 T2 S L I

Colias gigantea gigantea Strecker, 1900  TL: Csnada (Mt)  M S L Ha

Colias gigantea mayi F. Chermock & R. Chermock, 1940  TL: Csnada (Mt)  M T S

Colias gigantea harroweri Klots, 1940  [= kohleri; = gracemma]  M S Ab L

Colias pelidne Boisduval & Le Conte, [1830]  Pelidne Sulphur    iN

Colias pelidne pelidne Boisduval & Le Conte, [1830]    iN  M S L

Colias pelidne auroraborealis Hammond & McCorkle, 2017  TL: Canada

Colias pelidne philipi Hammond & McCorkle, 2017  TL: USA, AK

Colias pelidne pelhami Hammond & McCorkle, 2017  TL: Canada

Colias skinneri W. Barnes, 1897  Pelidne Sulphur    iN

Colias skinneri skinneri W. Barnes, 1897    iN  M S1 S2 L Ha

Colias skinneri hinchliffi Hammond & McCorkle, 2017  TL: USA, OR

Colias skinneri curtisi Hammond & McCorkle, 2017  TL: USA, ID

Colias interior Scudder, 1862  Pink-edged Sulphur    iN

Colias interior interior Scudder, 1862    iN  M T S1 S2 S3 L

Colias interior carolae Hammond & McCorkle, 2017  TL: USA, WV

Colias interior sigridae Hammond & McCorkle, 2017  TL: Canada

Colias interior dornfeldi Hammond & McCorkle, 2017  TL: USA, OR

Colias palaeno (Linnaeus, 1761)  Palaeno Sulphur    iN

Colias palaeno chippewa W. H. Edwards, 1870  TL: Canada (NT)  M S L I Ha

Colias palaeno baffinensis Ebner & Ferris, [1978]  TL: Canada (Nu)  M S

Colias behrii W. H. Edwards, 1866  Sierra Sulphur    iN  M T S L1 L2 Ha

Colias ponteni Wallengren, 1860  TL: USA (Hi)  M T

Colias vauthierii Guérin-Méneville, [1830]    iN  M T

Colias lesbia (Fabricius, 1775)    iN

Colias lesbia lesbia (Fabricius, 1775)  TL: Argentina  M T Ab

Colias lesbia mineira J. Zikán, 1940  TL: Brazil (MG)  M T

Colias lesbia andina Staudinger, 1894  TL: Bolivia  M T

Colias lesbia verhulsti Berger, 1983  TL: Peru

Colias lesbia dinora W. F. Kirby, 1881  TL: Ecuador  M T

Colias misti Kir'yanov, 2017  TL: to add later

Colias misti misti Kir'yanov, 2017  TL: to add later

Colias misti ccota Kir'yanov, 2019  TL: to add later

Colias euxanthe C. Felder & R. Felder, 1865    iN

Colias euxanthe alticola Godman & Salvin, 1891  TL: Ecuador  M T

Colias euxanthe euxanthe C. Felder & R. Felder, 1865  TL: Peru  M T

Colias euxanthe hermina (A. Butler, 1871)  TL: Peru  M T

Colias euxanthe stuebeli Reissinger, 1972  TL: Peru  M T

Colias flaveola Blanchard, 1852    iN

Colias flaveola mossi Rothschild, 1913  TL: Peru  M T

Colias flaveola erika Lamas, 1981  TL: Peru  M T

Colias flaveola weberbaueri Strand, 1912  TL: Peru  M T

Colias flaveola benyaminii Verhulst, 2016  TL: to add later

Colias flaveola cora Kir'yanov, 2017  TL: to add later

Colias flaveola laura Kir'yanov, 2017  TL: to add later

Colias flaveola harai Kir'yanov & Valencia, 2018  TL: to add later

Colias flaveola viuda Kir'yanov, 2020  TL: to add later

Colias flaveola blameyi Jörgensen, 1916  TL: Argentina  M T

Colias flaveola mendozina Breyer, 1939  TL: Argentina  M T

Colias flaveola flaveola Blanchard, 1852  TL: Chile

Colias dimera E. Doubleday, 1847    iN  M T

Zerene  Hübner, [1819]  

Zerene cesonia (Stoll, 1790)  Southern Dogface    iN

Zerene cesonia cynops (A. Butler, 1873)  Hispaniola  M T S1 S2

Zerene cesonia cesonia (Stoll, 1790)  S USA (strays far N) to Colombia, Cuba  M T S1 S2 S3 HS L1 L2 L3 I1 I2 H

Zerene cesonia patiensis Constantino, G. Rodríguez & Salazar, 2004  TL: Colombia  M T

Zerene cesonia boyacensis Le Crom, Winhard & Torres, 2004  TL: Colombia

Zerene cesonia inca (Tessmann, 1928)  TL: Peru  M T S

Zerene cesonia limonella Lamas, 1981  TL: Peru  M T S

Zerene cesonia therapis (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Zerene cesonia cesonides (Staudinger, 1894)  TL: Bolivia  M T S

Zerene eurydice (Boisduval, 1855)  California Dogface  Baja California (N) to N California  iN  M T1 T2 S L1 L2 I Ha

Anteos  Hübner, [1819]  

Anteos maerula (Fabricius, 1775)  Yellow Angled-Sulphur  SW USA (strays), S TX to Peru, S FL, Bahamas, Gr. Antilles  iN  M T S1 S2 L I Ha

Anteos clorinde (Godart, [1824])  White Angled-Sulphur  SW USA (strays), S TX to Argentina, Cuba, Jamaica, Hispaniola  iN  M T S1 S2 L1 L2 I

Anteos menippe (Hübner, [1818])  Orangetip Angled-Sulphur    iN  M T S

Phoebis  Hübner, [1819]  

Phoebis sennae (Linnaeus, 1758)  Cloudless Sulphur    iN

Phoebis sennae sennae (Linnaeus, 1758)  Caribbean Islands, blends with eubule S FL  M T S L I

Phoebis sennae eubule (Linnaeus, 1767)  SE USA (strays far N, NW)  M T S HS L1 L2 L3 L4 I G

Phoebis sennae marcellina (Cramer, 1777)  SW USA (strays far N), S TX through Panama, most S. America  M T S1 S2 HS L1 L2 I H

Phoebis sennae amphitrite (Feisthamel, 1839)  TL: Chile  M T

Phoebis bourkei (Dixey, 1933)  TL: Ecuador  M T

Phoebis editha (A. Butler, 1870)  Edith's Sulphur  Hispaniola  iN  M T S

Phoebis philea (Linnaeus, 1763)  Orange-barred Sulphur    iN

Phoebis philea philea (Linnaeus, 1763)  SW USA (strays far N) to S Brazil, Gulf Coast to FL, P. Rico, etc.  M T TI S1 S2 S3 Ab HS L1 L2 I Ha G

Phoebis philea thalestris (Illiger, 1801)  Hispaniola  M S G

Phoebis philea huebneri Fruhstorfer, 1907  Cuba [P. p. philea also in W Cuba, with intergrades]  M S

Phoebis avellaneda (Herrich-Schäffer, 1865)  Orange-washed Sulphur  Cuba  iN  M T S1 S2 G

Phoebis neocypris (Hübner, [1823])    iN

Phoebis neocypris virgo (A. Butler, 1870)  SE AZ & S TX (rare strays) to Brazil; ref.271  M T S1 S2 L I G

Phoebis neocypris rurina (C. Felder & R. Felder, 1861)  TL: Venezuela; Colombia  M T

Phoebis neocypris neocypris (Hübner, [1823])    iN  M T S

Phoebis neocypris vocator Brévignon, 2006  TL: French Guiana

Phoebis argante (Fabricius, 1775)  Dark Apricot Sulphur  S Mexico to S Brazil, Trinidad  iN

Phoebis argante argante (Fabricius, 1775)  TL: Brazil  M T S1 S2 S3 S4 S5 S6 S7 S8 S9 S10 S11 S12 S13 S14 S15 S16 S17 L I1 I2 G

Phoebis argante larra (Fabricius, 1798)  TL: America  M S

Phoebis argante chincha Lamas, 1976  TL: Peru  M T

Phoebis argante minuscula (A. Butler, 1869)  [= fornax], Cuba  M T S

Phoebis argante comstocki Avinoff, 1944  Jamaica  M T S L

Phoebis argante rorata (A. Butler, 1869)  Hispaniola  M T S1 S2

Phoebis argante martini W. Comstock, 1944  Puerto Rico  M T S

Phoebis agarithe (Boisduval, 1836)  Large Orange Sulphur    iN

Phoebis agarithe fisheri (H. Edwards, 1883)  Baja California  M S L

Phoebis agarithe agarithe (Boisduval, 1836)  S W USA (strays far N), S TX to Peru; Galapagos Is., Hawaii  M T S1 S2 S3 L1 L2 L3 I1 I2 I3 H

Phoebis agarithe maxima (Neumoegen, 1891)  Florida  M T S L

Phoebis agarithe antillia F. Brown, 1929  Bahamas, Greater & Lesser Antilles  M T S L G

Phoebis agarithe pupillata Dillon, 1947  Windward Islands, at least Guadeloupe to at least St. Lucia and Barbados  M T S

Phoebis agarithe tumbesina Lamas, 1981  TL: Peru  M T

Phoebis trite (Linnaeus, 1758)  ref.271  iN

Phoebis trite northern segregate ,  E & W Mexico to Panama  M T

Phoebis trite watsoni F. Brown, 1929  Hispaniola, P. Rico, St. Kitts, Montserrat, Guad. to St. Vincent  M T S

Phoebis trite trite (Linnaeus, 1758)  TL: Calidioribus regionibus  M T S

Phoebis trite banksi Breyer, 1939  TL: Brazil (SC)  M T S

Aphrissa  A. Butler, 1873  

Aphrissa boisduvalii (C. Felder & R. Felder, 1861)  Boisduval's Sulphur  E & S Mexico to W Peru  iN  M T1 T2 S1 S2

Aphrissa schausi (Avinoff, 1926)  Schaus' Sulphur  [taxonomic status uncertain] S Mexico and Guatemala  M T S1 S2

Aphrissa wallacei (C. Felder & R. Felder, 1862)  Wallace's Sulphur  [= fluminensis] Costa Rica to Argentina  M T1 T2 S1 S2 S3

Aphrissa statira (Cramer, 1777)  Statira Sulphur    iN

Aphrissa statira statira (Cramer, 1777)  S TX (strays N), E & W Mexico to Argentina, Lesser Antilles  M T1 T2 OI TI S1 S2 S3 S4 IL L I G

Aphrissa statira floridensis (Neumoegen, 1891)  Florida  M T S1 S2 L H

Aphrissa statira cubana R.F. d'Almeida, 1939  Cuba, Cayman Islands, Jamaica  M T S

Aphrissa statira hispaniolae (Munroe, 1947)  Hispaniola  M T S

Aphrissa neleis (Boisduval, 1836)  Pink-spot Sulphur  Cuba, Isla de Juventud, New Providence, N & S Andros, S FL  iN  M T S1 S2 IL L1 L2 G

Aphrissa godartiana (Swainson, 1821)  Godart's Sulphur  TL: ?

Aphrissa godartiana godartiana (Swainson, 1821)  Hispaniola  M T OI S1 S2 IL G

Aphrissa godartiana hartonia (A. Butler, 1870)  Jamaica  M T S IL G

Aphrissa orbis (Poey, 1832)  Orbis Sulphur  TL: Cuba

Aphrissa orbis orbis (Poey, 1832)  S FL (rare stray), Cuba, Isla de Juventud, Cayman Brac  M T S IL G

Aphrissa orbis browni (Munroe, 1947)  Hispaniola  M T S IL

  Subfamily Pierinae

  Tribe Anthocharidini

Anthocharis  Boisduval, Rambur & Graslin, [1833]  

Anthocharis midea (Hübner, [1809])  Falcate Orangetip    iN

Anthocharis midea midea (Hübner, [1809])  TL: USA (Ga)  M T S1 S2

Anthocharis midea annickae dos Passos & Klots, 1969  TL: USA (Ct)  M S1 S2 L1 L2 L3 L4 I H

Anthocharis midea texana Gatrelle, 1998  [far NE TAMPS]  M T1 T2 S L1 L2 L3 L4 L5 I1 I2 H F

Anthocharis limonea (A. Butler, 1871)  Mexican Orangetip  E & C Mexico  iN  M T S L I Ha

Anthocharis cethura C. Felder & R. Felder, 1865  Desert Orangetip    iN

Anthocharis cethura morrisoni W. H. Edwards, 1881  TL: USA (Ca)  M S L I H Ha

Anthocharis cethura bajacalifornica J. Emmel, T. Emmel & Mattoon, 1998  TL: Mexico (BCN)  M T S I H Ha

Anthocharis cethura catalina Meadows, 1937  TL: USA (Ca)  M T

Anthocharis cethura cethura C. Felder & R. Felder, 1865  TL: [USA (Ca)]  M T S1 S2 S3 L

Anthocharis cethura hadromarmorata J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Nv)  M T1 T2 S L

Anthocharis cethura mojavensis J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S1 S2 L I1 I2 I3 H Ha

Anthocharis cethura pima W. H. Edwards, 1888  TL: USA (Az)  M T S L I F

Anthocharis sara Lucas, 1852  Sara Orangetip    iN

Anthocharis sara Isla de Cedros segregate ,  Isla de Cedros, Baja California, Mexico  M S1 S2 I Ha

Anthocharis sara gunderi Ingham, 1933  TL: USA (Ca)  M T S L Ha

Anthocharis sara sara Lucas, 1852  TL: USA (Ca)  M T1 T2 S1 S2 Ab Hy L1 L2 L3 L4 I H Ha

Anthocharis sara sempervirens J. Emmel, T. Emmel & Mattoon, 2008  TL: USA (Ca)  M T

Anthocharis sara pseudothoosa Austin, 1998  TL: USA (Nv)  M T S L Ha

Anthocharis julia W. H. Edwards, 1872  Sara Orangetip    iN

Anthocharis julia alaskensis Gunder, 1932  TL: USA (Ak)  M S

Anthocharis julia flora W. G. Wright, 1892  TL: USA (Wa)  M T S1 S2

Anthocharis julia stella W. H. Edwards, 1879  TL: USA (Nv)  M T S1 S2 L

Anthocharis julia sulfuris Pelham, 2008  [= columbia]  M S1 S2 L I Ha

Anthocharis julia browningi Skinner, 1906  TL: USA (Ut)  M T S L I

Anthocharis julia julia W. H. Edwards, 1872  TL: USA (Co)  M S1 S2 L Ha

Anthocharis julia prestonorum Stout, 2012  W & SW CO, W of the divide  M T S

Anthocharis thoosa (Scudder, 1878)  Sara Orangetip    iN

Anthocharis thoosa colorado Scott & M. Fisher, 2008  SW CO, SE UT, NW NM, [NE AZ]  M T S

Anthocharis thoosa coriande M. Fisher & Scott, 2008  N NM, far S-central CO  M T S1 S2

Anthocharis thoosa inghami Gunder, 1932  TL: USA (Az)  M T S Ab L I H

Anthocharis thoosa thoosa (Scudder, 1878)  TL: USA (Az)  M T S L I Ha

Anthocharis lanceolata Lucas, 1852  Gray Marble  SW OR, N CA, far W NV, S through Sierra Nevada  iN

Anthocharis lanceolata lanceolata Lucas, 1852  SW OR, N CA, far W NV, S through Sierra Nevada  M T S L I Ha

Anthocharis lanceolata australis (F. Grinnell, 1908)  S CA Ranges  M T S L I Ha

Anthocharis desertolimbus J. Emmel, T. Emmel & Mattoon, 1998  Gray Marble  W edge Colorado Desert (Palm Spgs. area S to N BC)  M T S Hy L I H Ha

Euchloe  Hübner, [1819]  

Euchloe ochracea (Trybom, 1877)  Arctic Marble  TL: not specified

Euchloe ochracea kamchatkensis Back, 2008  TL: Russia  M S1 S2 L Ha

Euchloe ausonides (Lucas, 1852)  Large Marble    iN

Euchloe ausonides ogilvia Back, 1991  TL: Canada (YT)  M S L

Euchloe ausonides insulanus C. Guppy & J. Shepard, 2001  Puget Sound, WA, [BC]  M T L

Euchloe ausonides transmontana Austin & J. Emmel, 1998  CA Sierra Nevada to W UT, N CA, OR, E WA, S BC, W ID  M T S L I Ha

Euchloe ausonides ausonides (Lucas, 1852)  W CA (esp. Bay area), low elevations  M T S1 S2 L

Euchloe ausonides palaeoreios K. Johnson, 1976  TL: USA (SD)  M T Ha

Euchloe ausonides coloradensis (H. Edwards, 1881)  TL: USA (Co)  M S1 S2 L1 L2 L3 I H Ha

Euchloe ausonides mayi F. Chermock & R. Chermock, 1940  TL: Canada (Mt)  M T S

Euchloe creusa (E. Doubleday, [1847])  Northern Marble    iN

Euchloe creusa creusa (E. Doubleday, [1847])  N MT (vic. Glacier NP), E BC, W ALTA, N to Arctic  M S L Ha

Euchloe hyantis (W. H. Edwards, 1871)  California Marble    iN

Euchloe hyantis andrewsi Martin, 1958  San Bernardino Mts., CA  M T1 T2 S

Euchloe hyantis hyantis (W. H. Edwards, 1871)  SW OR (NW Siskiyous), W CA to Baja California (N)  M S1 S2 L Ha

Euchloe hyantis Illinois Valley segregate ,  Illinois Valley, Josephine Co., OR  M S

Euchloe lotta Beutenmüller, 1898  Desert Marble  SE BC, S to E CA, AZ, N SON-CHIH, far W TX, W NM, N to W MT  iN

Euchloe lotta lotta Beutenmüller, 1898  SE BC, S to E CA, AZ, N SON-CHIH, far W TX, W NM, N to W MT  M T S1 S2 L1 L2 I Ha

Euchloe lotta belioides Verity, [1909]  TL: USA (Az)  M T

Euchloe guaymasensis Opler, 1987  Sonoran Marble  N & C Sonora, rarely in SE AZ (Cochise Co.)  M T S L I

Euchloe olympia (W. H. Edwards, 1871)  Olympia Marble  S ALTA to NE NM & C TX, prairies E to S ONT + Appalachians  iN

Euchloe olympia olympia (W. H. Edwards, 1871)  TL: USA (WV)  M S1 S2 Ab L Ha

Euchloe olympia rosa (W. H. Edwards, 1882)  TL: USA (Tx)

Euchloe olympia wagneri Pavulaan, 2020  dunes bordering Great Lakes (WI, IL, IN, MI)  M S L

Euchloe olympia huron Pavulaan, 2020  TL: Canada (On)

Eroessa  E. Doubleday, 1847  

Eroessa chiliensis (Guérin-Méneville, [1830])    iN  M S

Hesperocharis  C. Felder, 1862  

Hesperocharis hirlanda (Stoll, 1790)  ref.12  iN

Hesperocharis hirlanda tirimbina (Chacón, DeVries & Penz, 2018)  TL: Costa Rica

Hesperocharis hirlanda hirlanda (Stoll, 1790)  TL: Bengale  M T

Hesperocharis hirlanda apicalis Fruhstorfer, 1907  TL: Ecuador  M T

Hesperocharis hirlanda fulvinota A. Butler, 1871  TL: Brazil (RJ)  M T

Hesperocharis hirlanda helvia (Latreille, [1813])  TL: ["Mexico"]  M T

Hesperocharis hirlanda minturna (Fruhstorfer, 1910)  TL: Venezuela  M T

Hesperocharis hirlanda ninguida Fruhstorfer, 1907  TL: Peru  M T

Hesperocharis hirlanda planasia (Fruhstorfer, 1910)  TL: Brazil (MT)  M T

Hesperocharis hirlanda praeclara Fruhstorfer, 1907  TL: Brazil (ES)  M T

Hesperocharis hirlanda serda Fruhstorfer, 1907  TL: Colombia  M T

Hesperocharis anguitia (Godart, 1819)    iN

Hesperocharis anguitia anguitia (Godart, 1819)  TL: Brazil  M T

Hesperocharis anguitia giesekingi Breyer, 1960  TL: Argentina  M T

Hesperocharis costaricensis H. Bates, 1866  Costa Rican White    iN

Hesperocharis costaricensis costaricensis H. Bates, 1866  Nicaragua to Panama  M T S

Hesperocharis costaricensis pasion (Reakirt, [1867])  E & W Mexico to Guatemala (at least)  M T S1 S2 L

Hesperocharis crocea H. Bates, 1866  Orange White    iN

Hesperocharis crocea crocea H. Bates, 1866  E Mexico to Panama  M T S

Hesperocharis crocea idiotica (A. Butler, 1869)  TL: ?  M T

Hesperocharis crocea jaliscana Schaus, 1898  W Mexico (Nayarit to Guerrero)  M T S L

Hesperocharis crocea [n. ssp.] Lamas, MS  Peru

Hesperocharis emeris (Boisduval, 1836)  TL: Brazil  M T

Hesperocharis erota (Lucas, 1852)    iN  M T

Hesperocharis graphites H. Bates, 1864  Marbled White    iN

Hesperocharis graphites graphites H. Bates, 1864  Chiapas to Panama  M T S1 S2 L

Hesperocharis graphites avivolans (A. Butler, 1865)  E, W & S Mexico (to Oaxaca)  M T S L1 L2

Hesperocharis leucania (Boisduval, 1836)    iN  M T

Hesperocharis marchalii (Guérin-Méneville, [1844])    iN  M T S L

Hesperocharis nera (Hewitson, 1852)    iN

Hesperocharis nera nera (Hewitson, 1852)  TL: Ecuador  M T

Hesperocharis nera aida Fruhstorfer, 1908  TL: Peru  M T

Hesperocharis nera amazonica Fruhstorfer, 1907  TL: Peru  M T

Hesperocharis nera lamonti Kaye, 1920  TL: Trinidad  M T

Hesperocharis nera nereis C. Felder & R. Felder, 1865  TL: Colombia  M T

Hesperocharis nera nerida J. Zikán, 1940  TL: Brazil (Am)  M T

Hesperocharis nera nymphaea Möschler, 1876  TL: Surinam  M T

Hesperocharis nereina Hopffer, 1874    iN  M T L

Hesperocharis paranensis Schaus, 1898    iN

Hesperocharis paranensis paranensis Schaus, 1898  TL: Brazil (Pr)  M T

Hesperocharis paranensis infrasignata Breyer, 1939  Argentina  iN  M T

Hesperocharis nambii Lamas, Constantino & Solarte, 2007  Colombia; ref.292  M T

Hesperocharis leucothea (J. Molina, 1782)  Chile; ref.12  iN  M T

Mathania  Oberthür, 1890  

Mathania agasicles (Hewitson, 1874)    iN

Mathania agasicles agasicles (Hewitson, 1874)  TL: Bolivia  M T

Mathania agasicles gaujoni (Dognin, 1887)  TL: Ecuador  M T

Mathania agasicles [n. ssp.#1] Lamas, MS  TL: Peru

Mathania agasicles [n. ssp.#2] Lamas, MS  TL: Peru

Mathania aureomaculata (Dognin, 1888)  TL: Ecuador

Mathania aureomaculata aureomaculata (Dognin, 1888)  TL: Ecuador  M T

Mathania aureomaculata esther Oberthür, 1890  TL: Peru  M T

Mathania aureomaculata pallida Röber, 1909  TL: Peru  M T

Mathania aureomaculata [n. ssp.] Lamas, MS  Peru

Mathania carrizoi Giacomelli, 1914    iN  M T

Mathania [n. sp.] Lamas, MS  Peru

  Tribe Pierini

Ganyra  Billberg, 1820  

Ganyra howarthi (Dixey, 1915)  Howarth's White  S AZ (as stray), NW Mexico (SIN, SON, BC, BCS)  iN  M T S1 S2 L

Ganyra josephina (Godart, 1819)  Giant White    iN

Ganyra josephina josepha (Salvin & Godman, 1868)  S TX (strays N), E & W Mexico to Panama  M T S1 S2 L1 L2 I

Ganyra josephina janeta (Dixey, 1915)  TL: [Venezuela]  M T

Ganyra josephina josephina (Godart, 1819)  Hispaniola  M T S

Ganyra josephina krugii (Dewitz, 1877)  Puerto Rico & Mona  M T L

Ganyra josephina paramaryllis (W. Comstock, 1943)  Jamaica  M S

Ganyra menciae (Ramsden, [1914])  Cuban White  Cuba, St. Lucia  iN  M T S

Ganyra phaloe (Godart, 1819)    iN

Ganyra phaloe phaloe (Godart, 1819)  TL: Brazil  M T

Ganyra phaloe amphissa (Fruhstorfer, 1907)  TL: Bolivia  M T

Ganyra phaloe confusa Llorente, Le Crom & Torres, 2004  TL: Colombia

Ganyra phaloe diana (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Ganyra phaloe endeis (Godart, 1819)  TL: Brazil  M T

Ganyra phaloe imperator (W. F. Kirby, 1881)  TL: Ecuador  M T

Ganyra phaloe lamonti (Kaye, 1919)  TL: Trinidad  M T

Ganyra phaloe limona (Schaus, 1913)  Costa Rica to Colombia  M T S I

Ganyra phaloe sevata (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Ganyra phaloe sincera (Weymer, 1890)  TL: Ecuador  M T

Ganyra phaloe sublineata (Schaus, 1902)  TL: Peru  M T

Ganyra phaloe tiburtia (Fruhstorfer, 1907)  E & S Mexico to El Salvador  M T S L

Ganyra phaloe [n. ssp.] Lamas, MS  Peru

Ascia  Scopoli, 1777  

Ascia monuste (Linnaeus, 1764)  Great Southern White    iN

Ascia monuste raza Klots, 1930  Baja California Sur  M T S

Ascia monuste monuste (Linnaeus, 1764)  SW USA (strays), W Mexico & S TX to N S. Am; S Lesser Antilles  M S1 S2 L1 L2 L3 L4 I H

Ascia monuste phileta (Fabricius, 1775)  SE USA (strays N), N Bahamas, (possibly sporadic on Cuba)  M S L I

Ascia monuste eubotea (Godart, 1819)  Cuba, Hispaniola to Virgin Islands  M T S L I

Ascia monuste virginia (Godart, 1819)  Lesser Antilles between (not incl.) Virgin Islands and St. Vincent  M T S

Ascia monuste suasa (Boisduval, 1836)  TL: "Chile" - Peru  M T

Ascia monuste orseis (Godart, 1819)  TL: Brazil  M T L

Ascia monuste automate (Burmeister, 1878)  TL: Argentina  M T

Phulia  Herrich-Schäffer, 1867  

Phulia autodice (Hübner, 1818)  ref.12  iN

Phulia autodice autodice (Hübner, 1818)  TL: Paraguay  M T

Phulia autodice blanchardii (A. Butler, 1881)  TL: Chile  M T

Phulia autodice ernestae (Herrera, 1954)  TL: Chile

Phulia distincta (Jörgensen, 1916)  ref.12  iN

Phulia distincta distincta (Jörgensen, 1916)  TL: Argentina  M T

Phulia distincta fieldi (Herrera, 1970)  TL: Chile  M T

Phulia homoeodice (Paravicini, 1910)  ref.12  M T

Phulia inversa (Hayward, 1949)  ref.12  iN

Phulia inversa inversa (Hayward, 1949)  TL: Argentina  M T

Phulia inversa razmilici (Herrera, 1970)  TL: Chile  M T

Phulia mariae (Herrera, 1970)  ref.12  iN  M T

Phulia mercedis (Eschscholtz, 1821)  ref.12  iN

Phulia mercedis mercedis (Eschscholtz, 1821)  TL: Chile  M T

Phulia mercedis arctodice (Staudinger, 1899)  TL: Ecuador  M T

Phulia mercedis fueguensis (W. D. Field, 1959)  TL: Argentina  M T

Phulia mercedis macrodice (Staudinger, 1899)  TL: Bolivia  M T

Phulia mercedis sterodice (Staudinger, 1899)  TL: Chile  M T

Phulia mercedis vanvolxemii (Capronnier, 1874)  TL: Argentina  M T

Phulia orthodice (Weymer, 1890)  ref.12  iN

Phulia orthodice orthodice (Weymer, 1890)  TL: Bolivia  M T

Phulia orthodice carabaya (Ackery, 1975)  TL: Peru  M T

Phulia orthodice [n. ssp.] Lamas, MS  Peru

Phulia sagittata (Röber, 1908)  ref.12  iN

Phulia sagittata sagittata (Röber, 1908)  TL: Peru  M T

Phulia sagittata herrerai (Lamas, 1981)  TL: Peru  M T

Phulia stigmadice (Staudinger, 1894)  ref.12  iN  M T

Phulia theodice (Boisduval, 1832)  ref.12  iN

Phulia theodice theodice (Boisduval, 1832)  TL: [Chile]  M T

Phulia theodice gymnodice (Staudinger, 1899)  TL: Chile  M T

Phulia xanthodice (Lucas, 1852)  ref.12  iN

Phulia xanthodice xanthodice (Lucas, 1852)  TL: Venezuela; "Peru"  M T

Phulia xanthodice nigrisulphurata (Le Crom, Torres & Salazar, 2004)  TL: Colombia

Phulia xanthodice paramosa (Ackery, 1975)  TL: Colombia  M T

Phulia xanthodice paucar (Lamas, 1981)  TL: Peru  M T

Phulia xanthodice pyrrhomma (Röber, 1908)  TL: Peru  M T

Phulia maenacte (Boisduval, 1836)  ref.12  iN

Phulia maenacte maenacte (Boisduval, 1836)  TL: Paraguay  M T

Phulia maenacte itatiayae (Foetterle, 1902)  TL: Brazil (RJ)  M T

Phulia coropunae (Dyar, 1913)  ref.12  M T

Phulia forsteri (W. D. Field & Herrera, 1977)  ref.12  M T

Phulia stoddardi T. Pyrcz & Cerdeña , 2023  ref.284

Phulia huanaco (Staudinger, 1894)  ref.12  iN  M T

Phulia isabela (W. D. Field & Herrera, 1977)  ref.12  M T

Phulia nysias Weymer, 1890  ref.12  iN  M T

Phulia rosea (Ureta, 1956)  ref.12  iN

Phulia rosea rosea (Ureta, 1956)  TL: Chile

Phulia rosea annamariea (W. D. Field & Herrera, 1977)  TL: Bolivia  M T

Phulia rosea maria (W. D. Field & Herrera, 1977)  TL: Chile  M T

Phulia argyrodice (Staudinger, 1899)  ref.12  iN  M T

Phulia galactodice (Ureta, 1955)  ref.12

Phulia huemul (L. Peña, 1964)  ref.12  iN  M T

Phulia microdice (Blanchard, 1852)  ref.12  iN  M T

Phulia penai (Ureta, 1955)  ref.12

Phulia wagenknechti (Ureta, 1938)  ref.12  iN

Phulia wagenknechti wagenknechti (Ureta, 1938)  TL: Chile

Phulia wagenknechti sulfurodice (Ureta, 1955)  TL: Chile  M S

Phulia pelambres (Benyamini, 2014)  ref.12

Phulia phantasma Lamas, Willmott & Boyer, 2023  ref.284

Phulia illimani Weymer, 1890  TL: Bolivia  M T

Phulia ilyodes Ureta, 1955    iN  M S

Phulia madeleinea (W. D. Field & Herrera, 1977)    iN  M T L

Phulia garleppi W. D. Field & Herrera, 1977  TL: Peru

Phulia garleppi garleppi W. D. Field & Herrera, 1977  TL: Peru  M S

Phulia garleppi [n. ssp.] Lamas, MS  Peru

Phulia nannophyes Dyar, 1913  TL: Peru

Phulia nannophyes nannophyes Dyar, 1913  TL: Peru  M T

Phulia nannophyes galera Lamas, 1981  TL: Peru

Phulia nymphula (Blanchard, 1852)    iN

Phulia nymphula nymphula (Blanchard, 1852)  TL: Chile  M T

Phulia nymphula nympha Staudinger, 1894  TL: Bolivia  M T

Phulia paranympha Staudinger, 1894  TL: Bolivia  M T

Pontia  Fabricius, 1807  

Pontia beckerii (W. H. Edwards, 1871)  Becker's White  BC to Baja C. Sur, N AZ, N NM, CO, WY, S & W MT; basins  iN  M T S1 S2 L1 L2 I H

Pontia santamarta (Ackery, 1975)  Colombia; ref.12  M T

Pontia occidentalis (Reakirt, 1866)  Western White    iN

Pontia occidentalis occidentalis (Reakirt, 1866)  BC to CA (S Sierra), W to N NM, N to MAN, SASK, ALTA  M S1 S2 L1 L2 L3 L4 I Ha

Pontia occidentalis nelsoni (W. H. Edwards, 1883)  Alaska  M S L

Pontia protodice (Boisduval & Le Conte, [1830])  Checkered White  widespread across N America to S Mexico; Cuba. Vagrant N  iN  M T S L1 L2 L3 L4 L5 L6 I H

Pontia sisymbrii (Boisduval, 1852)  Spring White    iN

Pontia sisymbrii sisymbrii (Boisduval, 1852)  Baja California, N along Sierra Nevada-Cascades to W-C OR  M T S L I Ha

Pontia sisymbrii nigravenosa Austin & J. Emmel, 1998  Mojave Desert: SE CA, W AZ, S NV, SW UT  M T1 T2 L

Pontia sisymbrii transversa R. Holland, 1995  S AZ, S NM, W TX, N SON, N CHIH  M T S L I

Pontia sisymbrii elivata (W. Barnes & Benjamin, 1926)  SE OR to C NV, E to CO, N NM  M S1 S2 L I

Pontia sisymbrii nordini (K. Johnson, 1977)  W NE, E WY, W SD, E MT  M S

Pontia sisymbrii flavitincta (J. A. Comstock, 1924)  S BC to NE OR, E to NW WY, W MT, S ALTA  M T S L

Pontia sisymbrii beringiensis C. Guppy & Kondla, 2001  N BC-S YK  M S

Pieris  Schrank, 1801  

Pieris angelika Eitschberger, [1984]  Arctic White  C AK to YK & N BC  iN

Pieris angelika angelika Eitschberger, [1984]  C AK to YK & N BC  iN  M T S1 S2 S3 L Ha

Pieris marginalis Scudder, 1861  Margined White    iN

Pieris marginalis pseudobryoniae W. Barnes & McDunnough, 1916  TL: USA (Ak)

Pieris marginalis shapiroi Eitschberger, [1984]  TL: USA (Ak)

Pieris marginalis hulda W. H. Edwards, 1869  AK coast from Kodiak Island to Aleutian Islands  M S L

Pieris marginalis meckyae Eitschberger, [1984]  S-central AK (e.g., Anchorage to Palmer)  M S

Pieris marginalis guppyi Eitschberger, [1984]  TL: USA (Ak)

Pieris marginalis tremblayi Eitschberger, [1984]  TL: USA (Ak), Canada (BC)

Pieris marginalis reicheli Eitschberger, [1984]  Rocky Mts. Of BC, ALTA, N ID, N MT  M S L

Pieris marginalis marginalis Scudder, 1861  W OR (N of Siskiyous), W Cascades to coast, N to SW BC  M S1 S2 L Ha

Pieris marginalis sequoia J. Emmel, T. Emmel & Mattoon, 1998  N CA coast: redwood forests Humboldt Co. (vic. Prairie Ck. S.P.)  M T1 T2 S

Pieris marginalis venosa Scudder, 1861  much of W CA  M S L1 L2

Pieris marginalis castoria Reakirt, 1866  Siskiyou Mts. (SW OR, NW CA), W Sierra Nevada  M T S L I

Pieris marginalis Warner Mts. segregate ,  Warner Mts., Lake Co. OR - Modoc Co. CA  M S

Pieris marginalis ziegleri Eitschberger, 1991  NW WY, W MT, ID, E WA, NE OR  M S1 S2 L

Pieris marginalis pallidissima W. Barnes & McDunnough, 1916  W CO, UT, far NW AZ, NE NV, S ID, SW WY; bivoltine  M S L I Ha

Pieris marginalis macdunnoughii C. Remington, 1954  [= macdunnoughii], CO, S WY, N NM; univoltine  M S L1 L2 Ha

Pieris marginalis mogollon Burdick, 1942  White Mts. (AZ) & Mogollon Rim (NM)  M T S L1 L2 I H

Pieris marginalis siblanca Grishin, 2022  S NM; ref.16

Pieris oleracea (T. Harris, 1829)  Mustard White    iN

Pieris oleracea oleracea (T. Harris, 1829)  TL: USA (Ma/NH)  M S1 S2 L

Pieris oleracea frigida Scudder, 1861  TL: Canada (NL)  M S

Pieris virginiensis W. H. Edwards, 1870  West Virginia White    iN

Pieris virginiensis virginiensis W. H. Edwards, 1870  TL: USA (WV)  M S L

Pieris virginiensis hyatti Eitschberger, [1984]  TL: Canada (On)  M S Ha

Pieris rapae (Linnaeus, 1758)  Cabbage White    iN

Pieris rapae rapae (Linnaeus, 1758)  widespread across N America to C Mexico, Hawaii; introduced  M T S HS L1 L2 L3 L4 I H

Pieris brassicae (Linnaeus, 1758)  Accidental rare escapee in US, not to be expected  iN  M T

Glennia  Klots, 1933  

Glennia pylotis (Godart, 1819)    iN  M T

Leptophobia  A. Butler, 1870  

Leptophobia aripa (Boisduval, 1836)    iN

Leptophobia aripa aripa (Boisduval, 1836)  TL: Venezuela  M T Ma L

Leptophobia aripa balidia (Boisduval, 1836)  TL: Brazil  M T

Leptophobia aripa elodia (Boisduval, 1836)  S TX (rare stray), E & W Mexico to Panama  M T S L I

Leptophobia aripa elodina (Röber, 1908)  TL: Bolivia  M T S

Leptophobia caesia (Lucas, 1852)    iN

Leptophobia caesia caesia (Lucas, 1852)  TL: Ecuador  M T

Leptophobia caesia phanokia Fruhstorfer, 1907  TL: Colombia  M T

Leptophobia caesia tenuicornis A. Butler & H. Druce, 1872  Costa Rica & Panama  M T S L

Leptophobia caesia [n. ssp.] Lamas, MS  Colombia

Leptophobia cinerea (Hewitson, 1867)    iN

Leptophobia cinerea cinerea (Hewitson, 1867)  TL: Ecuador  M T S

Leptophobia cinerea menthe (Hopffer, 1874)  TL: Peru  M T

Leptophobia diaguita Jörgensen, 1916    iN

Leptophobia diaguita diaguita Jörgensen, 1916  TL: Argentina  M T

Leptophobia diaguita latifascia Joicey & Talbot, 1928  TL: Peru; Ecuador  M T

Leptophobia diaguita mandor Lamas, 2003  TL: Peru

Leptophobia eleone (E. Doubleday, 1847)    iN

Leptophobia eleone eleone (E. Doubleday, 1847)  TL: [Colombia]  M T

Leptophobia eleone luca Fruhstorfer, 1907  TL: Bolivia  M T Ma L

Leptophobia eleone denigrata Le Crom, Andrade & Winhard, 2004  TL: Colombia

Leptophobia eleusis (Lucas, 1852)    iN

Leptophobia eleusis eleusis (Lucas, 1852)  TL: Colombia  M T L

Leptophobia eleusis mollitica Fruhstorfer, 1908  TL: Peru  M T

Leptophobia eleusis [n. ssp.] Lamas, MS  Peru

Leptophobia erinna (Hopffer, 1874)  TL: Peru  M T

Leptophobia eucosma (Erschoff, 1875)  TL: Peru

Leptophobia eucosma eucosma (Erschoff, 1875)  TL: Peru  M T

Leptophobia eucosma euremoides Röber, 1908  TL: Peru  M T

Leptophobia eucosma [n. ssp.#1] Lamas, MS  TL: Peru

Leptophobia eucosma [n. ssp.#2] Lamas, MS  TL: Peru

Leptophobia forsteri Baumann & Reissinger, 1969  TL: Peru  M T

Leptophobia gonzaga Fruhstorfer, 1908  TL: Ecuador

Leptophobia gonzaga sotara Lamas, G. Rodríguez & Constantino, 2004  TL: Colombia  M T

Leptophobia gonzaga gonzaga Fruhstorfer, 1908  TL: Ecuador  M T

Leptophobia gonzaga [n. ssp.] Lamas, MS  Ecuador

Leptophobia helena (Lucas, 1852)    iN

Leptophobia helena helena (Lucas, 1852)  TL: Ecuador  M T

Leptophobia helena doubledayi Röber, 1908  TL: Bolivia  M T

Leptophobia helena hughesi Lamas, 2003  TL: Peru

Leptophobia helena smithii (W. F. Kirby, 1881)  TL: Ecuador  M T

Leptophobia helena [n. ssp.#1] Lamas, MS  TL: Colombia

Leptophobia helena [n. ssp.#2] Lamas, MS  TL: Ecuador

Leptophobia micaia Lamas, Pyrcz & G. Rodríguez, 2004  TL: Colombia  M T

Leptophobia nephthis (Hopffer, 1874)  TL: Bolivia; Peru  M T

Leptophobia olympia (C. Felder & R. Felder, 1861)    iN

Leptophobia olympia olympia (C. Felder & R. Felder, 1861)  TL: Venezuela  M T L

Leptophobia olympia potoniei Baumann & Reissinger, 1969  TL: Peru  M T

Leptophobia olympia [n. ssp.] Lamas, MS  Peru

Leptophobia penthica (Kollar, 1850)    iN

Leptophobia penthica penthica (Kollar, 1850)  TL: Colombia  M T

Leptophobia penthica basiliola Fruhstorfer, 1908  TL: Bolivia  M T

Leptophobia penthica semicaesia (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Leptophobia penthica stamnata (Lucas, 1852)  TL: Venezuela  M T

Leptophobia penthica [n. ssp.#1] Lamas, MS  TL: Ecuador

Leptophobia penthica [n. ssp.#2] Lamas, MS  TL: Peru

Leptophobia penthica [n. ssp.#3] Lamas, MS  TL: Peru

Leptophobia penthica [n. ssp.#4] Lamas, MS  TL: Peru

Leptophobia penthica [n. ssp.#5] Lamas, MS  TL: Bolivia

Leptophobia philoma (Hewitson, 1870)    iN

Leptophobia philoma philoma (Hewitson, 1870)  TL: Ecuador  M T

Leptophobia philoma pastaza (Joicey & Talbot, 1928)  TL: Ecuador  M T

Leptophobia philoma subargentea A. Butler, 1898  TL: Peru  M T S L

Leptophobia philoma intermedia Llorente & Le Crom, 2004  TL: Colombia

Leptophobia philoma pseudopenthica Le Crom & Llorente, 2004  TL: Colombia

Leptophobia philoma [n. ssp.] Lamas, MS  Peru

Leptophobia pinara (C. Felder & R. Felder, 1865)  TL: Colombia

Leptophobia pinara pinara (C. Felder & R. Felder, 1865)  TL: Colombia  M T L

Leptophobia pinara [n. ssp.] Lamas, MS  Peru

Leptophobia tovaria (C. Felder & R. Felder, 1861)    iN

Leptophobia tovaria Panama segregate ,  Panama  M S

Leptophobia tovaria tovaria (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Leptophobia tovaria gina Fruhstorfer, 1908  TL: Peru  M T S

Leptophobia tovaria maruga Fruhstorfer, 1907  TL: Ecuador  M T L

Leptophobia tovaria subflavescens (W. F. Kirby, 1887)  TL: Colombia  M T

Leptophobia tovaria sanctipetri Andrade, Le Crom & Winhard, 2004  TL: Colombia

Leptophobia tovaria pseudolympia Le Crom, Llorente & Salazar, 2004  TL: Colombia

Leptophobia tovaria [n. ssp.#1] Lamas, MS  TL: Panama

Leptophobia tovaria [n. ssp.#2] Lamas, MS  TL: Bolivia

Leptophobia tovaria [n. ssp.#3] Lamas, MS  TL: Argentina

Leptophobia [n. sp.#1] Lamas, MS  TL: not specified

Leptophobia [n. sp.#1] [n. ssp.#1] Lamas, MS  TL: Venezuela

Leptophobia [n. sp.#1] [n. ssp.#2] Lamas, MS  TL: Colombia

Leptophobia [n. sp.#1] [n. ssp.#3] Lamas, MS  TL: Ecuador

Leptophobia [n. sp.#1] [n. ssp.#4] Lamas, MS  TL: Peru

Leptophobia [n. sp.#2] Lamas, MS  TL: Peru

Itaballia  Kaye, 1904  

Itaballia demophile (Linnaeus, 1763)    iN

Itaballia demophile demophile (Linnaeus, 1763)  TL: Indiis

Itaballia demophile calydonia (Boisduval, 1836)  TL: Colombia  M T

Itaballia demophile centralis Joicey & Talbot, 1928  S TX (one-time stray), E & S Mexico to Panama  M T S1 S2 L I

Itaballia demophile charopus (Fruhstorfer, 1907)  TL: Brazil (Am)  M T

Itaballia demophile huebneri (Fruhstorfer, 1907)  TL: [Brazil]

Itaballia demophile lucania (Fruhstorfer, 1907)  TL: Peru  M T S

Itaballia demophile nimietes (Fruhstorfer, 1907)  TL: Brazil (Ba)  M T

Itaballia demophile niphates (Fruhstorfer, 1907)  TL: Brazil (Pa)  M T

Itaballia demophile niseias (Fruhstorfer, 1907)  TL: Paraguay  M T

Itaballia marana (E. Doubleday, 1844)    iN  M T

Itaballia pandosia (Hewitson, 1853)    iN

Itaballia pandosia pandosia (Hewitson, 1853)  TL: Venezuela  M T

Itaballia pandosia kicaha (Reakirt, 1863)  S Mexico to Panama  M T S L Ha

Itaballia pandosia pisonis (Hewitson, 1861)  TL: "[Upper] Amazon"; Colombia  M T

Itaballia pandosia sabata (Fruhstorfer, 1907)  SE Panama, W Colombia  M T S L

Pieriballia  Klots, 1933  

Pieriballia viardi (Boisduval, 1836)  Viardi White    iN

Pieriballia viardi viardi (Boisduval, 1836)  S TX (rare stray), E & W Mexico to Honduras  M T S1 S2 Ab L1 L2

Pieriballia viardi apicalis (A. Butler, 1898)  TL: Ecuador  M T

Pieriballia viardi decorata (Joicey & Talbot, 1928)  TL: Peru  M T

Pieriballia viardi interposita (Joicey & Talbot, 1928)  TL: Peru  M T

Pieriballia viardi locusta (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Pieriballia viardi mandela (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Pieriballia viardi molione (Fruhstorfer, 1908)  TL: Paraguay  M T

Pieriballia viardi noctipennis (A. Butler & H. Druce, 1872)  Costa Rica & Panama  M T S L I

Pieriballia viardi rubecula (Fruhstorfer, 1907)  TL: Peru  M T

Pieriballia viardi tithoreides (A. Butler, 1898)  TL: Ecuador  M T

Perrhybris  Hübner, [1819]  

Perrhybris lorena (Hewitson, 1852)    iN  M T

Perrhybris lypera (Kollar, 1850)    iN

Perrhybris lypera lypera (Kollar, 1850)  TL: Colombia  M T

Perrhybris lypera septentrionalis Röber, 1928  Guatemala to Panama  M T S

Perrhybris lypera sulphuralis (A. Butler, 1896)  TL: Colombia  M T

Perrhybris pamela (Stoll, 1780)    iN

Perrhybris pamela pamela (Stoll, 1780)  TL: Surinam

Perrhybris pamela alethina (A. Butler, 1872)  Costa Rica & Panama  M T S1 S2 I

Perrhybris pamela amazonica Fruhstorfer, 1907  TL: Peru  M T

Perrhybris pamela bertha Lamas, 1981  TL: Peru

Perrhybris pamela bogotana (A. Butler, 1898)  TL: Colombia  M T

Perrhybris pamela boyi J. Zikán, 1940  TL: Brazil (Am)  M T

Perrhybris pamela carmenta Fruhstorfer, 1907  TL: Peru  M T S

Perrhybris pamela chajulensis J. Maza & R. G. Maza, 1989  N Oaxaca, Chiapas to Honduras  M T S

Perrhybris pamela eieidias Hübner, [1821]  TL: [Brazil]  M T

Perrhybris pamela flava (Oberthür, 1896)  TL: S. Amer.  M T

Perrhybris pamela fruhstorferi Röber, 1908  Panama  M S L

Perrhybris pamela glessaria Fruhstorfer, 1907  TL: Ecuador  M T

Perrhybris pamela incisa Fruhstorfer, 1907  TL: Brazil (Ba)  M T

Perrhybris pamela lucasi Fruhstorfer, 1907  TL: French Guiana; Brazil

Perrhybris pamela malenka (Hewitson, 1852)  TL: Venezuela  M T

Perrhybris pamela mapa J. Maza & R. G. Maza, 1989  SW Chiapas  M T S

Perrhybris pamela mazuka Lamas, 1981  TL: Peru

Perrhybris pamela [n. ssp.] Lamas, MS  Costa Rica

Charonias  Röber, 1908  

Charonias eurytele (Hewitson, [1853])    iN

Charonias eurytele eurytele (Hewitson, [1853])  TL: Ecuador  M T

Charonias eurytele caraca Kaye, 1913  TL: Venezuela

Charonias eurytele dismorphina Röber, 1908  TL: Colombia  M T

Charonias eurytele dismorphites (A. Butler, 1872)  Costa Rica & Panama  M T S

Charonias eurytele nigrescens (Salvin & Godman, 1868)  S Mexico to Guatemala (at least)  M T S

Charonias eurytele [n. ssp.#1] Lamas, MS  TL: Colombia  M T

Charonias eurytele [n. ssp.#2] Lamas, MS  TL: Venezuela

Charonias theano (Boisduval, 1836)    iN  M T

Archonias  Hübner, [1831]  [includes Catasticta]

Archonias brassolis (Fabricius, 1776)  ref.12  iN

Archonias brassolis brassolis (Fabricius, 1776)  TL: Surinam

Archonias brassolis approximata (A. Butler, 1873)  SE Mexico to Panama  M T S L

Archonias brassolis brevia G. Rodríguez, Le Crom & Salazar, 2004  TL: Colombia

Archonias brassolis critias (C. Felder & R. Felder, 1859)  TL: S. Amer.  M T

Archonias brassolis cutila Fruhstorfer, 1907  TL: Ecuador  M T

Archonias brassolis marcias Hübner, [1831]  TL: Brazil (Ba)

Archonias brassolis negrina (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Archonias brassolis nigripennis (A. Butler, 1873)  TL: Colombia  M T

Archonias brassolis rosacea (A. Butler, 1873)  TL: Ecuador  M T

Archonias brassolis rubrosparsa Stichel, 1908  TL: Ecuador  M T

Archonias brassolis semialba Constantino, Salazar & Rodríguez, 2004  TL: Colombia  M T

Archonias brassolis tereas (Godart, 1819)  TL: Brazil  M T

Archonias abiseo (Lamas & Bollino, 2004)  TL: Peru  M T

Archonias affinis (Röber, 1909)    iN

Archonias affinis affinis (Röber, 1909)  TL: Colombia  M T

Archonias affinis giga (F. Brown & Gabriel, 1939)  Colombia  iN  M T

Archonias albofasciata (Lathy & Rosenberg, 1912)    iN

Archonias albofasciata albofasciata (Lathy & Rosenberg, 1912)  TL: Colombia  M T

Archonias albofasciata rubroreducta (Reissinger, 1972)  TL: Ecuador  M T

Archonias amastris (Hewitson, 1874)  TL: Bolivia

Archonias amastris amastris (Hewitson, 1874)  TL: Bolivia  M T

Archonias amastris dentata (Lathy & Rosenberg, 1912)  TL: Peru  M T S

Archonias anaitis (Hewitson, 1869)    iN

Archonias anaitis anaitis (Hewitson, 1869)  TL: Ecuador  M T

Archonias anaitis felicitas (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias anaitis sordida (A. Butler, 1897)  TL: Bolivia  M T

Archonias anaitis suasella (Röber, 1908)  TL: Peru  M T

Archonias apaturina (A. Butler, 1901)    iN

Archonias apaturina apaturina (A. Butler, 1901)  TL: Ecuador  M T

Archonias apaturina citra (F. Brown, 1939)  TL: Peru  M T

Archonias apaturina subturina (Reissinger, 1972)  TL: Colombia  M T

Archonias apaturina supraturina (Reissinger, 1972)  TL: Peru  M T

Archonias arborardens (Reissinger, 1972)  TL: Peru  M T

Archonias atahuallpa (Eitschberger & T. Racheli, 1998)  TL: Peru  M S

Archonias aureomaculata (Lathy & Rosenberg, 1912)    iN

Archonias aureomaculata aureomaculata (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias aureomaculata gabrieli (Reissinger, 1972)  TL: Peru  M T

Archonias aureomaculata [n. ssp.] (Lamas, MS)  Peru

Archonias bithys (Hübner, [1831])    iN  M T

Archonias cerberus (Godman & Salvin, 1889)  Costa Rican Dartwhite  Costa Rica & Panama  iN  M T S

Archonias chelidonis (Hopffer, 1874)    iN

Archonias chelidonis chelidonis (Hopffer, 1874)  TL: Bolivia  M T S

Archonias chelidonis contrasta (Reissinger, 1972)  TL: Peru  M T

Archonias chelidonis igneata (Reissinger, 1972)  TL: Peru  M T

Archonias chelidonis jacinta (A. Butler, 1901)  TL: Bolivia  M T

Archonias chelidonis taminoides (Eitschberger & T. Racheli, 1998)  TL: Bolivia  M T

Archonias chrysolopha (Kollar, 1850)    iN

Archonias chrysolopha chrysolopha (Kollar, 1850)  TL: [Colombia]  M T

Archonias chrysolopha adamsi (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias chrysolopha beatrizae (Bollino & Costa, 2007)  ref.171  M T

Archonias chrysolopha quiroza (F. Brown, 1939)  TL: Peru  M T S

Archonias chrysolopha spectrum (Reissinger, 1972)  TL: Venezuela  M T

Archonias chrysolopha [n. ssp.] (Lamas, MS)  Peru

Archonias cinerea (A. Butler, 1897)    iN

Archonias cinerea cinerea (A. Butler, 1897)  TL: ["Ecuador"]  M T S

Archonias cinerea farallonensis (Rodríguez, 2017)  TL: Colombia

Archonias cinerea suprema (Fassl, 1915)  TL: Colombia  M T

Archonias coerulescens (Eitschberger & T. Racheli, 1998)    iN

Archonias coerulescens coerulescens (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias coerulescens intimpa (Bollino & Boyer, 2008)  ref.170

Archonias colla (E. Doubleday, 1847)    iN

Archonias colla colla (E. Doubleday, 1847)  TL: Bolivia  M T

Archonias colla philomene (Röber, 1908)  TL: Bolivia  M T

Archonias colla punctata (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias collina (F. Brown, 1939)    iN

Archonias collina collina (F. Brown, 1939)  TL: Peru  M T

Archonias collina carpis (Reissinger, 1972)  TL: Peru  M T

Archonias cora (Lucas, 1852)    iN  M T S

Archonias corcyra (C. Felder & R. Felder, 1859)    iN

Archonias corcyra corcyra (C. Felder & R. Felder, 1859)  TL: [Bolivia]  M T

Archonias corcyra staudingeri (A. Butler, 1897)  TL: Peru  M T

Archonias ctemene (Hewitson, 1869)    iN

Archonias ctemene ctemene (Hewitson, 1869)  TL: Ecuador  M T

Archonias ctemene actinotis (A. Butler, 1872)  Costa Rica & Panama  M T S L

Archonias ctemene alma (Hopffer, 1874)  TL: Bolivia  M T Ma L

Archonias ctemene grisella (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias ctemene pitalito (Rodríguez, 2017)  TL: Colombia

Archonias ctemene rubricata (Weymer, 1907)  TL: Colombia  M T

Archonias ctemene strigosa (A. Butler, 1896)  TL: Peru  M T S

Archonias ctemene zebrella (Fruhstorfer, 1912)  TL: Peru  M T

Archonias ctemene [n. ssp.] (Lamas, MS)  Peru

Archonias discalba (F. Brown & Gabriel, 1939)  TL: Ecuador  M T

Archonias distincta (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias duida (F. Brown, 1932)    iN  M T

Archonias eurigania (Hewitson, 1870)    iN

Archonias eurigania eurigania (Hewitson, 1870)  TL: Ecuador  M T S

Archonias eurigania straminea (A. Butler, 1896)  TL: Ecuador  M T S

Archonias ferra (F. Brown & Gabriel, 1939)    iN

Archonias ferra ferra (F. Brown & Gabriel, 1939)  TL: Ecuador  M T L

Archonias ferra ferruginosa (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias ferra orcus (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias ferra putumayo (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias flisa (Herrich-Schäffer, [1858])  Narrow-banded Dartwhite    iN

Archonias flisa flisa (Herrich-Schäffer, [1858])  [= arechiza], E & W Mexico to Guatemala  M T S L

Archonias flisa archoflisa (Eitschberger & T. Racheli, 1998)  Panama  M T

Archonias flisa arechiza (Reakirt, 1866)  TL: Mexico  M T

Archonias flisa briseis (Eitschberger & T. Racheli, 1998)  TL: Venezuela

Archonias flisa dilutior (Avinoff, 1926)  TL: Colombia  M T

Archonias flisa duna (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T L

Archonias flisa flisandra (Reissinger, 1972)  Chiapas & Guatemala  M T S L

Archonias flisa flisella (Reissinger, 1972)  SE Mexico to El Salvador  M T S

Archonias flisa flisoides (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias flisa melanisa (Eitschberger & T. Racheli, 1998)  Costa Rica  M T S L

Archonias flisa noakesi (Joicey & Rosenberg, 1915)  TL: Colombia  M T

Archonias flisa postaurea (F. Brown, 1933)  TL: Colombia  M T

Archonias flisa viloriai (Bollino & Costa, 2007)  ref.171  M T

Archonias oaxaca (Beutelspacher, 1984)  Oaxacan Dartwhite  Oaxaca and Chiapas, Mexico  M T L

Archonias frontina (F. Brown & Gabriel, 1939)    iN

Archonias frontina frontina (F. Brown & Gabriel, 1939)  TL: Colombia  M T

Archonias frontina muehlei (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias fulva (Joicey & Rosenberg, 1915)  TL: Peru

Archonias fulva fulva (Joicey & Rosenberg, 1915)  TL: Peru  M T S

Archonias fulva kentae (F. Brown & Goodson, 1940)  TL: Peru  M T

Archonias fulva anaranjada (Bollino & S. Padrón, 2016)  ref.172

Archonias grisea (Joicey & Rosenberg, 1915)    iN  M T

Archonias hebra (Lucas, 1852)    iN

Archonias hebra hebra (Lucas, 1852)  TL: Colombia  M T

Archonias hebra apollinari (Fassl, 1915)  TL: Colombia  M T

Archonias hebra crowleyi (A. Butler, 1901)  TL: Venezuela  M T

Archonias hebra mariae (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias hebra potamea (C. Felder & R. Felder, 1861)  TL: Venezuela; "Ecuador"  M T

Archonias hegemon (Godman & Salvin, 1889)  Frosted Dartwhite    iN

Archonias hegemon hegemon (Godman & Salvin, 1889)  Costa Rica & Panama  M T S L

Archonias hegemon helle (Röber, 1924)  TL: Colombia  M T

Archonias hegemon tatae (F. Brown & Gabriel, 1939)  TL: Ecuador  M T

Archonias huancabambensis (Joicey & Rosenberg, 1915)  TL: Peru

Archonias huancabambensis huancabambensis (Joicey & Rosenberg, 1915)  TL: Peru  M T

Archonias huancabambensis bamba (Reissinger, 1972)  TL: Peru  M T

Archonias huebneri (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias incerta Dognin, 1888  TL: Ecuador

Archonias incerta incerta Dognin, 1888  TL: Ecuador  M T

Archonias incerta concerta (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias lanceolata (Lathy & Rosenberg, 1912)    iN

Archonias lanceolata lanceolata (Lathy & Rosenberg, 1912)  TL: Colombia  M T

Archonias lanceolata alexandra (Bollino & L. Rodríguez, 2003)  TL: Colombia  M T

Archonias leucophaea (Lathy & Rosenberg, 1912)    iN

Archonias leucophaea leucophaea (Lathy & Rosenberg, 1912)  TL: Peru  M T S

Archonias leucophaea lathyi (Joicey & Talbot, 1918)  TL: Ecuador  M T

Archonias lisa (Baumann & Reissinger, 1969)  TL: Peru  M T

Archonias ludovici (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias lycurgus (Godman & Salvin, 1880)  TL: Colombia  M T

Archonias manco (E. Doubleday, 1848)    iN

Archonias manco manco (E. Doubleday, 1848)  TL: Bolivia  M T

Archonias manco capac (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias manco reissingeri (Eitschberger & T. Racheli, 1998)  TL: Peru  M T S

Archonias marcapita (Röber, 1909)  TL: Bolivia

Archonias marcapita marcapita (Röber, 1909)  TL: Bolivia  M T S

Archonias marcapita boettgeri (Bollino & Lamas, 2004)  TL: Peru  M T

Archonias marcapita roberti (Reissinger, 1972)  TL: Peru  M T

Archonias modesta (Lucas, 1852)    iN

Archonias modesta modesta (Lucas, 1852)  TL: Peru  M T S

Archonias modesta goodsoni (Reissinger, 1972)  TL: Peru  M T

Archonias modesta sachapuyo (Bollino & Vitale, 2003)  TL: Peru  M T

Archonias modesta [n. ssp.] (Bollino, MS)  Bolivia

Archonias nimbata (Joicey & Talbot, 1918)  TL: Peru

Archonias nimbata nimbata (Joicey & Talbot, 1918)  TL: Peru  M T

Archonias nimbata philobata (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias nimbata pleione (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias nimbice (Boisduval, 1836)  Mexican Dartwhite    iN

Archonias nimbice nimbice (Boisduval, 1836)  W TX, E & W Mexico to Oaxaca  M T S L1 L2

Archonias nimbice ochracea (H. Bates, 1864)  Chiapas and Guatemala to Nicaragua  M T S L

Archonias nimbice bryson (Godman & Salvin, 1889)  Costa Rica & Panama  M T S L

Archonias nimbice ligata (Eitschberger & T. Racheli, 1998)  Panama  M T S L

Archonias nimbice [n. ssp.] (Llorente, MS)  Los Tuxtlas region, Veracruz  M S

Archonias notha (E. Doubleday, 1847)    iN

Archonias notha notha (E. Doubleday, 1847)  TL: Venezuela  M T

Archonias notha caucana (Röber, 1908)  TL: Colombia  M T

Archonias notha pieridoides (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Archonias paucartambo (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias pharnakia (Fruhstorfer, 1907)    iN

Archonias pharnakia pharnakia (Fruhstorfer, 1907)  TL: Peru  M T S

Archonias pharnakia styx (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias philais (C. Felder & R. Felder, 1865)    iN

Archonias philais philais (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Archonias philais borgesi (Eitschberger & T. Racheli, 1998)  TL: Venezuela  M T

Archonias philodora (F. Brown, 1939)    iN

Archonias philodora philodora (F. Brown, 1939)  TL: Ecuador  M L

Archonias philodora maja (Reissinger, 1972)  TL: Ecuador  M T

Archonias philodora zamora (Reissinger, 1972)  TL: Ecuador  M T

Archonias philone (C. Felder & R. Felder, 1865)    iN

Archonias philone philone (C. Felder & R. Felder, 1865)  TL: Colombia; "Ecuador"  M T

Archonias philone ecuadora (F. Brown, 1939)  TL: Ecuador  M T

Archonias philone florida (Bollino & Vitale, 2003)  TL: Peru  M T

Archonias philone grossana (F. Brown & Gabriel, 1939)  TL: Colombia  M T

Archonias philone stabilis (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias philone variabilis (Röber, 1924)  TL: Colombia  M T S

Archonias philoscia (C. Felder & R. Felder, 1861)  TL: Venezuela

Archonias philoscia philoscia (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Archonias philoscia incertina (Röber, 1924)  TL: Colombia  M T

Archonias philoscia reyi (Bollino & Costa, 2007)  ref.171  M T

Archonias philothea (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Archonias pieris (Hopffer, 1874)    iN

Archonias pieris pieris (Hopffer, 1874)  TL: "Bolivia"; Peru  M T S

Archonias pieris innuba (Röber, 1908)  TL: Bolivia  M T Ma L

Archonias pieris intermedia (Lathy & Rosenberg, 1912)  TL: Peru  M T S

Archonias pillcopata (Bollino, 2008)  ref.169

Archonias pinava (E. Doubleday, 1847)    iN

Archonias pinava pinava (E. Doubleday, 1847)  TL: Bolivia  M T S

Archonias pinava lucida (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias pluvius (Tessmann, 1928)  TL: Peru

Archonias pluvius pluvius (Tessmann, 1928)  TL: Peru  M T

Archonias pluvius seminigra (Bollino & Vitale, 2004)  ref.173  M T

Archonias potameoides (Reissinger, 1972)  TL: Peru  M T

Archonias poujadei (Dognin, 1887)    iN

Archonias poujadei poujadei (Dognin, 1887)  TL: Ecuador  M T

Archonias poujadei condor (Radford & Willmott, 2013)  ref.287

Archonias poujadei clara (Röber, 1909)  TL: [Peru]  M T S

Archonias poujadei eximia (Röber, 1909)  TL: Bolivia  M T

Archonias poujadei lamasi (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias prioneris (Hopffer, 1874)    iN

Archonias prioneris prioneris (Hopffer, 1874)  TL: Peru  M T S

Archonias prioneris albescens (Röber, 1924)  TL: Colombia  M T

Archonias prioneris araguana (Eitschberger & T. Racheli, 1998)  TL: Venezuela  M T

Archonias prioneris estancia (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias prioneris nigrilineata (Bollino, L. Rodríguez & Constantino, 2004)  TL: Colombia  M T

Archonias prioneris parrishi (Bollino & Costa, 2007)  ref.171  M T

Archonias prioneris peruviana (Joicey & Talbot, 1918)  TL: Peru  M T

Archonias pyrczi (Bollino, 2008)  ref.169

Archonias radiata (Kollar, 1850)    iN

Archonias radiata radiata (Kollar, 1850)  TL: [Colombia]  M T

Archonias radiata julita (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias reducta (A. Butler, 1896)    iN

Archonias reducta reducta (A. Butler, 1896)  TL: Ecuador  M T

Archonias reducta boliviana (A. Butler, 1896)  TL: Bolivia  M T S Ma L

Archonias reducta butleria (F. Brown & Gabriel, 1939)  TL: Peru  M T

Archonias revancha (Rey & Pyrcz, 1996)  TL: Venezuela  M T

Archonias rileya (F. Brown & Gabriel, 1939)  TL: Peru  M T

Archonias rochereaui (Le Cerf, 1924)  TL: Colombia

Archonias rochereaui rochereaui (Le Cerf, 1924)  TL: Colombia  M T

Archonias rochereaui hollandi (Avinoff, 1926)  TL: Colombia  M T

Archonias rochereaui laurentina (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias rosea (Joicey & Rosenberg, 1915)  TL: Colombia  M T

Archonias scaeva (Röber, 1909)  TL: Peru

Archonias scaeva scaeva (Röber, 1909)  TL: Peru  M T S

Archonias scaeva restricta (F. Brown & Goodson, 1940)  TL: Bolivia  M T

Archonias scurra (Röber, 1924)    iN  M T

Archonias seitzi (Lathy & Rosenberg, 1912)    iN

Archonias seitzi seitzi (Lathy & Rosenberg, 1912)  TL: Colombia  M T

Archonias seitzi blanca (Reissinger, 1972)  TL: Ecuador  M T

Archonias seitzi clarita (Bollino & Costa, 2007)  ref.171  M T

Archonias seitzi zana (F. Brown, 1939)  TL: Colombia  M T

Archonias sella (Eitschberger & T. Racheli, 1998)  TL: Peru

Archonias sella sella (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias sella dognini (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias semiramis (Lucas, 1852)    iN

Archonias semiramis semiramis (Lucas, 1852)  TL: Colombia  M T

Archonias semiramis belmira (Bollino & L. Rodríguez, 2003)  TL: Colombia  M T

Archonias semiramis costarum (Bollino & Lamas, 2004)  TL: Venezuela  M T

Archonias semiramis palla (F. Brown, 1939)  TL: Colombia  M T

Archonias semiramis salomon (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias semiramis willmotti (Bollino & Lamas, 2004)  TL: Ecuador  M T

Archonias sibyllae (Nakahara, Padrón & MacDonald, 2018)  TL: Panama

Archonias similis (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias sinapina (A. Butler, 1896)    iN

Archonias sinapina sinapina (A. Butler, 1896)  TL: Peru  M T

Archonias sinapina subflava (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias sisamnus (Fabricius, 1793)    iN

Archonias sisamnus sisamnus (Fabricius, 1793)  TL: ?  M T

Archonias sisamnus ayanganna (Bollino & Costa, 2007)  ref.171  M T

Archonias sisamnus bithyna (Röber, 1924)  TL: Colombia  M T L

Archonias sisamnus merida (Eitschberger & T. Racheli, 1998)  TL: Venezuela  M T

Archonias sisamnus smalli (Eitschberger & T. Racheli, 1998)  Honduras to Panama  M T S L

Archonias sisamnus telasco (Lucas, 1852)  TL: Peru  M T S Ma L

Archonias smithia (F. Brown & Gabriel, 1939)    iN  M T

Archonias socorrensis (Fassl, 1915)  TL: Colombia

Archonias socorrensis socorrensis (Fassl, 1915)  TL: Colombia  M T

Archonias socorrensis cotopaxiensis (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias striata (Eitschberger & T. Racheli, 1998)  TL: Peru

Archonias striata striata (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias striata batesi (Eitschberger & T. Racheli, 1998)  TL: Peru  M T

Archonias striata jimbura (Bollino & Vitale, 2002)  TL: Ecuador

Archonias striata [n. ssp.#1] (Bollino & Lamas, MS)  TL: Ecuador

Archonias striata [n. ssp.#2] (Bollino & Lamas, MS)  TL: Ecuador

Archonias suadela (Hopffer, 1874)  TL: Peru  M T S

Archonias suasa (Röber, 1908)    iN

Archonias suasa suasa (Röber, 1908)  TL: Bolivia  M T S

Archonias suasa pacis (Eitschberger & T. Racheli, 1998)  TL: Bolivia  M T

Archonias suasa [n. ssp.] (Lamas, MS)  Peru

Archonias superba (Lathy & Rosenberg, 1912)    iN  M T S

Archonias susiana (Hopffer, 1874)    iN

Archonias susiana susiana (Hopffer, 1874)  TL: Peru  M T

Archonias susiana acomayo (Reissinger, 1972)  TL: Peru  M T

Archonias susiana amba (F. Brown, Gabriel & Goodson, 1940)  TL: Ecuador  M T

Archonias susiana galbinea (Eitschberger & T. Racheli, 1998)  TL: Peru  M T S

Archonias susiana roeberi (Reissinger, 1972)  TL: Peru  M T

Archonias susiana sebundoia (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias susiana tamboensis (Joicey & Talbot, 1918)  TL: not specified  M T

Archonias tamsa (F. Brown & Gabriel, 1939)  TL: Peru

Archonias tamsa tamsa (F. Brown & Gabriel, 1939)  TL: Peru  M T

Archonias tamsa yanganza (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias teutamis (Hewitson, 1860)    iN

Archonias teutamis teutamis (Hewitson, 1860)  TL: Peru  M T

Archonias teutamis epimene (Hewitson, 1870)  TL: Ecuador  M L

Archonias teutamis [n. ssp.] (Lamas, MS)  Peru

Archonias teutila (E. Doubleday, 1847)  Pure-banded Dartwhite    iN

Archonias teutila teutila (E. Doubleday, 1847)  C Mexico (Jalisco to Veracruz)  M T S L Ha

Archonias teutila flavifaciata (Beutelspacher, 1986)  S Mexico (Oaxaca)  M T S L

Archonias teutila [n. ssp.#1] (Llorente, MS)  S Mexico (Guerrero)  M S

Archonias teutila [n. ssp.#2] (Llorente, MS)  S Mexico (Chiapas) to Guatemala (at least)  M S

Archonias teutila flavomaculata (Lathy & Rosenberg, 1912)  Costa Rica & Panama  M T S L

Archonias theresa (A. Butler & H. Druce, 1874)  Small Dartwhite  Costa Rica & Panama  iN  M T S L

Archonias thomasorum (Jasinski, 1998)  TL: Ecuador

Archonias thomasorum thomasorum (Jasinski, 1998)  TL: Ecuador

Archonias thomasorum inexpectata (Bollino & Vitale, 2004)  TL: Ecuador  M T

Archonias toca (E. Doubleday, 1847)    iN

Archonias toca toca (E. Doubleday, 1847)  TL: Bolivia  M T

Archonias toca detrita (Röber, 1909)  TL: Peru  M T

Archonias tomyris (C. Felder & R. Felder, 1865)    iN

Archonias tomyris tomyris (C. Felder & R. Felder, 1865)  TL: Colombia  M T Ma L

Archonias tomyris barbara (Reissinger, 1969)  TL: Peru  M T

Archonias tomyris myris (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias tomyris subtamina (Eitschberger & T. Racheli, 1998)  TL: Venezuela  M T

Archonias tomyris subtomyris (Reissinger, 1972)  TL: Peru  M T

Archonias tomyris tamina (Röber, 1909)  TL: Bolivia  M T

Archonias tricolor (A. Butler, 1897)    iN

Archonias tricolor flava (E. Krüger, 1925)  TL: Colombia  M T

Archonias tricolor tricolor (A. Butler, 1897)  TL: Ecuador  M T

Archonias tricolor rodriguezi (Eitschberger & T. Racheli, 1998)  TL: Colombia  M T

Archonias tricolor tomasi (Wojtusiak & Rey, 1999)  TL: Venezuela  M T

Archonias tricolor perija (Bollino & Costa, 2017)  TL: Venezuela

Archonias troezene (C. Felder & R. Felder, 1865)    iN

Archonias troezene troezene (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Archonias troezene gelba (F. Brown & Gabriel, 1939)  TL: Venezuela  M T

Archonias troezene miza (Bollino & Costa, 2007)  ref.171  M T

Archonias troezene troezenides (Röber, 1908)  TL: Colombia  M T

Archonias truncata (Lathy & Rosenberg, 1912)    iN

Archonias truncata truncata (Lathy & Rosenberg, 1912)  TL: Venezuela  M T

Archonias truncata xanthotaenia (Röber, 1924)  TL: Ecuador  M T

Archonias uricoecheae (C. Felder & R. Felder, 1861)    iN

Archonias uricoecheae uricoecheae (C. Felder & R. Felder, 1861)  TL: Colombia  M T S

Archonias uricoecheae inopa (Wojtusiak & Rey, 1999)  TL: Venezuela  M T

Archonias vilcabamba (Lamas & Bollino, 2004)  TL: Peru  M T

Archonias vulnerata (A. Butler, 1897)  TL: Ecuador

Archonias vulnerata vulnerata (A. Butler, 1897)  TL: Ecuador  M T

Archonias vulnerata koenigi (Eitschberger & T. Racheli, 1998)  TL: Peru  M S

Archonias watkinsi (Lathy & Rosenberg, 1912)    iN

Archonias watkinsi watkinsi (Lathy & Rosenberg, 1912)  TL: Peru  M T S

Archonias watkinsi huanuco (Reissinger, 1972)  TL: Peru  M T

Archonias watkinsi suffusa (Lathy & Rosenberg, 1912)  TL: Peru  M T

Archonias zamorana (Eitschberger & T. Racheli, 1998)  TL: Ecuador  M T

Archonias [n. sp.#1] (Lamas, MS)  TL: Peru

Archonias [n. sp.#2] (Lamas, MS)  TL: Peru

Archonias [n. sp.#3] (Llorente, MS)  TL: Mexico

Eucheira  Westwood, 1834  

Eucheira socialis Westwood, 1834  Social White    iN

Eucheira socialis socialis Westwood, 1834  C Mexico to Oaxaca  M T S

Eucheira socialis westwoodi Beutelspacher, 1984  NW Mexico  M T1 T2 S I H

Neophasia  Behr, 1869  

Neophasia menapia (C. Felder & R. Felder, 1859)  Pine White    iN

Neophasia menapia tehachapina J. Emmel, T. Emmel & Mattoon, 1998  Tehachapi Mts., Kern Co., CA  M T S

Neophasia menapia menapia (C. Felder & R. Felder, 1859)  CA Sierra Nevada N to N OR/ S WA (or BC if tau treated as syn.)  M T S L1 L2 I Ha

Neophasia menapia melanica Scott, 1981  immediate coast, Mendocino Co., CA  M S L

Neophasia menapia tau (Scudder, 1861)  BC, WA, ID, W MT, NE WY, blending in OR with typical menapia  M S L

Neophasia menapia magnamenapia Austin, 1998  W NV, S & C UT, AZ, NM, CO, E WY, W NE, W SD  M T S1 S2 L1 L2 L3 I1 I2 H Ha

Neophasia inopinatus Durden, 2008  TL: Mexico

Neophasia terlooii Behr, 1869  Mexican Pine White  SE AZ, SW NM to central western Mexico (MICH), also NL  iN  M T S Ab L1 L2 L3 I Ha

Pereute  Herrich-Schäffer, 1867  

Pereute antodyca (Boisduval, 1836)    iN  M T

Pereute callinice (C. Felder & R. Felder, 1861)    iN

Pereute callinice callinice (C. Felder & R. Felder, 1861)  TL: Colombia; Venezuela  M T

Pereute callinice numbalensis Joicey & Talbot, 1928  TL: Ecuador  M T

Pereute callinira Staudinger, 1884    iN

Pereute callinira callinira Staudinger, 1884  TL: Peru; Colombia  M T L

Pereute callinira sabrina Fruhstorfer, 1907  TL: Colombia  M T

Pereute charops (Boisduval, 1836)  Darkened White    iN

Pereute charops leonilae Llorente, 1986  NW Mexico (Sinaloa to Colima)  M T S L

Pereute charops sphocra Draudt, 1931  SW Mexico (Pacific slope of Guerrero & Oaxaca)  M T S Ha

Pereute charops charops (Boisduval, 1836)  E Mexico  M T S Ab L

Pereute charops nigricans Joicey & Talbot, 1928  S Mexico (Chiapas) to Nicaragua  M T S L

Pereute charops Costa Rican segregate ,  Costa Rica & Panama  M S L

Pereute charops columbica Fruhstorfer, 1907  TL: Colombia  M T

Pereute charops peruviana (Hopffer, 1878)  TL: Peru  M T

Pereute charops venezuelana (Hopffer, 1878)  TL: [Venezuela]  M T

Pereute charops [n. ssp.] Lamas, MS  Costa Rica

Pereute cheops Staudinger, 1884  Dark White  Costa Rica & Panama  iN  M T S L

Pereute leucodrosime (Kollar, 1850)    iN

Pereute leucodrosime leucodrosime (Kollar, 1850)  TL: Colombia  M T

Pereute leucodrosime bellatrix Fruhstorfer, 1907  TL: Peru  M T S

Pereute leucodrosime beryllina Fruhstorfer, 1907  TL: Ecuador  M T

Pereute leucodrosime caesarea (Lucas, 1852)  TL: [Ecuador]  M T

Pereute lindemannae Reissinger, 1970  TL: Brazil (Am)

Pereute lindemannae lindemannae Reissinger, 1970  TL: Brazil (Am)  M T

Pereute lindemannae pemona de Marmels, Clavijo & M. Chacín, 2003  TL: Venezuela  M T

Pereute lindemannae piaroa de Marmels, Clavijo & M. Chacín, 2003  TL: Venezuela  M T

Pereute swainsoni (G. Gray, 1832)    iN  M T

Pereute telthusa (Hewitson, 1860)    iN  M T

Leodonta  A. Butler, 1870  

Leodonta dysoni (E. Doubleday, 1847)  Dyson's White    iN

Leodonta dysoni dysoni (E. Doubleday, 1847)  Costa Rica to Venezuela  M T S L

Leodonta dysoni batzebina Röber, 1924  TL: Colombia  M T

Leodonta dysoni marginata Schaus, 1902  TL: Venezuela  M T

Leodonta dysoni [n. ssp.] Lamas, MS  Peru

Leodonta tagaste (C. Felder & R. Felder, 1859)    iN

Leodonta tagaste tagaste (C. Felder & R. Felder, 1859)  TL: Peru  M T

Leodonta tagaste colombiana Constantino & Salazar, 2004  TL: Colombia  M T

Leodonta tagaste peruana Bollino, 2010  TL: Peru  M S

Leodonta tagaste calderoni Bollino, 2010  TL: Peru

Leodonta tagaste extrema Bollino, 2010  TL: Bolivia

Leodonta tellane (Hewitson, 1860)    iN

Leodonta tellane tellane (Hewitson, 1860)  TL: Colombia  M T

Leodonta tellane chiriquensis (Staudinger, 1884)  Costa Rica & Panama  M T S L

Leodonta tellane intermedia Röber, 1908  TL: Colombia  M T

Leodonta tellane tingomariae Baumann & Reissinger, 1969  TL: Peru  M T

Leodonta tellane triangulata Constantino & Rodríguez, 2004  TL: Colombia

Leodonta tellane [n. ssp.#1] Constantino, MS  TL: Colombia

Leodonta tellane [n. ssp.#2] Lamas, MS  TL: Ecuador

Leodonta tellane [n. ssp.#3] Lamas, MS  TL: Peru

Leodonta tellane [n. ssp.#4] Lamas, MS  TL: Peru

Leodonta zenobia (C. Felder & R. Felder, 1865)    iN

Leodonta zenobia zenobia (C. Felder & R. Felder, 1865)  TL: Colombia  M T S

Leodonta zenobia monticola Joicey & Talbot, 1918  TL: Peru  M T S

Leodonta zenobia angustata Constantino & Fagua, 2004  TL: Colombia

Leodonta zenobia [n. ssp.#1] Lamas, MS  TL: Venezuela

Leodonta zenobia [n. ssp.#2] Lamas, MS  TL: Peru

Leodonta zenobina (Hopffer, 1869)    iN  M T

Melete  Swainson, [1831]  

Melete calymnia (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Melete leucadia (C. Felder & R. Felder, 1862)    iN

Melete leucadia leucadia (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Melete leucadia reyi Neild & Costa, 2017  ref.190

Melete leucanthe (C. Felder & R. Felder, 1861)    iN

Melete leucanthe leucanthe (C. Felder & R. Felder, 1861)  TL: Colombia; ["Ecuador"]  M T S L

Melete leucanthe veronica Le Crom & Llorente, 2004  TL: Colombia  M L

Melete lycimnia (Cramer, 1777)    iN

Melete lycimnia lycimnia (Cramer, 1777)  TL: Surinam  M T

Melete lycimnia aelia (C. Felder & R. Felder, 1861)  TL: Colombia; ["Ecuador"]  M T S L

Melete lycimnia chagris (Staudinger, 1876)  Panama  M T IL

Melete lycimnia eurymnia (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Melete lycimnia flippantha (Fabricius, 1793)  TL: ?  M T

Melete lycimnia harti (A. Butler, 1896)  TL: Trinidad  M T

Melete lycimnia isandra (Boisduval, 1836)  S TX (as stray), E & W Mexico to Costa Rica  M T S1 S2 S3 L I

Melete lycimnia latilimbata (A. Butler, 1896)  TL: Ecuador  M T

Melete lycimnia monstrosa (A. Butler, 1875)  Panama  M T S

Melete lycimnia napona (Röber, 1909)  TL: Ecuador  M T

Melete lycimnia narmia Fruhstorfer, 1910  TL: [Brazil (MT)]  M T

Melete lycimnia paulista Fruhstorfer, 1908  TL: Brazil (SP)  M T

Melete lycimnia peruviana (Lucas, 1852)  TL: Peru  M T

Melete lycimnia petronia Fruhstorfer, 1907  TL: Brazil (SC)  M T

Melete lycimnia phazania Fruhstorfer, 1907  TL: Brazil (Ba)  M T

Melete lycimnia reducta Constantino, Le Crom & Torres, 2004  TL: Colombia  M T

Melete lycimnia theodori Fruhstorfer, 1907  TL: Brazil (Am)  M T

Melete polyhymnia (C. Felder & R. Felder, 1865)    iN

Melete polyhymnia polyhymnia (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Melete polyhymnia caesarea Fruhstorfer, 1912  TL: Colombia  M T

Melete polyhymnia florinda (A. Butler, 1875)  Chiapas to Colombia  M T S L

Melete polyhymnia serrana R. G. Maza, 1984  N Oaxaca, Mexico (Gulf Slope)  M T

Melete salacia (Godart, 1819)  Black-striped White    iN

Melete salacia salacia (Godart, 1819)  Hispaniola  M T S

Melete salacia cubana Fruhstorfer, 1908  Cuba  M T S L

Glutophrissa  A. Butler, 1887  

Glutophrissa drusilla (Cramer, 1777)  Florida White    iN

Glutophrissa drusilla noroesta Grishin, 2022  S Texas to Central America; ref.16  M S1 S2 Ab L I H

Glutophrissa drusilla drusilla (Cramer, 1777)  ref.16  M T

Glutophrissa drusilla boydi (W. Comstock, 1943)  Hispaniola, Mona, Puerto Rico, Culebra, Virgin Islands  M T S

Glutophrissa drusilla castalia (Fabricius, 1793)  Jamaica  M T S

Glutophrissa drusilla comstocki (Dillon, 1947)  Dominica & Martinique  M T S

Glutophrissa drusilla monomorpha (A. Hall, 1936)  Grenada  M T

Glutophrissa drusilla neumoegenii (Skinner, 1894)  S Florida  M T S L

Glutophrissa drusilla poeyi (A. Butler, 1872)  Bahamas, Cuba, Isla de Juventud, Caymans  M T S

Glutophrissa drusilla sofaia Brévignon, 2020  ref.178

Glutophrissa drusilla tenuis (Lamas, 1981)  W Mexico & S TX (strays far N) to Peru  M T

Glutophrissa punctifera (R.F. d'Almeida, 1939)  Antillean White  Hispaniola, Puerto Rico, St. Thomas & St. John  iN  M S

  Subfamily Dismorphiinae

Pseudopieris  Godman & Salvin, 1889  

Pseudopieris nehemia (Boisduval, 1836)    iN

Pseudopieris nehemia irma Lamas, 1979  S Mexico (Chiapas) & Guatemala  M T

Pseudopieris nehemia francisca Lamas, 1979  Honduras  M T

Pseudopieris nehemia luisa Lamas, 1979  Panama to Ecuador  M S L

Pseudopieris nehemia aequatorialis (C. Felder & R. Felder, 1861)  TL: Ecuador  M T

Pseudopieris nehemia melania Lamas, 1985  TL: Peru

Pseudopieris nehemia jessica Lamas, 2004  TL: Peru  M L

Pseudopieris nehemia mariana Lamas, 2004  TL: Peru (SM, HU)

Pseudopieris nehemia limbalis Röber, 1924  TL: Brazil (Pa)  M T S

Pseudopieris nehemia nehemia (Boisduval, 1836)  TL: Brazil  M T

Pseudopieris nehemia prasina Hayward, 1949  TL: Argentina  M T

Pseudopieris nehemia [n. ssp.] Lamas, MS  Peru

Pseudopieris viridula (C. Felder & R. Felder, 1861)    iN

Pseudopieris viridula viridula (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Pseudopieris viridula mauritia Lamas, 2004  TL: Venezuela (Merida) - W. Colombia  M T

Pseudopieris viridula mimaripa de Marmels, Clavijo & M. Chacín, 2003  TL: Venezuela  M T

Pseudopieris viridula autana Costa & Viloria, 2021  TL: Venezuela

Pseudopieris viridula zulma Lamas, 2004  TL: S. Peru-Bolivia  M T S

Lieinix  G. Gray, 1832  

Lieinix christa (Reissinger, 1970)  TL: Brazil (Am)

Lieinix christa bousquetsi Costa, Neild, Viloria & Grishin, 2021  TL: Venezuela

Lieinix christa christa (Reissinger, 1970)  TL: Brazil (Am)  M T

Lieinix cinerascens (Salvin, 1871)  Bluish Mimic-White  El Salvador to Panama  iN  M T S

Lieinix lala (Godman & Salvin, 1889)  Dark Mimic-White  TL: Guatemala

Lieinix lala lala (Godman & Salvin, 1889)  C Chiapas, Mexico to Guatemala  M T S

Lieinix lala turrenti J. Maza & R. G. Maza, 1984  SW Chiapas, Mexico  M T S

Lieinix neblina J. Maza & R. G. Maza, 1984  Guerrero Mimic-White  SW Mexico (Guerrero and Oaxaca)  M T

Lieinix nemesis (Latreille, [1813])    iN

Lieinix nemesis nemesis (Latreille, [1813])  TL: [S. Amer.]  M T S Ma L1 L2

Lieinix nemesis atthis (E. Doubleday, 1842)  E Mexico to Panama  M T S1 S2 L1 L2

Lieinix nemesis nayaritensis Llorente, 1984  W Mexico (Nayarit to Oaxaca)  M T S L

Lieinix viridifascia (A. Butler, 1872)  Greenish Mimic-White  Meseta Central & Volcan Chiriqui, Costa Rica & Panama  iN  M S L

Dismorphia  Hübner, 1816  

Dismorphia altis Fassl, 1910    iN

Dismorphia altis altis Fassl, 1910  TL: Colombia  M T

Dismorphia altis sophia Lamas, Rodríguez & Llorente, 2004  TL: Colombia  M T

Dismorphia altis negreti Pyrcz & G. Rodríguez, 2004  TL: Colombia  M T S

Dismorphia amphione (Cramer, 1779)    iN

Dismorphia amphione amphione (Cramer, 1779)  TL: Surinam

Dismorphia amphione astynome (Dalman, 1823)  TL: Brazil  M T L

Dismorphia amphione beroe (Lucas, 1852)  Panama-Colombia  M T S

Dismorphia amphione bertha Lamas, 2004  TL: Peru (LO)

Dismorphia amphione broomeae A. Butler, 1899  TL: Trinidad  M T

Dismorphia amphione daguana Bargmann, 1929  TL: Colombia  M T

Dismorphia amphione discrepans A. Butler, 1896  TL: Colombia  M T

Dismorphia amphione egaena (H. Bates, 1861)  TL: Brazil (Am)  M T

Dismorphia amphione fusca Winhard, 2019  ref.360

Dismorphia amphione isolda Llorente, 1984  SW Mexico, Pacific slope of Guerrero & Oaxaca  M T S L

Dismorphia amphione lupita Lamas, 1979  NW Mexico (Nayarit, Jalisco, Colima)  M T S L

Dismorphia amphione meridionalis Röber, 1909  TL: Bolivia  M T

Dismorphia amphione mora Lamas, 2004  TL: Peru (MD)  M S

Dismorphia amphione praxinoe (E. Doubleday, 1844)  E Mexico to Colombia  M T S L

Dismorphia amphione rhomboidea A. Butler, 1896  TL: Peru  M T

Dismorphia arcadia (C. Felder & R. Felder, 1862)    iN

Dismorphia arcadia arcadia (C. Felder & R. Felder, 1862)  TL: Colombia  M T L

Dismorphia arcadia lucilla A. Butler, 1899  TL: Ecuador  M T S L

Dismorphia arcadia heloisa Lamas, 2004  TL: N. Peru  M L

Dismorphia arcadia medorina (Hewitson, 1875)  TL: Bolivia  M T

Dismorphia hippotas (Hewitson, 1875)  TL: Ecuador

Dismorphia hippotas diana Lamas, Llorente & G. Rodríguez, 2004  TL: Colombia  M T

Dismorphia hippotas hippotas (Hewitson, 1875)  TL: Ecuador  M T L

Dismorphia astyocha (Hübner, [1831])    iN  M T L

Dismorphia boliviana Forster, 1955  TL: Bolivia  M T S L

Dismorphia crisia (Drury, 1782)    iN

Dismorphia crisia crisia (Drury, 1782)  TL: Brazil (RJ)  M T L

Dismorphia crisia alvarezi J. Maza & R. G. Maza, 1984  SW Chiapas, Mexico, to Guatemala  M T S L

Dismorphia crisia foedora (Lucas, 1852)  TL: Venezuela  M Ma L

Dismorphia crisia interrupta Talbot, 1932  TL: "Peru"; Colombia  M L

Dismorphia crisia lubina A. Butler, 1872  Costa Rica & Panama  M T S1 S2 L I

Dismorphia crisia neblina Reissinger, 1970  TL: Venezuela  M T

Dismorphia crisia roraimae A. Hall, 1939  TL: Guyana  M T

Dismorphia crisia saltensis Breyer, 1939  TL: Argentina  M T

Dismorphia crisia steinhauseri J. Maza & R. G. Maza, 1984  El Salvador  M T S L

Dismorphia crisia tolimensis Fassl, 1915  TL: Colombia  M T L

Dismorphia crisia virgo (H. Bates, 1864)  SE Mexico & Guatemala  M T S

Dismorphia crisia anamaria Lamas, 2004  TL: N.W. Peru  M Ma L

Dismorphia crisia sylvia Lamas, 2004  TL: N.E. Peru  M T

Dismorphia cubana (Herrich-Schäffer, 1862)  Cuban Mimic-White  Cuba  iN  M T S L

Dismorphia eunoe (E. Doubleday, 1844)  Eunoe Mimic-White    iN

Dismorphia eunoe eunoe (E. Doubleday, 1844)  S Mexico; Atlantic slope of Veracruz & Oaxaca  M T S

Dismorphia eunoe chamula Llorente & Luis, 1988  S Mexico (Chiapas)  M T S

Dismorphia eunoe desine (Hewitson, 1869)  Nicaragua & Costa Rica to Panama  M T S1 S2 L I

Dismorphia eunoe popoluca Llorente & Luis, 1988  Los Tuxtlas region, Veracruz, Mexico  M T S

Dismorphia eunoe noelia Lamas, 2004  Darien, Panama  M OI L

Dismorphia eunoe [n. ssp.] Lamas, MS  Panama

Dismorphia hyposticta (C. Felder & R. Felder, 1861)    iN

Dismorphia hyposticta hyposticta (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Dismorphia hyposticta manuelita Fassl, 1910  TL: Colombia  M T S L

Dismorphia hyposticta paulina Lamas, 2004  TL: Peru  M T

Dismorphia hyposticta ophelia Lamas, 2004  TL: Peru

Dismorphia laja (Cramer, 1779)  TL: Surinam

Dismorphia laja laja (Cramer, 1779)  TL: Surinam  M S

Dismorphia laja carthesis (Hewitson, 1869)  TL: Guyana  M T

Dismorphia laja jurua Röber, 1924  TL: Brazil (Am)  M T

Dismorphia laja koenigi Baumann & Reissinger, 1969  TL: Peru  M T S

Dismorphia laja lysianax (Hewitson, [1860])  TL: [Peru]  M T

Dismorphia laja spectabilis Avinoff, 1926  TL: Bolivia  M T

Dismorphia laja tapajona (H. Bates, 1861)  TL: Brazil (Pa)  M T

Dismorphia laja tricolor Grose-Smith & W. F. Kirby, 1897  TL: [Colombia]  M T

Dismorphia laja rosina Lamas, 2004  TL: Peru  M T

Dismorphia lelex (Hewitson, 1869)    iN

Dismorphia lelex lelex (Hewitson, 1869)  TL: Ecuador  M T Ma L

Dismorphia lelex valeria Lamas, Llorente & Salazar, 2004  TL: Colombia  M S L

Dismorphia lelex xiomara Lamas, 2004  TL: Ecuador  M T L

Dismorphia lewyi (Lucas, 1852)    iN

Dismorphia lewyi lewyi (Lucas, 1852)  TL: Colombia  M S

Dismorphia lewyi boliviensis Röber, 1909  TL: Bolivia  M T

Dismorphia lewyi dolorita Fassl, 1910  TL: Colombia  M T Ma L

Dismorphia lewyi leonora (Hewitson, 1869)  TL: Ecuador  M T L

Dismorphia lewyi nasua (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Dismorphia lewyi rebecca Lamas, 2004  TL: Peru  M T S

Dismorphia lua (Hewitson, 1869)    iN

Dismorphia lua lua (Hewitson, 1869)  TL: Ecuador  M T

Dismorphia lua costaricensis (Schaus, 1913)  Costa Rica  M T

Dismorphia lua garleppi Staudinger, 1894  TL: Bolivia  M T

Dismorphia lua idae Fassl, 1910  Panama - W Colombia  M T S L

Dismorphia lua roberta Lamas, 2004  TL: Peru  M T

Dismorphia lycosura (Hewitson, [1860])  TL: Peru

Dismorphia lycosura lycosura (Hewitson, [1860])  TL: Peru  M T

Dismorphia lycosura [n. ssp.] Lamas, MS  Peru

Dismorphia lygdamis (Hewitson, 1869)    iN

Dismorphia lygdamis lygdamis (Hewitson, 1869)  TL: Ecuador  M T

Dismorphia lygdamis doris Baumann & Reissinger, 1969  TL: Peru  M T

Dismorphia lygdamis beatrix Lamas, MS  TL: Peru  M S L

Dismorphia lysis (Hewitson, 1869)    iN

Dismorphia lysis lysis (Hewitson, 1869)  TL: Ecuador  M T L

Dismorphia lysis peruana Röber, 1909  TL: Peru  M T S L

Dismorphia lysis mariella Lamas, 2004  TL: Peru

Dismorphia medora (E. Doubleday, 1844)    iN

Dismorphia medora dorisa Llorente & Le Crom, 2014  TL: Colombia

Dismorphia medora medora (E. Doubleday, 1844)  TL: Colombia  M T S Ma L

Dismorphia medora reducta Llorente & Andrade, 2004  TL: Colombia  M T

Dismorphia medora lilianna Lamas, 2004  TL: Ecuador  M T Ma L

Dismorphia medora juditha Lamas, 2004  TL: Peru

Dismorphia medora [n. ssp.] Lamas, MS  Peru

Dismorphia medorilla (Hewitson, 1877)  TL: Ecuador

Dismorphia medorilla medorilla (Hewitson, 1877)  TL: Ecuador  M T S

Dismorphia medorilla buchtieni Fassl, 1915  TL: Bolivia  M T

Dismorphia medorilla sarita Lamas, 2004  TL: Peru  M T S

Dismorphia melia (Godart, [1824])    iN  M T

Dismorphia mirandola (Hewitson, 1878)    iN

Dismorphia mirandola mirandola (Hewitson, 1878)  TL: Ecuador  M T

Dismorphia mirandola discoloria Weymer, 1891  TL: Colombia  M T S

Dismorphia mirandola [n. ssp.] Lamas, MS  Colombia

Dismorphia niepelti Weymer, 1909    iN  M T

Dismorphia pseudolewyi Forster, 1955  TL: Bolivia  M T

Dismorphia spio (Godart, 1819)  Hispaniolan Mimic-White  Hispaniola and Puerto Rico  iN  M T S1 S2 L

Dismorphia teresa (Hewitson, 1869)    iN  M T S

Dismorphia thermesia (Godart, 1819)    iN

Dismorphia thermesia minima Llorente, Andrade & Salazar, 2004  TL: Colombia

Dismorphia thermesia thermesia (Godart, 1819)  TL: Brazil  M S L

Dismorphia thermesia [n. ssp.#1] Lamas, MS  TL: Peru

Dismorphia thermesia [n. ssp.#2] Lamas, MS  TL: Guyana

Dismorphia thermesina (Hopffer, 1874)    iN

Dismorphia thermesina thermesina (Hopffer, 1874)  TL: [Peru]  M T S

Dismorphia thermesina pimpla (Hopffer, 1874)  TL: Bolivia  M T L

Dismorphia theucharila (E. Doubleday, 1848)    iN

Dismorphia theucharila theucharila (E. Doubleday, 1848)  TL: Venezuela  M T

Dismorphia theucharila argochloe (H. Bates, 1861)  TL: Brazil (Am)  M T

Dismorphia theucharila avonia (Hewitson, 1867)  TL: Ecuador  M T

Dismorphia theucharila fortunata (Lucas, 1854)  E Mexico to NW Colombia  M T S1 S2 L I

Dismorphia theucharila leuconoe (H. Bates, 1861)  TL: Brazil ([Am])  M T L

Dismorphia theucharila lysinoe (Hewitson, [1853])  TL: [Brazil (Am)]  M T

Dismorphia theucharila lysinoides Staudinger, 1884  TL: Colombia  M T

Dismorphia theucharila siloe (Hewitson, [1858])  Panama and Colombia  M T L

Dismorphia theucharila theonoe (Hewitson, [1853])  TL: [Brazil (Pa)]  M T

Dismorphia theucharila vitrea E. Krüger, 1925  TL: Surinam  M T

Dismorphia theucharila xanthone Röber, 1924  TL: [Colombia]  M T L

Dismorphia theucharila marta G. Rodríguez, Llorente & Salazar, 2004  TL: Colombia

Dismorphia theucharila yolanda Lamas, 2004  TL: Colombia  M T

Dismorphia zaela (Hewitson, [1858])    iN

Dismorphia zaela zaela (Hewitson, [1858])  TL: [Ecuador]  M T Ma L

Dismorphia zaela abilene (Hewitson, [1872])  TL: Ecuador  M T S

Dismorphia zaela oreas (Salvin, 1871)  Costa Rica & Panama  M T S L

Dismorphia zaela laura Lamas, Llorente & Salazar, 2004  TL: Colombia  M T

Dismorphia zathoe (Hewitson, [1858])    iN

Dismorphia zathoe zathoe (Hewitson, [1858])  TL: Colombia  M T L

Dismorphia zathoe core (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Dismorphia zathoe demeter Röber, 1909  TL: Colombia  M T

Dismorphia zathoe othoe (Hewitson, 1867)  TL: Colombia  M T

Dismorphia zathoe walteri Llorente & Le Crom, 2014  TL: Colombia

Dismorphia zathoe pallidula A. Butler & H. Druce, 1874  Costa Rica & Panama  M T S1 S2 L1 L2

Dismorphia zathoe blanca Costa & Attal, 2021  TL: Venezuela

Dismorphia zathoe hernandi Costa & Neild, 2021  TL: Venezuela

Dismorphia zathoe proserpina Grose-Smith & W. F. Kirby, 1897  TL: Guyana  M T

Dismorphia [n. sp.] Lamas, MS  not specified

Dismorphia [n. sp.] [n. ssp.#1] Lamas, MS  TL: Peru

Dismorphia [n. sp.] [n. ssp.#2] Lamas, MS  TL: Peru

Patia  Klots, 1933  

Patia orise (Boisduval, 1836)    iN

Patia orise denigrata (Rosenberg & Talbot, 1914)  TL: Peru  M T

Patia orise orise (Boisduval, 1836)  TL: French Guiana  M T S

Patia rhetes (Hewitson, [1857])  TL: Colombia  M T

Patia cordillera (C. Felder & R. Felder, 1862)    iN

Patia cordillera sororna (A. Butler, 1872)  Costa Rica to Colombia  M T S

Patia cordillera thecla (Bargmann, 1929)  TL: Colombia  M T

Patia cordillera cordillera (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Patia cordillera larunda (Hewitson, 1869)  TL: Ecuador  M T

Enantia  Hübner, [1819]  

Enantia lina (Herbst, 1792)    iN

Enantia lina virna Lamas, 2003  S Mexico (SW Chiapas) to SW Guatemala  M T S

Enantia lina marion Godman & Salvin, 1889  SE Mexico to Panama  M T S L

Enantia lina agatha Lamas, Llorente & Constantino, 2004  TL: Colombia  M T

Enantia lina mercenaria (C. Felder & R. Felder, 1861)  TL: Venezuela  M T L

Enantia lina lina (Herbst, 1792)  TL: America

Enantia lina acutipennis A. Butler, 1896  TL: Trinidad  M T

Enantia lina galanthis (H. Bates, 1861)  TL: [Brazil (Am)]  M T L

Enantia lina aphrodite (C. Felder & R. Felder, 1865)  TL: Brazil  M T L

Enantia lina psamathe (Fabricius, 1793)  TL: America  M T

Enantia lina versicolora (Fruhstorfer, 1912)  TL: Brazil (Pe)  M T

Enantia lina [n. ssp.] Lamas, MS  Ecuador

Enantia limnorina (C. Felder & R. Felder, 1865)    iN  M T L

Enantia citrinella (C. Felder & R. Felder, 1861)    iN

Enantia citrinella citrinella (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Enantia citrinella [n. ssp.#1] Lamas, MS  TL: Colombia

Enantia citrinella [n. ssp.#2] Lamas, MS  TL: Peru

Enantia citrinella [n. ssp.#3] Lamas, MS  TL: Peru

Enantia melite (Linnaeus, 1763)    iN

Enantia melite melite (Linnaeus, 1763)  TL: Indiis  M T L

Enantia melite linealis (Prüffer, 1922)  TL: Peru  M T S

Enantia melite theugenis (E. Doubleday, 1848)  TL: Bolivia  M T

Enantia melite vilma Lamas, 2004  TL: Peru  M T

Enantia albania (H. Bates, 1864)  Costa-spotted Mimic-White    iN

Enantia albania albania (H. Bates, 1864)  S TX (one-time stray), E Mexico to Guatemala & El Salvador  M T S1 S2 L

Enantia albania amalia (Staudinger, 1884)  Nicaragua to Panama  M T S L

Enantia albania zoraida Lamas, Llorente & Constantino, 2004  TL: Colombia  M T S

Enantia albania sonya Lamas, Llorente & Winhard, 2004  TL: Colombia  M T

Enantia albania nuria Lamas, 2004  TL: Ecuador

Enantia mazai Llorente, 1984  De la Maza's Mimic-White    iN

Enantia mazai mazai Llorente, 1984  E Mexico to Chiapas  M T1 T2 S L

Enantia mazai diazi Llorente, 1984  W Mexico  M T S1 S2 Ab L H

Enantia aloikea Brévignon, 1993  TL: French Guiana  M S

Enantia clarissa (Weymer, 1895)    iN  M T

Enantia jethys (Boisduval, 1836)  Jethys Mimic-White  E Mexico to El Salvador & Nicaragua  iN  M T S1 S2 L

Moschoneura  A. Butler, 1870  

Moschoneura pinthous (Linnaeus, 1758)    iN

Moschoneura pinthous ithomia (Hewitson, 1867)  ref.269  M T

Moschoneura pinthous monica Lamas, 2004  TL: Peru  M S L

Moschoneura pinthous patricia Lamas, 2004  TL: Peru

Moschoneura pinthous amelina (Hopffer, 1874)  TL: [Peru]  M T

Moschoneura pinthous carmenae Neild & Losada, 2021  TL: Venezuela

Moschoneura pinthous pinthous (Linnaeus, 1758)  TL: Indiis  M T L

Moschoneura pinthous proxima (Röber, 1924)  TL: Brazil (Am)  M T L

Moschoneura pinthous cyra (E. Doubleday, 1844)  TL: [Brazil (Ba)]  M T

Moschoneura methymna (Godart, 1819)  TL: not specified

Moschoneura methymna [n. ssp.#1] Lamas, MS  TL: Venezuela

Moschoneura methymna [n. ssp.#2] Lamas, MS  TL: Brazil (Pa)

Moschoneura methymna [n. ssp.#3] Lamas, MS  TL: Brazil (Pa)

Moschoneura ela (Hewitson, 1877)    iN

Moschoneura ela ela (Hewitson, 1877)  ref.241  M T

Moschoneura ela winhardi Llorente, Castro & Le Crom, 2016  ref.241

Family NYMPHALIDAE Rafinesque, 1815 – BRUSHFOOTS

  Subfamily Libytheinae

Libytheana  Michener, 1943  

Libytheana carinenta (Cramer, 1777)  American Snout    iN

Libytheana carinenta bachmanii (Kirtland, 1851)  E North America, [rarely Cuba]  M T S L

Libytheana carinenta larvata (Strecker, [1878])  C & S TX, NE Mexico, strays far N (regularly to to E CO)  M T S1 S2 S3 Ma L1 L2 L3 L4 I H

Libytheana carinenta mexicana Michener, 1943  W & S Mexico to Panama  M T S L

Libytheana carinenta carinenta (Cramer, 1777)  South America, [Panama?]  M S HS L

Libytheana fulvescens (Lathy, 1904)  Dominican Snout  Dominica  M T S

Libytheana motya (Hübner, [1823])  Cuban Snout  Cuba, [S TX as rare stray]  iN  M S

Libytheana terena (Godart, 1819)  Antillean Snout  Hispaniola, Jamaica, Puerto Rico, [Cuba as stray?]  iN  M T S

Prolibythea  Scudder, [1889]  a fossil genus

Prolibythea vagabunda Scudder, 1889  Fossil from Colorado

  Subfamily Danainae

  Tribe Danaini

Archaeolycorea  Martins, 1989  a fossil genus

Archaeolycorea ferreirai Martins, 1989  Fossil from Brazil (SP)

Anetia  Hübner, [1823]  

Anetia briarea (Godart, 1819)  Many-spotted King    iN

Anetia briarea briarea (Godart, 1819)  Hispaniola  M T S L

Anetia briarea numidia Hübner, [1823]  Cuba & Isla de Juventud  M S L

Anetia cubana (Salvin, 1869)  Cuban King  E Cuba  M T S L

Anetia jaegeri (Ménétriés, 1832)  Hispaniolan King  Hispaniola, [Jamaica?]  iN  M S L I

Anetia pantheratus (Martyn, 1797)  Great King    iN

Anetia pantheratus pantheratus (Martyn, 1797)  Hispaniola  M T S L

Anetia pantheratus clarescens (A. Hall, 1925)  Cuba  M T S

Anetia thirza (Geyer, [1833])  Cloud-forest King    iN

Anetia thirza thirza (Geyer, [1833])  C Mexico to El Salvador  M T S L

Anetia thirza insignis (Salvin, 1869)  Costa Rica & Panama  M T S L

Lycorea  E. Doubleday, [1847]  

Lycorea halia (Hübner, 1816)    iN

Lycorea halia halia (Hübner, 1816)  TL: Surinam  M T

Lycorea halia atergatis E. Doubleday, [1847]  continental; S TX (rare stray) to N South America, [St. Lucia]  M T S Ab L

Lycorea halia cinnamomea Weymer, 1883  TL: Brazil (Am)  M T

Lycorea halia cleobaea (Godart, 1819)  Hispaniola, [Jamaica, Puerto Rico]  M T S L

Lycorea halia demeter C. Felder & R. Felder, 1865  Cuba  M T S

Lycorea halia discreta Haensch, 1909  TL: Brazil ([MG])  M T

Lycorea halia fasciata Haensch, 1909  TL: [Ecuador]  M T

Lycorea halia pales C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T S

Lycorea halia transiens N. Riley, 1919  TL: Brazil (Am)  M T

Lycorea halia [n. ssp.#1] Lamas, MS  TL: Peru

Lycorea halia [n. ssp.#2] Lamas, MS  TL: Trinidad

Lycorea ilione (Cramer, 1775)    iN

Lycorea ilione ilione (Cramer, 1775)  TL: Brazil

Lycorea ilione albescens (Distant, 1876)  S Mexico to Panama  M S L

Lycorea ilione decolorata (Haensch, 1909)  TL: Ecuador  M T

Lycorea ilione lamirus (Latreille, [1817])  TL: ["Peru"]  M T

Lycorea ilione suffusca Winhard, 2019  ref.360

Lycorea ilione phenarete (E. Doubleday, [1847])  TL: Bolivia  M T

Lycorea pasinuntia (Stoll, 1780)    iN  M T

Danaus  Kluk, 1780  

Danaus cleophile (Godart, 1819)  Caribbean Queen  Jamaica & Hispaniola  iN  M T S1 S2

Danaus eresimus (Cramer, 1777)  Soldier    iN

Danaus eresimus eresimus (Cramer, 1777)  Grenada & Barbados, possibly only as vagrants from S  M T S

Danaus eresimus dilucida W. Forbes, 1939  TL: Brazil (Pa)  M T

Danaus eresimus erginus (Godman & Salvin, 1897)  TL: Peru  M T

Danaus eresimus estevana Talbot, 1943  TL: Venezuela  M T

Danaus eresimus montezuma Talbot, 1943  continental; S TX to Colombia (strays N)  M T S1 S2 L1 L2

Danaus eresimus plexaure (Godart, 1819)  TL: Brazil

Danaus eresimus tethys W. Forbes, [1944]  S Florida, Gr. Antilles (except P. Rico), Grand Cayman  M T S L

Danaus eresimus [n. ssp.] Lamas, MS  Peru

Danaus erippus (Cramer, 1775)    iN  M T

Danaus gilippus (Cramer, 1775)    iN

Danaus gilippus thersippus (H. Bates, 1863)  continental; SW USA & TX (strays far N) to Colombia  M T S1 S2 Ab Hy L1 L2 L3 L4 I H F

Danaus gilippus berenice (Cramer, 1779)  SE USA, Bahamas, Cuba, Caymans  M S L I

Danaus gilippus jamaicensis (H. Bates, 1864)  Jamaica  M T S

Danaus gilippus cleothera (Godart, 1819)  Hispaniola, Mona, rarely in W Puerto Rico  M T S

Danaus gilippus S Lesser Antilles segregate ,  [nr. xanthippus], S Lesser Antilles  M L

Danaus gilippus hermippus (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Danaus gilippus wheeleri Talbot, 1943  TL: Colombia  M T

Danaus gilippus candidus A. Clark, 1941  TL: Peru  M T

Danaus gilippus xanthippus (C. Felder & R. Felder, 1860)  TL: Brazil  M T

Danaus gilippus nivosus (Godman & Salvin, 1897)  TL: Peru  M T

Danaus gilippus gilippus (Cramer, 1775)  TL: Brazil (RJ)

Danaus gilippus [n. ssp.] Lamas, MS  Grenada

Danaus plexippus (Linnaeus, 1758)  Monarch    iN

Danaus plexippus plexippus (Linnaeus, 1758)  N & C Am., Cuba, N Baham. (to Crooked I.); migratory  M T S1 S2 Hy HS L1 L2 L3 L4 L5 I1 I2 H F Ha1 Ha2

Danaus plexippus leucogyne (A. Butler, 1884)  Virgin Islands, N Leeward Is.; possibly a form of D. p. megalippe  M T S

Danaus plexippus megalippe (Hübner, [1826])  E Cuba, S Bahamas, Gr. Antilles, Virgin Is., L. Antilles, Caymans  M S L I

Danaus plexippus nigrippus (Haensch, 1909)  TL: [Ecuador]  M T

Danaus plexippus portoricensis A. Clark, 1941  Puerto Rico; possibly only a form of D. p. megalippe  M T S

Danaus plexippus tobagi A. Clark, 1941  TL: Tobago  M T

  Tribe Ithomiini

Elzunia  Bryk, 1937  

Elzunia humboldt (Latreille, [1809])    iN

Elzunia humboldt humboldt (Latreille, [1809])  TL: ["Peru"]  M S L

Elzunia humboldt flavomaculata (Staudinger, 1885)  TL: Colombia  M T

Elzunia humboldt albomaculata (Haensch, 1903)  TL: Ecuador  M T

Elzunia humboldt cassandrina (Srnka, 1885)  TL: Ecuador  M T L

Elzunia humboldt judsoni R. Fox, 1956  TL: Peru  M T S

Elzunia humboldt microguttata (Röber, 1927)  TL: Colombia  M T S

Elzunia humboldt carlosi Le Crom & Llorente, 2018  TL: Colombia

Elzunia humboldt willmotti Le Crom, Andrade & Llorente, 2018  TL: Colombia

Elzunia bomplandii (Guérin-Méneville, [1844])  TL: Colombia

Elzunia bomplandii bomplandii (Guérin-Méneville, [1844])  TL: Colombia  M S L

Elzunia bomplandii atahualpa R. Fox, 1956  TL: Peru  M T S

Elzunia bomplandii descandollesi (Staudinger, 1885)  TL: Colombia  M T S

Elzunia bomplandii tamasea (Hewitson, 1873)  TL: Colombia  M T

Elzunia bomplandii latreillei (Staudinger, 1885)  TL: Colombia  M T S

Elzunia bomplandii golondrina Willmott, Lamas & J. Hall, 2021  TL: Ecuador

Elzunia regalis (Stichel, 1903)  TL: Colombia

Elzunia regalis regalis (Stichel, 1903)  TL: Colombia  M T

Elzunia regalis joiceyi (Kaye, 1918)  TL: Colombia  M T

Elzunia regalis purace Constantino & Salazar, 2007  TL: Colombia

Elzunia regalis quinduensis Constantino & Salazar, 2007  TL: Colombia

Elzunia pavonii (A. Butler, 1873)    iN  M T S L1 L2

Tithorea  E. Doubleday, 1847  

Tithorea harmonia (Cramer, 1777)    iN

Tithorea harmonia harmonia (Cramer, 1777)  TL: Surinam  M T S

Tithorea harmonia brunnea Haensch, 1905  TL: Peru  M T

Tithorea harmonia caissara (J. Zikán, 1941)  TL: Brazil (ES)  M T

Tithorea harmonia cuparina H. Bates, 1862  TL: Brazil (Pa)  M T

Tithorea harmonia dorada K. Brown, 1977  TL: Venezuela  M T S

Tithorea harmonia egaensis A. Butler, 1873  TL: Brazil (Am)  M T

Tithorea harmonia flacilla Godman & Salvin, 1898  TL: Colombia  M T

Tithorea harmonia furia Staudinger, [1884]  TL: Venezuela  M T

Tithorea harmonia furina Godman & Salvin, 1898  TL: Colombia  M T S

Tithorea harmonia gilberti K. Brown, 1977  TL: Peru  M T

Tithorea harmonia helicaon Godman & Salvin, 1879  Nicaragua to Panama  M T S

Tithorea harmonia hermias Godman & Salvin, 1898  TL: Ecuador  M T

Tithorea harmonia hermina Haensch, 1903  TL: Ecuador  M T

Tithorea harmonia hippothous Godman & Salvin, 1879  SE Mexico & Guatemala  M T S L

Tithorea harmonia irene (Drury, 1782)  Panama & Colombia  M T S

Tithorea harmonia lateflava (Haensch, 1909)  TL: Bolivia  M T

Tithorea harmonia manabiana R. Fox, 1956  TL: Ecuador  M T

Tithorea harmonia martina R. Fox, 1956  TL: Peru  M T L

Tithorea harmonia megara (Godart, 1819)  TL: Antilles  M T

Tithorea harmonia melanina Haensch, 1905  TL: Peru  M T

Tithorea harmonia neitha Hopffer, 1874  TL: Peru  M T

Tithorea harmonia pseudethra A. Butler, 1873  TL: Brazil  M T

Tithorea harmonia pseudonyma Staudinger, 1894  TL: Bolivia  M T

Tithorea harmonia salvadoris Staudinger, 1885  S Chiapas to Honduras & El Salvador  M T S

Tithorea harmonia sulphurata (J. Zikán, 1941)  TL: Brazil (Am)  M T

Tithorea harmonia [n. ssp.#1] Lamas, MS  TL: Colombia

Tithorea harmonia [n. ssp.#2] K. Brown, MS  TL: Venezuela

Tithorea harmonia [n. ssp.#3] K. Brown, MS  TL: Brazil (Pe)

Tithorea harmonia [n. ssp.#4] K. Brown, MS  TL: Brazil (Pa)

Tithorea harmonia [n. ssp.#5] K. Brown, MS  TL: Brazil (Pa)

Tithorea harmonia [n. ssp.#6] K. Brown, MS  TL: Guyana

Tithorea tarricina Hewitson, [1858]    iN

Tithorea tarricina tarricina Hewitson, [1858]  TL: Colombia  M T

Tithorea tarricina bonita Haensch, 1903  TL: Ecuador  M T

Tithorea tarricina duenna H. Bates, 1864  SE Mexico & Guatemala  M T S L

Tithorea tarricina franciscoi K. Brown, 1977  TL: Venezuela  M T S

Tithorea tarricina hecalesina C. Felder & R. Felder, 1865  TL: Colombia  M T

Tithorea tarricina lecromi Vitale & Rodríguez, 2004  ref.298

Tithorea tarricina parola Godman & Salvin, 1898  TL: Colombia  M T

Tithorea tarricina pinthias Godman & Salvin, 1878  Honduras to Panama  M T S L

Tithorea tarricina rafaeli Neild, 2008  ref.266

Tithorea tarricina tagarma Hewitson, 1874  TL: Bolivia  M T L

Tithorea pacifica Willmott & Lamas, 2004  TL: Ecuador

Tithorea pacifica pacifica Willmott & Lamas, 2004  TL: Ecuador

Tithorea pacifica concordia Willmott & Lamas, 2004  E Panama  M T S

Tithorea pacifica euphonia Willmott & Lamas, 2004  TL: Colombia  M T

Aeria  Hübner, 1816  

Aeria elara (Hewitson, 1855)    iN

Aeria elara elara (Hewitson, 1855)  TL: Brazil  M T

Aeria elara elarina (Oberthür, 1879)  TL: Brazil  M T S

Aeria elara [n. ssp.#1] Lamas, MS  TL: Peru

Aeria elara [n. ssp.#2] K. Brown, MS  TL: Brazil (SP)

Aeria elara [n. ssp.#3] K. Brown, MS  TL: Colombia

Aeria eurimedia (Cramer, 1777)    iN

Aeria eurimedia Costa Rican segregate ,  Costa Rica & Panama  M L

Aeria eurimedia eurimedia (Cramer, 1777)  TL: Surinam

Aeria eurimedia agna Godman & Salvin, 1879  Panama to NW Venezuela  M T S L

Aeria eurimedia latistriga M. Hering, 1925  TL: Colombia

Aeria eurimedia negricola (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Aeria eurimedia pacifica Godman & Salvin, 1879  SE Mexico to Honduras  M T S1 S2 L

Aeria eurimedia sisenna Weymer, 1899  TL: Bolivia  M T

Aeria eurimedia [n. ssp.] Lamas, MS  Costa Rica

Aeria olena Weymer, 1875    iN

Aeria olena olena Weymer, 1875  TL: Brazil  M T

Aeria olena [n. ssp.#1] K. Brown, MS  TL: Brazil (MT)

Aeria olena [n. ssp.#2] K. Brown, MS  TL: Brazil (Pe)

Athesis  E. Doubleday, 1847  

Athesis acrisione Hewitson, 1869    iN

Athesis acrisione acrisione Hewitson, 1869  TL: Ecuador  M T S

Athesis acrisione deflavata Niepelt, 1928  TL: Colombia  M T

Athesis clearista E. Doubleday, 1847    iN

Athesis clearista clearista E. Doubleday, 1847  TL: Venezuela  M T S L

Athesis clearista colombiensis Kaye, 1918  TL: Colombia  M T

Athesis clearista [n. ssp.] Lamas, MS  Colombia

Athesis vitrala Kaye, 1918    iN  M T

Eutresis  E. Doubleday, 1847  

Eutresis dilucida Staudinger, 1885  Smoky Tigerwing  Costa Rica & W Panama  M T S

Eutresis hypereia E. Doubleday, 1847    iN

Eutresis hypereia theope Godman & Salvin, 1877  Costa Rica & Panama  M T OI S L

Eutresis hypereia Panamanian segregate ,  Panama  M S

Eutresis hypereia hypereia E. Doubleday, 1847  TL: Venezuela  M T S

Eutresis hypereia antioquensis Staudinger, 1885  TL: Colombia  M T

Eutresis hypereia banosana R. Fox, 1956  TL: Ecuador  M T

Eutresis hypereia hyspa Godman & Salvin, 1879  TL: Ecuador  M T

Eutresis hypereia imeriensis K. Brown, 1977  TL: Brazil (Am)

Eutresis hypereia imitatrix Staudinger, 1876  TL: Peru  M T

Eutresis hypereia [n. ssp.] Lamas, MS  Panama

Athyrtis  C. Felder & R. Felder, 1862  

Athyrtis mechanitis C. Felder & R. Felder, 1862    iN

Athyrtis mechanitis mechanitis C. Felder & R. Felder, 1862  TL: Colombia  M T

Athyrtis mechanitis oberthueri Srnka, 1885  TL: Ecuador  M T

Athyrtis mechanitis salvini Srnka, 1884  TL: Peru  M T

Paititia  Lamas, 1979  

Paititia neglecta Lamas, 1979  TL: Peru  M T

Olyras  E. Doubleday, 1847  

Olyras crathis E. Doubleday, 1847    iN

Olyras crathis staudingeri Godman & Salvin, 1897  Costa Rica & Panama  M T S

Olyras crathis Panamanian segregate ,  Panama  M S

Olyras crathis crathis E. Doubleday, 1847  TL: Venezuela  M T S

Olyras crathis browni Neild, 2008  ref.266

Olyras crathis montagui A. Butler, 1870  TL: Colombia  M T

Olyras crathis sticheli Haensch, 1905  TL: Peru  M T

Olyras crathis weeksi R. Fox, 1956  TL: Ecuador  M T

Olyras crathis [n. ssp.#1] Lamas, MS  TL: Panama

Olyras crathis [n. ssp.#2] Lamas, MS  TL: Colombia

Olyras insignis Salvin, 1869  Insignis Tigerwing    iN

Olyras insignis insignis Salvin, 1869  Costa Rica & Panama  M T S

Olyras insignis praestans Godman & Salvin, 1897  TL: Colombia  M T

Olyras insignis translucens Hewitson, 1872  TL: Ecuador  M T

Olyras theon H. Bates, 1866  Rusty Tigerwing  SE Mexico, Guatemala  iN  M T S L

Patricia  R. Fox, 1940  

Patricia demylus (Godman & Salvin, 1879)    iN

Patricia demylus demylus (Godman & Salvin, 1879)  TL: Ecuador  M T S

Patricia demylus gemellus R. Fox, 1960  TL: Bolivia  M T S

Patricia demylus nebulosa Lamas, 1979  TL: Peru  M T

Patricia demylus [n. ssp.] Lamas, MS  Peru

Patricia dercyllidas (Hewitson, 1864)    iN

Patricia dercyllidas dercyllidas (Hewitson, 1864)  TL: Colombia  M T S

Patricia dercyllidas hazelea R. Fox, 1956  TL: Ecuador  M T S

Patricia dercyllidas [n. ssp.#1] Lamas, MS  TL: Peru

Patricia dercyllidas [n. ssp.#2] Lamas, MS  TL: Peru

Patricia dercyllidas [n. ssp.#3] Lamas, MS  TL: Peru

Patricia hewitsonii (Srnka, 1885)    iN  M T

Patricia oligyrtis (Hewitson, 1877)  TL: Ecuador

Patricia oligyrtis oligyrtis (Hewitson, 1877)  TL: Ecuador  M T S

Patricia oligyrtis [n. ssp.] Lamas, MS  Peru

Melinaea  Hübner, 1816  

Melinaea ethra (Godart, 1819)    iN  M T S

Melinaea idae (C. Felder & R. Felder, 1862)  Idae Tigerwing    iN

Melinaea idae idae (C. Felder & R. Felder, 1862)  Panama, Colombia  M T S L

Melinaea idae vespertina R. Fox, 1945  TL: Ecuador  M T

Melinaea isocomma W. Forbes, 1948    iN

Melinaea isocomma isocomma W. Forbes, 1948  TL: Colombia

Melinaea isocomma simulator R. Fox, 1960  TL: Ecuador  M T

Melinaea isocomma [n. ssp.] Lamas, MS  Peru

Melinaea lilis (E. Doubleday, 1847)    iN

Melinaea lilis lilis (E. Doubleday, 1847)  TL: Venezuela  M T

Melinaea lilis dodona Hopffer, 1874  TL: Bolivia  M T

Melinaea lilis ezra R. Fox, 1939  TL: Colombia  M T S

Melinaea lilis flavicans C. Hoffmann, 1924  W Mexico (NAY to GRO)  M T S

Melinaea lilis imitata H. Bates, 1864  E & S Mexico to Costa Rica (Atlantic slope)  M T S L

Melinaea lilis kayei K. Brown, 1977  TL: Guyana  M T

Melinaea lilis lateapicalis A. Hall, 1935  TL: Venezuela  M T S

Melinaea lilis messatis (Hewitson, [1856])  Panama, Colombia  M T OI

Melinaea lilis parallelis A. Butler, 1873  SE Costa Rica & Panama (Atlantic slope)  M T S L

Melinaea lilis sola Kaye, 1925  TL: Trinidad  M T

Melinaea scylax Salvin, 1871  Scylax Tigerwing  Costa Rica to Colombia (Pacific slope)  iN  M T S L

Melinaea ludovica (Cramer, 1780)    iN

Melinaea ludovica ludovica (Cramer, 1780)  TL: Surinam  M T S

Melinaea ludovica manuelito Tessmann, 1928  TL: Peru  M T

Melinaea ludovica paraiya Reakirt, 1866  TL: Brazil (RJ/SC)

Melinaea marsaeus (Hewitson, 1860)    iN

Melinaea marsaeus marsaeus (Hewitson, 1860)  TL: Brazil ([Am])  M T S

Melinaea marsaeus clara Rosenberg & Talbot, 1914  TL: Peru  M T S

Melinaea marsaeus macaria Godman & Salvin, 1898  TL: Colombia  M T

Melinaea marsaeus messenina C. Felder & R. Felder, 1865  TL: Colombia  M T S

Melinaea marsaeus mothone (Hewitson, 1860)  TL: Colombia  M T

Melinaea marsaeus phasiana A. Butler, 1870  TL: Peru  M T

Melinaea marsaeus pothete R.F. d'Almeida, 1945  TL: Brazil (Rd)  M T S

Melinaea marsaeus rileyi R. Fox, 1942  TL: Peru  M T

Melinaea marsaeus [n. ssp.] Lamas, MS  Peru

Melinaea menophilus (Hewitson, [1856])    iN

Melinaea menophilus menophilus (Hewitson, [1856])  TL: Colombia  M T S

Melinaea menophilus cocana Haensch, 1903  TL: Ecuador  M T

Melinaea menophilus ernestoi K. Brown, 1977  TL: Colombia  M T

Melinaea menophilus hicetas Godman & Salvin, 1879  TL: Peru  M T

Melinaea menophilus maenius (Hewitson, 1860)  TL: Brazil ([Am])  M T

Melinaea menophilus mediatrix Weymer, 1891  TL: French Guiana  M T S

Melinaea menophilus orestes Salvin, 1871  TL: Peru  M T S

Melinaea menophilus zaneka A. Butler, 1870  TL: Ecuador  M T

Melinaea menophilus [n. ssp.#1] Lamas, MS  TL: Peru

Melinaea menophilus [n. ssp.#2] Lamas, MS  TL: Peru

Melinaea menophilus [n. ssp.#3] Lamas, MS  TL: Brazil (Rd)

Melinaea menophilus [n. ssp.#4] K. Brown, MS  TL: Brazil (Rd)

Melinaea mnasias (Hewitson, [1856])  TL: [Brazil (Pa)]

Melinaea mnasias mnasias (Hewitson, [1856])  TL: [Brazil (Pa)]  M T S

Melinaea mnasias abitagua K. Brown, 1977  TL: Ecuador  M T

Melinaea mnasias comma W. Forbes, 1927  TL: Peru

Melinaea mnasias eratosthenes A. Hall, 1935  TL: French Guiana  M T S

Melinaea mnasias lucifer H. Bates, 1862  TL: Brazil (Am)  M T

Melinaea mnasias lutzi R. Fox, 1942  TL: Peru  M T

Melinaea mnasias neblinae K. Brown, 1977  TL: Brazil (Am)  M T

Melinaea mnasias romualdo R. Fox, 1965  TL: Peru  M T

Melinaea mnasias rondonia K. Brown, 1977  TL: Brazil (Rd)  M T

Melinaea mnasias tecta Haensch, 1909  TL: Guyana  M T

Melinaea mnasias thera C. Felder & R. Felder, 1865  TL: ?  M T

Melinaea mnasias [n. ssp.#1] Lamas, MS  TL: Peru

Melinaea mnasias [n. ssp.#2] K. Brown, MS  TL: Venezuela

Melinaea mnasias [n. ssp.#3] K. Brown, MS  TL: Brazil (Pa)

Melinaea mneme (Linnaeus, 1763)    iN

Melinaea mneme mneme (Linnaeus, 1763)  TL: China  M T

Melinaea mneme mauensis Weymer, 1891  TL: Brazil (Am)  M T S

Melinaea mneme [n. ssp.#1] K. Brown, MS  TL: Venezuela

Melinaea mneme [n. ssp.#2] K. Brown, MS  TL: Brazil (Rd)

Melinaea mnemopsis Berg, 1897  TL: Peru

Melinaea mnemopsis mnemopsis Berg, 1897  TL: Peru  M T

Melinaea mnemopsis [n. ssp.] K. Brown, MS  Brazil (Rd)

Melinaea satevis (E. Doubleday, 1847)    iN

Melinaea satevis satevis (E. Doubleday, 1847)  TL: Bolivia  M T

Melinaea satevis aurantia W. Forbes, 1942  TL: Venezuela  M T S

Melinaea satevis crameri Godman & Salvin, 1898  TL: Guyana  M T S

Melinaea satevis cydon Godman & Salvin, 1879  TL: Brazil (Am); Peru  M T

Melinaea satevis flavomacula Weymer, 1894  TL: Brazil (Am)  M T

Melinaea satevis lamasi K. Brown, 1977  TL: Peru  M T S

Melinaea satevis maelus (Hewitson, 1860)  TL: Brazil ([Am])  M T

Melinaea satevis maeonis Hewitson, 1869  TL: Ecuador  M T

Melinaea satevis tarapotensis Haensch, 1909  TL: Peru  M T

Melinaea satevis zamora Haensch, 1909  TL: Peru  M T S

Methona  E. Doubleday, 1847  

Methona confusa A. Butler, 1873    iN

Methona confusa Panamanian segregate ,  Panama  M S L

Methona confusa confusa A. Butler, 1873  TL: Brazil (Pa)  M T S L

Methona confusa psamathe Godman & Salvin, 1898  TL: Ecuador  M T S

Methona confusa [n. ssp.#1] Lamas, MS  TL: Panama

Methona confusa [n. ssp.#2] Lamas, MS  TL: Panama

Methona curvifascia Weymer, 1883    iN  M T S

Methona grandior (W. Forbes, 1944)    iN

Methona grandior grandior (W. Forbes, 1944)  TL: French Guiana

Methona grandior incana (W. Forbes, 1944)  TL: Ecuador  M T

Methona grandior batesi Lamas & Willmott, 2021  ref.318

Methona maxima (W. Forbes, 1944)    iN

Methona maxima maxima (W. Forbes, 1944)  TL: Bolivia  M T

Methona maxima nigerrima (W. Forbes, 1944)  TL: Peru

Methona megisto C. Felder & R. Felder, 1860    iN  M T S

Methona singularis (Staudinger, [1884])    iN  M T S

Methona themisto (Hübner, 1818)    iN

Methona themisto themisto (Hübner, 1818)  TL: Brazil  M T S

Methona themisto [n. ssp.] Lamas, MS  Brazil (SP)

Thyridia  Hübner, 1816  

Thyridia psidii (Linnaeus, 1758)    iN

Thyridia psidii melantho H. Bates, 1866  S Mexico (Chiapas) to Panama  M T S L

Thyridia psidii aedesia E. Doubleday, 1847  TL: Venezuela  M T S

Thyridia psidii psidii (Linnaeus, 1758)  TL: America [Guianas]  M OD TI IL Txt

Thyridia psidii ino C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T S

Thyridia psidii cetoides (Rosenberg & Talbot, 1914)  TL: Peru  M T S

Thyridia psidii pallida Godman & Salvin, 1898  TL: Brazil (MT)  M T S

Thyridia psidii hippodamia (Fabricius, 1775)  TL: [Brazil]  M T

Scada  W. F. Kirby, 1871  

Scada karschina (Herbst, 1792)    iN

Scada karschina karschina (Herbst, 1792)  TL: Brazil (RJ)  M T

Scada karschina delicata Talbot, 1932  TL: Brazil?  M T

Scada kusa (Hewitson, 1872)    iN  M T

Scada reckia (Hübner, [1808])    iN

Scada reckia reckia (Hübner, [1808])  TL: ?

Scada reckia bernhardi Neild, 2008  ref.266

Scada reckia ethica (Hewitson, [1861])  TL: Ecuador  M T S

Scada reckia huascara R. Fox, 1967  TL: Colombia

Scada reckia junina Bryk, 1953  TL: Peru  M T

Scada reckia labyrintha Lamas, 1985  TL: Peru  M T S

Scada reckia majuscula Haensch, 1905  TL: Guyana  M T

Scada reckia ortygia (H. Druce, 1876)  TL: Peru  M T

Scada reckia philemon (C. Felder & R. Felder, 1865)  TL: Venezuela?  M T

Scada reckia theaphia (H. Bates, 1862)  TL: Brazil (Pa)  M T

Scada reckia ruiza Lamas & Willmott, 2021  ref.318

Scada reckia [n. ssp.#1] K. Brown, MS  TL: Brazil (Am)

Scada reckia [n. ssp.#2] K. Brown, MS  TL: Brazil (Rd)

Scada reckia [n. ssp.#3] K. Brown, MS  TL: Brazil (Pa)

Scada zemira (Hewitson, 1856)    iN  M T L

Scada zibia (Hewitson, 1856)    iN

Scada zibia zibia (Hewitson, 1856)  TL: Colombia  M T S

Scada zibia batesi Haensch, 1903  TL: Ecuador  M T S

Scada zibia perpuncta Kaye, 1918  TL: Colombia  M T

Scada zibia quotidiana Haensch, 1909  TL: Ecuador  M T

Scada zibia xanthina (H. Bates, 1866)  Nicaragua to Panama  M T S L

Scada zibia zeroca R. Fox, 1967  TL: Ecuador  M T L

Sais  Hübner, 1816  

Sais browni Takahashi, 1977  TL: Colombia

Sais browni browni Takahashi, 1977  TL: Colombia

Sais browni demarmelsi Neild, 2008  ref.266

Sais rosalia (Cramer, 1779)    iN

Sais rosalia rosalia (Cramer, 1779)  TL: Surinam  M T

Sais rosalia badia Haensch, 1905  TL: Upper Amazon  M T

Sais rosalia brasiliensis Talbot, 1928  TL: Brazil (MT)  M T

Sais rosalia camariensis Haensch, 1905  TL: Guyana  M T S

Sais rosalia klagesi Avinoff, 1926  TL: Brazil (Am)  M T

Sais rosalia mosella (Hewitson, 1867)  TL: Venezuela  M T

Sais rosalia promissa Weymer, 1883  TL: Brazil (Am)  M T

Sais rosalia rosalinde Weymer, 1890  TL: Brazil (Pa)  M T

Sais rosalia schatzi J. Zikán, 1941  TL: Brazil (Am)  M T S

Sais rosalia virchowii Dewitz, 1877  TL: Venezuela  M T S

Sais rosalia zitella (Hewitson, 1868)  TL: Peru  M T

Sais rosalia [n. ssp.#1] Lamas, MS  TL: Colombia

Sais rosalia [n. ssp.#2] Lamas, MS  TL: Venezuela

Sais rosalia [n. ssp.#3] Lamas, MS  TL: Guyana

Sais rosalia [n. ssp.#4] K. Brown, MS  TL: Brazil

Sais rosalia [n. ssp.#5] K. Brown, MS  TL: Brazil

Forbestra  R. Fox, 1967  

Forbestra equicola (Cramer, 1780)    iN

Forbestra equicola equicola (Cramer, 1780)  TL: Surinam  M T

Forbestra equicola equicoloides (Godman & Salvin, 1898)  TL: Ecuador; Peru; Brazil (Am)  M T S

Forbestra equicola [n. ssp.] K. Brown, MS  Brazil (Rd)

Forbestra olivencia (H. Bates, 1862)    iN

Forbestra olivencia olivencia (H. Bates, 1862)  TL: Brazil (Am)  M T S

Forbestra olivencia aeneola R. Fox, 1967  TL: Peru  M T S

Forbestra olivencia juntana (Haensch, 1903)  TL: Ecuador  M T

Forbestra olivencia oiticicai (R.F. d'Almeida, 1951)  TL: Brazil (Pa)  M T S

Forbestra olivencia truncata (A. Butler, 1877)  TL: Brazil (Am)  M T

Forbestra olivencia [n. ssp.] Lamas, MS  Peru

Forbestra olivencia boliviana Winhard, 2020  ref.361

Forbestra proceris (Weymer, 1883)    iN  M T

Mechanitis  Fabricius, 1807  

Mechanitis lysimnia (Fabricius, 1793)    iN

Mechanitis lysimnia lysimnia (Fabricius, 1793)  TL: ?  M T

Mechanitis lysimnia bipuncta W. Forbes, 1948  TL: Venezuela

Mechanitis lysimnia elisa (Guérin-Méneville, [1844])  TL: Bolivia  M T

Mechanitis lysimnia labotas Distant, 1876  Costa Rica  M T S L

Mechanitis lysimnia limnaea W. Forbes, 1930  TL: French Guiana  M T

Mechanitis lysimnia macrinus Hewitson, 1860  Panama & Colombia  M T S L

Mechanitis lysimnia menecles Hewitson, 1860  TL: Brazil (Am)  M T

Mechanitis lysimnia nesaea Hübner, [1820]  TL: ?  M T

Mechanitis lysimnia neukircheni Neild, 2008  TL: Venezuela

Mechanitis lysimnia ocona H. Druce, 1876  TL: Peru  M T S

Mechanitis lysimnia roqueensis Bryk, 1953  TL: Peru  M T

Mechanitis lysimnia solaria W. Forbes, 1948  TL: Venezuela

Mechanitis lysimnia utemaia Reakirt, 1866  E & SW Mexico to Costa Rica  M T S L

Mechanitis lysimnia [n. ssp.#1] K. Brown, MS  TL: Colombia

Mechanitis lysimnia [n. ssp.#2] K. Brown, MS  TL: Colombia

Mechanitis lysimnia [n. ssp.#3] Lamas, MS  TL: Peru

Mechanitis mazaeus Hewitson, 1860    iN

Mechanitis mazaeus mazaeus Hewitson, 1860  TL: Brazil (Am)  M T S

Mechanitis mazaeus beebei W. Forbes, 1948  TL: Venezuela

Mechanitis mazaeus egaensis H. Bates, 1862  TL: Brazil (Am)  M T

Mechanitis mazaeus fallax A. Butler, 1873  TL: Colombia  M T

Mechanitis mazaeus lanei R. Fox, 1967  TL: Brazil (Pa)  M T

Mechanitis mazaeus pannifera A. Butler, 1877  TL: Brazil (Am/Pa)  M T

Mechanitis mazaeus pothetoides R.F. d'Almeida, 1951  TL: Brazil (Rd)  M T S

Mechanitis mazaeus visenda A. Butler, 1877  TL: Brazil (Pa)  M T S

Mechanitis mazaeus [n. ssp.#1] K. Brown, MS  TL: Ecuador

Mechanitis mazaeus [n. ssp.#2] K. Brown, MS  TL: Brazil (Rd)

Mechanitis messenoides C. Felder & R. Felder, 1865    iN

Mechanitis messenoides messenoides C. Felder & R. Felder, 1865  TL: Colombia  M T S

Mechanitis messenoides holmgreni Bryk, 1953  TL: Peru  M T S

Mechanitis messenoides deceptus A. Butler, 1873  TL: Ecuador  M T S

Mechanitis menapis Hewitson, [1856]    iN

Mechanitis menapis menapis Hewitson, [1856]  TL: Colombia  M T S

Mechanitis menapis caribensis R. Fox, 1967  TL: Venezuela  M T

Mechanitis menapis dariensis K. Brown, 1977  Panama  M S

Mechanitis menapis doryssus H. Bates, 1864  SE Mexico & Guatemala  M T S L I

Mechanitis menapis mantineus Hewitson, 1869  TL: Ecuador  M T

Mechanitis menapis occasiva R. Fox, 1967  TL: Colombia  M T

Mechanitis menapis saturata Godman, 1901  Honduras to Costa Rica  M T S L I

Mechanitis menapis [n. ssp.] Lamas, MS  Colombia

Mechanitis polymnia (Linnaeus, 1758)    iN

Mechanitis polymnia polymnia (Linnaeus, 1758)  TL: Surinam  M T S HS

Mechanitis polymnia angustifascia Talbot, 1928  TL: Brazil (MT)  M T

Mechanitis polymnia apicenotata J. Zikán, 1941  TL: Brazil (Am)  M T

Mechanitis polymnia bolivarensis R. Fox, 1967  TL: Venezuela  M T S

Mechanitis polymnia casabranca Haensch, 1905  TL: Brazil (MG)  M T

Mechanitis polymnia caucaensis Haensch, 1909  TL: Colombia  M T

Mechanitis polymnia chimborazona H. Bates, 1864  TL: Ecuador  M T

Mechanitis polymnia dorissides Staudinger, [1884]  TL: Peru  M T

Mechanitis polymnia eurydice Haensch, 1905  TL: Peru  M T

Mechanitis polymnia isthmia H. Bates, 1863  Costa Rica & Panama  M T S L I

Mechanitis polymnia kayei R. Fox, 1967  TL: Trinidad  M T

Mechanitis polymnia lycidice H. Bates, 1864  SE Mexico to Honduras  M T S L I

Mechanitis polymnia mauensis W. Forbes, 1948  TL: Brazil (Pa)

Mechanitis polymnia proceriformis Bryk, 1953  TL: Peru  M T

Mechanitis polymnia veritabilis A. Butler, 1873  TL: Colombia; Venezuela  M T S

Mechanitis polymnia werneri M. Hering, 1925  TL: Colombia  M T

Mechanitis polymnia [n. ssp.#1] K. Brown, MS  TL: Colombia

Mechanitis polymnia [n. ssp.#2] Lamas, MS  TL: Colombia

Mechanitis polymnia [n. ssp.#3] Lamas, MS  TL: Peru

Mechanitis polymnia [n. ssp.#4] Lamas, MS  TL: Peru

Mechanitis polymnia [n. ssp.#5] Lamas, MS  TL: Ecuador

Mechanitis polymnia [n. ssp.#6] Lamas, MS  TL: Brazil

Epityches  R.F. d'Almeida, 1938  

Epityches eupompe (Geyer, 1832)  S Brazil; ref.16  iN  M T S

Epityches ferra (Haensch, 1909)  Peru; ref.16  M T S

Epityches thyridiana (Haensch, 1909)  Bolivia; ref.16  M T S

Hyalyris  Boisduval, 1870  

Hyalyris antea (Hewitson, 1869)    iN

Hyalyris antea antea (Hewitson, 1869)  TL: Ecuador  M T

Hyalyris antea achuar Vitale & Bollino, 2000  TL: Ecuador  M T

Hyalyris antea amarilla Vitale & Bollino, 2000  TL: Ecuador  M T L

Hyalyris antea atagalpa (Haensch, 1905)  TL: Peru  M T S

Hyalyris antea cana (Haensch, 1905)  TL: Colombia  M T

Hyalyris antea flebilis R. Fox, 1971  TL: Venezuela  M T

Hyalyris antea frater (Salvin, 1869)  TL: Peru  M T S L

Hyalyris antea lalimba Lamas & Willmott, 2021  ref.318

Hyalyris antea [n. ssp.#1] Lamas & Willmott, MS  TL: Venezuela

Hyalyris antea [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyalyris coeno (E. Doubleday, 1847)    iN

Hyalyris coeno coeno (E. Doubleday, 1847)  TL: Venezuela  M T

Hyalyris coeno acceptabilis (Weeks, 1902)  TL: Bolivia  M T

Hyalyris coeno angustior (Schaus & T. Cockerell, 1923)  TL: Colombia  M T

Hyalyris coeno atrata R. Fox, 1971  TL: Peru  M T

Hyalyris coeno avinoffi R. Fox, 1971  TL: Venezuela  M T

Hyalyris coeno florida (Röber, 1930)  TL: Colombia  M T

Hyalyris coeno norella (Hewitson, [1859])  TL: Ecuador  M T

Hyalyris coeno norellana (Haensch, 1903)  TL: Ecuador  M T

Hyalyris coeno [n. ssp.] Lamas & Willmott, MS  Peru

Hyalyris excelsa (C. Felder & R. Felder, 1862)    iN

Hyalyris excelsa decumana (Godman & Salvin, 1879)  Costa Rica & Panama  M T S

Hyalyris excelsa Panamanian s segregate ,  Panama  M S

Hyalyris excelsa excelsa (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Hyalyris excelsa robertus (Bargmann, 1929)  TL: Colombia

Hyalyris excelsa tachira (T. Racheli & L. Racheli, 1998)  TL: Venezuela

Hyalyris excelsa [n. ssp.#1] Lamas & Willmott, MS  TL: Panama

Hyalyris excelsa [n. ssp.#2] Lamas & Willmott, MS  TL: Panama

Hyalyris excelsa [n. ssp.#3] Willmott & Lamas, MS  TL: Ecuador

Hyalyris fiammetta (Hewitson, 1852)  TL: S. Amer.  M T S

Hyalyris juninensis R. Fox & Real, 1971  TL: Peru

Hyalyris juninensis juninensis R. Fox & Real, 1971  TL: Peru  M T

Hyalyris juninensis [n. ssp.] Lamas, MS  Peru

Hyalyris latilimbata (Weymer, 1890)    iN  M T

Hyalyris leptalina (C. Felder & R. Felder, 1865)  TL: Brazil

Hyalyris leptalina leptalina (C. Felder & R. Felder, 1865)  TL: Brazil  M T S

Hyalyris leptalina [n. ssp.] Lamas, MS  Brazil

Hyalyris mestra (Hopffer, 1874)    iN

Hyalyris mestra mestra (Hopffer, 1874)  TL: Peru  M T

Hyalyris mestra fassli R. Fox & Real, 1971  TL: Bolivia

Hyalyris mestra personata R. Fox, 1971  TL: Ecuador  M T

Hyalyris mestra [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyalyris mestra [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyalyris ocna (Herrich-Schäffer, 1865)    iN

Hyalyris ocna ocna (Herrich-Schäffer, 1865)  TL: Colombia  M T

Hyalyris ocna aurea Lamas & Willmott, 2021  ref.318

Hyalyris ocna equatoria Willmott & Lamas, 2021  ref.318

Hyalyris ocna [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyalyris ocna [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyalyris oulita (Hewitson, [1859])    iN

Hyalyris oulita oulita (Hewitson, [1859])  TL: [Peru]  M T L

Hyalyris oulita lurida (A. Butler, 1873)  TL: Peru  M T S

Hyalyris oulita metella (Hopffer, 1874)  TL: Peru  M T L

Hyalyris oulita [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyalyris oulita [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyalyris oulita [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hyalyris oulita [n. ssp.#4] Lamas & Willmott, MS  TL: Bolivia

Hyalyris yasunina Willmott, Lamas & J. Hall, 2020  ref.317

Hyalyris praxilla (Hewitson, 1870)    iN

Hyalyris praxilla praxilla (Hewitson, 1870)  TL: Ecuador  M T

Hyalyris praxilla leuca Willmott & Lamas, 2021  ref.318

Hyalyris praxilla abscondita Lamas, Radford & Willmott, 2021  ref.318

Hyalyris schlingeri Real, 1971  TL: Peru  M T

Hyalyris lactea Willmott, Lamas & J. Hall, 2020  ref.317

Hyalyris lactea lactea Willmott, Lamas & J. Hall, 2020  ref.317

Hyalyris lactea granatina Willmott & Lamas, 2020  ref.317

Hyalyris lactea crocata Willmott & Lamas, 2020  ref.317

Napeogenes  H. Bates, 1862  

Napeogenes aethra (Hewitson, 1869)  TL: Ecuador  M T

Napeogenes apulia (Hewitson, 1858)    iN

Napeogenes apulia apulia (Hewitson, 1858)  TL: Colombia  M T

Napeogenes apulia palmasensis R. Fox & Real, 1971  TL: Peru  M T

Napeogenes apulia woytkowskii R. Fox & Real, 1971  TL: Peru  M T

Napeogenes apulia ocnita Willmott & Vitale, 2008  ref.357

Napeogenes apulia sangay Vitale & Willmott, 2008  ref.357

Napeogenes apulia [n. ssp.] Lamas, MS  Peru

Napeogenes benigna Weymer, 1899  TL: Colombia

Napeogenes benigna benigna Weymer, 1899  TL: Colombia  M T

Napeogenes benigna sandra Vitale & Willmott, 2008  ref.357

Napeogenes cranto C. Felder & R. Felder, 1865    iN

Napeogenes cranto paedaretus Godman & Salvin, 1878  Costa Rica & Panama  M T S

Napeogenes cranto reali R. Fox, 1968  Panama  M T S

Napeogenes cranto Panamanian segregate ,  Panama  M S

Napeogenes cranto cranto C. Felder & R. Felder, 1865  TL: Colombia  M T

Napeogenes cranto marilenae Vitale & Bollino, 2001  TL: Colombia

Napeogenes cranto chocoana Vitale & Bollino, 2008  ref.357  M T

Napeogenes duessa (Hewitson, [1859])    iN

Napeogenes duessa duessa (Hewitson, [1859])  TL: [Peru]  M T S

Napeogenes duessa anitae R. Fox & Real, 1971  TL: Ecuador

Napeogenes duessa cunza Vitale & Willmott, 2008  ref.357

Napeogenes duessa incas Bryk, 1953  TL: Peru  M T

Napeogenes duessa jamariensis R.F. d'Almeida, 1951  TL: Brazil (Rd)  M T

Napeogenes duessa mastersi R. Fox & Real, 1971  TL: Bolivia  M T

Napeogenes duessa orellana Vitale & Willmott, 2008  ref.357

Napeogenes duessa [n. ssp.#1] Lamas, MS  TL: Peru

Napeogenes duessa [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes duessa [n. ssp.#3] Lamas, MS  TL: Peru

Napeogenes flossina A. Butler, 1873  TL: Colombia

Napeogenes flossina flossina A. Butler, 1873  TL: Colombia  M T

Napeogenes flossina baumanni R. Fox & Real, 1971  TL: Bolivia  M T

Napeogenes flossina hypsaea Staudinger, 1885  TL: Colombia  M T

Napeogenes flossina [n. ssp.#1] Lamas, MS  TL: Peru

Napeogenes flossina [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes garwoodae Willmott & J. Hall, 2020  ref.317  iN

Napeogenes glycera Godman, 1899    iN

Napeogenes glycera glycera Godman, 1899  TL: Ecuador  M T

Napeogenes glycera ellariformis Strand, 1912  TL: Peru  M T

Napeogenes glycera eunomia Godman, 1899  TL: Peru  M T

Napeogenes glycera nausica Weymer, 1899  TL: Ecuador  M T

Napeogenes glycera olyrina Haensch, 1905  TL: Bolivia  M T

Napeogenes glycera mirador Lamas, Willmott & Radford, 2021  ref.318

Napeogenes glycera [n. ssp.] Lamas, MS  Peru

Napeogenes gracilis Haensch, 1905  TL: Bolivia  M T

Napeogenes harbona (Hewitson, 1869)    iN

Napeogenes harbona chiguinda Willmott & Vitale, 2008  ref.357

Napeogenes harbona harbona (Hewitson, 1869)  TL: Ecuador  M T

Napeogenes harbona domiduca (Hewitson, 1876)  TL: Bolivia  M T S

Napeogenes harbona forsteri R. Fox & Real, 1971  TL: Peru  M T

Napeogenes harbona podocarpus Vitale & Willmott, 2008  ref.357

Napeogenes harbona [n. ssp.#1] Lamas, MS  TL: Peru

Napeogenes harbona [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes harbona [n. ssp.#3] Lamas, MS  TL: Peru

Napeogenes inachia (Hewitson, 1855)    iN

Napeogenes inachia inachia (Hewitson, 1855)  TL: [Brazil (Pa)]  M T

Napeogenes inachia johnsoni R. Fox & Real, 1971  TL: Colombia  M T

Napeogenes inachia moles Haensch, 1905  TL: Guyana  M T

Napeogenes inachia patientia Lamas, 1985  TL: Peru  M T L

Napeogenes inachia pozziana (Oberthür, 1879)  TL: [Ecuador]  M T L

Napeogenes inachia pyrois H. Bates, 1862  TL: Brazil (Pa)  M T

Napeogenes inachia sulphurina H. Bates, 1862  TL: Brazil (Ba)  M T S

Napeogenes inachia [n. ssp.#1] Lamas, MS  TL: Peru

Napeogenes inachia [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes inachia tijatawaman Brévignon, 2007  ref.176

Napeogenes inachia [n. ssp.#3] K. Brown, MS  TL: Brazil

Napeogenes inachia [n. ssp.#4] Lamas, MS  TL: Brazil (Am)

Napeogenes inachia [n. ssp.#5] K. Brown, MS  TL: Brazil (Am)

Napeogenes inachia [n. ssp.#6] K. Brown, MS  TL: Brazil (Rd)

Napeogenes inachia [n. ssp.#7] K. Brown, MS  TL: Brazil

Napeogenes inachia [n. ssp.#8] K. Brown, MS  TL: Brazil (Pe)

Napeogenes inachia [n. ssp.#9] K. Brown, MS  TL: Brazil (MG)

Napeogenes inachia juanjuiensis R. Fox & Real, 1971  TL: Peru  M T

Napeogenes inachia [n. ssp.#10] Lamas, MS  TL: Peru

Napeogenes inachia [n. ssp.#11] Lamas, MS  TL: Peru

Napeogenes paruensis R.F. d'Almeida, 1958  TL: Brazil (Pa)

Napeogenes paruensis paruensis R.F. d'Almeida, 1958  ref.305  M T

Napeogenes paruensis tawaman Brévignon, 2007  ref.304,176

Napeogenes larilla (Hewitson, 1877)    iN

Napeogenes larilla larilla (Hewitson, 1877)  TL: Ecuador  M T

Napeogenes larilla delavillae Neild, 2008  ref.266

Napeogenes larilla [n. ssp.#1] Lamas, MS  TL: Colombia

Napeogenes larilla pastazana Vitale & Willmott, 2008  ref.357

Napeogenes larilla reventador Willmott & Vitale, 2008  ref.357

Napeogenes larilla [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes larina (Hewitson, [1856])    iN  M T

Napeogenes lycora (Hewitson, 1870)  TL: Ecuador

Napeogenes lycora lycora (Hewitson, 1870)  TL: Ecuador  M T

Napeogenes lycora attali Vitale & Willmott, 2008  ref.357

Napeogenes peridia (Hewitson, [1854])    iN

Napeogenes peridia hemisticta Schaus, 1913  Costa Rica & W Panama  M T S L

Napeogenes peridia hemimelaena Godman & Salvin, 1877  Panama  M T S

Napeogenes peridia peridia (Hewitson, [1854])  TL: Colombia  M T S

Napeogenes peridia hoppi M. Hering, 1925  TL: Colombia  M T S

Napeogenes peridia decora Godman, 1899  TL: Ecuador  M T S

Napeogenes peridia willmotti Vitale & Constantino, 2012  TL: Colombia

Napeogenes pharo (C. Felder & R. Felder, 1862)    iN

Napeogenes pharo pharo (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S

Napeogenes pharo astarte R. Fox, 1943  TL: Brazil (Am)  M T

Napeogenes pharo azeka (Hewitson, 1869)  TL: Colombia  M T

Napeogenes pharo elva Godman, 1899  TL: Colombia  M T

Napeogenes pharo gryne R.F. d'Almeida, 1951  TL: Brazil (Am)  M T

Napeogenes pharo lamia (Hewitson, 1869)  TL: Ecuador  M T

Napeogenes pharo osuna (Hewitson, 1876)  TL: Bolivia  M T

Napeogenes pharo pheranthes H. Bates, 1862  TL: Brazil (Am)  M T

Napeogenes pharo [n. ssp.#1] Lamas, MS  TL: Ecuador

Napeogenes pharo [n. ssp.#2] Lamas, MS  TL: Peru

Napeogenes pharo [n. ssp.#3] K. Brown, MS  TL: Brazil

Napeogenes pharo [n. ssp.#4] Lamas, MS  TL: Brazil (Pa)

Napeogenes rhezia (Geyer, [1834])    iN

Napeogenes rhezia rhezia (Geyer, [1834])  TL: S. Amer.  M T

Napeogenes rhezia adelphe H. Bates, 1862  TL: Brazil (Pa)  M T

Napeogenes rhezia adulta Haensch, 1905  TL: Guyana  M T

Napeogenes rhezia catamelas W. Forbes, 1942  TL: Venezuela  M T

Napeogenes rhezia cyrianassa (E. Doubleday, [1847])  TL: Brazil (Pa)  M T

Napeogenes rhezia tunantina H. Bates, 1862  TL: Brazil (Am)  M T S

Napeogenes rhezia yanetta (Hewitson, 1867)  TL: Brazil (ES)  M T

Napeogenes rhezia [n. ssp.#1] K. Brown, MS  TL: Venezuela

Napeogenes rhezia itu Brévignon, 2007  ref.176

Napeogenes rhezia [n. ssp.#2] K. Brown, MS  TL: Brazil (Pa)

Napeogenes rhezia [n. ssp.#3] K. Brown, MS  TL: Brazil (Am)

Napeogenes rhezia [n. ssp.#4] K. Brown, MS  TL: Brazil (Rd)

Napeogenes rhezia achaea (Hewitson, 1869)  TL: Ecuador  M T

Napeogenes rhezia curvilutea Kaye, 1918  TL: Peru  M T

Napeogenes rhezia nicolayi R. Fox & Real, 1971  TL: Colombia  M T S

Napeogenes rhezia [n. ssp.#5] Lamas, MS  TL: Peru

Napeogenes rhezia [n. ssp.#6] Lamas, MS  TL: Peru

Napeogenes sodalis Haensch, 1905  TL: Peru  M T

Napeogenes stella (Hewitson, [1855])    iN

Napeogenes stella Panamanian segregate ,  Panama  M S L

Napeogenes stella stella (Hewitson, [1855])  TL: Colombia  M T S

Napeogenes stella aster Godman, 1899  TL: Ecuador  M T

Napeogenes stella browni T. Racheli & L. Racheli, 1998  TL: Venezuela

Napeogenes stella opacella Kaye, 1918  TL: Colombia  M T S

Napeogenes stella delgadoi Vitale & Constantino, 2012  ref.297  M T

Napeogenes stella caucaensis Constantino & Vitale, 2012  ref.297  M T

Napeogenes sulphureophila Bryk, 1937  TL: not specified  M T

Napeogenes sylphis (Guérin-Méneville, [1844])    iN

Napeogenes sylphis sylphis (Guérin-Méneville, [1844])  TL: Bolivia

Napeogenes sylphis acreana R.F. d'Almeida, 1958  TL: Brazil (Ac)  M T

Napeogenes sylphis corena (Hewitson, [1861])  TL: Brazil (Am)  M T

Napeogenes sylphis ercilla (Hewitson, 1858)  TL: Brazil (Am)  M T

Napeogenes sylphis ithra (Hewitson, 1855)  TL: [Brazil (Pa)]  M T

Napeogenes sylphis potaronus Kaye, 1905  TL: Guyana  M T

Napeogenes sylphis rindgei R. Fox & Real, 1971  TL: Peru  M T

Napeogenes sylphis thira (Hewitson, 1874)  TL: Peru  M T

Napeogenes sylphis [n. ssp.#1] Lamas, MS  TL: Peru

Napeogenes sylphis [n. ssp.#2] Lamas, MS  TL: Brazil (Am)

Napeogenes sylphis ehu Brévignon, 2007  ref.176

Napeogenes tolosa (Hewitson, 1855)  Tolosa Tigerwing    iN

Napeogenes tolosa tolosa (Hewitson, 1855)  SE Mexico, Guatemala, Honduras  M T S L

Napeogenes tolosa tepehua (R. de la Maza & J. de la Maza, 2018)  ref.245

Napeogenes tolosa mombachoensis Brabant & Maes, 1997  Nicaragua  M T L

Napeogenes tolosa amara Godman, 1899  Nicaragua to Panama  M T S L

Napeogenes tolosa Panamanian s segregate ,  Panama  M S

Napeogenes tolosa chrispina (Hewitson, 1874)  TL: Colombia  M T

Napeogenes tolosa diaphanosa Kaye, 1918  TL: Colombia  M T

Napeogenes tolosa panamensis Vitale, Constantino & Delgado, 2012  ref.297  M T

Napeogenes tolosa [n. ssp.] Lamas, MS  Panama

Napeogenes verticilla (Hewitson, 1874)  TL: Peru  M T

Napeogenes zurippa (Hewitson, [1876])    iN

Napeogenes zurippa zurippa (Hewitson, [1876])  TL: Bolivia  M T

Napeogenes zurippa deucalion Haensch, 1905  TL: Peru  M T S

Napeogenes zurippa iquitensis Staudinger, [1884]  TL: Peru  M T

Napeogenes zurippa otaxes Godman, 1899  TL: Peru  M T S

Napeogenes zurippa pyrrho H. Druce, 1876  TL: Peru  M T

Napeogenes zurippa [n. ssp.] K. Brown, MS  Brazil (Rd)

Napeogenes zurippa quadrilis Haensch, 1903  TL: Ecuador  M T S

Hypothyris  Hübner, 1821  

Hypothyris anastasia (H. Bates, 1862)    iN

Hypothyris anastasia anastasia (H. Bates, 1862)  TL: Brazil (Am)  M T

Hypothyris anastasia acreana R.F. d'Almeida, 1958  TL: Brazil (Ac)  M T

Hypothyris anastasia anastasina (Staudinger, 1885)  TL: Peru  M T

Hypothyris anastasia arpi (R.F. d'Almeida, 1958)  TL: Brazil (Pa)  M T

Hypothyris anastasia bicolora (Haensch, 1903)  TL: Ecuador  M T

Hypothyris anastasia castanea (A. Butler, 1877)  TL: Brazil (Am)  M T

Hypothyris anastasia honesta (Weymer, 1883)  TL: Ecuador  M T S

Hypothyris anastasia niphas R.F. d'Almeida, 1945  TL: Brazil (Rd)  M T

Hypothyris anastasia porsenna (Srnka, 1885)  TL: Peru  M T

Hypothyris anastasia [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris anastasia [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris cantobrica (Hewitson, 1876)    iN

Hypothyris cantobrica cantobrica (Hewitson, 1876)  TL: Bolivia  M T

Hypothyris cantobrica carvalhoi (R.F. d'Almeida, 1954)  TL: Brazil (Pa)  M T

Hypothyris cantobrica nundina (R.F. d'Almeida, 1945)  TL: Brazil (Rd)  M T

Hypothyris cantobrica schunkeae (Lamas, 1979)  TL: Peru  M T

Hypothyris cantobrica zamorita Willmott, Boyer & Vitale, 2021  ref.318

Hypothyris cantobrica [n. ssp.#1] K. Brown, MS  TL: Brazil

Hypothyris cantobrica [n. ssp.#2] K. Brown, MS  TL: Brazil (Pa)

Hypothyris cantobrica [n. ssp.#3] K. Brown, MS  TL: Brazil (Pa)

Hypothyris connexa (A. Hall, 1939)  TL: Guyana

Hypothyris connexa connexa (A. Hall, 1939)  TL: Guyana  M T

Hypothyris connexa lema K. Brown, 1977  TL: Venezuela  M T

Hypothyris connexa [n. ssp.] K. Brown, MS  Venezuela

Hypothyris daphnis R.F. d'Almeida, 1945    iN

Hypothyris daphnis daphnis R.F. d'Almeida, 1945  TL: Brazil (Rd)  M T S

Hypothyris daphnis amapaensis K. Brown, 1980  TL: Brazil (Ap)  M T

Hypothyris daphnis clenchi K. Brown, 1980  TL: Brazil (Pa)  M T

Hypothyris daphnis daphnoides R.F. d'Almeida, 1945  TL: Brazil (Pa)  M T

Hypothyris daphnis madeira K. Brown, 1980  TL: Brazil (Am)  M T

Hypothyris euclea (Godart, 1819)    iN

Hypothyris euclea valora (Haensch, 1909)  S Mexico to Panama  M T S L

Hypothyris euclea Costa Rican segregate ,  Costa Rica  M S L

Hypothyris euclea euclea (Godart, 1819)  TL: Antilles  M T S

Hypothyris euclea barii (H. Bates, 1862)  TL: Brazil (Pa)  M T

Hypothyris euclea caldasensis R. Fox, 1971  TL: Colombia  M T

Hypothyris euclea callanga (Haensch, 1905)  TL: Peru  M T

Hypothyris euclea forbesi R. Fox, 1941  TL: Venezuela  M T

Hypothyris euclea intermedia (A. Butler, 1873)  TL: Colombia  M T

Hypothyris euclea interrupta (J. Zikán, 1941)  TL: Brazil (Am)  M T

Hypothyris euclea laphria (E. Doubleday, 1847)  TL: Brazil  M S

Hypothyris euclea micheneri R. Fox, 1971  TL: Colombia  M T

Hypothyris euclea nina (Haensch, 1905)  TL: Bolivia  M T

Hypothyris euclea nonia R.F. d'Almeida, 1945  TL: Brazil (Ac)  M T

Hypothyris euclea peruviana (Staudinger, 1885)  TL: Peru  M T L

Hypothyris euclea philetaera (Hewitson, 1876)  TL: ?  M T

Hypothyris euclea pyrippe (Hopffer, 1874)  TL: Peru  M T

Hypothyris euclea [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris euclea [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris euclea [n. ssp.#3] Lamas, MS  TL: Peru

Hypothyris euclea [n. ssp.#4] Lamas, MS  TL: Peru

Hypothyris euclea [n. ssp.#5] K. Brown, MS  TL: Brazil (Rd)

Hypothyris fluonia (Hewitson, 1854)    iN

Hypothyris fluonia fluonia (Hewitson, 1854)  TL: Brazil (Am)  M T S

Hypothyris fluonia baiochii T. Racheli & L. Racheli, 1998  TL: Venezuela

Hypothyris fluonia berna (Haensch, 1903)  TL: Ecuador  M T S L

Hypothyris fluonia flavigera (N. Riley, 1919)  TL: Brazil (Am)  M T

Hypothyris fluonia fulvifascia (Talbot, 1932)  TL: Peru  M T

Hypothyris fluonia hygia (Godman, 1899)  TL: Surinam  M T

Hypothyris fluonia iberina R.F. d'Almeida, 1945  TL: Brazil (Rd)  M T S

Hypothyris fluonia manaos (H. Bates, 1862)  TL: Brazil (Am)  M T

Hypothyris fluonia pardalina (Hopffer, 1874)  TL: Peru  M T

Hypothyris fluonia rowena (Hewitson, [1857])  TL: Colombia  M T

Hypothyris fluonia seminigra (Rosenberg & Talbot, 1914)  TL: Peru  M T S

Hypothyris fluonia uchiza Lamas, 1979  TL: Peru  M T

Hypothyris fluonia viola (Haensch, 1905)  TL: Bolivia  M T

Hypothyris fluonia violantilla R.F. d'Almeida, 1952  TL: Brazil (MT)  M T

Hypothyris fluonia [n. ssp.#1] Lamas, MS  TL: Ecuador

Hypothyris fluonia [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris fluonia [n. ssp.#3] K. Brown, MS  TL: Brazil (Am)

Hypothyris fluonia [n. ssp.#4] K. Brown, MS  TL: Brazil (Pa)

Hypothyris fluonia [n. ssp.#5] K. Brown, MS  TL: Brazil (Pa)

Hypothyris gemella R. Fox, 1971  TL: Guyana

Hypothyris gemella gemella R. Fox, 1971  TL: Guyana  M T

Hypothyris gemella mossi Neild, 2008  ref.266

Hypothyris gemella foxi Neild, 2008  ref.266

Hypothyris gemella batesi Neild, 2008  ref.266

Hypothyris gemella anduzei Neild, 2008  ref.266

Hypothyris gemella pseudoglycon Neild, 2008  ref.266

Hypothyris gemella [n. ssp.] K. Brown, MS  Brazil

Hypothyris leprieuri (Feisthamel, 1835)    iN

Hypothyris leprieuri leprieuri (Feisthamel, 1835)  TL: French Guiana

Hypothyris leprieuri catilla (Hewitson, 1876)  TL: Bolivia  M T S

Hypothyris leprieuri ignorata (Haensch, 1905)  TL: Brazil (Pa)  M T

Hypothyris leprieuri michaelisi (Haensch, 1909)  TL: Brazil (Pa)  M T

Hypothyris leprieuri nemea (Weymer, 1899)  TL: Colombia  M T

Hypothyris leprieuri ninyas R.F. d'Almeida, 1945  TL: Brazil (Rd)  M T

Hypothyris leprieuri [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris leprieuri [n. ssp.#2] K. Brown, MS  TL: Brazil

Hypothyris leprieuri [n. ssp.#3] K. Brown, MS  TL: Brazil (Pa)

Hypothyris leprieuri [n. ssp.#4] K. Brown, MS  TL: Brazil (Pa)

Hypothyris leprieuri [n. ssp.#5] K. Brown, MS  TL: Brazil

Hypothyris leprieuri [n. ssp.#6] K. Brown, MS  TL: Brazil

Hypothyris lycaste (Fabricius, 1793)  Lycaste Tigerwing    iN

Hypothyris lycaste dionaea (Hewitson, 1854)  SE Mexico & Guatemala  M T S L

Hypothyris lycaste callispila (H. Bates, 1866)  Costa Rica & Panama  M T IL

Hypothyris lycaste Panamanian segregate ,  Panama  M S

Hypothyris lycaste lycaste (Fabricius, 1793)  Panama & Colombia  M T IL L

Hypothyris lycaste antonia (Hewitson, 1869)  TL: Ecuador  M T

Hypothyris lycaste fraterna (Haensch, 1909)  TL: Venezuela  M T

Hypothyris lycaste glabra (Godman, 1899)  TL: Colombia  M T

Hypothyris lycaste limosa R. Fox, 1971  TL: Colombia  M T

Hypothyris lycaste limpida (Haensch, 1905)  TL: Colombia  M T

Hypothyris lycaste mergelena (Hewitson, [1855])  TL: Colombia  M T

Hypothyris lycaste [n. ssp.] Lamas, MS  Panama

Hypothyris mamercus (Hewitson, 1869)    iN

Hypothyris mamercus mamercus (Hewitson, 1869)  TL: Ecuador  M T S

Hypothyris mamercus maenas (Haensch, 1909)  TL: [Peru]  M T

Hypothyris mamercus poemne R.F. d'Almeida, 1939  TL: Brazil (Pa)  M T

Hypothyris mamercus polymnides (Haensch, 1905)  TL: Colombia  M T

Hypothyris mamercus [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris mamercus [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris mamercus [n. ssp.#3] K. Brown, MS  TL: Brazil (Rd)

Hypothyris mamercus [n. ssp.#4] K. Brown, MS  TL: Brazil

Hypothyris mamercus [n. ssp.#5] K. Brown, MS  TL: Brazil (Pa)

Hypothyris mansuetus (Hewitson, 1860)    iN

Hypothyris mansuetus mansuetus (Hewitson, 1860)  TL: [Peru]  M T S

Hypothyris mansuetus amica (Weymer, 1883)  TL: Colombia  M T S

Hypothyris mansuetus deemae R. Fox, 1943  TL: Peru  M T

Hypothyris mansuetus klotsi R. Fox, 1941  TL: Peru  M T

Hypothyris mansuetus meterus (Hewitson, 1860)  TL: [Peru]  M T

Hypothyris mansuetus zephyrus R. Fox, 1945  TL: Ecuador  M T

Hypothyris mansuetus [n. ssp.] Lamas, MS  Peru

Hypothyris moebiusi (Haensch, 1903)    iN

Hypothyris moebiusi moebiusi (Haensch, 1903)  TL: Ecuador  M T

Hypothyris moebiusi unicolora (Tessmann, 1928)  TL: Peru  M T

Hypothyris moebiusi [n. ssp.] Lamas, MS  Peru

Hypothyris ninonia (Hübner, [1806])    iN

Hypothyris ninonia ninonia (Hübner, [1806])  TL: ?

Hypothyris ninonia aetha R. Fox & Real, 1971  TL: Bolivia  M T

Hypothyris ninonia antonina (Staudinger, [1884])  TL: Peru  M T

Hypothyris ninonia apollinis (Staudinger, [1884])  TL: Peru  M T

Hypothyris ninonia caquetacola R. Fox, 1971  TL: Colombia  M T

Hypothyris ninonia colophonia R.F. d'Almeida, 1945  TL: Brazil (Rr)  M T

Hypothyris ninonia completa (Haensch, 1909)  TL: Brazil (Pa)  M T

Hypothyris ninonia completomaculata (J. Zikán, 1941)  TL: Brazil (Am)  M T

Hypothyris ninonia cornelie (Guérin-Méneville, [1844])  TL: Bolivia  M T

Hypothyris ninonia daeta (Boisduval, 1836)  TL: Brazil  M T

Hypothyris ninonia daetina (Weymer, 1899)  TL: Brazil (Ba)  M T

Hypothyris ninonia diphes R. Fox, 1971  TL: Colombia  M T

Hypothyris ninonia fimbria (Hewitson, 1855)  TL: Colombia  M T

Hypothyris ninonia granadensis (Haensch, 1905)  TL: Colombia  M T

Hypothyris ninonia latefasciata (Haensch, 1909)  TL: Upper Amazon  M T

Hypothyris ninonia latipennis (Tessmann, 1928)  TL: Peru  M T

Hypothyris ninonia matouryensis Brévignon, 2007  ref.176

Hypothyris ninonia mutilla (Hewitson, 1867)  TL: Guyana  M T

Hypothyris ninonia mysotis (Haensch, 1909)  TL: Venezuela  M T

Hypothyris ninonia neimyi (N. Riley, 1919)  TL: Brazil (Rd)  M T

Hypothyris ninonia pellucida (Haensch, 1905)  TL: Guyana  M T

Hypothyris ninonia vallina (Haensch, 1909)  TL: Venezuela  M T

Hypothyris ninonia [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris ninonia [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris ninonia [n. ssp.#3] Lamas, MS  TL: Peru

Hypothyris ninonia [n. ssp.#4] K. Brown, MS  TL: Brazil (Pa)

Hypothyris ninonia [n. ssp.#5] K. Brown, MS  TL: Brazil (Am)

Hypothyris ninonia [n. ssp.#6] K. Brown, MS  TL: Brazil (Am)

Hypothyris ninonia [n. ssp.#7] K. Brown, MS  TL: Brazil (Am)

Hypothyris semifulva (Salvin, 1869)    iN

Hypothyris semifulva semifulva (Salvin, 1869)  TL: Ecuador  M T

Hypothyris semifulva angelina (Haensch, 1905)  TL: Peru  M T S

Hypothyris semifulva dalmeidai R. Fox & Real, 1971  TL: Brazil (Rd)  M T

Hypothyris semifulva fulminans (A. Butler, 1873)  TL: Colombia  M T

Hypothyris semifulva meteroides R. Fox, 1971  TL: Peru  M T

Hypothyris semifulva pallisteri R. Fox & Real, 1971  TL: Peru  M T S

Hypothyris semifulva putumayoensis R. Fox & Real, 1971  TL: Colombia  M T

Hypothyris semifulva satura (Haensch, 1903)  TL: Ecuador  M T

Hypothyris semifulva soror (Srnka, 1885)  TL: Peru  M T

Hypothyris semifulva virgilini (N. Riley, 1919)  TL: Brazil (Ac)  M T

Hypothyris semifulva [n. ssp.#1] Lamas, MS  TL: Ecuador

Hypothyris semifulva [n. ssp.#2] Lamas, MS  TL: Peru

Hypothyris thea (Hewitson, 1852)  TL: Brazil (Am)

Hypothyris thea thea (Hewitson, 1852)  TL: Brazil (Am)  M T

Hypothyris thea mayi R.F. d'Almeida, 1945  TL: Brazil (Pa)  M T S

Hypothyris thea theatina (Haensch, 1909)  TL: Upper Amazon  M T

Hypothyris thea [n. ssp.#1] Lamas, MS  TL: Peru

Hypothyris thea [n. ssp.#2] K. Brown, MS  TL: Bolivia

Hypothyris vallonia (Hewitson, [1853])  TL: Brazil (Pa)

Hypothyris vallonia vallonia (Hewitson, [1853])  TL: Brazil (Pa)  M T

Hypothyris vallonia glycon (Godman, 1899)  TL: Guyana; Surinam  M T

Hypothyris vallonia [n. ssp.#1] K. Brown, MS  TL: Brazil

Hypothyris vallonia [n. ssp.#2] K. Brown, MS  TL: Brazil (Pa)

Hypothyris xanthostola (H. Bates, 1862)  TL: Brazil (Pa)

Hypothyris xanthostola xanthostola (H. Bates, 1862)  TL: Brazil (Pa)  M T S

Hypothyris xanthostola bellatula (R.F. d'Almeida, 1945)  TL: Brazil (Rd)  M T

Hypothyris xanthostola desmora Bryk, 1937  TL: Brazil (Pa)  M T

Hypothyris xanthostola ipoti Brévignon, 2007  ref.176

Hypothyris [n. sp.] Lamas, MS  Peru

Pagyris  Boisduval, 1870  

Pagyris euryanassa (C. Felder & R. Felder, 1860)  ref.16  iN  M T

Pagyris cymothoe (Hewitson, [1855])    iN

Pagyris cymothoe cymothoe (Hewitson, [1855])  TL: Venezuela  M T S L

Pagyris cymothoe sylvella (Hewitson, 1868)  TL: [Ecuador]  M T S

Pagyris priscilla Lamas, 1986  TL: Bolivia  M T

Pagyris renelichyi Neild, 2008  Venezuela; ref.266

Pagyris ulla (Hewitson, [1857])    iN

Pagyris ulla ulla (Hewitson, [1857])  TL: Colombia  M T

Pagyris ulla carmilla Lamas, 1986  TL: Peru  M T

Pagyris ulla dimidiata (Staudinger, 1885)  TL: Colombia  M T

Pagyris ulla leonila Lamas, 1986  TL: Peru  M T

Pagyris ulla vanilla Lamas, 1986  TL: Ecuador  M T

Pagyris ulla zorrilla Lamas, 1986  TL: Ecuador  M T

Ithomia  Hübner, 1816  

Ithomia agnosia Hewitson, [1855]    iN

Ithomia agnosia agnosia Hewitson, [1855]  TL: Venezuela  M T S

Ithomia agnosia pellucida Weymer, 1875  TL: Trinidad  M T

Ithomia agnosia zikani R.F. d'Almeida, 1940  TL: Brazil (MG)  M T S

Ithomia agnosia zumba Lamas & Willmott, 2021  ref.318

Ithomia amarilla Haensch, 1903    iN  M T S L

Ithomia arduinna R.F. d'Almeida, 1952    iN

Ithomia arduinna arduinna R.F. d'Almeida, 1952  TL: Bolivia  M T S L

Ithomia arduinna ardua Lamas, Vitale & Boyer, 2021  ref.318

Ithomia arduinna [n. ssp.] Lamas, MS  Peru

Ithomia avella Hewitson, 1854    iN

Ithomia avella avella Hewitson, 1854  TL: Colombia  M T

Ithomia avella adamsi Neild, 2008  ref.266

Ithomia avella cesleria Hewitson, [1855]  TL: Colombia  M T

Ithomia avella deliciae R. Fox, 1941  TL: Colombia

Ithomia avella epona Hewitson, 1869  TL: Ecuador  M T S

Ithomia avella katherineae R. Fox, 1941  TL: Colombia  M T

Ithomia avella miraculosa Lamas, 2003  TL: Colombia  M T

Ithomia avella roberti Neild, 2008  ref.266

Ithomia avella salazari Vitale & Bollino, 2000  TL: Ecuador  M T

Ithomia celemia Hewitson, [1854]    iN

Ithomia celemia plaginota A. Butler & H. Druce, 1872  Costa Rica & Panama  M T S

Ithomia celemia Panamanian segregate ,  Panama  M S

Ithomia celemia celemia Hewitson, [1854]  TL: Colombia  M T S

Ithomia celemia lurida Haensch, 1909  TL: Colombia  M T

Ithomia celemia [n. ssp.] Lamas, MS  Panama

Ithomia cleora Hewitson, 1855    iN  M T S L

Ithomia diasia Hewitson, 1854    iN

Ithomia diasia diasia Hewitson, 1854  TL: Colombia  M T

Ithomia diasia galata Hewitson, 1855  Panama to Colombia  M T S

Ithomia diasia hippocrenis H. Bates, 1866  Nicaragua to Panama  M T S L

Ithomia diasia leucophaena M. Hering, 1925  TL: Colombia

Ithomia diasia morena Haensch, 1903  TL: Ecuador  M T S

Ithomia diasia ossuna Haensch, 1909  TL: Colombia  M T

Ithomia drymo Hübner, 1816    iN  M T

Ithomia eleonora Haensch, 1905    iN  M T S

Ithomia ellara Hewitson, 1874    iN  M T

Ithomia heraldica H. Bates, 1866  Heraldica Clearwing    iN

Ithomia heraldica heraldica H. Bates, 1866  Costa Rica & Panama  M T S L

Ithomia heraldica Panamanian segregate ,  Panama  M S L

Ithomia hyala Hewitson, [1856]    iN

Ithomia hyala Panamanian segregate ,  Costa Rica & Panama  M S

Ithomia hyala hyala Hewitson, [1856]  TL: Ecuador  M T S

Ithomia hyala sincerus Bargmann, 1929  TL: [Colombia]  M T

Ithomia hyala [n. ssp.] Lamas, MS  Panama

Ithomia iphianassa E. Doubleday, 1847    iN

Ithomia iphianassa iphianassa E. Doubleday, 1847  TL: Venezuela  M T S

Ithomia iphianassa alienassa Haensch, 1905  TL: Colombia  M T S

Ithomia iphianassa anaphissa Herrich-Schäffer, 1865  TL: Colombia  M T

Ithomia iphianassa ethilla Neustetter, 1929  TL: Colombia  M T

Ithomia iphianassa florencia Winhard, 2020  ref.361

Ithomia iphianassa panamensis H. Bates, 1863  Panama  M T OI S L

Ithomia iphianassa phanessa Herrich-Schäffer, 1865  TL: Colombia  M T S

Ithomia iphianassa occidentissa Lamas & Willmott, 2021  ref.318

Ithomia jucunda Godman & Salvin, 1878  Jucunda Clearwing    iN

Ithomia jucunda jucunda Godman & Salvin, 1878  Panama  M T S

Ithomia jucunda bolivari Schaus, 1913  Nicaragua to Panama (Atlantic slope)  M T S L

Ithomia jucunda centromaculata Weymer, 1899  Panama & Colombia  M T OI S

Ithomia jucunda lamasi Vitale & Bollino, 2000  TL: Ecuador  M T

Ithomia jucunda [n. ssp.] Lamas, MS  Colombia

Ithomia lagusa Hewitson, [1856]    iN

Ithomia lagusa lagusa Hewitson, [1856]  TL: Colombia  M T

Ithomia lagusa adelinda Hewitson, 1868  TL: Ecuador  M T

Ithomia lagusa linda Hewitson, 1872  TL: Ecuador  M T

Ithomia lagusa peruana Salvin, 1869  TL: Peru  M T S L

Ithomia lagusa rachelii Vitale & Bollino, 2000  TL: Ecuador  M T

Ithomia lagusa theuda Hewitson, 1872  TL: Ecuador  M T

Ithomia lagusa [n. ssp.#1] Lamas, MS  TL: Colombia

Ithomia lagusa [n. ssp.#2] Lamas, MS  TL: Colombia

Ithomia lagusa [n. ssp.#3] Lamas, MS  TL: Colombia

Ithomia hymettia (Staudinger, 1885)  ref.318

Ithomia hymettia hymettia (Staudinger, 1885)  ref.318  M T S

Ithomia hymettia petitia Lamas & Willmott, 2021  ref.318

Ithomia leila Hewitson, 1852  Leila’s Clearwing  SE Mexico & Guatemala  iN  M T S L

Ithomia lichyi R.F. d'Almeida, 1939    iN

Ithomia lichyi lichyi R.F. d'Almeida, 1939  TL: Brazil ([RJ])  M T

Ithomia lichyi dalmeidai R. Fox, 1941  TL: Peru  M T

Ithomia lichyi neivai R.F. d'Almeida, 1940  TL: Brazil (Ac)  M T S

Ithomia lichyi [n. ssp.] Lamas, MS  Brazil

Ithomia patilla Hewitson, 1852  Patilla Clearwing  SE Mexico to Panama  iN  M T S L

Ithomia praeithomia Vitale & Bollino, 2003  TL: Peru

Ithomia praeithomia praeithomia Vitale & Bollino, 2003  TL: Peru

Ithomia praeithomia [n. ssp.] Lamas, MS  Peru

Ithomia pseudoagalla Rebel, 1902    iN  M T S L

Ithomia salapia Hewitson, [1853]    iN

Ithomia salapia salapia Hewitson, [1853]  TL: Ecuador  M T S

Ithomia salapia aquinia Hopffer, 1874  TL: Bolivia  M T L

Ithomia salapia ardea Hewitson, 1855  TL: Bolivia  M T

Ithomia salapia derasa Hewitson, 1855  TL: Nicaragua  M T S

Ithomia salapia diaphana Bryk, 1937  TL: Peru  M T S

Ithomia terra Hewitson, [1853]    iN

Ithomia terra terra Hewitson, [1853]  TL: Colombia  M T S

Ithomia terra jasinskii Vitale & Bollino, 2000  TL: Ecuador  M T

Ithomia terra oenanthe Weymer, 1899  TL: Colombia  M T

Ithomia terra terrana Haensch, 1903  TL: Ecuador  M T

Ithomia terra vulcana Haensch, 1909  Costa Rica & Panama  M T S L

Ithomia terra [n. ssp.#1] Lamas, MS  TL: Colombia

Ithomia terra [n. ssp.#2] Lamas, MS  TL: Peru

Ithomia xenos (H. Bates, 1866)  Xenos Clearwing    iN

Ithomia xenos xenos (H. Bates, 1866)  Costa Rica & Panama  M T S L

Ithomia xenos Panamanian segregate ,  Panama  M S

Ithomia xenos [n. ssp.] Lamas, MS  Panama

Megoleria  Constantino, 1999  

Megoleria orestilla (Hewitson, 1867)    iN

Megoleria orestilla orestilla (Hewitson, 1867)  TL: Colombia  M T

Megoleria orestilla polymacula (Rosenberg & Talbot, 1914)  TL: Peru  M T S

Megoleria orestilla polylla Lamas & Willmott, 2008  ref.334

Megoleria susiana (C. Felder & R. Felder, 1862)    iN

Megoleria susiana susiana (C. Felder & R. Felder, 1862)  TL: Colombia  M T S L

Megoleria susiana sandra Constantino, 1999  TL: Colombia  M T

Megoleria susiana susanna (Staudinger, [1884])  TL: Colombia  M T S

Hyposcada  Godman & Salvin, 1879  

Hyposcada anchiala (Hewitson, 1868)    iN

Hyposcada anchiala Panamanian segregate ,  Panama  M S

Hyposcada anchiala anchiala (Hewitson, 1868)  TL: Peru  M T

Hyposcada anchiala abidula J. Zikán, 1941  TL: Brazil (Am)  M T

Hyposcada anchiala cynara R.F. d'Almeida, 1945  TL: Brazil (Am)  M T

Hyposcada anchiala ecuadorina Bryk, 1953  TL: Ecuador  M T S

Hyposcada anchiala fallax (Staudinger, [1884])  TL: Peru  M S

Hyposcada anchiala gallardi Brévignon, 1993  TL: French Guiana  M T

Hyposcada anchiala interrupta Tessmann, 1928  TL: Peru  M T S

Hyposcada anchiala kezia (Hewitson, 1868)  TL: Peru  M T S

Hyposcada anchiala mendax R. Fox, 1941  TL: Peru  M T

Hyposcada anchiala rezia Haensch, 1905  TL: Peru  M T S

Hyposcada anchiala richardsi R. Fox, 1941  TL: Peru  M T S

Hyposcada anchiala [n. ssp.#1] Lamas & Willmott, MS  TL: Panama

Hyposcada anchiala [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyposcada attilodes Kaye, 1918  TL: Bolivia  M T

Hyposcada [n. sp.] Lamas & Willmott, MS  not specified

Hyposcada [n. sp.] [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyposcada [n. sp.] [n. ssp.#2] Lamas & Willmott, MS  TL: Bolivia

Hyposcada dujardini Brévignon, 1993  TL: French Guiana

Hyposcada dujardini dujardini Brévignon, 1993  TL: French Guiana  M T S

Hyposcada dujardini humboldti Neild, 2008  ref.266

Hyposcada illinissa (Hewitson, [1852])    iN

Hyposcada illinissa illinissa (Hewitson, [1852])  TL: [Brazil (Am)]  M T S

Hyposcada illinissa abida (Hewitson, 1871)  TL: Colombia  M T

Hyposcada illinissa aesion (Godman & Salvin, 1878)  Panama  M T S L

Hyposcada illinissa brisotis (Haensch, 1909)  TL: Bolivia  M T S

Hyposcada illinissa cynthia R. Fox, 1941  TL: Ecuador  M T

Hyposcada illinissa dolabella (Hewitson, 1876)  TL: Bolivia  M T S

Hyposcada illinissa ida Haensch, 1903  TL: Ecuador  M T L

Hyposcada illinissa idina Haensch, 1905  TL: [Peru]  M T

Hyposcada illinissa ilerdinoides (Staudinger, [1884])  TL: Brazil (Am)  M T

Hyposcada illinissa margarita R. Fox, 1941  TL: Peru  M T

Hyposcada illinissa napirida J. Zikán, 1941  TL: Brazil (Am)  M T

Hyposcada illinissa napoensis Vitale & Bollino, 2001  TL: Ecuador  M T

Hyposcada illinissa roberto Neild, 2008  ref.266

Hyposcada illinissa sinilia (Herrich-Schäffer, 1865)  TL: Colombia  M T L

Hyposcada illinissa tundayme S. Padrón, 2008  ref.275

Hyposcada illinissa varia Winhard, 2018  ref.359

Hyposcada illinissa mocoa Winhard, 2018  ref.349,359

Hyposcada illinissa morona Lamas & Willmott, 2021  ref.318

Hyposcada illinissa [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#5] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#6] Lamas & Willmott, MS  TL: Peru

Hyposcada illinissa [n. ssp.#7] Lamas & Willmott, MS  TL: Brazil (Am)

Hyposcada schausi R. Fox, 1941  TL: Colombia

Hyposcada schausi schausi R. Fox, 1941  TL: Colombia  M T

Hyposcada schausi lactea Vitale & Bollino, 2001  TL: Ecuador

Hyposcada taliata (Hewitson, 1874)  TL: Peru

Hyposcada taliata taliata (Hewitson, 1874)  TL: Peru  M T

Hyposcada taliata laetitia Lamas & Willmott, 2021  ref.318

Hyposcada virginiana (Hewitson, [1855])  Virginiana Clearwing    iN

Hyposcada virginiana virginiana (Hewitson, [1855])  SE Mexico to Honduras  M T S L

Hyposcada virginiana adelphina (H. Bates, 1866)  Panama  M T TI L

Hyposcada virginiana consobrina (Godman & Salvin, 1892)  TL: Ecuador  M T

Hyposcada virginiana evanides Haensch, 1909  Nicaragua to Panama  M T S L

Hyposcada virginiana neustetteri Bargmann, 1928  TL: Colombia  M T

Hyposcada zarepha (Hewitson, 1869)    iN

Hyposcada zarepha zarepha (Hewitson, 1869)  TL: French Guiana  M T

Hyposcada zarepha bonplandi Neild, 2008  ref.266

Hyposcada zarepha pseudegra (J. Zikán, 1941)  TL: Brazil (Pa)  M T

Hyposcada kena (Hewitson, 1872)  ref.318

Hyposcada kena kena (Hewitson, 1872)  TL: Ecuador  M T

Hyposcada kena flexibilis (Haensch, 1909)  TL: Peru  M T S

Hyposcada kena luisa (Tessmann, 1928)  TL: Peru  M T

Hyposcada kena virina (Haensch, 1909)  TL: Bolivia  M T

Hyposcada kena unki Lamas, Willmott & Boyer, 2021  ref.318

Hyposcada kena [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hyposcada kena [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hyposcada kena [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hyposcada kena [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Hyposcada kena [n. ssp.#5] Lamas & Willmott, MS  TL: Colombia

Hyposcada kena [n. ssp.#6] Lamas & Willmott, MS  TL: Colombia

Oleria  Hübner, 1816  

Oleria [n. sp.#1] Willmott & Lamas, MS  TL: Guyana

Oleria aegle (Fabricius, 1776)    iN

Oleria aegle aegle (Fabricius, 1776)  TL: India

Oleria aegle divisa (Aurivillius, 1929)  TL: Brazil (Am)  M T S

Oleria aegle egra (Hewitson, [1852])  TL: [Brazil (Pa)]  M T

Oleria aegle [n. ssp.] Lamas & Willmott, MS  Brazil (Rd)

Oleria agarista (C. Felder & R. Felder, 1862)    iN

Oleria agarista agarista (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Oleria agarista idalie R. Fox, 1941  TL: Peru

Oleria agarista [n. ssp.] Willmott & Lamas, MS  Colombia

Oleria alexina (Hewitson, [1859])    iN  M T S

Oleria amalda (Hewitson, [1857])    iN

Oleria amalda amalda (Hewitson, [1857])  TL: Colombia  M T S

Oleria amalda amaldina (Haensch, 1909)  TL: Colombia  M T L

Oleria amalda faunula (Bargmann, 1929)  TL: Colombia  M T

Oleria amalda modesta (Haensch, 1903)  TL: Ecuador  M T S

Oleria amalda [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria amalda [n. ssp.#2] Lamas & Willmott, MS  TL: Colombia

Oleria amalda [n. ssp.#3] Neild, MS  TL: Venezuela

Oleria antaxis (Haensch, 1909)    iN

Oleria antaxis antaxis (Haensch, 1909)  TL: [Brazil (Pa)]  M T S

Oleria antaxis machadoi R.F. d'Almeida, 1962  TL: Brazil (Ap)  M T

Oleria antaxis saulensis Brévignon, 1993  TL: French Guiana  M T

Oleria aquata (Weymer, 1875)    iN  M T S

Oleria assimilis (Haensch, 1903)    iN

Oleria assimilis assimilis (Haensch, 1903)  TL: Ecuador  M T S L

Oleria assimilis [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria assimilis [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria assimilis [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Oleria assimilis [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Oleria astrea (Cramer, 1775)    iN

Oleria astrea astrea (Cramer, 1775)  TL: Brazil  M S

Oleria astrea burchelli (Sanders, 1904)  TL: Brazil (Pa)  M T S L

Oleria astrea chloris Lamas & Brévignon, 1993  TL: French Guiana  M T

Oleria astrea [n. ssp.#1] Willmott & Lamas, MS  TL: Brazil (RJ)

Oleria astrea [n. ssp.#2] Lamas & Willmott, MS  TL: Brazil (Rd)

Oleria thiemei (Oberthür, 1879)  Thieme's Clearwing  Panama to Colombia  iN  M T S

Oleria athalina (Staudinger, [1884])    iN

Oleria athalina athalina (Staudinger, [1884])  TL: Ecuador or Bolivia  M T

Oleria athalina banjana (Haensch, 1903)  TL: Ecuador  M T S

Oleria athalina epimakrena (Haensch, 1905)  TL: Bolivia  M T S

Oleria athalina [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria athalina [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria attalia (Hewitson, 1855)    iN

Oleria attalia attalia (Hewitson, 1855)  TL: Bolivia  M T S L

Oleria attalia tabera (Hewitson, 1869)  TL: Ecuador  M T

Oleria attalia [n. ssp.] Lamas & Willmott, MS  Peru

Oleria baizana (Haensch, 1903)    iN

Oleria baizana baizana (Haensch, 1903)  TL: Ecuador  M T

Oleria baizana munaycha T. Racheli & L. Racheli, 2003  ref.286

Oleria bifurcata Willmott & Lamas, 2020  ref.317

Oleria bioculata (Haensch, 1905)    iN

Oleria bioculata bioculata (Haensch, 1905)  TL: Bolivia  M T

Oleria bioculata tapio Bryk, 1953  TL: Peru  M T

Oleria boyeri Neild, 2008  ref.266

Oleria chimaera Willmott & Lamas, 2020  ref.317

Oleria cyrene (Latreille, [1809])    iN

Oleria cyrene cyrene (Latreille, [1809])  TL: Peru  M T

Oleria cyrene solida (Weymer, 1883)  TL: Ecuador  M T

Oleria cyrene susianoides Winhard, 2019  ref.351,360

Oleria deronda (Hewitson, 1876)    iN

Oleria deronda deronda (Hewitson, 1876)  TL: Bolivia  M T

Oleria deronda valida (Haensch, 1905)  TL: Peru  M T S

Oleria deronda [n. ssp.] Lamas & Willmott, MS  Peru

Oleria derondina (Haensch, 1909)    iN

Oleria derondina derondina (Haensch, 1909)  TL: Bolivia  M T S

Oleria derondina mandorina Lamas, 2003  TL: Peru  M T

Oleria derondina [n. ssp.] Lamas & Willmott, MS  Peru

Oleria didymaea (Hewitson, 1876)    iN

Oleria didymaea didymaea (Hewitson, 1876)  TL: Bolivia  M T S L

Oleria didymaea denuda (N. Riley, 1919)  TL: Brazil (Ac)  M T

Oleria didymaea ramona (Haensch, 1909)  TL: Peru  M T

Oleria didymaea [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria didymaea [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria didymaea [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Oleria didymaea [n. ssp.#4] Lamas & Willmott, MS  TL: Brazil (Rd)

Oleria enania (Haensch, 1909)  TL: Peru

Oleria enania enania (Haensch, 1909)  TL: Peru  M T

Oleria enania itacoaiensis R.F. d'Almeida, 1964  TL: Brazil (Am)  M T

Oleria enania [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria enania [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria estella (Hewitson, 1868)    iN

Oleria estella estella (Hewitson, 1868)  TL: Ecuador  M T S

Oleria estella subosa (Haensch, 1909)  TL: Bolivia  M T S

Oleria fasciata (Haensch, 1903)    iN

Oleria fasciata fasciata (Haensch, 1903)  TL: Ecuador  M T

Oleria fasciata melchori Lamas, 2003  TL: Peru  M T

Oleria fasciata [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria fasciata [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria flora (Cramer, 1779)  TL: Surinam

Oleria flora flora (Cramer, 1779)  TL: Surinam  M L

Oleria flora [n. ssp.] Willmott & Lamas, MS  Brazil (Pa)

Oleria fumata (Haensch, 1905)    iN

Oleria fumata fumata (Haensch, 1905)  TL: Colombia  M T L

Oleria fumata richardina Lamas, 1994  TL: Colombia  M T

Oleria gunilla (Hewitson, 1858)    iN

Oleria gunilla gunilla (Hewitson, 1858)  TL: [Brazil (Am)]  M T

Oleria gunilla borilis (Haensch, 1909)  TL: Peru  M T

Oleria gunilla escura (Haensch, 1903)  TL: Ecuador  M T

Oleria gunilla lerdina (Staudinger, 1885)  TL: Peru  M T

Oleria gunilla lota (Hewitson, 1872)  TL: Ecuador  M T L

Oleria gunilla lourdes Freitas & Willmott, 2016  TL: Brazil (MT)

Oleria gunilla lubilerda (Haensch, 1905)  TL: Colombia  M T S

Oleria gunilla praemona (Haensch, 1909)  TL: [Peru]  M T

Oleria gunilla serdolis (Haensch, 1909)  TL: [Peru]  M T S

Oleria gunilla [n. ssp.#1] Lamas & Willmott, MS  TL: Brazil (Pa)

Oleria gunilla [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria ilerdina (Hewitson, 1858)    iN

Oleria ilerdina ilerdina (Hewitson, 1858)  TL: Peru  M T

Oleria ilerdina hermieri Brévignon, 1993  TL: French Guiana  M T

Oleria ilerdina lerida (W. F. Kirby, 1878)  TL: Ecuador  M T

Oleria ilerdina priscilla (Hewitson, 1858)  TL: [Brazil (Am)]  M T

Oleria makrena (Hewitson, 1854)    iN

Oleria makrena Panamanian segregate ,  Panama  M S

Oleria makrena makrena (Hewitson, 1854)  TL: Venezuela  M S

Oleria makrena caucana (Staudinger, 1885)  TL: Colombia  M L

Oleria makrena [n. ssp.#1] Willmott & Lamas, MS  TL: Panama

Oleria makrena [n. ssp.#2] Lamas & Willmott, MS  TL: Colombia

Oleria makrena [n. ssp.#3] Lamas & Willmott, MS  TL: Colombia

Oleria onega (Hewitson, [1852])    iN

Oleria onega onega (Hewitson, [1852])  TL: [Brazil (Am)]  M T S

Oleria onega bocca (N. Riley, 1919)  TL: Brazil (Am)  M T

Oleria onega crispinilla (Hopffer, 1874)  TL: Peru  M S

Oleria onega epicharme (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Oleria onega ilerda (Hewitson, [1854])  TL: Colombia

Oleria onega janarilla (Hewitson, 1863)  TL: Ecuador

Oleria onega lentita Lamas, 1985  TL: Peru  M T S L

Oleria onega perspicua (A. Butler, 1877)  TL: Brazil (Am)  M T

Oleria onega astigara Willmott & Lamas, 2021  ref.318

Oleria onega [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria onega [n. ssp.#2] Willmott & Lamas, MS  TL: Colombia

Oleria onega [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Oleria padilla (Hewitson, 1863)    iN

Oleria padilla padilla (Hewitson, 1863)  TL: Ecuador  M T S

Oleria padilla cajamarcensis Baumann, 1985  TL: Peru

Oleria padilla gorkyi Lamas, 2003  TL: Peru  M T

Oleria padilla pseudmakrena Baumann, 1985  TL: Bolivia  M S

Oleria padilla [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria padilla [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria paula Weymer, 1883  Paula’s Clearwing    iN

Oleria paula paula (Weymer, 1883)  E & SW Mexico to Panama  M S L Ha

Oleria paula Panamanian segregate ,  Osa Peninsula, Costa Rica, to W Panama, Pacific slope  M S

Oleria paula [n. ssp.#1] Lamas & Willmott, MS  TL: Panama

Oleria paula [n. ssp.#2] Lamas & Willmott, MS  TL: Colombia

Oleria phenomoe (E. Doubleday, [1847])    iN

Oleria phenomoe phenomoe (E. Doubleday, [1847])  TL: Venezuela  M T S

Oleria phenomoe apexatrata Neild, 2008  ref.266

Oleria quadrata (Haensch, 1903)    iN

Oleria quadrata quadrata (Haensch, 1903)  TL: Ecuador  M S

Oleria quadrata [n. ssp.] Willmott & Lamas, MS  Colombia

Oleria quintina (C. Felder & R. Felder, 1865)    iN  M T S L

Oleria radina (Haensch, 1909)    iN

Oleria radina radina (Haensch, 1909)  TL: Colombia

Oleria radina bonita Willmott & Lamas, 2021  ref.318

Oleria radina [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria radina [n. ssp.#2] Willmott & Lamas, MS  TL: Colombia

Oleria rubescens (A. Butler & H. Druce, 1872)  Rubescens Clearwing  Nicaragua to Panama  iN  M T S

Oleria santineza (Haensch, 1903)    iN

Oleria santineza santineza (Haensch, 1903)  TL: Ecuador

Oleria santineza reducida Neild, 2008  ref.266  M S

Oleria santineza veneae Neild, 2008  ref.266

Oleria santineza [n. ssp.] Willmott & Lamas, MS  Colombia

Oleria sexmaculata (Haensch, 1903)    iN

Oleria sexmaculata sexmaculata (Haensch, 1903)  TL: Ecuador

Oleria sexmaculata lerda (Haensch, 1909)  TL: [Peru]

Oleria sexmaculata [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria sexmaculata [n. ssp.#2] Willmott & Lamas, MS  TL: Colombia

Oleria sexmaculata [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Oleria similigena R.F. d'Almeida, 1962  TL: Brazil (Ap)

Oleria similigena similigena R.F. d'Almeida, 1962  TL: Brazil (Ap)  M S

Oleria similigena [n. ssp.#1] Willmott & Lamas, MS  TL: Guyana

Oleria similigena [n. ssp.#2] Willmott & Lamas, MS  TL: Brazil (Pa)

Oleria synnova (Hewitson, [1859])  TL: [Brazil (Am)]  M T

Oleria tigilla (Weymer, 1899)    iN

Oleria tigilla tigilla (Weymer, 1899)  TL: Ecuador  M S

Oleria tigilla raya Lamas & Willmott, 2021  ref.318

Oleria tigilla [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Oleria tigilla [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria tremona (Haensch, 1909)    iN

Oleria tremona tremona (Haensch, 1909)  TL: Colombia

Oleria tremona ochoai Lamas, 2003  TL: Peru  M T

Oleria tremona schoenfelderi Baumann, 1985  TL: Bolivia

Oleria tremona benigna Lamas & Willmott, 2021  ref.318

Oleria tremona [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Oleria tremona [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Oleria vicina (Salvin, 1869)  Vicina Clearwing  Costa Rica & W Panama  iN  M T IL L

Oleria victorine (Guérin-Méneville, [1844])    iN

Oleria victorine victorine (Guérin-Méneville, [1844])  TL: Bolivia  M S

Oleria victorine graziella (Oberthür, 1879)  TL: Venezuela  M T S

Oleria victorine manora (Schaus, 1902)  TL: Brazil (RJ)  M S

Oleria victorine sarilis (Haensch, 1909)  TL: [Peru]  M L

Oleria victorine vicenteae Neild, 2008  ref.266

Oleria victorine [n. ssp.] Lamas & Willmott, MS  Brazil (Rd)

Oleria [n. sp.#2] Lamas & Willmott, MS  TL: Peru

Oleria zea (Hewitson, [1855])  Zea Clearwing    iN

Oleria zea zea (Hewitson, [1855])  SE Mexico to El Salvador & Honduras  M T S L

Oleria zea diazi J. Maza & Lamas, 1978  SW Mexico  M T S

Oleria zelica (Hewitson, 1856)    iN

Oleria zelica zelica (Hewitson, 1856)  TL: Ecuador  M T S

Oleria zelica daguana (Bargmann, 1928)  TL: Colombia  M T

Oleria zelica pagasa (H. Druce, 1875)  Costa Rica & Panama  M T S L

Ollantaya  K. Brown & A. Freitas, 1994  

Ollantaya aegineta (Hewitson, 1869)  TL: Ecuador

Ollantaya aegineta aegineta (Hewitson, 1869)  ref.194  M T

Ollantaya aegineta cleobulina (Hewitson, 1876)  ref.194  M T

Ollantaya aegineta inelegans (Hewitson, 1877)  ref.194  M T S

Ollantaya aegineta [n. ssp.] Lamas & Willmott, MS  Peru

Ollantaya canilla (Hewitson, 1874)  ref.194  iN

Ollantaya canilla canilla (Hewitson, 1874)  TL: Peru  M T

Ollantaya canilla [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Ollantaya canilla [n. ssp.#2] Willmott & Lamas, MS  TL: Bolivia

Ollantaya olerioides (R.F. d'Almeida, 1952)  TL: Bolivia

Ollantaya olerioides olerioides (R.F. d'Almeida, 1952)  ref.194  M T

Ollantaya olerioides baeza Willmott & Lamas, 2021  ref.318

Ollantaya olerioides [n. ssp.] Lamas & Willmott, MS  Peru

Ollantaya [n. sp.] Lamas & Willmott, MS  Peru

Ceratinia  Hübner, 1816  

Ceratinia cayana (Salvin, 1869)    iN

Ceratinia cayana cayana (Salvin, 1869)  TL: French Guiana  M T

Ceratinia cayana giparanaensis R.F. d'Almeida, 1964  TL: Brazil (Rd)  M T

Ceratinia iolaia (Hewitson, [1856])    iN

Ceratinia iolaia iolaia (Hewitson, [1856])  TL: Colombia  M T

Ceratinia iolaia conveniens Bryk, 1937  TL: Colombia  M T

Ceratinia iolaia rehni R. Fox, 1941  TL: Colombia  M T

Ceratinia neso (Hübner, [1806])    iN

Ceratinia neso neso (Hübner, [1806])  TL: [Brazil]  M T

Ceratinia neso espriella (Hewitson, 1868)  TL: Ecuador  M T

Ceratinia neso hamlini (Weeks, 1906)  TL: Venezuela  M T

Ceratinia neso immaculata Neild, 2008  ref.266

Ceratinia neso naranja Neild, 2008  ref.266

Ceratinia neso nisea (Godart, 1819)  TL: French Guiana; Surinam  M T

Ceratinia neso niselina (J. Zikán, 1941)  TL: Brazil (Am)  M T

Ceratinia neso peruensis (Haensch, 1905)  TL: Peru  M T

Ceratinia neso tarapotis (Haensch, 1909)  TL: [Peru]  M T

Ceratinia neso [n. ssp.#1] Lamas, MS  TL: Colombia

Ceratinia neso [n. ssp.#2] Lamas, MS  TL: Venezuela

Ceratinia neso [n. ssp.#3] Lamas, MS  TL: Venezuela

Ceratinia neso [n. ssp.#4] Lamas, MS  TL: Peru

Ceratinia neso [n. ssp.#5] Lamas, MS  TL: Bolivia

Ceratinia neso [n. ssp.#6] Lamas, MS  TL: Brazil (Rd)

Ceratinia neso [n. ssp.#7] Lamas, MS  TL: Brazil (Pa)

Ceratinia tutia (Hewitson, 1852)    iN

Ceratinia tutia tutia (Hewitson, 1852)  TL: Amazon  M T L

Ceratinia tutia Northern segregate ,  SE Mexico, Guatemala, Honduras  M S

Ceratinia tutia Costa Rican segregate ,  Costa Rica  M S L

Ceratinia tutia azara (Hewitson, [1854])  TL: Ecuador  M S

Ceratinia tutia dorilla (H. Bates, 1864)  Panama  M T S L1 L2

Ceratinia tutia singularis (Rebel, 1902)  TL: Ecuador  M T

Ceratinia tutia tosca (Schaus, 1902)  TL: Colombia  M T

Ceratinia tutia transversa (M. Hering, 1925)  TL: Colombia  M S

Ceratinia tutia [n. ssp.#1] Lamas, MS  TL: Mexico

Ceratinia tutia [n. ssp.#2] Lamas, MS  TL: Costa Rica

Ceratinia tutia [n. ssp.#3] Lamas, MS  TL: Panama

Ceratinia tutia [n. ssp.#4] Lamas, MS  TL: Colombia

Ceratinia tutia [n. ssp.#5] Lamas, MS  TL: Venezuela

Ceratinia poecila (H. Bates, 1862)  TL: Colombia

Ceratinia poecila poecila (H. Bates, 1862)  TL: Colombia  M T

Ceratinia poecila azarina (Weymer, 1899)  TL: Ecuador  M T

Ceratinia poecila callichroma (Staudinger, 1885)  TL: Ecuador  M T

Ceratinia poecila chanchamaya (Haensch, 1905)  TL: Peru  M T

Ceratinia poecila fuscens (Haensch, 1905)  TL: Bolivia  M T

Ceratinia poecila hopfferi (Weymer, 1899)  TL: [Peru]  M T

Ceratinia poecila poeciloides (N. Riley, 1919)  TL: Brazil (Am)  M T

Ceratinia poecila porrecta (Haensch, 1905)  TL: Bolivia  M T

Ceratinia poecila radiosa (Haensch, 1903)  TL: Ecuador  M T

Ceratinia poecila robusta (Haensch, 1905)  TL: Bolivia  M T

Ceratinia poecila selenides (Weymer, 1899)  TL: Peru  M T

Ceratinia poecila [n. ssp.#1] Lamas, MS  TL: Peru

Ceratinia poecila [n. ssp.#2] Lamas, MS  TL: Peru

Ceratinia poecila [n. ssp.#3] Lamas, MS  TL: Peru

Ceratinia poecila [n. ssp.#4] Lamas, MS  TL: Peru

Callithomia  H. Bates, 1862  

Callithomia alexirrhoe H. Bates, 1862    iN

Callithomia alexirrhoe alexirrhoe H. Bates, 1862  TL: Brazil (Am)  M T

Callithomia alexirrhoe butes Godman & Salvin, 1898  TL: [Peru]  M T S

Callithomia alexirrhoe schulzi Haensch, 1905  TL: Brazil (Pa)  M T

Callithomia alexirrhoe thornax H. Bates, 1862  TL: Brazil (Am)  M T

Callithomia alexirrhoe zeuxippe H. Bates, 1862  TL: Brazil (Pa)  M T

Callithomia alexirrhoe [n. ssp.#1] Lamas, MS  TL: Colombia

Callithomia alexirrhoe [n. ssp.#2] Lamas, MS  TL: Brazil (MT)

Callithomia hezia (Hewitson, [1854])  Hezia Clearwing    iN

Callithomia hezia hezia (Hewitson, [1854])  SE Mexico to Honduras  M T S L

Callithomia hezia beronilla (Hewitson, 1874)  TL: Colombia  M T

Callithomia hezia formosa Winhard, 2019  ref.360

Callithomia hezia hedila Godman & Salvin, 1879  SE Mexico to Guatemala  M T S

Callithomia hezia nigra Winhard, 2019  ref.360

Callithomia hezia perija Masters, 1973  TL: Venezuela  M T

Callithomia hezia phagesia (Hewitson, 1869)  TL: Ecuador  M T

Callithomia hezia pulchra Winhard, 2019  ref.360

Callithomia hezia renati Neild, 2008  ref.266

Callithomia hezia smalli R. Fox, 1968  Panama  M T

Callithomia hezia tridactyla Dewitz, 1877  TL: Colombia  M T

Callithomia hezia villula (Hewitson, 1858)  TL: Colombia  M T

Callithomia hezia wellingi R. Fox, 1968  SW Mexico (S Oaxaca; Sierra Madre del Sur)  M T S

Callithomia hezia [n. ssp.#1] Lamas, MS  TL: Colombia

Callithomia hezia [n. ssp.#2] Lamas, MS  TL: Colombia

Callithomia hydra C. Felder & R. Felder, 1865  TL: Venezuela

Callithomia hydra hydra C. Felder & R. Felder, 1865  TL: Venezuela  M T

Callithomia hydra megaleas Godman & Salvin, 1898  [Panama], W Costa Rica  M T S

Callithomia hydra panamensis Godman & Salvin, 1878  Panama  M T S1 S2

Callithomia lenea (Cramer, 1779)    iN

Callithomia lenea lenea (Cramer, 1779)  TL: Surinam  M T

Callithomia lenea agrippina (Hewitson, 1863)  Panama to Colombia  M T S

Callithomia lenea alpho (C. Felder & R. Felder, 1865)  TL: Venezuela  M T

Callithomia lenea epidero (H. Bates, 1862)  TL: Brazil ([Pa])  M T

Callithomia lenea fumantis (Haensch, 1909)  TL: Colombia  M T

Callithomia lenea inturna (R. Fox, 1941)  TL: Peru  M T

Callithomia lenea methonella (Weymer, 1875)  TL: Brazil (SC)  M T L

Callithomia lenea nikita (Haensch, 1909)  TL: Venezuela  M T

Callithomia lenea obfuscata (A. Butler, 1873)  TL: Brazil (Am)  M T

Callithomia lenea pulcheria (Hewitson, 1870)  TL: Ecuador  M T

Callithomia lenea skinneri Masters, 1973  TL: Venezuela

Callithomia lenea siparia (Kaye, 1922)  TL: Trinidad  M T

Callithomia lenea travassosi R.F. d'Almeida, 1958  TL: Brazil (MT)  M T

Callithomia lenea xantho (C. Felder & R. Felder, 1860)  TL: Brazil (Ba)  M T

Callithomia lenea zelie (Guérin-Méneville, [1844])  TL: Bolivia  M T S

Callithomia lenea bella Neild, 2008  ref.266

Callithomia lenea [n. ssp.#1] Lamas, MS  TL: Venezuela

Callithomia lenea [n. ssp.#2] Lamas, MS  TL: Venezuela

Dircenna  E. Doubleday, 1847  

Dircenna adina (Hewitson, [1855])    iN

Dircenna adina adina (Hewitson, [1855])  TL: Venezuela  M T S

Dircenna adina columbiana E. Krüger, 1925  TL: Colombia  M T

Dircenna adina lorica Weymer, 1875  TL: Guyana  M T S

Dircenna adina marica (C. Felder & R. Felder, 1865)  TL: Venezuela  M T S

Dircenna adina steinheili Staudinger, [1884]  TL: Colombia  M T S

Dircenna adina stevei Neild, 2008  ref.266

Dircenna adina suna Haensch, 1903  TL: Ecuador  M T S

Dircenna adina willmotti Neild, 2008  ref.266

Dircenna adina xanthophane Hopffer, 1874  TL: Peru  M T S

Dircenna adina [n. ssp.#1] Lamas, MS  TL: Brazil (Rr)

Dircenna adina [n. ssp.#2] Lamas, MS  TL: Colombia

Dircenna adina [n. ssp.#3] Lamas, MS  TL: Colombia

Dircenna adina [n. ssp.#4] Lamas, MS  TL: Peru

Dircenna dero (Hübner, 1823)    iN

Dircenna dero Northern segregate ,  SE Mexico & Guatemala  M S

Dircenna dero dero (Hübner, 1823)  TL: Surinam  M S

Dircenna dero caliginosa J. Zikán, 1941  TL: Brazil (Am)  M T L

Dircenna dero celtina Burmeister, 1878  TL: ?  M T

Dircenna dero christopheri Neild, 2008  ref.266

Dircenna dero euchytma (C. Felder & R. Felder, 1865)  Nicaragua to Colombia & Venezuela  M T S L

Dircenna dero euteles Erschoff, 1875  TL: French Guiana  M S

Dircenna dero rhoeo C. Felder & R. Felder, 1860  TL: Brazil ([RJ])  M T S

Dircenna dero [n. ssp.#1] Lamas, MS  TL: Mexico

Dircenna dero [n. ssp.#2] Lamas, MS  TL: Peru

Dircenna dero [n. ssp.#3] Lamas, MS  TL: Brazil (Pa)

Dircenna jemina (Geyer, 1837)    iN

Dircenna jemina Northern segregate ,  E Mexico to Panama  M S

Dircenna jemina jemina (Geyer, 1837)  TL: Venezuela  M T S

Dircenna jemina chiriquensis Haensch, 1909  Costa Rica & Panama (Pacific slope)  M T S

Dircenna jemina reynoldsi Neild, 2008  ref.266

Dircenna jemina varina (Hewitson, 1869)  TL: Ecuador  M T S

Dircenna jemina visina Haensch, 1903  TL: Ecuador  M T S

Dircenna jemina [n. ssp.#1] Lamas, MS  TL: Guatemala

Dircenna jemina [n. ssp.#2] Lamas, MS  TL: Panama

Dircenna jemina [n. ssp.#3] Lamas, MS  TL: Colombia

Dircenna jemina [n. ssp.#4] Lamas, MS  TL: Colombia

Dircenna jemina [n. ssp.#5] Lamas, MS  TL: Colombia

Dircenna jemina [n. ssp.#6] Lamas, MS  TL: Colombia

Dircenna jemina [n. ssp.#7] Lamas, MS  TL: Ecuador

Dircenna klugii (Geyer, 1837)  Klug’s Clearwing    iN

Dircenna klugii klugii (Geyer, 1837)  S TX (very rare stray), E & W Mexico to Nicaragua  M S L

Dircenna klugii Southern segregate ,  Costa Rica & Panama  M S L

Dircenna klugii [n. ssp.] Lamas, MS  Costa Rica

Dircenna loreta Haensch, 1903    iN

Dircenna loreta loreta Haensch, 1903  TL: Ecuador  M T

Dircenna loreta acreana R.F. d'Almeida, 1950  TL: Brazil (Ac)  M T

Dircenna loreta melini Bryk, 1953  TL: Peru  M T

Dircenna loreta [n. ssp.] Lamas, MS  Brazil (Pa)

Dircenna olyras (C. Felder & R. Felder, 1865)    iN

Dircenna olyras Panamanian segregate ,  Panama  M S L

Dircenna olyras olyras (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Dircenna olyras bairdii Reakirt, 1868  TL: Colombia  M T

Dircenna olyras relata A. Butler & H. Druce, 1872  Nicaragua to Panama  M T S L

Dircenna olyras [n. ssp.#1] Lamas, MS  TL: Panama

Dircenna olyras [n. ssp.#2] Lamas, MS  TL: Ecuador

Hyalenna  W. Forbes, 1942  

Hyalenna paradoxa (Staudinger, [1884])    iN

Hyalenna paradoxa paradoxa (Staudinger, [1884])  TL: Colombia  M T S

Hyalenna paradoxa praestigiosa (Haensch, 1903)  TL: Ecuador  M T S

Hyalenna paradoxa catenata (Kaye, 1918)  TL: Peru  M T

Hyalenna paradoxa incachaca Lamas & Willmott, 2006  ref.332

Hyalenna hugia (Schaus, 1902)  TL: Bolivia  M T S

Hyalenna perasippa (Hewitson, 1877)  TL: Ecuador

Hyalenna perasippa valencia Willmott & Lamas, 2006  ref.332  M S

Hyalenna perasippa ortygiosa Willmott & Lamas, 2006  ref.332  M S

Hyalenna perasippa perasippa (Hewitson, 1877)  TL: Ecuador  M T

Hyalenna perasippa solitaria Lamas & Willmott, 2006  ref.332  M T

Hyalenna buckleyi Willmott & Lamas, 2006  TL: Ecuador

Hyalenna buckleyi buckleyi Willmott & Lamas, 2006  ref.332  M T

Hyalenna buckleyi pomacocha Lamas & Willmott, 2006  ref.332

Hyalenna pascua (Schaus, 1902)    iN  M T S

Hyalenna alidella (Hewitson, 1869)  TL: Colombia

Hyalenna alidella exsulans Lamas & Willmott, 2006  Panama (Darien); ref.332  M T

Hyalenna alidella vesca Lamas & Willmott, 2006  ref.332  M T S

Hyalenna alidella alidella (Hewitson, 1869)  TL: Colombia  M T S

Hyalenna alidella cinereola Lamas & Willmott, 2006  ref.332  M S

Hyalenna alidella scantilla (Hewitson, 1877)  TL: Ecuador  M T S

Hyalenna alidella dirama (Haensch, 1905)  TL: Bolivia  M T

Hyalenna alidella minna (Schaus, 1902)  TL: Bolivia  M T S

Hyalenna sulmona (Hewitson, 1877)    iN

Hyalenna sulmona tersa Willmott & Lamas, 2006  ref.332  M T

Hyalenna sulmona hyalina Lamas & Willmott, 2006  ref.332  M T S

Hyalenna sulmona lobusa (Haensch, 1909)  TL: Colombia  M T

Hyalenna sulmona sulmona (Hewitson, 1877)  TL: Ecuador  M T S

Hyalenna sulmona balsamica Lamas & Willmott, 2006  ref.332  M T

Episcada  Godman & Salvin, 1879  

Episcada apuleia (Hewitson, 1868)    iN

Episcada apuleia apuleia (Hewitson, 1868)  TL: Ecuador  M T

Episcada apuleia santanella (Haensch, 1903)  TL: Ecuador  M T

Episcada apuleia cora Haensch, 1909  TL: Bolivia  M T

Episcada apuleia [n. ssp.] Lamas & Willmott, MS  Peru

Episcada clausina (Hewitson, 1876)    iN

Episcada clausina clausina (Hewitson, 1876)  TL: Bolivia  M T S

Episcada clausina bomboiza Willmott & Lamas, 2021  ref.318

Episcada doto (Hübner, [1806])  TL: Brazil ("RJ")

Episcada doto doto (Hübner, [1806])  TL: Brazil ("RJ")  M T

Episcada doto paquito Neild, 2008  ref.266

Episcada doto [n. ssp.] Lamas & Willmott, MS  Peru

Episcada hemixanthe (C. Felder & R. Felder, 1865)    iN

Episcada hymen Haensch, 1905  TL: Bolivia

Episcada hymen hymen Haensch, 1905  TL: Bolivia  M T

Episcada hymen [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Episcada hymen [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Episcada hymenaea (Prittwitz, 1865)    iN

Episcada hymenaea hymenaea (Prittwitz, 1865)  TL: Brazil (RJ)  M T S

Episcada hymenaea caucana (K. Brown, 1970)  TL: Colombia

Episcada hymenaea centralis (K. Brown & O. Mielke, 1970)  TL: Brazil (MT)  M T

Episcada hymenaea guianensis Neild, 2008  ref.266

Episcada hymenaea mirifica Lamas & Willmott, 2021  ref.318

Episcada [n. sp.#1] Lamas & Willmott, MS  TL: not specified

Episcada [n. sp.#1] [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Episcada [n. sp.#1] [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Episcada [n. sp.#1] [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Episcada [n. sp.#1] [n. ssp.#4] Lamas & Willmott, MS  TL: Bolivia

Episcada mira (Hewitson, 1877)    iN  M T

Episcada philoclea (Hewitson, [1855])    iN  M T S

Episcada pichita Lamas & Willmott, 2020  ref.317  iN

Episcada polita Weymer, 1899    iN

Episcada polita polita Weymer, 1899  TL: Colombia  M T

Episcada polita cabenis Haensch, 1905  TL: Colombia  M T

Episcada polita aprica Neild, 2008  ref.266

Episcada polita raymondi Neild, 2008  ref.266

Episcada polita archaia Neild, 2008  ref.266

Episcada polita pichincha Willmott & Lamas, 2021  ref.318

Episcada polita flava Winhard, 2021  ref.318

Episcada [n. sp.#2] Lamas & Willmott, MS  TL: Peru

Episcada salvinia (H. Bates, 1864)  Salvin's Clearwing    iN

Episcada salvinia salvinia (H. Bates, 1864)  E Mexico, Guatemala  M T L

Episcada salvinia apia (C. Felder & R. Felder, 1865)  TL: Colombia  M T L

Episcada salvinia opleri Lamas, 1978  Costa Rica & Panama  M T S L

Episcada salvinia portilla J. Maza & Lamas, 1978  W Mexico (NAY to OAX)  M T S

Episcada striposis Haensch, 1909    iN  M T S

Episcada sulphurea Haensch, 1905  TL: Bolivia  M T S

Episcada arcadia Lamas & Willmott, 2021  ref.318

Episcada arcadia arcadia Lamas & Willmott, 2021  ref.318

Episcada arcadia navarro Lamas & Willmott, 2021  ref.318

Episcada sylpha Haensch, 1905  TL: Venezuela  M T S

Episcada sylvo (Geyer, 1832)  ref.319  iN  M T S

Episcada ticidella (Hewitson, 1869)    iN

Episcada ticidella ticidella (Hewitson, 1869)  TL: Ecuador  M T

Episcada ticidella [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Episcada ticidella [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Episcada trapezula Brabant & Bischler, 2003  TL: Peru  M S

Episcada vitrea R.F. d'Almeida & O. Mielke, 1967  TL: Brazil (RJ)  M T

Episcada zajciwi R.F. d'Almeida & O. Mielke, 1967    iN

Episcada zajciwi zajciwi R.F. d'Almeida & O. Mielke, 1967  TL: Brazil (Ce)  M T

Episcada zajciwi canaria (K. Brown & R.F. d'Almeida, 1970)  TL: Brazil (ES)  M T S

Episcada zajciwi [n. ssp.] Lamas & Willmott, MS  Brazil (Ce)

Episcada [n. sp.#3] Lamas & Willmott, MS  TL: Peru

Haenschia  Lamas, 2004  

Haenschia derama (Haensch, 1905)    iN  M T

Haenschia sidonia (Haensch, 1905)  TL: Peru  M T

Haenschia [n. sp.#1] Lamas, MS  TL: Peru

Haenschia [n. sp.#2] Lamas, MS  TL: Peru

Pteronymia  A. Butler & H. Druce, 1872  

Pteronymia alcmena (Godman & Salvin, 1877)  Alcmena Clearwing    iN

Pteronymia alcmena alcmena (Godman & Salvin, 1877)  S Mexico (Chiapas), Guatemala, Honduras, Nicaragua  M T S L

Pteronymia alcmena Panamanian segregate ,  Panama  M S

Pteronymia alcmena [n. ssp.] Lamas & Willmott, MS  Panama

Pteronymia aletta (Hewitson, [1855])    iN

Pteronymia aletta agalla Godman & Salvin, 1879  Costa Rica & Panama (Pacific slope)  M T S

Pteronymia aletta Panamanian segregate ,  Panama  M S

Pteronymia aletta aletta (Hewitson, [1855])  TL: Venezuela  M T S

Pteronymia aletta lilla (Hewitson, 1856)  TL: Ecuador  M T

Pteronymia aletta [n. ssp.#1] Lamas & Willmott, MS  TL: Panama

Pteronymia aletta [n. ssp.#2] Willmott & Lamas, MS  TL: Colombia

Pteronymia aletta [n. ssp.#3] Willmott & Lamas, MS  TL: Colombia

Pteronymia alicia Neild, 2008  ref.266  M S

Pteronymia alida (Hewitson, 1855)    iN

Pteronymia alida alida (Hewitson, 1855)  TL: ?  M T S

Pteronymia alida dispaena (Hewitson, 1876)  TL: Bolivia  M T S

Pteronymia alida zabina (Hewitson, [1857])  TL: Colombia  M T

Pteronymia alida georgei Neild, 2008  ref.266

Pteronymia alida cosanga Willmott & Lamas, 2021  ref.318

Pteronymia alida francisca Willmott & Lamas, 2021  ref.318

Pteronymia alida luctuosa Willmott & Lamas, 2021  ref.318

Pteronymia alida [n. ssp.#1] Lamas & Willmott, MS  TL: Colombia

Pteronymia alida [n. ssp.#2] Willmott & Lamas, MS  TL: Ecuador

Pteronymia alida [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Pteronymia alida [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Pteronymia alida [n. ssp.#5] Lamas & Willmott, MS  TL: Peru

Pteronymia alina Haensch, 1909    iN

Pteronymia alina alina Haensch, 1909  TL: Bolivia  M T

Pteronymia alina machay T. Racheli & L. Racheli, 2003  ref.286

Pteronymia alina mielkei Lamas, 2003  TL: Peru  M T

Pteronymia alina sumaq T. Racheli & L. Racheli, 2003  ref.286

Pteronymia alina [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pteronymia alina [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia alina [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Pteronymia alina [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Pteronymia alissa (Hewitson, 1869)    iN

Pteronymia alissa alissa (Hewitson, 1869)  TL: Ecuador  M T L

Pteronymia alissa amandes Kaye, 1921  TL: Trinidad  M T S

Pteronymia alissa [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Pteronymia alissa [n. ssp.#2] Willmott & Lamas, MS  TL: Colombia

Pteronymia andreas (Weeks, 1901)  TL: Bolivia

Pteronymia andreas andreas (Weeks, 1901)  TL: Bolivia  M T L

Pteronymia andreas [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia artena (Hewitson, [1855])  Artena Clearwing    iN

Pteronymia artena praedicta J. Maza & Lamas, 1982  SW Mexico (GRO, S OAX)  M T S

Pteronymia artena artena (Hewitson, [1855])  E Mexico, Guatemala, Honduras  M T

Pteronymia artena Costa Rican segregate ,  Costa Rica  M S

Pteronymia artena Panamanian segregate ,  Panama  M S L

Pteronymia artena beebei R. Fox & J. Fox, 1947  TL: Venezuela  M T S

Pteronymia artena afrania (Hopffer, 1874)  TL: Bolivia  M T L

Pteronymia artena [n. ssp.#1] Lamas & Willmott, MS  TL: Costa Rica

Pteronymia artena [n. ssp.#2] Lamas & Willmott, MS  TL: Panama

Pteronymia artena [n. ssp.#3] Willmott & Lamas, MS  TL: Colombia

Pteronymia artena [n. ssp.#4] Willmott & Lamas, MS  TL: Colombia

Pteronymia asopo (C. Felder & R. Felder, 1865)  TL: Venezuela  M T S

Pteronymia calgiria Schaus, 1902    iN  M T S

Pteronymia mariannae Lamas, Willmott & J. Hall, 2020  ref.317

Pteronymia mariannae mariannae Lamas, Willmott & J. Hall, 2020  ref.317

Pteronymia mariannae chisinche Willmott & Lamas, 2020  ref.317

Pteronymia carlia Schaus, 1902    iN  M T

Pteronymia cotytto (Guérin-Méneville, [1844])  Cotytto Clearwing    iN

Pteronymia cotytto cotytto (Guérin-Méneville, [1844])  E & S Mexico to Nicaragua, [Costa Rica?]  M T S L

Pteronymia cotytto Panamanian segregate ,  Panama  M S

Pteronymia cotytto [n. ssp.] Lamas & Willmott, MS  Panama

Pteronymia cuneata Willmott, Lamas & J. Hall, 2020  ref.317

Pteronymia dispar Haensch, 1905  TL: Colombia  M T

Pteronymia donella (C. Felder & R. Felder, 1865)    iN

Pteronymia donella Panamanian segregate ,  Costa Rica & Panama  M S

Pteronymia donella donella (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Pteronymia donella donata Haensch, 1909  Costa Rica & Panama (Pacific slope)  M T S

Pteronymia donella mariagabrielae Neild & De Sousa, 2008  ref.266

Pteronymia donella [n. ssp.] Lamas & Willmott, MS  Panama

Pteronymia dorothyae Neild, 2008  TL: Venezuela

Pteronymia dorothyae dorothyae Neild, 2008  ref.266

Pteronymia dorothyae marjorieae Neild, 2008  ref.266

Pteronymia euritea (Cramer, 1780)    iN  M S

Pteronymia [n. sp.#1] Lamas & Willmott, MS  TL: Panama

Pteronymia forsteri Baumann, 1985    iN  M T S

Pteronymia fulvimargo A. Butler & H. Druce, 1872  Fulvimargo Clearwing  Costa Rica & W Panama  M T S

Pteronymia fumida Schaus, 1913  Fumida Clearwing    iN

Pteronymia fumida fumida Schaus, 1913  Costa Rica & Panama  M T S

Pteronymia fumida pacifica E. Krüger, 1925  TL: Colombia

Pteronymia fumida ortizi T. Racheli & L. Racheli, 2003  ref.286

Pteronymia gertschi R. Fox, 1945  TL: Ecuador

Pteronymia gertschi gertschi R. Fox, 1945  TL: Ecuador  M T

Pteronymia gertschi [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia glauca Haensch, 1903    iN  M T

Pteronymia granica (Hewitson, 1877)  TL: Ecuador

Pteronymia granica granica (Hewitson, 1877)  TL: Ecuador  M T

Pteronymia granica [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia [n. sp.#2] Lamas & Willmott, MS  TL: Mexico (Gue)

Pteronymia hara (Hewitson, 1877)    iN

Pteronymia hara Costa Rican segregate ,  Costa Rica  M S

Pteronymia hara Panamanian segregate ,  Panama  M S

Pteronymia hara hara (Hewitson, 1877)  TL: Ecuador  M T

Pteronymia hara brunneata Bryk, 1937  TL: Ecuador  M T

Pteronymia hara semonis Haensch, 1909  TL: Ecuador  M T

Pteronymia hara ancha Neild, 2008  ref.266

Pteronymia hara albida Neild, 2008  ref.266

Pteronymia hara estrecha Neild, 2008  ref.266

Pteronymia hara olerina Willmott & Lamas, 2021  ref.318

Pteronymia hara [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia inania Haensch, 1903  TL: Ecuador

Pteronymia inania inania Haensch, 1903  TL: Ecuador  M T

Pteronymia inania evonia Haensch, 1905  TL: Colombia  M T S

Pteronymia inania bethana Willmott & Lamas, 2021  ref.318

Pteronymia inania [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia latilla (Hewitson, [1855])    iN

Pteronymia latilla fulvescens Godman & Salvin, 1879  Costa Rica & W Panama  M T S L

Pteronymia latilla Panamanian segregate ,  Panama  M S L

Pteronymia latilla latilla (Hewitson, [1855])  TL: Venezuela  M T S

Pteronymia latilla barilla Haensch, 1903  TL: Ecuador  M T

Pteronymia latilla nigricans Bryk, 1937  TL: Colombia

Pteronymia latilla [n. ssp.] Lamas & Willmott, MS  Colombia

Pteronymia laura (Staudinger, 1885)    iN  M T

Pteronymia lonera (A. Butler & H. Druce, 1872)  Lonera Clearwing  Costa Rica & W Panama  M T S

Pteronymia medellina Haensch, 1905    iN

Pteronymia medellina medellina Haensch, 1905  TL: Colombia  M T S

Pteronymia medellina [n. ssp.] Willmott & Lamas, MS  Colombia

Pteronymia obscuratus (Fabricius, 1793)  Obscure Clearwing  Panama  M T S

Pteronymia olimba Haensch, 1905  TL: Peru  M T

Pteronymia oneida (Hewitson, 1855)    iN

Pteronymia oneida oneida (Hewitson, 1855)  TL: Colombia  M T

Pteronymia oneida lisae Salazar, 1992  TL: Colombia

Pteronymia oneida sebastiani Neild, 2008  ref.266

Pteronymia oneida merida Neild, 2008  ref.266  M S

Pteronymia oneida ricardi Neild, 2008  ref.266

Pteronymia oneida [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pteronymia oneida [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia ozia (Hewitson, 1870)    iN

Pteronymia ozia ozia (Hewitson, 1870)  TL: Ecuador  M T

Pteronymia ozia browni R. Fox, 1945  TL: Ecuador  M T

Pteronymia ozia tanampaya Haensch, 1909  TL: Bolivia  M T S

Pteronymia parva (Salvin, 1869)  Parva Clearwing  [S Mexico?], Costa Rica & Panama  iN  M T S

Pteronymia peteri Neild, 2008  ref.266

Pteronymia picta (Salvin, 1869)    iN

Pteronymia picta notilla A. Butler & H. Druce, 1872  Costa Rica to Panama  M T S

Pteronymia picta Panamanian segregate ,  Panama  M S

Pteronymia picta picta (Salvin, 1869)  TL: Colombia  M T

Pteronymia picta [n. ssp.#1] Lamas & Willmott, MS  TL: Panama

Pteronymia picta [n. ssp.#2] Lamas & Willmott, MS  TL: Colombia

Pteronymia primula (H. Bates, 1862)    iN

Pteronymia primula primula (H. Bates, 1862)  TL: Brazil (Am)  M T

Pteronymia primula auricula Haensch, 1905  TL: Colombia  M T S

Pteronymia primula crocea Neild, 2008  ref.266

Pteronymia primula [n. ssp.] Lamas & Willmott, MS  Bolivia

Pteronymia rufocincta (Salvin, 1869)  West-Mexican Clearwing  W Mexico (NAY to OAX)  iN  M T S L

Pteronymia sao (Hübner, [1813])    iN

Pteronymia sao sao (Hübner, [1813])  TL: [Brazil]

Pteronymia sao antisao (H. Bates, 1862)  TL: [Brazil (Am)]  M T S

Pteronymia sao guntheri Lamas, 1985  TL: Peru  M T S

Pteronymia sao loreto Lamas & Willmott, 2021  ref.318

Pteronymia sao [n. ssp.#1] Lamas & Willmott, MS  TL: Brazil (Pa)

Pteronymia sao [n. ssp.#2] Lamas & Willmott, MS  TL: Bolivia

Pteronymia serrata Haensch, 1903    iN

Pteronymia serrata serrata Haensch, 1903  TL: Ecuador  M T

Pteronymia serrata amplificata Kaye, 1918  TL: Ecuador  M T

Pteronymia serrata triangula Winhard, 2019  ref.360

Pteronymia serrata [n. ssp.] Lamas & Willmott, MS  Peru

Pteronymia sexpunctata (Bryk, 1953)  TL: Peru

Pteronymia sexpunctata sexpunctata (Bryk, 1953)  TL: Peru  M T

Pteronymia sexpunctata [n. ssp.#1] Willmott & Lamas, MS  TL: Colombia

Pteronymia sexpunctata [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia simplex (Salvin, 1869)  Simple Clearwing    iN

Pteronymia simplex simplex (Salvin, 1869)  Costa Rica & Panama  M T S L

Pteronymia simplex fenochioi Lamas, 1978  E Mexico to Nicaragua  M T S L

Pteronymia simplex timagenes Godman & Salvin, 1889  SW Mexico (GRO, S OAX)  M T S

Pteronymia tamina Haensch, 1909    iN

Pteronymia tamina tamina Haensch, 1909  TL: Peru  M T

Pteronymia tamina [n. ssp.] Willmott & Lamas, MS  Ecuador

Pteronymia teresita (Hewitson, 1863)    iN  M T

Pteronymia thabena (Hewitson, 1869)    iN

Pteronymia thabena thabena (Hewitson, 1869)  TL: Ecuador  M T

Pteronymia thabena denticulata Haensch, 1905  TL: Bolivia  M T S

Pteronymia thabena [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pteronymia thabena [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia ticida (Hewitson, 1869)    iN

Pteronymia ticida ticida (Hewitson, 1869)  TL: Ecuador  M T

Pteronymia ticida citrina Neild, 2008  ref.266

Pteronymia ticida yungava Haensch, 1905  TL: Bolivia  M T S

Pteronymia ticida melanoptera (Hewitson, 1877)  TL: Ecuador  M T S

Pteronymia ticida [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pteronymia ticida [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia ticida [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Pteronymia tucuna (H. Bates, 1862)    iN  M T S

Pteronymia veia (Hewitson, [1853])    iN

Pteronymia veia veia (Hewitson, [1853])  TL: ?  M T S

Pteronymia veia brunnea Bryk, 1937  TL: ?  M T S

Pteronymia veia florea Neild, 2008  ref.266  M L

Pteronymia veia linzera (Herrich-Schäffer, 1865)  TL: Colombia  M T S

Pteronymia veia pomacocha T. Racheli & L. Racheli, 2003  TL: Peru

Pteronymia veia [n. ssp.] Lamas & Willmott, MS  Colombia

Pteronymia luisa Willmott, Lamas & J. Hall, 2020  Ecuador; ref.317

Pteronymia vestilla (Hewitson, [1853])    iN

Pteronymia vestilla vestilla (Hewitson, [1853])  TL: Brazil (Am)  M T

Pteronymia vestilla sparsa Haensch, 1903  TL: Ecuador  M T

Pteronymia vestilla ucaya Haensch, 1909  TL: Peru  M T

Pteronymia vestilla [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pteronymia vestilla [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Pteronymia zerlina (Hewitson, [1856])    iN

Pteronymia zerlina zerlina (Hewitson, [1856])  TL: Colombia  M T

Pteronymia zerlina amarillana Neild, 2008  ref.266

Pteronymia zerlina cristalina Neild, 2008  ref.266

Pteronymia zerlina nubivaga R. Fox & J. Fox, 1947  TL: Venezuela  M T S

Pteronymia zerlina pronuba (Hewitson, 1870)  TL: Ecuador  M T

Pteronymia zerlina [n. ssp.] Lamas & Willmott, MS  Colombia

Velamysta  Haensch, 1909  

Velamysta peninna (Hewitson, 1855)    iN

Velamysta peninna peninna (Hewitson, 1855)  TL: Bolivia  M T

Velamysta peninna [n. ssp.] Lamas, MS  Peru

Velamysta phengites R. Fox, 1945  TL: Ecuador

Velamysta phengites phengites R. Fox, 1945  TL: Ecuador  M T

Velamysta phengites perlucens Winhard, 2017  ref.317

Velamysta phengites [n. ssp.#1] Lamas, MS  TL: Colombia

Velamysta phengites [n. ssp.#2] Lamas, MS  TL: Peru

Velamysta phengites [n. ssp.#3] Lamas, MS  TL: Peru

Velamysta pupilla (Hewitson, 1874)    iN

Velamysta pupilla pupilla (Hewitson, 1874)  TL: Bolivia  M T

Velamysta pupilla anomala (Staudinger, [1884])  TL: Colombia  M T

Velamysta cruxifera (Hewitson, 1877)  TL: Ecuador

Velamysta cruxifera cruxifera (Hewitson, 1877)  ref.335  M T S

Velamysta cruxifera greeneyi Vitale & Bollino, 2003  ref.335

Velamysta desmondi Kell & Willmott, 2020  ref.317  iN

Godyris  Boisduval, 1870  

Godyris crinippa (Hewitson, 1874)    iN

Godyris crinippa crinippa (Hewitson, 1874)  TL: Bolivia  M T

Godyris crinippa [n. ssp.] Lamas, MS  Peru

Godyris dircenna (C. Felder & R. Felder, 1865)    iN  M T

Godyris duillia (Hewitson, 1854)    iN  M T S

Godyris hewitsoni (Haensch, 1903)  TL: ["Bolivia"]

Godyris hewitsoni hewitsoni (Haensch, 1903)  TL: ["Bolivia"]  M T

Godyris hewitsoni [n. ssp.#1] Lamas, MS  TL: Colombia

Godyris hewitsoni [n. ssp.#2] Lamas, MS  TL: Peru

Godyris hewitsoni [n. ssp.#3] Lamas, MS  TL: Peru

Godyris hewitsoni [n. ssp.#4] Lamas, MS  TL: Peru

Godyris kedema (Hewitson, [1855])    iN

Godyris kedema furina (Godman & Salvin, [1879])  Panama  M T S L

Godyris kedema Panamanian segregate ,  Panama  M S

Godyris kedema kedema (Hewitson, [1855])  TL: Venezuela  M T S

Godyris kedema albinotata (A. Butler, 1873)  TL: Colombia  M T

Godyris kedema [n. ssp.] Lamas, MS  Panama

Godyris lauta (Haensch, 1910)  TL: [Ecuador]

Godyris lauta lauta (Haensch, 1910)  TL: [Ecuador]  M T

Godyris lauta [n. ssp.] Lamas, MS  Peru

Godyris nepos (Weymer, 1875)    iN  M T

Godyris nero (Hewitson, [1855])  Nero Clearwing    iN

Godyris nero nero (Hewitson, [1855])  E Mexico to Nicaragua  M T S L

Godyris nero Southern segregate ,  Costa Rica  M S

Godyris nero [n. ssp.] Lamas, MS  Costa Rica

Godyris panthyale (C. Felder & R. Felder, 1862)    iN

Godyris panthyale panthyale (C. Felder & R. Felder, 1862)  TL: Colombia  M T S

Godyris panthyale quinta (Staudinger, 1885)  TL: Colombia  M T

Godyris panthyale [n. ssp.] Lamas, MS  Peru

Godyris sappho (Haensch, 1910)  TL: Bolivia

Godyris sappho sappho (Haensch, 1910)  TL: Bolivia  M T

Godyris sappho tika (T. Racheli & L. Racheli, 2003)  TL: Peru

Godyris sappho [n. ssp.] Lamas, MS  Peru

Godyris zavaleta (Hewitson, [1855])    iN

Godyris zavaleta sosunga (Reakirt, [1866])  SE Mexico to Nicaragua  M S L

Godyris zavaleta caesiopicta (Niepelt, 1915)  [= sorites], Costa Rica & Panama (Atlantic slope)  M T S1 S2 L

Godyris zavaleta zygia (Godman & Salvin, 1877)  Costa Rica & Panama (Pacific slope)  M T OI S

Godyris zavaleta Panamanian s segregate ,  Panama  M S L

Godyris zavaleta zavaleta (Hewitson, [1855])  TL: [Colombia]  M T

Godyris zavaleta christiani Neild, 2008  ref.266

Godyris zavaleta eutelina Brévignon, 1993  TL: French Guiana  M S

Godyris zavaleta gonussa (Hewitson, [1856])  TL: Colombia  M T

Godyris zavaleta huallaga R. Fox, 1941  TL: Peru  M T

Godyris zavaleta matronalis (Weymer, 1883)  TL: Ecuador  M T S

Godyris zavaleta petersii (Dewitz, 1877)  TL: Colombia  M T

Godyris zavaleta rosata Vitale & G. Rodriguez, 2004  ref.298

Godyris zavaleta telesilla (Hewitson, 1863)  TL: Ecuador  M T

Godyris zavaleta [n. ssp.#1] Lamas, MS  TL: Panama

Godyris zavaleta [n. ssp.#2] Lamas, MS  TL: Panama

Godyris zavaleta [n. ssp.#3] Lamas, MS  TL: Peru

Godyris zavaleta [n. ssp.#4] Lamas, MS  TL: Peru

Godyris zavaleta [n. ssp.#5] Lamas, MS  TL: Peru

Godyris zavaleta [n. ssp.#6] Lamas, MS  TL: Peru

Godyris zavaleta [n. ssp.#7] Lamas, MS  TL: Peru

Godyris zavaleta [n. ssp.#8] Lamas, MS  TL: Bolivia

Godyris zavaleta [n. ssp.#9] Lamas, MS  TL: Brazil (Rd)

Pachacutia  Willmott & Lamas, 2007  

Pachacutia baroni Willmott & Lamas, 2007  ref.333  M S

Pachacutia mantura (Hewitson, 1876)  TL: Bolivia

Pachacutia mantura joroni Lamas & Willmott, 2007  ref.333  M T

Pachacutia mantura honrathi (Srnka, 1885)  TL: Peru  M T

Pachacutia mantura mantura (Hewitson, 1876)  TL: Bolivia  M T

Pachacutia cleomella (Hewitson, 1874)  TL: Bolivia  M T

Pachacutia germaini Lamas & Willmott, 2007  ref.333  M T S

Veladyris  R. Fox, 1945  

Veladyris electrea (M. Brabant, 2004)  TL: Peru

Veladyris cytharista (Hewitson, 1874)    iN  M T S

Veladyris pardalis (Salvin, 1869)    iN

Veladyris pardalis pardalis (Salvin, 1869)  TL: Ecuador  M T

Veladyris pardalis aurea Lamas, 1980  TL: Ecuador  M T S L

Veladyris pardalis christina Lamas, 1980  TL: Peru  M T L

Veladyris pardalis totumbra (Kaye, 1919)  TL: Peru  M T

Veladyris pardalis [n. ssp.] Lamas & Vitale, MS  Peru

Veladyris [n. sp.] Lamas & Vitale, MS  Peru

Hypoleria  Godman & Salvin, 1879  

Hypoleria adasa (Hewitson, [1855])    iN

Hypoleria adasa adasa (Hewitson, [1855])  TL: Brazil  M T

Hypoleria adasa parcilimbata R.F. d'Almeida, 1943  TL: Brazil (SP)  M T

Hypoleria alema (Hewitson, [1857])    iN

Hypoleria alema alema (Hewitson, [1857])  TL: Colombia  M T

Hypoleria alema cajona Haensch, 1905  TL: Peru  M T S L

Hypoleria alema chrysodonia (H. Bates, 1862)  TL: Brazil (Am)  M T S L

Hypoleria alema consimilis Talbot, 1928  TL: Brazil (MT)  M T

Hypoleria alema garleppi Haensch, 1905  TL: Peru  M T

Hypoleria alema ina (Hewitson, [1859])  TL: Ecuador  M T

Hypoleria alema inconspicua Neild, 2008  ref.266

Hypoleria alema indecora Haensch, 1905  TL: Guyana  M T

Hypoleria alema karschi Haensch, 1903  TL: Ecuador  M T

Hypoleria alema niclae Neild, 2008  ref.266

Hypoleria alema oreas Weymer, 1899  TL: Brazil (ES)  M T

Hypoleria alema pastaza Willmott & Lamas, 2021  ref.318

Hypoleria alema proxima Weymer, 1899  TL: Brazil (SP)  M T

Hypoleria alema [n. ssp.#1] Lamas, MS  TL: Brazil (Am)

Hypoleria alema [n. ssp.#2] Lamas, MS  TL: Brazil (Rd)

Hypoleria alema [n. ssp.#3] Lamas, MS  TL: French Guiana

Hypoleria asellia (Hopffer, 1874)  TL: Peru  M T

Hypoleria aureliana (H. Bates, 1862)  TL: Brazil (Am)  M T

Hypoleria lavinia (Hewitson, [1855])    iN

Hypoleria lavinia lavinia (Hewitson, [1855])  TL: [Ecuador]  M T

Hypoleria lavinia cassotis (H. Bates, 1864)  S Mexico to Honduras  M T S L

Hypoleria lavinia libera Godman & Salvin, 1879  Panama  M T S L

Hypoleria lavinia meridana R. Fox, 1948  TL: Venezuela  M T S

Hypoleria lavinia rhene (Godman & Salvin, 1878)  Nicaragua to W Panama  M T S1 S2 L

Hypoleria lavinia riffarthi Haensch, 1905  TL: Ecuador  M T

Hypoleria lavinia vanilia (Herrich-Schäffer, 1865)  Panama to Colombia  M T S L

Hypoleria lavinia vaniliana Kaye, 1919  TL: Colombia  M T

Hypoleria lavinia aquilonalis Winhard, 2019  ref.360

Hypoleria mulviana R.F. d'Almeida, 1958  TL: Brazil (Pa)

Hypoleria ocalea (E. Doubleday, 1847)    iN

Hypoleria ocalea ocalea (E. Doubleday, 1847)  TL: Venezuela; "Brazil"  M T S

Hypoleria ocalea gephira (Hewitson, [1856])  TL: Colombia  M T

Hypoleria ocalea [n. ssp.#1] Lamas, MS  TL: Colombia

Hypoleria ocalea [n. ssp.#2] Lamas, MS  TL: Colombia

Hypoleria sarepta (Hewitson, [1852])    iN

Hypoleria sarepta sarepta (Hewitson, [1852])  TL: [Brazil (Am)]  M T S

Hypoleria sarepta cidonia (Hewitson, [1857])  TL: Colombia  M T S

Hypoleria sarepta famina Haensch, 1909  TL: Peru  M T

Hypoleria sarepta goiana R.F. d'Almeida, 1951  TL: Brazil (Go)  M T S

Hypoleria sarepta olerioides R.F. d'Almeida, 1951  TL: Brazil (Rd)  M T

Hypoleria sarepta oriana (Hewitson, [1859])  TL: Brazil (Am)  M T

Hypoleria sarepta orianula Bryk, 1953  TL: Peru

Hypoleria sarepta virginia (Hewitson, [1853])  TL: Brazil (Am)  M T

Hypoleria sarepta vitiosa Lamas, 1985  TL: Peru  M T L

Hypoleria sarepta [n. ssp.#1] Lamas, MS  TL: Peru

Hypoleria sarepta [n. ssp.#2] Lamas, MS  TL: Peru

Hypoleria xenophis Haensch, 1909  TL: Peru  M T

Brevioleria  Lamas, 2004  

Brevioleria aelia (Hewitson, 1852)    iN

Brevioleria aelia aelia (Hewitson, 1852)  TL: Brazil (Am)  M T

Brevioleria aelia brevicula (R.F. d'Almeida, 1951)  TL: Peru  M T

Brevioleria aelia jamariensis (R.F. d'Almeida, 1951)  TL: Brazil (Rd)  M T L

Brevioleria aelia orolina (Hewitson, [1861])  TL: Brazil (Am)  M T

Brevioleria aelia pachiteae (Tessmann, 1928)  TL: Peru  M T L

Brevioleria aelia plisthenes (R.F. d'Almeida, 1958)  TL: Brazil (MG)  M T

Brevioleria aelia yawara (J. Zikán, 1942)  TL: Brazil (Am)  M T

Brevioleria aelia [n. ssp.] Lamas, MS  Colombia

Brevioleria arzalia (Hewitson, 1876)    iN

Brevioleria arzalia arzalia (Hewitson, 1876)  TL: Bolivia  M T

Brevioleria arzalia loronia Lamas & Willmott, 2001  ref.318

Brevioleria coenina (Hewitson, 1869)  TL: Ecuador  M T

Brevioleria seba (Hewitson, 1872)    iN

Brevioleria seba seba (Hewitson, 1872)  TL: ?  M T

Brevioleria seba emyra (Haensch, 1905)  TL: Brazil (MG)  M T

Brevioleria seba oculata (Haensch, 1903)  TL: Ecuador  M T

Brevioleria seba [n. ssp.#1] Lamas, MS  TL: Peru

Brevioleria seba [n. ssp.#2] Lamas, MS  TL: Peru

Brevioleria seba [n. ssp.#3] Lamas, MS  TL: Peru

Brevioleria [n. sp.] Lamas, MS  Brazil (Pa)

Mcclungia  R. Fox, 1940  

Mcclungia cymo (Hübner, [1806])    iN

Mcclungia cymo cymo (Hübner, [1806])  TL: Brazil  M T

Mcclungia cymo arcuata (Tessmann, 1928)  TL: Peru  M T S

Mcclungia cymo collenettei (Talbot, 1928)  TL: Brazil (MT)  M T

Mcclungia cymo fallens (Haensch, 1905)  TL: Brazil (MG)  M T

Mcclungia cymo salonina (Hewitson, 1855)  TL: Bolivia  M T L

Mcclungia cymo subtilis (Haensch, 1903)  TL: Ecuador  M T L

Mcclungia cymo trepotis (Haensch, 1910)  TL: Bolivia  M T S

Mcclungia cymo wana (A. Hall, 1930)  TL: Guyana  M T S

Mcclungia cymo [n. ssp.] Lamas, MS  Brazil (Rd)

Mcclungia cymo iguazu Winhard, 2020  ref.361

Greta  Hemming, 1934  

Greta andromica (Hewitson, [1855])    iN

Greta andromica lyra (Salvin, 1869)  E Mexico to Panama  M T S L

Greta andromica Panamanian segregate ,  Panama  M S

Greta andromica andromica (Hewitson, [1855])  TL: Venezuela  M T S L

Greta andromica andania (Hopffer, 1874)  TL: Peru  M T S L

Greta andromica dromica (Haensch, 1910)  TL: Colombia  M T

Greta andromica nerina (Haensch, 1905)  TL: Colombia  M T

Greta andromica trifenestra R. Fox, 1941  TL: Trinidad  M T

Greta andromica [n. ssp.] Lamas & Willmott, MS  Peru

Greta annette (Guérin-Méneville, [1844])  White-spotted Clearwing    iN

Greta annette annette (Guérin-Méneville, [1844])  E Mexico, Guatemala, Honduras  M T S L

Greta annette championi Lamas, 1978  Nicaragua to Panama  M T S L

Greta annette moschion (Godman, 1901)  W Mexico (NAY to S OAX)  M T S L Ha

Greta clavijoi Neild, 2008  ref.266  M S

Greta cubana (Herrich-Schäffer, 1862)  Cuban Clearwing  Cuba  iN  M T S I

Greta diaphanus (Drury, 1773)    iN  M T S1 S2 L I

Greta morgane (Geyer, 1837)  Rusty Clearwing    iN

Greta morgane morgane (Geyer, 1837)  W Mexico (NAY to S OAX)  M S L

Greta morgane oto (Hewitson, [1855])  E Mexico to Panama  M T S L1 L2

Greta [n. sp.] Lamas & Willmott, MS  Peru

Hypomenitis  R. Fox, 1945  

Hypomenitis alphesiboea (Hewitson, 1869)  ref.184  iN  M T S

Hypomenitis candida Lamas & Willmott, 2020  ref.317

Hypomenitis [n. sp.#1] Lamas & Willmott, MS  TL: Peru

Hypomenitis [n. sp.#2] Lamas & Willmott, MS  TL: Peru

Hypomenitis depauperata (Boisduval, 1870)    iN

Hypomenitis depauperata depauperata (Boisduval, 1870)  ref.184  M T S

Hypomenitis depauperata umbrosa (Haensch, 1903)  ref.184  M T

Hypomenitis dercetis (E. Doubleday, 1847)    iN

Hypomenitis dercetis dercetis (E. Doubleday, 1847)  ref.184  M T S

Hypomenitis dercetis [n. ssp.] Lamas & Willmott, MS  Peru

Hypomenitis enigma (Haensch, 1905)  TL: Bolivia

Hypomenitis enigma enigma (Haensch, 1905)  ref.184  M T S

Hypomenitis enigma koechlini (Lamas, 2003)  ref.184  M T

Hypomenitis enigma pseudortygia (Neild & Willmott, 2008)  ref.266  M T

Hypomenitis enigma [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hypomenitis enigma [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hypomenitis enigma [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hypomenitis enigma [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Hypomenitis jamesiana Willmott, Lamas & J. Hall, 2020  ref.317

Hypomenitis esula (Hewitson, 1855)  TL: Colombia

Hypomenitis esula esula (Hewitson, 1855)  ref.184  M T

Hypomenitis esula [n. ssp.] Lamas & Willmott, MS  Colombia

Hypomenitis [n. sp.#3] Lamas & Willmott, MS  TL: Peru

Hypomenitis gabiglooris Brabant & Bischler, 2005  Gabi's Clearwing  Nicaragua to W Panama; ref.184  iN  M S

Hypomenitis gardneri (Weeks, 1901)    iN

Hypomenitis gardneri gardneri (Weeks, 1901)  ref.184  M T S

Hypomenitis gardneri devriesi (D'Abrera, 1984)  ref.184  M T

Hypomenitis gardneri [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hypomenitis gardneri [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hypomenitis gardneri [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hypomenitis gardneri [n. ssp.#4] Lamas & Willmott, MS  TL: Peru

Hypomenitis hermana (Haensch, 1903)    iN

Hypomenitis hermana hermana (Haensch, 1903)  ref.184  M T

Hypomenitis hermana joiceyi (Kaye, 1918)  ref.184  M T

Hypomenitis hermana [n. ssp.] Willmott & Lamas, MS  Colombia

Hypomenitis hermana serafina Willmott & Lamas, 2021  ref.318

Hypomenitis hermana chamba Willmott & Lamas, 2021  ref.318

Hypomenitis libethris (C. Felder & R. Felder, 1865)    iN

Hypomenitis libethris libethris (C. Felder & R. Felder, 1865)  ref.184  M T S L

Hypomenitis libethris [n. ssp.] Willmott & Lamas, MS  Colombia

Hypomenitis lojana Vitale & Bollino, 2001  ref.184  M T

Hypomenitis lydia (Weymer, 1899)  [= flava]; ref.184  iN  M T

Hypomenitis ochretis (Haensch, 1903)    iN

Hypomenitis ochretis ochretis (Haensch, 1903)  ref.184  M T

Hypomenitis ochretis [n. ssp.] Willmott & Lamas, MS  Colombia

Hypomenitis oneidodes (Kaye, 1918)  TL: Ecuador

Hypomenitis oneidodes oneidodes (Kaye, 1918)  ref.184  M T S

Hypomenitis oneidodes guarumales Boyer, Petit & Willmott, 2021  ref.318

Hypomenitis oneidodes nicolasi Willmott & Lamas, 2021  ref.318

Hypomenitis ortygia (Weymer, 1890)    iN

Hypomenitis ortygia ortygia (Weymer, 1890)  [= panthyaloides]; ref.184  M T

Hypomenitis ortygia pyrczi Vitale & Bollino, 2001  ref.184

Hypomenitis ortygia [n. ssp.#1] Lamas & Willmott, MS  TL: Colombia

Hypomenitis ortygia [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hypomenitis ortygia [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hypomenitis [n. sp.#4] Willmott & Lamas, MS  TL: Colombia

Hypomenitis polissena (Hewitson, 1863)    iN

Hypomenitis polissena polissena (Hewitson, 1863)  ref.184  M T

Hypomenitis polissena colombiana Vitale & Bollino, 2001  ref.184

Hypomenitis polissena umbrana (Haensch, 1908)  Costa Rica & Panama; ref.184  M T S L

Hypomenitis polissena bolivariana Neild, 2008  ref.266

Hypomenitis polissena [n. ssp.] Lamas & Willmott, MS  Colombia

Hypomenitis theudelinda (Hewitson, [1861])    iN

Hypomenitis theudelinda theudelinda (Hewitson, [1861])  ref.184  M T

Hypomenitis theudelinda bonita Vitale & Bollino, 2001  ref.184

Hypomenitis theudelinda cyrcilla (Hewitson, 1874)  ref.184  M T

Hypomenitis theudelinda zalmunna (Hewitson, 1869)  ref.184  M T L

Hypomenitis theudelinda [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Hypomenitis theudelinda [n. ssp.#2] Lamas & Willmott, MS  TL: Peru

Hypomenitis theudelinda [n. ssp.#3] Lamas & Willmott, MS  TL: Peru

Hypomenitis [n. sp.#5] Lamas & Willmott, MS  Undescribed Clearwing (Costa Rica)  Costa Rica

Heterosais  Godman & Salvin, 1880  

Heterosais edessa (Hewitson, [1855])    iN  M T S

Heterosais giulia (Hewitson, [1855])    iN

Heterosais giulia cadra (Godman & Salvin, 1878)  Nicaragua to Panama  M T S

Heterosais giulia Panamanian segregate ,  Panama  M S L

Heterosais giulia giulia (Hewitson, [1855])  TL: Colombia  M T S

Heterosais giulia pallidula Haensch, 1903  TL: Ecuador  M T

Heterosais giulia caucaensis Constantino, Gil & Valencia, 2005  TL: Colombia  M T

Heterosais giulia [n. ssp.#1] Lamas, MS  TL: Colombia

Heterosais giulia [n. ssp.#2] Lamas, MS  TL: Brazil (Am)

Heterosais nephele (H. Bates, 1862)  TL: Brazil (Am)

Heterosais nephele nephele (H. Bates, 1862)  TL: Brazil (Am)  M T S

Heterosais nephele gedera (Hewitson, 1869)  TL: [Colombia]  M T

Pseudoscada  Godman & Salvin, 1879  

Pseudoscada acilla (Hewitson, 1867)    iN

Pseudoscada acilla acilla (Hewitson, 1867)  TL: Brazil (MG/[RJ])  M T

Pseudoscada acilla quadrifasciata Talbot, 1928  TL: Brazil (MT)  M T

Pseudoscada acilla [n. ssp.] Lamas & Willmott, MS  Brazil (Rd)

Pseudoscada erruca (Hewitson, 1855)    iN  M T S

Pseudoscada florula (Hewitson, [1855])    iN

Pseudoscada florula florula (Hewitson, [1855])  TL: French Guiana  M T

Pseudoscada florula aureola (H. Bates, 1862)  TL: Brazil (Am)  M T S V

Pseudoscada florula exornata Haensch, [1908]  TL: Guyana  M T

Pseudoscada florula genetyllis (R.F. d'Almeida, 1922)  TL: Brazil (RJ)  M T S

Pseudoscada florula gracilis Tessmann, 1928  TL: Peru  M T S V

Pseudoscada florula [n. ssp.#1] Lamas & Willmott, MS  TL: Peru

Pseudoscada florula [n. ssp.#2] Lamas & Willmott, MS  TL: Brazil (Rd)

Pseudoscada timna (Hewitson, [1855])    iN

Pseudoscada timna timna (Hewitson, [1855])  TL: Venezuela  M T S L

Pseudoscada timna utilla (Hewitson, 1856)  TL: Colombia  M T

Pseudoscada pusio (Godman & Salvin, 1877)  Timna Clearwing  Nicaragua to Panama  iN  M T S L

Pseudoscada troetschi (Staudinger, [1884])    iN

Pseudoscada troetschi troetschi (Staudinger, [1884])  TL: Colombia  M T

Pseudoscada troetschi saturata (Staudinger, 1885)  Panama and Colombia  M T S L

Pseudoscada troetschi colombiana Winhard, 2020  ref.237,361

Pseudoscada troetschi litoralis Winhard, 2020  ref.236,361

  Subfamily Heliconiinae

  Tribe Acraeini

Actinote  Hübner, [1819]  

Actinote abana (Hewitson, 1868)  ref.12  iN

Actinote abana abana (Hewitson, 1868)  TL: Ecuador  M T

Actinote abana capnodes Jordan, 1913  TL: Colombia  M T

Actinote erinome (C. Felder & R. Felder, 1861)  ref.12  iN

Actinote erinome erinome (C. Felder & R. Felder, 1861)  TL: Ecuador  M T S

Actinote erinome mathani Oberthür, 1917  TL: Peru  M T

Actinote erinome testacea (Salvin & Godman, 1868)  TL: Bolivia  M T S

Actinote erinome [n. ssp.] Lamas, MS  Ecuador

Actinote euryleuca Jordan, 1910  ref.12  M T L

Actinote hylonome (E. Doubleday, 1844)  ref.12  iN

Actinote hylonome hylonome (E. Doubleday, 1844)  TL: Colombia  M T

Actinote hylonome basilia (Lamas, 1995)  TL: Venezuela

Actinote roseus Winhard, 2017  ref.12,Winhard_2017

Actinote intensa Jordan, 1910  ref.12  M T S

Actinote johncoulsoni Willmott, Lamas & J. Hall, 2017  ref.12

Actinote alcione (Hewitson, 1852)  ref.12  iN

Actinote alcione alcione (Hewitson, 1852)  TL: Ecuador  M T S

Actinote alcione corduba (Hewitson, 1874)  TL: Bolivia  M T S

Actinote alcione elatus H. Druce, 1903  TL: Ecuador  M T

Actinote alcione flaugeri Neild & Romero, 2008  TL: Venezuela

Actinote alcione miguelangeli Neild & Romero, 2008  TL: Venezuela

Actinote alcione sodalis A. Butler, 1877  TL: Peru  M T S L

Actinote alcione theophila (Dognin, 1887)  TL: Ecuador  M T

Actinote alcione varians Jordan, 1910  TL: Colombia  M T

Actinote anaxo (Hopffer, 1874)  ref.12  iN

Actinote anaxo anaxo (Hopffer, 1874)  TL: Peru  M T

Actinote anaxo comta Jordan, 1910  TL: Peru  M T

Actinote anaxo [n. ssp.] Lamas, MS  Peru

Actinote callianthe (C. Felder & R. Felder, 1862)  ref.12  iN

Actinote callianthe callianthe (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Actinote callianthe alla Jordan, 1910  TL: Venezuela  M T

Actinote callianthe amoena Jordan, 1913  TL: Colombia  M T

Actinote callianthe borgesi Neild & Romero, 2008  TL: Venezuela

Actinote callianthe rufa Jordan, 1910  TL: Ecuador  M T

Actinote dicaeus (Latreille, [1817])  ref.12  iN

Actinote dicaeus dicaeus (Latreille, [1817])  TL: ["Peru"]  M T

Actinote dicaeus albofasciata (Hewitson, 1869)  TL: Ecuador  M T S L

Actinote dicaeus amida (Hewitson, 1854)  TL: Colombia  M T

Actinote dicaeus callianira Geyer, 1837  TL: Mexico?  M T

Actinote dicaeus carica (Weymer, 1890)  TL: Peru  M T

Actinote dicaeus eulalia Oberthür, 1917  TL: Colombia  M T

Actinote dicaeus euris Jordan, 1910  TL: Peru  M T S

Actinote dicaeus jucunda Jordan, 1910  TL: Bolivia  M T

Actinote dicaeus laverna (E. Doubleday, [1847])  TL: Venezuela  M T

Actinote dicaeus luisi Neild & Romero, 2008  TL: Venezuela

Actinote dicaeus ozinta (Schaus, 1902)  TL: Venezuela  M T

Actinote eresia (C. Felder & R. Felder, 1862)  ref.12  iN

Actinote eresia eresia (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Actinote eresia binghamae Dyar, 1913  TL: Peru  M T

Actinote eresia eresina (Hopffer, 1874)  TL: Bolivia  M T

Actinote eresia lacrymosa Oberthür, 1917  TL: Colombia  M T

Actinote eresia leptogramma Jordan, 1913  TL: Ecuador  M T

Actinote eresia [n. ssp.#1] Lamas, MS  TL: Peru

Actinote eresia [n. ssp.#2] Lamas, MS  TL: Peru

Actinote eresia [n. ssp.#3] Lamas, MS  TL: Peru

Actinote griseata A. Butler, 1873  TL: Peru  M T

Actinote hilaris Jordan, 1910  ref.12  iN

Actinote hilaris hilaris Jordan, 1910  TL: Peru  M T S L

Actinote hilaris desmiala Jordan, 1913  TL: Ecuador  M T S

Actinote hilaris [n. ssp.] Lamas, MS  Peru

Actinote momina Jordan, 1910  ref.12  M T

Actinote negra (C. Felder & R. Felder, 1862)  ref.12  iN

Actinote negra negra (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T L

Actinote negra avuncula Oberthür, 1917  TL: Peru  M T

Actinote negra demonica (Hopffer, 1874)  TL: Bolivia  M T S L

Actinote negra euclia (Dognin, 1887)  TL: Ecuador  M T S

Actinote negra menoetes Oberthür, 1917  TL: Ecuador  M T

Actinote negra scotosis Jordan, 1910  TL: Ecuador  M T

Actinote negra sobrina Jordan, 1910  TL: Peru  M T

Actinote negra [n. ssp.] Lamas, MS  Peru

Actinote neleus (Latreille, [1813])  ref.12  iN

Actinote neleus neleus (Latreille, [1813])  TL: [S. Amer.]  M T S

Actinote neleus bermudezi Neild & Romero, 2008  TL: Venezuela

Actinote ozomene (Godart, 1819)  ref.12  iN

Actinote ozomene ozomene (Godart, 1819)  TL: S. Amer.  M T

Actinote ozomene calimene Rebel, 1902  TL: [Colombia]  M T

Actinote ozomene carbonaria (Hopffer, 1874)  TL: Bolivia  M T

Actinote ozomene cleasa (Hewitson, 1868)  TL: Ecuador  M T

Actinote ozomene gabrielae Rebel, 1902  TL: Ecuador  M T

Actinote ozomene nox (H. Bates, 1864)  [= leucomelas], E & S Mexico to Panama  M T S1 S2 L I

Actinote rubrocellulata Hayward, 1960  ref.12  iN

Actinote rubrocellulata rubrocellulata Hayward, 1960  TL: Peru  M T S

Actinote rubrocellulata [n. ssp.] Lamas, MS  Peru

Actinote stratonice (Latreille, [1813])  ref.12  iN

Actinote stratonice stratonice (Latreille, [1813])  TL: [Colombia]  M T L

Actinote stratonice acipha (Hewitson, 1861)  TL: Colombia  M T

Actinote stratonice adoxa Jordan, 1910  TL: Colombia  M T

Actinote stratonice aereta Jordan, 1913  TL: Ecuador  M T

Actinote stratonice marthae Jordan, 1910  TL: Colombia  M T

Actinote stratonice meridana Jordan, 1910  TL: Venezuela  M T

Actinote stratonice oaxaca J. Miller & L. Miller, 1979  S Oaxaca (Sierra Madre del Sur)  M T S L

Actinote tenebrosa (Hewitson, 1868)  ref.12  iN

Actinote tenebrosa tenebrosa (Hewitson, 1868)  TL: Ecuador  M T S

Actinote tenebrosa erebia Oberthür, 1917  TL: Peru  M T L

Actinote tenebrosa [n. ssp.] Lamas, MS  Peru

Actinote trinacria (C. Felder & R. Felder, 1862)  ref.12  iN

Actinote trinacria trinacria (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Actinote trinacria chea H. Druce, 1903  TL: Colombia  M T

Actinote trinacria naura (H. Druce, 1875)  TL: Venezuela  M T

Actinote trinacria santamarta (Winhard, 2017)  ref.12,Winhard_2017

Actinote trinacria tenebrarum Oberthür, 1917  TL: Colombia  M T

Actinote trinacria vivasi Neild & Romero, 2008  TL: Venezuela

Actinote [n. sp.#1] Lamas, MS  TL: Peru

Actinote [n. sp.#2] Lamas, MS  TL: Peru

Actinote alalia (C. Felder & R. Felder, 1860)    iN  M T

Actinote alberti Neild & Romero, 2008  TL: Venezuela

Actinote alberti alberti Neild & Romero, 2008  ref.266  M T

Actinote alberti skinneri Romero & Neild, 2008  ref.266  M T

Actinote anteas (E. Doubleday, [1847])    iN

Actinote anteas Central American segregate ,  S Mexico to Panama  M S L

Actinote anteas anteas (E. Doubleday, [1847])  TL: Venezuela  M T

Actinote anteas brettia Oberthür, 1917  TL: Colombia  M T

Actinote anteas byssa Oberthür, 1917  TL: Venezuela  M T

Actinote anteas cedestis Jordan, 1913  TL: Ecuador  M T

Actinote anteas crassinia (Hopffer, 1874)  TL: Peru  M T S

Actinote anteas pierrei Neild, 2008  ref.266

Actinote anteas suspecta Jordan, 1913  TL: Ecuador  M T

Actinote anteas terpsinoe (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Actinote ballettae Neild & Romero, 2008  TL: Venezuela

Actinote ballettae ballettae Neild & Romero, 2008  ref.266  M T

Actinote ballettae bonyae Neild & Romero, 2008  ref.266

Actinote bonita Penz, 1996  TL: Brazil (MG)  M T

Actinote brylla Oberthür, 1917    iN  M T

Actinote canutia (Hopffer, 1874)    iN  M T

Actinote carycina Jordan, 1913    iN  M T S

Actinote catarina Penz, 1996    iN  M T

Actinote conspicua Jordan, 1913    iN  M T

Actinote dalmeidai Francini, 1996    iN  M T

Actinote discrepans R.F. d'Almeida, 1958    iN  M T

Actinote eberti Francini, Freitas & Penz, 2004  ref.205

Actinote genitrix R.F. d'Almeida, 1922    iN

Actinote genitrix genitrix R.F. d'Almeida, 1922  TL: Brazil ([RJ])  M T

Actinote genitrix costae Neild, 2008  ref.266

Actinote guatemalena (H. Bates, 1864)  Guatemalan Actinote    iN

Actinote guatemalena guatemalena (H. Bates, 1864)  Chiapas to Costa Rica  M T S L

Actinote guatemalena guerrerensis J. Maza, 1982  SW Mexico (Sierra Madre del Sur)  M T S

Actinote guatemalena veraecrucis Jordan, 1913  SE Mexico  M T S1 S2

Actinote kennethi Freitas, Willmott & J. Hall, 2009  ref.312  M S

Actinote lapitha (Staudinger, 1885)  Pale Actinote    iN

Actinote lapitha lapitha (Staudinger, 1885)  C Chiapas to Costa Rica & Panama  M T S

Actinote lapitha calderoni Schaus, 1920  W Chiapas to El Salvador  M T S

Actinote latior Jordan, 1913  TL: Colombia  M T

Actinote mamita (Burmeister, 1861)    iN

Actinote mamita mamita (Burmeister, 1861)  TL: Argentina  M T S

Actinote mamita elena A. Hall, 1921  TL: Colombia  M T

Actinote mamita mitama (Schaus, 1902)  TL: Brazil (Pr)  M T

Actinote melampeplos Godman & Salvin, 1881  Bow-winged Actinote  S Mexico (S Oaxaca), Ecuador  iN

Actinote melampeplos melampeplos Godman & Salvin, 1881  TL: Costa Rica  M T S L

Actinote melampeplos [n. ssp.] Lamas, MS  Ecuador

Actinote melanisans Oberthür, 1917    iN  M T

Actinote morio Oberthür, 1917  TL: Brazil (MG)  M T

Actinote parapheles Jordan, 1913    iN  M T

Actinote pellenea Hübner, [1821]    iN

Actinote pellenea pellenea Hübner, [1821]  TL: [Brazil]  M T

Actinote pellenea adriana (Hopffer, 1874)  TL: Bolivia  M T

Actinote pellenea calymma Jordan, 1913  TL: Paraguay  M T1 T2

Actinote pellenea crucis Jordan, 1913  TL: Bolivia  M T

Actinote pellenea epiphaea Jordan, 1913  TL: Peru  M T

Actinote pellenea equatoria (H. Bates, 1864)  TL: Ecuador  M T

Actinote pellenea fernandezi Neild & Romero, 2008  ref.266

Actinote pellenea hahneli Jordan, 1913  TL: Venezuela  M T

Actinote pellenea hyalina Jordan, 1913  TL: [Peru]  M T

Actinote pellenea limbata Jordan, 1913  TL: [Colombia]  M T

Actinote pellenea mucia (Hopffer, 1874)  TL: Peru  M T

Actinote pellenea perfulva Jordan, 1913  TL: Colombia  M T

Actinote pellenea subbadia Jordan, 1913  TL: Venezuela  M T

Actinote pellenea subhyalina (Staudinger, 1885)  TL: Peru  M T

Actinote pellenea trinitatis Jordan, 1913  TL: Trinidad  M T

Actinote pellenea [n. ssp.#1] Lamas, MS  TL: Ecuador

Actinote pellenea [n. ssp.#2] Lamas, MS  TL: Peru

Actinote pellenea [n. ssp.#3] Lamas, MS  TL: Peru

Actinote pellenea [n. ssp.#4] Lamas, MS  TL: Bolivia

Actinote pratensis Francini, Freitas & Penz, 2004  ref.205  iN

Actinote quadra (Schaus, 1902)    iN  M T

Actinote rhodope R.F. d'Almeida, 1923    iN  M T S

Actinote romeroi Neild & Costa, 2008  ref.266  M T

Actinote rufina Oberthür, 1917    iN  M T S

Actinote surima (Schaus, 1902)    iN

Actinote surima surima (Schaus, 1902)  TL: Brazil (Pr)  M T

Actinote surima perisa Jordan, 1913  TL: Argentina  M T

Actinote thalia (Linnaeus, 1758)    iN

Actinote thalia thalia (Linnaeus, 1758)  TL: Indiis  M T S

Actinote thalia eupelia Jordan, 1913  TL: Bolivia  M T

Actinote thalia pyrrha (Fabricius, 1775)  TL: Brazil ([RJ])  M T S

Actinote thalia sainorum Neild & Romero, 2008  ref.266  M S

Actinote zikani R.F. d'Almeida, 1951  TL: Brazil (SP)  M T

Actinote [n. sp.#3] Lamas, MS  TL: Peru

Actinote [n. sp.#4] Lamas, MS  TL: Peru

Actinote [n. sp.#5] Lamas, MS  TL: Peru

Actinote [n. sp.#6] Lamas, MS  TL: Peru

Actinote [n. sp.#7] Lamas, MS  TL: Peru

Actinote [n. sp.#8] Penz, MS  TL: Brazil

  Tribe Heliconiini

Dione  Hübner, [1819]  

Dione incarnata N. Riley, 1926  Gulf Fritillary  ref.11,88, iN.312743

Dione incarnata incarnata N. Riley, 1926  SW USA, C & S TX (strays far N) through C Amer.; HI; ref.11,88  M T S1 S2 L I H

Dione incarnata nigrior (Michener, 1942)  SE USA (strays far N); ref.11,88  iN  M T S1 S2 Ab HS L1 L2 I H

Dione insularis (Maynard, 1889)  Gulf Fritillary  Bahamas, West Indies except southern Lesser Antilles; ref.88  M T S L

Dione vanillae (Linnaeus, 1758)  Gulf Fritillary  S Lesser Antilles (St. Lucia, Barbados), Trinidad, N S. Amer. ...; ref.10,88  iN  M S HS L

Dione forbesi (Michener, 1942)  ref.88

Dione galapagensis (W. Holland, 1890)  ref.88  M T S

Dione lucina (C. Felder & R. Felder, 1862)  ref.88  M T

Dione maculosa Stichel, [1908]  ref.88  M T

Dione dodona Lamas & Farfán, 2022  Peru; ref.89  M T

Dione moneta Hübner, [1825]    iN

Dione moneta poeyii A. Butler, 1873  E & W Mexico (strays N into USA) to Panama  M T S L I H

Dione moneta butleri Stichel, [1908]  TL: Colombia  M T

Dione moneta moneta Hübner, [1825]  TL: ?

Dione glycera (C. Felder & R. Felder, 1861)    iN  M T S

Dione juno (Cramer, 1779)  Juno Longwing    iN

Dione juno huascuma (Reakirt, 1866)  E & W Mexico through Central America  M S Ab L1 L2 I

Dione juno juno (Cramer, 1779)  Lesser Antilles from Martinique southward, NE South America  M S

Dione juno andicola (H. Bates, 1864)  TL: Ecuador  M T

Dione juno miraculosa M. Hering, 1926  TL: Peru  M T L

Dione juno suffumata K. Brown & O. Mielke, 1972  TL: Paraguay

Podotricha  Michener, 1942  

Podotricha judith (Guérin-Méneville, [1844])    iN

Podotricha judith judith (Guérin-Méneville, [1844])  TL: Colombia  M T

Podotricha judith caucana (N. Riley, 1926)  TL: Colombia  M T

Podotricha judith mellosa (Stichel, 1906)  TL: Ecuador  M T L

Podotricha judith straminea (N. Riley, 1926)  TL: Ecuador

Podotricha telesiphe (Hewitson, 1867)    iN

Podotricha telesiphe telesiphe (Hewitson, 1867)  TL: Ecuador  M T S

Podotricha telesiphe tithraustes (Salvin, 1871)  TL: Ecuador  M T

Dryadula  Michener, 1942  

Dryadula phaetusa (Linnaeus, 1758)  Banded Longwing  S TX (rare stray), E & S Mexico to Brazil  iN  M T S HS L1 L2

Dryas  Hübner, [1807]  

Dryas iulia (Fabricius, 1775)  Julia    iN

Dryas iulia largo Clench, 1975  S FL (strays N)  M S L I

Dryas iulia nudeola (M. Bates, 1934)  Cuba  M T S L

Dryas iulia carteri (N. Riley, 1926)  N Bahamas  M T S L

Dryas iulia zoe L. Miller & Steinhauser, 1992  Grand Cayman & Cayman Brac  M T

Dryas iulia delila (Fabricius, 1775)  Jamaica  M S

Dryas iulia fucatus (Boddaert, 1783)  [= hispaniola], Hispaniola & satellite islands  M S1 S2 L

Dryas iulia iulia (Fabricius, 1775)  Mona, Puerto Rico, Tortola, St. John, St. Thomas, St. Croix  M T S L

Dryas dominicana (A. Hall, 1917)  Julia  Guadeloupe & Dominica; ref.88

Dryas dominicana warneri (A. Hall, 1936)  St. Kitts, Montserrat, Saba  M T S

Dryas dominicana dominicana (A. Hall, 1917)  Guadeloupe & Dominica  M T S

Dryas lucia (N. Riley, 1926)  Julia  St. Lucia; ref.88

Dryas lucia martinica Enrico & Pinchon, 1969  Martinique  M T S

Dryas lucia lucia (N. Riley, 1926)  St. Lucia  M T S

Dryas alcionea (Cramer, 1779)  ref.88

Dryas alcionea framptoni (N. Riley, 1926)  St. Vincent, some Grenadines, Grenada, [Barbados?]  M T S L

Dryas alcionea alcionea (Cramer, 1779)  TL: Surinam  M T S

Dryas alcionea moderata (N. Riley, 1926)  S TX (strays far N), E & W Mexico through Central America  M S L1 L2 I H Ha

Philaethria  Billberg, 1820  

Philaethria diatonica (Fruhstorfer, 1912)  Northern Green Longwing  [= Central American "dido" records], S Mexico to Panama (strays)  iN  M T S L I

Philaethria dido (Linnaeus, 1763)    iN

Philaethria dido panamensis Constantino & Salazar, 2010  TL: Panama  M T

Philaethria dido chocoensis Constantino, 1999  TL: Colombia  M T

Philaethria dido dido (Linnaeus, 1763)  TL: S. Amer.  M S HS

Philaethria constantinoi Salazar, 1991  TL: Colombia  M T S

Philaethria browni Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria romeroi Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria neildi Constantino & Salazar, 2010  TL: Venezuela

Philaethria neildi winhardi Constantino & Salazar, 2010  TL: Colombia  M T

Philaethria neildi neildi Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria neildi tachiraensis Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria ostara (Röber, 1906)    iN

Philaethria ostara araguensis Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria ostara meridensis Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria ostara ostara (Röber, 1906)  TL: Colombia  M S

Philaethria andrei Brévignon, 2002    iN

Philaethria andrei orinocensis Constantino & Salazar, 2010  TL: Venezuela  M T

Philaethria andrei andrei Brévignon, 2002  TL: French Guiana  M T

Philaethria pygmalion (Fruhstorfer, 1912)    iN

Philaethria pygmalion metaensis Constantino & Salazar, 2010  TL: Colombia  M T

Philaethria pygmalion pygmalion (Fruhstorfer, 1912)  TL: Brazil (Pa)  M T S

Philaethria wernickei (Röber, 1906)    iN  M S

Neruda  J. Turner, 1976  

Neruda aoede (Hübner, [1813])    iN

Neruda aoede aoede (Hübner, [1813])  TL: [Brazil]

Neruda aoede aliciae Neukirchen, 2000  TL: Ecuador  M T

Neruda aoede astydamia (Erichson, [1849])  TL: Guyana  M T

Neruda aoede auca Neukirchen, 1997  TL: Ecuador  M T

Neruda aoede ayacuchensis (Neukirchen, 1992)  TL: Venezuela  M T

Neruda aoede bartletti (H. Druce, 1876)  TL: Peru  M T

Neruda aoede centurius (Neukirchen, 1994)  TL: French Guiana  M T

Neruda aoede cupidineus (Stichel, 1906)  TL: Peru  M T

Neruda aoede emmelina (Oberthür, 1902)  TL: Guyana  M T

Neruda aoede eurycleia (K. Brown, 1973)  TL: Brazil (MT)  M T

Neruda aoede faleria (Fruhstorfer, 1910)  TL: Brazil (MT)

Neruda aoede lucretius (Weymer, 1891)  TL: Brazil (Am)  M T

Neruda aoede manu (Lamas, 1976)  TL: Peru

Neruda aoede philipi (K. Brown, 1976)  TL: Bolivia  M T

Neruda aoede [n. ssp.] K. Brown, MS  Brazil

Neruda godmani (Staudinger, 1882)  TL: Colombia  M T

Neruda metharme (Erichson, [1849])    iN

Neruda metharme metharme (Erichson, [1849])  TL: Guyana  M T S

Neruda metharme adelinae (Cast, 2019)  ref.182

Neruda metharme makiritare K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Neruda metharme perseis (Stichel, 1923)  TL: Colombia  M T

Neruda metis Moreira & O. Mielke, 2010  ref.248

Eueides  Hübner, 1816  

Eueides aliphera (Godart, 1819)    iN

Eueides aliphera aliphera (Godart, 1819)  TL: Brazil  M T S

Eueides aliphera cyllenella Seitz, 1912  TL: ?  M T

Eueides aliphera gracilis Stichel, 1903  E & S Mexico to Panama  M T S L Ha

Eueides emsleyi K. Brown, 1976  TL: Colombia

Eueides emsleyi emsleyi K. Brown, 1976  TL: Colombia

Eueides emsleyi esmeraldensis Cast, 2013  TL: Ecuador  M T

Eueides heliconioides C. Felder & R. Felder, 1861    iN

Eueides heliconioides heliconioides C. Felder & R. Felder, 1861  TL: Ecuador  M T S L

Eueides heliconioides eanes Hewitson, 1861  TL: Peru  M T

Eueides heliconioides koenigi H. Holzinger & R. Holzinger, 1993  TL: Peru

Eueides isabella (Stoll, 1781)    iN

Eueides isabella isabella (Stoll, 1781)  TL: Surinam

Eueides isabella arquata Stichel, 1903  TL: Colombia  M T

Eueides isabella cleobaea Geyer, 1832  Cuba  M S

Eueides isabella dianasa (Hübner, [1806])  TL: Brazil (Ba)  M T

Eueides isabella dissoluta Stichel, 1903  TL: Peru  M T

Eueides isabella dynastes C. Felder & R. Felder, 1861  TL: Venezuela; Colombia  M T

Eueides isabella ecuadorensis Strand, 1912  TL: Ecuador

Eueides isabella eva (Fabricius, 1793)  [= zoracon], E Mexico (strays far N) through Central America  M T S L1 L2 I

Eueides isabella hippolinus A. Butler, 1873  TL: Peru  M T

Eueides isabella huebneri Ménétriés, 1857  TL: Brazil  M T

Eueides isabella melphis (Godart, 1819)  Hispaniola  M T S

Eueides isabella nigricornis R. G. Maza, 1982  SW Mexico  M T

Eueides isabella [n. ssp.] K. Brown, MS  Brazil (Ce)

Eueides lampeto H. Bates, 1862    iN

Eueides lampeto lampeto H. Bates, 1862  TL: Brazil (Am)  M T

Eueides lampeto acacetes Hewitson, 1869  TL: Ecuador  M T

Eueides lampeto apicalis Röber, 1927  TL: Ecuador  M T

Eueides lampeto brownsbergensis Gernaat & Beckles, 2010  TL: Suriname

Eueides lampeto carbo Seitz, 1913  TL: Ecuador  M T

Eueides lampeto concisa Lamas, 1985  TL: Peru  M T

Eueides lampeto nigrofulva Kaye, 1906  TL: Guyana  M T

Eueides lampeto [n. ssp.#1] K. Brown, MS  TL: French Guiana

Eueides lampeto [n. ssp.#2] K. Brown, MS  TL: Brazil (Am)

Eueides libitina Staudinger, 1885    iN

Eueides libitina libitina Staudinger, 1885  TL: French Guiana  M T

Eueides libitina malleti Lamas, 1998  TL: Peru  M T

Eueides lineata Salvin & Godman, 1868  Thick-edged Longwing  S Mexico to Panama  iN  M T S L

Eueides lybia (Fabricius, 1775)    iN

Eueides lybia lybia (Fabricius, 1775)  TL: India

Eueides lybia lybioides Staudinger, 1876  Pacific slope of Costa Rica and Panama  M T S L

Eueides lybia olympia (Fabricius, 1793)  Atlantic slope from Nicaragua to Colombia  M T S L

Eueides lybia orinocensis K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Eueides lybia otelloi K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Eueides lybia salcedoi K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Eueides pavana Ménétriés, 1857    iN  M T

Eueides procula E. Doubleday, [1847]    iN

Eueides procula procula E. Doubleday, [1847]  TL: Venezuela  M T

Eueides procula asidia Schaus, 1920  SE Mexico to Nicaragua  M T S Ab L

Eueides procula browni H. Holzinger & R. Holzinger, 1974  TL: Venezuela

Eueides procula edias Hewitson, 1861  TL: Colombia  M T

Eueides procula eurysaces Hewitson, 1864  TL: Ecuador  M T

Eueides procula kuenowii Dewitz, 1877  TL: Colombia  M T

Eueides procula vulgiformis A. Butler & H. Druce, 1872  Costa Rica & Panama  M T S

Eueides tales (Cramer, 1775)    iN

Eueides tales tales (Cramer, 1775)  TL: Surinam

Eueides tales barcellinus J. Zikán, 1937  TL: Brazil (Am)  M T

Eueides tales calathus Stichel, 1909  TL: Ecuador  M T

Eueides tales cognata Weymer, 1890  TL: Colombia  M T

Eueides tales franciscus K. Brown & H. Holzinger, 1973  TL: Venezuela  M T

Eueides tales llanera Winhard, 2019  ref.360

Eueides tales michaeli J. Zikán, 1937  TL: Peru  M T

Eueides tales pseudeanes Boullet & Le Cerf, 1910  TL: Venezuela  M T

Eueides tales pythagoras W. F. Kirby, [1900]  TL: Brazil (Pa)  M T S

Eueides tales surdus Stichel, 1903  TL: Brazil (Pa); Guyana  M T

Eueides tales tabernula Lamas, 1985  TL: Peru

Eueides tales xenophanes C. Felder & R. Felder, 1865  TL: Colombia  M T

Eueides tales [n. ssp.#1] K. Brown, MS  TL: French Guiana

Eueides tales [n. ssp.#2] K. Brown, MS  TL: Brazil (Pa)

Eueides tales guyanensis Lacomme, 2011  TL: not specified

Eueides vibilia (Godart, 1819)    iN

Eueides vibilia vibilia (Godart, 1819)  TL: Brazil  M T

Eueides vibilia mereaui (Hübner, 1823)  TL: Brazil  M T

Eueides vibilia unifasciatus A. Butler, 1873  TL: Brazil (Am)  M T

Eueides vibilia vialis Stichel, 1903  SE Mexico to Panama  M T S L

Eueides vibilia vicinalis Stichel, 1903  TL: Ecuador  M T

Eueides vibilia [n. ssp.] K. Brown, MS  Colombia

Heliconius  Kluk, 1780  

Heliconius antiochus (Linnaeus, 1767)    iN

Heliconius antiochus antiochus (Linnaeus, 1767)  TL: Indiis  M T

Heliconius antiochus aranea (Fabricius, 1793)  TL: ?

Heliconius antiochus araneides Staudinger, 1897  TL: Venezuela  M T

Heliconius antiochus salvinii Dewitz, 1877  TL: [Venezuela]  M T

Heliconius antiochus [n. ssp.] Lamas, MS  Peru

Heliconius astraea Staudinger, 1897  TL: Brazil (Am)

Heliconius astraea astraea Staudinger, 1897  TL: Brazil (Am)  M T

Heliconius astraea rondonia K. Brown, 1973  TL: Brazil (Rd)  M T S

Heliconius atthis (E. Doubleday, 1847)    iN  M T L

Heliconius besckei (Ménétriés, 1857)    iN  M T

Heliconius burneyi (Hübner, [1831])    iN

Heliconius burneyi burneyi (Hübner, [1831])  TL: Brazil  M T

Heliconius burneyi ada Neustetter, 1925  TL: Brazil (Rd)  M T

Heliconius burneyi anjae Neukirchen, 1995  TL: Brazil (Am)  M T

Heliconius burneyi boliviensis Neukirchen, 1995  TL: Bolivia  M T

Heliconius burneyi catharinae Staudinger, 1885  TL: [Brazil (Pa)]  M T

Heliconius burneyi huebneri Staudinger, 1897  TL: [Peru]  M T

Heliconius burneyi jamesi Neukirchen, 1995  TL: Peru  M T

Heliconius burneyi koenigi Neukirchen, 1995  TL: Peru  M T

Heliconius burneyi lindigii C. Felder & R. Felder, 1865  TL: Colombia  M T

Heliconius burneyi skinneri K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius charithonia (Linnaeus, 1767)  Zebra Longwing    iN

Heliconius charithonia vazquezae W. Comstock & F. Brown, 1950  S TX (strays far N), E & W Mexico through Central America  M T S Ab L1 L2 I H Ha

Heliconius charithonia tuckeri W. Comstock & F. Brown, 1950  E TX to FL (strays N), Grand Bahama, Great Abaco  M T S L I

Heliconius charithonia ramsdeni W. Comstock & F. Brown, 1950  Cuba, Andros, [New Providence?]  M T S

Heliconius charithonia simulator Röber, 1921  Jamaica  M S L

Heliconius charithonia churchi W. Comstock & F. Brown, 1950  Hispaniola & satellite islands  M T S L

Heliconius charithonia charithonia (Linnaeus, 1767)  Mona, Puerto Rico, Virgin Islands, some Leeward Is. to Montserrat  M T S L

Heliconius charithonia antiquus Lamas, 1988  St Kitts & Antigua; as an occasional form on other islands  M T S

Heliconius charithonia bassleri W. Comstock & F. Brown, 1950  TL: Colombia  M T

Heliconius clysonymus Latreille, [1817]    iN

Heliconius clysonymus clysonymus Latreille, [1817]  TL: [Colombia]  M T

Heliconius clysonymus hygiana (Hewitson, 1867)  TL: Ecuador  M T

Heliconius clysonymus montanus Salvin, 1871  Costa Rica & Panama  M T TI L

Heliconius clysonymus tabaconas K. Brown, 1976  TL: Peru  M T

Heliconius congener Weymer, 1890    iN

Heliconius congener congener Weymer, 1890  TL: Ecuador  M T L

Heliconius congener aquilionaris K. Brown, 1976  TL: Colombia  M T

Heliconius congener ocannensis Stichel, 1905  TL: Colombia  M T

Heliconius cydno (E. Doubleday, 1847)    iN

Heliconius cydno cydno (E. Doubleday, 1847)  TL: Colombia  M T

Heliconius cydno alithea (Hewitson, 1869)  TL: Ecuador  M T L

Heliconius cydno barinasensis Masters, 1973  TL: Venezuela

Heliconius cydno chioneus H. Bates, 1864  SE Costa Rica to Colombia  M T S L

Heliconius cydno cordula Neustetter, 1913  TL: Venezuela  M T

Heliconius cydno cydnides Staudinger, 1885  TL: Colombia  M T

Heliconius cydno gadouae K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius cydno galanthus H. Bates, 1864  SE Mexico to E Costa Rica  M T S L

Heliconius cydno hermogenes (Hewitson, [1858])  TL: [Colombia]  M T

Heliconius cydno lisethae Neukirchen, 1995  TL: Colombia  M T

Heliconius cydno wanningeri Neukirchen, 1991  TL: Colombia  M T

Heliconius cydno weymeri Staudinger, 1897  TL: Colombia  M T Ab

Heliconius cydno zelinde A. Butler, 1869  TL: W coast of America  M T Ab

Heliconius pachinus Salvin, 1871  Pachinus Longwing  Pacific slope of Costa Rica and Panama  iN  M T S L I

Heliconius demeter Staudinger, 1897    iN

Heliconius demeter demeter Staudinger, 1897  TL: Peru  M T

Heliconius demeter angeli Neukirchen, 1997  TL: Peru

Heliconius demeter beebei J. Turner, 1966  TL: Guyana  M T

Heliconius demeter bouqueti Nöldner, 1901  TL: French Guiana  M T

Heliconius demeter karinae Neukirchen, 1990  TL: Brazil (Pa)  M T

Heliconius demeter neildi Neukirchen, 1997  TL: Ecuador  M T

Heliconius demeter terrasanta K. Brown & Benson, 1975  TL: Brazil (Pa)  M T

Heliconius demeter titan Neukirchen, 1995  TL: Brazil (Am)  M T

Heliconius demeter turneri K. Brown & Benson, 1975  TL: Brazil (Am)  M T

Heliconius demeter zikani K. Brown & Benson, 1975  TL: Brazil (Am)  M T

Heliconius demeter joroni Lamas & Rosser, 2018  Zoological Journal of the Linnean Society 186(1): 233-249)

Heliconius eratosignis (Joicey & Talbot, 1925)    iN

Heliconius eratosignis eratosignis (Joicey & Talbot, 1925)  ref.291  M T

Heliconius eratosignis tambopata Lamas, 1985  ref.291

Heliconius eratosignis ucayalensis H. Holzinger & R. Holzinger, 1975  ref.291

Heliconius eratosignis ulysses K. Brown & Benson, 1975  ref.291  M T

Heliconius doris (Linnaeus, 1771)    iN

Heliconius doris viridis Staudinger, 1885  SE Mexico to Panama  M T S L

Heliconius doris doris (Linnaeus, 1771)  TL: Surinam  M T

Heliconius doris dives (Oberthür, 1920)  TL: Colombia  M T

Heliconius doris obscurus Weymer, 1891  TL: Colombia  M T

Heliconius doris [n. ssp.] Neukirchen, MS  Trinidad

Heliconius egeria (Cramer, 1775)  TL: Surinam

Heliconius egeria egeria (Cramer, 1775)  TL: Surinam  M T

Heliconius egeria christiani Neukirchen, 1997  TL: Brazil (Pa)  M T

Heliconius egeria egerides Staudinger, 1897  TL: Brazil (Am)  M T

Heliconius egeria homogena Bryk, 1953  TL: Brazil (Am)  M T

Heliconius egeria hyas Weymer, 1883  TL: [Brazil (Am)]  M T

Heliconius egeria keithbrowni Neukirchen, 1992  TL: Venezuela  M T

Heliconius egeria mariasibyllae Neukirchen, 1991  TL: Brazil (Pa)  M T

Heliconius eleuchia (Hewitson, [1854])  White-edged Longwing    iN

Heliconius eleuchia eleuchia (Hewitson, [1854])  Costa Rica to Colombia  M T L1 L2 Ha

Heliconius eleuchia eleusinus Staudinger, 1885  TL: Colombia  M T

Heliconius eleuchia primularis A. Butler, 1869  TL: Ecuador  M T L

Heliconius elevatus Nöldner, 1901    iN

Heliconius elevatus elevatus Nöldner, 1901  TL: Amazonas  M T

Heliconius elevatus bari (Oberthür, 1902)  TL: French Guiana  M T

Heliconius elevatus lapis Lamas, 1976  TL: Peru

Heliconius elevatus perchlora Joicey & Kaye, 1917  TL: Bolivia  M T

Heliconius elevatus pseudocupidineus Neustetter, 1931  TL: Peru  M T

Heliconius elevatus roraima J. Turner, 1966  TL: Guyana  M T

Heliconius elevatus schmassmanni Joicey & Talbot, 1925  TL: Brazil (MT)  M T

Heliconius elevatus sonjae Neukirchen, 1997  TL: Brazil (Pa)  M T

Heliconius elevatus taracuanus Bryk, 1953  TL: Brazil (Am)

Heliconius elevatus tumatumari Kaye, 1906  TL: Guyana  M T

Heliconius elevatus willmotti Neukirchen, 1997  TL: Ecuador  M T

Heliconius elevatus zoelleri Neukirchen, 1990  TL: Venezuela  M T

Heliconius erato (Linnaeus, 1758)    iN

Heliconius erato erato (Linnaeus, 1758)  TL: Surinam  M T1 T2 HS

Heliconius erato adana J. Turner, 1967  TL: [Trinidad]  M T

Heliconius erato amalfreda Riffarth, 1901  TL: Brazil (Pa)  M T

Heliconius erato amazona Staudinger, 1897  TL: Brazil (Pa)  M T

Heliconius erato amphitrite Riffarth, 1901  TL: Peru  M T

Heliconius erato chestertonii (Hewitson, 1872)  TL: Colombia  M T

Heliconius erato colombina Staudinger, 1897  TL: Colombia  M T

Heliconius erato cruentus Lamas, 1998  W Mexico  M T S L

Heliconius erato cyrbia (Godart, 1819)  TL: America  M T

Heliconius erato demophoon Ménétriés, 1855  Nicaragua  M T OI S

Heliconius erato dignus Stichel, 1923  TL: Colombia  M T

Heliconius erato emma Riffarth, 1901  TL: Peru; "Ecuador"  M T

Heliconius erato estrella H. Bates, 1862  TL: Brazil (Pa)  M T

Heliconius erato etylus Salvin, 1871  TL: Ecuador  M T

Heliconius erato favorinus Hopffer, 1874  TL: Bolivia  M T

Heliconius erato guarica Reakirt, 1868  TL: Colombia  M T

Heliconius erato hydara (Hewitson, 1867)  Panama to Brazil  M T1 T2 L

Heliconius erato lativitta A. Butler, 1877  TL: Brazil (Am); Ecuador  M T

Heliconius erato lichyi K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius erato luscombei Lamas, 1976  TL: Peru

Heliconius erato magnifica Riffarth, 1900  TL: Guyana

Heliconius erato microclea Kaye, 1907  TL: Peru  M T

Heliconius erato notabilis Salvin & Godman, 1868  TL: Ecuador  M T

Heliconius erato petiverana (E. Doubleday, 1847)  S TX, E Mexico to Panama  M T S Ab L

Heliconius erato phyllis (Fabricius, 1775)  TL: Brazil  M T

Heliconius erato reductimacula Bryk, 1953  TL: Brazil (Am)

Heliconius erato tobagoensis Barcant, 1982  TL: Tobago  M T

Heliconius erato venus Staudinger, 1882  SE Panama (Darien), W Colombia (Choco)  M T L

Heliconius erato venustus Salvin, 1871  TL: Bolivia  M T

Heliconius ethilla (Godart, 1819)    iN

Heliconius ethilla ethilla (Godart, 1819)  TL: Antilles  M T

Heliconius ethilla adela Neustetter, 1912  TL: Peru  M T

Heliconius ethilla aerotome C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Heliconius ethilla cephallenia C. Felder & R. Felder, 1865  TL: Surinam  M T

Heliconius ethilla chapadensis K. Brown, 1973  TL: Brazil (MT)  M T

Heliconius ethilla claudia Godman & Salvin, 1881  Panama  M T S

Heliconius ethilla eucoma (Hübner, [1831])  TL: Brazil

Heliconius ethilla flavofasciatus Weymer, 1894  TL: Brazil (Pa)  M T

Heliconius ethilla flavomaculatus Weymer, 1894  TL: Brazil (Pe)  M T

Heliconius ethilla hyalina Neustetter, 1928  TL: Brazil (Rr)  M T

Heliconius ethilla jaruensis K. Brown, 1976  TL: Brazil (Rd)  M T

Heliconius ethilla latona Neukirchen, 1998  TL: Colombia  M T

Heliconius ethilla mentor Weymer, 1883  TL: Colombia  M T

Heliconius ethilla metalilis A. Butler, 1873  TL: Venezuela  M T

Heliconius ethilla michaelianius Lamas, 1988  TL: Peru  M T

Heliconius ethilla narcaea (Godart, 1819)  TL: Antilles  M T

Heliconius ethilla nebulosa Kaye, 1916  TL: Peru  M T

Heliconius ethilla neukircheni Lamas, 1998  TL: Peru  M T

Heliconius ethilla numismaticus Weymer, 1894  TL: Brazil (Pa)  M T

Heliconius ethilla penthesilea Neukirchen, 1994  TL: Brazil (Am)  M T

Heliconius ethilla polychrous C. Felder & R. Felder, 1865  TL: Brazil ([SP])  M T

Heliconius ethilla semiflavidus Weymer, 1894  TL: Colombia  M T

Heliconius ethilla thielei Riffarth, 1900  TL: French Guiana  M T

Heliconius ethilla tyndarus Weymer, 1897  TL: Bolivia  M T

Heliconius ethilla yuruani K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius hecale (Fabricius, 1776)    iN

Heliconius hecale hecale (Fabricius, 1776)  TL: Surinam  M T

Heliconius hecale anderida (Hewitson, [1853])  TL: [Venezuela]  M T

Heliconius hecale annetta Riffarth, 1900  TL: Colombia  M T

Heliconius hecale australis K. Brown, 1976  TL: Ecuador  M T

Heliconius hecale barcanti K. Brown, 1976  TL: Venezuela  M T

Heliconius hecale clearei A. Hall, 1930  TL: Guyana  M T

Heliconius hecale discomaculatus Weymer, 1891  Honduras  M T

Heliconius hecale ennius Weymer, 1891  TL: Brazil (Am)

Heliconius hecale felix Weymer, 1894  TL: Peru  M T

Heliconius hecale fornarina (Hewitson, 1854)  C & SW Chiapas to Guatemala  M T S

Heliconius hecale holcophorus Staudinger, 1897  TL: Colombia  M T

Heliconius hecale humboldti Neustetter, 1928  TL: Peru  M T

Heliconius hecale ithaca C. Felder & R. Felder, 1862  TL: Colombia  M T

Heliconius hecale latus Riffarth, 1900  TL: Brazil (Pa)  M T

Heliconius hecale melicerta H. Bates, 1866  Panama & Colombia  M T S L

Heliconius hecale metellus Weymer, 1894  TL: Brazil (Pa)  M T

Heliconius hecale naxos Neukirchen, 1998  TL: Brazil (Pa)

Heliconius hecale nigrofasciatus Weymer, 1894  TL: Brazil (Am)  M T S

Heliconius hecale novatus H. Bates, 1867  TL: Brazil (Pa/Ma)  M T

Heliconius hecale paraensis Riffarth, 1900  TL: Brazil (Pa)  M T

Heliconius hecale paulus Neukirchen, 1998  TL: Brazil (Pa)

Heliconius hecale quitalena (Hewitson, [1853])  TL: Ecuador  M T

Heliconius hecale rosalesi K. Brown & F. Fernández, 1976  TL: Venezuela  M T

Heliconius hecale shanki Lamas & K. Brown, 1976  TL: Peru

Heliconius hecale sisyphus Salvin, 1871  TL: Peru  M T

Heliconius hecale sulphureus Weymer, 1894  TL: Brazil (Am)  M T

Heliconius hecale vetustus A. Butler, 1873  TL: Guyana  M T

Heliconius hecale zeus Neukirchen, 1995  TL: Bolivia  M T

Heliconius hecale zuleika (Hewitson, 1854)  SE Mexico to Panama  M T S1 S2 S3 L I H

Heliconius hecalesia (Hewitson, [1854])    iN

Heliconius hecalesia hecalesia (Hewitson, [1854])  TL: Colombia

Heliconius hecalesia eximius Stichel, 1923  TL: Colombia

Heliconius hecalesia formosus H. Bates, 1866  Costa Rica & Panama  M T S L

Heliconius hecalesia gynaesia (Hewitson, 1875)  TL: ?  M T

Heliconius hecalesia longarena (Hewitson, 1875)  TL: Colombia  M T

Heliconius hecalesia octavia H. Bates, 1866  SE Mexico to Nicaragua  M T S L

Heliconius hecalesia romeroi K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius hecuba (Hewitson, [1858])    iN

Heliconius hecuba hecuba (Hewitson, [1858])  TL: Colombia  M T

Heliconius hecuba bonplandi Neukirchen, 1991  TL: Ecuador  M T

Heliconius hecuba cassandra C. Felder & R. Felder, 1862  TL: Colombia  M T

Heliconius hecuba choarina (Hewitson, 1872)  TL: Ecuador  M T

Heliconius hecuba creusa H. Holzinger & R. Holzinger, 1989  TL: Colombia  M T

Heliconius hecuba crispus Staudinger, 1885  TL: Colombia

Heliconius hecuba flava K. Brown, 1979  TL: Ecuador  M T

Heliconius hecuba lamasi Neukirchen, 1991  TL: Ecuador

Heliconius hecuba salazari Neukirchen, 1993  TL: Colombia  M T

Heliconius hecuba tolima Fassl, 1912  TL: Colombia  M T

Heliconius hecuba walteri Salazar, 1998  TL: Colombia  M T

Heliconius hermathena (Hewitson, [1854])    iN

Heliconius hermathena hermathena (Hewitson, [1854])  TL: [Brazil]  M T

Heliconius hermathena duckei K. Brown & Benson, 1977  TL: Brazil (Pa)  M T

Heliconius hermathena renatae K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius hermathena sabinae Neukirchen, 1992  TL: Brazil (Am)  M T

Heliconius hermathena sheppardi K. Brown & Benson, 1977  TL: Brazil (Am)  M T

Heliconius hermathena vereatta Stichel, 1912  TL: Brazil (Pa)  M T

Heliconius heurippa (Hewitson, [1854])    iN  M S

Heliconius hierax (Hewitson, 1869)    iN  M T

Heliconius himera (Hewitson, 1867)    iN  M T

Heliconius hortense (Guérin-Méneville, [1844])  Mexican Longwing  E & W Mexico to El Salvador & Honduras  iN  M S Ab L

Heliconius ismenius Latreille, [1817]    iN

Heliconius ismenius ismenius Latreille, [1817]  TL: [S. Amer.]  M T

Heliconius ismenius boulleti Neustetter, 1928  TL: French Guiana  M T

Heliconius ismenius clarescens A. Butler, 1875  Costa Rica & Panama  M T S L

Heliconius ismenius fasciatus Godman & Salvin, 1877  Panama  M T TI

Heliconius ismenius metaphorus Weymer, 1883  TL: Ecuador  M T

Heliconius ismenius occidentalis Neustetter, 1928  TL: Colombia  M T

Heliconius ismenius telchinia (E. Doubleday, 1847)  E Mexico to Panama  M T S L

Heliconius ismenius tilletti K. Brown & F. Fernández, 1976  TL: Venezuela  M T

Heliconius lalitae Brévignon, 1996  TL: French Guiana  M T

Heliconius leucadia H. Bates, 1862    iN

Heliconius leucadia leucadia H. Bates, 1862  TL: Brazil (Am)  M T

Heliconius leucadia andromeda Neukirchen, 1996  TL: Ecuador  M T

Heliconius leucadia birgitae Neukirchen, 1996  TL: Peru  M T

Heliconius leucadia pseudorhea Staudinger, 1897  TL: Brazil (Am)  M T S

Heliconius luciana Lichy, 1960  TL: Venezuela

Heliconius luciana luciana Lichy, 1970  TL: Venezuela  M T

Heliconius luciana watunna Lichy, 1960  TL: Venezuela

Heliconius melpomene (Linnaeus, 1758)    iN

Heliconius melpomene melpomene (Linnaeus, 1758)  TL: America  M T1 T2 T3

Heliconius melpomene aglaope C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Heliconius melpomene amandus Grose-Smith & W. F. Kirby, 1892  TL: Bolivia  M T

Heliconius melpomene amaryllis C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Heliconius melpomene anduzei K. Brown & F. Fernández, 1985  TL: Venezuela  M T

Heliconius melpomene bellula J. Turner, 1971  TL: Colombia  M T

Heliconius melpomene burchelli Poulton, 1910  TL: Brazil (Go)  M T

Heliconius melpomene cythera (Hewitson, 1869)  TL: Ecuador  M T

Heliconius melpomene ecuadorensis Emsley, 1964  TL: Ecuador  M T

Heliconius melpomene euryades Riffarth, 1900  TL: Peru  M T

Heliconius melpomene flagrans Stichel, 1919  TL: Trinidad  M T

Heliconius melpomene intersectus Neustetter, 1928  TL: Brazil ([Pa])  M T

Heliconius melpomene madeira N. Riley, 1919  TL: Brazil (Am)  M T

Heliconius melpomene malleti Lamas, 1988  TL: Ecuador  M T

Heliconius melpomene martinae Cast & Le Crom, 2012  TL: Colombia  M T

Heliconius melpomene meriana J. Turner, 1967  TL: Surinam  M T

Heliconius melpomene michellae Neukirchen, 1997  TL: Peru  M T

Heliconius melpomene nanna Stichel, 1899  TL: Brazil (ES/MG)  M T

Heliconius melpomene penelope Staudinger, 1894  TL: Bolivia  M T

Heliconius melpomene plesseni Riffarth, 1907  TL: Ecuador  M T

Heliconius melpomene pyrforus Kaye, 1907  TL: Guyana  M T

Heliconius melpomene rosina (Boisduval, 1870)  Costa Rica & Panama  M T S L

Heliconius melpomene schunkei Lamas, 1976  TL: Peru  M T L

Heliconius melpomene tessa Barcant, 1982  TL: Tobago  M T

Heliconius melpomene thelxiope (Hübner, [1806])  TL: [Brazil]

Heliconius melpomene thelxiopeia Staudinger, 1897  TL: French Guiana  M T

Heliconius melpomene vicina Ménétriés, 1857  TL: Brazil  M T

Heliconius melpomene vulcanus A. Butler, 1865  Panama & Colombia  M T L

Heliconius melpomene xenoclea (Hewitson, [1853])  TL: ?  M T

Heliconius nattereri C. Felder & R. Felder, 1865  TL: Brazil ([Ba])  M T

Heliconius numata (Cramer, 1780)    iN

Heliconius numata numata (Cramer, 1780)  TL: Surinam  M T

Heliconius numata arcuella H. Druce, 1874  TL: Peru  M T

Heliconius numata aristiona (Hewitson, [1853])  TL: Colombia  M T

Heliconius numata aulicus Weymer, 1883  TL: [Venezuela]  M T

Heliconius numata aurora H. Bates, 1862  TL: Brazil (Am)  M T

Heliconius numata bicoloratus A. Butler, 1873  TL: Peru  M T L

Heliconius numata ethra (Hübner, [1831])  TL: Brazil  M T

Heliconius numata euphone C. Felder & R. Felder, 1862  TL: Colombia  M T

Heliconius numata geminatus Weymer, 1894  TL: Brazil (Am)  M T

Heliconius numata holzingeri F. Fernández & K. Brown, 1976  TL: Venezuela

Heliconius numata ignotus Joicey & Kaye, 1917  TL: Peru  M T

Heliconius numata illustris Weymer, 1894  TL: Peru  M T

Heliconius numata jiparanaensis Neustetter, 1931  TL: Brazil (Rd)  M T S

Heliconius numata lenaeus Weymer, 1891  TL: Colombia

Heliconius numata lyrcaeus Weymer, 1891  TL: Peru?  M T

Heliconius numata mavors Weymer, 1894  TL: [Brazil (Am)]  M T

Heliconius numata messene C. Felder & R. Felder, 1862  TL: Colombia  M T

Heliconius numata mirus Weymer, 1894  TL: Bolivia  M T

Heliconius numata nubifer A. Butler, 1875  TL: Brazil (Am)  M T

Heliconius numata peeblesi Joicey & Talbot, 1925  TL: Venezuela  M T

Heliconius numata pratti Joicey & Kaye, 1917  TL: Peru  M T

Heliconius numata robigus Weymer, 1875  TL: Venezuela  M T

Heliconius numata silvana (Stoll, 1781)  TL: Surinam  M T

Heliconius numata sourensis K. Brown, 1976  TL: Brazil (Pa)  M T

Heliconius numata superioris A. Butler, 1875  TL: Brazil (Am/Pa)  M T

Heliconius numata talboti Joicey & Kaye, 1917  TL: Peru  M T

Heliconius numata tarapotensis Riffarth, 1901  TL: Peru  M T

Heliconius numata zobrysi Fruhstorfer, 1910  TL: Brazil (MT)  M T

Heliconius pardalinus H. Bates, 1862    iN

Heliconius pardalinus pardalinus H. Bates, 1862  TL: Brazil (Am)  M T

Heliconius pardalinus ariadne Neukirchen, 1995  TL: Bolivia  M T

Heliconius pardalinus butleri K. Brown, 1976  TL: Peru  M T

Heliconius pardalinus dilatus Weymer, 1894  TL: Peru  M T

Heliconius pardalinus gloria Winhard, 2019  ref.360

Heliconius pardalinus julia Neukirchen, 2000  TL: Ecuador

Heliconius pardalinus lucescens Weymer, 1894  TL: Brazil (Pa)  M T

Heliconius pardalinus maeon Weymer, 1891  TL: ?  M T L

Heliconius pardalinus orteguaza K. Brown, 1976  TL: Colombia  M T

Heliconius pardalinus radiosus A. Butler, 1873  TL: Brazil (Pa)  M T

Heliconius pardalinus sergestus Weymer, 1894  TL: Peru  M T

Heliconius pardalinus tithoreides Staudinger, 1900  TL: Peru  M T

Heliconius peruvianus (C. Felder & R. Felder, 1859)    iN  M S

Heliconius ricini (Linnaeus, 1758)    iN

Heliconius ricini ricini (Linnaeus, 1758)  TL: America  M HS

Heliconius ricini insulanus (Stichel, 1909)  TL: Trinidad  M T

Heliconius sapho (Drury, 1782)  Sapho Longwing    iN

Heliconius sapho sapho (Drury, 1782)  Panama  M OI IL L

Heliconius sapho candidus K. Brown, 1976  TL: Ecuador  M T

Heliconius sapho chocoensis K. Brown & Benson, 1975  TL: Colombia  M T

Heliconius sapho leuce (E. Doubleday, 1847)  SE Mexico to Costa Rica; Atlantic slope  M S L I

Heliconius hewitsoni (Hewitson, 1875)  Hewitson's Longwing  Costa Rica & Panama; Pacific slope  iN  M T S L

Heliconius sara (Fabricius, 1793)    iN

Heliconius sara sara (Fabricius, 1793)  TL: Guyana  M T

Heliconius sara apseudes (Hübner, [1813])  TL: Brazil  M S

Heliconius sara brevimaculata Riffarth, 1901  TL: Colombia  M T

Heliconius sara elektra Neukirchen, 1998  TL: Colombia  M T

Heliconius sara fulgidus Stichel, 1906  Guatemala to Panama; Atlantic slope  M T L

Heliconius sara magdalena H. Bates, 1864  Panama to Colombia & Venezuela  M T S L

Heliconius sara sprucei H. Bates, 1864  TL: Ecuador  M T L

Heliconius sara theudela (Hewitson, 1874)  Costa Rica & Panama; Pacific slope  M T S L I H

Heliconius sara veraepacis H. Bates, 1864  SE Mexico to Costa Rica (Atlantic Slope)  M T S L

Heliconius sara williami Neukirchen, 1994  TL: Trinidad  M T

Heliconius telesiphe (E. Doubleday, 1847)    iN

Heliconius telesiphe telesiphe (E. Doubleday, 1847)  TL: Bolivia  M T L

Heliconius telesiphe cretacea Neustetter, 1916  TL: French Guiana  M T

Heliconius telesiphe sotericus Salvin, 1871  TL: Ecuador  M T

Heliconius timareta (Hewitson, 1867)    iN

Heliconius timareta timareta (Hewitson, 1867)  TL: Ecuador  M T

Heliconius timareta timoratus Lamas, 1998  TL: Peru  M T

Heliconius tristero A. V.Z. Brower, 1996  TL: Colombia  M T

Heliconius wallacei Reakirt, 1866    iN

Heliconius wallacei wallacei Reakirt, 1866  TL: [Brazil]  M T

Heliconius wallacei araguaia K. Brown, 1976  TL: Brazil (Go)  M T

Heliconius wallacei colon Weymer, 1891  TL: [Brazil (Am)]  M T

Heliconius wallacei flavescens Weymer, 1891  TL: Ecuador  M T L

Heliconius wallacei kayei Neustetter, 1929  TL: Trinidad  M T

Heliconius wallacei mimulinus A. Butler, 1873  TL: Colombia  M T

Heliconius xanthocles H. Bates, 1862    iN

Heliconius xanthocles xanthocles H. Bates, 1862  TL: Guyana; "French Guiana"  M T

Heliconius xanthocles buechei Neukirchen, 1992  TL: Venezuela  M T

Heliconius xanthocles cleoxanthe H. Holzinger & R. Holzinger, 1972  TL: Venezuela

Heliconius xanthocles donatia Fruhstorfer, 1910  TL: Brazil (MT)  M T

Heliconius xanthocles explicata K. Brown, 1976  TL: Colombia  M T

Heliconius xanthocles hippocrene H. Holzinger & K. Brown, 1982  TL: Bolivia

Heliconius xanthocles melete C. Felder & R. Felder, 1865  TL: Colombia  M T

Heliconius xanthocles melior Staudinger, 1897  TL: Peru  M T

Heliconius xanthocles melittus Staudinger, 1897  TL: Peru  M T

Heliconius xanthocles napoensis H. Holzinger & K. Brown, 1982  TL: Ecuador  M S

Heliconius xanthocles paraplesius H. Bates, 1867  TL: Brazil (Ma)  M T

Heliconius xanthocles quindecim Lamas, 1976  TL: Peru

Heliconius xanthocles rindgei H. Holzinger & K. Brown, 1982  TL: Colombia  M T

Heliconius xanthocles similatus J. Zikán, 1937  TL: Brazil (Am)  M T

Heliconius xanthocles vala Staudinger, 1885  TL: French Guiana  M T

Heliconius xanthocles zamora H. Holzinger & K. Brown, 1982  TL: Ecuador

  Tribe Argynnini

Euptoieta  E. Doubleday, 1848  

Euptoieta bogotana Staudinger, 1885    iN  M T

Euptoieta claudia (Cramer, 1775)  Variegated Fritillary  S N Am. (strays far N) to Panama, Bahamas, Gr. Antilles, Barbuda  iN

Euptoieta claudia claudia (Cramer, 1775)  TL: Jamaica  M S1 S2 HS L1 L2 L3 L4 L5 L6 I1 I2 I3 H F

Euptoieta claudia daunius (Herbst, 1798)  TL: East Indies  M T

Euptoieta hegesia (Cramer, 1779)  Mexican Fritillary    iN

Euptoieta hegesia hegesia (Cramer, 1779)  Bahamas, Caymans, Jamaica, Cuba, Hispaniola, Mona  M T S Ab L

Euptoieta hegesia meridiania Stichel, 1938  [= hoffmanni], S TX, W MX (strays N), E Mexico to Argentina  M T S L1 L2 I1 I2

Euptoieta hegesia watsoni W. Comstock, 1944  Puerto Rico  M T S L

Euptoieta hortensia (Blanchard, 1852)    iN  M T

Euptoieta perdistincta A. Hall, 1930  TL: Peru  M T

Euptoieta poasina Schaus, 1913  TL: Costa Rica  M T

Euptoieta sunides (Hewitson, 1877)  TL: Ecuador  M T

Euptoieta thekla A. Hall, 1919    iN  M T

Euptoieta [n. sp.] Lamas, MS  Peru

Yramea  Reuss, 1920  

Yramea cytheris (Drury, 1773)    iN

Yramea cytheris cytheris (Drury, 1773)  TL: Falkland Is.  M T

Yramea cytheris siga (Geyer, 1832)  TL: Java  M T

Yramea inca (Staudinger, 1894)    iN

Yramea inca inca (Staudinger, 1894)  TL: Bolivia  M T

Yramea inca cora (Lehmann, 1913)  TL: Peru  M T

Yramea inca [n. ssp.] Lamas, MS  Argentina

Yramea lathonioides (Blanchard, 1852)    iN  M T

Yramea modesta (Blanchard, 1852)    iN  M T

Yramea sobrina (Weymer, 1890)    iN  M T

Yramea [n. sp.] Lamas & Grados, MS  Peru

Boloria  Moore, 1900  

Boloria alaskensis (W. Holland, 1900)  Mountain Fritillary    iN

Boloria alaskensis alaskensis (W. Holland, 1900)  Nunavut W to AK, S to C BC  M S L Ha

Boloria alaskensis halli Klots, 1940  Wind River Mountains, NW WY  M S1 S2 L Ha

Boloria eunomia (Esper, 1800)  Bog Fritillary    iN

Boloria eunomia triclaris (Hübner, [1821])  Labrador & vic.  M S

Boloria eunomia dawsoni (W. Barnes & McDunnough, 1916)  N WI, MI, MN, S boreal Canada to Yukon in taiga habitats  M T S1 S2

Boloria eunomia nichollae (W. Barnes & Benjamin, 1926)  Rocky Mountains of C BC & ALTA  M S

Boloria eunomia denali (Klots, 1940)  AK, N montane YK  M S L

Boloria eunomia ursadentis Ferris & Groothuis, 1971  Beartooth Plateau of N WY and S MT  M S L Ha

Boloria eunomia laddi (Klots, 1940)  Snowy Range of Albany and Carbon Cos., WY, similar in Big Horn Mts of N-central WY  M S1 S2

Boloria eunomia caelestis (Hemming, 1933)  north and central Colorado Rocky Mountains  M S L Ha

Boloria myrina (Cramer, 1777)  Silver-bordered Fritillary  [former selene]; ref.16

Boloria myrina myrina (Cramer, 1777)  TL: USA (NY)  M S L Ha

Boloria myrina nebraskensis (W. Holland, 1928)  TL: USA (Ne)  M S

Boloria myrina sabulocollis Kohler, 1977  TL: USA (Ne)  M S

Boloria myrina tollandensis (W. Barnes & Benjamin, 1925)  TL: USA (Co)  M S1 S2 L I Ha

Boloria myrina Pacific Northwestern s segregate ,  E OR, E WA, S ID, NW WY  M S1 S2 L

Boloria myrina albequina (W. Holland, 1928)  TL: not specified  M S

Boloria myrina atrocostalis (Huard, 1927)  TL: not specified  M S Ab L

Boloria myrina terraenovae (W. Holland, 1928)  TL: not specified  M S

Boloria natazhati (A. Gibson, 1920)  Cryptic Fritillary    iN

Boloria natazhati natazhati (A. Gibson, 1920)  Wrangell & St. Elias Mts., Montana Mt., YK  M T

Boloria natazhati nabokovi D. Stallings & Turner, 1947  N BC, probably S YK, White Mt., MacKenzie Mts., Bernard H., NWT  M S Ha

Boloria natazhati bankslandia J. Shepard, 1998  Victoria Island, NWT (and adjacent mainland)  M S

Boloria freija (Thunberg, 1791)  Freija Fritillary    iN

Boloria freija freija (Thunberg, 1791)  TL: Sweden  M T S1 S2 L Ha

Boloria freija tarquinius (J. Curtis, 1835)  E Canadian arctic tundra  M S

Boloria freija browni (Higgins, 1953)  TL: USA (Co)  M S L Ha

Boloria chariclea (Schneider, 1794)  Purplish Fritillary    iN

Boloria chariclea montinus (Scudder, 1863)  TL: not specified  M S L Ha

Boloria chariclea boisduvalii (Duponchel, [1832])  SE Canadian arctic (occurrs between B. c. arctica & B. c. grandis)  M S1 S2

Boloria chariclea arctica (Zetterstedt, [1839])  NE Canadian arctic to N YK  M S

Boloria chariclea grandis (W. Barnes & McDunnough, 1916)  taiga from Labrador to YK & BC, [to MT?]  M S1 S2 L Ha

Boloria chariclea butleri (W. H. Edwards, 1883)  montane AK & YK to N BC  M S1 S2 L Ha

Boloria chariclea rainieri (W. Barnes & McDunnough, 1913)  TL: not specified  M T S L

Boloria chariclea ingens (W. Barnes & McDunnough, 1918)  TL: USA (Wy)  M S L Ha

Boloria chariclea helena (W. H. Edwards, 1871)  TL: USA (Co)  M T S1 S2 Ab L1 L2 L3 Ha

Boloria kriemhild (Strecker, 1879)  Relict Fritillary    iN  M S L1 L2 Ha

Boloria epithore (W. H. Edwards, 1864)  Pacific Fritillary    iN

Boloria epithore sigridae (J. Shepard, 2001)  TL: not specified  M T

Boloria epithore chermocki E. Perkins & S. Perkins, 1966  TL: USA (Or)  M T S1 S2 L1 L2 Ha

Boloria epithore NE Oregon segregate ,  NE Oregon and surrounding areas  M S Ab L

Boloria epithore epithore (W. H. Edwards, 1864)  TL: USA (Ca)  M S L H

Boloria epithore sierra E. Perkins, 1973  TL: USA (Ca)  M T S L

Boloria improba (A. Butler, 1877)  Dingy Fritillary    iN

Boloria improba improba (A. Butler, 1877)  Scandanavia, Nunavut, NWT  M S

Boloria improba youngi (W. Holland, 1900)  AK, YK  M S L Ha

Boloria improba nunatak Scott, 1998  Rocky Mountains of Alberta & British Columbia  M T S

Boloria improba harryi (Ferris, 1984)  NW Wyoming  M T S1 S2 S3

Boloria improba acrocnema Gall & Sperling, 1980  SW Colorado Rocky Mountains  M T S L

Boloria bellona (Fabricius, 1775)  Meadow Fritillary    iN

Boloria bellona bellona (Fabricius, 1775)  TL: not specified  M S L1 L2

Boloria bellona toddi (W. Holland, 1928)  TL: not specified  M S1 S2 L

Boloria bellona jenistai D. Stallings & Turner, 1947  boreal habitats from YK to Labrador  M T S

Boloria bellona S Rocky Mountain segregate ,  Wyoming, NW Colorado  M S

Boloria frigga (Thunberg, 1791)  Frigga Fritillary    iN

Boloria frigga saga (Staudinger, 1861)  TL: Canada  M S1 S2 L Ha

Boloria frigga gibsoni (W. Barnes & Benjamin, 1926)  arctic; AK to Baffin Island  M S L I Ha

Boloria frigga sagata (W. Barnes & Benjamin, 1923)  TL: USA (Co)  M S1 S2 L Ha

Boloria alberta (W. H. Edwards, 1890)  Alberta Fritillary    iN  M S Ha

Boloria polaris (Boisduval, [1828])  Polaris Fritillary    iN

Boloria polaris polaris (Boisduval, [1828])  TL: Norway  M S L

Boloria polaris stellata Masters, 1972  vic. Churchill, Manitoba  M S

Boloria astarte (E. Doubleday, [1847])  Astarte Fritillary    iN

Boloria astarte tschukotkensis (Wyatt, 1961)  Seward Peninsula, AK

Boloria astarte alaskia Grishin, 2024  Seward Peninsula, AK; ref.154  M T S

Boloria astarte distincta (A. Gibson, 1920)  most of AK, YK, NW BC, W NWT  M S Ha

Boloria astarte astarte (E. Doubleday, 1847)  far N WA, MT, Rocky Mountains of BC-Alberta  M S

Argynnis  Fabricius, 1807  

Argynnis diana (Cramer, 1777)  Diana Fritillary    iN

Argynnis diana arkansana Pavulaan, 2021  TL: USA (Ar)

Argynnis diana diana (Cramer, 1777)  TL: USA (Va)  M S Ab L1 L2 I F Ha

Argynnis cybele (Fabricius, 1775)  Great Spangled Fritillary  ref.16  iN

Argynnis cybele cybele (Fabricius, 1775)  TL: USA (NY)  M S L1 L2 L3 L4 LAb Ha

Argynnis cybele novascotiae McDunnough, 1935  TL: not specified  M S

Argynnis cybele krautwurmi W. Holland, 1931  TL: USA (Mi)  M S L

Argynnis cybele pseudocarpenteri F. Chermock & R. Chermock, 1940  TL: not specified  M T S

Argynnis cybele carpenterii W. H. Edwards, 1876  TL: USA (NM)  M T

Argynnis cybele neomexicana Grishin, 2022  [former carpenterii] SW CO & NC NM; ref.16  M S

Argynnis cybele charlottii W. Barnes, 1897  TL: USA (Co)  M S L I Ha

Argynnis leto Behr, 1862  Great Spangled Fritillary  ref.16

Argynnis leto letona (dos Passos & Grey, 1945)  TL: USA (Ut)  M S I Ha

Argynnis leto leto Behr, 1862  TL: USA (Ca)  M T S L1 L2 Ha

Argynnis leto pugetensis (F. Chermock & Frechin, 1947)  TL: USA (Wa)  M T1 T2 S L1 L2 Ha

Argynnis leto eileenae (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S I

Argynnis aphrodite (Fabricius, 1787)  Aphrodite Fritillary    iN

Argynnis aphrodite cullasaja (Gatrelle, 1998)  TL: USA (NC)  M T

Argynnis aphrodite aphrodite (Fabricius, 1787)  TL: USA (NY)  M S L

Argynnis aphrodite alcestis W. H. Edwards, [1877]  TL: USA (Il)  M T S Ab L

Argynnis aphrodite ethne Hemming, 1933  TL: not specified  M T S1 S2 Ab L1 L2 L3 L4 Ha

Argynnis aphrodite W Colorado segregate (, )  W Colorado  M S

Argynnis aphrodite byblis (W. Barnes & Benjamin, 1926)  TL: USA (Az)  M S L

Argynnis aphrodite columbia H. Edwards, 1877  TL: USA (Ak)  M T S

Argynnis aphrodite whitehousei Gunder, 1932  TL: not specified  M T S

Argynnis aphrodite manitoba F. Chermock & R. Chermock, 1940  TL: not specified  M T1 T2

Argynnis aphrodite winni Gunder, 1932  TL: not specified  M S

Argynnis idalia (Drury, 1773)  Regal Fritillary    iN

Argynnis idalia idalia (Drury, 1773)  TL: USA (NY)  M T S L I

Argynnis idalia occidentalis (B. Williams, 2002)  TL: USA (Il)  M S L1 L2 L3 L4 Ha

Argynnis nokomis W. H. Edwards, 1862  Nokomis Fritillary    iN

Argynnis nokomis coerulescens W. Holland, 1900  NW Mexico, extirpated from SE AZ  M T S L I

Argynnis nokomis nitocris W. H. Edwards, 1874  TL: USA (Az)  M T S L1 L2 H

Argynnis nokomis nokomis W. H. Edwards, 1862  TL: USA (Co)  M T S

Argynnis nokomis tularosa R. Holland, 2010  TL: USA (NM)  M T OD

Argynnis nokomis apacheana Skinner, 1918  TL: USA (Ca)  M T S1 S2 L1 L2 I1 I2 Ha

Argynnis nokomis carsonensis (Austin, 1998)  TL: USA (Nv)  M S L Ha

Argynnis nokomis wenona (dos Passos & Grey, 1945)  NE Mexico  M T

Argynnis nokomis melaena (Mooser & García, 1980)  C Mexico (Aguascalientes)  M S

Argynnis edwardsii Reakirt, 1866  Edwards' Fritillary    iN  M T S1 S2 Ab L1 L2 Ha

Argynnis coronis Behr, 1864  Coronis Fritillary    iN

Argynnis coronis hennei Gunder, 1934  TL: USA (Ca)  M T S

Argynnis coronis coronis Behr, 1864  TL: not specified  M T S L1 L2

Argynnis coronis simaetha (dos Passos & Grey, 1945)  TL: USA (Wa)  M T S L Ha

Argynnis coronis snyderi Skinner, 1897  TL: USA (Ut)  M T S1 S2 L

Argynnis coronis halcyone W. H. Edwards, [1869]  TL: USA (Co)  M T S1 S2 S3 L1 L2 Ha

Argynnis coronis carolae dos Passos & Grey, 1942  USA (Nv); ref.11, iN.1267545  M T S L Ha

Argynnis coronis semiramis W. H. Edwards, 1886  TL: USA (Ca)  M T S L

Argynnis zerene Boisduval, 1852  Zerene Fritillary    iN

Argynnis zerene sitka (Hammond, Harry & McCorkle, 2001)  TL: USA (Ak)  M T

Argynnis zerene bremnerii W. H. Edwards, 1872  TL: USA (Wa)  M T S

Argynnis zerene Willamette Valley, W OR segregate (, )  apparently extinct  M S

Argynnis zerene hippolyta W. H. Edwards, 1879  TL: USA (Or)  M T S L1 L2 L3 I Ha

Argynnis zerene gloriosa (Moeck, 1957)  TL: USA (Or)  M S

Argynnis zerene S OR coastal segregate (, )  M S

Argynnis zerene behrensii W. H. Edwards, 1869  TL: USA (Ca)  M T S I

Argynnis zerene sonomensis (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Argynnis zerene puntareyes (J. Emmel & T. Emmel, 1998)  TL: USA (Ca)  M T S1 S2 L

Argynnis zerene myrtleae (dos Passos & Grey, 1945)  TL: USA (Ca)  M T

Argynnis zerene monticola Behr, 1863  TL: USA (Ca)  M T S

Argynnis zerene malcolmi J. A. Comstock, 1920  TL: USA (Ca)  M T S L

Argynnis zerene zerene Boisduval, 1852  TL: USA (Ca)  M T S L Ha

Argynnis zerene E-central OR Cascades segregate (, )  M L Ha

Argynnis zerene gunderi J. A. Comstock, 1925  TL: USA (Ca)  M T S1 S2 L1 L2 Ha

Argynnis zerene picta McDunnough, 1924  TL: not specified  M T S L

Argynnis zerene platina Skinner, 1897  TL: USA (Ut)  M S L

Argynnis zerene sinope (dos Passos & Grey, 1945)  TL: USA (Co)  M S1 S2 S3 L1 L2 Ha

Argynnis zerene atossa W. H. Edwards, 1890  USA (Ca); ref.11  M T S1 S2 S3

Argynnis callippe Boisduval, 1852  Callippe Fritillary    iN

Argynnis callippe adiasteoides (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T1 T2

Argynnis callippe liliana H. Edwards, 1877  TL: USA (Ca)  M S L

Argynnis callippe callippe Boisduval, 1852  TL: USA (Ca)  M T S

Argynnis callippe comstocki Gunder, 1925  TL: USA (Ca)  M T S L1 L2 H

Argynnis callippe macaria W. H. Edwards, 1877  TL: USA (Ca)  M T S L

Argynnis callippe laurina W. G. Wright, 1905  TL: USA (Ca)  M T S Ab L

Argynnis callippe rupestris Behr, 1863  TL: USA (Ca)  M T S

Argynnis callippe inornata W. H. Edwards, 1872  TL: USA (Ca)  M S

Argynnis callippe juba Boisduval, 1869  TL: USA (Ca)  M T S

Argynnis callippe hagemani (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T1 T2 S

Argynnis callippe shasta (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S L

Argynnis callippe elaine (dos Passos & Grey, 1945)  TL: USA (Or)  M T S L1 L2 Ha

Argynnis callippe N Siskiyous & W Cascades, low elevations segregate (, )  N Siskiyous, W Cascades, Oregon, below 3500'  M L

Argynnis callippe W OR Cascades, high elevations segregate (, )  W OR Cascades, above 3500'  M S

Argynnis callippe semivirida McDunnough, 1924  TL: not specified  M S1 S2 L1 L2 Ha

Argynnis callippe chilcotinensis (C. Guppy & J. Shepard, 2001)  TL: not specified  M S

Argynnis callippe nevadensis W. H. Edwards, 1870  TL: USA (Nv)  M S L

Argynnis callippe harmonia (dos Passos & Grey, 1945)  TL: USA (Nv)  M S L

Argynnis callippe calgariana McDunnough, 1924  TL: not specified  M S

Argynnis callippe gallatini McDunnough, 1929  TL: not specified  M S1 S2 Ab L1 L2 Ha

Argynnis callippe meadii W. H. Edwards, 1872  TL: USA (Co)  M T S1 S2 L1 L2 Ha

Argynnis egleis Behr, 1862  Great Basin Fritillary    iN

Argynnis egleis tehachapina J. A. Comstock, 1920  TL: USA (Ca)  M T S L

Argynnis egleis egleis Behr, 1862  TL: USA (Ca)  M T S L1 L2 L3

Argynnis egleis Warner Mts. segregate (, )  Warner Mts., Lake Co. OR / Modoc Co., CA  M S L

Argynnis egleis oweni W. H. Edwards, 1892  TL: USA (Ca)  M T S L

Argynnis egleis mattooni (J. Emmel & T. Emmel, 1998)  TL: USA (Ca)  M T S L Ha

Argynnis egleis yolaboli (J. Emmel & T. Emmel, 1998)  TL: USA (Ca)  M T

Argynnis egleis moecki (Hammond & Dornfeld, 1983)  TL: USA (Or)  M S L Ha

Argynnis egleis toiyabe (Howe, 1975)  TL: USA (Nv)  M T S

Argynnis egleis reidi (Austin, 1998)  TL: USA (Nv)  M T1 T2 L Ha

Argynnis egleis linda dos Passos & Grey, 1942  TL: USA (Id)  M S

Argynnis egleis Stansbury Mts. segregate (, )  Stansbury Mts., Tooele Co., Utah  M S

Argynnis egleis utahensis Skinner, 1919  TL: USA (Ut)  M S

Argynnis egleis secreta (dos Passos & Grey, 1945)  TL: USA (Co)  M S1 S2 L Ha

Argynnis egleis macdunnoughi Gunder, 1932  TL: not specified  M S

Argynnis egleis morrellensis Kohler, 2020  TL: USA (Mt)

Argynnis egleis kutoyisiks Kohler, 2020  TL: USA (Mt)

Argynnis egleis albrighti Gunder, 1932  TL: not specified  M S

Argynnis adiaste W. H. Edwards, 1864  Unsilvered Fritillary    iN

Argynnis adiaste adiaste W. H. Edwards, 1864  TL: USA (Ca)  M T S L

Argynnis adiaste clemencei J. A. Comstock, 1925  TL: USA (Ca)  M T S L

Argynnis atlantis W. H. Edwards, 1862  Atlantis Fritillary    iN

Argynnis atlantis atlantis W. H. Edwards, 1862  TL: USA (NY)  M T S L

Argynnis atlantis canadensis dos Passos, 1935  TL: not specified  M T S

Argynnis atlantis brittanae Pavulaan, 2022  TL: USA (WV)

Argynnis atlantis hollandi F. Chermock & R. Chermock, 1940  TL: not specified  M T1 T2 S

Argynnis atlantis pahasapa (Spomer, Scott & Kondla, 1998)  TL: USA (SD)  M T

Argynnis atlantis sorocko (Scott, Kondla & Spomer, 1998)  TL: USA (Co)  M T S L

Argynnis irene Boisduval, 1869  Northwestern Fritillary  ref.11

Argynnis irene irene Boisduval, 1869  TL: USA (Ca)  M T S L

Argynnis irene hanseni (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Argynnis irene cottlei J. A. Comstock, 1925  TL: USA (Ca)  M S

Argynnis irene dodgei Gunder, 1931  TL: USA (Or)  M T S L1 L2 Ha

Argynnis nausicaa W. H. Edwards, 1874  Northwestern Fritillary  ref.11

Argynnis nausicaa elko (Austin, 1984)  TL: USA (Nv)  M T S L Ha

Argynnis nausicaa greyi (Moeck, 1950)  TL: USA (Nv)  M T S Ha

Argynnis nausicaa viola (dos Passos & Grey, 1945)  TL: USA (Id)  M T S

Argynnis nausicaa tetonia (dos Passos & Grey, 1945)  TL: USA (Wy)  M T S1 S2 L

Argynnis nausicaa chitone W. H. Edwards, 1879  TL: USA (Ut)  M T S L

Argynnis nausicaa schellbachi (Garth, 1949)  TL: USA (Az)  M T S L

Argynnis nausicaa electa W. H. Edwards, 1878  TL: USA (Co)  M T S L Ha

Argynnis nausicaa dorothea (Moeck, 1947)  TL: USA (NM)  M T S L

Argynnis nausicaa capitanensis (R. Holland, 1988)  TL: USA (NM)  M T S

Argynnis nausicaa nausicaa W. H. Edwards, 1874  TL: USA (Az)  M T S L1 L2 L3 L4 L5 L6

Argynnis hesperis W. H. Edwards, 1864  Northwestern Fritillary    iN

Argynnis hesperis hutchinsi (dos Passos & Grey, 1947)  TL: not specified  M S L

Argynnis hesperis brico (Kondla, Scott & Spomer, 1998)  TL: not specified  M T S

Argynnis hesperis beani (W. Barnes & Benjamin, 1926)  TL: not specified  M T S

Argynnis hesperis dennisi (dos Passos & Grey, 1945)  TL: not specified  M T S1 S2

Argynnis hesperis lurana (dos Passos & Grey, 1945)  TL: USA (SD)  M T S1 S2 L Ha

Argynnis hesperis hesperis W. H. Edwards, 1864  TL: USA (Co)  M T S1 S2 S3 S4 Ab L Ha

Argynnis hesperis ratonensis (Scott, 1981)  TL: USA (NM)  M T S

Argynnis hydaspe Boisduval, 1869  Hydaspe Fritillary    iN

Argynnis hydaspe viridicornis J. A. Comstock, 1925  TL: not specified  M T S L1 L2

Argynnis hydaspe hydaspe Boisduval, 1869  TL: USA (Ca)  M T S L

Argynnis hydaspe tehama (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S I

Argynnis hydaspe davisi (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Argynnis hydaspe rhodope W. H. Edwards, 1874  TL: Canada (BC)  M T S1 S2 L1 L2 Ha

Argynnis hydaspe NW OR & SW WA Coast Range segregate (, )  M S L Ha

Argynnis hydaspe minor (dos Passos & Grey, 1947)  TL: not specified  M T S

Argynnis mormonia Boisduval, 1869  Mormon Fritillary  ref.16  iN

Argynnis mormonia mormonia Boisduval, 1869  TL: USA (Nv)  M T S L Ha

Argynnis mormonia obsidiana (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S L Ha

Argynnis bischoffii W. H. Edwards, 1870  Mormon Fritillary  ref.16

Argynnis bischoffii bischoffii W. H. Edwards, 1870  TL: USA (Ak)  M S L

Argynnis bischoffii opis W. H. Edwards, 1874  TL: not specified  M S

Argynnis bischoffii washingtonia W. Barnes & McDunnough, 1913  TL: not specified  M S L

Argynnis bischoffii erinna W. H. Edwards, 1883  TL: USA (Wa)  M T S L Ha

Argynnis bischoffii kimimela Marrone, Spomer & Scott, 2008  Black Hills of SD & WY  M S L Ha

Argynnis bischoffii eurynome W. H. Edwards, 1872  TL: USA (Co)  M T S Ab L1 L2 L3 Ha

Argynnis bischoffii Warner Mts. segregate (, )  M S

Argynnis bischoffii artonis W. H. Edwards, 1881  TL: USA (Nv)  M T S L Ha

Argynnis bischoffii luski W. Barnes & McDunnough, 1913  TL: USA (Az)  M S L1 L2 H

  Subfamily Limenitidinae

  Tribe Limenitidini

Limenitis  Fabricius, 1807  

Limenitis arthemis (Drury, 1773)  White Admiral    iN

Limenitis arthemis rubrofasciata (W. Barnes & McDunnough, 1916)  TL: not specified  M S1 S2 S3

Limenitis arthemis arthemis (Drury, 1773)  TL: USA (NY)  M S1 S2 L1 L2

Limenitis arthemis astyanax (Fabricius, 1775)  TL: not specified  M S1 S2 S3 S4 S5 Hy HS L1 L2 L3 L4 L5 I H F

Limenitis arthemis arizonensis W. H. Edwards, 1882  TL: USA (Az)  M T S1 S2 L I H

Limenitis weidemeyerii W. H. Edwards, 1861  Weidemeyer’s Admiral    iN

Limenitis weidemeyerii weidemeyerii W. H. Edwards, 1861  TL: USA (Co)  M S1 S2 S3 L Ha

Limenitis weidemeyerii oberfoelli F. Brown, 1960  TL: USA (ND)  M S1 S2 L Ha

Limenitis weidemeyerii latifascia S. Perkins & E. Perkins, 1967  TL: USA (Id)  M S1 S2 Hy L

Limenitis weidemeyerii nevadae (W. Barnes & Benjamin, 1924)  TL: USA (Nv)  M S1 S2

Limenitis weidemeyerii angustifascia (W. Barnes & McDunnough, 1912)  TL: USA (Az)  M S1 S2 L1 L2 I Ha

Limenitis weidemeyerii siennafascia Austin & Mullins, 1984  TL: USA (Az)  M T S L

Limenitis weidemeyerii Coahuila segregate ,  N Coahuila, Mexico  M S L Ha

Limenitis lorquini Boisduval, 1852  Lorquin's Admiral    iN

Limenitis lorquini powelli Austin & J. Emmel, 1998  TL: not specified  M T S1 S2 L I

Limenitis lorquini pallidafacies Austin & J. Emmel, 1998  TL: USA (Nv)  M T S L Ha

Limenitis lorquini lorquini Boisduval, 1852  TL: USA (Ca)  M T S1 S2 S3 Hy L1 L2 L3 Ha

Limenitis lorquini ilgae C. Guppy, 2001  TL: not specified  M S L Ha

Limenitis lorquini burrisonii Maynard, 1891  TL: not specified  M S Hy L Ha

Limenitis archippus (Cramer, 1775)  Viceroy    iN

Limenitis archippus floridensis Strecker, 1878  vagrant in Cuba  M T S1 S2 L H

Limenitis archippus watsoni (dos Passos, 1938)  TL: USA (La)  M S1 S2 L

Limenitis archippus archippus (Cramer, 1775)  TL: USA (NY)  M S1 S2 S3 S4 S5 Hy L1 L2 I

Limenitis archippus idaho Austin, 1998  TL: USA (Nv)  M T S1 S2 Ha

Limenitis archippus lahontani Herlan, 1971  TL: USA (Nv)  M T S1 S2

Limenitis archippus obsoleta W. H. Edwards, 1882  NW Mexico  M T S1 S2 L I

Limenitis archippus hoffmanni R. Chermock, 1947  E Mexico  M T S1 S2 L H

Limenitis archippus Baja California Sur segregate ,  Baja California Sur - only 1 specimen known  M S

Adelpha  Hübner, [1819]  

Adelpha alala (Hewitson, 1847)    iN

Adelpha alala alala (Hewitson, 1847)  TL: Venezuela  M T

Adelpha alala completa Fruhstorfer, 1907  TL: Colombia  M T

Adelpha alala negra (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S L

Adelpha aricia (Hewitson, 1847)    iN

Adelpha aricia aricia (Hewitson, 1847)  TL: Bolivia  M T

Adelpha aricia portunus A. Hall, 1938  TL: Peru  M T

Adelpha aricia serenita Fruhstorfer, 1915  TL: Peru  M T

Adelpha aricia [n. ssp.] Lamas, MS  Peru

Adelpha corcyra (Hewitson, 1847)    iN

Adelpha corcyra corcyra (Hewitson, 1847)  TL: Colombia  M T

Adelpha corcyra aretina Fruhstorfer, 1907  TL: Ecuador  M T

Adelpha corcyra collina (Hewitson, 1847)  TL: Ecuador  M T

Adelpha corcyra dognini Willmott, 2003  TL: Ecuador  M T S

Adelpha corcyra salazari Willmott, 2003  TL: Colombia  M T

Adelpha pithys (H. Bates, 1864)  Pithy Sister  SW & S Mexico to El Salvador, [Honduras?]  iN  M T S L

Adelpha donysa (Hewitson, 1847)  Donysa Sister    iN

Adelpha donysa SW Mexican segregate ,  SW Mexico (MICH to OAX)  M S

Adelpha donysa donysa (Hewitson, 1847)  SE Mexico to Guatemala  M T S

Adelpha donysa albifilum Steinhauser, 1974  El Salvador, Honduras  M T

Adelpha tracta (A. Butler, 1872)  Tracta Sister  Nicaragua to W Panama  iN  M T S1 S2 L I

Adelpha bredowii Geyer, 1837  Mexican Sister  SW & S-central Mexico, local in NW Mexico; cloud forest  iN  M T S L

Adelpha californica (A. Butler, 1865)  California Sister  Baja California (N) to S WA, Cascade-Sierra Nevada, westward  iN  M T S L1 L2 I Ha

Adelpha eulalia (E. Doubleday, [1848])  Arizona Sister  far SE CA, S NV, UT, AZ, NM, S CO, TX (strays N), Mx to Guat.  iN  M T S1 S2 L1 L2 L3 I1 I2 I3 I4 H

Adelpha diocles Godman & Salvin, 1878  Short-tailed Sister    iN

Adelpha diocles diocles Godman & Salvin, 1878  E Costa Rica & W Panama (Chiriqui massif)  M T S

Adelpha diocles creton Godman, 1901  E-central & S Mexico to Guatemala & El Salvador  M T S L

Adelpha herbita Weymer, 1907  TL: Brazil (SC)

Adelpha herbita herbita Weymer, 1907  TL: Brazil (SC)

Adelpha herbita perdita Willmott & Freitas, 2016  ref.313

Adelpha zea (Hewitson, 1850)    iN  M T

Adelpha nea (Hewitson, 1847)  Nea Sister    iN

Adelpha nea nea (Hewitson, 1847)  Costa Rica to Venezuela, to S Peru, Guianas, Amazonian Brazil  M T S

Adelpha nea sentia Godman & Salvin, 1884  SE Mexico (OAX, VER), Belize  M T S

Adelpha paroeca (H. Bates, 1864)  Paroeca Sister    iN

Adelpha paroeca paroeca (H. Bates, 1864)  [= emathia], E & W Mexico to W Panama  M T S L

Adelpha paroeca pseudodonysa Salazar, 2000  TL: Colombia  M T

Adelpha paraena (H. Bates, 1865)    iN

Adelpha paraena paraena (H. Bates, 1865)  TL: Brazil (Pa)  M T

Adelpha paraena lecromi Willmott, 2003  C Panama to N Colombia  M S

Adelpha paraena massilia (C. Felder & R. Felder, 1867)  E & W Mexico to C Panama  M T S L I Ha

Adelpha paraena reyi Neild, 1996  TL: Venezuela  M T

Adelpha radiata Fruhstorfer, 1915    iN

Adelpha radiata radiata Fruhstorfer, 1915  TL: Brazil (SC)  M T

Adelpha radiata aiellae Willmott & J. Hall, 1999  NW Costa Rica to NW Ecuador  M T S I

Adelpha radiata explicator Willmott & J. Hall, 1999  TL: Ecuador

Adelpha radiata gilletella Brévignon, 1995  TL: French Guiana

Adelpha radiata myrlea Fruhstorfer, 1915  TL: Brazil (ES)  M T

Adelpha radiata romeroi Willmott & Neild, 2003  TL: Venezuela

Adelpha serpa (Boisduval, 1836)    iN

Adelpha serpa serpa (Boisduval, 1836)  TL: Brazil  M T

Adelpha serpa celerio (H. Bates, 1864)  E & W Mexico to W Colombia, NW Venezuela,  M T S1 S2 L I1 I2 I3

Adelpha serpa diadochus Fruhstorfer, 1915  TL: Peru  M T S

Adelpha serpa duiliae Fruhstorfer, 1913  TL: Ecuador  M T

Adelpha seriphia (C. Felder & R. Felder, 1867)    iN

Adelpha seriphia seriphia (C. Felder & R. Felder, 1867)  TL: Venezuela  M T

Adelpha seriphia aquillia Fruhstorfer, 1915  TL: Colombia  M T

Adelpha seriphia naryce Fruhstorfer, 1915  TL: Peru  M T

Adelpha seriphia barcanti Willmott, 2003  TL: Trinidad  M T

Adelpha seriphia egregia Röber, 1927  TL: Colombia  M T

Adelpha seriphia pione Godman & Salvin, 1884  TL: Panama  M T

Adelpha seriphia therasia Fruhstorfer, 1915  TL: Bolivia  M T

Adelpha hyas (Doyère, [1840])    iN

Adelpha hyas hyas (Doyère, [1840])  TL: ?  M S

Adelpha hyas hewitsoni Willmott & J. Hall, 1999  TL: Ecuador

Adelpha hyas viracocha A. Hall, 1938  TL: Peru  M T

Adelpha godmani Fruhstorfer, 1913  Godman's Sister  Chiapas to W Ecuador  iN  M T S L

Adelpha margarita Willmott & J. Hall, 2013    iN

Adelpha margarita margarita Willmott & J. Hall, 2013  ref.321

Adelpha margarita garleppi Willmott, 2013  ref.321

Adelpha abia (Hewitson, 1850)    iN  M T

Adelpha abyla (Hewitson, 1850)  Jamaican Sister  Jamaica  iN  M T S

Adelpha adstricta Fruhstorfer, 1915    iN

Adelpha amazona Austin & Jasinski, 1999    iN

Adelpha argentea Willmott & J. Hall, 1995    iN  M T

Adelpha atlantica Willmott, 2003  TL: Brazil (RJ)  M T

Adelpha attica (C. Felder & R. Felder, 1867)    iN

Adelpha attica attica (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Adelpha attica hemileuca Willmott & J. Hall, 1999  NW Ecuador, W Colombia, 1 Panama specimen nr. nominate form  M T

Adelpha barnesia Schaus, 1902    iN

Adelpha barnesia barnesia Schaus, 1902  TL: Bolivia  M T

Adelpha barnesia leucas Fruhstorfer, 1915  E & W Mexico to NW Venezuela & W Ecuador  M T S I

Adelpha barnesia trinita Kaye, 1914  TL: Trinidad  M T

Adelpha basiloides (H. Bates, 1865)  Spot-celled Sister  S TX, E & W Mexico to N Venezuela & W Ecuador  iN  M T S1 S2 L I

Adelpha boeotia (C. Felder & R. Felder, 1867)  Oberthur's Sister    iN

Adelpha boeotia boeotia (C. Felder & R. Felder, 1867)  E Panama to Bolivia, SW Brazil, Guianas  M T S L

Adelpha boeotia fidicula Fruhstorfer, 1915  TL: Brazil (ES)  M T

Adelpha boeotia oberthurii (Boisduval, 1870)  [= jacquelinae], SE Mexico to W Panama  M T L

Adelpha boreas (A. Butler, 1866)  Boreas Sister    iN

Adelpha boreas boreas (A. Butler, 1866)  E Costa Rica to Venezuela & Bolivia  M T S L

Adelpha boreas kayei A. Hall, 1939  TL: Guyana  M T

Adelpha calliphane Fruhstorfer, 1915    iN  M T

Adelpha capucinus (Walch, 1775)    iN

Adelpha capucinus capucinus (Walch, 1775)  TL: East Indies  M T S

Adelpha capucinus gutierrezi Neild, 1996  TL: Venezuela  M T

Adelpha capucinus velia (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Adelpha cocala (Cramer, 1779)    iN

Adelpha cocala cocala (Cramer, 1779)  TL: Surinam  M T S

Adelpha cocala caninia Fruhstorfer, 1915  TL: Brazil (SC/SP)  M T

Adelpha cocala didia Fruhstorfer, 1915  TL: Brazil (ES)  M T

Adelpha cocala lorzae (Boisduval, 1870)  SE Mexico (OAX, CHIS) to NW Venezuela & W Ecuador  M T S1 S2 I

Adelpha cocala orellanae Neild, 1996  TL: Venezuela  M T

Adelpha coryneta (Hewitson, 1874)    iN  M T

Adelpha cytherea (Linnaeus, 1758)    iN

Adelpha cytherea cytherea (Linnaeus, 1758)  TL: Indiis  M T S

Adelpha cytherea aea (C. Felder & R. Felder, 1867)  TL: Brazil?  M T

Adelpha cytherea daguana Fruhstorfer, 1913  E Costa Rica (as intergrades with marcia) to NW Venez. & W Ecuad.  M T S L

Adelpha cytherea insularis Fruhstorfer, 1913  TL: Trinidad  M T

Adelpha cytherea marcia Fruhstorfer, 1913  S Mexico through E Costa Rica  M T S1 S2 L I

Adelpha cytherea nahua Grose-Smith, 1898  TL: Venezuela  M T

Adelpha cytherea olbia (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Adelpha delinita Fruhstorfer, 1913    iN

Adelpha delinita delinita Fruhstorfer, 1913  TL: Colombia  M T S L

Adelpha delinita utina A. Hall, 1938  [= escalantei], SE Mexico to W Panama  M T S

Adelpha demialba (A. Butler, 1872)  White-spotted Sister  Costa Rica to E Panama; cloud forest  iN  M T S L I1 I2

Adelpha diazi Beutelspacher, 1975  Diaz's Sister  E Mexico  iN  M T S

Adelpha epione (Godart, [1824])    iN

Adelpha epione epione (Godart, [1824])  TL: Brazil

Adelpha epione agilla Fruhstorfer, 1907  TL: Colombia  M T S

Adelpha epizygis Fruhstorfer, 1915    iN

Adelpha epizygis epizygis Fruhstorfer, 1915  TL: Brazil (SP)  M T

Adelpha epizygis epona Fruhstorfer, 1915  TL: [Brazil (Ba)]  M T

Adelpha erotia (Hewitson, 1847)  Erotia Sister    iN

Adelpha erotia erotia (Hewitson, 1847)  SE Mexico (Chiapas) to Bolivia, Amazonian Brazil & Guianas  M T S L

Adelpha erotia caphira (Hewitson, 1869)  TL: Venezuela  M T

Adelpha erymanthis Godman & Salvin, 1884  Erymanthis Sister    iN

Adelpha erymanthis erymanthis Godman & Salvin, 1884  S Mexico (Chiapas) to C Panama  M T S I

Adelpha erymanthis esperanza Balcázar & Willmott, 2003  Sierra de Juárez, Oaxaca, Mexico  M S HS

Adelpha erymanthis fortunata Willmott, 2003  TL: Ecuador

Adelpha ethelda (Hewitson, 1867)    iN

Adelpha ethelda Northern segregate ,  SE Mexico  M S

Adelpha ethelda ethelda (Hewitson, 1867)  TL: Ecuador  M T

Adelpha ethelda eponina Staudinger, 1886  TL: Colombia  M T

Adelpha ethelda galbao Brévignon, 1995  TL: French Guiana

Adelpha ethelda sophax Godman & Salvin, 1878  E Costa Rica to C Panama; cloud forest  M T L

Adelpha ethelda zalmona (Hewitson, 1871)  TL: Colombia  M T

Adelpha ethelda [n. ssp.] Willmott, MS  Mexico

Adelpha fabricia Fruhstorfer, 1913    iN  M T

Adelpha falcipennis Fruhstorfer, 1915    iN  M T S

Adelpha felderi (Boisduval, 1870)  Felder's Sister  E Mexico to Honduras  iN  M T S L

Adelpha fessonia (Hewitson, 1847)  Band-celled Sister    iN

Adelpha fessonia fessonia (Hewitson, 1847)  S TX, E & W Mexico to W Panama  M T S1 S2 L1 L2 I Ha

Adelpha fessonia cestus (Hewitson, 1847)  TL: Venezuela  M T

Adelpha fessonia ernestoi Willmott, 2003  TL: Colombia  M T

Adelpha lapitha A. Hall, 1929  Hispaniolan Sister  Hispaniola  M S

Adelpha gavina Fruhstorfer, 1915    iN  M T

Adelpha gelania (Godart, [1824])  Antillean Sister    iN

Adelpha gelania gelania (Godart, [1824])  Hispaniola  M S

Adelpha gelania arecosa (Hewitson, 1847)  Puerto Rico  M T S L

Adelpha heraclea (C. Felder & R. Felder, 1867)  Heraclea Sister    iN

Adelpha heraclea heraclea (C. Felder & R. Felder, 1867)  Guatemala to SE Peru, Brazil  M T L

Adelpha heraclea antonii Neild, 1996  TL: Venezuela

Adelpha heraclea makkeda (Hewitson, 1871)  TL: Brazil (Pa)  M T

Adelpha hesterbergi Willmott & J. Hall, 1999  Hesterberg's Sister  Costa Rica, NW Ecuador, [Panama?, W Colombia?]  iN  M T

Adelpha iphicleola (H. Bates, 1864)  Iphicleola Sister    iN

Adelpha iphicleola iphicleola (H. Bates, 1864)  [= massilides], E & W Mexico to E Panama  M T S1 S2 L I

Adelpha iphicleola gortyna Fruhstorfer, 1915  Costa Rica to W Panama, possibly only as intergrades with above  M T S L

Adelpha iphicleola iphimedia Fruhstorfer, 1915  Cuba & Isla de Juventud  M T S

Adelpha iphicleola leucates Fruhstorfer, 1915  TL: Brazil (Ba)  M T

Adelpha iphicleola thessalita Willmott & J. Hall, 1999  TL: Ecuador

Adelpha iphiclus (Linnaeus, 1758)  Iphiclus Sister    iN

Adelpha iphiclus iphiclus (Linnaeus, 1758)  W & S Mexico to Bolivia, Guianas, Trinidad  M T S1 S2 HS L I

Adelpha iphiclus ephesa (Ménétriés, 1857)  TL: Brazil  M T

Adelpha iphiclus estrecha Willmott & J. Hall, 1999  TL: Ecuador  M T S

Adelpha irmina (E. Doubleday, [1848])    iN

Adelpha irmina irmina (E. Doubleday, [1848])  TL: Venezuela  M T

Adelpha irmina tumida (A. Butler, 1873)  TL: Peru; Colombia  M T S L

Adelpha irmina arcana Benmesbah & Costa, 2019  ref.192

Adelpha irmina [n. ssp.] Lamas, MS  Peru

Adelpha jordani Fruhstorfer, 1913    iN  M T S

Adelpha justina (C. Felder & R. Felder, 1861)    iN

Adelpha justina justina (C. Felder & R. Felder, 1861)  TL: Colombia  M T S

Adelpha justina inesae Orellana, 1996  TL: Venezuela  M T

Adelpha justina justinella Fruhstorfer, 1907  TL: Venezuela  M T

Adelpha justina maira Orellana, 1996  TL: Venezuela  M T

Adelpha justina pichincha Willmott & J. Hall, 2013  TL: Ecuador

Adelpha justina valentina Fruhstorfer, 1915  TL: Colombia; Bolivia  M T S

Adelpha lamasi Willmott & J. Hall, 1999    iN  M T

Adelpha leuceria (H. Druce, 1874)    iN  M T S L

Adelpha juanna Grose-Smith, 1898  TL: Colombia  M T

Adelpha leucerioides Beutelspacher, 1975  Tawny-striped Sister  E Mexico to Guatemala  iN

Adelpha leucerioides SW Mexican segregate ,  SW Mexico  M S

Adelpha leucerioides leucerioides Beutelspacher, 1975  TL: Mexico (Ver)  M T S

Adelpha leucophthalma (Latreille, [1809])  Leucophthalma Sister    iN

Adelpha leucophthalma leucophthalma (Latreille, [1809])  Nicaragua to Colombia  M T S1 S2 L I

Adelpha leucophthalma irminella Strand, 1918  TL: Ecuador  M T

Adelpha leucophthalma smalli Willmott, 2003  Cerro Montuoso, Azuero Peninsula, Herrera, Panama  M S

Adelpha levona Steinhauser & L. Miller, 1977    iN  M T

Adelpha melanthe (H. Bates, 1864)  Rayed Sister  E & S Mexico to C Panama  iN  M T S1 S2 L I1 I2

Adelpha lycorias (Godart, [1824])    iN

Adelpha lycorias melanippe Godman & Salvin, 1884  TL: Colombia  M T

Adelpha lycorias spruceana (H. Bates, 1864)  TL: Ecuador  M T

Adelpha lycorias wallisii (Dewitz, 1877)  TL: Colombia  M T

Adelpha lycorias lara (Hewitson, 1850)  TL: Venezuela  M T S L

Adelpha lycorias lycorias (Godart, [1824])  TL: Brazil  M T

Adelpha malea (C. Felder & R. Felder, 1861)    iN

Adelpha malea malea (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Adelpha malea aethalia (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Adelpha malea fundania Fruhstorfer, 1915  E Mexico to NW Venezuela  M T S I

Adelpha malea goyama Schaus, 1902  TL: Brazil (Pr)  M T

Adelpha malea ixia (C. Felder & R. Felder, 1867)  TL: Venezuela  M T

Adelpha melona (Hewitson, 1847)    iN

Adelpha melona melona (Hewitson, 1847)  TL: Colombia  M T

Adelpha melona deborah Weeks, 1901  TL: Colombia  M T

Adelpha melona leucocoma Fruhstorfer, 1915  TL: Upper Amazon  M T S

Adelpha melona neildi Willmott, 2003  C Panama  M L

Adelpha melona pseudarete Fruhstorfer, 1915  TL: ?  M T

Adelpha mesentina (Cramer, 1777)    iN  M T

Adelpha messana (C. Felder & R. Felder, 1867)  Messana Sister    iN

Adelpha messana messana (C. Felder & R. Felder, 1867)  Guatemala to Colombia, NW Ecuador, NW Venezuela  M T S L

Adelpha messana delphicola Fruhstorfer, 1910  TL: [Peru]  M T

Adelpha messana seminivea Freitas, Willmott & Woodbury, 2019  ref.218

Adelpha milleri Beutelspacher, 1976  Miller’s Sister  SE Mexico & Belize  M T S

Adelpha mythra (Godart, [1824])    iN  M T

Adelpha naxia (C. Felder & R. Felder, 1867)  Naxia Sister    iN

Adelpha naxia naxia (C. Felder & R. Felder, 1867)  W & S Mexico to Bolivia, Brazilian Amazon, Guianas  M T S L I Ha

Adelpha naxia hieronica Fruhstorfer, 1913  TL: Trinidad  M T

Adelpha naxia oteroi Neild, 1996  TL: Venezuela  M T

Adelpha olynthia (C. Felder & R. Felder, 1867)    iN  M T S

Adelpha phylaca (H. Bates, 1866)  Phylaca Sister    iN

Adelpha phylaca phylaca (H. Bates, 1866)  E & W Mexico to W Honduras  M T S1 S2 L

Adelpha phylaca joffrei Neild, 1996  TL: Venezuela  M T

Adelpha phylaca pseudaethalia A. Hall, 1938  S Mexico (Chiapas), to NW Venezuela & W Ecuador  M T S I1 I2

Adelpha plesaure Hübner, 1823    iN

Adelpha plesaure plesaure Hübner, 1823  TL: Brazil  M T

Adelpha plesaure phliassa (Godart, [1824])  TL: French Guiana; Brazil  M T S

Adelpha plesaure pseudomalea A. Hall, 1938  TL: Venezuela  M T

Adelpha plesaure symona Kaye, 1925  TL: Trinidad  M T

Adelpha pollina Fruhstorfer, 1915    iN  M T S

Adelpha poltius A. Hall, 1938    iN  M T

Adelpha rothschildi Fruhstorfer, 1913    iN  M T

Adelpha salmoneus (A. Butler, 1866)    iN

Adelpha salmoneus salmoneus (A. Butler, 1866)  TL: Venezuela  M T

Adelpha salmoneus colada (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Adelpha salmoneus emilia Fruhstorfer, 1908  TL: Colombia  M T

Adelpha salmoneus salmonides A. Hall, 1938  SE Mexico to C Panama  M T S1 S2 L I

Adelpha salus A. Hall, 1935    iN

Adelpha salus Northern segregate ,  SE Mexico to W Panama  M S

Adelpha salus salus A. Hall, 1935  TL: Colombia  M T L

Adelpha salus emmeli Willmott & J. Hall, 1999  TL: Ecuador

Adelpha salus [n. ssp.] Willmott, MS  Mexico

Adelpha saundersii (Hewitson, 1867)    iN

Adelpha saundersii saundersii (Hewitson, 1867)  TL: Ecuador  M T S L

Adelpha saundersii frontina A. Hall, 1935  TL: Colombia  M T

Adelpha saundersii helepecki Weeks, 1901  TL: Bolivia  M T S

Adelpha shuara Willmott & J. Hall, 1995    iN  M T S

Adelpha sichaeus (A. Butler, 1866)    iN  M T S L

Adelpha stilesiana DeVries & I. Chacón, 1982  Talamanca Sister  Talamanca massif, Costa Rica & W Panama, also Vol. Chiriqui  M T S

Adelpha syma (Godart, [1824])    iN  M T

Adelpha thesprotia (C. Felder & R. Felder, 1867)    iN  M T

Adelpha thessalia (C. Felder & R. Felder, 1867)    iN

Adelpha thessalia thessalia (C. Felder & R. Felder, 1867)  TL: Brazil (Am)  M T S

Adelpha thessalia cesilas Fruhstorfer, 1915  TL: Bolivia  M T

Adelpha thessalia indefecta Fruhstorfer, 1913  TL: Paraguay  M T

Adelpha thoasa (Hewitson, 1850)    iN

Adelpha thoasa thoasa (Hewitson, 1850)  TL: [Brazil (Pa)]  M T

Adelpha thoasa calliphiclea (A. Butler, [1870])  TL: Surinam  M T

Adelpha thoasa gerona (Hewitson, 1867)  TL: Brazil (MG)  M T

Adelpha thoasa manilia Fruhstorfer, 1915  TL: Bolivia  M T

Adelpha viola Fruhstorfer, 1913  TL: Brazil (SP)

Adelpha viola viola Fruhstorfer, 1913  TL: Brazil (SP)  M T

Adelpha viola pseudococala A. Hall, 1933  TL: Brazil (Pa); Guyana; Peru  M T

Adelpha ximena (C. Felder & R. Felder, 1862)    iN

Adelpha ximena ximena (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Adelpha ximena mossi A. Hall, 1933  TL: Brazil (Pa)  M T

Adelpha zina (Hewitson, 1867)  Zina Sister    iN

Adelpha zina zina (Hewitson, 1867)  E Panama to SW Ecuador & Venezuela  M T S L

Adelpha zina desousae Neild & Orellana, 1996  TL: Venezuela

Adelpha zina enope A. Hall, 1938  TL: Colombia  M T

Adelpha zina inachia Staudinger, 1886  TL: Colombia  M T

Adelpha zina irma Fruhstorfer, 1907  TL: Peru  M T S L

Adelpha zina lacina (A. Butler, 1872)  Nicaragua to W Panama  M T OI S1 S2 L I

Adelpha zina pyrczi Neild & Orellana, 1996  TL: Venezuela  M T

  Subfamily Apaturinae

Asterocampa  Röber, 1916  

Asterocampa celtis (Boisduval & Le Conte, [1835])  Hackberry Emperor    iN

Asterocampa celtis celtis (Boisduval & Le Conte, [1835])  E North America (except as others)  M T S Ab L1 L2 L3 L4

Asterocampa celtis reinthali Friedlander, 1987  SE USA coastal plain (SC to FL, W to E TX)  M T S1 S2 L

Asterocampa celtis antonia (W. H. Edwards, [1878])  [= jeffermont] SW USA (SE CA to TX, N to SE CO), N & C Mexico  M T S1 S2 L1 L2 I H

Asterocampa leilia (W. H. Edwards, 1874)  Empress leilia  S & C AZ, SW NM, S & W TX, E & NW Mexico, Baja California  iN  M T S L1 L2 L3 I H

Asterocampa clyton (Boisduval & Le Conte, [1835])  Tawny Emperor    iN

Asterocampa clyton clyton (Boisduval & Le Conte, [1835])  E North America (except as others)  M T S1 S2 Ab L1 L2 L3

Asterocampa clyton flora (W. H. Edwards, 1876)  SE USA coastal plain (SC to FL, W to E TX)  M T S1 S2 L I

Asterocampa clyton texana (Skinner, 1911)  S AZ, S NM, W & C TX, NW Mexico (SON, CHIH)  M T S L I

Asterocampa clyton louisa D. Stallings & J. Turner, 1947  S TX, NE Mexico (to SLP)  M T S L1 L2 I

Asterocampa idyja (Geyer, [1828])  Dusky Emperor    iN

Asterocampa idyja idyja (Geyer, [1828])  Cuba & Isla de Juventud, Hispaniola, Puerto Rico  M T S1 S2

Asterocampa idyja argus (H. Bates, 1864)  SE AZ (as vagrant), E & W Mexico to Costa Rica  M T S L1 L2 I

Doxocopa  Hübner, [1819]  

Doxocopa agathina (Cramer, 1777)    iN

Doxocopa agathina agathina (Cramer, 1777)  TL: Surinam  M T

Doxocopa agathina vacuna (Godart, [1824])  TL: Brazil  M T

Doxocopa callianira (Ménétriés, 1855)  Nicaraguan Emperor  S Mexico to Costa Rica  iN  M T S L

Doxocopa clothilda (C. Felder & R. Felder, 1867)  Clothilda Emperor  Costa Rica to Colombia  iN  M T S L

Doxocopa cyane (Latreille, [1813])    iN

Doxocopa cyane cyane (Latreille, [1813])  TL: [Peru]  M T S L

Doxocopa cyane burmeisteri (Godman & Salvin, 1884)  TL: Argentina  M T S

Doxocopa cyane mexicana Bryk, 1953  E & S Mexico to Colombia  M T S

Doxocopa cyane vespertina Lamas, 1999  TL: Peru  M T

Doxocopa elis (C. Felder & R. Felder, 1861)    iN  M T S

Doxocopa excelsa (Gillott, 1927)  Broad-banded Emperor  Nicaragua & Costa Rica  M T S

Doxocopa kallina (Staudinger, 1886)    iN  M T

Doxocopa laure (Drury, 1773)  Silver Emperor  ref.133  iN

Doxocopa laure laure (Drury, 1773)  continental; MX to Panama  M T S

Doxocopa laure laura (Hübner, [1823])  Jamaica  M T S

Doxocopa druryi (Hübner, [1825])  Silver Emperor  Cuba; ref.133

Doxocopa druryi druryi (Hübner, [1825])  Cuba  M S

Doxocopa druryi acca (C. Felder & R. Felder, 1867)  continental; S TX, SE AZ (uncommon) to Panama  M T S L1 L2 L3 I

Doxocopa druryi mima (Fruhstorfer, 1907)  TL: Colombia  M T

Doxocopa druryi griseldis (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Doxocopa druryi laurona (Schaus, 1902)  TL: Brazil (RJ)  M T

Doxocopa laurentia (Godart, [1824])    iN

Doxocopa laurentia laurentia (Godart, [1824])  TL: Brazil  M T Ab

Doxocopa laurentia cherubina (C. Felder & R. Felder, 1867)  S Mexico to Colombia  M T S L

Doxocopa laurentia thalysia (Fruhstorfer, 1907)  TL: Ecuador  M T

Doxocopa lavinia (A. Butler, 1866)    iN

Doxocopa lavinia ducaudi Brévignon, 2008  TL: French Guiana

Doxocopa lavinia lavinia (A. Butler, 1866)    iN  M T

Doxocopa linda (C. Felder & R. Felder, 1862)    iN

Doxocopa linda linda (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Doxocopa linda carwa Lamas, 1999  TL: Peru  M T

Doxocopa linda godmani (Dannatt, 1904)  TL: Venezuela

Doxocopa linda mileta (Boisduval, 1870)  TL: Brazil  M T

Doxocopa linda plesaurina (A. Butler & H. Druce, 1872)  Costa Rica & Panama  M T S

Doxocopa pavon (Latreille, [1809])    iN

Doxocopa pavon pavon (Latreille, [1809])  TL: Ecuador  M T

Doxocopa pavon theodora (Lucas, 1857)  S TX, E & W Mexico to Colombia  M T OI S L1 L2

Doxocopa thoe (Godart, [1824])  Hispaniolan Emperor  Hispaniola  iN  M T S

Doxocopa zalmunna (A. Butler, 1869)  TL: Brazil  M T

Doxocopa zunilda (Godart, [1824])    iN

Doxocopa zunilda zunilda (Godart, [1824])  TL: Brazil  M T

Doxocopa zunilda felderi (Godman & Salvin, 1884)  Costa Rica to Colombia  M T S L

Doxocopa zunilda floris (Fruhstorfer, 1907)  TL: Peru  M T

Doxocopa wilmattae (T. Cockerell, 1907)  Fossil from Colorado

  Subfamily Biblidinae

  Tribe Biblidini

Biblis  Fabricius, 1807  

Biblis aganisa Boisduval, 1836  Red Rim  SW USA & S TX (strays N), E & W Mexico at least to Costa Rica  M T S1 S2 L I

Biblis hyperia (Cramer, 1779)  Red Rim    iN

Biblis hyperia hyperia (Cramer, 1779)  Hispaniola, Mona, Puerto Rico, Culebra, Virgin Is. to St. Lucia  M T S L

Biblis hyperia laticlavia (Thieme, 1904)  TL: Ecuador  M T

Biblis hyperia rosanti Brévignon, 2006  TL: French Guiana

Biblis hyperia nectanabis (Fruhstorfer, 1909)  TL: Brazil (RS)  M T

Biblis hyperia pacifica (A. Hall, 1928)  TL: Ecuador  M T

Biblis hyperia [n. ssp.] Lamas, MS  Peru

Vila  W. F. Kirby, 1871  

Vila azeca (E. Doubleday, [1848])    iN

Vila azeca azeca (E. Doubleday, [1848])  TL: Bolivia  M T

Vila azeca cacica Staudinger, 1886  TL: Ecuador  M T

Vila azeca mariana (H. Bates, 1865)  TL: Brazil (Am)  M T

Vila azeca semistalachtis A. Hall, 1928  TL: Colombia  M T

Vila azeca stalachtoides (H. Bates, 1867)  TL: Brazil (Ma/Pa)  M T

Vila emilia (Cramer, 1779)    iN

Vila emilia emilia (Cramer, 1779)  TL: Surinam

Vila emilia caecilia (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S

Vila emilia carmenae Le Crom, 1990  TL: Colombia  M T

Vila emilia matteii Neild, 1996  TL: Venezuela  M T

Vila emilia sinefascia A. Hall, 1935  TL: [Peru; Brazil (Am)]  M T

Vila eueidiformis Joicey & Talbot, 1918    iN  M T

Mestra  Hübner, [1825]  

Mestra amymone (Ménétriés, 1857)  Northern Mestra  S TX (strays far N), E & W Mexico to Costa Rica  iN  M S Ma L1 L2 I1 I2 I3 H

Mestra dorcas (Fabricius, 1775)  Jamaican Mestra  Jamaica  iN  M T S1 S2 L

Mestra hersilia (Fabricius, 1776)  South American Mestra    iN

Mestra hersilia hersilia (Fabricius, 1776)  N S America, Trinidad, Tobago, Barbados, Grenada to Dominica  M T S L

Mestra hersilia latimargo (A. Hall, 1929)  TL: Ecuador  M T

Mestra hersilia semifulva (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Mestra hersilia apicalis (Staudinger, 1886)  TL: Brazil (SP)  M T

Mestra hersilia hypermestra Hübner, [1825]  TL: ?  M T

Archimestra  Munroe, 1949  

Archimestra teleboas (Ménétriés, 1832)  Archimestra  Hispaniola  iN  M S L

  Tribe Epicaliini

Catonephele  Hübner, [1819]  ref.198

Catonephele acontius (Linnaeus, 1771)    iN

Catonephele acontius acontius (Linnaeus, 1771)  TL: China  M T S

Catonephele acontius caeruleus Jenkins, 1985  TL: Bolivia

Catonephele antinoe (Godart, [1824])    iN  M T

Catonephele chromis E. Doubleday, [1848]    iN

Catonephele chromis chromis (E. Doubleday, [1848])  TL: Honduras  M T S L

Catonephele chromis godmani Stichel, 1901  Honduras to Panama  M T S

Catonephele cortesi R. G. Maza, 1982  West-Mexican Banner  W Mexico  iN  M T S L Ha

Catonephele mexicana Jenkins & R. G. Maza, 1985  East-Mexican Banner  E Mexico to Costa Rica  iN  M S1 S2 L I

Catonephele numilia (Cramer, 1775)    iN

Catonephele numilia numilia (Cramer, 1775)  TL: Surinam

Catonephele numilia esite (R. Felder, 1869)  E Mexico to W Ecuador, Venezuela & Trinidad  M T S1 S2 L1 L2 I1 I2

Catonephele numilia immaculata Jenkins, 1985  SW Mexico  M T S L Ha

Catonephele numilia neogermanica Stichel, 1899  TL: Paraguay  M T

Catonephele numilia penthia (Hewitson, 1852)  TL: Brazil (RJ)  M T S

Catonephele nyctimus (Westwood, 1850)    iN  M T

Catonephele orites Stichel, 1899  Broken Banner  Costa Rica to NW Ecuador  iN  M T S L

Catonephele sabrina (Hewitson, 1852)    iN  M T

Catonephele salacia (Hewitson, 1852)    iN  M T

Catonephele salambria (C. Felder & R. Felder, 1861)    iN  M T

Nessaea  Hübner, [1819]  ref.198

Nessaea aglaura (E. Doubleday, [1848])  Northern Nessaea    iN

Nessaea aglaura aglaura (E. Doubleday, [1848])  S Mexico to W Ecuador  M T S L

Nessaea aglaura regina (Salvin, 1869)  TL: Venezuela  M T

Nessaea aglaura thalia Bargmann, 1928  TL: [Colombia]  M T S

Nessaea batesii (C. Felder & R. Felder, 1860)    iN

Nessaea batesii batesii (C. Felder & R. Felder, 1860)  TL: [Brazil (Pa)]  M T

Nessaea batesii magniplaga Röber, 1928  TL: Bolivia

Nessaea hewitsonii (C. Felder & R. Felder, 1859)    iN

Nessaea hewitsonii hewitsonii (C. Felder & R. Felder, 1859)  TL: [Brazil (Am)]  M T S

Nessaea hewitsonii boliviensis Jenkins, 1989  TL: Bolivia  M T S

Nessaea obrinus (Linnaeus, 1758)    iN

Nessaea obrinus obrinus (Linnaeus, 1758)  TL: India  M S

Nessaea obrinus faventia Fruhstorfer, 1910  TL: Brazil (MT)  M T

Nessaea obrinus lesoudieri Le Moult, 1933  TL: Brazil (Am)  M T

Myscelia  E. Doubleday, [1845]  ref.198

Myscelia aracynthia (Dalman, 1823)  Hispaniolan Bluewing  Hispaniola  iN  M T S

Myscelia capenas (Hewitson, [1857])    iN

Myscelia capenas capenas (Hewitson, [1857])  TL: Brazil (Am)  M T

Myscelia capenas octomaculata (A. Butler, 1873)  TL: Peru  M T S

Myscelia cyananthe C. Felder & R. Felder, 1867  Blackened Bluewing    iN

Myscelia cyananthe cyananthe C. Felder & R. Felder, 1867  E & S Mexico  M T S Ab L

Myscelia cyananthe diaziana R. G. Maza & J. Maza, 1985  Chiapas  M T

Myscelia cyananthe skinneri Mengel, 1894  NW Mexico, stray to S AZ, S CA, S NM  M T S L

Myscelia cyananthe streckeri Skinner, 1889  Baja California  M T S L

Myscelia cyaniris E. Doubleday, [1848]  Whitened Bluewing    iN

Myscelia cyaniris cyaniris E. Doubleday, [1848]  E Mexico to Panama  M T S L

Myscelia cyaniris albescens A. Hall, 1935  TL: Venezuela  M T

Myscelia cyaniris alvaradia R. G. Maza & Díaz, 1982  W Mexico  M T S Ha

Myscelia cyaniris millerorum Jenkins, 1984  TL: Ecuador

Myscelia ethusa (Doyère, [1840])  Mexican Bluewing    iN

Myscelia ethusa ethusa (Doyère, [1840])  S TX, E & W Mexico  M T S L1 L2 I H

Myscelia ethusa chiapensis Jenkins, 1984  Chiapas  M T

Myscelia ethusa cyanecula C. Felder & R. Felder, 1867  TL: Mexico  M T S

Myscelia ethusa pattenia A. Butler & H. Druce, 1872  Guatemala to Costa Rica  M T S L

Myscelia hypatia Strecker, 1900  TL: [Brazil (Pa)]

Myscelia leucocyana C. Felder & R. Felder, 1861  Pale Bluewing    iN

Myscelia leucocyana leucocyana C. Felder & R. Felder, 1861  Panama to Venezuela  M T S L

Myscelia leucocyana smalli Jenkins, 1984  Nicaragua & Costa Rica  M T L

Myscelia milloi Oberthür, 1916    iN  M S

Myscelia orsis (Drury, 1782)    iN  M S

  Tribe Cybdelini

Cybdelis  Boisduval, 1836  ref.198

Cybdelis boliviana Salvin, 1869    iN

Cybdelis boliviana boliviana Salvin, 1869  TL: Bolivia  M T

Cybdelis boliviana [n. ssp.] Lamas, MS  Peru

Cybdelis mnasylus E. Doubleday, [1848]    iN  M T

Cybdelis petronita Burmeister, 1861  ref.198  M T

Cybdelis thrasylla C. Felder & R. Felder, 1859  ref.198  M T

Cybdelis phaesyla (Hübner, [1831])    iN  M S

Sea  Hayward, 1950  ref.198

Sea sophronia (Godart, [1824])  TL: Brazil

Sea sophronia sophronia (Godart, [1824])    iN  M S

Sea sophronia jenkinsi (F.M.S. Dias, R.R. Siewert & Freitas, 2019)  ref.198

  Tribe Eunicini

Eunica  Hübner, [1819]  ref.198

Eunica alcmena (E. Doubleday, [1847])  Dark Purplewing    iN

Eunica alcmena alcmena (E. Doubleday, [1847])  S Mexico to Colombia & W Venezuela  M T S L

Eunica alcmena flora C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T S

Eunica alpais (Godart, [1824])    iN

Eunica alpais alpais (Godart, [1824])  TL: Brazil  M T S

Eunica alpais excelsa Godman & Salvin, 1877  S Mexico (Oaxaca, Chiapas) to E Panama  M T S L

Eunica araucana C. Felder & R. Felder, 1862  Araucana Purplewing  W Panama, Colombia, W Venezuela, Ecuador  iN  M T S

Eunica amelia (Cramer, 1777)    iN

Eunica amelia amelia (Cramer, 1777)  TL: Surinam

Eunica amelia erroneata Oberthür, 1916  TL: Peru; Brazil (Am)  M T S

Eunica amycla (Godart, [1824])    iN  M T

Eunica anna (Cramer, 1780)    iN  M T S

Eunica bechina (Hewitson, 1852)    iN

Eunica bechina bechina (Hewitson, 1852)  TL: Brazil (Am)  M T

Eunica bechina magnipunctata Talbot, 1928  TL: Brazil (MT)  M T

Eunica brunnea Salvin, 1869  TL: Peru  M T

Eunica caelina (Godart, [1824])    iN

Eunica caelina caelina (Godart, [1824])  TL: Brazil  M T S

Eunica caelina agustina R. G. Maza, 1982  W Mexico  M T S

Eunica caelina alycia Fruhstorfer, 1909  TL: Upper Amazon  M T S

Eunica caelina augusta H. Bates, 1866  SE Mexico to W Panama  M T S

Eunica caelina olympias C. Felder & R. Felder, 1862  E Panama to Peru & Venezuela  M T S

Eunica caralis (Hewitson, [1857])  Caralis Purplewing    iN

Eunica caralis caralis (Hewitson, [1857])  Honduras, Panama, Colombia, Venezuela  M T S

Eunica caralis ariba Fruhstorfer, 1908  TL: Brazil?  M S

Eunica carias (Hewitson, [1857])  Carias Purplewing    iN

Eunica carias carias (Hewitson, [1857])  Costa Rica to Colombia  M T S L

Eunica carias cabira C. Felder & R. Felder, 1861  TL: Venezuela  M S

Eunica carias tenebrosa Salvin, 1869  TL: Peru  M T S

Eunica chlororhoa Salvin, 1869    iN

Eunica chlororhoa chlororhoa Salvin, 1869  TL: Peru  M T

Eunica chlororhoa mira Godman & Salvin, 1877  Honduras to Colombia  M T S L

Eunica clytia (Hewitson, 1852)  Clytia Purplewing  W Panama to Peru & C Brazil  iN  M T S L

Eunica concordia (Hewitson, 1852)    iN  M T S

Eunica cuvierii (Godart, 1819)    iN  M S

Eunica eburnea Fruhstorfer, 1907    iN  M T

Eunica eurota (Cramer, 1775)    iN

Eunica eurota eurota (Cramer, 1775)  TL: West Indies  M T

Eunica eurota dolores (Prittwitz, 1871)  TL: [Brazil (RJ)]  M T

Eunica evelide H. Bates, 1864    iN

Eunica evelide evelide H. Bates, 1864  TL: [Colombia]  M T S

Eunica evelide elegans Salvin, 1869  TL: Bolivia; Peru  M T S

Eunica heraclitus Gundlach, 1881  Cuban Purplewing  Cuba; apparently extinct  M IL

Eunica incognita Jenkins, 1990    iN  M T

Eunica ingens Seitz, 1915    iN  M T

Eunica interphasis Jenkins, 1990  TL: Brazil (Rd)  M T

Eunica macris (Godart, [1824])    iN

Eunica macris macris (Godart, [1824])  TL: Brazil  M T

Eunica macris [n. ssp.] Lamas, MS  Peru

Eunica maja (Fabricius, 1775)    iN

Eunica maja maja (Fabricius, 1775)  TL: Brazil  M T S

Eunica maja noerina A. Hall, 1935  TL: [Brazil (Am)]  M T S

Eunica malvina H. Bates, 1864    iN

Eunica malvina malvina H. Bates, 1864  TL: [Brazil (Pa)]  M T

Eunica malvina albida Jenkins, 1990  SE Mexico to Panama  M T S

Eunica malvina almae Vargas, Llorente & Luis, 1996  W Mexico  M T

Eunica margarita (Godart, [1824])    iN  M T

Eunica marsolia (Godart, [1824])    iN

Eunica marsolia marsolia (Godart, [1824])  TL: Brazil  M S

Eunica marsolia fasula Fruhstorfer, 1909  TL: [Peru]  M T S

Eunica marsolia paraensis Jenkins, 1990  TL: Brazil (Pa)  M T

Eunica monima (Stoll, 1782)  Dingy Purplewing  SE AZ & S TX (strays), E & W Mexico to Venez.; Gr. Antilles, S FL  iN  M T S L1 L2

Eunica mygdonia (Godart, [1824])    iN

Eunica mygdonia mygdonia (Godart, [1824])  TL: Brazil  M T S

Eunica mygdonia omoa A. Hall, 1919  S Mexico to Panama  M T S L

Eunica norica (Hewitson, 1852)  Midnight Purplewing    iN

Eunica norica norica (Hewitson, 1852)  Costa Rica to Colombia, Venezuela & W Ecuador  M T S L

Eunica norica occia Fruhstorfer, 1909  TL: Peru  M T S

Eunica orphise (Cramer, 1775)    iN  M T S

Eunica phasis C. Felder & R. Felder, 1862    iN  M T

Eunica pomona (C. Felder & R. Felder, 1867)    iN

Eunica pomona pomona (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Eunica pomona amata H. Druce, 1874  S Mexico (S Chiapas) to E Panama  M T TI S L

Eunica pusilla H. Bates, 1864  Pusilla Purplewing  Costa Rica to Bolivia & SE Brazil  iN  M T S L

Eunica sophonisba (Cramer, 1780)    iN

Eunica sophonisba sophonisba (Cramer, 1780)  TL: Surinam  M T S

Eunica sophonisba agele Seitz, 1915  TL: Brazil (Am); Peru; Colombia  M T

Eunica sydonia (Godart, [1824])    iN

Eunica sydonia sydonia (Godart, [1824])  TL: Brazil  M T S

Eunica sydonia caresa (Hewitson, [1857])  SE Mexico to E Panama  M T S L

Eunica tatila (Herrich-Schäffer, [1855])  Florida Purplewing    iN

Eunica tatila tatila (Herrich-Schäffer, [1855])  E & W Mexico (strays far N from E MX) to Costa Rica  M T S L

Eunica tatila bellaria Fruhstorfer, 1908  Panama to S Brazil & Argentina  M T S

Eunica tatila tatilina Fruhstorfer, 1908  TL: Upper Amazon?  M T

Eunica tatila tatilista Kaye, 1926  S FL, Bahamas, Greater Antilles, Mona  M S L H

Eunica veronica H. Bates, 1864    iN  M T

Eunica viola H. Bates, 1864    iN  M T S

Eunica violetta Staudinger, [1885]  TL: Peru  M T

Eunica volumna (Godart, [1824])    iN

Eunica volumna volumna (Godart, [1824])  TL: Brazil  M T S

Eunica volumna celma (Hewitson, 1852)  TL: [Brazil (Am)]  M T S

Eunica volumna venusia (C. Felder & R. Felder, 1867)  SE Mexico (Chiapas) to Colombia  M T S L

  Tribe Ageroniini

Batesia  C. Felder & R. Felder, 1862  

Batesia hypochlora C. Felder & R. Felder, 1862    iN

Batesia hypochlora hypochlora C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Batesia hypochlora hypoxantha Salvin & Godman, 1868  TL: Peru  M T S

Ectima  E. Doubleday, [1848]  

Ectima erycinoides C. Felder & R. Felder, 1867    iN

Ectima erycinoides Northern segregate ,  SE Mexico to Panama  M S L

Ectima erycinoides erycinoides C. Felder & R. Felder, 1867  TL: Colombia  M T

Ectima erycinoides ecuadorensis Jenkins, 1985  TL: Ecuador

Ectima erycinoides [n. ssp.] Lamas, MS  Mexico (Chia)

Ectima iona E. Doubleday, [1848]    iN  M T

Ectima lirides Staudinger, [1885]    iN  M T S

Ectima thecla (Fabricius, 1796)    iN

Ectima thecla thecla (Fabricius, 1796)  TL: Indiis  M T

Ectima thecla lirina C. Felder & R. Felder, 1867  TL: Brazil?  M T

Ectima thecla peruviana Bryk, 1953  TL: Peru

Hamadryas  Hübner, [1806]  

Hamadryas albicornis (Staudinger, [1885])  TL: Peru  M T

Hamadryas alicia (H. Bates, 1865)  TL: Brazil (Am)  M T

Hamadryas amphichloe (Boisduval, 1870)    iN

Hamadryas amphichloe amphichloe (Boisduval, 1870)  TL: Ecuador  M T

Hamadryas amphichloe lamasi Jenkins, 1983  TL: Peru  M T

Hamadryas amphinome (Linnaeus, 1767)    iN

Hamadryas amphinome amphinome (Linnaeus, 1767)  TL: S. Amer.  M T

Hamadryas amphinome fumosa (Fruhstorfer, 1915)  TL: Colombia  M T

Hamadryas amphinome mazai Jenkins, 1983  W Mexico  M S L Ha

Hamadryas amphinome mexicana (Lucas, 1853)  E Mexico to Colombia, Cuba  M T S L

Hamadryas arete (E. Doubleday, 1847)    iN  M T

Hamadryas arinome (Lucas, 1853)    iN

Hamadryas arinome arinome (Lucas, 1853)  TL: French Guiana  M T

Hamadryas arinome arienis (Godman & Salvin, 1883)  Costa Rica to Bolivia  M T S L

Hamadryas arinome obnubila (Fruhstorfer, 1916)  TL: Brazil (ES)  M T

Hamadryas atlantis (H. Bates, 1864)  Black-patched Cracker    iN

Hamadryas atlantis atlantis (H. Bates, 1864)  S Mexico (Chiapas), Guatemala, Honduras  M T S

Hamadryas atlantis lelaps (Godman & Salvin, 1883)  W Mexico  M T S L

Hamadryas belladonna (H. Bates, 1865)    iN  M T

Hamadryas chloe (Stoll, 1787)    iN

Hamadryas chloe chloe (Stoll, 1787)  TL: Surinam  M T

Hamadryas chloe daphnis (Staudinger, 1886)  TL: Peru  M T

Hamadryas chloe obidona (Fruhstorfer, 1914)  TL: Brazil (Pa)  M T

Hamadryas chloe rhea (Fruhstorfer, 1907)  TL: Brazil (ES/MG)  M T

Hamadryas epinome (C. Felder & R. Felder, 1867)    iN  M T

Hamadryas februa (Hübner, [1823])    iN

Hamadryas februa NW Mexican segregate ,  NW Mexico  M S L I

Hamadryas februa ferentina (Godart, [1824])  S TX, E & S Mexico to Colombia, Trinidad, Guianas & Amazon  M T S L1 L2 I

Hamadryas februa ferox (Staudinger, 1886)  S FL (rare stray), Caribbean isls., Pan to Venez; ref.135,136  M T S1 S2 L

Hamadryas februa februa (Hübner, [1823])  TL: ?  M T

Hamadryas feronia (Linnaeus, 1758)    iN

Hamadryas feronia feronia (Linnaeus, 1758)  TL: India  M T HS

Hamadryas feronia farinulenta (Fruhstorfer, 1916)  S TX (rare stray), E & W Mexico to W Peru, Venezuela, Trinidad  M T S L

Hamadryas fornax (Hübner, [1823])    iN

Hamadryas fornax fornax (Hübner, [1823])  TL: ?

Hamadryas fornax fornacalia (Fruhstorfer, 1907)  E Mexico to Colombia & Venezuela  M T S L

Hamadryas glauconome (H. Bates, 1864)  Glaucous Cracker    iN

Hamadryas glauconome glauconome (H. Bates, 1864)  S TX (rare stray), E & S Mexico to Costa Rica (Panama?)  M T S L

Hamadryas glauconome grisea Jenkins, 1983  NW Mexico  M T S L

Hamadryas guatemalena (H. Bates, 1864)  Guatamalan Cracker    iN

Hamadryas guatemalena guatemalena (H. Bates, 1864)  SE Mexico (much of Chiapas, Yucatan Peninsula) to Costa Rica  M T S L

Hamadryas guatemalena marmarice (Fruhstorfer, 1916)  S TX, most of Mexico  M T S L I

Hamadryas iphthime (H. Bates, 1864)  Brownish Cracker    iN

Hamadryas iphthime joannae Jenkins, 1983  S TX (rare stray), E & S Mexico to Costa Rica  M S1 S2 L1 L2

Hamadryas iphthime iphthime (H. Bates, 1864)  Costa Rica, Panama to Guyana, Colombia, Peru, Bolivia, S Brazil  M T L

Hamadryas iphthime joannae Jenkins, 1983  S TX (rare stray), E & S Mexico to Costa Rica  M S1 S2 L1 L2

Hamadryas iphthime griffoni Brévignon, 2006  TL: French Guiana

Hamadryas julitta (Fruhstorfer, 1914)  Yucatan Cracker  [= honorina], Yucatan Peninsula  iN  M T S L

Hamadryas laodamia (Cramer, 1777)    iN

Hamadryas laodamia laodamia (Cramer, 1777)  TL: Surinam  M T

Hamadryas laodamia saurites (Fruhstorfer, 1916)  SE Mexico to Colombia, N Venezuela, Trinidad  M T S L I

Hamadryas rosandra (Fruhstorfer, 1916)  TL: Brazil (Am)  M T

Hamadryas velutina (H. Bates, 1865)    iN

Hamadryas velutina velutina (H. Bates, 1865)  TL: [Brazil (Am)]  M T

Hamadryas velutina browni Jenkins, 1983  TL: Brazil (MT)  M T

Panacea  Godman & Salvin, 1883  

Panacea procilla (Hewitson, [1854])    iN

Panacea procilla procilla (Hewitson, [1854])  TL: Colombia  M T S

Panacea procilla bleuzeni Plantrou & Attal, 1986  TL: French Guiana  M T

Panacea procilla divalis (H. Bates, 1868)  TL: Peru  M T S

Panacea procilla lysimache Godman & Salvin, 1883  Costa Rica & Panama  M T

Panacea procilla mamorensis A. Hall, 1917  TL: Brazil (Rd)  M T

Panacea prola (E. Doubleday, [1848])    iN

Panacea prola prola (E. Doubleday, [1848])  TL: Colombia  M T S

Panacea prola amazonica Fruhstorfer, 1915  TL: [Peru]  M T S

Panacea regina (H. Bates, 1864)    iN

Panacea regina regina (H. Bates, 1864)  TL: Brazil (Am)  M T S

Panacea regina chalcothea (H. Bates, 1868)  TL: Colombia; [Peru]  M T S

  Tribe Epiphilini

Asterope  Hübner, [1819]  

Asterope batesii (Hewitson, 1850)  TL: Brazil (Am)  M T

Asterope buckleyi (Hewitson, 1869)  TL: Ecuador  M T

Asterope degandii (Hewitson, 1858)  Red-based Banner    iN

Asterope degandii degandii (Hewitson, 1858)  Peru  M T OI S

Asterope degandii adamsi (Lathy, 1903)  TL: Peru  M T

Asterope degandii allyni Jenkins, 1987  TL: Peru  M T

Asterope degandii bartletti (Godman & Salvin, 1878)  TL: Peru  M T

Asterope degandii caelestis Attal & Hiltbrand, 2006  ref.160  M T

Asterope degandii orientalis Attal, 1999  TL: Brazil (Pa)  M T

Asterope degandii mocoensis Attal, Pacl & Hiltbrand, 2020  ref.161

Asterope leprieuri (Feisthamel, 1835)    iN

Asterope leprieuri leprieuri (Feisthamel, 1835)  TL: [Brazil (Pa)]  M T

Asterope leprieuri depuiseti (C. Felder & R. Felder, 1861)  TL: Ecuador  M T S

Asterope leprieuri optima (A. Butler, 1869)  TL: Peru  M T

Asterope leprieuri philotima (Rebel, 1912)  TL: Peru  M T S

Asterope markii (Hewitson, 1857)    iN

Asterope markii markii (Hewitson, 1857)  TL: [Brazil (Am)]  M T S

Asterope markii ackeryi Jenkins, 1987  TL: Brazil (Pa)  M T

Asterope markii boyi (Röber, 1924)  TL: Brazil ([Pa])  M T

Asterope markii davisii (A. Butler, 1877)  TL: Peru  M T

Asterope markii gallardi Neukirchen, 1996  TL: French Guiana

Asterope markii hewitsoni (Staudinger, 1886)  TL: Colombia  M T

Asterope markii werneri Neukirchen, 1995  TL: Brazil (Am)  M T

Asterope sapphira (Hübner, [1816])  TL: Brazil  M T

Asterope whitelyi (Salvin, 1869)  TL: Peru

Asterope whitelyi whitelyi (Salvin, 1869)  TL: Peru  M T

Asterope whitelyi cyaneogyna Attal & Hiltbrand, 2006  ref.160  M T

Asterope whitelyi glennae Attal & Hiltbrand, 2006  ref.160  M T

Asterope whitelyi srnkai (Honrath, 1884)  TL: Peru  M T

Bolboneura  Godman & Salvin, 1877  

Bolboneura sylphis (H. Bates, 1864)  Variable Banner    iN

Bolboneura sylphis sylphis (H. Bates, 1864)  C & W Chiapas,Guatemala, Honduras  M T S L

Bolboneura sylphis beatrix R. G. Maza, 1985  SW Mexico (MICH, GRO, MOR, SW PUE, C & S OAX)  M T L

Bolboneura sylphis lacandona R. G. Maza & J. Maza, 1985  E Chiapas  M T

Bolboneura sylphis veracruzana (Draudt, 1931)  E Mexico  M T S L

Epiphile  E. Doubleday, [1845]  

Epiphile adrasta Hewitson, 1861  Common Banner    iN

Epiphile adrasta adrasta Hewitson, 1861  S TX (as vagrant), E Mexico to W Panama  M T S L

Epiphile adrasta bandusia Fruhstorfer, 1912  C Panama to W Ecuador  M T S L

Epiphile adrasta calixto Attal & Crosson, 2003  ref.159

Epiphile adrasta escalantei Descimon & Mast, 1979  W Mexico (SIN to S OAX)  M S L

Epiphile boliviana Röber, 1914    iN

Epiphile boliviana boliviana Röber, 1914  TL: Bolivia  M T

Epiphile boliviana grosi Attal & Crosson, 2003  ref.159  M T

Epiphile boliviana lamasi Neild, 1996  TL: Venezuela  M T

Epiphile chrysites (Latreille, [1809])  Chrysites Banner    iN

Epiphile chrysites chrysites (Latreille, [1809])  [W Panama?], Venezuela to Bolivia  M T S L

Epiphile chrysites [n. ssp.] Lamas, MS  Bolivia  M T

Epiphile dilecta Röber, 1914    iN

Epiphile dilecta dilecta Röber, 1914  TL: Bolivia  M T

Epiphile dilecta jenkinsi Lamas, 1995  TL: Peru

Epiphile dilecta venezuelensis Jenkins, 1986  TL: Venezuela  M T

Epiphile epicaste Hewitson, 1857    iN

Epiphile epicaste epicaste Hewitson, 1857  TL: Colombia  M T

Epiphile epicaste [n. ssp.] Lamas, MS  Peru

Epiphile epimenes Hewitson, 1857    iN

Epiphile epimenes epimenes Hewitson, 1857  TL: Colombia  M T

Epiphile epimenes electra Staudinger, [1885]  TL: Venezuela  M T

Epiphile epimenes kalbreyeri Fassl, 1912  TL: Colombia  M T

Epiphile eriopis Hewitson, 1857    iN

Epiphile eriopis eriopis Hewitson, 1857  TL: Colombia  M T

Epiphile eriopis devriesi Jenkins, 1986  Nicaragua to Colombia  M T S

Epiphile grandis A. Butler, 1872  Grand Banner  Costa Rica & Panama  M T OI S

Epiphile hermosa J. Maza & Díaz, 1978  Beautiful Banner  SE Mexico, Guatemala  iN  M T S L

Epiphile hubneri Hewitson, 1861    iN

Epiphile iblis C. Felder & R. Felder, 1861    iN

Epiphile iblis iblis C. Felder & R. Felder, 1861  TL: Venezuela; Colombia  M T

Epiphile iblis plusios Godman & Salvin, 1883  Costa Rica & Panama  M T S L

Epiphile iblis plutonia H. Bates, 1864  SE Mexico & Guatemala  M T S

Epiphile iblis pyrczi Attal, 1999  TL: Colombia  M T

Epiphile imperator Attal, 2005  TL: Ecuador

Epiphile imperator imperator Attal, 2005  ref.157  M T

Epiphile imperator duponti Attal, 2005  ref.157  M T

Epiphile imperator willmotti Attal, 2005  ref.157

Epiphile lampethusa E. Doubleday, [1848]    iN

Epiphile lampethusa lampethusa E. Doubleday, [1848]  TL: Bolivia  M T

Epiphile lampethusa zipa Mengel, 1899  TL: not specified  M T

Epiphile latifasciata Röber, 1914    iN

Epiphile latifasciata latifasciata Röber, 1914  TL: Bolivia  M T

Epiphile latifasciata obscurior Attal, 1997  TL: Ecuador  M T

Epiphile orea (Hübner, [1823])    iN

Epiphile orea orea (Hübner, [1823])  TL: ?

Epiphile orea distalis Attal, 1997  TL: Ecuador  M T

Epiphile orea helios Attal & Crosson, 2003  ref.159

Epiphile orea negrina C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T S

Epiphile neildi Attal, 2005    iN

Epiphile neildi neildi Attal, 2005  ref.157  M T

Epiphile neildi bakuru Attal & G. Rodríguez, 2006  ref.160  M T

Lucinia  Hübner, [1823]  

Lucinia sida Hübner, [1823]  Caribbean Banner    iN

Lucinia sida sida Hübner, [1823]  Cuba  M T S

Lucinia sida albomaculata Rindge, 1955  Bahamas  M T S

Lucinia sida torrebia (Ménétriés, 1832)  Hispaniola  M S

Lucinia cadma (Drury, 1773)  Jamaican Banner  Jamaica  M T S

Nica  Hübner, [1826]  

Nica flavilla (Godart, [1824])    iN

Nica flavilla bachiana (R. G. Maza & J. Maza, 1985)  W Mexico  M T S L

Nica flavilla E Mexican segregate ,  E Mexico  M S L

Nica flavilla canthara (E. Doubleday, 1849)  Nicaragua to Venezuela (at least)  M T S L1 L2

Nica flavilla flavilla (Godart, [1824])  TL: Brazil  M T

Nica flavilla lucillae Neild, 1996  TL: Venezuela  M T

Nica flavilla sylvestris H. Bates, 1864  TL: Brazil (Am)  M T

Nica flavilla [n. ssp.] Lamas, MS  Mexico

Peria  W. F. Kirby, 1871  

Peria lamis (Cramer, 1779)    iN  M S

Pyrrhogyra  Hübner, [1819]  

Pyrrhogyra amphiro H. Bates, 1865    iN

Pyrrhogyra amphiro amphiro H. Bates, 1865  TL: Brazil (Am)  M T S

Pyrrhogyra amphiro agilis Brévignon, 1995  TL: French Guiana

Pyrrhogyra amphiro juani Staudinger, 1886  TL: Colombia  M T

Pyrrhogyra amphiro typhoeus C. Felder & R. Felder, 1867  TL: Colombia  M T

Pyrrhogyra crameri Aurivillius, 1882    iN

Pyrrhogyra crameri Northern segregate ,  Nicaragua to Panama  M L

Pyrrhogyra crameri nautaca Fruhstorfer, 1908  TL: Colombia; "Upper Amazon"  M T

Pyrrhogyra crameri undine Bargmann, 1929  TL: Colombia  M T

Pyrrhogyra crameri hagnodorus Fruhstorfer, 1908  TL: Peru  M T

Pyrrhogyra crameri crameri Aurivillius, 1882  TL: Surinam  M T

Pyrrhogyra crameri [n. ssp.] Lamas, MS  Panama

Pyrrhogyra edocla (E. Doubleday, [1848])  Green-spotted Banner    iN

Pyrrhogyra edocla edocla (E. Doubleday, [1848])  E Mexico to Colombia & Venezuela  M T S L

Pyrrhogyra edocla cuparina H. Bates, 1865  TL: Brazil (Pa)  M T

Pyrrhogyra edocla lysanias C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T S

Pyrrhogyra edocla paradisea R. G. Maza & J. Maza, 1985  W Mexico  M T S

Pyrrhogyra edocla perscita Brévignon, 1996  TL: French Guiana

Pyrrhogyra neaerea (Linnaeus, 1758)    iN

Pyrrhogyra neaerea neaerea (Linnaeus, 1758)  TL: Indiis  M T HS

Pyrrhogyra neaerea arge Gosse, 1880  TL: Paraguay  M T L

Pyrrhogyra neaerea argina Fruhstorfer, 1908  TL: Upper Amazon  M T

Pyrrhogyra neaerea hypsenor Godman & Salvin, 1884  E & W Mexico to Panama  M T S L

Pyrrhogyra neaerea kheili Fruhstorfer, 1908  TL: Colombia  M T

Pyrrhogyra neaerea ophni A. Butler, 1870  TL: Brazil (MG)  M T

Pyrrhogyra neaerea susarion Fruhstorfer, 1912  TL: Brazil (Pe)  M T

Pyrrhogyra otolais H. Bates, 1864  Double-banded Banner    iN

Pyrrhogyra otolais otolais H. Bates, 1864  E & W Mexico to Panama  M T S L

Pyrrhogyra otolais nasica Staudinger, 1886  TL: Colombia  M T

Pyrrhogyra otolais olivenca Fruhstorfer, 1908  TL: Brazil (Am)  M T S

Pyrrhogyra otolais placita Brévignon, 1996  TL: French Guiana

Pyrrhogyra otolais seitzi Fruhstorfer, 1908  TL: Bolivia  M T

Pyrrhogyra stratonicus Fruhstorfer, 1908    iN

Pyrrhogyra stratonicus stratonicus Fruhstorfer, 1908  TL: Upper Amazon  M T S

Pyrrhogyra stratonicus matthias Brévignon, 1995  TL: French Guiana

Temenis  Hübner, [1819]  

Temenis huebneri Fruhstorfer, 1907    iN

Temenis huebneri huebneri Fruhstorfer, 1907  TL: Brazil (MG)  M T

Temenis huebneri korallion Fruhstorfer, 1912  TL: Brazil (SP)  M T

Temenis huebneri pedania Fruhstorfer, 1912  TL: Brazil (Pe)  M T

Temenis laothoe (Cramer, 1777)    iN

Temenis laothoe quilapayunia R. G. Maza & Turrent, 1985  W Mexico  M T S L Ha

Temenis laothoe E Mexican segregate ,  S TX (as vagrant), E Mexico  M S L

Temenis laothoe hondurensis Fruhstorfer, 1907  S Mexico (Chiapas) to Colombia  M T S L I

Temenis laothoe laothoe (Cramer, 1777)  TL: Surinam  M T

Temenis laothoe bahiana Fruhstorfer, 1907  TL: Brazil (Ba)  M T

Temenis laothoe columbiana Fruhstorfer, 1907  TL: Colombia  M T

Temenis laothoe meridionalis Ebert, 1965  TL: Brazil (SP)  M T S

Temenis laothoe santina Fruhstorfer, 1907  TL: Brazil (ES); Paraguay  M T

Callicorina  Smart, 1975  

Callicorina pulchra (Hewitson, 1861)  ref.12  iN

Callicorina pulchra Northern segregate ,  Costa Rica & Panama

Callicorina pulchra pulchra (Hewitson, 1861)  TL: Colombia  M T

Callicorina pulchra amazonica Fruhstorfer, 1907  TL: [Brazil (Am)]

Callicorina pulchra pallidior (Oberthür, 1901)  TL: Peru; Bolivia; Brazil (Pa)  M T

Callicorina pulchra [n. ssp.] Lamas, MS  Panama

  Tribe Eubagini

Dynamine  Hübner, [1819]  

Dynamine aerata (A. Butler, 1877)    iN

Dynamine aerata aerata (A. Butler, 1877)  TL: Brazil (Am)  M T S

Dynamine aerata [n. ssp.#1] Lamas, MS  TL: Venezuela

Dynamine aerata [n. ssp.#2] Lamas, MS  TL: Paraguay

Dynamine agacles (Dalman, 1823)    iN

Dynamine agacles agacles (Dalman, 1823)  TL: Brazil  M T

Dynamine agacles core Röber, 1915  Costa Rica to Colombia  M T S L

Dynamine agatha (Oberthür, 1916)  TL: Brazil (Am)  M T

Dynamine amplias (Hewitson, 1859)    iN  M T

Dynamine anubis (Hewitson, 1859)    iN

Dynamine anubis anubis (Hewitson, 1859)  TL: Brazil (Am)  M T

Dynamine anubis pieridoides (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Dynamine arene Hübner, [1823]    iN  M S

Dynamine artemisia (Fabricius, 1793)    iN

Dynamine artemisia Northern segregate ,  E & S Mexico to Panama  M S Ab L

Dynamine artemisia artemisia (Fabricius, 1793)  TL: America  M S

Dynamine artemisia ackeryi Neild, 1996  TL: Venezuela  M T

Dynamine artemisia glauce (H. Bates, 1865)  TL: Brazil (Am); "Guatemala"  M T S

Dynamine artemisia [n. ssp.#1] Lamas, MS  TL: Mexico

Dynamine artemisia [n. ssp.#2] Lamas, MS  TL: Venezuela

Dynamine ate (Godman & Salvin, 1883)  Little Sailor  SE Mexico (SE Chiapas), Guatemala, Honduras, Costa Rica  iN  M T S

Dynamine athemon (Linnaeus, 1758)    iN

Dynamine athemon athemon (Linnaeus, 1758)  TL: America

Dynamine athemon amazonica Röber, 1915  TL: Brazil (Am)  M S

Dynamine athemon athemaena (Hübner, [1824])  TL: Brazil

Dynamine athemon barreiroi A. Fernández, 1928  TL: Peru  M T L

Dynamine athemon isolda A. Hall, 1919  TL: Colombia  M T

Dynamine athemon maeon (E. Doubleday, 1849)  Panama & Colombia  M T OI L

Dynamine athemon niveata (A. Butler, 1877)  TL: Brazil (Am)  M T

Dynamine chiquita Willmott & J. Hall, 2010  ref.320

Dynamine chryseis (H. Bates, 1865)  Thick-rimmed Sailor  Nicaragua to Amazon Basin  iN  M T S L

Dynamine coenus (Fabricius, 1793)    iN

Dynamine coenus coenus (Fabricius, 1793)  TL: Indiis  M T

Dynamine coenus albidula Weeks, 1901  TL: Bolivia  M T

Dynamine coenus leucothea (H. Bates, 1865)  TL: Brazil (Am)  M T

Dynamine colombiana Talbot, 1932  Colombian Sailor  Nicaragua to Colombia & NW Venezuela  iN  M T L

Dynamine dyonis Geyer, 1837  Blue-eyed Sailor  S TX, E & W Mexico to Panama  iN  M S L

Dynamine erchia (Hewitson, 1852)    iN  M T

Dynamine gisella (Hewitson, 1857)    iN  M T

Dynamine haenschi A. Hall, 1917    iN  M T

Dynamine hecuba (Schaus, 1913)  Costa Rican Sailor  Costa Rica  M T S

Dynamine ines (Godart, [1824])    iN

Dynamine ines ines (Godart, [1824])  TL: Brazil  M T

Dynamine ines geta (Godman & Salvin, 1878)  TL: Bolivia  M S

Dynamine ines [n. ssp.] Lamas, MS  Colombia

Dynamine intermedia Talbot, 1932  TL: Peru  M T

Dynamine laugieri (Oberthür, 1916)  TL: French Guiana  M T

Dynamine meridionalis Röber, 1915    iN

Dynamine myrrhina (E. Doubleday, 1849)    iN  M T

Dynamine myrson (E. Doubleday, 1849)    iN

Dynamine myrson myrson (E. Doubleday, 1849)  TL: Brazil (Pa)  M T S

Dynamine myrson manriquei Neild, 1996  TL: Peru  M T

Dynamine neoris (Hewitson, 1859)  TL: [Brazil (Am)]  M T

Dynamine onias (Hewitson, 1857)    iN  M T

Dynamine paulina (H. Bates, 1865)    iN

Dynamine paulina paulina (H. Bates, 1865)  TL: Brazil (Am)  M T

Dynamine paulina thalassina (Boisduval, 1870)  Guatemala to Colombia  M T S L

Dynamine pebana Staudinger, [1885]    iN

Dynamine pebana pebana Staudinger, [1885]  TL: Peru  M T

Dynamine pebana elisa Brévignon, 1995  TL: French Guiana

Dynamine perpetua (H. Bates, 1865)  TL: Brazil (Am)  M T

Dynamine persis (Hewitson, 1859)  TL: Brazil (Am)  M T

Dynamine postverta (Cramer, 1779)    iN

Dynamine postverta postverta (Cramer, 1779)  TL: Surinam  M S

Dynamine postverta mexicana R.F. d'Almeida, 1952  [= mylitta], S TX (rare stray), E & W Mexico, C America, Cuba  M T S L1 L2

Dynamine racidula (Hewitson, 1852)    iN

Dynamine racidula racidula (Hewitson, 1852)  TL: Brazil (Am)  M T S L

Dynamine racidula trembathi Neild, 1996  TL: Venezuela  M T

Dynamine sara (H. Bates, 1865)    iN  M T S

Dynamine serina (Fabricius, 1775)  Caribbean Sailor    iN

Dynamine serina serina (Fabricius, 1775)  [= egaea], Jamaica  M T S

Dynamine serina calais M. Bates, 1936  Cuba  M T IL

Dynamine serina zetes (Ménétriés, 1832)  Hispaniola  M S

Dynamine setabis (E. Doubleday, 1849)    iN

Dynamine setabis setabis (E. Doubleday, 1849)  TL: Venezuela; Colombia  M T

Dynamine setabis agnes Röber, 1915  TL: Peru  M T S

Dynamine sosthenes (Hewitson, 1869)  Thin-rimmed Sailor    iN

Dynamine sosthenes sosthenes (Hewitson, 1869)  Nicaragua & Costa Rica  M T S

Dynamine sosthenes smerdis Tessmann, 1928  TL: Peru  M T S

Dynamine sosthenes venezolensis Tessmann, 1928  TL: Venezuela  M T

Dynamine theseus (C. Felder & R. Felder, 1861)  White Sailor  S Mexico to Colombia & Venezuela  iN  M T S L

Dynamine tithia (Hübner, 1823)    iN

Dynamine tithia tithia (Hübner, [1823])  TL: Brazil  M T

Dynamine tithia dentilinea Talbot, 1932  TL: Ecuador  M T

Dynamine tithia salpensa (C. Felder & R. Felder, 1862)  Andes and upper Amazon basin, Colombia to Bolivia (hypothetical for Central America)  M T S

Dynamine tithia [n. ssp.] Lamas, MS  Colombia

Dynamine vicaria (H. Bates, 1865)    iN

Dynamine vicaria vicaria (H. Bates, 1865)  TL: Brazil (Am)  M T

Dynamine vicaria gillotti A. Hall, 1930  Costa Rica  M T S

Dynamine vicaria hoppi M. Hering, 1926  TL: Colombia  M T

Dynamine zenobia (H. Bates, 1865)  TL: Brazil (Am)

Dynamine zenobia zenobia (H. Bates, 1865)  TL: Brazil (Am)  M T S

Dynamine zenobia ampliata J. Zikán, 1937  TL: Colombia  M T

Dynamine [n. sp.] Lamas, MS  Peru

  Tribe Callicorini

Haematera  E. Doubleday, 1849  

Haematera pyrame (Hübner, [1819])    iN

Haematera pyrame Northern segregate ,  Nicaragua to Colombia  M S L

Haematera pyrame pyrame (Hübner, [1819])  TL: French Guiana  M T

Haematera pyrame rubra Kaye, 1904  TL: Trinidad

Haematera pyrame thysbe E. Doubleday, 1849  TL: Venezuela; Colombia  M T

Haematera pyrame [n. ssp.#1] Lamas, MS  TL: Panama

Haematera pyrame [n. ssp.#2] Lamas, MS  TL: Peru

Haematera pyrame [n. ssp.#3] Lamas, MS  TL: Peru

Antigonis  C. Felder, 1861  

Antigonis pharsalia (Hewitson, 1852)  TL: [Brazil (Am)]

Antigonis pharsalia pharsalia (Hewitson, 1852)  TL: [Brazil (Am)]  M T

Antigonis pharsalia felderi H. Bates, 1864  TL: Brazil (Am)  M T

Callicore  Hübner, [1819]  

Callicore astarte (Cramer, 1779)    iN

Callicore astarte astarte (Cramer, 1779)  TL: Surinam  M T

Callicore astarte antillena (Kaye, 1914)  TL: Trinidad; "St. Lucia"

Callicore astarte astartoides (Dillon, 1948)  TL: Bolivia  M T

Callicore astarte casta (Salvin, 1869)  Veracruz & Oaxaca, Mexico  M T S

Callicore astarte codomannus (Fabricius, 1781)  TL: Brazil  M T

Callicore astarte lilliputa (Dillon, 1948)  TL: Colombia  M T

Callicore astarte otheres (Fruhstorfer, 1916)  TL: Colombia

Callicore astarte panamaensis Kariya, 2006  TL: Panama

Callicore astarte patelina (Hewitson, 1853)  Chiapas to Costa Rica  M T S L

Callicore astarte selima (Guenée, 1872)  TL: Brazil (MG)  M T

Callicore astarte staudingeri (Dillon, 1948)  Panama (Darien), Colombia (Antioquia), Venezuela (Aragua, Trujillo, Perija)  M T S

Callicore astarte stratiotes (C. Felder & R. Felder, 1861)  TL: Ecuador  M T S

Callicore astarte [n. ssp.] Lamas, MS  Brazil

Callicore ines (Hopp, 1922)    iN  M T

Callicore cynosura (E. Doubleday, [1847])    iN

Callicore cynosura cynosura (E. Doubleday, [1847])  TL: Bolivia  M T S

Callicore cynosura amazona (H. Bates, 1864)  TL: Brazil (Pa)  M T

Callicore cynosura fulva (Dillon, 1948)  TL: Brazil (Am)  M T

Callicore excelsior (Hewitson, [1858])    iN

Callicore excelsior excelsior (Hewitson, [1858])  TL: Brazil (Am)  M T

Callicore excelsior arirambae (Ducke, 1913)  TL: Brazil (Pa)  M T

Callicore excelsior elatior (Oberthür, 1916)  TL: Ecuador

Callicore excelsior excelsissima (Staudinger, [1885])  TL: Brazil (Am)  M T

Callicore excelsior inferior (A. Butler, 1877)  TL: Peru  M T

Callicore excelsior marisolae Neukirchen, 1995  TL: Brazil (Am)  M T

Callicore excelsior mauensis (Fassl, 1922)  TL: Brazil (Am)

Callicore excelsior michaeli (Staudinger, 1890)  TL: Brazil (Am)  M T

Callicore excelsior micheneri (Dillon, 1948)  TL: Colombia  M T

Callicore excelsior ockendeni (Oberthür, 1916)  TL: Peru  M T

Callicore excelsior pastazza (Staudinger, 1886)  TL: Peru  M T

Callicore excelsior splendida (Weymer, 1890)  TL: Peru  M T

Callicore sorana (Godart, [1824])    iN

Callicore sorana sorana (Godart, [1824])  TL: Brazil  M T L

Callicore sorana horstii (Mengel, 1916)  TL: Bolivia  M T

Callicore cyclops (Staudinger, 1891)    iN  M T

Callicore pitheas (Latreille, [1813])  Two-eyed Eighty-eight  W Chiapas to Venezuela & W Ecuador  iN  M T S1 S2 L I1 I2

Callicore texa (Hewitson, [1855])    iN

Callicore texa texa (Hewitson, [1855])  TL: Colombia  M T

Callicore texa aretas (Hewitson, 1857)  TL: Venezuela  M T

Callicore texa bari (Oberthür, 1916)  TL: French Guiana; "Peru"  M T

Callicore texa boyeri Attal, 1999  TL: Ecuador  M T

Callicore texa fassli (Röber, 1915)  TL: Bolivia  M T

Callicore texa heroica (Fruhstorfer, 1916)  Veracruz, N Oaxaca  M T S

Callicore texa kayei (A. Hall, 1917)  TL: Trinidad  M T

Callicore texa loxicha R. G. Maza & J. Maza, 1983  S Oaxaca (Sierra Madre del Sur)  M T S L

Callicore texa maimuna (Hewitson, 1858)  TL: [Peru]  M T

Callicore texa maximilla (Fruhstorfer, 1916)  TL: Brazil (ES)  M T

Callicore texa sigillata (Kotzsch, 1939)  TL: Peru  M T

Callicore texa skinneri Neild, 1996  TL: Venezuela

Callicore texa strympli (Fassl, 1922)  TL: Brazil (Pa)

Callicore texa tacana R. G. Maza & J. Maza, 1983  W Chiapas to Guatemala  M T S

Callicore texa titania (Salvin, 1869)  far SE Mexico (E Chiapas) to Colombia  M T S L I

Callicore hydaspes (Drury, 1782)    iN  M T

Callicore lyca (E. Doubleday, [1847])  Six-spotted Eighty-eight    iN

Callicore lyca lyca (E. Doubleday, [1847])  S Mexico to Honduras  M T S

Callicore lyca aegina (C. Felder & R. Felder, 1861)  TL: Ecuador  M T S L

Callicore lyca aerias (Godman & Salvin, 1883)  Nicaragua to Panama  M T S1 S2 L I1 I2

Callicore lyca bella (Röber, 1924)  TL: Colombia  M T

Callicore lyca exultans (Fruhstorfer, 1916)  TL: Bolivia  M T

Callicore lyca mena (Staudinger, 1886)  TL: Peru  M T

Callicore lyca mionina (Hewitson, 1855)  TL: Colombia  M T

Callicore lyca odilia (Oberthür, 1916)  TL: Colombia  M T

Callicore lyca salamis (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Callicore lyca sticheli (Dillon, 1948)  TL: Honduras  M T

Callicore brome (Doyère, [1840])    iN

Callicore brome brome (Doyère, [1840])  TL: ?  M T

Callicore brome angusta Attal, 2000  Costa Rica  M T L I

Callicore brome danieli Attal & Crosson, 2003  Panama (Darien); ref.159  M T L

Callicore maronensis (Oberthür, 1916)    iN  M T

Callicore hesperis (Guérin-Méneville, [1844])    iN  M T

Callicore atacama (Hewitson, 1852)    iN

Callicore atacama folschveilleri (Attal, 2008)  Costa Rica & W Panama  M T

Callicore atacama manova (Fruhstorfer, 1916)  TL: Colombia  M T

Callicore atacama faustina (H. Bates, 1866)  Costa Rica (Atlantic slope Talamancas) & Panama  M T

Callicore atacama atacama (Hewitson, 1852)  TL: Ecuador  M T

Callicore felderi (Hewitson, 1864)    iN

Callicore felderi felderi (Hewitson, 1864)  TL: [Peru]  M T

Callicore felderi cajetani (Guenée, 1872)  TL: Peru  M T

Catagramma  Boisduval, 1836  

Catagramma eunomia Hewitson, 1853    iN

Catagramma eunomia eunomia Hewitson, 1853  ref.213  M T

Catagramma eunomia alani (Attal & Crosson, 2003)  ref.213  M T

Catagramma eunomia atahualpa (Attal, 2006)  ref.213

Catagramma eunomia ferrerorum Attal, 2000  ref.213

Catagramma eunomia incarnata Röber, 1915  ref.213  M T

Catagramma eunomia valeriae (Neild, 1996)  ref.213  M T

Catagramma hydarnis (Godart, [1824])  ref.213  iN  M T

Catagramma hystaspes (Fabricius, 1781)    iN

Catagramma hystaspes hystaspes (Fabricius, 1781)  ref.213  M T

Catagramma hystaspes discrepans Stichel, 1936  ref.213  M T

Catagramma hystaspes zelphanta Hewitson, 1858  ref.213  M T S

Catagramma pygas (Godart, [1824])    iN

Catagramma pygas pygas (Godart, [1824])  ref.213  M T

Catagramma pygas aphidna Hewitson, 1869  ref.213  M T

Catagramma pygas concolor Talbot, 1928  ref.213  M T

Catagramma pygas cyllene E. Doubleday, [1847]  ref.213  M T S

Catagramma pygas eucale Fruhstorfer, 1916  ref.213  M T

Catagramma pygas lalannensis Brévignon, 1995  ref.213

Catagramma pygas rondoni Ribeiro, 1931  ref.213  M T

Catagramma pygas thamyras Ménétriés, 1857  ref.213  M T

Catagramma tolima Hewitson, 1852    iN

Catagramma tolima tolima Hewitson, 1852  ref.213  M T

Catagramma tolima bugaba Staudinger, 1876  W Costa Rica & Panama; ref.213  M T S1 S2 L I

Catagramma tolima chimana Oberthür, 1916  ref.213  M T

Catagramma tolima aurantiaca Oberthür, 1916  TL: Colombia  M T

Catagramma tolima denina Hewitson, 1858  ref.213  M T

Catagramma tolima guatemalena H. Bates, 1866  Guatemala; ref.213  M T TI L

Catagramma tolima pacifica H. Bates, 1866  C & S Chiapas to W Guatemala; ref.213  M T S

Catagramma tolima peralta Dillon, 1948  Honduras to E Costa Rica & Panama; ref.213  M T S1 S2 L I

Catagramma tolima tehuana (R. G. Maza & J. Maza, 1983)  Veracruz, Tabasco, Oaxaca, N Chiapas (vic. Palenque); ref.213  M T S L

Catagramma pyracmon (Godart, [1824])    iN

Catagramma pyracmon pyracmon (Godart, [1824])  ref.213  M T

Catagramma pyracmon isaura J. Zikán, 1937  ref.213  M T

Catagramma pyracmon peristera Hewitson, 1853  ref.213  M T

Catagramma pyracmon piraya (Dillon, 1948)  ref.213  M T

Catagramma pyracmon pujoli Oberthür, 1916  ref.213  M T

Archaeogramma  Attal & Viloria, 2014  

Archaeogramma claritae Costa, 2014  ref.188

Diaethria  Billberg, 1820  

Diaethria anna (Guérin-Méneville, [1844])  Anna’s Eighty-eight    iN

Diaethria anna anna (Guérin-Méneville, [1844])  E Mexico to Costa Rica  M T S L1 L2 I

Diaethria anna mixteca J. Maza, 1977  SW Mexico  M T S Ab L

Diaethria anna salvadorensis (Franz & Schröder, 1954)  SW Chiapas to El Salvador  M T S

Diaethria astala (Guérin-Méneville, [1844])  Astala Eighty-eight    iN

Diaethria astala astala (Guérin-Méneville, [1844])  E Mexico to Colombia  M T S L1 L2

Diaethria astala asteroide R. G. Maza & R. F. Maza, 1985  SW Mexico  M T

Diaethria asteria (Godman & Salvin, 1894)  West-Mexican Eighty-eight  W Mexico  iN  M T S

Diaethria bacchis (E. Doubleday, 1849)  White-patched Eighty-eight  E & W Mexico & Guatemala  iN  M T S L

Diaethria candrena (Godart, [1824])    iN

Diaethria candrena candrena (Godart, [1824])  TL: Brazil  M T

Diaethria candrena longfieldae (Talbot, 1928)  TL: Brazil (MT)  M T

Diaethria ceryx (Hewitson, 1864)    iN  M T

Diaethria clymena (Cramer, 1775)    iN

Diaethria clymena clymena (Cramer, 1775)  TL: Guyana  M T

Diaethria clymena aurelia (Guenée, 1872)  TL: ?  M T

Diaethria clymena beleses (Godman & Salvin, 1889)  TL: Panama  M T

Diaethria clymena bourcieri (Guenée, 1872)  TL: Ecuador  M T

Diaethria clymena colombiana (Viette, 1958)  TL: Colombia  M T

Diaethria clymena consobrina (Guérin-Méneville, [1844])  TL: Colombia  M T

Diaethria clymena dodone (Guenée, 1872)  TL: Colombia  M T

Diaethria clymena janeira (C. Felder, 1862)  TL: Brazil (RJ)  M T

Diaethria clymena juani Neild, 1996  TL: Venezuela  M T

Diaethria clymena marchalii (Guérin-Méneville, [1844])  Nicaragua to Brazil  M T S L

Diaethria clymena meridionalis (H. Bates, 1864)  TL: Brazil (RS)  M T

Diaethria clymena peruviana (Guenée, 1872)  TL: Peru  M T

Diaethria clymena seropina (Röber, 1924)  TL: Brazil (Pa)

Diaethria eluina (Hewitson, [1855])    iN

Diaethria eluina eluina (Hewitson, [1855])  TL: [Brazil (MG)]  M T

Diaethria eluina lidwina (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Diaethria eluina splendidus (Oberthür, 1916)  TL: Brazil (Go)  M T

Diaethria euclides (Latreille, [1809])    iN

Diaethria euclides euclides (Latreille, [1809])  TL: Peru

Diaethria euclides artemis (Röber, 1915)  TL: Colombia  M T

Diaethria euclides gueneei (Röber, 1915)  TL: Ecuador  M T

Diaethria euclides metiscus (E. Doubleday, 1849)  TL: Venezuela  M T

Diaethria euclides phlogea (Salvin & Godman, 1868)  TL: Colombia  M T

Diaethria euclides phlogeides (Röber, 1915)  TL: Colombia  M T

Diaethria gabaza (Hewitson, [1855])    iN

Diaethria gabaza gabaza (Hewitson, [1855])  TL: Colombia  M T

Diaethria gabaza eupepla (Salvin & Godman, 1868)  Guatemala to Colombia  M T TI

Diaethria gabaza gabazina (Oberthür, 1916)  TL: Colombia  M T

Diaethria kolyma (Hewitson, 1852)    iN

Diaethria kolyma kolyma (Hewitson, 1852)  TL: Ecuador  M T

Diaethria kolyma pasithea (Hewitson, 1864)  TL: Ecuador

Diaethria neglecta (Salvin, 1869)    iN

Diaethria neglecta neglecta (Salvin, 1869)  TL: Bolivia; Peru; Ecuador; Colombia; "Guatemala"  M T S

Diaethria neglecta merida (Honrath, 1884)  TL: Venezuela  M T

Diaethria nystographa (Guenée, 1872)    iN

Diaethria nystographa nystographa (Guenée, 1872)  TL: Ecuador  M T

Diaethria nystographa aliciae Neild, 1996  TL: Venezuela  M T

Diaethria nystographa charis (Oberthür, 1916)  TL: Colombia  M T

Diaethria nystographa panthalis (Honrath, 1884)  TL: Venezuela  M T

Diaethria nystographa perezi Lamas, 1995  TL: Peru

Diaethria pandama (E. Doubleday, [1848])  Orange-striped Eighty-eight  E & W Mexico to Panama  iN  M T S L

Perisama  E. Doubleday, 1849  

Perisama barnesi Schaus, 1913  Costa Rican Eighty-eight  Costa Rica (Carrillo Belt, Atlantic slope, to Talamancas); ref.12,213  M T S

Perisama vaninka (Hewitson, [1855])  ref.12  iN

Perisama vaninka vaninka (Hewitson, [1855])  ref.12,213

Perisama vaninka delafuentei (Neild, 1996)  ref.12

Perisama vaninka doris (C. Felder & R. Felder, 1861)  ref.12,213  M S

Perisama vaninka gonalia Fruhstorfer, 1916  ref.12,213

Perisama vaninka volara (Hewitson, 1868)  ref.12,213

Perisama vaninka [n. ssp.] Lamas, MS  Colombia

Perisama cardases (Hewitson, 1869)  ref.12,375,213  iN  M T

Perisama campaspe (Hewitson, 1869)  ref.12,375,213  iN  M T

Perisama diotima (Hewitson, 1852)  ref.12,375,213  iN  M T S

Perisama cecidas (Hewitson, 1869)  ref.12,375,213  iN  M T

Perisama alicia (Hewitson, 1868)    iN  M T S

Perisama antioquia Attal & Crosson, 1996    iN  M T

Perisama bomplandii (Guérin-Méneville, [1844])    iN

Perisama bomplandii bomplandii (Guérin-Méneville, [1844])  TL: Colombia  M T S

Perisama bomplandii albipennis A. Butler, 1873  TL: Peru  M T

Perisama bomplandii equatorialis (Guenée, 1872)  TL: Ecuador  M T

Perisama bomplandii parabomplandii Dognin, 1899  TL: Colombia  M T

Perisama bomplandii reyi Attal & Crosson, 2003  ref.159

Perisama bomplandii velasteguii Lamas, 1995  TL: Ecuador  M T

Perisama cabirnia (Hewitson, 1874)    iN  M T

Perisama calamis (Hewitson, 1869)    iN  M T S

Perisama camelita (Hewitson, [1876])    iN  M T S

Perisama canoma H. Druce, 1874    iN

Perisama canoma canoma H. Druce, 1874  TL: Peru  M T S

Perisama canoma cotyora (Hewitson, 1874)  TL: Bolivia  M T

Perisama canoma cyaneis Attal & Crosson, 1997  TL: Ecuador  M T

Perisama canoma lamasi Descimon & Mast, 1995  TL: Peru  M S

Perisama canoma magaly Attal & Crosson, 1999  TL: Ecuador  M T

Perisama canoma thryoessa Fruhstorfer, 1916  TL: Peru  M T

Perisama clisithera (Hewitson, 1874)    iN  M T S

Perisama cloelia (Hewitson, 1868)  TL: Ecuador  M T

Perisama comnena (Hewitson, 1868)    iN

Perisama comnena comnena (Hewitson, 1868)  TL: Bolivia  M T S

Perisama comnena attali Neild, 1996  TL: Venezuela  M T

Perisama comnena bertha Oberthür, 1916  TL: Colombia  M T

Perisama comnena bolivar Attal & Crosson, 1996  TL: Venezuela  M T

Perisama comnena comnenoides Oberthür, 1916  TL: Peru  M T

Perisama comnena divergens A. Butler, 1873  TL: Ecuador  M T

Perisama comnena ismaeli Attal & Crosson, 1996  TL: Ecuador

Perisama dorbignyi (Guérin-Méneville, [1844])    iN

Perisama dorbignyi dorbignyi (Guérin-Méneville, [1844])  TL: Colombia  M T S

Perisama dorbignyi aldasi Attal & Crosson, 1996  TL: Ecuador  M T

Perisama dorbignyi aristoteles Oberthür, 1916  TL: Ecuador  M T

Perisama dorbignyi jurinei (Guenée, 1872)  TL: Peru  M T S

Perisama dorbignyi klugi Descimon & Mast, 1995  TL: Peru  M S

Perisama dorbignyi mola Dognin, 1899  TL: Ecuador  M T

Perisama dorbignyi paula Oberthür, 1916  TL: Colombia  M T

Perisama euriclea (E. Doubleday, [1847])  TL: Venezuela  M T

Perisama gisco Godman & Salvin, 1880    iN  M T L

Perisama goeringi H. Druce, 1875  TL: Venezuela  M T

Perisama hazarma (Hewitson, 1877)  TL: Ecuador  M T S

Perisama humboldtii (Guérin-Méneville, [1844])    iN

Perisama humboldtii humboldtii (Guérin-Méneville, [1844])  TL: Colombia  M T

Perisama humboldtii intermedia Oberthür, 1916  TL: Ecuador  M T

Perisama humboldtii tringa (Guenée, 1872)  TL: Peru  M T S

Perisama humboldtii tristis Oberthür, 1916  TL: Venezuela  M T

Perisama humboldtii [n. ssp.] Lamas, MS  Colombia

Perisama lanice (Hewitson, 1868)  TL: Ecuador

Perisama lanice lanice (Hewitson, 1868)  TL: Ecuador  M T

Perisama lanice latimargo Oberthür, 1916  TL: Peru  M T

Perisama lanice picteti (Guenée, 1872)  TL: Peru  M T

Perisama lebasii (Guérin-Méneville, [1844])    iN

Perisama lebasii lebasii (Guérin-Méneville, [1844])  TL: Colombia  M T

Perisama lebasii ambatensis Oberthür, 1916  TL: Ecuador  M T

Perisama lebasii astuta Dognin, 1899  TL: Colombia  M T

Perisama lebasii hilara Salvin, 1869  TL: Peru  M T S

Perisama lebasii koenigi Descimon & Mast, 1995  TL: Peru  M T

Perisama lebasii selva Attal & Crosson, 1997  TL: Ecuador  M T

Perisama morona (Hewitson, 1868)    iN

Perisama morona morona (Hewitson, 1868)  TL: Bolivia  M T

Perisama morona moronina Röber, 1915  TL: Peru  M T

Perisama nevada Attal & Crosson, 1996    iN  M T Ma

Perisama oppelii (Latreille, [1809])    iN

Perisama oppelii oppelii (Latreille, [1809])  TL: Peru  M T S

Perisama oppelii amalia Oberthür, 1916  TL: Colombia

Perisama oppelii angustifasciata Röber, 1924  TL: Colombia

Perisama oppelii bleuzeni Attal & Crosson, 1996  TL: Venezuela  M T

Perisama oppelii cristal Attal & Crosson, 1996  TL: Ecuador  M T S

Perisama oppelii eminens Oberthür, 1881  TL: Peru  M T

Perisama oppelii erebina Oberthür, 1916  TL: Colombia  M T

Perisama oppelii iphigenia Oberthür, 1916  TL: Bolivia  M T

Perisama oppelii viridinota A. Butler, 1873  TL: Peru  M T

Perisama oppelii xanthica (Hewitson, 1868)  TL: Peru  M T

Perisama ouma Dognin, 1891    iN  M T S

Perisama paralicia Fruhstorfer, 1916    iN

Perisama paralicia paralicia Fruhstorfer, 1916  TL: Peru  M T S

Perisama paralicia bonita Attal & Crosson, 1997  TL: Ecuador  M T S

Perisama patara (Hewitson, [1855])  TL: Colombia

Perisama patara patara (Hewitson, [1855])  TL: Colombia

Perisama patara arletta Descimon & Mast, 1995  TL: Peru  M S

Perisama patara gallardi Attal & Crosson, 1996  TL: Colombia  M T

Perisama philinus E. Doubleday, 1849    iN

Perisama philinus philinus E. Doubleday, 1849  TL: Bolivia  M T S

Perisama philinus descimoni Mast, 1995  TL: Peru  M S

Perisama philinus nyctimene (Hewitson, 1868)  TL: Ecuador  M T S

Perisama philinus saussurei (Guenée, 1872)  TL: Brazil

Perisama satanas Attal & Crosson, 1996  TL: Colombia  M T

Perisama tristrigosa A. Butler, 1873    iN  M T S

Perisama tryphena (Hewitson, [1857])    iN

Perisama tryphena tryphena (Hewitson, [1857])  TL: Colombia  M T S

Perisama tryphena affinis Crosson & Attal, 1995  TL: Peru  M T

Perisama tryphena asormani Attal & Crosson, 1996  TL: Venezuela  M T

Perisama tryphena cauca Attal & Crosson, 1996  TL: Colombia  M T

Perisama tryphena fernandezi Attal & Crosson, 1996  TL: Venezuela

Perisama tryphena pallida Attal & Crosson, 1996  TL: Ecuador  M T

Perisama tryphena pastaza Attal & Crosson, 1996  TL: Ecuador

Perisama typhania Oberthür, 1916    iN  M T

Perisama vitringa (Hewitson, 1858)    iN

Perisama vitringa vitringa (Hewitson, 1858)  TL: [Peru]  M T

Perisama vitringa huallaga Attal & Crosson, 1996  TL: Peru  M T S

Perisama vitringa ochreipennis A. Butler, 1873  TL: Bolivia  M T S

Perisama yeba (Hewitson, [1857])    iN

Perisama yeba yeba (Hewitson, [1857])  TL: Colombia  M T

Perisama yeba emma Oberthür, 1916  TL: Venezuela  M T

Perisama yeba guerini C. Felder & R. Felder, 1867  TL: Colombia  M T

Perisama yeba johannes Oberthür, 1916  TL: Ecuador  M T

Perisama yeba malvina Kretzschmar, [1894]  TL: Colombia  M T

Perisama yeba nimbala Attal & Crosson, 1996  TL: Ecuador  M T

Perisama [n. sp.] Lamas, MS  Peru

  Subfamily Cyrestinae

  Tribe Cyrestini

Marpesia  Hübner, 1818  

Marpesia berania (Hewitson, 1852)    iN

Marpesia berania berania (Hewitson, 1852)  TL: Ecuador; Brazil (Am)  M T S

Marpesia berania fruhstorferi (Seitz, 1914)  Honduras to Panama  M T S L

Marpesia chiron (Fabricius, 1775)  Many-banded Daggerwing  S TX (strays N), E & W Mexico to Argentina, Greater Antilles  iN

Marpesia chiron chiron (Fabricius, 1775)  TL: [Haiti]  M T S HS L1 L2

Marpesia chiron chironides (Staudinger, 1886)  TL: Cuba  M T

Marpesia chiron marius (Cramer, 1779)  TL: Surinam

Marpesia corinna (Latreille, [1813])    iN  M T S

Marpesia corita (Westwood, 1850)  Orange-banded Daggerwing    iN

Marpesia corita corita (Westwood, 1850)  E & S Mexico, Guatemala, Honduras  M T S L Ha

Marpesia corita phiale (Godman & Salvin, 1878)  SW Mexico (S Chiapas) & W Guatemala  M T S L

Marpesia crethon (Fabricius, 1776)    iN  M T S

Marpesia egina (H. Bates, 1865)    iN  M T

Marpesia eleuchea Hübner, 1818  Caribbean Daggerwing    iN

Marpesia eleuchea eleuchea Hübner, 1818  Cuba & Isla de Juventud, rare stray to S FL Keys  M S

Marpesia eleuchea bahamensis Munroe, 1971  Bahamas  M T S

Marpesia eleuchea dospassosi Munroe, 1971  Hispaniola  M T S

Marpesia eleuchea pellenis (Godart, [1824])  Jamaica  M S L

Marpesia furcula (Fabricius, 1793)    iN

Marpesia furcula Northern segregate ,  [= "iole" records for Central America], Nicaragua to Panama  M L

Marpesia furcula furcula (Fabricius, 1793)  TL: Jamaica; India

Marpesia furcula violetta (A. Hall, 1929)  TL: Ecuador  M T

Marpesia furcula oechalia (Westwood, 1850)  TL: Bolivia  M T

Marpesia furcula [n. ssp.] Lamas, MS  Panama

Marpesia harmonia (Klug, 1836)  Pale Daggerwing  SE Mexico & Guatemala  iN  M S L

Marpesia livius (W. F. Kirby, 1871)    iN

Marpesia livius livius (W. F. Kirby, 1871)  TL: Ecuador  M T S

Marpesia livius alcibiades (Staudinger, 1876)  Costa Rica & Panama  M T S

Marpesia marcella (C. Felder & R. Felder, 1861)    iN

Marpesia marcella marcella (C. Felder & R. Felder, 1861)  TL: Colombia; "Ecuador"  M T S

Marpesia marcella valetta (A. Butler & H. Druce, 1872)  Guatemala to Costa Rica  M T S

Marpesia marcella [n. ssp.] Lamas, MS  Panama

Marpesia merops (Doyère, [1840])  Spot-banded Daggerwing  Guatemala to Brazil  iN  M T S L1 L2

Marpesia orsilochus (Fabricius, 1776)    iN  M S

Marpesia petreus (Cramer, 1776)  Ruddy Daggerwing    iN

Marpesia petreus Northern segregate ,  S FL, S TX, E & W Mexico (strays far N) to W Ecuador  M S1 S2 L1 L2 L3 I

Marpesia petreus petreus (Cramer, 1776)  Guianas, Amazonian Colombia to Bolivia  M T S L

Marpesia petreus damicorum Brévignon, 2001  Mona, Puerto Rico, Lesser Antilles (St. Kitts, Guadeloupe, etc.)  M S L

Marpesia petreus rheophila Lamas, 1995  TL: Peru

Marpesia petreus [n. ssp.] Lamas, MS  Mexico

Marpesia themistocles (Fabricius, 1793)    iN

Marpesia themistocles themistocles (Fabricius, 1793)  TL: ?

Marpesia themistocles norica (Hewitson, 1852)  TL: [Brazil (Am)]  M T S

Marpesia tutelina (Hewitson, 1852)    iN  M T

Marpesia zerynthia Hübner, [1823]    iN

Marpesia zerynthia zerynthia Hübner, [1823]  TL: ?  M T

Marpesia zerynthia dentigera (Fruhstorfer, 1907)  [= coresia], E & S Mexico to Ecuador, one-time stray to S TX  M T S L1 L2

Marpesia pantepuiana Costa, Attal & Viloria, 2016  ref.189

  Subfamily Nymphalinae

  Tribe Coeini

Pycina  E. Doubleday, [1849]  

Pycina zamba E. Doubleday, [1849]    iN

Pycina zamba zamba E. Doubleday, [1849]  TL: Venezuela  M T S

Pycina zamba zelys Godman & Salvin, 1884  S Mexico to Panama  M T S

Historis  Hübner, [1819]  

Historis acheronta (Fabricius, 1775)  Tailed Cecropian    iN

Historis acheronta acheronta (Fabricius, 1775)  continental; S TX & AZ (as stray), E & W Mexico to Brazil  M T S L1 L2 I

Historis acheronta cadmus (Cramer, 1775)  Jamaica  M OI

Historis acheronta semele (M. Bates, 1939)  Cuba and Hispaniola  M S

Historis odius (Fabricius, 1775)  Orion Cecropian    iN

Historis odius odius (Fabricius, 1775)  Greater Antilles  M T S L

Historis odius caloucaera Brévignon, 2003  Windward and Leeward Islands  M OI

Historis odius dious Lamas, 1995  continental; S TX (as stray), E & W Mexico to Argentina  M T S L

Baeotus  Hemming, 1939  

Baeotus aeilus (Stoll, 1780)    iN  M T S

Baeotus beotus (E. Doubleday, [1849])  Graphic Beauty  SE Mexico to Amazon Basin  iN  M T S1 S2 L I

Baeotus deucalion (C. Felder & R. Felder, 1860)    iN  M T S

Baeotus japetus (Staudinger, [1885])    iN  M T S

  Tribe Nymphalini

Colobura  Billberg, 1820  

Colobura annulata Willmott, Constantino & J. Hall, 2001  New Beauty  S Mexico to W Ecuador, Venezuela to Bolivia & Guianas, Trinidad  iN  M T S L I

Colobura dirce (Linnaeus, 1758)  Dirce Beauty    iN

Colobura dirce dirce (Linnaeus, 1758)  E & W Mexico to Argentina  M T S HS L I

Colobura dirce wolcotti W. Comstock, 1942  Greater Antilles (Jamaica, Cuba, Hispaniola, Puerto Rico)  M T S L

Tigridia  Hübner, [1819]  

Tigridia acesta (Linnaeus, 1758)    iN

Tigridia acesta Northern segregate ,  SE Mexico to Panama  M S L I

Tigridia acesta acesta (Linnaeus, 1758)  TL: Indiis

Tigridia acesta columbina (Neustetter, 1929)  TL: Colombia  M T

Tigridia acesta fulvescens (A. Butler, 1873)  TL: Peru  M T

Tigridia acesta latifascia (A. Butler, 1873)  TL: Colombia  M T

Tigridia acesta ochracea (Bryk, 1953)  TL: Peru

Tigridia acesta tapajona (A. Butler, 1873)  TL: Brazil (Pa)  M T

Tigridia acesta [n. ssp.#1] Lamas, MS  TL: Mexico (Chia)

Tigridia acesta [n. ssp.#2] Lamas, MS  TL: Venezuela

Tigridia acesta [n. ssp.#3] Lamas, MS  TL: Bolivia

Smyrna  Hübner, [1823]  

Smyrna blomfildia (Fabricius, 1781)    iN

Smyrna blomfildia blomfildia (Fabricius, 1781)  TL: Brazil  M T

Smyrna blomfildia datis Fruhstorfer, 1908  S TX, E & W Mexico to Panama  M T S L I

Smyrna karwinskii Geyer, [1833]  Karwinski’s Beauty  E & W Mexico to Nicaragua  iN  M S L

Hypanartia  Hübner, [1821]  

Hypanartia bella (Fabricius, 1793)    iN  M T

Hypanartia celestia Lamas, Willmott & J. Hall, 2001  TL: Peru  M T S

Hypanartia charon (Hewitson, 1878)  TL: Ecuador  M T L

Hypanartia christophori Jasinski, 1998    iN  M S

Hypanartia cinderella Lamas, Willmott & J. Hall, 2001    iN  M T L

Hypanartia dione (Latreille, [1813])    iN

Hypanartia dione dione (Latreille, [1813])  TL: [Colombia]  M T S

Hypanartia dione arcaei (Salvin, 1871)  Costa Rica & Panama  M T S L

Hypanartia dione disjuncta Willmott, J. Hall & Lamas, 2001  E & W Mexico, Guatemala, Honduras  M T S L

Hypanartia fassli Willmott, J. Hall & Lamas, 2001  TL: Colombia  M T

Hypanartia godmanii (H. Bates, 1864)  Godman's Mapwing  E & W Mexico to Colombia  iN  M T S L

Hypanartia kefersteini (E. Doubleday, [1847])    iN  M T

Hypanartia lethe (Fabricius, 1793)  Orange Mapwing  S TX (rare stray), E & S Mexico to Brazil  iN

Hypanartia lethe lethe (Fabricius, 1793)  TL: Indiis  M S L

Hypanartia lethe rosamariae Neild, 2008  ref.266

Hypanartia lindigii (C. Felder & R. Felder, 1862)    iN  M T L

Hypanartia paullus (Fabricius, 1793)  Antillean Mapwing  Greater Antilles (Jamaica, Cuba, Hispaniola, Puerto Rico)  iN  M T S L

Hypanartia splendida Rothschild, 1903  TL: Peru  M T

Hypanartia trimaculata Willmott, J. Hall & Lamas, 2001    iN

Hypanartia trimaculata trimaculata Willmott, J. Hall & Lamas, 2001  TL: Colombia  M T

Hypanartia trimaculata autumna Willmott, J. Hall & Lamas, 2001  [Mexican "kefersteini" records], E & S Mexico to Panama  M T S L

Hypanartia trimaculata fountaineae Neild, 2008  ref.266

Vanessa  Fabricius, 1807  

Vanessa altissima (Rosenberg & Talbot, 1914)    iN  M T S

Vanessa annabella (W. D. Field, 1971)  West Coast Lady    iN  M T S Ab L1 L2 I Ha

Vanessa atalanta (Linnaeus, 1758)  Red Admiral    iN

Vanessa atalanta rubria (Fruhstorfer, 1909)  Gr. Antilles (not PR), N Bahamas (strays), N Am. to Guat., Venez.  M T S Ab L1 L2 L3 I H

Vanessa braziliensis (Moore, 1883)    iN  M T

Vanessa cardui (Linnaeus, 1758)  Painted Lady  nearly cosmopolitan as vagrant; uncommon or vagrant in W. Indies  iN  M T S L1 L2 L3 I

Vanessa carye (Hübner, [1812])    iN  M T

Vanessa myrinna (E. Doubleday, 1849)    iN  M T HS

Vanessa tameamea Escholtz, 1821  Kameamea  Endemic to Hawaii  iN  M S L Ha

Vanessa terpsichore (Philippi, 1859)    iN  M S L

Vanessa virginiensis (Drury, 1773)  American Lady  N Bahamas, Gr. Antilles, Hawaii, Canary Is.; N. Am. to Colombia  iN  M T S Ab L1 L2 L3 L4 I H

Vanessa amerindica J. Miller & F. Brown, 1989  a fossil from Colorado

Aglais  Dalman, 1816  

Aglais io (Linnaeus, 1758)  TL: Sweden

Aglais milberti (Godart, 1819)  Milbert's Tortoiseshell    iN

Aglais milberti viola dos Passos, 1938  TL: Canada (NL)  M S

Aglais milberti milberti (Godart, 1819)  TL: USA (Pa)  M S1 S2 L

Aglais milberti subpallida (T. Cockerell, 1889)  TL: USA (Co)  M S1 S2 L I Ha

Aglais milberti pullum (Austin, 1998)  TL: USA (Az)  M T S L

Nymphalis  Kluk, 1780  

Nymphalis l-album (Esper, 1781)  Compton Tortoiseshell    iN

Nymphalis l-album j-album (Boisduval & Le Conte, [1835])  TL: USA (In)  M S L

Nymphalis l-album watsoni (G. Hall, 1924)  TL: Canada (BC)  M S

Nymphalis californica (Boisduval, 1852)  California Tortoiseshell    iN  M T S1 S2 L I Ha1 Ha2

Nymphalis antiopa (Linnaeus, 1758)  Mourning Cloak    iN

Nymphalis antiopa antiopa (Linnaeus, 1758)  TL: Sweden; America  M T S1 S2 S3 Ab L1 L2 L3 I H Txt

Nymphalis cyanomelas (E. Doubleday, [1848])  Mexican Tortoiseshell  S Mexico to El Salvador  M T S

Polygonia  Hübner, [1819]  

Polygonia interrogationis (Fabricius, 1798)  Question Mark    iN  M T S1 S2 L1 L2 L3 L4 I1 I2 H F

Polygonia comma (T. Harris, 1841)  Comma Anglewing    iN  M S1 S2 L1 L2

Polygonia g-argenteum (E. Doubleday, 1848)  Mexican Anglewing  Mexico  iN  M T S L

Polygonia satyrus (W. H. Edwards, 1869)  Satyr Anglewing    iN

Polygonia satyrus satyrus (W. H. Edwards, 1869)  TL: USA (Co)  M T S1 S2 S3 L1 L2 I H

Polygonia satyrus neomarsyas dos Passos, 1969  [= transcanada]  M T S1 S2 L Ha

Polygonia progne (Cramer, 1775)  Gray Anglewing    iN  M T S1 S2 L

Polygonia oreas (W. H. Edwards, 1869)  Oreas Anglewing    iN

Polygonia oreas oreas (W. H. Edwards, 1869)  TL: USA (Ca)  M S1 S2

Polygonia oreas silenus (W. H. Edwards, 1870)  TL: USA (Or)  M S L Ha

Polygonia oreas threatfuli C. Guppy & J. Shepard, 2001  TL: Canada (BC)  M S1 S2

Polygonia oreas satellow Scott, 2006  TL: USA (Co)  M S

Polygonia oreas nigrozephyrus Scott, 1984  TL: USA (Co)  M S

Polygonia gracilis (Grote & Robinson, 1867)  Hoary Anglewing    iN

Polygonia gracilis gracilis (Grote & Robinson, 1867)  TL: USA (NH)  M S1 S2 L

Polygonia gracilis zephyrus (W. H. Edwards, 1870)  TL: USA (Nv)  M T S1 S2 L1 L2 L3 L4 I1 I2 H Ha

Polygonia haroldii (Dewitz, 1877)  Spotless Anglewing  Mexico  iN  M T S1 S2 L

Polygonia faunus (W. H. Edwards, 1862)  Green Anglewing    iN

Polygonia faunus smythi A. Clark, 1937  TL: USA (Va)  M S1 S2

Polygonia faunus faunus (W. H. Edwards, 1862)  TL: USA (NY)  M S1 S2 L

Polygonia faunus arcticus Leussler, 1935  [= cenveray]  M T S1 S2 L

Polygonia faunus rusticus (W. H. Edwards, 1874)  TL: USA (Ca)  M S L

Polygonia faunus fulvescens J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S1 S2

Polygonia faunus hylas (W. H. Edwards, 1872)  TL: USA (Co)  M S1 S2 L

Jupitellia  , []  a fossil genus

Jupitellia charon (Scudder, 1889)  Fossil from Colorado

  Tribe Victorinini

Anartia  Hübner, [1819]  

Anartia amathea (Linnaeus, 1758)  Red Peacock    iN

Anartia amathea amathea (Linnaeus, 1758)  Panama to Argentina; Grenada, Barbados, Antigua  M S Hy HS L

Anartia amathea roeselia (Eschscholtz, 1821)  TL: Brazil  M T

Anartia amathea sticheli Fruhstorfer, 1907  TL: Bolivia  M T

Anartia amathea [n. ssp.] Lamas, MS  Lesser Antilles

Anartia chrysopelea Hübner, [1831]  Cuban Peacock  Cuba & Isla de Juventud, rare stray to S FL  iN  M T S

Anartia fatima (Fabricius, 1793)  Banded Peacock    iN

Anartia fatima fatima (Fabricius, 1793)  S TX, E & S Mexico to far NW Colombia  M T S Hy L1 L2 I H

Anartia fatima colima Lamas, 1995  NW Mexico  M T S L

Anartia jatrophae (Linnaeus, 1763)  White Peacock    iN

Anartia jatrophae jatrophae (Linnaeus, 1763)  South America, Lesser Antilles N to Barbuda & St. Kitts  M T S1 S2 HS L

Anartia jatrophae guantanamo Munroe, 1942  S FL (strays N), Bahamas, Cuba, Turks & Caicos  M T S L1 L2

Anartia jatrophae intermedia Munroe, 1942  St. Croix  M T

Anartia jatrophae jamaicensis Möschler, 1886  Jamaica, Grand Cayman, [Cayman Brac?]  M S1 S2 L

Anartia jatrophae luteipicta Fruhstorfer, 1907  S TX (strays far N), E & W Mexico through Central America  M T S Ab Ma L1 L2 L3 L4 I H

Anartia jatrophae saturata Staudinger, [1885]  Hispaniola, Great Inagua  M T S

Anartia jatrophae semifusca Munroe, 1942  Puerto Rico, Mona, E to Virgin Islands  M T S L

Anartia lytrea (Godart, 1819)  Hispaniolan Peacock  Hispaniola (Dominican Republic); rare stray to Cuba  iN  M T S

Metamorpha  Hübner, [1819]  

Metamorpha elissa Hübner, [1819]  Metamorpha    iN

Metamorpha elissa elissa Hübner, [1819]  Panama to Guianas and Peru  M S L

Metamorpha elissa alberti Neild, 2008  ref.266

Metamorpha elissa pulsitia R. Fox & A. Forbes, 1971  TL: Bolivia  M T

Napeocles  H. Bates, 1864  

Napeocles jucunda (Hübner, [1808])    iN

Napeocles jucunda jucunda (Hübner, [1808])  TL: ?  M T

Napeocles jucunda caesari Neild, 2008  ref.266

Siproeta  Hübner, [1823]  

Siproeta epaphus (Latreille, [1813])  Rusty-tipped Page    iN

Siproeta epaphus epaphus (Latreille, [1813])  S TX, S NM (rare strays), E & W Mexico to Peru  M T S L1 L2 I H

Siproeta epaphus gadoui Masters, 1967  TL: Venezuela  M T

Siproeta epaphus trayja Hübner, [1823]  TL: Brazil  M L

Siproeta stelenes (Linnaeus, 1758)  Malachite    iN

Siproeta stelenes stelenes (Linnaeus, 1758)  Jamaica, Hispaniola, Puerto Rico, St. Kitts, St. Croix  M S1 S2 HS L

Siproeta stelenes biplagiata (Fruhstorfer, 1907)  S TX (strays N), E & W Mexico to Amazon, S FL, Cuba, Caymans  M T S L1 L2 I H

Siproeta stelenes meridionalis (Fruhstorfer, 1909)  TL: Brazil (RS)  M T S L

Siproeta stelenes sophene (Fruhstorfer, 1907)  TL: Ecuador  M T

Siproeta superba (H. Bates, 1864)  Broad-banded Page    iN

Siproeta superba superba (H. Bates, 1864)  S Mexico, Guatemala, Honduras  M T S L

Siproeta superba euoe R. Fox & A. Forbes, 1971  Costa Rica  M T S

  Tribe Junoniini

Hypolimnas  Hübner, [1819]  

Hypolimnas misippus (Linnaeus, 1764)  Mimic  NE S. Am, Gr. & Lesser Antilles, N continental and Yucatan coasts (as st ...  iN  M T S HS

Junonia  Hübner, [1819]  

Junonia coenia Hübner, [1822]  Northern Buckeye  E USA (strays N), Bermuda, Cuba & Isla de Juventud  iN  M T S1 S2 Ab L1 L2 L3 L4 L5 L6 L7 I H

Junonia grisea Austin & J. Emmel, 1998  Northern Buckeye  W USA (strays N & NE), Baja California to SW Oregon to W TX  iN  M T S L1 L2 Ha

Junonia nigrosuffusa W. Barnes & McDunnough, 1916  Dark Buckeye  SW USA & W TX to SW Mexico (strays N)  iN  M T S L

Junonia stemosa Grishin, 2020  Twintip buckeye  C & S TX (strays N); ref.6  iN  M S1 S2 L1 L2

Junonia zonalis C. Felder & R. Felder, 1867  Tropical Buckeye    iN

Junonia zonalis zonalis C. Felder & R. Felder, 1867  S FL, Bahamas, Cuba, Cayman Is., Jamaica, Hispaniola; C. Am.?  M S IL L

Junonia zonalis michaelisi Fruhstorfer, 1907  Puerto Rico  M T L

Junonia zonalis swifti Brévignon, 2004  Guadeloupe and nearby islands; possibly other I. in Less. Antilles  M S

Junonia pacoma Grishin, 2020  Pacific mangrove buckeye  Son to Jal & BCS, coastal; ref.6,16  iN  M S L

Junonia neildi Brévignon, 2004  Mangrove Buckeye  Guadeloupe; possibly elsewhere in Lesser Antilles  iN

Junonia neildi varia Grishin, 2020  Gulf Coast mangroves, S TX to Belize (at least), ? along Pacific; ref.6  M S L

Junonia neildi Caribbean segregate ,  S FL, Bahamas, Greater Antilles, Cayman Islands  M S1 S2 L

Junonia neildi neildi Brévignon, 2004  Guadeloupe; possibly elsewhere in Lesser Antilles  M S

Junonia litoralis Brévignon, 2009  Litoral Buckeye  French Guiana & Trinidad  iN

Junonia wahlbergi Brévignon, 2008  Wahlberg's Buckeye  French Guiana & Trinidad

Junonia evarete (Cramer, 1779)  Tropical Buckeye    iN

Junonia evarete evarete (Cramer, 1779)  NE South America (at least)  M T OI L

Junonia evarete flirtea (Fabricius, 1793)  TL: India

Junonia evarete fuscescens (A. Butler, 1901)  TL: Ecuador  M L

Junonia evarete huacapistana W. Forbes, [1929]  TL: Peru

Junonia evarete lima W. Forbes, [1929]  TL: Peru

Junonia evarete oscura Neild, 2008  ref.266  M T

Junonia evarete occidentalis C. Felder & R. Felder, 1862  TL: Brazil (Am)

Junonia evarete [n. ssp.] Lamas, MS  Mexico (Oax)

Junonia divaricata C. Felder & R. Felder, 1867    iN

Junonia genoveva (Cramer, 1780)  Mangrove Buckeye    iN

Junonia genoveva genoveva (Cramer, 1780)  NE South America, possibly Central America  M T OI

Junonia genoveva constricta C. Felder & R. Felder, 1867  TL: Colombia  M T L

Junonia genoveva hilaris C. Felder & R. Felder, 1867  TL: Paraguay

Junonia genoveva incarnata C. Felder & R. Felder, 1867  TL: Colombia  M T

Junonia genoveva infuscata C. Felder & R. Felder, 1867  TL: Brazil (Ba)  M T

Junonia genoveva vivida W. Forbes, [1929]  TL: Guyana; Surinam  M T

Junonia genoveva [n. ssp.#1] Lamas, MS  TL: Mexico (Oax)

Junonia genoveva [n. ssp.#2] Lamas, MS  TL: Peru

Junonia vestina C. Felder & R. Felder, 1867    iN

Junonia vestina vestina C. Felder & R. Felder, 1867  TL: ?

Junonia vestina livia Fruhstorfer, 1912  TL: Bolivia  M T

  Tribe Melitaeini

Euphydryas  Scudder, 1872  

Euphydryas gillettii (W. Barnes, 1897)  Gillett's Checkerspot    iN  M S1 S2 L I Ha

Euphydryas editha (Boisduval, 1852)  Edith's Checkerspot    iN

Euphydryas editha taylori (W. H. Edwards, 1888)  TL: Canada (BC)  M S1 S2 S3 S4 Ab L

Euphydryas editha editha (Boisduval, 1852)  TL: USA (Ca)  M T S1 S2 L

Euphydryas editha insularis T. Emmel & J. Emmel, 1974  TL: USA (Ca)  M T

Euphydryas editha augustina (W. G. Wright, 1905)  TL: USA (Ca)  M T S1 S2 L I

Euphydryas editha quino (Behr, 1863)  TL: USA (Ca)  M T S1 S2 L I H Ha

Euphydryas editha ehrlichi Baughman & Murphy, 1998  TL: USA (Ca)  M S

Euphydryas editha nubigena (Behr, 1863)  TL: USA (Ca)  M T S L Ha

Euphydryas editha tahoensis Austin & Murphy, 1998  TL: USA (Nv)  M S L

Euphydryas editha aurilacus Gunder, 1928  TL: USA (Ca)  M S L

Euphydryas editha lawrencei Gunder, 1931  TL: USA (Or)  M T S1 S2 L Ha

Euphydryas editha colonia (W. G. Wright, 1905)  TL: USA (Or)  M T S L

Euphydryas editha rubicunda (H. Edwards, 1881)  TL: USA (Ca)  M S1 S2 L H

Euphydryas editha baroni (W. H. Edwards, 1879)  TL: USA (Ca)  M S Ab L I

Euphydryas editha karinae Baughman & Murphy, 1998  TL: USA (Ca)  M S

Euphydryas editha luestherae Murphy & P. Ehrlich, 1980  TL: USA (Ca)  M T S I

Euphydryas editha monoensis Gunder, 1928  TL: USA (Ca)  M S Ab

Euphydryas editha edithana (Strand, 1915)  TL: USA (Ca)  M T S L Ha

Euphydryas editha C Oregon segregate ,  Pine Mtn. area, Deschutes Co., OR  M S L Ha

Euphydryas editha Ochoco Mts. segregate ,  vic. Big Summit Prairie, Ochoco Mts., OR  M S

Euphydryas editha NE Oregon segregate ,  Blue Mountains, NE Oregon  M S L

Euphydryas editha bingi Baughman & Murphy, 1998  S end Warner Mts., CA, high elevation  M S L

Euphydryas editha mattooni Austin & Murphy, 1998  TL: USA (Ca)  M T1 T2 T3

Euphydryas editha owyheensis Austin & Murphy, 1998  TL: USA (Nv)  M T1 T2 Ha

Euphydryas editha lehmani Gunder, 1929  TL: USA (Nv)  M T S Ha

Euphydryas editha gunnisonensis F. Brown, [1971]  TL: USA (Co)  M S1 S2

Euphydryas editha alebarki Ferris, [1971]  TL: USA (Wy)  M T S1 S2 S3 L Ha

Euphydryas editha montanus McDunnough, 1928  TL: USA (Wy)  M S L

Euphydryas editha hutchinsi McDunnough, 1928  TL: USA (Mt)  M S

Euphydryas editha beani (Skinner, 1897)  TL: Canada (Ab)  M S

Euphydryas chalcedona (E. Doubleday, [1847])  Chalcedon Checkerspot    iN

Euphydryas chalcedona klotsi dos Passos, 1938  TL: USA (Az)  M S L1 L2 I

Euphydryas chalcedona kingstonensis T. Emmel & J. Emmel, 1973  TL: USA (Ca)  M T S1 S2 L

Euphydryas chalcedona corralensis T. Emmel & J. Emmel, 1973  TL: USA (Ca)  M T S1 S2 L H Ha

Euphydryas chalcedona hennei Scott, 1981  TL: USA (Ca)  M T S1 S2 L1 L2 L3 L4 I H Ha

Euphydryas chalcedona sierra (W. G. Wright, 1905)  TL: USA (Ca)  M T S1 S2 L H

Euphydryas chalcedona olancha (W. G. Wright, 1905)  TL: USA (Ca)  M T S Ha

Euphydryas chalcedona macglashanii (Rivers, 1888)  TL: USA (Ca)  M S1 S2

Euphydryas chalcedona dwinellei (H. Edwards, 1881)  TL: USA (Ca)  M S1 S2 S3 Ab

Euphydryas chalcedona trinitina Baughman & Murphy, 1998  TL: USA (Ca)  M L I Ha

Euphydryas chalcedona chalcedona (E. Doubleday, [1847])  [= sinecat]  M T1 T2 S1 S2 S3 Ab L1 L2 L3 L4 L5 I Ha

Euphydryas colon (W. H. Edwards, 1881)  Snowberry Checkerspot    iN

Euphydryas colon colon (W. H. Edwards, 1881)  TL: USA (Wa)  M S Ab L1 L2 Ha

Euphydryas colon svihlae Pelham, 2008  TL: USA (Wa)  M S

Euphydryas colon E-central OR Cascades segregate ,   M L Ha

Euphydryas colon SE Cascades, Warner Mts. segregate ,  [nr. wallacensis]  M S1 S2 L1 L2 L3 Ha

Euphydryas colon wallacensis Gunder, 1928  TL: USA (Id)  M T S1 S2 S3 Ab L Ha

Euphydryas colon nevadensis Bauer, 1975  TL: USA (Nv)  M S Ha

Euphydryas colon central WA Cascades segregate ,  [often called paradoxa in literature] C WA Cascades from C Yakima to far S Chelan Co.  M L1 L2 L3 I Ha

Euphydryas colon paradoxa McDunnough, 1927  TL: Canada (BC)  M S

Euphydryas anicia (E. Doubleday, [1847])  Anicia Checkerspot    iN

Euphydryas anicia helvia (Scudder, 1869)  TL: USA (Ak)  M S1 S2

Euphydryas anicia anicia (E. Doubleday, [1847])  TL: Canada (Ab)  M S1 S2

Euphydryas anicia howlandi D. Stallings & Turner, 1947  TL: USA (Mt)  M T S1 S2

Euphydryas anicia hopfingeri Gunder, 1934  TL: USA (Wa)  M S

Euphydryas anicia N WA Cascades segregate ,   M Ab L Ha

Euphydryas anicia NE OR, high elevations segregate ,  [nr. howlandi]  M S

Euphydryas anicia bakeri D. Stallings & Turner, 1945  TL: USA (Or)  M T S1 S2

Euphydryas anicia effi D. Stallings & Turner, 1945  TL: USA (Id)  M T S1 S2

Euphydryas anicia windi Gunder, 1932  TL: USA (Wy)  M S Ab L Ha

Euphydryas anicia maria (Skinner, 1899)  TL: USA (Ut)  M S1 S2

Euphydryas anicia brucei (W. H. Edwards, 1888)  TL: USA (Co)  M T S L1 L2 L3

Euphydryas anicia eurytion (Mead, 1875)  TL: USA (Co)  M T S Ab L1 L2 H

Euphydryas anicia wecoeut M. Fisher, Spomer & Scott, 2006  TL: USA (Co)  M T S

Euphydryas anicia alena W. Barnes & Benjamin, 1926  TL: USA (Ut)  M S L

Euphydryas anicia magdalena W. Barnes & McDunnough, 1918  TL: USA (Az)  M S L1 L2 L3 H

Euphydryas anicia hermosa (W. G. Wright, 1905)  TL: USA (Az)  M T S L1 L2 I

Euphydryas anicia chuskae (Ferris & R. Holland, 1980)  TL: USA (NM)  M T S

Euphydryas anicia carmentis W. Barnes & Benjamin, 1926  TL: USA (Co)  M S1 S2

Euphydryas anicia cloudcrofti (Ferris & R. Holland, 1980)  TL: USA (NM)  M T S L

Euphydryas anicia capella (W. Barnes, 1897)  TL: USA (Co)  M S1 S2 S3 L

Euphydryas anicia bernadetta Leussler, 1920  [= rorina]  M T S L1 L2 L3 L4 Ha

Euphydryas anicia veazieae Fender & Jewett, 1953  TL: USA (Or)  M S1 S2 Ab L Ha

Euphydryas anicia macyi Fender & Jewett, 1953  TL: USA (Or)  M S1 S2

Euphydryas anicia wheeleri (H. Edwards, 1881)  TL: USA (Nv)  M S L Ha

Euphydryas anicia morandi (Gunder, 1928)  TL: USA (Nv)  M T S

Euphydryas anicia variicolor Baughman & Murphy, 1998  TL: USA (Ca)  M S L

Euphydryas phaeton (Drury, 1773)  Baltimore Checkerspot    iN

Euphydryas phaeton borealis F. Chermock & R. Chermock, 1940  TL: USA (Me)

Euphydryas phaeton phaeton (Drury, 1773)  TL: USA (NY)  M T S Ab L1 L2 I H

Euphydryas phaeton clarki Pavulaan, 2021  TL: USA (Md)  M T

Euphydryas phaeton ozarkae Masters, 1968  TL: USA (Mo)  M S1 S2 Ab L

Poladryas  Bauer, 1961  

Poladryas arachne (W. H. Edwards, 1869)  Arachne Checkerspot    iN

Poladryas arachne arachne (W. H. Edwards, 1869)  TL: USA (Co)  M T S1 S2 S3 L Ha

Poladryas arachne gilensis (W. Holland, 1930)  TL: USA (Az)  M T S

Poladryas arachne expedita Austin, 1998  TL: USA (Nv)  M T I

Poladryas arachne nympha (W. H. Edwards, 1884)  TL: USA (Az)  M T S Ab L1 L2

Poladryas arachne monache (J. A. Comstock, 1918)  TL: USA (Ca)  M T S

Poladryas minuta (W. H. Edwards, 1861)  Dotted Checkerspot    iN

Poladryas minuta minuta (W. H. Edwards, 1861)  [= simador]; far SE CO, E NM, N & W TX, NE Mexico  M T S I

Poladryas minuta NW Mexican segregate ,  NW Mexico (SON, CHIH, DGO)  M S

Chlosyne  A. Butler, 1870  

Chlosyne gaudialis (H. Bates, 1864)  Gaudy Checkerspot    iN

Chlosyne gaudialis gaudialis (H. Bates, 1864)  SE Mexico to Panama  M T S L

Chlosyne gaudialis wellingi L. Miller & Rotger, 1979  S Oaxaca (Sierra Madre del Sur)  M S

Chlosyne narva (Fabricius, 1793)  Narva Checkerspot    iN

Chlosyne narva narva (Fabricius, 1793)  Nicaragua to Colombia and Venezuela  M S

Chlosyne narva bonpland (Latreille, [1809])  Costa Rica to W Ecuador  M T S1 S2 L

Chlosyne janais (Drury, 1782)  Crimson-patch Checkerspot    iN

Chlosyne janais janais (Drury, 1782)  C & S TX, T & S Mexico to Colombia  M S L1 L2 L3 L4 I H

Chlosyne janais gloriosa Bauer, 1960  W Mexico  M T S1 S2 L

Chlosyne janais marianna Röber, [1914]  S-C Mexico (Balsas Basin)  M T S L

Chlosyne ehrenbergii (Geyer, [1833])  White-rayed Checkerspot  Mexico  iN  M S L

Chlosyne definita (E. Aaron, [1885])  Definite Checkerspot    iN

Chlosyne definita definita (E. Aaron, [1885])  S NM, S & W TX, NE Mexico  M T S L I H

Chlosyne definita dolosa Grishin, 2023  NW Mexico; ref.151

Chlosyne anastasia (Hemming, 1934)  Definite Checkerspot  C & NW Mexico; ref.151  M T S L

Chlosyne eumeda (Godman & Salvin, 1894)  Eumeda Checkerspot  W Mexico (Nayarit to Guerrero)  iN

Chlosyne eumeda NW Mexican segregate ,  Sonora & Sinaloa, NW Mexico  M S L I

Chlosyne eumeda eumeda (Godman & Salvin, 1894)  TL: Mexico (Gue)  M T S L

Chlosyne marina (Geyer, 1837)  Marina Checkerspot  Sonora, Chihuahua  iN

Chlosyne marina marina (Geyer, 1837)  TL: Mexico  M T S

Chlosyne marina SW Mexico segregate ,  Guerrero and Oaxaca  M S L

Chlosyne marina Chiapas segregate ,  Chiapas  M S

Chlosyne melitaeoides (C. Felder & R. Felder, 1867)  Melitaeoides Checkerspot  S TX (as vagrant), E Mexico  iN  M T S L

Chlosyne erodyle (H. Bates, 1864)  Erodyle Checkerspot    iN

Chlosyne erodyle Oaxaca segregate ,  Oaxaca  M S L

Chlosyne erodyle erodyle (H. Bates, 1864)  Chiapas to NW Costa Rica  M T S L

Chlosyne poecile (C. Felder & R. Felder, 1867)  Poecile Checkerspot  SW Costa Rica to Venezuela  iN

Chlosyne poecile poecile (C. Felder & R. Felder, 1867)  TL: Colombia  M T S L

Chlosyne poecile rubrigutta Röber, [1914]  TL: Colombia  M T

Chlosyne poecile rodriguezi Neild, 2008  TL: Venezuela

Chlosyne poecile intermedia Neild, 2008  TL: Venezuela

Chlosyne poecile murphyi Neild, 2008  TL: Venezuela

Chlosyne pardelina Grishin, 2023  Banded Checkerspot  [infrasub. of Higgins 1960] S TX, NE Mexico; ref.151  M T S L H

Chlosyne endeis (Godman & Salvin, 1894)  Banded Checkerspot  W Mexico  iN  M T S

Chlosyne melanarge (H. Bates, 1864)  Cream-banded Checkerspot  E Mexico to Costa Rica  iN  M T S L

Chlosyne hippodrome (Geyer, 1837)  Simple Checkerspot    iN

Chlosyne hippodrome hippodrome (Geyer, 1837)  E & W Mexico to Colombia  M S L1 L2

Chlosyne hippodrome fabricii Higgins, 1960  TL: Indiis

Chlosyne rosita A. Hall, 1924  Rosita Checkerspot    iN

Chlosyne rosita rosita A. Hall, 1924  Chiapas, Guatemala, El Salvador  M T S1 S2 S3 S4

Chlosyne rosita browni Bauer, 1961  S TX (as stray), E Mexico  M T S L I

Chlosyne rosita mazarum L. Miller & Rotger, 1979  TL: Mexico (Mor)

Chlosyne rosita montana A. Hall, 1924  C and NW Mexico, SE AZ (as stray)  M T S Ab L

Chlosyne rosita riobalsensis Bauer, 1961  SW Mexico (Rio Balsas basin) [mazarum = form of riobalsensis]  M T S1 S2 L Ha

Chlosyne perlula (C. Felder & R. Felder, 1861)  Perula Checkerspot  Venezuela & Colombia  iN  M T S

Chlosyne ezra (Hewitson, 1864)  Ezra Checkerspot  Costa Rica & Panama  iN  M T S L

Chlosyne theona (Ménétriés, 1855)  Theona Checkerspot    iN

Chlosyne theona costaricensis (Austin & M. Smith, 1998)  NW Costa Rica, SW Nicaragua  M S L

Chlosyne theona theona (Ménétriés, 1855)  Chiapas to Nicaragua  M S L

Chlosyne theona minimus (Austin & M. Smith, 1998)  Yucatan Peninsula to Honduras  M S L

Chlosyne theona mullinsi (Austin & M. Smith, 1998)  SW & C Mexico  M S L I

Chlosyne theona brocki (Austin & M. Smith, 1998)  NW Mexico (Sonora to N Nayarit)  M S L I

Chlosyne theona thekla (W. H. Edwards, 1870)  far NW Mexico (N Sonora, SE AZ, SW NM  M T S L1 L2 I H

Chlosyne theona hondana (Weymer, 1890)  TL: Colombia  M T

Chlosyne bollii (W. H. Edwards, [1878])  Theona Checkerspot  C Texas, NE Mexico; ref.151  M T S Ab L1 L2 L3 I H

Chlosyne chinatiensis (Tinkham, 1944)  Theona Checkerspot  W TX, N-C Mexico (Dur); ref.11,151  iN  M S I

Chlosyne cyneas (Godman & Salvin, 1878)    iN  M T S1 S2 Hy L1 L2 L3 I H

Chlosyne cynisca (Godman & Salvin, 1882)  Oaxacan Checkerspot  Oaxaca, Puebla  iN  M T S L

Chlosyne fulvia (W. H. Edwards, 1879)  Fulvia Checkerspot  W NE, W KS, E CO, E NM, W TX, far NE Mexico; ref.11  iN  M T S1 S2 L F

Chlosyne coronado (M. Smith & Brock, 1988)  Fulvia Checkerspot  AZ, SW NM, far NW Mexico; ref.11  iN

Chlosyne coronado pariaensis (M. Smith & Brock, 1988)  S-central UT, N-central AZ; ref.11  M T S L I Ha

Chlosyne coronado coronado (M. Smith & Brock, 1988)  AZ, SW NM, far NW Mexico; ref.11  M T S L1 L2 L3 I H

Chlosyne leanira (C. Felder & R. Felder, 1860)  Leanira Checkerspot    iN

Chlosyne leanira oregonensis Bauer, 1975  TL: USA (Or)  M S L Ha

Chlosyne leanira leanira (C. Felder & R. Felder, 1860)  TL: USA (Ca)  M T S L I

Chlosyne leanira obsoleta (H. Edwards, 1877)  TL: USA (Ca)  M T S

Chlosyne leanira nebularum (Austin & M. Smith, 1998)  TL: USA (Ca)  M T S L

Chlosyne leanira elegans (Priestaf & J. Emmel, 1998)  TL: USA (Ca)  M T S L1 I F Ha

Chlosyne leanira wrightii (W. H. Edwards, 1886)  TL: USA (Ca)  M T S L1 L2 I H

Chlosyne leanira alma (Strecker, [1878])  TL: USA (Az)  M T S L1 L2 I1 I2 H Ha

Chlosyne leanira basinensis (Austin & M. Smith, 1998)  TL: USA (Ca)  M T S Ha

Chlosyne leanira flavodorsalis (Austin & M. Smith, 1998)  TL: USA (Ut)  M T S

Chlosyne leanira austrima (Austin & M. Smith, 1998)  TL: Mexico (BCN)  M T S

Chlosyne nycteis (E. Doubleday, [1847])  Silvery Checkerspot    iN

Chlosyne nycteis pastoron Pelham, 2008  [= obsoleta]  M T L

Chlosyne nycteis nycteis (E. Doubleday, [1847])  TL: USA  M T OI S1 S2 L1 L2 L3 L4 L5 L6 L7 I F

Chlosyne nycteis reversa (F. Chermock & R. Chermock, 1940)  TL: Canada (Mt)  M S

Chlosyne nycteis drusius (W. H. Edwards, 1884)  TL: USA (Co)  M S1 S2 L1 L2 Ha

Chlosyne gorgone (Hübner, [1810])  Gorgone Checkerspot    iN

Chlosyne gorgone gorgone (Hübner, [1810])  TL: ?  M T S

Chlosyne gorgone carlota (Reakirt, 1866)  TL: USA (Mo)  M T S Ab HS L1 L2 L3 I F

Chlosyne californica (W. G. Wright, 1905)  California Patch    iN  M T S L1 L2 I H

Chlosyne lacinia (Geyer, 1837)  Bordered Patch    iN

Chlosyne lacinia lacinia (Geyer, 1837)  C Mexico to Panama  M T S L1 L2

Chlosyne lacinia adjutrix Scudder, 1875  NE Mexico, S & C Texas (strays N)  M T S Ab L1 L2 L3 L4 L5 I1 I2 H

Chlosyne lacinia crocale (W. H. Edwards, 1874)  NW Mexico, SW USA (strays N)  M T S L1 L2 I

Chlosyne lacinia saundersi (E. Doubleday, [1847])  TL: Venezuela  M T L

Chlosyne kendallorum Opler, 1999  Kendall's Checkerspot  NE Mexico  iN  M T S

Chlosyne harrisii (Scudder, 1863)  Harris's Checkerspot    iN

Chlosyne harrisii harrisii (Scudder, 1863)  TL: USA (Me/NH/Ma)  M S L I Ha

Chlosyne harrisii albimontana (Avinoff, 1930)  TL: USA (NH)  M S

Chlosyne harrisii liggetti (Avinoff, 1930)  TL: USA (Pa)  M S

Chlosyne harrisii hanhami (Fletcher, 1903)  TL: USA (Mn)  M S

Chlosyne hoffmanni (Behr, 1863)  Hoffmann's Checkerspot    iN

Chlosyne hoffmanni hoffmanni (Behr, 1863)  TL: USA (Ca)  M T S L1 L2

Chlosyne hoffmanni segregata (W. Barnes & McDunnough, 1918)  TL: USA (Or)  M T S L Ha

Chlosyne hoffmanni manchada Bauer, 1960  TL: USA (Wa)  M T L Ha

Chlosyne whitneyi (Behr, 1863)  Sierra Nevada Checkerspot    iN  M T S L

Chlosyne acastus (W. H. Edwards, 1874)  Sagebrush Checkerspot    iN

Chlosyne acastus sabina (W. G. Wright, 1905)  TL: USA (Az)  M T S L I

Chlosyne acastus neumoegeni (Skinner, 1895)  TL: USA (Ut)  M T S L1 L2 L3 I H Ha

Chlosyne acastus acastus (W. H. Edwards, 1874)  TL: USA (Ut)  M S I

Chlosyne acastus arkanyon M. Fisher & Scott, 2006  TL: USA (Co)  M T S1 S2 L1 L2 Ha

Chlosyne acastus vallismortis (J. Johnson, 1938)  TL: USA (Ca)  M T S

Chlosyne acastus robusta Austin, 1998  TL: USA (Nv)  M T S

Chlosyne acastus waucoba J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T

Chlosyne acastus far N Great Basin segregate ,  [nr. acastus]  M S

Chlosyne acastus dorothyi Bauer, 1975  TL: USA (Or)  M S

Chlosyne gabbii (Behr, 1863)  Gabb’s Checkerspot    iN

Chlosyne gabbii gabbii (Behr, 1863)  TL: USA (Ca)  M T S L1 L2 I Ha

Chlosyne gabbii atrifasciata Hawks & J. Emmel, 1998  TL: USA (Ca)

Chlosyne palla (Boisduval, 1852)  Northern Checkerspot    iN

Chlosyne palla sterope (W. H. Edwards, 1870)  NW US & SW Can; ref.16  M T L Ha

Chlosyne palla E Columbia Gorge segregate ,   M S L Ha

Chlosyne palla E-central OR Cascades segregate ,   M L Ha

Chlosyne palla eremita (W. G. Wright, 1905)  TL: USA (Ca)  M T S L Ha

Chlosyne palla palla (Boisduval, 1852)  TL: USA (Ca)  M S L Ha

Chlosyne palla altasierra J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T L

Chlosyne palla pola (Boisduval, 1869)  ref.20  M T

Chlosyne palla australomontana J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L1 L2 L3

Chlosyne flavula (W. Barnes & McDunnough, 1918)  Northern Checkerspot  ref.20

Chlosyne flavula blackmorei Pelham, 2008  TL: not specified  M S L I Ha

Chlosyne flavula flavula (W. Barnes & McDunnough, 1918)  TL: USA (Co)  M S1 S2 L1 L2 L3 L4 Ha

Chlosyne flavula calydon (W. Holland, 1931)  TL: USA (Co)  M S

Chlosyne damoetas (Skinner, 1902)  Rockslide Checkerspot    iN

Chlosyne damoetas damoetas (Skinner, 1902)  TL: USA (Co)  M S1 S2 L1 L2 I Ha

Chlosyne damoetas windriver Scott, 1998  TL: USA (Wy)  M S L

Chlosyne damoetas altalus Scott, 1998  TL: not specified  M T

Microtia  H. Bates, 1864  

Microtia elvira Grishin, 2023  Elfoid  S AZ & W Mexico; ref.146  M T S L I

Microtia elva H. Bates, 1864  Elf  ref.146  iN

Microtia elva elva H. Bates, 1864  S TX (as stray), E Mexico to Costa Rica  M T S1 S2 L

Microtia elva horni Rebel, 1906  S Mexico & Guatemala  M T S L

Microtia elva bogotana Grishin, 2024  E Colombia; ref.154  M T

Microtia dymas (W. H. Edwards, 1877)  Tiny Checkerspot  ref.12  iN

Microtia dymas dymas (W. H. Edwards, 1877)  C TX to NE Mexico  M T S L1 L2 I1 I2 I3

Microtia dymas chara (W. H. Edwards, [1884])  S AZ, S NM, W TX, NW Mexico, Baja California Sur  M T S L1 L2 I H

Microtia dymas imperialis Bauer, 1959  S CA, Baja California Norte  M T S L Ha

Microtia perse (W. H. Edwards, 1882)  Arizona Checkerspot  S AZ, SW NM, N Sonora, NW Chihuahua; ref.12  iN  M T S L I H

Microtia elada (Hewitson, 1868)  Elada Checkerspot  ref.12  iN

Microtia elada ulrica (W. H. Edwards, 1877)  C & W TX, NE Mexico  M T S L1 L2 L3 I H

Microtia elada elada (Hewitson, 1868)  C & S Mexico  M T S L

Microtia elada hepburni (Godman, 1901)  NW Mexico  M T S L

Microtia anomalus (Godman & Salvin, 1897)  Anomalus Checkerspot  W Mexico (Nay to Mich); ref.12  iN

Microtia anomalus anomalus (Godman & Salvin, 1897)  TL: Mexico (Col)  M T S

Microtia coracara (Dyar, 1912)  Coracara Checkerspot  SW Mexico (Mor, Gue, Oax); ref.12  iN  M T S L

Gnathotriche  C. Felder & R. Felder, 1862  

Gnathotriche exclamationis (Kollar, 1850)    iN

Gnathotriche exclamationis exclamationis (Kollar, 1850)  TL: Venezuela  M T

Gnathotriche exclamationis elaea (Hewitson, 1869)  TL: Ecuador  M T

Gnathotriche exclamationis sodalis Staudinger, 1885  TL: Colombia  M T

Gnathotriche mundina (H. Druce, 1876)    iN

Gnathotriche mundina mundina (H. Druce, 1876)  TL: Peru  M T

Gnathotriche mundina erebia (Röber, [1914])  TL: Peru  M T

Gnathotriche mundina fallax (Staudinger, 1885)  TL: Peru  M T

Gnathotriche mundina hopfferi (Röber, [1914])  TL: [Bolivia]  M T

Gnathotriche mundina rosina (Dognin, 1888)  TL: Ecuador  M T

Gnathotriche mundina steinii (Dewitz, 1877)  TL: Colombia  M T

Higginsius  Hemming, 1964  

Higginsius fasciata (Hopffer, 1874)    iN  M T

Higginsius miriam (Dognin, 1888)  TL: Ecuador  M T

Antillea  Higgins, 1959  

Antillea pelops (Drury, 1773)  Antillean Crescent    iN

Antillea pelops pelops (Drury, 1773)  Hispaniola, Puerto Rico, St. Kitts  M T S L

Antillea pelops anocaona (Herrich-Schäffer, 1865)  Cuba  M T S

Antillea pelops pygmaea (Godart, 1819)  Jamaica  M T S

Antillea proclea (E. Doubleday, [1847])  Jamaican Crescent  Jamaica  iN  M T S L

Atlantea  Higgins, 1959  

Atlantea cryptadia Sommer & A. Schwartz, 1980  Hispaniolan Checkerspot  Hispaniola  M T S

Atlantea pantoni (Kaye, 1906)  Jamaican Checkerspot  Jamaica  iN  M T S L

Atlantea perezi (Herrich-Schäffer, 1862)  Cuban Checkerspot  NE Cuba  iN  M T S L

Atlantea tulita (Dewitz, 1877)  Puerto Rican Checkerspot  Puerto Rico  iN  M S L I

Phystis  Higgins, 1981  

Phystis simois (Hewitson, 1864)  ref.12  iN  M T

Phystis pratti (A. Hall, 1935)  ref.12

Phystis pratti pratti (A. Hall, 1935)  TL: Peru  M T

Phystis pratti chinchipensis (Hayward, 1964)  TL: Peru  M T

Phystis variegata (Röber, 1913)  ref.12  M T

Phystis fontus (A. Hall, 1928)  ref.12

Phystis fontus fontus (A. Hall, 1928)  TL: Guyana  M T

Phystis fontus schmitzorum (Austin & O. Mielke, 1993)  TL: Brazil (Rd)  M T

Mazia  Higgins, 1981  

Mazia amazonica (H. Bates, 1864)    iN

Mazia amazonica amazonica (H. Bates, 1864)  TL: Brazil (Pa)  M T S

Mazia amazonica cocha Lamas, 1995  TL: Peru

Mazia amazonica tambopata Lamas, 1995  TL: Peru

Tisona  Higgins, 1981  

Tisona saladillensis (Giacomelli, 1911)  TL: Argentina

Tisona saladillensis saladillensis (Giacomelli, 1911)  TL: Argentina  M T

Tisona saladillensis clarior Higgins, 1981  TL: Bolivia  M T

Ortilia  Higgins, 1981  

Ortilia gentina Higgins, 1981    iN  M T

Ortilia liriope (Cramer, 1775)    iN  M S

Notilia  Grishin, 2021  

Notilia dicoma (Hewitson, 1864)  ref.12  iN  M T

Notilia orthia (Hewitson, 1864)  ref.12  iN  M T

Notilia orticas (Schaus, 1902)  ref.12  iN

Notilia orticas orticas (Schaus, 1902)  TL: Brazil (Pr)  M T L

Notilia orticas zamora (A. Hall, 1917)  TL: Venezuela  M T

Notilia polinella (A. Hall, 1928)  ref.12  M T

Notilia sejona (Schaus, 1902)  ref.12  iN  M T

Notilia velica (Hewitson, 1864)  ref.12  iN

Notilia velica velica (Hewitson, 1864)  TL: ?  M T

Notilia velica durnfordi (Godman & Salvin, 1878)  TL: Argentina  M T

Levinata  Grishin, 2021  

Levinata levina (Hewitson, 1872)  ref.12  iN  M T

Anthanassa  Scudder, 1875  

Anthanassa acesas (Hewitson, 1864)    iN

Anthanassa acesas acesas (Hewitson, 1864)  TL: Venezuela  M T

Anthanassa acesas castianira (Godman & Salvin, 1880)  TL: Colombia  M T

Anthanassa annulata Higgins, 1981    iN  M T

Anthanassa ardys (Hewitson, 1864)  Ardys Crescent    iN

Anthanassa ardys ardys (Hewitson, 1864)  E & W Mexico through Oaxaca  M T S L

Anthanassa ardys [n. ssp.] Lamas, MS  Nicaragua to Colombia  M S L

Anthanassa subota (Godman & Salvin, 1878)  Cryptic Crescent  Chiapas & Guatemala  iN

Anthanassa subota [n. ssp.] Lamas, MS  W Mexico (Nayarit to Oaxaca)  M S

Anthanassa subota subota (Godman & Salvin, 1878)  TL: Guatemala  M T S

Anthanassa argentea (Godman & Salvin, 1882)  Chestnut Crescent  S TX (as stray), E & S Mexico, Guatemala, Nicaragua  iN  M T S L

Anthanassa atronia (H. Bates, 1866)  Brown Crescent  E & S Mexico to Panama  iN  M T S L

Anthanassa crithona (Salvin, 1871)  Orange-banded Crescent  Nicaragua to Panama  iN  M T S L

Anthanassa dracaena (C. Felder & R. Felder, 1867)    iN

Anthanassa dracaena dracaena (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Anthanassa dracaena phlegias (Godman, 1901)  S Mexico to Costa Rica  M T S L

Anthanassa drusilla (C. Felder & R. Felder, 1861)  Orange-patched Crescent    iN

Anthanassa drusilla drusilla (C. Felder & R. Felder, 1861)  Panama, Colombia, Venezuela, Ecuador  M T S L

Anthanassa drusilla alceta (Hewitson, 1869)  TL: Ecuador  M T L

Anthanassa drusilla halli Neild, 2008  ref.266

Anthanassa drusilla forbesi Neild, 2008  ref.266

Anthanassa drusilla higginsi Neild, 2008  ref.266

Anthanassa drusilla lelex (H. Bates, 1864)  S TX (rare stray), E & S Mexico to Panama  M T S L

Anthanassa drusilla verena (Hewitson, 1864)  TL: Bolivia  M T

Anthanassa drymaea (Godman & Salvin, 1878)  Weak-banded Crescent  S Mexico (Chiapas), Guatemala, Honduras  iN  M T S L

Anthanassa frisia (Poey, 1832)  Cuban Crescent  S FL (strays N), N Bahamas, Cuba, Hispaniola, Jamaica  iN  M T S L

Anthanassa hermas (Hewitson, 1864)    iN

Anthanassa hermas dubia (A. Hall, 1929)  TL: Venezuela  M T

Anthanassa hermas hermas (Hewitson, 1864)  TL: [Brazil (Pe)]  M T

Anthanassa hermas taeniata (Röber, 1913)  TL: Peru  M T L

Anthanassa tulcis (H. Bates, 1864)  Pale-banded Crescent  SE AZ (as stray), S TX, E & W Mexico to Panama  iN  M T S L1 L2 L3 I

Anthanassa nebulosa (Godman & Salvin, 1878)  Nebulosa Crescent    iN

Anthanassa nebulosa nebulosa (Godman & Salvin, 1878)  Chiapas & Guatemala  M T S

Anthanassa nebulosa alexon (Godman & Salvin, 1889)  [= subconcolor], E & W Mexico  M T S L

Anthanassa otanes (Hewitson, 1864)  Blackened Crescent    iN

Anthanassa otanes otanes (Hewitson, 1864)  [= sopolis], Chiapas, W Guatemala to Costa Rica; also W Mexico?  M T S L

Anthanassa otanes cyno (Godman & Salvin, 1889)  E Mexico  M T S

Anthanassa otanes fulviplaga (A. Butler, 1872)  Costa Rica & Panama  M T S L

Anthanassa otanes oaxaca Beutelspacher, 1990  vic. Sierra de Juarez, Oaxaca, Mexico  M T S L

Anthanassa ptolyca (H. Bates, 1864)  Darkened Crescent    iN

Anthanassa ptolyca E Mexican segregate ,  S TX (rare stray), E Mexico  M S L

Anthanassa ptolyca ptolyca (H. Bates, 1864)  Chiapas to Venezuela  M T S L

Anthanassa ptolyca amator (A. Hall, 1929)  W Mexico  M T S L

Anthanassa sitalces (Godman & Salvin, 1882)  Montane Crescent    iN

Anthanassa sitalces sitalces (Godman & Salvin, 1882)  Chiapas & Guatemala  M T S

Anthanassa sitalces cortes (A. Hall, 1917)  W Mexico to Oaxaca  M T S L

Anthanassa sosis (Godman & Salvin, 1878)  Sosis Crescent  [= dora], Costa Rica to Colombia  iN  M T S L

Anthanassa texana (W. H. Edwards, 1863)  Texan Crescent  AZ to TX (strays far N), Baja, E & W Mexico to Guatemala  iN  M T S Ab L I H

Anthanassa seminole (Skinner, 1911)  Texan Crescent  SC to FL, E to LA & far E TX; ref.146  M S L H

Castilia  Higgins, 1981  

Castilia angusta (Hewitson, 1868)    iN  M T

Castilia castilla (C. Felder & R. Felder, 1862)    iN

Castilia castilla castilla (C. Felder & R. Felder, 1862)  TL: Colombia  M T S

Castilia castilla occidentalis (Fassl, 1912)  TL: Colombia

Castilia chiapaensis (Beutelspacher, 1990)  Chiapas Crescent  S Mexico (Chiapas)  iN  M T S

Castilia chinantlensis (R. F. Maza, 1978)  Chinantlan Crescent  SE Mexico (Sierra de Juarez, Oaxaca)  iN  M T S

Castilia eranites (Hewitson, 1857)  Mimic Crescent  S Mexico to Colombia & Venezuela  iN  M T S L

Castilia fulgora (Godman & Salvin, 1878)  Fulgora Crescent  Costa Rica (Atlantic slope, Cordillera Central) & W Panama  iN  M T TI L

Castilia griseobasalis (Röber, 1913)  Gray-based Crescent  E & W Mexico to Costa Rica  iN  M T S L

Castilia guaya (A. Hall, 1929)  TL: Ecuador  M T

Castilia myia (Hewitson, [1864])  Mayan Crescent  E & W Mexico to Panama  iN  M T S L

Castilia neria (Hewitson, 1869)    iN  M T

Castilia nortbrundii (Weeks, 1901)  TL: Bolivia  M T

Castilia ofella (Hewitson, [1864])  White-dotted Crescent  S Mexico, Guatemala to Colombia, Venezuela, Trinidad  iN  M S L1 L2

Castilia perilla (Hewitson, 1852)    iN  M T L

Telenassa  Higgins, 1981  

Telenassa abas (Hewitson, 1864)    iN  M T

Telenassa berenice (C. Felder & R. Felder, 1862)    iN

Telenassa berenice berenice (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S

Telenassa berenice drusinilla (Röber, 1913)  TL: Argentina  M T

Telenassa delphia (C. Felder & R. Felder, 1861)    iN

Telenassa delphia delphia (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Telenassa delphia alani Neild & Orellana, 2008  ref.266

Telenassa delphia austini Neild, 2008  ref.266

Telenassa delphia catenaria (Godman & Salvin, 1880)  TL: Colombia  M T

Telenassa delphia elaphina (Röber, 1913)  TL: Bolivia  M T

Telenassa delphia flavocincta (Dognin, 1887)  TL: Ecuador  M T

Telenassa delphia gaujoni (Dognin, 1887)  TL: Ecuador  M T

Telenassa delphia nana (H. Druce, 1874)  TL: Peru  M T

Telenassa delphia nussia (H. Druce, 1876)  TL: Peru  M T

Telenassa delphia trimaculata (Hewitson, 1869)  TL: Ecuador  M T L

Telenassa jana (C. Felder & R. Felder, 1867)    iN  M T

Telenassa notus (A. Hall, 1917)  TL: Peru  M T

Telenassa sepultus (A. Hall, 1928)  TL: Peru  M T

Telenassa teletusa (Godart, [1824])    iN

Telenassa teletusa teletusa (Godart, [1824])  TL: Brazil  M T

Telenassa teletusa burchelli (Moulton, 1909)  TL: Brazil (Go)  M T

Ithra  Grishin, 2021  

Ithra ithra (W. F. Kirby, 1900)  ref.12  iN  M T

Dagon  Higgins, 1981  

Dagon catula (Hopffer, 1874)    iN  M T

Dagon morena (Röber, 1913)    iN  M T

Dagon pusilla (Salvin, 1869)    iN  M T

Eresia  Boisduval, 1836  

Eresia actinote Salvin, 1869    iN

Eresia actinote actinote Salvin, 1869  TL: Peru  M T

Eresia actinote limbata (Röber, 1913)  TL: Bolivia  M T

Eresia actinote [n. ssp.] Lamas, MS  Peru

Eresia carme E. Doubleday, [1847]    iN

Eresia carme carme E. Doubleday, [1847]  TL: Venezuela  M T

Eresia carme mejiai Neild & De Sousa, 2008  ref.266

Eresia carme judithae Neild, 2008  ref.266

Eresia casiphia Hewitson, 1869    iN  M T

Eresia clio (Linnaeus, 1758)  Clio Crescent    iN

Eresia clio clio (Linnaeus, 1758)  [= clara], S Mexico to Brazil & Bolivia  M T S L

Eresia clio carmenae Neild, 2008  ref.266

Eresia clio reducta (A. Hall, 1929)  TL: Ecuador  M T

Eresia datis Hewitson, [1864]    iN

Eresia datis datis Hewitson, [1864]  TL: [Peru]  M T

Eresia datis corybassa Hewitson, 1874  TL: Bolivia  M T

Eresia datis fassli (Röber, 1913)  TL: Colombia  M T

Eresia datis fenestrata (Röber, 1913)  TL: Peru  M T

Eresia datis magniplaga (Röber, 1927)  TL: Colombia  M T

Eresia datis manto (A. Hall, 1930)  TL: Colombia  M T

Eresia datis margaretha Hewitson, 1872  TL: Colombia  M T

Eresia datis moesta Salvin & Godman, 1868  TL: Ecuador  M T L

Eresia datis phaedima Salvin & Godman, 1868  TL: Peru  M T

Eresia datis [n. ssp.] Lamas, MS  Colombia

Eresia emerantia Hewitson, 1857    iN

Eresia emerantia emerantia Hewitson, 1857  TL: Colombia  M T

Eresia emerantia coela H. Druce, 1874  Costa Rica & NE Panama  M T S L

Eresia emerantia sestia Hewitson, 1869  TL: Ecuador  M T

Eresia erysice (Geyer, 1832)  TL: S. Amer.

Eresia erysice erysice (Geyer, 1832)  TL: S. Amer.

Eresia erysice etesiae (A. Hall, 1928)  TL: French Guiana  M T

Eresia estebana (A. Hall, 1929)  TL: Venezuela

Eresia estebana estebana (A. Hall, 1929)  TL: Venezuela  M T

Eresia estebana camachoi Neild, 2008  ref.266

Eresia eunice (Hübner, [1807])    iN

Eresia eunice eunice (Hübner, [1807])  TL: [Brazil]  M T

Eresia eunice drypetis Godman & Salvin, 1878  Panama, Colombia  M T OI L

Eresia eunice esora Hewitson, 1857  TL: Brazil  M T

Eresia eunice gudruna Röber, 1913  TL: [Brazil (Am)]  M T

Eresia eunice klagesii (Weeks, 1906)  TL: Venezuela  M T

Eresia eunice mechanitis Godman & Salvin, 1878  Nicaragua to Panama  M T L

Eresia eunice polymnia (Röber, 1913)  TL: Colombia

Eresia eunice wallacei Neild, 2008  ref.266

Eresia eunice [n. ssp.] Lamas, MS  Peru

Eresia ithomioides Hewitson, [1864]    iN

Eresia ithomioides ithomioides Hewitson, [1864]  TL: Colombia  M T

Eresia ithomioides alsina Hewitson, 1869  Nicaragua, Costa Rica, W Panama  M T S L

Eresia ithomioides anomala Higgins, 1981  TL: Colombia  M T

Eresia ithomioides cissia (A. Hall, 1928)  TL: Colombia  M T

Eresia ithomioides eutropia Hewitson, 1874  [= melania], Costa Rica, Panama, W Colombia  M T S L

Eresia ithomioides mimas (Staudinger, 1885)  TL: Colombia  M T

Eresia ithomioides poecilina H. Bates, 1866  Costa Rica & Panama  M T S

Eresia ithomioides quintilla Hewitson, 1872  TL: Ecuador  M T

Eresia ithomioides [n. ssp.] Lamas, MS  Colombia

Eresia lansdorfi (Godart, 1819)    iN  M T

Eresia letitia Hewitson, 1869    iN

Eresia letitia letitia Hewitson, 1869  TL: Ecuador  M T

Eresia letitia ocellata (Röber, 1913)  TL: Peru  M T

Eresia nauplius (Linnaeus, 1758)    iN

Eresia nauplius nauplius (Linnaeus, 1758)  TL: Indiis

Eresia nauplius extensa (A. Hall, 1929)  TL: Brazil (MT)  M T

Eresia nauplius plagiata (Röber, 1913)  TL: Peru  M T

Eresia olivencia H. Bates, 1864  TL: Brazil (Am)  M T

Eresia pelonia Hewitson, 1852    iN

Eresia pelonia pelonia Hewitson, 1852  TL: Ecuador  M T L

Eresia pelonia callonia (Staudinger, 1885)  TL: Peru  M T

Eresia pelonia callonioides Strand, 1912  TL: Peru  M T

Eresia pelonia ithomiola Salvin, 1869  TL: Peru  M T L

Eresia perna Hewitson, 1852    iN

Eresia perna perna Hewitson, 1852  TL: Brazil (RJ)  M T

Eresia perna aveyrona H. Bates, 1864  TL: Brazil (Pa)  M T

Eresia perna mylitta Hewitson, 1869  TL: Ecuador  M T

Eresia perna [n. ssp.] Lamas, MS  Brazil

Eresia phillyra Hewitson, 1852  Longwing Crescent    iN

Eresia phillyra phillyra Hewitson, 1852  E & S Mexico, Guatemala  M T S L

Eresia phillyra nigripennis Salvin, 1869  Costa Rica, [Panama?]  M T OI

Eresia polina Hewitson, 1852    iN

Eresia polina polina Hewitson, 1852  TL: Ecuador  M T

Eresia polina joaquini Neild, 2008  ref.266

Eresia polina laias Godman & Salvin, 1879  TL: Colombia  M T

Eresia polina lycus (A. Hall, 1928)  TL: Colombia  M T

Eresia polina oblita (Staudinger, 1885)  TL: Venezuela  M T

Eresia sticta Schaus, 1913  Costa Rican Crescent  Costa Rica (Carrillo belt, Atlantic slope), [Colombia?]  iN  M T S L

Janatella  Higgins, 1981  

Janatella fellula (Schaus, 1902)    iN  M T

Janatella hera (Cramer, 1779)    iN  M T

Janatella leucodesma (C. Felder & R. Felder, 1861)  Whitened Crescent  Nicaragua to Venezuela & Trinidad  iN  M T S L

Tegosa  Higgins, 1981  

Tegosa anieta (Hewitson, 1864)  Black-bordered Crescent    iN

Tegosa anieta anieta (Hewitson, 1864)  Nicaragua to Venezuela & Trinidad  M T S L

Tegosa anieta cluvia (Godman & Salvin, 1882)  Chiapas & Guatemala  M T S L

Tegosa anieta lirina (Röber, 1913)  TL: Bolivia  M T

Tegosa anieta luka Higgins, 1981  S TX (rare stray), E Mexico to Honduras  M T S L

Tegosa claudina (Eschscholtz, 1821)  Claudina Crescent  SE Mexico to Argentina  iN  M T OI S L

Tegosa etia (Hewitson, 1868)    iN  M T

Tegosa flavida (Hewitson, 1868)    iN  M T

Tegosa fragilis (H. Bates, 1864)    iN  M T

Tegosa guatemalena (H. Bates, 1864)  Guatemalan Crescent  E & W Mexico, Guatemala, Belize  iN  M T S L

Tegosa infrequens Higgins, 1981    iN  M T

Tegosa nazaria (C. Felder & R. Felder, 1867)    iN  M T

Tegosa nigrella (H. Bates, 1866)  Nigrella Crescent    iN

Tegosa nigrella nigrella (H. Bates, 1866)  S Mexico (Chiapas), Guatemala, Honduras  M T S L

Tegosa nigrella niveonotis (A. Butler & H. Druce, 1872)  Costa Rica (Atlantic slope)  M T S L

Tegosa orobia (Hewitson, 1864)    iN

Tegosa orobia orobia (Hewitson, 1864)  TL: ?  M T

Tegosa orobia ursula (Staudinger, 1894)  TL: Bolivia  M T

Tegosa pastazena (H. Bates, 1864)    iN  M T

Tegosa selene (Röber, 1913)    iN  M T

Tegosa serpia Higgins, 1981    iN  M T

Tegosa tissoides (A. Hall, 1928)    iN  M T

Phyciodes  Hübner, [1819]  

Phyciodes graphica (R. Felder, 1869)  Vesta Crescent  [= vesta]  iN

Phyciodes graphica graphica (R. Felder, 1869)  TL: Mexico (Oax)  M T S Ma L1 L2 L3 I H

Phyciodes graphica vesta (W. H. Edwards, 1869)  TL: USA (Tx)  M T

Phyciodes pallescens (R. Felder, 1869)  Mexican Crescent  E & W Mexico, SE AZ as rare stray  iN  M T S L

Phyciodes picta (W. H. Edwards, 1865)  Painted Crescent    iN

Phyciodes picta picta (W. H. Edwards, 1865)  TL: USA (Nb)  M S L1 L2

Phyciodes picta canace W. H. Edwards, 1871  TL: USA (Az)  M T S L1 L2

Phyciodes orseis W. H. Edwards, 1871  California Crescent    iN

Phyciodes orseis orseis W. H. Edwards, 1871  TL: USA (Ca)  M S1 S2 L I Ha

Phyciodes orseis Warner Mts. segregate ,  Warner Mts. (OR/CA) W to SE Siskiyou Co., CA  M S

Phyciodes orseis herlani Bauer, 1975  TL: USA (Nv)  M T S L

Phyciodes pallida (W. H. Edwards, 1864)  Pale Crescent    iN

Phyciodes pallida pallida (W. H. Edwards, 1864)  TL: USA (Co)  M S L Ha

Phyciodes pallida barnesi Skinner, 1897  TL: USA (Co)  M S1 S2

Phyciodes mylitta (W. H. Edwards, 1861)  Mylitta Crescent    iN

Phyciodes mylitta mylitta (W. H. Edwards, 1861)  BC to MT, S to S CA  M T S1 S2 L1 L2 Ha

Phyciodes mylitta arizonensis Bauer, 1975  AZ, NM, W TX, N to E CO, far NW Mexico  M S Ab L1 L2 I

Phyciodes mylitta mexicana A. Hall, 1928  TL: Mexico (Ver)  M T S L Ha

Phyciodes mylitta thebais Godman & Salvin, 1878  [= arida]  M T S L

Phyciodes phaon (W. H. Edwards, 1864)  Phaon Crescent    iN

Phyciodes phaon phaon (W. H. Edwards, 1864)  E & C Mexico, TX to FL & VA (strays far N), Cuba, Grand Cayman; ref.16  M T S Ab L1 L2 L3 I F

Phyciodes phaon maya A. Hall, 1928  Chiapas & Guatemala  M T

Phyciodes jalapeno Scott, 1998  Phaon Crescent  NW Mexico & Baja (N), S CA, S AZ; ref.16  M T S L Ha

Phyciodes orantain Scott, 1998  Pearl Crescent  ref.16  M T S1 S2 S3 Ha

Phyciodes tharos (Drury, 1773)  Pearl Crescent    iN

Phyciodes tharos tharos (Drury, 1773)  TL: [USA (NY)]  M T S1 S2 S3 Ab L1 L2 L3 L4 L5 L6 L7 L8 Ha

Phyciodes tharos distincta Bauer, 1975  CA, AZ

Phyciodes tharos riocolorado Scott, 1992  TL: USA (Ut)  M T S

Phyciodes incognitus Gatrelle, 2004  Incognito Crescent  ref.16  M T L

Phyciodes diminutor Scott, 1998  Diminutor Crescent  ref.16  M T L

Phyciodes cocyta (Cramer, 1777)  Northern Crescent    iN

Phyciodes cocyta cocyta (Cramer, 1777)  TL: not specified  M S L Ha

Phyciodes cocyta selenis (W. Kirby, 1837)  TL: not specified  M T S1 S2 L1 L2 Ha

Phyciodes cocyta pascoensis W. G. Wright, 1905  TL: USA (Wa)  M T S

Phyciodes cocyta arenacolor Austin, 1998  TL: USA (Nv)  M T S

Phyciodes cocyta apache Scott & Opler, 2017  TL: USA (NM)

Phyciodes anasazi Scott, 1994  Anasaz Crescent  ref.16

Phyciodes anasazi anasazi Scott, 1994  TL: USA (Co)  M T S

Phyciodes anasazi apsaalooke Scott, 1994  ref.16  M S

Phyciodes batesii (Reakirt, [1866])  Tawny Crescent    iN

Phyciodes batesii lakota Scott, 1994  TL: USA (Ne)  M T S1 S2 L Ha

Phyciodes batesii saskatchewan Scott, 2006  TL: not specified  M T

Phyciodes batesii batesii (Reakirt, [1866])  TL: USA (Va)  M T OD S

Phyciodes batesii maconensis Gatrelle, 1998  TL: USA (NC)  M T L1 L2

Phyciodes pulchella (Boisduval, 1852)  Field Crescent    iN

Phyciodes pulchella tutchone Scott, 1994  TL: not specified  M T S L

Phyciodes pulchella owimba Scott, 1998  TL: not specified  M S Ab L1 L2 Ha

Phyciodes pulchella Siskiyou segregate ,  Siskiyou Mts., SW OR, NW CA  M L Ha

Phyciodes pulchella Willamette Valley segregate ,  Willamette Valley, W Oregon  M S L

Phyciodes pulchella pulchella (Boisduval, 1852)  TL: USA (Ca)  M T S L1 L2 L3 H F Ha

Phyciodes pulchella deltarufa Scott, 1998  TL: USA (Ca)  M T

Phyciodes pulchella S CA Sierra Nevada, low elevations segregate ,   M S L1 L2

Phyciodes pulchella montana (Behr, 1863)  TL: USA (Ca)  M T S1 S2 L Ha

Phyciodes pulchella OR Cascades, high elevations segregate ,  [nr. montana]  M S L Ha

Phyciodes pulchella vallis Austin, 1998  TL: USA (Nv)  M S

Phyciodes pulchella inornatus Austin, 1998  TL: USA (Nv)  M S L1 L2

Phyciodes pulchella shoshoni Scott, 1994  TL: USA (Nv)  M S L Ha

Phyciodes pulchella camillus W. H. Edwards, 1871  TL: USA (Co)  M T S1 S2 L1 L2 L3 Ha

  Tribe Nymphalinae-incertae-sedis

Prodryas  , []  a fossil genus

Prodryas persephone Scudder, 1878  Fossil from Colorado

  Subfamily Charaxinae

  Tribe Anaeini

Coenophlebia  C. Felder & R. Felder, 1862  

Coenophlebia archidona (Hewitson, 1860)    iN  M T

Consul  Hübner, [1807]  

Consul electra (Westwood, 1850)  Pearly Leafwing    iN

Consul electra electra (Westwood, 1850)  E Mexico to Panama  M T S Ab L

Consul electra adustus Lamas, 2003  W Mexico (NAY to MICH)  M T S

Consul excellens (H. Bates, 1864)  Black-veined Leafwing    iN

Consul excellens excellens (H. Bates, 1864)  E Chiapas to Guatemala  M T S L

Consul excellens genini (Le Cerf, 1922)  S Mexico (Oaxaca, W Chiapas) to Guatemala  M T S L

Consul fabius (Cramer, 1776)    iN

Consul fabius fabius (Cramer, 1776)  TL: Surinam  M T

Consul fabius albinotatus (A. Butler, 1874)  TL: Colombia  M T

Consul fabius bogotanus (A. Butler, 1874)  TL: Colombia  M T

Consul fabius castaneus (A. Butler, 1874)  TL: Brazil (Am)  M T

Consul fabius cecrops (E. Doubleday, [1849])  E & W Mexico to Bolivia  M T S Ab L1 L2

Consul fabius diffusus (A. Butler, 1875)  TL: Ecuador  M T

Consul fabius divisus (A. Butler, 1874)  TL: Peru  M T

Consul fabius drurii (A. Butler, 1874)  TL: Brazil  M T

Consul fabius fassli (Röber, 1916)  TL: Colombia  M T

Consul fabius fulvus (A. Butler, 1875)  TL: Peru  M T

Consul fabius ochraceus (A. Butler, 1874)  TL: French Guiana  M T

Consul fabius quadridentatus (A. Butler, 1874)  TL: Bolivia  M T

Consul fabius semifulvus (A. Butler, 1875)  TL: Ecuador  M T

Consul fabius superba (Niepelt, 1923)  TL: Colombia  M T

Consul fabius [n. ssp.] Lamas, MS  Ecuador

Consul panariste (Hewitson, 1856)    iN

Consul panariste panariste (Hewitson, 1856)  TL: Colombia  M T S

Consul panariste jansoni (Salvin, 1871)  Guatemala to Panama  M T S IL

Consul panariste ludmilla (Fassl, 1912)  TL: Colombia  M T

Consul panariste pandrosa (Niepelt, 1927)  TL: Colombia  M T

Hypna  Hübner, [1819]  

Hypna clytemnestra (Cramer, 1777)    iN

Hypna clytemnestra clytemnestra (Cramer, 1777)  TL: Surinam  M T

Hypna clytemnestra corumbaensis Talbot, 1928  TL: Brazil (MT)  M T

Hypna clytemnestra forbesi Godman & Salvin, 1884  TL: Brazil (Pe)  M T

Hypna clytemnestra huebneri A. Butler, 1866  TL: Brazil  M T

Hypna clytemnestra iphigenia Herrich-Schäffer, 1862  Cuba  M T S

Hypna clytemnestra mexicana A. Hall, 1917  W Mexico, Guatemala  M T S L Ha

Hypna clytemnestra negra C. Felder & R. Felder, 1862  Nicaragua to Bolivia  M T S L

Hypna clytemnestra rufescens A. Butler, 1866  TL: Venezuela  M T

Polygrapha  Staudinger, 1887  

Polygrapha cyanea (Salvin & Godman, 1868)    iN

Polygrapha cyanea cyanea (Salvin & Godman, 1868)  TL: Ecuador  M T S

Polygrapha cyanea silvaorum Constantino & Salazar, 1998  TL: Colombia  M T S

Polygrapha suprema (Schaus, 1920)    iN  M T S

Siderone  Hübner, [1823]  

Siderone galanthis (Cramer, 1775)  Red-striped Leafwing    iN

Siderone galanthis Mexican segregate ,  E & W Mexico to Central America  M S L

Siderone galanthis nemesis (Illiger, 1801)  Cuba & Isla de Juventud, Hispaniola, Puerto Rico  M T S

Siderone galanthis galanthis (Cramer, 1775)  Central America to Amazon Basin  M T S L

Siderone galanthis thebais C. Felder & R. Felder, 1862  TL: Colombia  M T

Siderone galanthis [n. ssp.] Lamas, MS  Mexico

Siderone syntyche Hewitson, [1854]  Red-patched Leafwing    iN

Siderone syntyche syntyche Hewitson, [1854]  S Mexico through Central America  M T S

Siderone syntyche angustifascia A. Hall, 1917  TL: Ecuador  M T

Siderone syntyche mars H. Bates, 1860  TL: Brazil (Am)  M T

Phantos  Dias, [2018]  

Phantos callidryas (R. Felder, 1869)  Pale Leafwing  E & W Mexico to Panama  iN  M T S1 S2 L

Phantos opalina (Godman & Salvin, 1884)  ref.197  M T

Zaretis  Hübner, [1819]  

Zaretis delassisei X. Choimet, 2009  ref.185

Zaretis ellops (Ménétriés, 1855)  Seasonal Leafwing  [= anzuletta], E & W Mexico to Colombia & NW Venezuela  iN  M T S L

Zaretis isidora (Cramer, 1779)  Isidora Leafwing  [S Mexico] to Ecuador & Argentina  iN  M T S L

Zaretis strigosus (Gmelin, [1790])  ref.197  iN  M T

Zaretis falcis Dias, Casagrande & O. Mielke, 2012  ref.196

Zaretis crawfordhilli Dias, 2018  ref.197  iN

Zaretis elianahenrichae Dias, 2018  ref.197  iN

Zaretis hurin Dias, 2018  ref.197  iN

Zaretis mirandahenrichae Dias, 2018  ref.197  iN

Zaretis itys (Cramer, 1777)  Itys Leafwing  S Mexico (SE Chiapas) to Amazon Basin and Bolivia  iN  M T S

Zaretis itylus (Westwood, 1850)  ref.196  M T

Zaretis syene (Hewitson, 1856)    iN  M T S

Zaretis pythagoras Willmott & J. Hall, 2004    iN  M S

Anaea  Hübner, [1819]  

Anaea aidea (Guérin-Méneville, [1844])  Tropical Leafwing  S TX (strays far N), E & W Mexico to Costa Rica  iN  M T S1 S2 S3 L1 L2 I H

Anaea andria Scudder, 1875  Goatweed Leafwing  SE AZ, far N Mexico, to E CO, E to IL, N FL, strays N  iN  M T S L1 L2 L3 I1 I2 H

Anaea troglodyta (Fabricius, 1775)  Florida Leafwing    iN

Anaea troglodyta floridalis F. Johnson & W. Comstock, 1941  S FL  M T S L

Anaea troglodyta cubana (H. Druce, 1905)  Cuba & Isla de Juventud, Grand Cayman  M T S

Anaea troglodyta portia (Fabricius, 1775)  Jamaica  M T S1 S2 L

Anaea troglodyta troglodyta (Fabricius, 1775)  Hispaniola  M T S L

Anaea troglodyta borinquenalis F. Johnson & W. Comstock, 1941  Mona Island & Puerto Rico, [Culebra]  M T S L

Anaea troglodyta astina (Fabricius, 1793)  Virgin Islands (St. Thomas, St. Croix, Tortola)  M T S

Anaea troglodyta minor A. Hall, 1936  N Lesser Antilles (St. Bartholomew & St. Kitts to Guadeloupe)  M T S

Fountainea  Rydon, 1971  

Fountainea centaurus (C. Felder & R. Felder, 1867)    iN  M T S

Fountainea eurypyle (C. Felder & R. Felder, 1862)    iN

Fountainea eurypyle eurypyle (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Fountainea eurypyle confusa (A. Hall, 1929)  E Mexico to Panama  M T S L I

Fountainea eurypyle glanzi (Rotger, Escalante & Coronado, 1965)  W Mexico (SIN to S OAX)  M T S Ha

Fountainea glycerium (E. Doubleday, [1849])  Angled Leafwing    iN

Fountainea glycerium glycerium (E. Doubleday, [1849])  E & W Mexico to Panama  M T S L1 L2

Fountainea glycerium comstocki (Witt, 1972)  TL: Colombia  M T

Fountainea glycerium cratais (Hewitson, 1874)  TL: Bolivia  M T

Fountainea glycerium venezuelana (F. Johnson & W. Comstock, 1941)  TL: Venezuela  M T

Fountainea glycerium yucatanum (Witt, 1980)  Yucatan Peninsula  M OI

Fountainea johnsoni (Avinoff & Shoumatoff, 1941)  Jamaican Leafwing  Jamaica, Hispaniola  M T S

Fountainea halice (Godart, [1824])    iN

Fountainea halice halice (Godart, [1824])  TL: Brazil

Fountainea halice chrysophana (H. Bates, 1866)  Costa Rica & Panama  M T L

Fountainea halice evelina (W. Comstock, 1961)  TL: Bolivia  M T

Fountainea halice fumata (A. Hall, 1935)  TL: Venezuela  M T

Fountainea halice martinezi (J. Maza & Díaz, 1978)  SE Mexico (VER, N OAX)  M T S

Fountainea halice maya (Witt, 1980)  Yucatan Peninsula  M T

Fountainea halice moretta (H. Druce, 1877)  TL: Brazil (Pe)  M T

Fountainea halice tehuana (A. Hall, 1917)  W Mexico (JAL to S CHIS)  M T S

Fountainea nessus (Latreille, [1813])  Cindy's Leafwing    iN  M T S L

Fountainea rayoensis (J. Maza & Díaz, 1978)  Rayo Leafwing  SW Mexico (NAY to OAX)  iN  M T S

Fountainea nobilis (H. Bates, 1864)  Noble Leafwing    iN

Fountainea nobilis nobilis (H. Bates, 1864)  S Mexico (SW Chiapas) & Guatemala  M T S

Fountainea nobilis caudata (Röber, 1916)  TL: Peru  M T S

Fountainea nobilis pacifica (Vélez & Salazar, 1991)  TL: Colombia  M T

Fountainea nobilis peralta (A. Hall, 1929)  Costa Rica & Panama  M T S L

Fountainea nobilis romeroi (Descimon, 1988)  TL: Venezuela

Fountainea nobilis titan (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Fountainea ryphea (Cramer, 1775)  Ryphea Leafwing    iN

Fountainea ryphea ryphea (Cramer, 1775)  E Mexico to Amazon Basin  M T S L

Fountainea ryphea ecuadoralis (F. Johnson & W. Comstock, 1941)  TL: Ecuador  M T

Fountainea ryphea phidile (Geyer, 1837)  TL: ?  M T

Fountainea sosippus (Hopffer, 1874)    iN  M T

Fountainea tyrianthina (Salvin & Godman, 1868)  ref.295  iN  M T S

Memphis  Hübner, [1819]  

Memphis acaudata (Röber, 1916)    iN  M T

Memphis acidalia (Hübner, [1819])    iN

Memphis acidalia acidalia (Hübner, [1819])  TL: Surinam  M T

Memphis acidalia memphis (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Memphis acidalia victoria (H. Druce, 1877)  TL: Brazil (RJ)  M T

Memphis alberta (H. Druce, 1876)    iN  M T S

Memphis ambrosia (H. Druce, 1874)  Ambrosia Leafwing  TL: Panama

Memphis ambrosia ambrosia (H. Druce, 1874)  Costa Rica & Panama  M T S L

Memphis ambrosia phoebe (H. Druce, 1877)  TL: Bolivia  M T

Memphis anassa (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Memphis anna (Staudinger, 1897)  TL: Brazil (Am); Peru

Memphis anna anna (Staudinger, 1897)  TL: Brazil (Am); Peru  M T

Memphis anna elina (Staudinger, 1897)  TL: Colombia  M T S

Memphis appias (Hübner, [1825])    iN  M S

Memphis arginussa (Geyer, 1832)    iN

Memphis arginussa arginussa (Geyer, 1832)  TL: Brazil  M T S

Memphis eubaena (Boisduval, 1870)  Mottled Leafwing  SE Mexico to Panama; ref.ref.Dias_et_el_2021  M T S

Memphis onophis (C. Felder & R. Felder, 1861)  ref.199  M T S

Memphis artacaena (Hewitson, 1869)  White-patched Leafwing  SE Mexico to Colombia  iN  M T S L I

Memphis aulica (Röber, 1916)  Aulica Leafwing  Costa Rica & Panama  M T S

Memphis aureola (H. Bates, 1866)  Aureola Leafwing  E & S Mexico to Colombia  iN

Memphis aureola aureola (H. Bates, 1866)  E & S Mexico to Colombia  M T S L

Memphis aureola pueblaensis Dottax & Salazar, 2011  Colombia

Memphis basilia (Stoll, 1780)    iN

Memphis basilia basilia (Stoll, 1780)  TL: Surinam  M T

Memphis basilia bella (W. Comstock, 1961)  TL: Colombia

Memphis basilia drucei (Staudinger, 1887)  TL: [Peru]  M T

Memphis beatrix (H. Druce, 1874)  Beatrix Leafwing  Costa Rica & Panama  iN  M T S

Memphis boliviana (H. Druce, 1877)  TL: Bolivia  M T

Memphis catinka (H. Druce, 1877)  TL: ?  M T

Memphis cerealia (H. Druce, 1877)  TL: Peru  M T

Memphis cleomestra (Hewitson, 1869)    iN

Memphis cleomestra cleomestra (Hewitson, 1869)  TL: Colombia  M T

Memphis cleomestra ada (A. Butler, 1875)  Costa Rica & Panama  M T S

Memphis cluvia (Hopffer, 1874)  TL: Bolivia  M T

Memphis dia (Godman & Salvin, 1884)  Dia Leafwing    iN

Memphis dia dia (Godman & Salvin, 1884)  SE Mexico to Panama  M T S

Memphis dia divina (Röber, 1916)  TL: Bolivia  M T

Memphis falcata (Hopffer, 1874)  TL: Peru  M S

Memphis forreri (Godman & Salvin, 1884)  Forrer's Leafwing  S TX (rare vagrant), E & W Mexico to Costa Rica  iN  M T S L

Memphis glauce (C. Felder & R. Felder, 1862)    iN

Memphis glauce glauce (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T S

Memphis glauce centralis (Röber, 1916)  Honduras to Panama  M T S

Memphis glauce cicla (Möschler, 1877)  TL: Surinam  M T

Memphis glauce felderi (Röber, 1916)  TL: Ecuador  M T

Memphis grandis (H. Druce, 1877)  TL: ?  M T S

Memphis hedemanni (R. Felder, 1869)  Hedemann's Leafwing  SE Mexico, Guatemala  M T S

Memphis herbacea (A. Butler & H. Druce, 1872)  Herbacea Leafwing  SE Mexico to Costa Rica  iN  M T S

Memphis hirta (Weymer, 1907)    iN  M T

Memphis iphis (Latreille, [1813])  TL: ?  M T S

Memphis juliani Constantino, 1999  TL: Colombia  M T

Memphis laertes (Cramer, 1775)    iN  M T S

Memphis laura (H. Druce, 1877)    iN

Memphis laura laura (H. Druce, 1877)  TL: Panama  M T

Memphis laura balboa (A. Hall, 1927)  Costa Rica & Panama  M T S

Memphis laura caucana (Joicey & Talbot, 1922)  TL: Colombia  M T S

Memphis laura rosae (Fassl, 1909)  TL: Colombia  M T S

Memphis lemnos (H. Druce, 1877)  TL: Peru  M T L

Memphis leonida (Stoll, 1782)    iN

Memphis leonida leonida (Stoll, 1782)  TL: Surinam  M T S L

Memphis leonida editha (W. Comstock, 1961)  TL: Brazil (SC)  M T S

Memphis lineata (Salvin, 1869)    iN  M T

Memphis lorna (H. Druce, 1877)    iN  M T

Memphis lyceus (H. Druce, 1877)  Lyceus Leafwing  Costa Rica to Bolivia  iN  M T S

Memphis maria Pyrcz & Neild, 1996  TL: Venezuela  M T

Memphis moeris (C. Felder & R. Felder, 1867)    iN  M T

Memphis montesino Pyrcz, 1995  TL: Venezuela  M T

Memphis mora (H. Druce, 1874)    iN

Memphis mora mora (H. Druce, 1874)  TL: Colombia  M T

Memphis mora annetta (W. Comstock, 1961)  TL: Ecuador  M T

Memphis mora montana (Röber, 1916)  TL: Peru  M T S L

Memphis mora orthesia (Godman & Salvin, 1884)  SE Mexico to Panama  M T S

Memphis moruus (Fabricius, 1775)    iN

Memphis moruus moruus (Fabricius, 1775)  TL: India  M T

Memphis moruus boisduvali (W. Comstock, 1961)  E Mexico to Panama  M T S L

Memphis moruus leonila (W. Comstock, 1961)  TL: Ecuador  M T S

Memphis moruus morpheus (Staudinger, [1886])  TL: Brazil (Am); Peru  M T S L

Memphis moruus phila (H. Druce, 1877)  TL: Colombia  M T

Memphis moruus stheno (Prittwitz, 1865)  TL: Brazil (RJ)  M T

Memphis neidhoeferi (Rotger, Escalante & Coronado, 1965)  Wavy-edged Leafwing  SE Mexico to Costa Rica  M T S

Memphis nenia (H. Druce, 1877)  TL: Brazil (SP)

Memphis nenia nenia (H. Druce, 1877)  TL: Brazil (SP)  M T

Memphis nenia threnodion (Bargmann, 1929)  TL: Colombia  M T

Memphis oenomais (Boisduval, 1870)  Edge Leafwing  E Mexico through Central America  iN  M T S L

Memphis offa (H. Druce, 1877)    iN

Memphis offa offa (H. Druce, 1877)  TL: Ecuador  M T S

Memphis offa gudrun (Niepelt, 1924)  TL: Colombia  M T

Memphis otrere (Hübner, [1825])    iN  M T S

Memphis paulus Orellana & Costa, 2014  ref.188

Memphis pasibula (E. Doubleday, [1849])    iN  M T S

Memphis perenna (Godman & Salvin, 1884)  Perenna Leafwing    iN

Memphis perenna perenna (Godman & Salvin, 1884)  E & SW Mexico & Guatemala  M T S L

Memphis perenna austrina (W. Comstock, 1961)  TL: Colombia  M S

Memphis lankesteri (A. Hall, 1935)  Lankester's Leafwing  Costa Rica; ref.199  M T S

Memphis phantes (Hopffer, 1874)    iN

Memphis phantes phantes (Hopffer, 1874)  TL: Peru  M T

Memphis phantes vicinia (Staudinger, 1887)  TL: Peru  M T S

Memphis philumena (E. Doubleday, [1849])    iN

Memphis philumena philumena (E. Doubleday, [1849])  TL: Bolivia  M T

Memphis philumena chaeronea (C. Felder & R. Felder, 1861)  TL: Colombia  M T

Memphis philumena corita (Fruhstorfer, 1916)  TL: Brazil (ES)  M T S

Memphis philumena indigotica (Salvin, 1869)  Costa Rica & Panama  M T S

Memphis philumena xenica (H. Bates, 1864)  E Mexico & Guatemala  M T S

Memphis pithyusa (R. Felder, 1869)  Pale-spotted Leafwing  S TX, E & W Mexico to Bolivia; ref.199  iN  M T S L

Memphis polycarmes (Fabricius, 1775)    iN  M S

Memphis polyxo (H. Druce, 1874)    iN  M T

Memphis praxias (Hopffer, 1874)    iN

Memphis praxias praxias (Hopffer, 1874)  TL: Peru

Memphis praxias oblita (A. Hall, 1929)  TL: Ecuador  M T

Memphis schausiana (Godman & Salvin, 1894)  Great Leafwing  C Veracruz  M T S

Memphis proserpina (Salvin, 1869)  Proserpina Leafwing  SE Mexico to Costa Rica  iN  M T S

Memphis elara (Godman & Salvin, 1897)  Elara Leafwing  Costa Rica & W Panama  M T S

Memphis kingi (L. Miller & Nicolay, 1971)  King's Leafwing  Panama (Cerro Campana, Chiriqui, Canal Zone)  M T

Memphis pseudiphis (Staudinger, 1887)    iN  M T S

Memphis salinasi Pyrcz, 1993  TL: Venezuela  M T

Memphis smalli T.J. Riley & F.M.S. Dias, 2021  ref.199

Memphis verticordia (Hübner, [1831])  Hispaniolan Leafwing  Hispaniola  iN

Memphis verticordia verticordia (Hübner, [1831])  TL: Cuba  M T S

Memphis echemus (E. Doubleday, [1849])  Cuban Leafwing    iN

Memphis echemus echemus (E. Doubleday, [1849])  Cuba & Isla de Juventud; one-time stray to Texas  M T S

Memphis echemus bahamae (Witt, 1972)  Bahamas: New Providence & Andros  M T S

Memphis echemus danieliana (Witt, 1972)  Cayman Islands  M T S

Memphis intermedia (Witt, 1972)  Bahaman Leafwing    iN

Memphis intermedia intermedia (Witt, 1972)  Islands of Great Inagua Bank & Turks, Caicos  M T S

Memphis intermedia venus L. Miller, Simon & Harvey, 1992  Islands of Crooked - Acklins Bank  M T1 T2

Memphis intermedia mayaguanae L. Miller, Simon & Harvey, 1992  Mayaguana Bank  M T1 T2

Memphis dominicana (Godman & Salvin, 1884)  Dominican Leafwing    iN

Memphis dominicana dominicana (Godman & Salvin, 1884)  Dominica  M T S

Memphis dominicana luciana (A. Hall, 1929)  St. Lucia  M T S

Memphis viloriae Pyrcz & Neild, 1996  TL: Venezuela  M T

Memphis wellingi L. Miller & J. Miller, 1976  Welling's Leafwing  SW Mexico (S OAX, S CHIS)  M T S

Memphis xenippa (A. Hall, 1935)  TL: Colombia  M T

Memphis xenocles (Westwood, 1850)    iN

Memphis xenocles xenocles (Westwood, 1850)  TL: Bolivia  M T S

Memphis xenocles carolina (W. Comstock, 1961)  SE Mexico to Panama  M S L

Memphis xenocles fisilis (A. Hall, 1935)  TL: Colombia  M T

Memphis xenocles marginalis (A. Hall, 1935)  TL: Brazil (RJ/MG/SP)  M T S

Memphis xenocrates (Westwood, 1850)    iN

Memphis xenocrates xenocrates (Westwood, 1850)  ref.295  M T

Memphis xenocrates ernestoi (Salazar & Constantino, 2001)  ref.295  M S

Memphis xenocrates punctimarginale (Kaye, 1918)  ref.295  M T

  Tribe Anaemorphini

Anaeomorpha  Rothschild, 1894  

Anaeomorpha splendida Rothschild, 1894  TL: Peru

Anaeomorpha splendida splendida Rothschild, 1894  TL: Peru  M T S

Anaeomorpha splendida esmeralda Attal & M. Büche, 2008  ref.158

Anaeomorpha mirifica M. Simon & Willmott, 2017  ref.314

  Tribe Preponini

Archaeoprepona  Fruhstorfer, 1915  

Archaeoprepona chromus (Guérin-Méneville, [1844])  ref.274  iN

Archaeoprepona chromus chromus (Guérin-Méneville, [1844])  TL: Colombia  M T

Archaeoprepona chromus macaraoensis (Schäffler & Frankenbach, 2011)  TL: not specified

Archaeoprepona chromus marankiariensis (Schäffler & Frankenbach, 2011)  TL: not specified

Archaeoprepona priene (Hewitson, 1859)  ref.274  M T

Archaeoprepona licomedes (Cramer, 1777)    iN

Archaeoprepona licomedes licomedes (Cramer, 1777)  TL: Surinam  M T

Archaeoprepona licomedes pacifica Constantino & Salazar, 1998  TL: Colombia  M T

Archaeoprepona chalciope (Hübner, [1823])    iN  M T

Archaeoprepona amphimachus (Fabricius, 1775)    iN

Archaeoprepona amphimachus amphimachus (Fabricius, 1775)  TL: Indiis  M T

Archaeoprepona amphimachus amphiktion (Fruhstorfer, 1916)  E Mexico to Colombia  M T S1 S2 L I1 I2 I3

Archaeoprepona amphimachus baroni J. Maza, 1982  SW Mexico (GRO, S OAX)  M T S

Archaeoprepona amphimachus pseudomeander (Fruhstorfer, 1906)  TL: Brazil (RS)  M T

Archaeoprepona amphimachus symaithus (Fruhstorfer, 1916)  TL: Ecuador  M T

Archaeoprepona amphimachus costaricana Schäffler & Frankenbach, 2011  TL: not specified

Archaeoprepona amphimachus esmeraldica Schäffler & Frankenbach, 2011  TL: not specified

Archaeoprepona amphimachus araguaensis Schäffler & Frankenbach, 2011  TL: not specified

Archaeoprepona meander (Cramer, 1775)    iN

Archaeoprepona meander meander (Cramer, 1775)  TL: Surinam

Archaeoprepona meander castorina (E. May, 1932)  TL: Brazil (RJ)  M T

Archaeoprepona meander megabates (Fruhstorfer, 1916)  Panama to Bolivia  M T S L

Archaeoprepona meander phoebus (Boisduval, 1870)  SE Mexico to Honduras  M T S

Archaeoprepona camilla (Godman & Salvin, 1884)  Pale Prepona    iN

Archaeoprepona camilla camilla (Godman & Salvin, 1884)  Nicaragua to Panama  M T S1 S2 L I1 I2

Archaeoprepona camilla metabus (Fruhstorfer, 1916)  TL: Colombia  M T

Archaeoprepona demophon (Linnaeus, 1758)    iN

Archaeoprepona demophon demophon (Linnaeus, 1758)  TL: Indiis  M HS

Archaeoprepona demophon centralis (Fruhstorfer, 1905)  S TX (rare stray), E Mexico through Panama  M T S L I1 I2

Archaeoprepona demophon muson (Fruhstorfer, 1905)  TL: Colombia; Ecuador; Bolivia  M T

Archaeoprepona demophon occidentalis (Stoffel & Descimon, 1974)  W Mexico (SIN to S OAX)  M T S L

Archaeoprepona demophon thalpius (Hübner, [1814])  TL: ?  M T

Archaeoprepona demophoon (Hübner, [1814])    iN

Archaeoprepona demophoon demophoon (Hübner, [1814])  TL: ?  M T

Archaeoprepona demophoon andicola (Fruhstorfer, 1904)  TL: Ecuador  M T

Archaeoprepona demophoon antimache (Hübner, [1819])  TL: Indien

Archaeoprepona demophoon crassina (Fruhstorfer, 1904)  Cuba  M T S

Archaeoprepona demophoon gulina (Fruhstorfer, 1904)  E Mexico to Amazon basin  M T S1 S2 L I1 I2

Archaeoprepona demophoon insulicola (Fruhstorfer, 1897)  Hispaniola  M T S L

Archaeoprepona demophoon ramosorum K. Johnson & Descimon, 1989  Puerto Rico  M T S L I

Archaeoprepona demophoon mexicana Llorente, Descimon & K. Johnson, 1993  W Mexico (NAY to S OAX)  M T S

Archaeoprepona demophoon dominici Schäffler & Frankenbach, 2011  TL: not specified

Archaeoprepona demophoon niklasi Schäffler & Frankenbach, 2011  TL: not specified

Archaeoprepona phaedra (Godman & Salvin, 1884)  Falcate Prepona    iN

Archaeoprepona phaedra SW Mexican segregate ,  SW Mexico (GRO, S OAX)  M S

Archaeoprepona phaedra aelia (Godman & Salvin, 1889)  SE Mexico possibly to Nicaragua  M T S L

Archaeoprepona phaedra phaedra (Godman & Salvin, 1884)  Costa Rica & Panama  M T

Mesoprepona  Bonfantti, Casagrande & O. Mielke, 2017  

Mesoprepona pheridamas (Cramer, 1777)  ref.273  iN  M T

Prepona  Boisduval, 1836  

Prepona dexamenus Hopffer, 1874  Least Prepona    iN

Prepona dexamenus dexamenus Hopffer, 1874  Panama to Amazon Basin  M T S

Prepona dexamenus leuctra Fruhstorfer, 1904  TL: Brazil (ES)  M T

Prepona dexamenus medinai Beutelspacher, 1981  SE Mexico to Honduras  M T S

Prepona laertes (Hübner, [1811])    iN

Prepona laertes laertes (Hübner, [1811])  TL: ?  M T

Prepona laertes demodice (Godart, [1824])  TL: French Guiana; Brazil  M T1 T2 T3 T4

Prepona laertes louisa A. Butler, 1870  TL: Cuba  M T

Prepona laertes octavia Fruhstorfer, 1905  E & W Mexico through Central America, recorded introduction to USA: Florida  M T1 T2 S1 S2 L

Prepona werneri M. Hering & Hopp, 1925    iN  M S

Prepona pylene Hewitson, [1854]    iN

Prepona pylene pylene Hewitson, [1854]  TL: Brazil (RJ)  M T

Prepona eugenes H. Bates, 1865  TL: Brazil (Pa)

Prepona eugenes eugenes H. Bates, 1865  TL: Brazil (Pa)  M T S

Prepona eugenes bahiana Fruhstorfer, 1897  TL: Brazil (Ba)  M T

Prepona eugenes laertides Staudinger, 1898  TL: Bolivia  M T

Prepona gnorima H. Bates, 1865  Mottled Prepona  Honduras to Colombia  iN

Prepona gnorima gnorima H. Bates, 1865  Honduras to Colombia  M T S

Prepona gnorima jordani Fruhstorfer, 1905  TL: Ecuador  M T S

Prepona gnorima philetas Fruhstorfer, 1904  SE Mexico to Honduras  M T S

Prepona deiphile (Godart, [1824])    iN

Prepona deiphile Nayarit segregate ,  Sierra de San Juan, Nayarit, Mexico  M S L

Prepona deiphile lambertoana Llorente, Luis & F. González, 1992  Michoacán, Estado de Mexico  M T1 T2 S

Prepona deiphile brooksiana Godman & Salvin, 1889  E Mexico (SLP to N OAX)  M T TI S L I

Prepona deiphile escalantiana Stoffel & Mast, 1973  SE Mexico (vic. Los Tuxtlas, Veracruz)  M S1 S2

Prepona deiphile NW Chiapas segregate ,  NW Chiapas  M S

Prepona deiphile diaziana L. Miller & J. Miller, 1976  E Chiapas  M T S

Prepona deiphile salvadora Llorente, Luis & F. González, 1992  El Salvador  M T S

Prepona deiphile lygia Fruhstorfer, 1904  Costa Rica & Panama  M T S

Prepona deiphile photidia Fruhstorfer, 1912  TL: Colombia

Prepona deiphile sphacteria Fruhstorfer, 1916  TL: Peru

Prepona deiphile neoterpe Honrath, 1884  TL: Peru  M T

Prepona deiphile deiphile (Godart, [1824])  TL: Brazil

Prepona deiphile xenagoras Hewitson, 1875  TL: Bolivia  M T

Prepona ibarra Beutelspacher, 1982  Guerrero, SW Mexico  M T S

Prepona silvana Turrent & García, 2019  Turrent & García (2019, Revista de la Sociedad Mexicana de Lepidopterolog ...

Prepona praeneste Hewitson, 1859    iN

Prepona praeneste praeneste Hewitson, 1859  TL: Colombia  M T

Prepona praeneste abrupta Biedermann, 1936  TL: Peru  M T

Prepona praeneste buckleyana Hewitson, 1876  TL: Bolivia  M T

Prepona praeneste choroniensis Lichy, 1975  TL: Venezuela  M T

Prepona praeneste confusa Niepelt, 1913  TL: Peru  M T

Prepona praeneste isabelae Orellana, 2000  TL: Venezuela

Prepona praeneste virago Biedermann, 1938  TL: Colombia  M T S

Prepona aedon (Hewitson, 1848)    iN

Prepona aedon aedon (Hewitson, 1848)  ref.274  M T

Prepona aedon pepitoensis (Michael, 1930)  ref.274  M T

Prepona aedon rodriguezi (Schaus, 1918)  S Mexico (SE Chiapas) to Costa Rica; ref.274  M T S L

Prepona amydon (Hewitson, [1854])    iN

Prepona amydon amydon (Hewitson, [1854])  ref.274  M T S

Prepona amydon amydonius (Staudinger, [1885])  ref.274  M T

Prepona amydon aristoxenus (Niepelt, 1913)  ref.274  M T

Prepona amydon aurantiaca (Fruhstorfer, 1897)  ref.274  M T

Prepona amydon bogotana (Fruhstorfer, 1895)  ref.274  M T

Prepona amydon boliviensis (Fruhstorfer, 1895)  ref.274  M T

Prepona amydon excelsior (Lathy, 1924)  ref.274  M T

Prepona amydon ferdinandi (Fruhstorfer, 1895)  ref.274  M T

Prepona amydon frontina (Fruhstorfer, 1895)  ref.274  M T

Prepona amydon lacandona (R. G. Maza & J. Maza, 1999)  SE Mexico (SE Chiapas) to Honduras; ref.274  M T S

Prepona amydon oaxacata (Kruck, 1931)  S Mexico (N OAX) to Guatemala; ref.274  M T S L

Prepona amydon phalcidon (Hewitson, 1855)  ref.274  M T1 T2 T3 T4 T5 T6

Prepona amydon philatelica (DeVries, 1980)  Nicaragua to Panama; ref.274  M T S L I1 I2

Prepona amydon rubella (Michael, 1930)  ref.274  M T

Prepona amydon smalli (L. Miller & Nicolay, 1971)  Panama (vic. Canal Zone); ref.274  M T S

Prepona amydon uniformis (Michael, 1929)  ref.274  M T

Prepona amydon zenodorus (Hewitson, 1870)  ref.274  M T

Prepona amydon [n. ssp.] (Neild, MS)  Venezuela

Prepona hewitsonius (H. Bates, 1860)    iN

Prepona hewitsonius hewitsonius (H. Bates, 1860)  ref.274  M T

Prepona hewitsonius beata (Staudinger, [1885])  ref.274  M T

Prepona hewitsonius beatifica (Hewitson, 1869)  ref.274  M T

Prepona hewitsonius stuarti (Godman & Salvin, 1882)  ref.274  M T S

Prepona claudina (Godart, [1824])    iN

Prepona claudina claudina (Godart, [1824])  ref.274  M T1 T2 T3 T4 T5 S

Prepona claudina annetta (G. Gray, 1832)  ref.274  M T

Prepona claudina croesus (Staudinger, 1896)  ref.274  M T

Prepona claudina delavillae (Neild, 1996)  ref.274

Prepona claudina godmani (Fruhstorfer, 1895)  ref.274  M T

Prepona claudina lugens (Staudinger, 1886)  ref.274  M T S

Prepona claudina patriciae (Attal, 2000)  ref.274

Prepona claudina sardanapalus (H. Bates, 1860)  ref.274  M T1 T2

Prepona narcissus (Staudinger, [1885])    iN

Prepona narcissus narcissus (Staudinger, [1885])  ref.274  M T

Prepona narcissus stoffeli (Mast & Descimon, 1972)  ref.274

Prepona narcissus tapajonus (Fassl, 1921)  ref.274  M T1 T2

  Subfamily Satyrinae

  Tribe Morphini

Antirrhea  Hübner, [1822]  

Antirrhea adoptiva (Weymer, 1909)    iN

Antirrhea adoptiva adoptiva (Weymer, 1909)  TL: Colombia  M T

Antirrhea adoptiva porphyrosticta (Watkins, 1928)  TL: Ecuador  M T

Antirrhea adoptiva [n. ssp.#1] Orellana, MS  TL: Venezuela

Antirrhea adoptiva [n. ssp.#2] Lamas, MS  TL: Peru

Antirrhea archaea Hübner, [1822]    iN

Antirrhea geryon (C. Felder & R. Felder, 1862)    iN

Antirrhea geryon geryon (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Antirrhea geryon geryonides (Weymer, 1909)  TL: Ecuador  M T

Antirrhea geryon weymeri Salazar, Constantino & López, 1998  TL: Colombia  M T S

Antirrhea hela (C. Felder & R. Felder, 1862)    iN  M T

Antirrhea ornata (A. Butler, 1870)    iN  M T

Antirrhea phasiana (A. Butler, 1870)    iN  M T S

Antirrhea philoctetes (Linnaeus, 1758)    iN

Antirrhea philoctetes philoctetes (Linnaeus, 1758)  TL: Indiis  M T HS

Antirrhea philoctetes murena (Staudinger, [1886])  TL: Brazil (Am)  M T S

Antirrhea philaretes (C. Felder & R. Felder, 1862)    iN

Antirrhea philaretes casta (H. Bates, 1865)  SE Mexico, Guatemala  M T S

Antirrhea philaretes tomasia (A. Butler, 1875)  Costa Rica & Panama (Pacific slope)  M T S L

Antirrhea philaretes philaretes (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Antirrhea philaretes lindigii (C. Felder & R. Felder, 1862)  [= miltiades], Guatemala to Amazon Basin; Atlantic slope  M T S1 S2 L I1 I2

Antirrhea philaretes intermedia Salazar, Constantino & López, 1998  TL: Colombia  M T

Antirrhea philaretes avernus (Hopffer, 1874)  TL: Peru  M T

Antirrhea philaretes theodori (Fruhstorfer, 1907)  TL: Brazil (Am)  M T

Antirrhea philaretes lumejia Orellana, 2004  TL: Venezuela

Antirrhea philaretes piaroa Orellana & Neild, 2008  ref.266

Antirrhea ulei Strand, 1912    iN

Antirrhea pterocopha (Salvin & Godman, 1868)  Brilliant Antirrhea    iN

Antirrhea pterocopha pterocopha (Salvin & Godman, 1868)  Costa Rica & W Panama (Atlantic slope; Carrillo Belt)  M T S

Antirrhea pterocopha chocoensis Salazar, 1997  TL: Colombia  M T S

Antirrhea taygetina (A. Butler, 1868)    iN

Antirrhea taygetina taygetina (A. Butler, 1868)  TL: Brazil (Am)  M T S

Antirrhea taygetina rodwayi (A. Hall, 1939)  TL: Guyana  M T

Antirrhea taygetina [n. ssp.] Vane-Wright, MS  Ecuador  M S

Antirrhea undulata (M. Hering & Hopp, 1925)    iN  M T

Antirrhea watkinsi Rosenberg & Talbot, 1914    iN

Antirrhea watkinsi watkinsi Rosenberg & Talbot, 1914  TL: Peru  M T S

Antirrhea watkinsi callephebus (Tessmann, 1928)  TL: Peru  M T

Caerois  Hübner, [1819]  

Caerois chorinaeus (Fabricius, 1775)    iN

Caerois chorinaeus chorinaeus (Fabricius, 1775)  TL: Surinam  M T HS

Caerois chorinaeus protonoe Fruhstorfer, 1912  TL: Peru  M T

Caerois chorinaeus rufomarginata Lathy, 1918  TL: Peru  M T

Caerois chorinaeus [n. ssp.] Lamas, MS  Peru

Caerois gerdrudtus (Fabricius, 1793)  False Antirrhea  Costa Rica to W Ecuador  iN  M S L I1 I2

Morpho  Fabricius, 1807  

Morpho absoloni E. May, 1924  TL: Brazil (Ac)  M T

Morpho achilles (Linnaeus, 1758)    iN

Morpho achilles achilles (Linnaeus, 1758)  TL: America  M T S HS

Morpho achilles agamedes Fruhstorfer, 1912  TL: Peru  M T

Morpho achilles fischeri Weber, 1962  TL: Peru  M T

Morpho achilles glaisi Le Moult & Réal, 1962  TL: Venezuela  M T

Morpho achilles guaraunos Le Moult, 1925  TL: Venezuela  M T

Morpho achilles patroclus C. Felder & R. Felder, 1861  TL: Colombia  M T

Morpho achilles phokylides Fruhstorfer, 1912  TL: Bolivia  M T

Morpho achilles vitrea A. Butler, 1866  TL: Bolivia  M T

Morpho aega (Hübner, [1822])    iN

Morpho aega aega (Hübner, [1822])  TL: ?  M T Ab

Morpho aega amargosensis Otero & Soares, 1996  TL: Brazil (Ba)  M T

Morpho amphitryon Staudinger, 1887    iN

Morpho amphitryon amphitryon Staudinger, 1887  TL: Peru  M T

Morpho amphitryon cinereus Duchêne, 1985  TL: Peru  M T

Morpho amphitryon susarion Fruhstorfer, 1913  TL: Bolivia  M T

Morpho anaxibia (Esper, [1801])    iN  M T

Morpho athena Otero, 1966    iN  M T

Morpho aurora Westwood, 1851    iN

Morpho aurora aurora Westwood, 1851  TL: Bolivia  M T

Morpho aurora aureola Fruhstorfer, 1913  TL: Peru

Morpho cisseis C. Felder & R. Felder, 1860    iN

Morpho cisseis cisseis C. Felder & R. Felder, 1860  TL: Brazil (Am)  M T

Morpho cisseis gahua Blandin, 1988  TL: Peru  M T

Morpho cisseis phanodemus Hewitson, 1869  TL: Ecuador  M T

Morpho cypris Westwood, 1851    iN

Morpho cypris cypris Westwood, 1851  TL: Colombia  M T

Morpho cypris aphrodite Le Moult & Réal, 1962  Honduras, Nicaragua  M T S

Morpho cypris bugaba Staudinger, 1887  Costa Rica & Panama  M T S

Morpho cypris chrysonicus Fruhstorfer, 1913  TL: Venezuela  M T

Morpho cypris limonensis Schäffler, Frankenbach & Dufek, 2011  TL: not specified

Morpho cypris ceibaensis Schäffler, Frankenbach & Dufek, 2011  TL: not specified

Morpho deidamia (Hübner, [1819])    iN

Morpho deidamia deidamia (Hübner, [1819])  TL: Surinam  M T HS

Morpho deidamia diffusa Le Moult & Réal, 1962  TL: Brazil (Ma)  M T

Morpho deidamia diomedes Weber, 1944  TL: [Peru]

Morpho deidamia electra Röber, 1903  TL: Bolivia  M T

Morpho deidamia granadensis C. Felder & R. Felder, 1867  Panama to Colombia  M T S L

Morpho deidamia guaraura Le Cerf, 1925  TL: Venezuela  M T

Morpho deidamia hermione Röber, 1903  TL: Colombia  M T

Morpho deidamia jacki Neild, 2008  ref.266

Morpho deidamia lycanor Fruhstorfer, 1907  TL: Ecuador  M T

Morpho deidamia neoptolemus Wood, 1863  TL: [Brazil (Am)]  M T

Morpho deidamia polybaptus A. Butler, 1875  Nicaragua to Panama; Atlantic slope  M T S

Morpho deidamia pyrrhus Staudinger, 1887  TL: Peru  M T

Morpho deidamia steinbachi Le Moult & Réal, 1962  TL: Bolivia  M T

Morpho epistrophus (Fabricius, 1796)    iN

Morpho epistrophus epistrophus (Fabricius, 1796)  TL: Brazil (RJ)  M T

Morpho epistrophus argentinus Fruhstorfer, 1907  TL: Argentina  M T

Morpho epistrophus catenaria Perry, 1811  TL: Brazil  M T

Morpho epistrophus titei Le Moult & Réal, 1962  TL: Paraguay  M T

Morpho eugenia Deyrolle, 1860    iN  M T

Morpho hecuba (Linnaeus, 1771)    iN

Morpho hecuba hecuba (Linnaeus, 1771)  TL: French Guiana  M S

Morpho hecuba obidonus Fruhstorfer, 1905  TL: Brazil (Pa)  M T

Morpho hecuba polyidos Fruhstorfer, 1912  TL: Venezuela  M T

Morpho hecuba werneri Hopp, 1921  TL: Colombia  M T

Morpho helenor (Cramer, 1776)    iN

Morpho helenor helenor (Cramer, 1776)  TL: Surinam  M T

Morpho helenor achillaena (Hübner, [1823])  TL: ?  M T1 T2

Morpho helenor achillides C. Felder & R. Felder, 1867  TL: Brazil  M T1 T2

Morpho helenor anakreon Fruhstorfer, 1910  TL: Brazil (Pe)  M T

Morpho helenor charapensis Le Moult & Réal, 1962  TL: Peru  M T

Morpho helenor coelestis A. Butler, 1866  TL: Brazil  M T

Morpho helenor cortone Fruhstorfer, 1913  TL: Colombia  M T

Morpho helenor corydon Guenée, 1859  TL: "Colombia"; Venezuela  M T

Morpho helenor guerrerensis Le Moult & Réal, 1962  SW Mexico (MICH to S OAX)  M T S L

Morpho helenor insularis Fruhstorfer, 1912  TL: Trinidad  M T

Morpho helenor leontius C. Felder & R. Felder, 1867  TL: Colombia  M T

Morpho helenor macrophthalmus Fruhstorfer, 1913  TL: Colombia  M T

Morpho helenor maculata Röber, 1903  TL: Ecuador  M T

Morpho helenor marajoensis Le Moult & Réal, 1962  TL: Brazil (Pa)  M T

Morpho helenor marinita A. Butler, 1872  SW Costa Rica to Panama (Pacific slope)  M T S L I

Morpho helenor montezuma Guenée, 1859  E Mexico to Honduras  M T S L

Morpho helenor narcissus Staudinger, 1887  Nicaragua to Panama  M T S L I

Morpho helenor octavia H. Bates, 1864  SW Mexico (Chiapas) to El Salvador  M T S L

Morpho helenor packeri Neild, 2008  ref.266

Morpho helenor papirius Hopffer, 1874  TL: Peru  M T

Morpho helenor peleides Kollar, 1850  Panama to Colombia  M T S L

Morpho helenor peleus Röber, 1903  TL: Venezuela  M T

Morpho helenor pindarus Fruhstorfer, 1910  TL: Brazil (MT)  M T

Morpho helenor popilius Hopffer, 1874  TL: "Bolivia"; "Venezuela"; [Colombia]  M T

Morpho helenor rugitaeniatus Fruhstorfer, 1907  TL: Ecuador  M T

Morpho helenor telamon Röber, 1903  TL: Colombia  M T

Morpho helenor theodorus Fruhstorfer, 1907  TL: Brazil (Am)  M T

Morpho helenor tucupita Le Moult, 1925  TL: Venezuela  M T

Morpho helenor ululina Le Moult & Réal, 1962  TL: Venezuela  M T

Morpho helenor violaceus Fruhstorfer, 1912  TL: Brazil (SC)  M T

Morpho helenor zonaras Fruhstorfer, 1912  Panama  M T

Morpho helenor prometa Gareca & Blandin, 2011  TL: not specified

Morpho helenor coibaensis Jongerling, 2011  TL: not specified

Morpho helenor caldarai Schäffler & Frankenbach, 2011  TL: not specified

Morpho hercules (Dalman, 1823)    iN  M T

Morpho lympharis A. Butler, 1873    iN

Morpho lympharis lympharis A. Butler, 1873  TL: Peru  M T

Morpho lympharis descimokoenigi Blandin, 1993  TL: Peru  M T

Morpho lympharis eros Staudinger, 1892  TL: Bolivia  M T

Morpho lympharis stoffeli Le Moult & Réal, 1962  TL: Peru  M T

Morpho marcus (Schaller, 1785)    iN

Morpho marcus marcus (Schaller, 1785)  TL: ?  M T

Morpho marcus intermedia Kaye, 1917  TL: Peru  M T

Morpho marcus major Lathy, 1905  TL: Peru  M T

Morpho menelaus (Linnaeus, 1758)    iN

Morpho menelaus menelaus (Linnaeus, 1758)  TL: [Surinam]  M T HS

Morpho menelaus alexandrovna H. Druce, 1874  TL: Peru  M T

Morpho menelaus argentiferus Fruhstorfer, 1913  TL: Peru  M T

Morpho menelaus assarpai Röber, 1903  TL: Peru  M T

Morpho menelaus coeruleus (Perry, 1810)  TL: ?  M T

Morpho menelaus didius Hopffer, 1874  TL: Peru  M T1 T2

Morpho menelaus eberti Weber, 1963  TL: Brazil (Pe)  M T

Morpho menelaus godartii Guérin-Méneville, [1844]  TL: Bolivia  M T

Morpho menelaus titogilberti Blandin & Gareca, 2011  TL: not specified

Morpho menelaus julanthiscus Fruhstorfer, 1907  TL: Ecuador  M T

Morpho menelaus kesselringi H. Fischer, 1962  TL: Brazil (Pa)  M T

Morpho menelaus laurellae Neild, 2008  ref.266

Morpho menelaus neildi Blandin, 2008  ref.266

Morpho menelaus occidentalis C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Morpho menelaus orinocensis Le Moult, 1925  TL: Venezuela  M T

Morpho menelaus terrestris A. Butler, 1866  TL: Brazil (Am)  M T

Morpho menelaus verae Weber, 1951  TL: Brazil (Pa)  M T

Morpho menelaus zischkai H. Fischer, 1962  TL: Bolivia  M T

Morpho amathonte Deyrolle, 1860  Amathonte Morpho  Nicaragua to Panama; ref.168  M T S L

Morpho niepelti Röber, 1927    iN  M T

Morpho polyphemus Westwood, [1850]  White Morpho    iN

Morpho polyphemus polyphemus Westwood, [1850]  SE AZ (rare stray), W Mexico to El Salvador  M T S1 S2 L I

Morpho polyphemus catalina Corea & I. Chacón, 1984  Honduras to NW Costa Rica  M T S I

Morpho polyphemus luna A. Butler, 1869  SE Mexico to Guatemala  M T S L

Morpho portis (Hübner, [1821])    iN

Morpho portis portis (Hübner, [1821])  TL: ?  M T

Morpho portis thamyris C. Felder & R. Felder, 1867  TL: Brazil  M T

Morpho rhetenor (Cramer, 1775)    iN

Morpho rhetenor rhetenor (Cramer, 1775)  TL: Surinam  M T S

Morpho rhetenor cacica Staudinger, 1876  TL: Peru  M T

Morpho rhetenor columbianus E. Krüger, 1925  TL: Colombia  M T1 T2

Morpho rhetenor equatenor Le Moult & Réal, 1962  TL: Ecuador  M T

Morpho rhetenor hightoni Neild, 2008  ref.266

Morpho rhetenor helena Staudinger, 1890  TL: Peru  M T

Morpho rhetenor subtusmurina Le Moult & Réal, 1962  TL: Brazil (MT)  M T

Morpho rhodopteron Godman & Salvin, 1880    iN  M T

Morpho richardus Fruhstorfer, 1898  TL: Brazil (MG)  M T

Morpho sulkowskyi Kollar, 1850    iN

Morpho sulkowskyi sulkowskyi Kollar, 1850  TL: Colombia  M T

Morpho sulkowskyi hoppiana Niepelt, 1923  TL: Colombia  M T

Morpho sulkowskyi selenaris Le Moult & Réal, 1962  TL: Peru  M T

Morpho sulkowskyi sirene Niepelt, 1911  TL: Ecuador  M T

Morpho sulkowskyi [n. ssp.#1] Lamas, MS  TL: Peru  M T

Morpho sulkowskyi [n. ssp.#2] Lamas, MS  TL: Peru

Morpho telemachus (Linnaeus, 1758)    iN

Morpho telemachus telemachus (Linnaeus, 1758)  TL: [Surinam]  M T

Morpho telemachus iphiclus C. Felder & R. Felder, 1862  TL: Colombia  M T

Morpho telemachus lilianae Le Moult, 1927  TL: Venezuela  M T

Morpho telemachus [n. ssp.] Ebert, MS  Brazil (Pe)

Morpho justitiae Salvin & Godman, 1868  Stub-tailed Morpho  SE Mexico (E OAX, far SE VER, N & C CHIS) to Guatemala & Honduras; ref.144

Morpho justitiae schweizeri (R. F. Maza, 1987)  SE Mexico; vic. Los Tuxtlas, SE Veracruz; ref.144  M T S

Morpho justitiae oaxacensis Le Moult & Réal, 1962  SE Mexico (N Oaxaca); ref.144  M T S

Morpho justitiae neoescalantei Schäffler, 2009  ref.144

Morpho justitiae mercedes J. Maza, 2023  ref.144

Morpho justitiae justitiae Salvin & Godman, 1868  SE Mexico (E OAX, far SE VER, N & C CHIS) to Guatemala & Honduras; ref.144  M T S L

Morpho malintzin R. G. Maza & J. Maza, 2023  ref.144

Morpho aquarius A. Butler, 1872  ref.144

Morpho aquarius nebulosus R. G. Maza, 2023  ref.144

Morpho aquarius monica J. Maza, 2023  ref.144

Morpho aquarius aquarius A. Butler, 1872  ref.144  M T

Morpho theseus Deyrolle, 1860    iN

Morpho theseus theseus Deyrolle, 1860  TL: Colombia  M T

Morpho theseus claritae Neild, 2008  ref.266

Morpho theseus heraldica Niepelt, 1927  Costa Rica & Panama  M T S

Morpho theseus juturna A. Butler, 1870  TL: Colombia  M T

Morpho theseus pacificus E. Krüger, 1925  TL: Colombia  M T

Morpho theseus perlmani Neild, 2001  TL: Venezuela  M T

Morpho theseus triangulifera Le Moult & Réal, 1962  TL: Ecuador  M T

Morpho theseus yaritanus Fruhstorfer, 1913  TL: Venezuela  M T

Morpho theseus [n. ssp.] Llorente & Luis, MS  Mexico

Morpho theseus chirripoensis Schäffler, Frankenbach & Dufek, 2011  TL: not specified

Morpho uraneis H. Bates, 1865    iN  M T

Morpho zephyritis A. Butler, 1873  TL: Peru  M T

  Tribe Brassolini

Bia  Hübner, [1819]  

Bia actorion (Linnaeus, 1763)    iN

Bia actorion actorion (Linnaeus, 1763)  TL: Indiis  M HS

Bia actorion decaerulea Weymer, 1911  TL: [Brazil (Am)]  M T

Bia actorion rebeli Bryk, 1953  TL: Peru  M T S

Bia peruana Röber, 1904  TL: [Peru]  M T

Blepolenis  Röber, 1906  

Blepolenis bassus (C. Felder & R. Felder, 1867)    iN  M T

Blepolenis batea (Hübner, [1821])    iN

Blepolenis batea batea (Hübner, [1821])  TL: [Brazil]  M T

Blepolenis batea didymaon (C. Felder & R. Felder, 1867)  TL: Brazil  M T

Blepolenis batea praegrandis (Fruhstorfer, 1907)  TL: Paraguay  M T

Blepolenis catharinae (Stichel, 1902)    iN  M T

Brassolis  Fabricius, 1807  

Brassolis astyra Godart, [1824]    iN

Brassolis astyra astyra Godart, [1824]  TL: Brazil  M T

Brassolis astyra philocala Stichel, 1904  TL: Brazil (RS)  M T

Brassolis haenschi Stichel, 1902    iN

Brassolis haenschi haenschi Stichel, 1902  TL: Ecuador

Brassolis haenschi maritimus Stichel, 1903  TL: Venezuela  M T

Brassolis haenschi rufescens Rothschild, 1916  TL: Peru  M T

Brassolis isthmia H. Bates, 1864  Small-spotted Owl-Butterfly  Guatemala to Colombia; Atlantic slope  iN  M T S

Brassolis granadensis Stichel, 1902  Colombian Owl-Butterfly    iN

Brassolis granadensis granadensis Stichel, 1902  NW Costa Rica to W Ecuador; Pacific slope  M T S I

Brassolis granadensis boliviana Rothschild, 1916  TL: Bolivia  M T

Brassolis sophorae (Linnaeus, 1758)    iN

Brassolis sophorae sophorae (Linnaeus, 1758)  TL: America  M T HS

Brassolis sophorae ardens Stichel, 1903  TL: Peru  M T

Brassolis sophorae dinizi R.F. d'Almeida, 1956  TL: Brazil (Pb)  M T

Brassolis sophorae laurentii Stichel, 1925  TL: Brazil (MG)  M T

Brassolis sophorae luridus Stichel, 1902  TL: Colombia  M T

Brassolis sophorae philomela Stichel, 1925  TL: Ecuador  M T

Brassolis sophorae vulpeculus Stichel, 1902  TL: Paraguay  M T

Caligo  Hübner, [1819]  

Caligo arisbe Hübner, [1822]    iN

Caligo arisbe arisbe Hübner, [1822]  TL: ?  M T

Caligo arisbe fulgens Rothschild, 1916  TL: Brazil (RJ/SP)  M T

Caligo atreus (Kollar, 1850)    iN

Caligo atreus atreus (Kollar, 1850)  TL: Colombia  M T

Caligo atreus agesilaus H. Druce, 1902  TL: Ecuador  M T

Caligo atreus ajax (E. Doubleday, [1849])  TL: Venezuela; "Ecuador"  M T

Caligo atreus dentina H. Druce, 1874  TL: Peru  M T

Caligo atreus dionysos Fruhstorfer, 1912  [Chiapas?], Costa Rica & Panama  M T S1 S2 L I1 I2

Caligo bellerophon Stichel, 1903    iN  M T

Caligo beltrao (Illiger, 1801)    iN  M S

Caligo brasiliensis (C. Felder, 1862)    iN

Caligo brasiliensis brasiliensis (C. Felder, 1862)  TL: Brazil (RJ)  M T

Caligo brasiliensis caesius Stichel, 1903  TL: Venezuela

Caligo brasiliensis galba (Deyrolle, 1874)  TL: Colombia  M T

Caligo brasiliensis minor Kaye, 1904  TL: Trinidad  M T

Caligo brasiliensis morpheus Stichel, 1903  TL: Ecuador; Colombia  M T

Caligo brasiliensis sulanus Fruhstorfer, 1904  SE Mexico to Panama  M T S1 S2 L I1 I2

Caligo euphorbus (C. Felder & R. Felder, 1862)    iN

Caligo euphorbus euphorbus (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T

Caligo euphorbus menoetius Staudinger, 1887  TL: Brazil (Am)  M T

Caligo eurilochus (Cramer, 1775)    iN

Caligo eurilochus eurilochus (Cramer, 1775)  TL: Surinam  M T

Caligo eurilochus delectans Joicey & Kaye, 1917  TL: Venezuela  M T

Caligo eurilochus livius Staudinger, [1886]  TL: "Panama"; Peru; Brazil (Pa)  M T

Caligo eurilochus mattogrossensis Ribeiro, 1931  TL: Brazil (Rd)  M T

Caligo eurilochus pallidus Fruhstorfer, 1912  TL: Bolivia  M T

Caligo idomeneus (Linnaeus, 1758)    iN

Caligo idomeneus idomeneus (Linnaeus, 1758)  TL: S. Amer.  M T HS

Caligo idomeneus ariphron Fruhstorfer, 1910  TL: Brazil (ES)  M T

Caligo idomeneus idomenides Fruhstorfer, 1903  TL: Peru  M T

Caligo idomeneus rhoetus Staudinger, [1886]  TL: Brazil (Pa/Am)  M T

Caligo illioneus (Cramer, 1775)    iN

Caligo illioneus illioneus (Cramer, 1775)  TL: Surinam  M T

Caligo illioneus oberon A. Butler, 1870  Costa Rica & Panama  M T S1 S2 L I

Caligo illioneus pampeiro Fruhstorfer, 1904  TL: Paraguay  M T

Caligo illioneus pheidriades Fruhstorfer, 1912  TL: Bolivia  M T

Caligo illioneus praxsiodus Fruhstorfer, 1912  TL: Peru  M T

Caligo martia (Godart, [1824])    iN

Caligo oberthurii (Deyrolle, 1872)    iN

Caligo oberthurii oberthurii (Deyrolle, 1872)  TL: Colombia  M T

Caligo oberthurii phokilides Fruhstorfer, 1912  TL: Peru

Caligo oedipus Stichel, 1903  Fruhstorfer's Owl-Butterfly    iN

Caligo oedipus oedipus Stichel, 1903  Panama to Colombia  M T S1 S2 L I

Caligo oedipus fruhstorferi Stichel, 1904  SE Mexico to Honduras  M T S L

Caligo oileus C. Felder & R. Felder, 1861    iN

Caligo oileus oileus C. Felder & R. Felder, 1861  TL: Venezuela; Colombia  M T

Caligo oileus scamander (Boisduval, 1870)  Guatemala to Panama  M T S

Caligo oileus umbratilis Stichel, 1903  TL: Peru  M T

Caligo placidianus Staudinger, 1887    iN  M T S

Caligo prometheus (Kollar, 1850)    iN

Caligo prometheus prometheus (Kollar, 1850)  TL: Colombia  M T

Caligo prometheus atlas Röber, 1904  TL: Ecuador  M T

Caligo superbus Staudinger, 1887    iN

Caligo superbus superbus Staudinger, 1887  TL: Peru  M T S

Caligo superbus agamemnon Weymer, 1890  TL: Ecuador  M T

Caligo suzanna (Deyrolle, 1872)    iN  M T

Caligo telamonius (C. Felder & R. Felder, 1862)    iN

Caligo telamonius telamonius (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Caligo telamonius memnon (C. Felder & R. Felder, 1867)  E & SW Mexico to Nicaragua  M T S L I

Caligo telamonius menus Fruhstorfer, 1903  Costa Rica to Colombia  M T S1 S2 L I1 I2

Caligo telamonius modestus Niepelt, 1934  TL: Surinam

Caligo teucer (Linnaeus, 1758)    iN

Caligo teucer teucer (Linnaeus, 1758)  TL: America  M T HS

Caligo teucer ecuadora Joicey & Kaye, 1917  TL: Ecuador  M T L

Caligo teucer insulanus Stichel, 1904  TL: Trinidad  M T

Caligo teucer japetus Stichel, 1903  TL: Paraguay  M T

Caligo teucer nubilus Fruhstorfer, 1907  TL: Guyana  M T

Caligo teucer obidonus Fruhstorfer, 1904  TL: Brazil (Pa)  M T

Caligo teucer phorkys Fruhstorfer, 1912  TL: Bolivia  M T

Caligo teucer semicaerulea Joicey & Kaye, 1917  TL: Peru  M T

Caligo uranus Herrich-Schäffer, 1850  Gold-edged Owl-Butterfly  E Mexico to Honduras  iN  M T S L

Caligo zeuxippus H. Druce, 1902    iN

Caligo zeuxippus zeuxippus H. Druce, 1902  TL: Ecuador  M T

Caligo zeuxippus obscurus Niepelt, 1934  TL: Colombia  M T

Caligopsis  Seydel, 1924  

Caligopsis seleucida (Hewitson, 1877)    iN

Caligopsis seleucida seleucida (Hewitson, 1877)  TL: Bolivia  M T

Caligopsis seleucida juruana (Fruhstorfer, 1912)  TL: Brazil (Ac)  M T

Caligopsis [n. sp.] Lamas, MS  Peru  M T

Catoblepia  Stichel, 1901  

Catoblepia amphirhoe (Hübner, [1825])    iN  M T

Catoblepia berecynthia (Cramer, 1777)    iN

Catoblepia berecynthia berecynthia (Cramer, 1777)  TL: Surinam  M T S

Catoblepia berecynthia adjecta Stichel, 1906  TL: Bolivia  M T S

Catoblepia berecynthia berecynthina (Hopffer, 1874)  TL: Peru  M T

Catoblepia berecynthia halli Bristow, 1981  TL: Guyana  M T

Catoblepia berecynthia luxuriosus Stichel, 1902  TL: Colombia  M T

Catoblepia berecynthia meridaensis Bristow, 1981  TL: Venezuela

Catoblepia berecynthia midas Stichel, 1908  TL: Ecuador  M T

Catoblepia berecynthia unditaenia Fruhstorfer, 1907  TL: Brazil (RJ)  M T

Catoblepia generosa Stichel, 1902  TL: Ecuador  M T

Catoblepia orgetorix (Hewitson, 1870)  Orange-rimmed Owl-Butterfly    iN

Catoblepia orgetorix orgetorix (Hewitson, 1870)  Honduras & Nicaragua  M T TI

Catoblepia orgetorix championi Bristow, 1981  Nicaragua to Panama; Atlantic slope  M T S1 S2 L I1 I2

Catoblepia orgetorix magnalis Stichel, 1902  TL: Ecuador  M T

Catoblepia orgetorix rothschildi Casagrande & Lamas, 2004  TL: Colombia  M T

Catoblepia soranus (Westwood, 1851)    iN  M T

Catoblepia versitincta Stichel, 1901    iN

Catoblepia versitincta versitincta Stichel, 1901  TL: Surinam; French Guiana  M T

Catoblepia versitincta mossi Bristow, 1981  TL: Brazil (Pa)  M T

Catoblepia xanthicles (Godman & Salvin, 1881)  Xanthicles Owl-Butterfly    iN

Catoblepia xanthicles xanthicles (Godman & Salvin, 1881)  Costa Rica & Panama  M T S

Catoblepia xanthicles belisar Stichel, 1904  TL: Bolivia  M T

Catoblepia xanthicles cyparissa Fruhstorfer, 1907  TL: Colombia  M T

Catoblepia xanthicles mercedensis Bristow, 1981  TL: Peru  M T

Catoblepia xanthicles occidentalis Bristow, 1981  TL: Ecuador  M T

Catoblepia xanthicles orientalis Bristow, 1981  TL: Peru  M T

Catoblepia xanthicles paraensis Bristow, 1981  TL: Brazil (Pa)  M T

Catoblepia xanthicles sosigenes (Fruhstorfer, 1913)  TL: French Guiana  M T

Catoblepia xanthus (Linnaeus, 1758)    iN

Catoblepia xanthus xanthus (Linnaeus, 1758)  TL: ?

Catoblepia xanthus rivalis Niepelt, 1911  TL: Ecuador  M T

Dasyophthalma  Westwood, 1851  

Dasyophthalma creusa (Hübner, [1821])    iN

Dasyophthalma creusa creusa (Hübner, [1821])  TL: Brazil  M S

Dasyophthalma creusa baronesa Stichel, 1904  TL: Brazil (ES)  M T S

Dasyophthalma geraensis Rebel, 1922    iN  M T

Dasyophthalma rusina (Godart, [1824])    iN

Dasyophthalma rusina rusina (Godart, [1824])  TL: Brazil  M T

Dasyophthalma rusina delanira Hewitson, 1862  TL: Brazil  M T

Dasyophthalma rusina principesa Stichel, 1904  TL: Brazil (ES)  M T

Dasyophthalma vertebralis A. Butler, 1869  TL: Brazil ("Pa")  M T S

Dynastor  E. Doubleday, [1849]  

Dynastor darius (Fabricius, 1775)    iN

Dynastor darius darius (Fabricius, 1775)  TL: Brazil ([RJ])  M T

Dynastor darius anaxarete (Cramer, 1776)  TL: Surinam  M T

Dynastor darius faenius Fruhstorfer, 1912  TL: Brazil (RS)  M T

Dynastor darius ictericus Stichel, 1904  TL: Paraguay  M T

Dynastor darius stygianus A. Butler, 1872  S Mexico to W Ecuador  M T S L

Dynastor macrosiris (Westwood, 1851)    iN

Dynastor macrosiris macrosiris (Westwood, 1851)  TL: French Guiana  M T

Dynastor macrosiris hannibal Oberthür, 1881  TL: Colombia  M T

Dynastor macrosiris pharnaces Stichel, 1908  TL: Bolivia

Dynastor macrosiris strix (H. Bates, 1864)  E Mexico, Guatemala, Costa Rica  M T S L

Dynastor napoleon E. Doubleday, [1849]    iN  M T

Eryphanis  Boisduval, 1870  

Eryphanis aesacus (Herrich-Schäffer, 1850)  Double-spotted Owl-Butterfly  E & S Mexico to Honduras & Nicaragua  iN  M T S L

Eryphanis bubocula (A. Butler, 1872)  Bubocula Owl-Butterfly  Costa Rica to Colombia and Amazon Basin  iN  M T S

Eryphanis automedon (Cramer, 1775)    iN

Eryphanis automedon automedon (Cramer, 1775)  TL: Surinam  M T

Eryphanis automedon amphimedon (C. Felder & R. Felder, 1867)  TL: Brazil  M T

Eryphanis automedon cheiremon Fruhstorfer, 1912  TL: Bolivia  M T

Eryphanis lycomedon (C. Felder & R. Felder, 1862)  Split-spotted Owl-Butterfly  Guatemala, Costa Rica, Colombia, Ecuador, Bolivia, southern Brazil  iN  M T S L

Eryphanis gerhardi (Weeks, 1902)    iN  M T

Eryphanis reevesii (E. Doubleday, [1849])    iN

Eryphanis reevesii reevesii (E. Doubleday, [1849])  TL: [Brazil]  M T

Eryphanis reevesii pusillus Stichel, 1904  TL: Paraguay  M T

Eryphanis zolvizora (Hewitson, 1877)    iN

Eryphanis zolvizora isabelae Neild & De Sousa, 2014  Cordillera de la Costa in N Venezuela

Eryphanis zolvizora reyi Bristow, Neild, De Sousa & Huertas, 2014  Cordillera de Mérida in NW Venezuela & N Colombia

Eryphanis zolvizora opimus Staudinger, 1887  E slopes of W Andes in Colombia

Eryphanis zolvizora casagrande Bristow, 2014  SW Colombia & NW Ecuador

Eryphanis zolvizora greeneyi Penz & DeVries, 2008  E Ecuador

Eryphanis zolvizora chachapoya Blandin, 2014  N & central Peru  M T

Eryphanis zolvizora inca Blandin, 2014  central & S Peru

Eryphanis zolvizora zolvizora (Hewitson, 1877)  Bolivia  M T

Mielkella  Casagrande, 1982  

Mielkella singularis (Weymer, 1907)  Singular Owl-Butterfly  S Mexico (Chiapas), Guatemala  iN  M T S

Opoptera  Aurivillius, 1882  

Opoptera staudingeri (Godman & Salvin, 1894)  Staudinger's Owl-Butterfly    iN

Opoptera staudingeri staudingeri (Godman & Salvin, 1894)  Costa Rica & W Panama  M T TI L

Opoptera staudingeri mexicana J. Maza & R. G. Maza, 1989  S Mexico (Chiapas), Guatemala  M T

Opoptera aorsa (Godart, [1824])    iN

Opoptera aorsa aorsa (Godart, [1824])  TL: Brazil  M T

Opoptera aorsa colombiana (Rothschild, 1916)  TL: Colombia  M T

Opoptera aorsa fuscata Stichel, 1908  TL: Brazil (Am)  M T

Opoptera aorsa hilaris Stichel, 1901  TL: Ecuador

Opoptera arsippe (Hopffer, 1874)    iN

Opoptera arsippe arsippe (Hopffer, 1874)  TL: Peru

Opoptera arsippe bracteolata Stichel, 1901  TL: Bolivia  M T

Opoptera arsippe [n. ssp.] Lamas, MS  Peru

Opoptera fruhstorferi (Röber, 1896)    iN  M T S

Opoptera sulcius (Staudinger, 1887)    iN  M T

Opoptera syme (Hübner, [1821])    iN  M T

Opoptera [n. sp.] Casagrande, MS  Peru  M T

Opsiphanes  E. Doubleday, [1849]  

Opsiphanes bogotanus Distant, 1875    iN

Opsiphanes bogotanus bogotanus Distant, 1875  TL: Colombia  M T

Opsiphanes bogotanus alajuela Bristow, 1991  Costa Rica; cloud forests 700-1600m  M T S

Opsiphanes bogotanus blandini Bristow, 1991  TL: Ecuador  M T

Opsiphanes bogotanus phrataphernes Fruhstorfer, 1912  TL: Colombia  M T

Opsiphanes blythekitzmillerae Austin & A. Warren, 2007  Minerva's Owl-Butterfly  NW Mexico  iN  M T S L Ha

Opsiphanes boisduvallii E. Doubleday, [1849]  Orange Owl-Butterfly  E & W Mexico (include. BCS) to Nicaragua  iN  M T S L I

Opsiphanes camena Staudinger, [1886]    iN  M T S

Opsiphanes cassiae (Linnaeus, 1758)    iN

Opsiphanes cassiae cassiae (Linnaeus, 1758)  TL: America  M T HS

Opsiphanes cassiae castaneus Stichel, 1904  Honduras to Colombia  M T S L

Opsiphanes cassiae crameri C. Felder & R. Felder, 1862  TL: Brazil (Ba)  M T

Opsiphanes cassiae kleisthenes Fruhstorfer, 1912  TL: Colombia  M T

Opsiphanes cassiae mexicana Bristow, 1991  SE Mexico, Guatemala  M T S L

Opsiphanes cassiae rubigatus Stichel, 1904  TL: Ecuador  M T S

Opsiphanes cassiae strophios Fruhstorfer, 1907  TL: Bolivia  M T

Opsiphanes cassina C. Felder & R. Felder, 1862    iN

Opsiphanes cassina cassina C. Felder & R. Felder, 1862  TL: Brazil (Am)  M T

Opsiphanes cassina barkeri Bristow, 1991  TL: Ecuador  M T

Opsiphanes cassina caliensis Bristow, 1991  TL: Colombia  M T

Opsiphanes cassina chiriquensis Stichel, 1902  [= aiellae], Nicaragua through Panama  M T S L

Opsiphanes cassina fabricii (Boisduval, 1870)  E & W Mexico to Honduras & El Salvador  M T S L

Opsiphanes cassina merianae Stichel, 1902  TL: Surinam  M T

Opsiphanes cassina milesi Bristow, 1991  TL: Brazil (Pa)  M T

Opsiphanes cassina notanda Stichel, 1904  TL: Peru  M T

Opsiphanes cassina numatius Fruhstorfer, 1912  TL: Colombia  M T

Opsiphanes invirae (Hübner, [1808])    iN

Opsiphanes invirae invirae (Hübner, [1808])  TL: ?

Opsiphanes invirae aequatorialis Stichel, 1902  TL: Ecuador  M T

Opsiphanes invirae agasthenes Fruhstorfer, 1907  TL: Peru  M T

Opsiphanes invirae amplificatus Stichel, 1904  TL: Paraguay  M T

Opsiphanes invirae camposi Bristow, 1991  TL: Ecuador  M T

Opsiphanes invirae cuspidatus Stichel, 1904  Nicaragua to W Panama  M T S

Opsiphanes invirae intermedius Stichel, 1902  TL: Brazil (Am)  M T

Opsiphanes invirae pseudophilon Fruhstorfer, 1907  TL: Brazil (ES)  M T

Opsiphanes invirae remoliatus Fruhstorfer, 1907  TL: Brazil (SC?)  M T

Opsiphanes invirae relucens Fruhstorfer, 1907  S Mexico (Chiapas) to Honduras  M T S

Opsiphanes invirae roraimaensis Bristow, 1991  TL: Guyana  M T

Opsiphanes invirae sieberti Bristow, 1991  TL: French Guiana  M T

Opsiphanes invirae sticheli Röber, 1906  E Panama (CZ) to W Venezuela  M S

Opsiphanes jacobsorum I. Chacón & Montero, 2012  ref.183

Opsiphanes mutatus Stichel, 1902    iN

Opsiphanes mutatus mutatus Stichel, 1902  TL: Ecuador  M T

Opsiphanes mutatus farrago Stichel, 1903  TL: Peru  M T

Opsiphanes mutatus parodizi Bristow, 1991  TL: Bolivia  M T S

Opsiphanes tiberius Attal & Blandin, 2020  ref.193

Opsiphanes quiteria (Stoll, 1780)    iN

Opsiphanes quiteria quiteria (Stoll, 1780)  TL: Surinam  M T

Opsiphanes quiteria angostura Bristow, 1979  TL: Ecuador  M T

Opsiphanes quiteria badius Stichel, 1902  Panama (CZ) to Colombia & Venezuela  M T S L

Opsiphanes quiteria bolivianus Stichel, 1902  TL: Bolivia  M T

Opsiphanes quiteria cardenasi Bristow, 1991  TL: Brazil (Pa)  M T

Opsiphanes quiteria cauca Röber, 1906  TL: Colombia  M T

Opsiphanes quiteria erebus Röber, 1927  TL: Peru

Opsiphanes quiteria meridionalis Staudinger, 1887  TL: Brazil ([SC])  M T

Opsiphanes quiteria phylas Fruhstorfer, 1912  TL: Colombia  M T

Opsiphanes quiteria quaestor Stichel, 1902  TL: Ecuador  M T

Opsiphanes quiteria quirinalis Staudinger, 1887  TL: Peru  M T

Opsiphanes quiteria quirinus Godman & Salvin, 1881  SE Mexico to Panama (Atlantic slope)  M T S L

Opsiphanes quiteria talamancensis Bristow, 1991  Costa Rica to W Panama  M T S

Opsiphanes sallei E. Doubleday, [1849]    iN

Opsiphanes sallei sallei E. Doubleday, [1849]  TL: Venezuela  M T

Opsiphanes sallei colombiana Bristow, 1991  TL: Colombia  M T S

Opsiphanes sallei kennerleyi Bristow, 1991  TL: Peru  M T S

Opsiphanes sallei nicandrus Fruhstorfer, 1912  TL: Bolivia  M T

Opsiphanes tamarindi C. Felder & R. Felder, 1861  Narrow-banded Owl-Butterfly    iN

Opsiphanes tamarindi tamarindi C. Felder & R. Felder, 1861  E & W Mexico to Colombia  M T S L I

Opsiphanes tamarindi corrosus Stichel, 1904  TL: Ecuador  M T S

Opsiphanes tamarindi incolumis Stichel, 1904  TL: Peru  M T

Opsiphanes tamarindi latifascia Rothschild, 1916  TL: Peru  M T

Opsiphanes tamarindi xiphos Fruhstorfer, 1907  TL: Venezuela  M T

Opsiphanes zelotes Hewitson, 1873    iN

Opsiphanes zelotes zelotes Hewitson, 1873  TL: Colombia  M T

Opsiphanes zelotes zelus Stichel, 1908  Panama  M S

Orobrassolis  Casagrande, 1982  

Orobrassolis ornamentalis (Stichel, 1906)    iN  M T

Orobrassolis latusoris Penz & Simosen, 2011  TL: not specified

Penetes  E. Doubleday, [1849]  

Penetes pamphanis E. Doubleday, [1849]    iN  M T

Selenophanes  Staudinger, 1887  

Selenophanes cassiope (Cramer, 1775)    iN

Selenophanes cassiope cassiope (Cramer, 1775)  TL: Surinam; Guyana

Selenophanes cassiope amplior Stichel, 1902  TL: Colombia  M T

Selenophanes cassiope andromeda Stichel, 1901  TL: Bolivia  M T

Selenophanes cassiope cassiopeia (Staudinger, [1886])  TL: Brazil (Am)  M T

Selenophanes cassiope guarany Casagrande, 1992  TL: Brazil (SP)  M T

Selenophanes cassiope haraposa Bristow, 1982  TL: Brazil (Pa)  M T S

Selenophanes cassiope mapiriensis Bristow, 1982  TL: Bolivia  M T

Selenophanes cassiope perenensis Bristow, 1982  TL: Peru  M T

Selenophanes cassiope placentia (Fruhstorfer, 1912)  TL: Bolivia

Selenophanes cassiope theognis Fruhstorfer, 1910  TL: Brazil (MT)  M T

Selenophanes josephus (Godman & Salvin, 1881)  Josephus Owl-Butterfly    iN

Selenophanes josephus josephus (Godman & Salvin, 1881)  Guatemala  M T OI

Selenophanes josephus excisus (Rothschild, 1916)  TL: Colombia  M T

Selenophanes josephus excultus Stichel, 1902  E Panama to Colombia  M T S

Selenophanes supremus Stichel, 1901  TL: Ecuador

Selenophanes supremus supremus Stichel, 1901  TL: Ecuador  M T

Selenophanes supremus ditatus Thieme, 1902  TL: Peru  M S

Aponarope  Casagrande, 1982  

Aponarope sutor (Stichel, 1916)  TL: Brazil (MT)  M T S

Narope  E. Doubleday, [1849]  

Narope albopunctum Stichel, 1904  TL: Peru  M T

Narope anartes Hewitson, 1874    iN  M T S

Narope cauca Casagrande, 2002    iN  M T

Narope cyllabarus Westwood, 1851    iN  M T S

Narope cyllarus Westwood, 1851    iN  M T

Narope cyllastros E. Doubleday, [1849]    iN  M T

Narope cyllene C. Felder & R. Felder, 1859    iN  M T

Narope denticulatus Talbot, 1928    iN  M T

Narope guilhermei Casagrande, 1989  TL: Brazil (SC)  M T S

Narope marmorata Schaus, 1902    iN  M T S

Narope minor Casagrande, 2002  Small Owl-Butterfly  S Mexico (VER, OAX), Guatemala  M T OI G

Narope nesope Hewitson, 1869  TL: Ecuador  M T S

Narope obidos Casagrande, 2002  TL: Brazil (Pa)  M T S

Narope panniculus Stichel, 1904    iN  M T S

Narope syllabus Staudinger, 1887    iN

Narope syllabus syllabus Staudinger, 1887  TL: Peru  M T S

Narope syllabus stygius Staudinger, 1887  TL: Brazil (Am)  M T S

Narope testacea Godman & Salvin, 1878  Brown Owl-Butterfly  S Mexico to Panama  iN  M T S

Narope ybyra Casagrande, 2002  TL: Bolivia  M T

  Tribe Elymniini

Neorinella  Martins, Kucera-Santos, Vieira & Fragoso, 1993  a fossil genus

Neorinella garciae Martins, Kucera-Santos, Vieira & Fragoso, 1993  Fossil from Brazil (SP)

  Tribe Haeterini

Cithaerias  Hübner, [1819]  

Cithaerias andromeda (Fabricius, 1775)    iN

Cithaerias andromeda andromeda (Fabricius, 1775)  TL: India  M T

Cithaerias andromeda azurina (J. Zikán, 1942)  TL: Brazil (Am)  M T

Cithaerias andromeda bandusia Staudinger, [1887]  TL: Brazil (Am)  M T

Cithaerias andromeda esmeralda (E. Doubleday, 1845)  TL: Brazil (Pa)  M T

Cithaerias pyropina (Salvin & Godman, 1868)    iN

Cithaerias pyropina pyropina (Salvin & Godman, 1868)  TL: Peru  M T S

Cithaerias pyropina julia Willmott, 2015  ref.301

Cithaerias pyropina songoana (Langer, 1944)  TL: Bolivia  M T S

Cithaerias pireta (Stoll, 1780)  Rusted Clearwing-Satyr    iN

Cithaerias pireta pireta (Stoll, 1780)  [= menander], Guatemala to Colombia  M T S L

Cithaerias pireta aurora (C. Felder & R. Felder, 1862)  TL: [Peru]  M T

Cithaerias pireta magdalenensis Constantino, 1995  TL: Colombia  M T

Cithaerias aurorina (Weymer, 1910)    iN

Cithaerias aurorina aurorina (Weymer, 1910)  TL: Upper Amazon  M T S

Cithaerias aurorina tambopata Lamas, 1998  TL: Peru  M T S

Cithaerias cliftoni Constantino, 1995    iN  M T

Cithaerias pyritosa (J. Zikán, 1942)    iN  M T

Dulcedo  R.F. d'Almeida, 1951  

Dulcedo polita (Hewitson, 1869)  Uncolored Clearwing-Satyr  Nicaragua to Colombia  iN  M T S L

Haetera  Fabricius, 1807  

Haetera macleannania (H. Bates, 1865)  Reddish Clearwing-Satyr  Costa Rica to W Ecuador  iN  M T TI S IL L

Haetera piera (Linnaeus, 1758)    iN

Haetera piera sanguinolenta Constantino & Salazar, 2007  TL: Colombia  M T

Haetera piera piera (Linnaeus, 1758)  TL: Indiis  M HS

Haetera piera diaphana Lucas, 1857  TL: Cuba  M T

Haetera piera negra C. Felder & R. Felder, 1862  TL: [Peru]  M T

Haetera piera pakitza Lamas, 1998  TL: Peru  M T

Haetera piera unocellata Weymer, 1910  TL: Bolivia  M T

Pierella  Westwood, 1851  

Pierella amalia Weymer, 1885    iN  M T

Pierella astyoche (Erichson, [1849])    iN

Pierella astyoche astyoche (Erichson, [1849])  TL: Guyana  M T

Pierella astyoche bernhardina Bryk, 1953  TL: Brazil (Am)  M T

Pierella astyoche stollei Ribeiro, 1931  TL: Brazil (Rd)  M T

Pierella helvina (Hewitson, 1859)    iN

Pierella helvina helvina (Hewitson, 1859)  TL: Colombia  M T S

Pierella helvina hymettia Staudinger, [1886]  TL: Colombia

Pierella helvina incanescens Godman & Salvin, 1877  Guatemala to Panama  M T S L

Pierella helvina ocreata Salvin & Godman, 1868  Panama to NW Colombia  M T S L

Pierella helvina pacifica Niepelt, 1924  TL: Colombia  M T

Pierella hortona (Hewitson, 1854)    iN

Pierella hortona hortona (Hewitson, 1854)  TL: Brazil (Am)  M T

Pierella hortona albofasciata Rosenberg & Talbot, 1914  TL: Peru  M T

Pierella hortona [n. ssp.] Lamas, MS  Peru

Pierella hyalinus (Gmelin, [1790])    iN

Pierella hyalinus hyalinus (Gmelin, [1790])  TL: Surinam  M T

Pierella hyalinus extincta Weymer, 1910  TL: Brazil (Am)  M T S

Pierella hyalinus schmidti Constantino, 1995  TL: Colombia  M T

Pierella hyalinus velezi Constantino, 1995  TL: Colombia  M T

Pierella hyceta (Hewitson, 1859)    iN

Pierella hyceta hyceta (Hewitson, 1859)  TL: [Peru]  M T

Pierella hyceta ceryce (Hewitson, 1874)  TL: Bolivia  M T

Pierella hyceta latona (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Pierella hyceta [n. ssp.#1] Lamas, MS  TL: Peru

Pierella hyceta [n. ssp.#2] Lamas, MS  TL: Peru

Pierella lamia (Sulzer, 1776)    iN

Pierella lamia lamia (Sulzer, 1776)  TL: America  M T

Pierella lamia boliviana F. Brown, 1948  TL: Bolivia  M T

Pierella lamia chalybaea Godman, 1905  TL: Brazil (MT)  M T S

Pierella lena (Linnaeus, 1767)    iN

Pierella lena lena (Linnaeus, 1767)  TL: S. Amer.  M T

Pierella lena brasiliensis (C. Felder & R. Felder, 1862)  TL: [Peru]  M T S

Pierella lena salma Constantino, 2007  TL: Colombia  M T

Pierella lucia Weymer, 1885    iN  M T

Pierella luna (Fabricius, 1793)  Moon Satyr    iN

Pierella luna luna (Fabricius, 1793)  Nicaragua to Colombia  M T S L

Pierella luna lesbia Staudinger, 1887  TL: Colombia  M S

Pierella luna pallida (Salvin & Godman, 1868)  Honduras & Nicaragua  M T IL L

Pierella luna rubecula Salvin & Godman, 1868  E & S Mexico to Honduras  M T S L

Pierella nereis (Drury, 1782)    iN

Pseudohaetera  F. Brown, [1943]  

Pseudohaetera hypaesia (Hewitson, 1854)    iN  M T

Pseudohaetera mimica (Rosenberg & Talbot, 1914)    iN  M T L

  Tribe Melanitini

Manataria  W. F. Kirby, [1902]  

Manataria maculata (Hopffer, 1874)  White-spotted Satyr  E & W Mexico to Colombia  iN

Manataria maculata maculata (Hopffer, 1874)  E & W Mexico to Colombia  M S HS L I Ha

Manataria maculata daguana (Neustetter, 1929)  TL: Colombia  M T

Manataria hercyna (Hübner, [1821])    iN

Manataria hercyna hyrnethia Fruhstorfer, 1912  TL: Bolivia; Peru  M T

Manataria hercyna distincta (Lathy, 1918)  TL: French Guiana  M T

Manataria hercyna hercyna (Hübner, [1821])  TL: [Brazil]  M OI

  Tribe Satyrini, Subtribe Lethina

Lethe  Hübner, [1819]  

Lethe anthedon (A. Clark, 1936)  Northern Pearly-eye    iN

Lethe anthedon borealis (A. Clark, 1936)  TL: not specified  M T S L

Lethe anthedon anthedon (A. Clark, 1936)  TL: USA (NY)  M T S L1 L2 L3 L4

Lethe creola (Skinner, 1897)  Creole Pearly-eye    iN  M T S L1 L2

Lethe portlandia (Fabricius, 1781)  Southern Pearly-eye    iN

Lethe portlandia missarkae J. Heitzman & dos Passos, 1974  TL: USA (Ar)  M S L I H

Lethe portlandia portlandia (Fabricius, 1781)  TL: USA (Ga)  M S L

Lethe portlandia floralae J. Heitzman & dos Passos, 1974  TL: USA (Fl)  M T S L

Lethe eurydice (Linnaeus, 1763)  Eyed Brown    iN

Lethe eurydice fumosus (Leussler, 1916)  TL: USA (Ne)  M T S

Lethe eurydice eurydice (Linnaeus, 1763)  TL: USA (Pa)  M T S L

Lethe appalachia R. Chermock, 1947  Appalachian Brown    iN

Lethe appalachia leeuwi Gatrelle & Arbogast, 1974  TL: USA (Mi)  M T L

Lethe appalachia appalachia R. Chermock, 1947  TL: USA (NC)  M S L1 L2 Ha

  Tribe Satyrini, Subtribe Calistina

Calisto  Hübner, [1823]  

Calisto herophile Hübner, [1823]  Cuban Calisto    iN

Calisto herophile herophile Hübner, [1823]  much of Cuba and Isla de Juventud; lowlands  M S IL L I

Calisto herophile parsonsi Clench, 1943  Cuba: Sierra de Escambray, 500-1000m  M T IL

Calisto bruneri Michener, 1949  Bruner's Calisto  Nipe-Sagua-Baracoa Mts., E Cuba  iN  M T IL I

Calisto smintheus M. Bates, 1935  Smintheus Calisto  [= biocellatus, brocheri], Sierra de Maestra, E Cuba, 1000-2500m  iN  M T IL L I

Calisto muripetens M. Bates, 1939  Muripetens Calisto  C Cuba; ca. 1000m  M T S I

Calisto bradleyi Munroe, 1951  Bradley's Calisto  W Cuba; 500m  M S

Calisto brochei Torre, 1973  Broche's Calisto  Nipe-Sagua-Baracoa Mts., E Cuba  iN  M I

Calisto israeli Torre, 1973  Israeli Calisto  Nipe-Sagua-Baracoa Mts., E Cuba

Calisto occulta Núñez, 2012  Cryptic Calisto  Nipe-Sagua-Baracoa Mts., E Cuba  M I

Calisto torrei Núñez, 2013  Guamuhaya Mts., Cuba  M T L I

Calisto aquilum Núñez, 2013  Guamuhaya Mts., Cuba  iN

Calisto aquilum aquilum Núñez, 2013  Guamuhaya Mts., Cuba  M T L

Calisto aquilum occidentalis Núñez, 2013  Guaniguanico Mts., Cuba

Calisto dissimulatum Núñez, 2013  Sierra Cristal, Nipe-Sagua-Baracoa & middle of Sierra Maestra, Cuba  M T L I

Calisto sibylla M. Bates, 1934  Woodland Calisto  Bahamas: New Providence, N & S Andros  M T S

Calisto apollinis M. Bates, 1934  Cuban Calisto  Bahamas  M T S L

Calisto zangis (Fabricius, 1775)  Jamaican Calisto  Jamaica  iN  M T S1 S2 L

Calisto nubila Lathy, 1899  Puerto Rican Calisto  Puerto Rico  iN  M T S1 S2 L I

Calisto anegadensis D. Smith, L. Miller & McKenzie, 1991  Anegada Calisto  Anegada (British Virgin Islands)  M T

Calisto pulchella Lathy, 1899  Sugar cane Calisto    iN

Calisto pulchella pulchella Lathy, 1899  Hispaniola (widespread, except in Cordillera Central; see below)  M T S I

Calisto pulchella darlingtoni Clench, 1943  Cordillera Central, W Dominican Republic; montane  M T S

Calisto raburni Gali, 1985  Raburn's Calisto  NE of El Aguacate, Independencia, Dominican Republic  M T S

Calisto arcas M. Bates, 1939  Tawny-washed Calisto  Cordillera Central, Pico Duarte massif, Dominican Republic  M T S

Calisto chrysaoros M. Bates, 1935  White-banded Calisto  Massif de la Hotte, Haiti; Sierra de Baoruco, Dom. Rep.  iN  M T S L

Calisto galii A. Schwartz, 1983  Gali's Calisto  TL: Dominican Rep.

Calisto galii galii A. Schwartz, 1983  Cordillera Central, W Dominican Republic  M T S L

Calisto galii choneupsilon A. Schwartz, 1985  Sierra de Neiba, Dominican Republic  M T

Calisto clenchi A. Schwartz & Gali, 1984  Clench's Calisto  Massif de la Selle (Haiti-Dominican Republic)  M T S

Calisto wetherbeei A. Schwartz & F. González, 1988  Wetherbee's Calisto  Cordillera Central, W Dominican Republic  M T

Calisto neochma A. Schwartz, 1991  Neochma Calisto  Dominican Republic

Calisto clydoniata A. Schwartz & Gali, 1984  Yellow-stained Calisto  Sierra de Neiba, Dominican Republic  M T

Calisto archebates (Ménétriés, 1832)  Yellow-banded Calisto  Massif de la Selle (Haiti-D.R.), W Sierra de Baoruco, Dom. Rep.  iN  M S L I

Calisto eleleus M. Bates, 1935  Eleleus Calisto  Hispaniola; most abundant in Sierra de Baoruco  iN  M T S

Calisto loxias M. Bates, 1935  Dark Calisto  Massif de la Hotte, Haiti; nr. Las Abejas, SW Dom. Republic  M T S

Calisto woodsi K. Johnson & Hedges, 1998  Woods' Calisto  Massif de la Hotte, Haiti  M OI

Calisto tasajera F. González, A. Schwartz & Wetherbee, 1991  Tasajera Calisto  Cordillera Central, Dominican Republic  M S L I

Calisto schwartzi Gali, 1985  Schwartz's Calisto  nr. Aceitillar, Sierra de Baoruco, Pedernales, SW Dom. Republic  M T S

Calisto lyceius M. Bates, 1935  Saona Calisto  Isla Saona & Isla Catalina, Dominican Republic  iN  M T S

Calisto franciscoi Gali, 1985  Francisco's Calisto  SW Dom. Republic (Valle de Neiba & Barahona Pen. to Azua)  iN  M T S

Calisto crypta Gali, 1985  Cryptic Calisto  vic. Monte Cristi, Dominican Republic  iN  M T S L I Ha

Calisto confusa Lathy, 1899  Confused Calisto  Hispaniola (widespread)  iN  M T S L I

Calisto debarriera Clench, 1943  Debarriera Calisto  [= gonzalezi, montana, neiba] Haiti, Dom. Rep. (Sierra de Nieba)  iN  M T1 T2 S

Calisto hysius (Godart, [1824])  Godart's Calisto    iN

Calisto hysius hysius (Godart, [1824])  Hispaniola (S palaeoisland)  M T S L I

Calisto hysius aleucosticha Correa & A. Schwartz, 1986  N Cordillera Central, Dominican Republic  M T

Calisto batesi Michener, 1943  Bates' Calisto  Hispaniola (most common in N Dominican Republic)  iN  M T S L

Calisto pauli K. Johnson & Hedges, 1998  Paul's Calisto  Massif de la Hotte, Haiti  M OI

Calisto obscura Michener, 1943  Obscure Calisto  Hispaniola (widespread)  iN  M T S L I

Calisto ainigma K. Johnson, Quinter & Matusik, 1987  Enigmatic Calisto  Cordillera Central, Dominican Republic, possilby an aberration of C. obscura  M T

Calisto grannus M. Bates, 1939  Grannus Calisto    iN

Calisto grannus dilemma F. González, 1987  nr. Constanza, La Vega, Dominican Republic  M T S L I

Calisto grannus dystacta F. González, 1987  Loma Leonor, Santiago Rodriguez, Dominican Republic  M T

Calisto grannus micheneri Clench, 1944  Cordillera Central, Dominican Republic  iN  M T S

Calisto grannus micrommata A. Schwartz & Gali, 1984  S range Sierra de Neiba, SW Dominican Republic, 1600-1900m  M T

Calisto grannus grannus M. Bates, 1939  Cordillera Central, Dominican Republic, 1160-2300m  M T S L I

Calisto grannus amazona F. González, 1987  vic. Pico Duarte, Cordillera Central, Dom. Rep., 1200-1900m  M T S

Calisto grannus phoinix F. González, 1987  La Palma, E Cordillera Central, La Vega, Dominican Republic  M T S

Calisto grannus sommeri A. Schwartz & Gali, 1984  E Massif de la Selle, Haiti, to W Sierra de Baoruco, Dom. Rep.  M T S

Calisto thomasi K. Johnson & Hedges, 1998  Thomas' Calisto  Massif de la Hotte, Haiti  M OI

Calisto tragius M. Bates, 1935  Tragic Calisto  Massif de la Selle (Haiti); W Sierra de Baoruco, Dom. Republic  M T S

Calisto hendersoni Gali, 1985  Henderson's Calisto  nr. El Limon, Independencia, Dominican Republic  M T S

  Tribe Satyrini, Subtribe Gyrocheilina

Gyrocheilus  A. Butler, 1867  

Gyrocheilus patrobas (Hewitson, 1862)  Red-bordered Satyr    iN

Gyrocheilus patrobas tritonia (W. H. Edwards, 1874)  C & SE AZ, SW NM, far W TX, NW Mexico  M T S L I

Gyrocheilus patrobas patrobas (Hewitson, 1862)  C & S Mexico  M T S L

Gyrocheilus [n. sp.] N/A, N/A  Rust-bordered Satyr  NE Mexico  M S

  Tribe Satyrini, Subtribe Pronophilina

Pronophila  E. Doubleday, [1849]  

Pronophila unifasciata Lathy, 1906    iN

Pronophila unifasciata unifasciata Lathy, 1906  TL: Ecuador  M T S

Pronophila unifasciata deverra Thieme, 1907  TL: Ecuador

Pronophila unifasciata brennus Thieme, 1907  TL: Colombia  M T

Pronophila unifasciata bogotensis Jurriaanse, 1926  TL: Colombia  M T S

Pronophila unifasciata australis Hayward, 1949  TL: Argentina  M T

Pronophila unifasciata donachui Adams & Bernard, 1977  TL: Colombia  M T

Pronophila unifasciata nanegalito Pyrcz, 2000  TL: Ecuador

Pronophila unifasciata deceptoria Pyrcz, 2005  TL: not specified

Pronophila unifasciata eugeni Pyrcz, 2005  TL: Colombia

Pronophila unifasciata kennethi Pyrcz, 2005  TL: Ecuador  M S

Pronophila unifasciata antioquiana Pyrcz, 2005  TL: Peru

Pronophila unifasciata [n. ssp.] Pyrcz, MS  Bolivia

Pronophila juliani Adams & Bernard, 1977    iN  M T

Pronophila timanthes Salvin, 1871  Timanthes Satyr  Costa Rica to W Ecuador  iN  M T OI S L

Pronophila obscura A. Butler, 1868    iN

Pronophila obscura obscura A. Butler, 1868  TL: Venezuela  M T

Pronophila obscura cuchillaensis Viloria & Camacho, 1999  TL: Venezuela

Pronophila epidipnis Thieme, 1907    iN

Pronophila epidipnis epidipnis Thieme, 1907  TL: Venezuela  M T

Pronophila epidipnis orchewitsoni Adams & Bernard, 1979  TL: Colombia  M T S

Pronophila epidipnis lathyi Pyrcz, 2000  TL: Ecuador

Pronophila epidipnis orcidipnis Pyrcz & Viloria, 2007  ref.285

Pronophila epidipnis perplexa Pyrcz, 2004  TL: Peru

Pronophila epidipnis cana Pyrcz, MS  TL: Peru

Pronophila epidipnis [n. ssp.#1] Pyrcz, MS  TL: Colombia

Pronophila epidipnis [n. ssp.#2] Pyrcz, MS  TL: Bolivia

Pronophila tremocrata Pyrcz, 2004    iN  M S

Pronophila cordillera Westwood, 1851    iN

Pronophila cordillera cordillera Westwood, 1851  TL: [Bolivia]  M T

Pronophila cordillera variabilis A. Butler, 1873  TL: Peru  M T S

Pronophila cordillera colocasia Thieme, 1907  TL: Peru  M T

Pronophila cordillera [n. ssp.] Pyrcz & Boyer, MS  Peru

Pronophila orcus (Latreille, [1813])    iN

Pronophila orcus orcus (Latreille, [1813])  TL: ["Peru"]  M T S

Pronophila orcus locuples Weymer, 1912  TL: Bolivia  M S

Pronophila orcus nepete Thieme, 1907  TL: Peru  M T

Pronophila thelebe E. Doubleday, [1849]    iN  M T

Pronophila isobelae Pyrcz, 2000  TL: Ecuador

Pronophila isobelae isobelae Pyrcz, 2000  TL: Ecuador

Pronophila isobelae benevola Pyrcz, 2000  TL: Ecuador

Pronophila attali Pyrcz, 2000  TL: Peru

Pronophila rosenbergi Lathy, 1906    iN

Pronophila rosenbergi rosenbergi Lathy, 1906  TL: Ecuador  M T

Pronophila rosenbergi puyango Pyrcz, 2000  TL: Ecuador

Pronophila bernardi Pyrcz, 2004    iN

Pronophila intercidona Thieme, 1907    iN

Pronophila intercidona intercidona Thieme, 1907  TL: Ecuador  M T

Pronophila intercidona thelebina Thieme, 1907  TL: Bolivia  M T

Pronophila intercidona [n. ssp.] Pyrcz & Prieto, MS  Colombia

Pronophila margarita Pyrcz, 2000  TL: Ecuador

Corades  E. Doubleday, [1849]  

Corades enyo Hewitson, [1849]    iN

Corades enyo enyo Hewitson, [1849]  TL: Venezuela  M T

Corades enyo almo Thieme, 1907  TL: Colombia  M T L

Corades enyo [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Corades enyo [n. ssp.#2] Pyrcz, MS  TL: Bolivia

Corades enyo [n. ssp.#3] Pyrcz, MS  TL: Ecuador

Corades pax Watkins, 1939  TL: Venezuela  M T

Corades chelonis Hewitson, 1863    iN

Corades chelonis chelonis Hewitson, 1863  TL: Colombia  M T

Corades chelonis rubeta Thieme, 1907  TL: Venezuela

Corades chelonis rectilinea Pyrcz & Viloria, 2005  TL: Colombia

Corades lactefusa Thieme, 1907    iN  M S

Corades iduna Hewitson, 1850    iN

Corades iduna iduna Hewitson, 1850  TL: Bolivia  M T

Corades iduna procellaria Thieme, 1907  TL: Argentina

Corades iduna marginalis A. Butler, 1873  TL: Peru  M T S

Corades iduna peruviana A. Butler, 1873  TL: Peru  M T S

Corades iduna gradosi Lamas, 2003  TL: Peru

Corades iduna [n. ssp.] Pyrcz, MS  Peru

Corades pannonia Hewitson, 1850    iN

Corades pannonia pannonia Hewitson, 1850  TL: Venezuela  M T

Corades pannonia ploas Thieme, 1907  TL: Venezuela; Colombia  M T

Corades pannonia condorita Lamas, [1997]  TL: Peru

Corades pannonia albomaculata Staudinger, [1887]  TL: Peru  M T S

Corades pannonia [n. ssp.] Pyrcz, MS  Ecuador

Corades argentata A. Butler, 1868    iN  M T S L

Corades sareba Hewitson, 1850    iN

Corades sareba sareba Hewitson, 1850  TL: Bolivia  M T

Corades sareba fusciplaga A. Butler, 1873  TL: Peru  M T

Corades cybele A. Butler, 1866    iN

Corades cybele cybele A. Butler, 1866  TL: Colombia  M T

Corades cybele semiplena Thieme, 1907  TL: Peru  M T

Corades cybele [n. ssp.#1] Pyrcz, MS  TL: Colombia

Corades cybele [n. ssp.#2] Pyrcz, MS  TL: Peru

Corades cybele [n. ssp.#3] Pyrcz, MS  TL: Peru

Corades medeba Hewitson, 1850    iN

Corades medeba medeba Hewitson, 1850  TL: Bolivia  M T S L

Corades medeba columbina Staudinger, 1894  TL: Colombia  M T L

Corades medeba pittieri Pyrcz, 2010  TL: Venezuela

Corades melania Staudinger, 1894    iN

Corades melania melania Staudinger, 1894  TL: Bolivia  M T

Corades melania carlos Pyrcz, 2006  TL: Venezuela

Corades cistene Hewitson, 1863    iN

Corades cistene cistene Hewitson, 1863  TL: [Bolivia]  M T

Corades cistene generosa Thieme, 1907  TL: Peru; Bolivia  M T L

Corades cistene [n. ssp.] Pyrcz, MS  Ecuador

Corades dymantis Thieme, 1907    iN

Corades dymantis dymantis Thieme, 1907  TL: Colombia  M T

Corades dymantis [n. ssp.#1] Pyrcz, MS  TL: Colombia

Corades dymantis [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Corades dymantis [n. ssp.#3] Pyrcz, MS  TL: Ecuador

Corades chirone Hewitson, 1863    iN  M T

Corades tripunctata Weymer, 1890  TL: Peru

Corades tripunctata tripunctata Weymer, 1890  TL: Peru  M T

Corades tripunctata necrufa Pyrcz, 2004  TL: Peru

Corades tripunctata malambo Pyrcz, 2006  TL: Venezuela

Corades ulema Hewitson, 1850    iN

Corades ulema ulema Hewitson, 1850  TL: Bolivia  M T S L

Corades ulema mirianae Lamas, [1997]  TL: Peru

Corades anfortas (Thieme, 1907)    iN

Corades anfortas anfortas (Thieme, 1907)  TL: Peru  M T

Corades anfortas celestini Pyrcz, 2005  TL: Venezuela

Corades tricordatus Hewitson, 1874  TL: Bolivia  M T

Corades callipolis Hewitson, 1874  TL: Bolivia  M T

Corades liliaceus Pyrcz, MS  TL: Ecuador

Corades [n. sp.#1] Pyrcz, MS  TL: Peru

Corades [n. sp.#2] Lamas, MS  TL: Peru

Corades [n. sp.#3] Pyrcz, MS  TL: not specified

Corades [n. sp.#3] [n. ssp.#1] Pyrcz, MS  TL: Venezuela

Corades [n. sp.#3] [n. ssp.#2] Pyrcz, MS  TL: Venezuela

Corades [n. sp.#4] Pyrcz, MS  TL: Bolivia

Daedalma  Hewitson, 1858  

Daedalma dinias Hewitson, 1858    iN

Daedalma dinias dinias Hewitson, 1858  TL: Colombia  M T

Daedalma dinias salmoni Pyrcz, 2011  ref.281

Daedalma dinias emma Pyrcz & Greeney, 2011  ref.281  M L

Daedalma dinias leticia Pyrcz, 2011  ref.281

Daedalma dinias romeliana Pyrcz & Prieto, 2011  ref.281

Daedalma dinias granadillas Pyrcz & Willmott, 2011  ref.281

Daedalma dinias radfordi Pyrcz & Willmott, 2011  TL: not specified

Daedalma boliviana Staudinger, 1897    iN

Daedalma boliviana boliviana Staudinger, 1897  TL: Bolivia; Peru  M T

Daedalma boliviana peruviana Pyrcz, 2004  TL: Peru

Daedalma fraudata Pyrcz, 2004  TL: Peru  M S

Daedalma rubroreducta Pyrcz & Willmott, 2011  ref.281

Daedalma drusilla Hewitson, 1858    iN

Daedalma drusilla drusilla Hewitson, 1858  TL: Colombia  M T

Daedalma drusilla tinta Pyrcz, 2011  ref.281

Daedalma drusilla krugeriana Pyrcz, 2011  ref.281

Daedalma inconspicua A. Butler, 1866    iN

Daedalma inconspicua inconspicua A. Butler, 1866  TL: Ecuador  M T

Daedalma inconspicua blancae Pyrcz, 2011  ref.281

Daedalma inconspicua cuencana Pyrcz, 2011  ref.281

Daedalma inconspicua onorei Pyrcz, 2011  ref.281

Daedalma inconspicua orientalis Pyrcz, 2011  ref.281

Daedalma inconspicua variegata Pyrcz, 2011  ref.281

Daedalma inconspicua palacio Dognin, 1891  TL: Ecuador  M T

Daedalma inconspicua petitia Pyrcz, 2011  ref.281

Daedalma inconspicua tapichalaca Pyrcz & Willmott, 2011  ref.281

Daedalma vertex Pyrcz, 2004  TL: Peru

Daedalma vertex vertex Pyrcz, 2004  TL: Peru  M T

Daedalma vertex milleri Pyrcz, 2011  ref.281  M T

Daedalma parvomaculata E. Krüger, 1924  TL: Colombia  M T S

Daedalma adamsi D'Abrera, 1988  TL: Peru  M T

Daedalma eliza Pyrcz & Willmott, 2011  ref.281  iN

Daedalma dognini Pyrcz, 2011  TL: Peru

Daedalma dognini dognini Pyrcz, 2011  ref.281

Daedalma dognini mirianae Pyrcz, 2011  ref.281  M T

Thiemeia  Weymer, 1912  

Thiemeia phoronea (E. Doubleday, [1849])    iN

Thiemeia phoronea phoronea (E. Doubleday, [1849])  TL: [Venezuela]  M T

Thiemeia phoronea obscurata E. Krüger, 1924  TL: Colombia

Thiemeia phoronea ortruda (Thieme, 1907)  TL: Bolivia  M T

Junea  Hemming, 1964  

Junea dorinda (C. Felder & R. Felder, 1862)    iN

Junea dorinda dorinda (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Junea dorinda quasinegra Pyrcz, 2004  TL: Peru

Junea dorinda whitelyi (H. Druce, 1876)  TL: Peru  M T S

Junea dorinda [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Junea dorinda [n. ssp.#2] Pyrcz, MS  TL: Peru

Junea doraete (Hewitson, 1858)    iN

Junea doraete doraete (Hewitson, 1858)  TL: Colombia  M T

Junea doraete antissa (Thieme, 1907)  TL: Peru

Junea doraete gideon (Thieme, 1907)  TL: Peru  M T S

Junea doraete enipeus (Thieme, 1907)  TL: Bolivia  M T

Junea doraete [n. ssp.#1] Pyrcz & Clavijo, MS  TL: Colombia

Junea doraete [n. ssp.#2] Pyrcz & Clavijo, MS  TL: Colombia

Junea doraete [n. ssp.#3] Pyrcz, MS  TL: Ecuador

Junea doraete [n. ssp.#4] Pyrcz, MS  TL: Ecuador

Oxeoschistus  A. Butler, 1867  

Oxeoschistus protogenia (Hewitson, 1862)    iN  M T

Oxeoschistus pronax (Hewitson, 1859)    iN  M T S

Oxeoschistus puerta (Westwood, 1851)    iN

Oxeoschistus puerta submaculatus A. Butler & H. Druce, 1874  Costa Rica & Panama  M T S L

Oxeoschistus puerta pervius Thieme, 1907  TL: Colombia  M T

Oxeoschistus puerta puerta (Westwood, 1851)  TL: "Colombia"; Venezuela  M T

Oxeoschistus puerta magna Pyrcz & Viloria, 2007  TL: Venezuela

Oxeoschistus simplex A. Butler, 1868    iN

Oxeoschistus simplex simplex A. Butler, 1868  TL: Colombia  M T

Oxeoschistus simplex fuscus R. Krüger, 1929  TL: Colombia  M T

Oxeoschistus simplex isolda Thieme, 1907  TL: Ecuador  M T

Oxeoschistus simplex triplex Pyrcz & Rodriguez, MS  TL: Venezuela

Oxeoschistus simplex duplex Godman, 1905  TL: Bolivia  M T

Oxeoschistus simplex garleppi Lamas, 2003  TL: Peru

Oxeoschistus iphigenia Pyrcz, 2004    iN  M T

Oxeoschistus euriphyle A. Butler, 1872  Dot-banded Satyr  Costa Rica & Panama  iN  M T S L

Oxeoschistus hilara (H. Bates, 1865)  Dot-banded Satyr    iN

Oxeoschistus hilara hilara (H. Bates, 1865)  S Mexico (Chiapas) & Guatemala  M T S L

Oxeoschistus hilara [n. ssp.#1] Pyrcz, MS  TL: Mexico

Oxeoschistus hilara [n. ssp.#2] Pyrcz, MS  TL: Honduras

Oxeoschistus hilara [n. ssp.#3] Pyrcz & Marabuto, MS  TL: Honduras

Oxeoschistus cothonides Grose-Smith, 1896  Rust-patched Satyr  Costa Rica & Panama  iN  M T S

Oxeoschistus cothon Salvin, 1871  Lemon-patched Satyr  Costa Rica & W Panama  iN  M T S L

Oxeoschistus tauropolis (Westwood, [1850])  Yellow-patched Satyr    iN

Oxeoschistus tauropolis tauropolis (Westwood, [1850])  E Mexico to Panama  M T S L

Oxeoschistus tauropolis W Mexican segregate ,  SW Mexico  M S

Oxeoschistus tauropolis [n. ssp.] Pyrcz, MS  Mexico

Oxeoschistus leucospilos Staudinger, 1876    iN

Oxeoschistus leucospilos leucospilos Staudinger, 1876  TL: Peru  M T S

Oxeoschistus leucospilos pugil (Thieme, 1907)  TL: Bolivia  M T

Oxeoschistus romeo Pyrcz & Fratello, 2005  TL: not specified

Oxeoschistus [n. sp.] Pyrcz, MS  Guatemala

Mygona  Thieme, 1907  

Mygona irmina (E. Doubleday, [1849])    iN

Mygona irmina irmina (E. Doubleday, [1849])  TL: [Venezuela]  M T

Mygona irmina [n. ssp.#1] Pyrcz, MS  TL: Colombia

Mygona irmina [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Mygona prochyta (Hewitson, 1862)    iN  M T

Mygona chyprota (Grose-Smith, 1900)    iN  M T

Mygona thammi (Staudinger, 1876)    iN  M T

Mygona poeania (Hewitson, 1870)  TL: Ecuador

Mygona poeania poeania (Hewitson, 1870)  TL: Ecuador  M T

Mygona poeania magalyae Pyrcz, 2004  TL: Peru

Proboscis  Thieme, 1907  

Proboscis propylea (Hewitson, 1857)  TL: Colombia  M T

Proboscis pomarancia Pyrcz, 2004  TL: Peru  M T S

Apexacuta  Pyrcz, 2004  

Apexacuta orsedice (Hewitson, 1878)    iN

Apexacuta orsedice orsedice (Hewitson, 1878)  TL: Ecuador  M T

Apexacuta orsedice colombiana (E. Krüger, 1924)  TL: Colombia  M T S

Apexacuta orsedice violacea (Weymer, 1912)  TL: Ecuador  M T

Apexacuta orsedice [n. ssp.] Pyrcz & Clavijo, MS  Colombia

Apexacuta improvisa Pyrcz, 2004  TL: Peru

Apexacuta improvisa improvisa Pyrcz, 2004  TL: Peru  M L

Apexacuta improvisa [n. ssp.] Pyrcz & Boyer, MS  Peru

Apexacuta superior Pyrcz, 2004  TL: Peru

Apexacuta superior superior Pyrcz, 2004  TL: Peru  M T

Apexacuta superior [n. ssp.] Pyrcz & Boyer, MS  Peru

Apexacuta astoreth (Thieme, 1907)    iN

Apexacuta astoreth astoreth (Thieme, 1907)  TL: Bolivia  M T

Apexacuta astoreth [n. ssp.] Pyrcz & Boyer, MS  Peru

Apexacuta [n. sp.] Pyrcz & Boyer, MS  Peru

Lasiophila  C. Felder & R. Felder, 1859  

Lasiophila cirta C. Felder & R. Felder, 1859    iN

Lasiophila cirta cirta C. Felder & R. Felder, 1859  TL: Peru  M T

Lasiophila cirta atropurpurea Pyrcz, 2004  TL: Peru

Lasiophila cirta [n. ssp.#1] Pyrcz, MS  TL: Peru

Lasiophila cirta [n. ssp.#2] Pyrcz, MS  TL: Peru

Lasiophila lucuma Pyrcz, 2011  TL: not specified

Lasiophila lucuma lucuma Pyrcz, 2011  TL: not specified

Lasiophila lucuma [n. ssp.] Pyrcz, MS  Peru

Lasiophila piscina Thieme, 1903    iN

Lasiophila piscina piscina Thieme, 1903  TL: [Peru]  M T S

Lasiophila piscina [n. ssp.] Pyrcz, MS  Peru

Lasiophila luna Pyrcz, 2006    iN  M S

Lasiophila hewitsonia A. Butler, 1868  TL: Bolivia

Lasiophila hewitsonia hewitsonia A. Butler, 1868  TL: Bolivia  M T

Lasiophila hewitsonia neda Thieme, 1907  TL: Peru  M T

Lasiophila zapatoza (Westwood, 1851)    iN

Lasiophila zapatoza zapatoza (Westwood, 1851)  TL: "Bolivia"; Venezuela  M T

Lasiophila zapatoza meridae Adams & Bernard, 1981  TL: Venezuela  M T

Lasiophila zapatoza elusiva Pyrcz & Viloria, 2007  ref.285

Lasiophila zapatoza manaurera Adams & Bernard, 1979  TL: Colombia  M T

Lasiophila zapatoza sombra Thieme, 1907  TL: Colombia

Lasiophila zapatoza [n. ssp.] Pyrcz, MS  Venezuela

Lasiophila semipartita Weymer, 1912  TL: Colombia

Lasiophila prosymna (Hewitson, 1857)    iN

Lasiophila prosymna prosymna (Hewitson, 1857)  TL: Colombia  M T

Lasiophila prosymna dirempta Thieme, 1907  TL: Ecuador  M T

Lasiophila parthyene (Hewitson, 1872)    iN  M T

Lasiophila ciris Thieme, 1907    iN

Lasiophila ciris ciris Thieme, 1907  TL: Ecuador  M S

Lasiophila ciris [n. ssp.] Pyrcz, MS  Ecuador

Lasiophila circe C. Felder & R. Felder, 1859    iN

Lasiophila circe circe C. Felder & R. Felder, 1859  TL: Venezuela  M T S

Lasiophila circe farallonensis Pyrcz & Rodriguez, 2008  TL: not specified

Lasiophila circe cnephas Thieme, 1907  TL: Colombia

Lasiophila circe arithmetica Pyrcz & Viloria, 2007  ref.285

Lasiophila circe zarathustra Thieme, 1907  TL: Ecuador

Lasiophila circe [n. ssp.#1] Pyrcz, MS  TL: Colombia

Lasiophila circe [n. ssp.#2] Pyrcz, MS  TL: Colombia

Lasiophila persepolis (Hewitson, 1872)    iN

Lasiophila persepolis persepolis (Hewitson, 1872)  TL: Ecuador  M T

Lasiophila persepolis [n. ssp.] Pyrcz, MS  Ecuador

Lasiophila palades (Hewitson, 1872)    iN

Lasiophila palades palades (Hewitson, 1872)  TL: Ecuador  M T

Lasiophila palades limes Pyrcz, 1999  TL: Ecuador

Lasiophila phalaesia (Hewitson, 1868)    iN

Lasiophila phalaesia phalaesia (Hewitson, 1868)  TL: Ecuador  M T

Lasiophila phalaesia alce Pyrcz, 1999  TL: Ecuador

Lasiophila phalaesia confusa Staudinger, 1897  TL: [Ecuador]  M T

Lasiophila phalaesia kroli Pyrcz, 1999  TL: Ecuador

Lasiophila regia Staudinger, 1897  TL: Bolivia  M T

Lasiophila orbifera A. Butler, 1868    iN

Lasiophila orbifera orbifera A. Butler, 1868  TL: "Bolivia"; "Colombia"; Peru  M T S

Lasiophila orbifera munda Thieme, 1907  TL: Peru

Lasiophila orbifera intercepta Thieme, 1907  TL: Ecuador

Lasiophila orbifera pura Thieme, 1907  TL: Argentina

Lasiophila alkaios Tessmann, 1928  TL: Peru

Lasiophila alkaios alkaios Tessmann, 1928  TL: Peru  M T

Lasiophila alkaios gita Smart, 1975  TL: Peru

Lasiophila alkaios [n. ssp.] Pyrcz, MS  Peru

Lasiophila [n. sp.] Pyrcz & Gareca, MS  Bolivia

Drucina  A. Butler, 1872  

Drucina championi Godman & Salvin, 1881  Blue-spotted Satyr    iN

Drucina championi championi Godman & Salvin, 1881  S Mexico (Chiapas) & Guatemala  M T S IL L

Drucina championi [n. ssp.] Pyrcz, MS  Mexico

Drucina leonata A. Butler, 1872  Leonata Satyr  Costa Rica & Panama  iN  M T S L

Druphila  Pyrcz, 2004  

Druphila venerata (A. Butler, 1873)    iN

Druphila venerata venerata (A. Butler, 1873)  TL: Peru  M T

Druphila venerata [n. ssp.] Pyrcz, MS  Peru

Druphila [n. sp.] Pyrcz, MS  Peru

Foetterleia  Viloria, 2004  

Foetterleia schreineri (Foetterle, 1902)    iN  M T

Pseudomaniola  Röber, 1889  

Pseudomaniola phaselis (Hewitson, 1862)    iN

Pseudomaniola phaselis phaselis (Hewitson, 1862)  TL: Venezuela  M T S

Pseudomaniola phaselis pholoe (Staudinger, 1887)  TL: Colombia  M T S

Pseudomaniola phaselis rogersi (Godman & Salvin, 1878)  Costa Rica & W Panama  M T S IL L

Pseudomaniola phaselis macasica (Strand, 1912)  TL: Ecuador  M T

Pseudomaniola phaselis argyritis (Thieme, 1907)  TL: Bolivia  M T

Pseudomaniola gigas (Godman & Salvin, 1877)  Orange-bordered Satyr  S Mexico & Guatemala  iN  M T S

Pseudomaniola loxo (Dognin, 1891)    iN

Pseudomaniola loxo loxo (Dognin, 1891)  TL: Colombia  M T

Pseudomaniola loxo [n. ssp.] Pyrcz, MS  Ecuador

Pseudomaniola ilsa (Thieme, 1907)  TL: Colombia  M T

Pseudomaniola mirabilis (A. Butler, 1873)    iN

Pseudomaniola mirabilis mirabilis (A. Butler, 1873)  TL: Peru  M T

Pseudomaniola mirabilis extrema Pyrcz, 2004  TL: Peru

Pseudomaniola gerlinda (Thieme, 1907)    iN  M T S

Pseudomaniola mena (Grose-Smith, 1900)  TL: Bolivia  M T

Pseudomaniola clethra (Thieme, 1907)  TL: Peru  M T

Pseudomaniola helche (Thieme, 1907)  TL: Bolivia  M T

Pseudomaniola asuba (Thieme, 1907)  TL: Bolivia  M S

Pseudomaniola [n. sp.] Lamas, MS  Peru

Arhuaco  Adams & Bernard, 1977  

Arhuaco ica Adams & Bernard, 1977  TL: Colombia  M T

Arhuaco dryadina (Schaus, 1913)  Dryadina Satyr  Costa Rica & W Panama  M T

Cheimas  Thieme, 1907  

Cheimas opalinus (Staudinger, 1897)  TL: Venezuela

Cheimas opalinus opalinus (Staudinger, 1897)  TL: Venezuela  M T

Cheimas opalinus cristalinus Pyrcz & Boyer, 2018  ref.283

Cheimas opalinus dominici Pyrcz & Boyer, 2018  ref.283

Cheimas opalinus iosephi Pyrcz & Boyer, 2018  ref.283

Cheimas opalinus rosalinus Pyrcz, 2018  ref.283

Steroma  Westwood, [1850]  

Steroma bega Westwood, [1850]    iN

Steroma bega bega Westwood, [1850]  TL: Venezuela  M T

Steroma bega andensis C. Felder & R. Felder, 1867  TL: Colombia; "Ecuador"  M T S L

Steroma modesta Weymer, 1912    iN  M T S

Steroma polyxo (Godman & Salvin, 1880)    iN  M T

Steroma superba A. Butler, 1868    iN

Steroma superba superba A. Butler, 1868  TL: Bolivia  M T S

Steroma superba [n. ssp.] Pyrcz, MS  Peru

Steremnia  Thieme, 1905  

Steremnia lucillae Pyrcz, 2004  TL: not specified  M S

Steremnia monachella (Weymer, 1911)    iN

Steremnia monachella monachella (Weymer, 1911)  TL: Peru  M T S

Steremnia monachella adamsiella Pyrcz & Boyer, 2011  TL: Colombia

Steremnia monachella [n. ssp.#1] Pyrcz, MS  TL: Peru

Steremnia monachella [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Steremnia pronophila (C. Felder & R. Felder, 1867)    iN

Steremnia pronophila pronophila (C. Felder & R. Felder, 1867)  TL: Colombia; "Ecuador"  M T

Steremnia pronophila fluminea Pyrcz & Viloria, 2007  ref.285

Steremnia selva Adams, 1986  TL: Colombia

Steremnia selva selva Adams, 1986  TL: Colombia  M T

Steremnia selva [n. ssp.] Pyrcz, MS  Colombia

Steremnia umbracina (A. Butler, 1873)    iN

Steremnia umbracina umbracina (A. Butler, 1873)  TL: Peru  M T

Steremnia umbracina misella Thieme, 1905  TL: Peru

Steremnia agraulis (Weymer, 1912)  TL: Bolivia

Steremnia agraulis agraulis (Weymer, 1912)  TL: Bolivia  M T

Steremnia agraulis agraulina Pyrcz, MS  TL: Peru

Steremnia rugilas Thieme, 1905  TL: [Peru]  M T

Steremnia [n. sp.#1] Pyrcz, MS  TL: Ecuador

Steremnia [n. sp.#2] Pyrcz, MS  TL: not specified

Steremnia [n. sp.#2] [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Steremnia [n. sp.#2] [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Steremnia [n. sp.#3] Pyrcz, MS  TL: Ecuador

Eteona  E. Doubleday, 1848  

Eteona tisiphone (Boisduval, 1836)    iN  M T

Eretris  Thieme, 1905  

Eretris oculata (C. Felder & R. Felder, 1867)    iN  M T

Eretris suzannae DeVries, 1980  Suzanne's Satyr  Costa Rica & W Panama  iN  M T S

Eretris lecromi Pyrcz, 1999    iN  M S

Eretris depresissima Pyrcz, 1999  TL: Colombia

Eretris depresissima depresissima Pyrcz, 1999  TL: Colombia

Eretris depresissima niambii Pyrcz, 1999  TL: Colombia

Eretris agata Pyrcz & Fratello, 2005  TL: not specified

Eretris encycla (C. Felder & R. Felder, 1867)  TL: Venezuela

Eretris encycla encycla (C. Felder & R. Felder, 1867)  TL: Venezuela  M T

Eretris encycla [n. ssp.#1] Pyrcz, MS  TL: Venezuela

Eretris encycla [n. ssp.#2] Pyrcz, MS  TL: Colombia

Eretris encycla [n. ssp.#3] Pyrcz & Willmott, MS  TL: Ecuador

Eretris neocycla Pyrcz & Viloria, 2009  TL: not specified

Eretris calisto (C. Felder & R. Felder, 1867)    iN

Eretris calisto calisto (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Eretris calisto kogui Adams & Bernard, 1977  TL: Colombia  M T

Eretris calisto [n. ssp.] Pyrcz, MS  Colombia

Eretris cuaoensis Viloria & Attal, 2019  ref.192

Eretris ocellifera (C. Felder & R. Felder, 1867)    iN

Eretris ocellifera ocellifera (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Eretris ocellifera [n. ssp.#1] Pyrcz & Willmott, MS  TL: Colombia

Eretris ocellifera [n. ssp.#2] Pyrcz & Willmott, MS  TL: Colombia

Eretris hulda (A. Butler & H. Druce, 1872)  Hulda Satyr  Costa Rica & Panama  iN  M T S IL

Eretris maria (Schaus, 1920)  Maria’s Satyr  S Mexico & Guatemala  iN  M T S

Eretris centralis E. Krüger, 1924  TL: Colombia  M T S

Eretris porphyria (C. Felder & R. Felder, 1867)    iN

Eretris porphyria porphyria (C. Felder & R. Felder, 1867)  TL: Venezuela  M T

Eretris porphyria decorata (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Eretris porphyria catargyrea (Staudinger, 1887)  TL: Colombia  M T

Eretris porphyria perija Adams & Bernard, 1979  TL: Colombia  M T S

Eretris porphyria recta Pyrcz & Viloria, 2007  ref.285

Eretris porphyria pseudoperija Pyrcz, 1999  TL: Colombia

Eretris porphyria transmaraniona Pyrcz, 2004  TL: Peru

Eretris porphyria [n. ssp.#1] Pyrcz, MS  TL: Venezuela

Eretris porphyria [n. ssp.#2] Pyrcz, MS  TL: Venezuela

Eretris porphyria [n. ssp.#3] Pyrcz, MS  TL: Colombia

Eretris porphyria [n. ssp.#4] Pyrcz, MS  TL: Colombia

Eretris apuleja (C. Felder & R. Felder, 1867)    iN

Eretris apuleja apuleja (C. Felder & R. Felder, 1867)  TL: Venezuela  M T

Eretris apuleja subrufescens (Grose-Smith & W. F. Kirby, 1895)  TL: Costa Rica  M T

Eretris apuleja bogotana E. Krüger, 1924  TL: [Colombia]  M T S

Eretris apuleja ochrea Thieme, 1905  TL: Ecuador  M T

Eretris apuleja altamira Adams & Bernard, 1979  TL: Colombia  M T

Eretris apuleja remotissima Pyrcz & Viloria, 2007  ref.285

Eretris apuleja socorrensis Pyrcz & Rodriguez, 2008  TL: not specified

Eretris apuleja [n. ssp.#1] Pyrcz & Willmott, MS  TL: Ecuador

Eretris apuleja [n. ssp.#2] Pyrcz & Willmott, MS  TL: Ecuador

Eretris apuleja [n. ssp.#3] Pyrcz & Willmott, MS  TL: Ecuador

Eretris apuleja [n. ssp.#4] Pyrcz, MS  TL: Colombia

Eretris apuleja [n. ssp.#5] Pyrcz, MS  TL: Colombia

Eretris microrufescens Pyrcz & Wojtusiak, 2010  TL: not specified

Eretris suprarufescens Pyrcz & Wojtusiak, 2010  TL: not specified

Eretris apuleina (Pyrcz, 2004)    iN

Eretris apuleina apuleina (Pyrcz, 2004)  TL: Ecuador  M S

Eretris apuleina [n. ssp.] Pyrcz, MS  Peru

Eretris truncatina (Pyrcz, 2004)  TL: Peru

Eretris truncatina truncatina (Pyrcz, 2004)  TL: Peru  M T

Eretris truncatina [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris truncatina [n. ssp.#2] Pyrcz, MS  TL: Peru

Eretris subpunctata (Grose-Smith & W. F. Kirby, 1895)    iN

Eretris subpunctata subpunctata (Grose-Smith & W. F. Kirby, 1895)  TL: Bolivia  M T S L

Eretris subpunctata [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris subpunctata [n. ssp.#2] Pyrcz, MS  TL: Peru

Eretris mendoza (Pyrcz, 2004)    iN

Eretris mendoza mendoza (Pyrcz, 2004)  TL: Peru  M T

Eretris mendoza [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris mendoza [n. ssp.#2] Pyrcz, MS  TL: Peru

Eretris julieta Pyrcz & Gareca, 2010  TL: not specified

Eretris julieta julieta Pyrcz & Gareca, 2010  TL: not specified

Eretris julieta [n. ssp.] Pyrcz, MS  Bolivia

Eretris [n. sp.#1] Pyrcz, MS  TL: Bolivia

Eretris [n. sp.#2] Pyrcz, MS  TL: Ecuador

Eretris [n. sp.#3] Pyrcz, MS  TL: not specified

Eretris [n. sp.#3] [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris [n. sp.#3] [n. ssp.#2] Pyrcz, MS  TL: Bolivia

Eretris [n. sp.#3] [n. ssp.#3] Pyrcz, MS  TL: Peru

Eretris [n. sp.#4] Pyrcz, MS  TL: not specified

Eretris [n. sp.#4] [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris [n. sp.#4] [n. ssp.#2] Pyrcz, MS  TL: Peru

Eretris [n. sp.#4] [n. ssp.#3] Pyrcz, MS  TL: Peru

Eretris [n. sp.#4] [n. ssp.#4] Pyrcz, MS  TL: Peru

Eretris [n. sp.#4] [n. ssp.#5] Pyrcz, MS  TL: Bolivia

Eretris [n. sp.#5] Pyrcz, MS  TL: Bolivia

Eretris [n. sp.#6] Pyrcz, MS  TL: not specified

Eretris [n. sp.#6] [n. ssp.#1] Pyrcz, MS  TL: Peru

Eretris [n. sp.#6] [n. ssp.#2] Pyrcz, MS  TL: Peru

Eretris [n. sp.#6] [n. ssp.#3] Pyrcz, MS  TL: Peru

Eretris [n. sp.#6] [n. ssp.#4] Pyrcz, MS  TL: Bolivia

Manerebia  Staudinger, 1897  

Manerebia leaena (Hewitson, 1861)    iN

Manerebia leaena leaena (Hewitson, 1861)  TL: [Ecuador]  M T

Manerebia leaena lanassa (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Manerebia leaena gonzalezi Pyrcz & Viloria, 2006  ref.280

Manerebia inderena (Adams, 1986)    iN

Manerebia inderena inderena (Adams, 1986)  TL: Colombia  M T S

Manerebia inderena antioquiana Pyrcz & Willmott, 2006  ref.280

Manerebia inderena fina Pyrcz & Willmott, 2006  ref.280

Manerebia inderena similis Pyrcz & Willmott, 2006  ref.280

Manerebia inderena clara Pyrcz & Willmott, 2006  ref.280  M L

Manerebia inderena leaeniva Pyrcz & Willmott, 2006  ref.280

Manerebia inderena mirena Pyrcz & Willmott, 2006  ref.280

Manerebia inderena [n. ssp.] Pyrcz, MS  Ecuador

Manerebia prattorum Pyrcz & Willmott, 2006  ref.280

Manerebia prattorum prattorum Pyrcz & Willmott, 2006  TL: Peru  M S

Manerebia prattorum udimaense Pyrcz, MS  TL: Peru

Manerebia golondrina Pyrcz & Willmott, 2006  ref.280

Manerebia trimaculata (Hewitson, 1870)  TL: Ecuador

Manerebia trimaculata trimaculata (Hewitson, 1870)  TL: Ecuador  M T

Manerebia trimaculata [n. ssp.] Pyrcz, MS  Ecuador

Manerebia undulata Pyrcz & J. Hall, 2006  TL: Ecuador

Manerebia undulata undulata Pyrcz & J. Hall, 2006  ref.280

Manerebia undulata milaena Pyrcz & Willmott, 2006  ref.280

Manerebia interrupta (F. Brown, 1944)  TL: Ecuador  M T

Manerebia rufanalis Pyrcz & J. Hall, 2006  TL: Ecuador

Manerebia rufanalis rufanalis Pyrcz & J. Hall, 2006  ref.280

Manerebia rufanalis fernandina Pyrcz & Willmott, 2006  ref.280

Manerebia quinterae (Adams & Bernard, 1979)  TL: Venezuela  M T S

Manerebia navarrae (Adams & Bernard, 1979)  TL: Colombia  M T S

Manerebia satura (Weymer, 1911)  TL: Peru; "Colombia"

Manerebia satura satura (Weymer, 1911)  TL: Peru; "Colombia"

Manerebia satura keithi (Dyar, 1913)  TL: Peru  M T

Manerebia satura pauperata Pyrcz & Willmott, 2006  ref.280

Manerebia satura lamasi Pyrcz & Willmott, 2006  ref.280

Manerebia germaniae Pyrcz & J. Hall, 2006  TL: Ecuador

Manerebia germaniae germaniae Pyrcz & J. Hall, 2006  ref.280

Manerebia germaniae vitalei Pyrcz & Willmott, 2006  ref.280

Manerebia mammuthus Pyrcz & Willmott, 2006  ref.280

Manerebia franciscae (Adams & Bernard, 1981)  TL: Venezuela

Manerebia franciscae franciscae (Adams & Bernard, 1981)  TL: Venezuela  M T

Manerebia franciscae rodriguezi Pyrcz & Willmott, 2006  ref.280

Manerebia nevadensis E. Krüger, 1925    iN  M T S

Manerebia mycalesoides (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Manerebia magnifica Pyrcz & Willmott, 2006  ref.280  M T

Manerebia levana (Godman, 1905)  TL: Colombia

Manerebia levana levana (Godman, 1905)  TL: Colombia  M T

Manerebia levana pervaga Pyrcz & Viloria, 2007  TL: Venezuela

Manerebia apiculata (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Manerebia ignilineata (Dognin, 1896)  TL: Ecuador

Manerebia ignilineata ignilineata (Dognin, 1896)  TL: Ecuador  M T

Manerebia ignilineata neglecta (F. Brown, 1944)  TL: Ecuador  M T

Manerebia ignilineata jalca Pyrcz, 2004  TL: Peru

Manerebia seducta Pyrcz & Willmott, 2006  ref.280  M T

Manerebia diffusa Pyrcz, 2004  TL: Peru  M T

Manerebia zoippus (H. Druce, 1876)  TL: Peru  M T

Manerebia benigni Pyrcz, 2004    iN

Manerebia benigni benigni Pyrcz, 2004  TL: Peru  M T

Manerebia benigni tessmanni Pyrcz, 2004  TL: Peru  M T

Manerebia trirufa Pyrcz, 2004  TL: Peru

Manerebia trirufa trirufa Pyrcz, 2004  TL: Peru  M T

Manerebia trirufa [n. ssp.] Pyrcz, MS  Peru

Manerebia haywardi Pyrcz, 2004  TL: Peru

Manerebia haywardi haywardi Pyrcz, 2004  TL: Peru  M S

Manerebia haywardi [n. ssp.] Pyrcz, MS  Peru

Manerebia rubescens (A. Butler, 1873)    iN  M T

Manerebia cyclopella Staudinger, 1897    iN  M T S

Manerebia cyclops Staudinger, 1897  TL: Bolivia  M T

Manerebia cyclopina Staudinger, 1897    iN

Manerebia cyclopina cyclopina Staudinger, 1897  TL: Peru  M T

Manerebia cyclopina staudingeri Forster, 1964  TL: Bolivia  M T S

Manerebia monops Hayward, 1949  TL: Argentina  M T

Manerebia insulsa (Hewitson, 1874)  TL: Bolivia  M T

Manerebia typhlops Staudinger, 1897  TL: Bolivia  M T

Manerebia pluviosa Pyrcz & Viloria, 2007  ref.280

Manerebia reducta (F. Brown, 1944)  TL: Peru  M T

Manerebia [n. sp.#1] Pyrcz, MS  TL: not specified

Manerebia [n. sp.#1] [n. ssp.#1] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#1] [n. ssp.#2] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#2] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#3] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#4] Pyrcz, MS  TL: not specified

Manerebia [n. sp.#4] [n. ssp.#1] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#4] [n. ssp.#2] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#5] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#6] Pyrcz, MS  TL: not specified

Manerebia [n. sp.#6] [n. ssp.#1] Pyrcz, MS  TL: Peru

Manerebia [n. sp.#6] [n. ssp.#2] Pyrcz, MS  TL: Bolivia

Manerebia [n. sp.#7] Pyrcz, MS  TL: Argentina

Diaphanos  Adams & Bernard, 1981  

Diaphanos huberi Adams & Bernard, 1981  TL: Venezuela

Diaphanos huberi huberi Adams & Bernard, 1981  TL: Venezuela  M T

Diaphanos huberi [n. ssp.] Pyrcz & Viloria, MS  Venezuela

Diaphanos curvignathos Viloria, 1994  TL: Venezuela

Diaphanos curvignathos curvignathos Viloria, 1994  TL: Venezuela

Diaphanos curvignathos [n. ssp.] Pyrcz & Viloria, MS  Venezuela

Diaphanos fuscus Viloria, 1994  TL: Venezuela

Diaphanos [n. sp.] Viloria, MS  Venezuela

Idioneurula  Strand, 1932  

Idioneurula erebioides (C. Felder & R. Felder, 1867)    iN

Idioneurula erebioides erebioides (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Idioneurula erebioides socorroi Villalobos-Moreno & Salazar, 2013  TL: not specified

Idioneurula donegani (Huertas & Arias, 2008)    iN

Idioneurula eremita Pyrcz & Viloria, 2007  ref.285

Idioneurula jacquelinae (Pyrcz, 1995)  TL: Venezuela  M T

Lymanopoda  Westwood, 1851  

Lymanopoda panacea (Hewitson, 1869)    iN

Lymanopoda panacea panacea (Hewitson, 1869)  TL: Ecuador  M T

Lymanopoda panacea gortyna (Weymer, 1890)  TL: Colombia  M T

Lymanopoda panacea ocellifera A. Butler, 1873  TL: Peru  M T S

Lymanopoda panacea venusia Hopffer, 1874  TL: Peru

Lymanopoda affineola Weymer, 1911    iN  M T

Lymanopoda albomaculata Hewitson, 1861    iN

Lymanopoda albomaculata albomaculata Hewitson, 1861  TL: [Bolivia]  M T S

Lymanopoda albomaculata [n. ssp.] Pyrcz, MS  Peru

Lymanopoda albocincta Hewitson, 1861    iN

Lymanopoda albocincta albocincta Hewitson, 1861  TL: Colombia  M T

Lymanopoda albocincta issacha A. Butler, 1870  TL: Cuba  M T

Lymanopoda albocincta intermedia Pyrcz, 2004  TL: Peru

Lymanopoda albocincta [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Lymanopoda albocincta [n. ssp.#2] Pyrcz, MS  TL: Peru

Lymanopoda florenciaensis Salazar, Henao & Vargas, 2004    iN

Lymanopoda apulia Hopffer, 1874    iN  M S

Lymanopoda caucana Weymer, 1911    iN

Lymanopoda caucana caucana Weymer, 1911  TL: Colombia

Lymanopoda caucana orientalis Viloria & Camacho, 1999  TL: Venezuela

Lymanopoda caeruleata Godman & Salvin, 1880    iN  M T

Lymanopoda euopis Godman & Salvin, 1878  Costa Rican Satyr  Costa Rica  iN  M T TI S L

Lymanopoda cinna Godman & Salvin, 1889  Blue-stained Satyr  S Mexico (Chiapas) & Guatemala  M T S L

Lymanopoda acraeida A. Butler, 1868    iN

Lymanopoda acraeida acraeida A. Butler, 1868  TL: Bolivia  M T

Lymanopoda acraeida zigomala (Hewitson, 1877)  TL: Ecuador  M T

Lymanopoda acraeida chavezi Pyrcz, 2004  TL: Peru

Lymanopoda acraeida malia Godman, 1905  TL: Peru  M T S

Lymanopoda acraeida [n. ssp.] Pyrcz, MS  Peru

Lymanopoda venosa A. Butler, 1868    iN  M T S

Lymanopoda obsoleta (Westwood, 1851)    iN  M T

Lymanopoda altis Weymer, 1890    iN  M T

Lymanopoda maletera Adams & Bernard, 1979  TL: Venezuela  M T

Lymanopoda dietzi Adams & Bernard, 1981  TL: Venezuela

Lymanopoda dietzi dietzi Adams & Bernard, 1981  TL: Venezuela  M T S

Lymanopoda dietzi neildi Pyrcz, 2010  TL: not specified

Lymanopoda dietzi vetula Viloria & Pyrcz, 2010  TL: not specified

Lymanopoda dietzi oliva Pyrcz & Boyer, 2010  TL: not specified

Lymanopoda dietzi rosanna Viloria & Pyrcz, 2010  TL: Peru

Lymanopoda dietzi argentata Pyrcz, 2010  TL: not specified

Lymanopoda dietzi [n. ssp.] Pyrcz, MS  Venezuela

Lymanopoda confusa F. Brown, 1943    iN  M T

Lymanopoda lecromi Pyrcz & Viloria, 2007  ref.285

Lymanopoda confusiana Pyrcz & Boyer, 2010  TL: not specified

Lymanopoda magna Pyrcz, 2004  TL: Peru

Lymanopoda caudalis Rosenberg & Talbot, 1914  TL: Peru

Lymanopoda caudalis caudalis Rosenberg & Talbot, 1914  TL: Peru  M T

Lymanopoda caudalis raymondae Pyrcz & Boyer, 2010  TL: Venezuela

Lymanopoda balintiana Pyrcz & Boyer, 2010  TL: not specified

Lymanopoda inaudita Pyrcz, 2010    iN

Lymanopoda necalba Pyrcz & Boyer, 2010  TL: not specified

Lymanopoda marianna Staudinger, 1897  TL: Venezuela

Lymanopoda marianna marianna Staudinger, 1897  TL: Venezuela  M T

Lymanopoda marianna veronicae Pyrcz, 2010  TL: not specified

Lymanopoda marianna giselae Pyrcz, 2010  TL: not specified

Lymanopoda marianna ioanae Pyrcz, 2010  TL: not specified

Lymanopoda marianna jacquelinae Pyrcz, 2010  TL: not specified

Lymanopoda marianna valentinae Pyrcz, 2010  TL: not specified

Lymanopoda labda Hewitson, 1861    iN

Lymanopoda labda labda Hewitson, 1861  TL: Colombia  M T

Lymanopoda labda moritzi Pyrcz & Viloria, 2007  TL: Venezuela

Lymanopoda labda lebbaea C. Felder & R. Felder, 1867  TL: Colombia  M T

Lymanopoda labda bilinskii Pyrcz, 2011  TL: not specified

Lymanopoda altaselva Adams & Bernard, 1979  TL: Venezuela  M T

Lymanopoda nadia Pyrcz, 1999    iN  M T

Lymanopoda ionius Westwood, 1851    iN

Lymanopoda ionius ionius Westwood, 1851  TL: Colombia  M T

Lymanopoda ionius excisa Weymer, 1911  TL: Colombia

Lymanopoda ionius browni Pyrcz, 1999  TL: Ecuador

Lymanopoda ionius gargantua Pyrcz & Rodriguez, 2008  TL: not specified

Lymanopoda ionius [n. ssp.#1] Pyrcz, MS  TL: Colombia

Lymanopoda ionius [n. ssp.#2] Pyrcz, MS  TL: Colombia

Lymanopoda lactea Hewitson, 1861  TL: Colombia  M T S

Lymanopoda schmidti Adams, 1986  TL: Colombia  M T S

Lymanopoda labineta Hewitson, 1870    iN

Lymanopoda labineta labineta Hewitson, 1870  TL: Ecuador  M T

Lymanopoda labineta piniasi Pyrcz, 1999  TL: Ecuador

Lymanopoda labineta decorata Seydel, 1924  TL: Colombia  M T

Lymanopoda labineta wojtusiaki Pyrcz, 2011  TL: Ecuador

Lymanopoda pieridina Röber, 1927    iN

Lymanopoda pieridina pieridina Röber, 1927  TL: Colombia  M T

Lymanopoda pieridina albicosta Pyrcz, 1999  TL: Colombia

Lymanopoda paisa Pyrcz & Rodriguez, 2006  TL: not specified

Lymanopoda nivea Staudinger, 1887    iN

Lymanopoda nivea nivea Staudinger, 1887  TL: Ecuador  M T

Lymanopoda nivea sororcula Thieme, 1904  TL: Ecuador

Lymanopoda nivea bingo Pyrcz, 1999  TL: Ecuador

Lymanopoda nivea bonita Pyrcz, 1999  TL: Ecuador

Lymanopoda samius Westwood, 1851    iN

Lymanopoda samius samius Westwood, 1851  TL: Colombia  M T

Lymanopoda samius lineana Pyrcz & Viloria, 2007  ref.285

Lymanopoda ferruginosa A. Butler, 1868    iN

Lymanopoda ferruginosa ferruginosa A. Butler, 1868  TL: Bolivia  M T S

Lymanopoda ferruginosa [n. ssp.] Pyrcz, MS  Peru

Lymanopoda prusia (Heimlich, 1973)    iN  M S

Lymanopoda rana Weymer, 1911    iN

Lymanopoda rana rana Weymer, 1911  TL: Peru  M S

Lymanopoda rana marrana Pyrcz & Boyer, 2011  TL: not specified

Lymanopoda hyagnis Weymer, 1911    iN  M T S

Lymanopoda translucida Weymer, 1912  TL: Bolivia

Lymanopoda translucida translucida Weymer, 1912  TL: Bolivia

Lymanopoda translucida umbratilis Rosenberg & Talbot, 1914  TL: Peru  M T

Lymanopoda translucida leucotecta Rosenberg & Talbot, 1914  TL: Peru  M T

Lymanopoda hockingiana Pyrcz, 2011  TL: not specified

Lymanopoda araneola Pyrcz, 2004  TL: Peru  M S

Lymanopoda galactea Staudinger, 1897  TL: Bolivia  M T

Lymanopoda shefteli Dyar, 1913    iN

Lymanopoda shefteli shefteli Dyar, 1913  TL: Peru  M T

Lymanopoda shefteli siviae Fuchs, 1954  TL: Peru

Lymanopoda shefteli broweri Pyrcz & Boyer, 2011  TL: not specified

Lymanopoda mariposa Pyrcz & Boyer, 2011    iN

Lymanopoda ingasayana Pyrcz, 2004  TL: Peru

Lymanopoda ingasayana ingasayana Pyrcz, 2004  TL: Peru

Lymanopoda ingasayana [n. ssp.] Pyrcz, MS  Peru

Lymanopoda dyari Pyrcz, 2004  TL: Peru

Lymanopoda inde Pyrcz, 2004  TL: Peru

Lymanopoda inde inde Pyrcz, 2004  TL: Peru  M S

Lymanopoda inde [n. ssp.] Pyrcz, MS  Peru

Lymanopoda eubagioides A. Butler, 1873    iN  M T S

Lymanopoda hazelana F. Brown, 1943  TL: Ecuador

Lymanopoda hazelana hazelana F. Brown, 1943  TL: Ecuador  M T

Lymanopoda hazelana summa Pyrcz, 1999  TL: Ecuador

Lymanopoda hazelana viridia Pyrcz, 2011  TL: Ecuador

Lymanopoda melia Weymer, 1911    iN

Lymanopoda melia melia Weymer, 1911  TL: Colombia  M T

Lymanopoda melia alba E. Krüger, 1924  TL: Colombia  M T S

Lymanopoda melia [n. ssp.#1] Pyrcz, MS  TL: Colombia

Lymanopoda melia [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Lymanopoda pascua Pyrcz, 2005  TL: not specified

Lymanopoda tolima Weymer, 1911    iN

Lymanopoda huilana Weymer, 1890    iN

Lymanopoda huilana huilana Weymer, 1890  TL: Colombia  M T

Lymanopoda huilana cajamarca Adams, 1986  TL: Colombia  M T S

Lymanopoda huilana dominicae Pyrcz, MS  TL: not specified

Lymanopoda huilana [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Lymanopoda huilana [n. ssp.#2] Pyrcz, MS  TL: Ecuador

Lymanopoda huilana [n. ssp.#3] Pyrcz, MS  TL: Ecuador

Lymanopoda huilana [n. ssp.#4] Pyrcz, MS  TL: Colombia

Lymanopoda caracara Pyrcz, Willmott & J. Hall, 1999  TL: Ecuador

Lymanopoda caracara caracara Pyrcz, Willmott & J. Hall, 1999  TL: Ecuador

Lymanopoda caracara ichu Pyrcz, Willmott & J. Hall, 1999  TL: Ecuador  M S

Lymanopoda caracara [n. ssp.] Pyrcz, Willmott, MS  Ecuador

Lymanopoda viventieni (Apolinar, 1924)  TL: Colombia  M S

Lymanopoda mirabilis (Staudinger, 1897)  TL: Colombia  M T

Lymanopoda paramera Adams & Bernard, 1979  TL: Colombia  M T

Lymanopoda nevada E. Krüger, 1924    iN

Lymanopoda nevada nevada E. Krüger, 1924  TL: Colombia  M T S

Lymanopoda nevada nevadensis (A. Schultze, 1931)  TL: Colombia

Lymanopoda melendeza Adams, 1986  TL: Colombia  M T S

Lymanopoda [n. sp.#1] Pyrcz, MS  TL: Ecuador

Lymanopoda [n. sp.#2] Pyrcz, MS  TL: Ecuador

Lymanopoda [n. sp.#3] Huertas, MS  TL: Colombia

Lymanopoda [n. sp.#4] Pyrcz & Clavijo, MS  TL: Colombia

Lymanopoda [n. sp.#5] Pyrcz, MS  TL: Peru

Ianussiusa  Pyrcz & Viloria, 2004  

Ianussiusa maso (Godman, 1905)    iN

Ianussiusa maso maso (Godman, 1905)  TL: Colombia  M T

Ianussiusa maso santa Pyrcz & Viloria, 2007  ref.285

Ianussiusa maso polyommatus (Röber, 1927)  TL: Colombia  M T

Ianussiusa maso [n. ssp.#1] Pyrcz, MS  TL: Ecuador

Ianussiusa maso [n. ssp.#2] Pyrcz, MS  TL: Colombia

Redonda  Adams & Bernard, 1981  

Redonda empetrus (Thieme, 1905)  TL: Venezuela

Redonda empetrus empetrus (Thieme, 1905)  TL: Venezuela

Redonda empetrus bolivari Adams & Bernard, 1981  TL: Venezuela  M T

Redonda empetrus [n. ssp.] Pyrcz & Viloria, MS  Venezuela

Redonda bordoni Viloria & Pyrcz, 2003    iN

Redonda [n. sp.#1] Viloria & Pyrcz, MS  TL: not specified

Redonda [n. sp.#1] [n. ssp.#1] Pyrcz & Viloria, MS  TL: Venezuela

Redonda [n. sp.#1] [n. ssp.#2] Pyrcz & Viloria, MS  TL: Venezuela

Redonda [n. sp.#1] [n. ssp.#3] Pyrcz & Viloria, MS  TL: Venezuela

Redonda [n. sp.#2] Viloria & Pyrcz, MS  TL: Venezuela

Redonda [n. sp.#3] Pyrcz, MS  TL: Venezuela

Dangond  Adams & Bernard, 1979  

Dangond dangondi Adams & Bernard, 1979    iN  M T

Altopedaliodes  Forster, 1964  

Altopedaliodes cocytia (C. Felder & R. Felder, 1867)    iN

Altopedaliodes cocytia cocytia (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Altopedaliodes cocytia tamaensis Viloria & Pyrcz, 2007  TL: Colombia

Altopedaliodes nebris (Thieme, 1905)    iN  M T

Altopedaliodes reissi (Weymer, 1890)  TL: Colombia

Altopedaliodes reissi reissi (Weymer, 1890)  TL: Colombia  M T

Altopedaliodes reissi flavomaculata (E. Krüger, 1924)  TL: Colombia  M T S

Altopedaliodes reissi salazari Le Crom, 1994  TL: Colombia  M T

Altopedaliodes reissi [n. ssp.] Pyrcz & Willmott, MS  Ecuador

Altopedaliodes kruegeri Pyrcz, 1999  TL: Colombia

Altopedaliodes kruegeri kruegeri Pyrcz, 1999  TL: Colombia  M T

Altopedaliodes kruegeri belmira Pyrcz & Rodriguez, 2004  TL: not specified

Altopedaliodes scydmaena (Hayward, 1968)  TL: Ecuador  M T S

Altopedaliodes tena (Hewitson, 1869)    iN

Altopedaliodes tena tena (Hewitson, 1869)  TL: Ecuador  M T

Altopedaliodes tena [n. ssp.] Pyrcz, MS  Ecuador

Altopedaliodes nucea Pyrcz & Viloria, 1999  TL: Ecuador

Altopedaliodes pasicles (Hewitson, 1872)  TL: Ecuador  M T

Altopedaliodes perita (Hewitson, 1868)  TL: Ecuador

Altopedaliodes perita perita (Hewitson, 1868)  TL: Ecuador  M T

Altopedaliodes perita sorda Pyrcz, 2004  TL: Ecuador

Altopedaliodes zsolti Pyrcz & Viloria, 1999  TL: Ecuador

Altopedaliodes zsolti zsolti Pyrcz & Viloria, 1999  TL: Ecuador  M T

Altopedaliodes zsolti citra Pyrcz, 2004  TL: Ecuador

Altopedaliodes kurti Pyrcz & Viloria, 1999  TL: Ecuador  M T

Altopedaliodes [n. sp.#1] Pyrcz, MS  TL: Colombia

Altopedaliodes [n. sp.#2] Pyrcz & Willmott, MS  TL: Ecuador

Punapedaliodes  Forster, 1964  

Punapedaliodes flavopunctata (Staudinger, 1894)    iN

Punapedaliodes flavopunctata flavopunctata (Staudinger, 1894)  TL: Bolivia  M T

Punapedaliodes flavopunctata minima Pyrcz, 2004  TL: Peru

Punapedaliodes flavopunctata [n. ssp.] Pyrcz, MS  Ecuador

Antopedaliodes  Forster, 1964  

Antopedaliodes albopunctata (Weymer, 1890)    iN  M T S

Antopedaliodes antonia (Staudinger, 1897)    iN

Antopedaliodes antonia antonia (Staudinger, 1897)  TL: Bolivia  M T

Antopedaliodes antonia anina (Staudinger, 1897)  TL: Bolivia  M T

Antopedaliodes antonia quincedis (Thieme, 1905)  TL: Peru  M T

Antopedaliodes antonia [n. ssp.] Lamas, Viloria & Pyrcz, MS  Peru

Corderopedaliodes  Forster, 1964  

Corderopedaliodes corderoi (Dognin, 1893)    iN

Corderopedaliodes corderoi corderoi (Dognin, 1893)  TL: Ecuador  M T S

Corderopedaliodes corderoi exornata Pyrcz, 2004  TL: Peru

Corderopedaliodes buda Pyrcz, 2013  TL: not specified

Corderopedaliodes luzangelae Pyrcz & Warren-Gash, 2013  TL: not specified

Corderopedaliodes pandates (Hewitson, 1874)    iN

Corderopedaliodes pandates pandates (Hewitson, 1874)  TL: Bolivia  M T S

Corderopedaliodes pandates olivieri Pyrcz, 2013  TL: not specified

Panyapedaliodes  Forster, 1964  

Panyapedaliodes drymaea (Hewitson, 1858)    iN

Panyapedaliodes drymaea drymaea (Hewitson, 1858)  TL: [Ecuador]  M T S

Panyapedaliodes drymaea atropulla Pyrcz, 2005  TL: not specified

Panyapedaliodes jephtha (Thieme, 1905)    iN  M T

Panyapedaliodes mara (Thieme, 1905)    iN

Panyapedaliodes monticola (Tessmann, 1928)    iN  M T

Panyapedaliodes muscosa (Thieme, 1905)    iN

Panyapedaliodes muscosa muscosa (Thieme, 1905)  TL: Bolivia  M T

Panyapedaliodes muscosa sumaco Pyrcz & Rodriguez, 2005  TL: Ecuador

Panyapedaliodes muscosa tambito Pyrcz & Rodriguez, 2005  TL: not specified

Panyapedaliodes cocorna Pyrcz & Rodriguez, MS  TL: Peru

Panyapedaliodes panyasis (Hewitson, 1862)    iN  M T S

Panyapedaliodes stellata Pyrcz, 2004  TL: Peru

Panyapedaliodes phila (Hewitson, 1862)    iN

Panyapedaliodes phila phila (Hewitson, 1862)  TL: Colombia  M T

Panyapedaliodes phila certa Pyrcz, 2004  TL: Peru

Panyapedaliodes phila combeima (E. Krüger, 1924)  TL: Colombia  M T S

Panyapedaliodes phila philaenis (Thieme, 1905)  TL: Ecuador

Panyapedaliodes phila pilla Lamas, 2003  TL: Peru

Panyapedaliodes phila [n. ssp.] Pyrcz, MS  Ecuador

Panyapedaliodes puma (Thieme, 1905)    iN

Panyapedaliodes puma puma (Thieme, 1905)  TL: Bolivia  M T

Panyapedaliodes puma ernsti Pyrcz, 2006  TL: not specified

Panyapedaliodes rahab (Thieme, 1905)  TL: Peru  M T S

Panyapedaliodes silpa (Thieme, 1905)    iN

Panyapedaliodes silpa silpa (Thieme, 1905)  TL: Bolivia; Peru  M T

Panyapedaliodes silpa tomentosa (Weymer, 1912)  TL: Colombia  M S

Panyapedaliodes traceyannae Pyrcz & Viloria, 1999    iN

Panyapedaliodes [n. sp.#1] Pyrcz, MS  TL: Peru

Panyapedaliodes [n. sp.#2] Viloria & Pyrcz, MS  TL: Peru

Panyapedaliodes [n. sp.#3] Pyrcz, MS  TL: Peru

Panyapedaliodes [n. sp.#4] Lamas & Pyrcz, MS  TL: Peru

Panyapedaliodes [n. sp.#5] Pyrcz & Clavijo, MS  TL: Colombia

Paramo  Adams & Bernard, 1977  

Paramo oculata (E. Krüger, 1924)    iN  M T S

Parapedaliodes  Forster, 1964  

Parapedaliodes parepa (Hewitson, 1862)    iN

Parapedaliodes parepa parepa (Hewitson, 1862)  TL: Ecuador  M T

Parapedaliodes parepa milvia (Thieme, 1905)  TL: Peru  M T

Parapedaliodes parepa [n. ssp.#1] Pyrcz & Viloria, MS  TL: Peru

Parapedaliodes parepa [n. ssp.#2] Pyrcz & Viloria, MS  TL: Ecuador

Pherepedaliodes  Forster, 1964  

Pherepedaliodes naevia (Thieme, 1905)    iN

Pherepedaliodes naevia naevia (Thieme, 1905)  TL: Ecuador

Pherepedaliodes naevia [n. ssp.] Pyrcz, MS  Peru

Pherepedaliodes nubilia Pyrcz & Viloria, 1999  TL: Ecuador

Pherepedaliodes angela Pyrcz & Rodriguez, 2005  TL: not specified

Pherepedaliodes pheretiades (Grose-Smith & W. F. Kirby, 1894)    iN  M T

Pherepedaliodes [n. sp.#1] Pyrcz, MS  TL: Colombia

Pherepedaliodes [n. sp.#2] Pyrcz, MS  TL: Bolivia

Pherepedaliodes [n. sp.#3] Pyrcz, MS  TL: Peru

Physcopedaliodes  Forster, 1964  

Physcopedaliodes physcoa (Hewitson, 1862)    iN

Physcopedaliodes physcoa physcoa (Hewitson, 1862)  TL: Bolivia  M T

Physcopedaliodes physcoa marulla (Thieme, 1905)  TL: Peru  M T

Physcopedaliodes physcoa micromaculata Forster, 1964  TL: Bolivia  M T

Praepedaliodes  Forster, 1964  

Praepedaliodes amussis (Thieme, 1905)  TL: Colombia

Praepedaliodes granulata (A. Butler, 1868)  TL: Colombia  M T

Praepedaliodes phanias (Hewitson, 1862)    iN  M T

Praepedaliodes exul (Thieme, 1905)  TL: Brazil (RJ)  M T

Praepedaliodes duartei Dias, Dolibaina & Pyrcz, 2018  ref.282

Praepedaliodes francinii Freitas & Pyrcz, 2018  ref.282  iN

Praepedaliodes landryi Pyrcz & Freitas, 2018  ref.282  iN

Praepedaliodes pawlaki Pyrcz & Boyer, 2018  ref.282

Praepedaliodes sequeirae Pyrcz, Dias & Dolbaina, 2018  ref.282  iN

Praepedaliodes zaccae Dolibaina, Dias & Pyrcz, 2018  ref.282

Praepronophila  Forster, 1964  

Praepronophila emma (Staudinger, 1897)  TL: Bolivia  M T

Praepronophila perperna (Hewitson, 1862)    iN

Praepronophila perperna Panamanian segregate ,  Costa Rica & Panama  M S L

Praepronophila perperna perperna (Hewitson, 1862)  TL: [S. Amer.]  M T S

Praepronophila perperna smalli Pyrcz & Viloria, 2007  TL: Panama

Praepronophila perperna rodriguezi Pyrcz & Viloria, 2007  TL: Peru

Praepronophila petronius (Grose-Smith, 1900)  Toothed Satyr    iN

Praepronophila petronius kerrianna (L. Miller, 1986)  Costa Rica & Panama  M T S L

Praepronophila petronius petronius (Grose-Smith, 1900)  Panama & Colombia  M T S Ma L

Steromapedaliodes  Forster, 1964  

Steromapedaliodes albonotata (Godman, 1905)    iN

Steromapedaliodes albonotata albonotata (Godman, 1905)  TL: Venezuela  M T

Steromapedaliodes albonotata privigna (Thieme, 1905)  TL: Venezuela

Steromapedaliodes albonotata sanchezi Viloria & Pyrcz, 2001  TL: Venezuela

Steromapedaliodes albonotata schuberti Viloria & Pyrcz, 2001  TL: Venezuela  M T

Steromapedaliodes albonotata [n. ssp.#1] Pyrcz, MS  TL: Venezuela

Steromapedaliodes albonotata [n. ssp.#2] Pyrcz, MS  TL: Venezuela

Steromapedaliodes albonotata [n. ssp.#3] Pyrcz, MS  TL: Venezuela

Steromapedaliodes albonotata [n. ssp.#4] Pyrcz, MS  TL: Venezuela

Steromapedaliodes albarregas (Adams & Bernard, 1981)  TL: not specified

Steromapedaliodes albarregas albarregas (Adams & Bernard, 1981)  TL: not specified  M T

Steromapedaliodes albarregas [n. ssp.#1] Pyrcz, MS  TL: Venezuela

Steromapedaliodes albarregas [n. ssp.#2] Pyrcz, MS  TL: Venezuela

Steromapedaliodes [n. sp.] Pyrcz, MS  Venezuela

Neopedaliodes  Viloria, L. Miller & J. Miller, 2004  

Neopedaliodes chingazaensis (Torres & Le Crom, 1997)  TL: Colombia

Neopedaliodes zipa (Adams, 1986)  TL: Colombia  M T

Neopedaliodes nora (Adams, 1986)    iN

Neopedaliodes nora nora (Adams, 1986)  TL: Colombia  M T

Neopedaliodes nora stepheni Viloria, L. Miller & J. Miller, 2008  TL: Colombia

Neopedaliodes margaretha (Adams, 1986)  TL: Colombia

Neopedaliodes margaretha margaretha (Adams, 1986)  TL: Colombia  M T

Neopedaliodes margaretha allyni Viloria, L. Miller & J. Miller, 2008  TL: Colombia

Neopedaliodes philotera (Hewitson, 1868)  TL: ?  M T

Neopedaliodes lefebvreae Viloria, L. Miller & J. Miller, 2008  TL: Ecuador

Neopedaliodes parrhoebia (Hewitson, 1872)    iN

Neopedaliodes parrhoebia parrhoebia (Hewitson, 1872)  TL: Ecuador  M T

Neopedaliodes parrhoebia phintia (Hewitson, 1877)  TL: Ecuador  M T

Neopedaliodes entella (Thieme, 1905)  TL: Ecuador

Neopedaliodes juba (Staudinger, 1887)    iN

Neopedaliodes juba juba (Staudinger, 1887)  TL: Ecuador  M T

Neopedaliodes juba jubilosa Viloria, L. Miller & J. Miller, 2008  TL: Ecuador

Neopedaliodes michaeli Viloria, L. Miller & J. Miller, 2008  TL: Ecuador

Neopedaliodes michaeli michaeli Viloria, L. Miller & J. Miller, 2008  TL: Ecuador

Neopedaliodes michaeli innupta Pyrcz, 2008  TL: Ecuador

Neopedaliodes michaeli citrica Viloria, L. Miller & J. Miller, 2008  TL: Ecuador

Neopedaliodes proculeja (Thieme, 1905)  TL: Peru

Neopedaliodes proculeja malaga Pyrcz, 2008  TL: not specified

Neopedaliodes proculeja proculeja (Thieme, 1905)  TL: Peru  M T

Neopedaliodes phoenicusa (Hewitson, 1868)  TL: Ecuador

Neopedaliodes phoenicusa phoenicusa (Hewitson, 1868)  TL: Ecuador  M T

Neopedaliodes phoenicusa [n. ssp.] Pyrcz, MS  Ecuador

Neopedaliodes yeyo Pyrcz & Rodriguez, 2008  TL: not specified

Neopedaliodes [n. sp.] Pyrcz & Prieto, MS  Colombia

Pedaliodes  A. Butler, 1867  

Pedaliodes acjanaco Pyrcz, Viloria & Lamas, 2010  TL: Peru  M S

Pedaliodes adamsi D'Abrera, 1988  TL: Ecuador  M T

Pedaliodes albicilia Pyrcz, 2004  TL: Peru

Pedaliodes albutia Thieme, 1905  TL: Peru  M T

Pedaliodes [n. sp.#1] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#2] Lamas & Viloria, MS  TL: Peru

Pedaliodes alusana (Hewitson, 1869)    iN  M T

Pedaliodes amafania Thieme, 1905    iN  M T

Pedaliodes anchiphilonis Hayward, 1968  TL: Ecuador  M T

Pedaliodes antigua Adams & Bernard, 1981    iN  M T

Pedaliodes antulla Thieme, 1905    iN  M T

Pedaliodes arnotti Adams, 1986  TL: Colombia  M T

Pedaliodes arturi Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes asconia Thieme, 1905    iN  M T

Pedaliodes auraria Thieme, 1905    iN  M T

Pedaliodes aureola Pyrcz, 2004  TL: Peru  M S

Pedaliodes auristriga Thieme, 1905  TL: Peru  M S

Pedaliodes baccara Thieme, 1905    iN

Pedaliodes baccara baccara Thieme, 1905  TL: Colombia  M T

Pedaliodes baccara allopatra Viloria & Pyrcz, 2007  ref.285

Pedaliodes balnearia Pyrcz & Viloria, 1999  TL: Ecuador  M S

Pedaliodes bernardi Adams, 1986  TL: Colombia  M T

Pedaliodes [n. sp.#3] Viloria, MS  TL: Colombia

Pedaliodes boettgeri Pyrcz, 2004    iN

Pedaliodes caeca Pyrcz & Viloria, 1999  TL: Colombia

Pedaliodes canela Pyrcz & Viloria, 1999  TL: Colombia

Pedaliodes [n. sp.#4] Lamas & Viloria, MS  TL: Peru

Pedaliodes cebolleta Adams & Bernard, 1977  TL: Colombia  M T

Pedaliodes cesarense Adams & Bernard, 1979    iN  M T

Pedaliodes chaconi Viloria, 1998  TL: Venezuela  M T

Pedaliodes chrysotaenia (Hopffer, 1874)  TL: Peru  M T

Pedaliodes circumducta Thieme, 1905  Circumducta Satyr  SE Mexico, Guatemala  iN  M T L

Pedaliodes cledonia Thieme, 1905    iN

Pedaliodes cledonia cledonia Thieme, 1905  TL: Bolivia  M T

Pedaliodes cledonia molesta Pyrcz, 2004  TL: Peru  M S

Pedaliodes coca Staudinger, 1894    iN  M T

Pedaliodes [n. sp.#5] Viloria, MS  TL: Colombia

Pedaliodes costipunctata Weymer, 1912  TL: Colombia

Pedaliodes cremera Godman & Salvin, 1878  Cremera Satyr  Costa Rica & Panama  iN  M T OI S L

Pedaliodes croizatorum Viloria & Camacho, 1999  TL: Venezuela

Pedaliodes daulis Thieme, 1905  TL: Peru  M T

Pedaliodes dejecta (H. Bates, 1865)  Dejected Satyr    iN

Pedaliodes dejecta SW Mexican segregate ,  SW Mexico

Pedaliodes dejecta dejecta (H. Bates, 1865)  SE Mexico to Colombia  M T S L

Pedaliodes dejecta [n. ssp.] L. Miller & Llorente, MS  Mexico

Pedaliodes demathani Pyrcz, 2004    iN

Pedaliodes demathani demathani Pyrcz, 2004  TL: Peru  M S

Pedaliodes demathani pisca Lamas & Viloria, 2008  TL: Peru  M S

Pedaliodes demathani kuni Pyrcz & Boyer, 2008  TL: Peru  M S

Pedaliodes demathani chaneli Pyrcz & Boyer, 2008  TL: Peru

Pedaliodes demathani benedeki Pyrcz, 2008  TL: Peru

Pedaliodes demathani alfa Pyrcz & Boyer, 2008  TL: Peru

Pedaliodes demathani ockendeni Lamas & Viloria, 2008  TL: Peru

Pedaliodes demathani kalinowskii Pyrcz & Boyer, 2008  TL: Peru

Pedaliodes demathani horsti Pyrcz & Viloria, 2008  TL: Peru

Pedaliodes demathani nussi Pyrcz, 2008  TL: Bolivia

Pedaliodes demarmelsi Viloria, 1995  TL: Venezuela  M T

Pedaliodes dracula Pyrcz & Viloria, 1999    iN  M S

Pedaliodes empusa (C. Felder & R. Felder, 1867)  TL: Colombia

Pedaliodes empusa empusa (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Pedaliodes empusa medusa Pyrcz & Viloria, 2007  ref.285

Pedaliodes ereiba (C. Felder & R. Felder, 1867)    iN  M T

Pedaliodes erschoffi Pyrcz, 2004  TL: Peru  M S

Pedaliodes exanima (Erschoff, 1875)  TL: Peru  M S

Pedaliodes fassli Weymer, 1912  TL: Colombia  M T

Pedaliodes ferratilis A. Butler, 1873    iN  M T S

Pedaliodes fuscata (C. Felder & R. Felder, 1867)    iN  M T

Pedaliodes [n. sp.#6] Viloria, MS  TL: Colombia

Pedaliodes [n. sp.#7] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#8] Lamas & Viloria, MS  TL: Peru

Pedaliodes guicana Adams, 1986  TL: Colombia  M T

Pedaliodes gustavi Viloria, L. Miller & J. Miller, 2003  TL: Ecuador

Pedaliodes hardyi Adams, 1986    iN  M T

Pedaliodes [n. sp.#9] Viloria & Pyrcz, MS  TL: Venezuela

Pedaliodes hebena Pyrcz & Viloria, 1999    iN

Pedaliodes hewitsoni Staudinger, 1897    iN

Pedaliodes hewitsoni hewitsoni Staudinger, 1897  TL: Bolivia  M T

Pedaliodes hewitsoni primera Weeks, 1901  TL: Bolivia  M T S

Pedaliodes [n. sp.#10] Lamas & Viloria, MS  TL: Peru

Pedaliodes hopfferi Staudinger, 1887    iN  M T S

Pedaliodes [n. sp.#11] Lamas & Viloria, MS  TL: Peru

Pedaliodes japhleta A. Butler, 1870  TL: [Venezuela]  M T

Pedaliodes jelskii Pyrcz, 2004  TL: Peru

Pedaliodes [n. sp.#12] Viloria, MS  TL: Peru

Pedaliodes leucocheilus Godman & Salvin, 1880    iN  M T

Pedaliodes luperca Thieme, 1905  TL: Ecuador

Pedaliodes [n. sp.#13] Lamas & Viloria, MS  TL: Peru

Pedaliodes manis (C. Felder & R. Felder, 1867)  Manis Satyr  Costa Rica to Venezuela  iN  M T S L

Pedaliodes manneja Thieme, 1905  TL: Colombia; Venezuela  M T

Pedaliodes maruda Pyrcz, 2004  TL: Peru  M S

Pedaliodes [n. sp.#14] Viloria, MS  TL: Ecuador

Pedaliodes melaleuca Weymer, 1890    iN

Pedaliodes melaleuca melaleuca Weymer, 1890  TL: Colombia; Ecuador  M T

Pedaliodes melaleuca [n. ssp.] Viloria, MS  Ecuador

Pedaliodes [n. sp.#15] Pyrcz & Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#16] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#17] Lamas & Viloria, MS  TL: Peru

Pedaliodes montagna Adams & Bernard, 1981    iN  M T S

Pedaliodes morenoi Dognin, 1887    iN

Pedaliodes morenoi morenoi Dognin, 1887  TL: Ecuador  M T

Pedaliodes morenoi pilaloensis Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes napaea (H. Bates, 1865)  Napaea Satyr  S Mexico & Guatemala  iN  M T S L

Pedaliodes [n. sp.#18] L. Miller & Llorente, MS  TL: Mexico

Pedaliodes negreti Pyrcz, 1999  TL: Colombia

Pedaliodes niveonota A. Butler, 1873  TL: Peru  M T

Pedaliodes obscura Grose-Smith & W. F. Kirby, 1894  TL: Bolivia  M T

Pedaliodes obstructa Pyrcz & Viloria, 1999  TL: Colombia  M T

Pedaliodes occulta Pyrcz & Viloria, 1999  TL: Colombia

Pedaliodes ochrotaenia (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Pedaliodes ornata Grose-Smith & W. F. Kirby, 1895    iN  M T

Pedaliodes pacifica E. Krüger, 1924    iN

Pedaliodes pactyes (Hewitson, 1874)  TL: Bolivia  M T

Pedaliodes paeonides (Hewitson, 1874)  TL: Ecuador  M T S

Pedaliodes palaepolis (Hewitson, 1878)    iN

Pedaliodes palaepolis palaepolis (Hewitson, 1878)  TL: Bolivia  M T S

Pedaliodes palaepolis [n. ssp.#1] Lamas & Viloria, MS  TL: Peru

Pedaliodes palaepolis [n. ssp.#2] Lamas & Viloria, MS  TL: Peru

Pedaliodes palaepolis [n. ssp.#3] Viloria, MS  TL: Ecuador

Pedaliodes pallantis (Hewitson, 1862)  TL: Colombia  M T

Pedaliodes palpita Adams, 1986    iN

Pedaliodes palpita palpita Adams, 1986  TL: Colombia  M T

Pedaliodes palpita bedonica Pyrcz, 1999  TL: Colombia

Pedaliodes pammenes (Hewitson, 1874)  TL: Bolivia  M T

Pedaliodes paneis (Hewitson, 1862)  TL: [Peru]  M T

Pedaliodes panthides (Hewitson, 1874)  TL: Bolivia  M T

Pedaliodes parakana Viloria & Costa, 2019  ref.192

Pedaliodes parma Thieme, 1905    iN  M T

Pedaliodes parranda Adams, 1986  TL: Colombia  M T

Pedaliodes patizathes (Hewitson, 1874)  TL: [Bolivia]  M T

Pedaliodes pausia (Hewitson, 1862)    iN

Pedaliodes pausia pausia (Hewitson, 1862)  TL: Bolivia  M T S

Pedaliodes pausia uncus Thieme, 1905  TL: Argentina  M T S

Pedaliodes pedacia (Hewitson, 1868)  TL: Ecuador  M T

Pedaliodes pelinaea (Hewitson, 1878)    iN  M T

Pedaliodes pelinna (Hewitson, 1870)    iN  M T

Pedaliodes perisades (Hewitson, 1874)  TL: Bolivia  M T

Pedaliodes peruda (Hewitson, 1862)    iN  M T S

Pedaliodes peruviana A. Butler, 1873  TL: Peru  M T

Pedaliodes petri Pyrcz & Viloria, 1999    iN

Pedaliodes petri petri Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes petri maasseni Pyrcz, 2004  TL: Peru

Pedaliodes peucestas (Hewitson, 1862)    iN

Pedaliodes peucestas peucestas (Hewitson, 1862)  TL: Colombia  M T

Pedaliodes peucestas restricta Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes peucestas [n. ssp.] Viloria, MS  Ecuador

Pedaliodes phaea (Hewitson, 1862)    iN  M T

Pedaliodes phaedra (Hewitson, 1870)    iN

Pedaliodes phaedra phaedra (Hewitson, 1870)  TL: Ecuador  M T

Pedaliodes phaedra niphoessa Thieme, 1905  TL: Colombia  M S

Pedaliodes phaedra [n. ssp.#1] Viloria, MS  TL: Ecuador

Pedaliodes phaedra [n. ssp.#2] Viloria, MS  TL: Ecuador

Pedaliodes phaedra [n. ssp.#3] Viloria, MS  TL: Ecuador

Pedaliodes phaedra [n. ssp.#4] Viloria, MS  TL: Ecuador

Pedaliodes phaedra [n. ssp.#5] Viloria, MS  TL: Ecuador

Pedaliodes phaedra [n. ssp.#6] Viloria, MS  TL: Ecuador

Pedaliodes phaeina Staudinger, 1897    iN  M T

Pedaliodes [n. sp.#19] Lamas & Viloria, MS  TL: Peru

Pedaliodes phanoclea (Hewitson, 1877)  TL: Ecuador  M T

Pedaliodes phazania Grose-Smith, 1900    iN

Pedaliodes phazania phazania Grose-Smith, 1900  TL: Ecuador  M T

Pedaliodes phazania takahashii Adams & Bernard, 1977  TL: Colombia  M T

Pedaliodes pheres Thieme, 1905    iN  M S

Pedaliodes pheretias (Hewitson, 1872)  TL: Ecuador  M T

Pedaliodes philonis (Hewitson, 1872)  TL: Ecuador  M T

Pedaliodes phoenissa (Hewitson, 1862)    iN  M T S

Pedaliodes phoenix Lamas, 1999  TL: Peru  M T S

Pedaliodes phrasa Grose-Smith & W. F. Kirby, 1894  TL: Bolivia  M T

Pedaliodes phrasicla (Hewitson, 1874)    iN

Pedaliodes phrasicla phrasicla (Hewitson, 1874)  TL: Ecuador  M T

Pedaliodes phrasicla galaxias Weymer, 1912  TL: Peru; Bolivia  M T S

Pedaliodes phrasicla immaculata Pyrcz, 1999  TL: Colombia

Pedaliodes phrasicla rufa Pyrcz, 2004  TL: Peru

Pedaliodes phrasiclea Grose-Smith, 1900    iN  M T

Pedaliodes phrasis Grose-Smith, 1900  TL: Bolivia  M T S

Pedaliodes phthiotis (Hewitson, 1874)  TL: Ecuador  M T

Pedaliodes piletha (Hewitson, 1862)  TL: Venezuela  M T

Pedaliodes pimienta Adams, 1986    iN  M T

Pedaliodes piscolabis Pyrcz, 1999  TL: Colombia

Pedaliodes pisonia (Hewitson, 1862)    iN  M T

Pedaliodes plotina (Hewitson, 1862)    iN

Pedaliodes plotina plotina (Hewitson, 1862)  TL: Venezuela  M T

Pedaliodes plotina lucipara Weymer, 1912  TL: Colombia

Pedaliodes plotina pharnaspes (Hewitson, 1874)  TL: Venezuela  M T

Pedaliodes plotina rapha A. Butler, 1870  TL: [Venezuela]  M T S

Pedaliodes plotina ruricola Pyrcz & Viloria, 2007  ref.285

Pedaliodes plotina [n. ssp.#1] Viloria, MS  TL: Venezuela

Pedaliodes plotina [n. ssp.#2] Pyrcz, MS  TL: Venezuela

Pedaliodes poema Pyrcz & Viloria, 1999  TL: Colombia

Pedaliodes poesia (Hewitson, 1862)    iN  M T S

Pedaliodes polla Thieme, 1905    iN  M T S

Pedaliodes pollonia Adams, 1986    iN  M T S

Pedaliodes polusca (Hewitson, 1862)    iN  M T S

Pedaliodes [n. sp.#20] Lamas & Viloria, MS  TL: Peru

Pedaliodes pomponia (Hewitson, 1869)    iN  M T

Pedaliodes porcia (Hewitson, 1869)    iN

Pedaliodes porcia porcia (Hewitson, 1869)  TL: Ecuador  M T

Pedaliodes porcia pallantias (Hewitson, 1874)  TL: Ecuador  M T

Pedaliodes porcia [n. ssp.#1] Viloria, MS  TL: Colombia

Pedaliodes porcia [n. ssp.#2] Viloria, MS  TL: Colombia

Pedaliodes porcia [n. ssp.#3] Viloria, MS  TL: Colombia

Pedaliodes porima Grose-Smith & W. F. Kirby, 1894    iN  M T

Pedaliodes porina (Hewitson, 1862)    iN

Pedaliodes porina porina (Hewitson, 1862)  TL: ?  M T S

Pedaliodes porina [n. ssp.] Viloria & Pyrcz, MS  Venezuela

Pedaliodes praemontagna Pyrcz & Viloria, 2007  ref.285

Pedaliodes praxia (Hewitson, 1877)  TL: Ecuador

Pedaliodes praxia praxia (Hewitson, 1877)  TL: Ecuador  M T

Pedaliodes praxia [n. ssp.] Viloria, MS  Ecuador

Pedaliodes praxithea (Hewitson, 1870)    iN  M T S

Pedaliodes proerna (Hewitson, 1862)  TL: Colombia

Pedaliodes proerna proerna (Hewitson, 1862)  TL: Colombia  M T

Pedaliodes proerna fumaria Thieme, 1905  TL: Venezuela  M T

Pedaliodes prosa Staudinger, 1894  TL: Bolivia  M T

Pedaliodes prytanis (Hewitson, 1862)    iN  M T

Pedaliodes puracana E. Krüger, 1924  TL: Colombia  M T

Pedaliodes pylas (Hewitson, 1862)    iN  M T

Pedaliodes [n. sp.#21] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#22] Lamas & Viloria, MS  TL: Peru

Pedaliodes ralphi Adams, 1986  TL: Colombia  M T

Pedaliodes reyi Pyrcz & Viloria, 2007  ref.285  iN

Pedaliodes roraimae Strand, 1912    iN  M T

Pedaliodes rudnyi Pyrcz, 1999  TL: Colombia

Pedaliodes rumba Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes [n. sp.#23] Lamas & Viloria, MS  TL: Peru

Pedaliodes simmias Thieme, 1905  TL: Bolivia  M T S

Pedaliodes simpla Thieme, 1905    iN

Pedaliodes simplissima Pyrcz, 2004    iN  M S

Pedaliodes socorrae Adams, 1986  TL: Colombia  M T

Pedaliodes sonata Pyrcz & Viloria, 1999    iN

Pedaliodes sophismata Pyrcz, 2004  TL: Peru  M S

Pedaliodes spina Weymer, 1912    iN

Pedaliodes [n. sp.#24] Lamas & Viloria, MS  TL: Peru

Pedaliodes stuebeli Pyrcz, 2004  TL: Peru  M S

Pedaliodes suspiro Adams & Bernard, 1979    iN  M T

Pedaliodes syleus Thieme, 1905  TL: Peru

Pedaliodes symmachus Godman & Salvin, 1880    iN  M T

Pedaliodes sztolcmani Pyrcz, 2004  TL: Peru

Pedaliodes sztolcmani sztolcmani Pyrcz, 2004  TL: Peru

Pedaliodes sztolcmani gilvaecosta Pyrcz, 2004  TL: Peru

Pedaliodes tabaconas Pyrcz & Viloria, 1999  TL: Ecuador

Pedaliodes terramaris Viloria & Pyrcz, 2000  TL: Venezuela  M T

Pedaliodes thiemei Staudinger, 1897    iN  M T

Pedaliodes transmontana Pyrcz & Viloria, 1999    iN

Pedaliodes triaria Godman & Salvin, 1878  Ochre-banded Satyr  Costa Rica & Panama  iN  M T OI S

Pedaliodes tucca Thieme, 1905    iN  M S

Pedaliodes tyro Thieme, 1905  TL: Peru  M T

Pedaliodes tyrrheoides Adams & Bernard, 1979    iN  M T

Pedaliodes tyrrheus Godman & Salvin, 1880    iN

Pedaliodes tyrrheus tyrrheus Godman & Salvin, 1880  TL: Colombia  M T

Pedaliodes tyrrheus tairona Adams & Bernard, 1977  TL: Colombia  M T

Pedaliodes uaniuna Pyrcz & Viloria, 2004  TL: Peru  M S

Pedaliodes uniformis Weymer, 1912  TL: Bolivia  M T

Pedaliodes uniplaga Weymer, 1912  TL: not specified  M T S

Pedaliodes valencia Pyrcz & Viloria, 2007  ref.285

Pedaliodes vallenata Adams & Bernard, 1979  TL: Venezuela  M T S

Pedaliodes wilhelmi Pyrcz, 1999  TL: Colombia

Pedaliodes woytkowskii Pyrcz, 2004  TL: Peru  M S

Pedaliodes xanthosphenisca Hayward, 1968  TL: Ecuador  M T

Pedaliodes yutajeana Viloria & Pyrcz, 2000  TL: Venezuela  M T

Pedaliodes zuleta Adams & Bernard, 1979    iN  M T S

Pedaliodes zingara Heredia & Viloria , 2004  ref.200  iN

Pedaliodes haydoni Pyrcz & Viloria, 2013  TL: Colombia

Pedaliodes adrianae Pyrcz & Prieto, 2013  TL: Colombia

Pedaliodes rodriguezi Pyrcz & Andrade, 2013    iN

Pedaliodes philinae Pyrcz & Prieto, 2013  TL: Colombia

Pedaliodes [n. sp.#25] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#26] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#27] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#28] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#29] Viloria, MS  TL: Colombia

Pedaliodes [n. sp.#30] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#31] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#32] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#33] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#34] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#35] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#36] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#37] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#38] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#39] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#40] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#41] Viloria, MS  TL: Bolivia

Pedaliodes [n. sp.#42] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#43] Viloria, MS  TL: Brazil (SC)

Pedaliodes [n. sp.#44] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#45] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#46] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#47] Viloria, MS  TL: Bolivia

Pedaliodes [n. sp.#48] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#49] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#50] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#51] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#52] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#53] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#54] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#55] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#56] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#57] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#58] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#59] Viloria, MS  TL: Colombia

Pedaliodes [n. sp.#60] Viloria, MS  TL: ?

Pedaliodes [n. sp.#61] Viloria, MS  TL: Bolivia

Pedaliodes [n. sp.#62] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#63] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#64] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#65] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#66] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#67] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#68] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#69] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#70] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#71] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#72] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#73] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#74] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#75] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#76] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#77] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#78] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#79] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#80] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#81] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#82] Viloria, MS  TL: Colombia

Pedaliodes [n. sp.#83] Lamas & Viloria, MS  TL: Peru

Pedaliodes [n. sp.#84] Pyrcz, MS  TL: Ecuador

Pedaliodes [n. sp.#85] Viloria, MS  TL: Bolivia

Pedaliodes [n. sp.#86] Pyrcz, MS  TL: Ecuador

Pedaliodes [n. sp.#87] Viloria, MS  TL: Ecuador

Pedaliodes [n. sp.#88] Viloria, MS  TL: Bolivia

Pedaliodes [n. sp.#89] L. Miller & Llorente, MS  TL: Mexico (Gue)

Pedaliodes [n. sp.#90] Lamas, MS  TL: Peru

Pedaliodes [n. sp.#91] Lamas, MS  TL: Peru

Pedaliodes [n. sp.#92] Lamas, MS  TL: Peru

Protopedaliodes  Viloria & Pyrcz, 1994  

Protopedaliodes kukenani Viloria & Pyrcz, 1994  TL: Venezuela  M T

Protopedaliodes ridouti Viloria & Pyrcz, 2000  TL: Venezuela  M T

Protopedaliodes profauna Viloria & Pyrcz, 2000  TL: Venezuela  M T

Protopedaliodes arekuna Pyrcz & Stachowicz, 2021  TL: Venezuela

Argyrophorus  Blanchard, 1852  

Argyrophorus argenteus Blanchard, 1852    iN  M T

Argyrophorus blanchardi Pyrcz & Wojtusiak, MS  TL: Peru

Argyrophorus blanchardi blanchardi Pyrcz & Wojtusiak, MS  TL: Peru

Argyrophorus blanchardi celendeni Pyrcz & Wojtusiak, MS  TL: Peru

Argyrophorus blanchardi [n. ssp.] Lamas, MS  Peru

Argyrophorus chiliensis (Guérin-Méneville, [1830])    iN

Argyrophorus chiliensis chiliensis (Guérin-Méneville, [1830])  TL: Chile  M T

Argyrophorus chiliensis magallanicus (Herrera, 1965)  TL: Chile  M T

Argyrophorus chiliensis elwesi (Bryk, 1944)  TL: Argentina

Argyrophorus chiliensis [n. ssp.] Pyrcz, MS  Argentina

Argyrophorus tandilensis (Köhler, 1935)    iN  M T

Argyrophorus antarcticus (Mabille, 1885)  TL: [Argentina]  M T

Argyrophorus gustavi (Staudinger, 1898)    iN

Argyrophorus gustavi gustavi (Staudinger, 1898)  TL: Bolivia  M T

Argyrophorus gustavi penai (Hayward, 1967)  TL: Chile  M T

Argyrophorus monticolens (A. Butler, 1881)    iN

Argyrophorus monticolens monticolens (A. Butler, 1881)  TL: Chile  M T

Argyrophorus monticolens [n. ssp.] Pyrcz, MS  Argentina

Argyrophorus poaoeneis (Heimlich, 1959)  TL: Chile  M T

Argyrophorus angusta Weymer, 1911  TL: Bolivia

Argyrophorus angusta angusta Weymer, 1911  TL: Bolivia  M S

Argyrophorus angusta picota Fuchs, 1954  TL: Peru

Argyrophorus lamna Thieme, 1904    iN

Argyrophorus lamna lamna Thieme, 1904  TL: Bolivia  M S

Argyrophorus lamna [n. ssp.#1] Pyrcz, MS  TL: Peru  M T

Argyrophorus lamna [n. ssp.#2] Pyrcz, MS  TL: Peru

Argyrophorus lamna [n. ssp.#3] Pyrcz & Cerdeńa, MS  TL: Peru

Argyrophorus williamsianus A. Butler, 1868  TL: [Chile]  M T

Auca  Hayward, 1953  

Auca barrosi (C. Silva, 1917)    iN

Auca barrosi barrosi (C. Silva, 1917)  TL: Chile  M T

Auca barrosi [n. ssp.] Pyrcz, MS  Chile

Auca coctei (Guérin-Méneville, [1838])    iN

Auca coctei coctei (Guérin-Méneville, [1838])  TL: Chile  M T S

Auca coctei [n. ssp.] Pyrcz, MS  Chile

Auca pales (Philippi, 1859)  TL: Chile

Cosmosatyrus  C. Felder & R. Felder, 1867  

Cosmosatyrus leptoneuroides C. Felder & R. Felder, 1867    iN

Cosmosatyrus leptoneuroides leptoneuroides C. Felder & R. Felder, 1867  TL: Chile  M T

Cosmosatyrus leptoneuroides plumbeola (A. Butler, 1868)  TL: Chile  M T

Cosmosatyrus leptoneuroides [n. ssp.] Pyrcz, MS  Argentina

Elina  Blanchard, 1852  

Elina montrolii (Feisthamel, 1839)    iN  M T S

Elina vanessoides Blanchard, 1852    iN  M T S

Faunula  C. Felder & R. Felder, 1867  

Faunula leucoglene C. Felder & R. Felder, 1867    iN  M T

Faunula eleates (Weymer, 1890)    iN  M T

Faunula patagonica (Mabille, 1885)    iN  M T

Faunula dubii Pyrcz, 2012    iN

Faunula euripides (Weymer, 1890)  TL: Bolivia

Faunula euripides euripides (Weymer, 1890)  TL: Bolivia  M T

Faunula euripides salomonis (Köhler, 1939)  TL: Argentina  M T

Faunula euripides [n. ssp.] Pyrcz & Cerdeńa, MS  Peru

Haywardella  Herrera, 1966  

Haywardella edmondsii (A. Butler, 1881)    iN  M T

Homoeonympha  C. Felder & R. Felder, 1867  

Homoeonympha boisduvalii (Blanchard, 1852)    iN

Homoeonympha boisduvalii boisduvalii (Blanchard, 1852)  TL: Chile  M T S

Homoeonympha boisduvalii pusilla C. Felder & R. Felder, 1867  TL: Chile  M T

Homoeonympha vesagus (E. Doubleday, [1849])  TL: USA?  M T

Nelia  Hayward, 1953  

Nelia calvertii (Elwes, 1903)    iN  M T

Nelia nemyroides (Blanchard, 1852)    iN  M T

Nelia [n. sp.] Pyrcz, MS  Chile

Neomaenas  Wallengren, 1858  

Neomaenas coenonymphina A. Butler, 1881    iN  M T

Neomaenas edmondsii (A. Butler, 1881)  TL: Chile  M T

Neomaenas fractifascia A. Butler, 1881    iN  M T

Neomaenas inornata (Elwes, 1903)    iN  M T

Neomaenas poliozona (C. Felder & R. Felder, 1867)    iN  M T

Neomaenas [n. sp.] Pyrcz, MS  Chile

Neomaenas servilia Wallengren, 1858    iN  M T

Neomaenas wallengrenii A. Butler, 1881    iN  M T

Neomaenas ambiorix Wallengren, 1858    iN

Neomaenas ambiorix ambiorix Wallengren, 1858  TL: Chile  M T

Neomaenas ambiorix [n. ssp.#1] Pyrcz, MS  TL: Chile

Neomaenas ambiorix [n. ssp.#2] Pyrcz, MS  TL: Chile

Neomaenas simplex (A. Butler, 1881)  TL: Chile  M T

Neomaenas humilis (C. Felder & R. Felder, 1867)    iN

Neomaenas humilis humilis (C. Felder & R. Felder, 1867)  TL: Chile  M T

Neomaenas humilis [n. ssp.] Pyrcz, MS  Chile

Neomaenas schajovskoii (Hayward, 1954)    iN  M T

Pampasatyrus  Hayward, 1953  

Pampasatyrus glaucope (C. Felder & R. Felder, 1867)    iN

Pampasatyrus glaucope glaucope (C. Felder & R. Felder, 1867)  TL: Brazil  M T

Pampasatyrus glaucope boenninghauseni (Foetterle, 1902)  TL: Brazil (RJ)  M T

Pampasatyrus gyrtone (Berg, 1877)    iN  M T

Pampasatyrus quies (Berg, 1877)    iN  M T

Pampasatyrus reticulata (Weymer, 1907)    iN

Pampasatyrus yacantoensis (Köhler, 1939)    iN  M T

Pampasatyrus nilesi (Weeks, 1902)    iN

Pampasatyrus nilesi nilesi (Weeks, 1902)  TL: Bolivia  M T

Pampasatyrus nilesi johanna (Weymer, 1911)  TL: Argentina

Pampasatyrus nilesi [n. ssp.] Pyrcz, MS  Peru

Pampasatyrus [n. sp.#1] Zacca & Pyrcz, MS  TL: Brazil

Pampasatyrus [n. sp.#2] Zacca & Pyrcz, MS  TL: Argentina

Quilaphoetosus  Herrera, 1966  

Quilaphoetosus janirioides (Blanchard, 1852)    iN  M T

Quilaphoetosus monachus (Blanchard, 1852)    iN

Quilaphoetosus monachus monachus (Blanchard, 1852)  TL: Chile  M T

Quilaphoetosus monachus valdiviae (C. Felder & R. Felder, 1867)  TL: Chile  M T

Spinantenna  Hayward, 1953  

Spinantenna tristis (Guérin-Méneville, [1830])    iN  M T

Tetraphlebia  C. Felder & R. Felder, 1867  

Tetraphlebia germainii C. Felder & R. Felder, 1867    iN

Tetraphlebia germainii germainii C. Felder & R. Felder, 1867  TL: Chile  M T

Tetraphlebia germainii argentina Hayward, 1962  TL: Argentina  M T

Chillanella  Herrera, 1966  

Chillanella stelligera (A. Butler, 1881)    iN  M T

  Tribe Satyrini, Subtribe Euptychiina

Archeuptychia  Forster, 1964  

Archeuptychia cluena (Drury, 1782)    iN  M S

Caeruleuptychia  Forster, 1964  

Caeruleuptychia aegrota (A. Butler, 1867)    iN  M T S

Caeruleuptychia aetherialis (A. Butler, 1877)  ref.258  iN  M T

Caeruleuptychia helios (Weymer, 1911)    iN  M T

Caeruleuptychia trembathi Willmott, Nakahara, J. Hall & Neild, 2017  ref.258

Caeruleuptychia pilata (A. Butler, 1867)    iN  M T

Caeruleuptychia scripta Nakahara, Zacca & Huertas, 2017  ref.260

Caeruleuptychia maryzenderae Lamas & Nakahara, 2017  ref.258

Caeruleuptychia coelestis (A. Butler, 1867)    iN  M T

Caeruleuptychia coelica (Hewitson, 1869)    iN  M T

Caeruleuptychia divina (Weymer, 1911)    iN  M T

Caeruleuptychia cyanites (A. Butler, 1871)    iN  M T S

Caeruleuptychia glauca (Weymer, 1911)    iN  M T

Caeruleuptychia mare (A. Butler, 1869)  TL: Brazil (Pa)  M T

Caeruleuptychia urania (A. Butler, 1867)    iN  M T S

Caeruleuptychia ziza (A. Butler, 1869)  TL: Peru  M T

Caeruleuptychia lobelia (A. Butler, 1870)    iN  M T S

Caeruleuptychia probata (Weymer, 1911)    iN  M T

Caeruleuptychia francisca (A. Butler, 1870)    iN  M T S

Caeruleuptychia scopulata (Godman, 1905)    iN  M T S

Caeruleuptychia saul Brévignon & M. Benmesbah, 2011  ref.179

Caeruleuptychia tenera (Weymer, 1911)    iN  M T S

Caeruleuptychia twalela Brévignon, 2005  ref.174  iN

Caeruleuptychia romani (Aurivillius, 1929)  ref.201  iN  M T

Caeruleuptychia penicillata (Godman, 1905)  TL: Brazil (Pa)  M T S

Caeruleuptychia helena (Anken, 1994)    iN  M OI OD G

Caeruleuptychia umbrosa (A. Butler, 1870)    iN  M T S

Caeruleuptychia [n. sp.#1] Lamas, MS  TL: Peru

Caeruleuptychia [n. sp.#2] Lamas, MS  TL: Peru

Caeruleuptychia [n. sp.#3] Lamas, MS  TL: Peru

Caeruleuptychia [n. sp.#4] Lamas, MS  TL: Peru

Caeruleuptychia [n. sp.#5] Lamas, MS  TL: Peru

Caeruleuptychia [n. sp.#6] Lamas, MS  TL: Peru

Capronnieria  Forster, 1964  

Capronnieria galesus (Godart, [1824])    iN  M T S

Capronnieria narapa (Schaus, 1902)  ref.201  iN  M T S

Carminda  Ebert & Dias, 1998  

Carminda griseldis (Weymer, 1911)  ref.195  iN  M S

Carminda paeon (Godart, [1824])  ref.195  iN  M T

Carminda surpresa Barbosa, Aguiar, Rosa, Zacca & A. Freitas, 2020    iN

Carminda umuarama Ebert & Dias, 1997  ref.195  iN  M T

Cepheuptychia  Forster, 1964  

Cepheuptychia angelica (A. Butler, 1874)  TL: Brazil (RJ)  M T

Cepheuptychia cephus (Fabricius, 1775)    iN  M T S

Cepheuptychia glaucina (H. Bates, 1864)  Dirty-blue Satyr  SE Mexico, Guatemala  iN  M T S L

Cepheuptychia [n. sp.] Lamas, MS  Peru

Chloreuptychia  Forster, 1964  

Chloreuptychia agatha (A. Butler, 1867)    iN  M T S

Chloreuptychia amethysta Brévignon & M. Benmesbah, 2011  ref.179  iN

Chloreuptychia chlorimene (Hübner, [1819])    iN  M T S

Chloreuptychia gordonsmalli Thurman, MacDonald & Nakahara, 2021  TL: Panama

Chloreuptychia [n. sp.#1] Lamas, MS  TL: Peru

Chloreuptychia [n. sp.#2] Lamas, MS  TL: Peru

Lazulina  Willmott, Nakahara & Espeland, 2023  

Lazulina catharina (Staudinger, [1886])  ref.201  iN  M T

Lazulina hewitsonii (A. Butler, 1867)  ref.201  iN  M T

Lazulina tolumnia (Cramer, 1777)  ref.201  iN  M S

Lazulina [n. sp.] Lamas, MS  Peru

Amiga  Nakahara, Willmott & Espeland, 2019  

Amiga arnaca (Fabricius, 1776)  Blue-smudged Satyr    iN

Amiga arnaca arnaca (Fabricius, 1776)  Venezuela to Bolivia, Brazil, Guianas; ref.264  M T S L

Amiga arnaca adela Nakahara & Espeland, 2019  ref.264

Amiga arnaca indianacristoi Nakahara & Marín, 2019  ref.201,309,272,264

Amiga sericeella (H. Bates, 1864)  Blue-topped Satyr  S Mexico-Honduras; ref.264  M T S

Cissia  E. Doubleday, 1848  

Cissia penelope (Fabricius, 1775)    iN  M T S

Cissia pompilia (C. Felder & R. Felder, 1867)  Plain Satyr  [= usitata], E & S Mexico to Colombia  iN  M T S L1 L2

Cissia proba (Weymer, 1911)    iN  M T

Cissia cleophes (Godman & Salvin, 1889)  Cleophes Satyr  S Mexico; ref.11  iN  M T S L Ha

Cissia eous (A. Butler, 1867)  ref.367  iN  M T

Cissia phronius (Godart, [1824])  ref.367  iN

Cissia rubricata (W. H. Edwards, 1871)  Red Satyr    iN

Cissia rubricata rubricata (W. H. Edwards, 1871)  far NE Mexico; ref.367  M T S

Cissia rubricata smithorum (Wind, 1946)  N-central Mexico; ref.367  M T S L

Cissia cheneyorum (R. Chermock, 1949)  Red Satyr  NW Mexico; ref.16  M S L I

Cissia anabelae (L. Miller, 1976)  Red Satyr  Chiapas; ref.16  M T

Cissia wahala Grishin, 2022  Mexico (Oax, Pue); ref.16

Cissia pseudocleophes (L. Miller, 1976)  Red Satyr  W Mexico; ref.16  M T S L

Cissia joyceae Singer, DeVries & P. Ehrlich, 1983  Joyce's Satyr  Costa Rica  M T

Vareuptychia  Forster, 1964  

Vareuptychia similis (A. Butler, 1867)  One-pupil Satyr  W Mexico to Colombia; ref.370  iN  M T S L

Vareuptychia themis (A. Butler, 1867)  Two-pupil Satyr  [= undina], W & SE Mexico to Nicaragua  iN  M T S1 S2 L Ha

Vanima  Zacca, Casagrande & O. Mielke, 2020  

Vanima labe (A. Butler, 1870)  Labe Satyr  S Mexico to W Ecuador; ref.370  iN  M T S L

Vanima lesbia (Staudinger, [1886])  ref.370  M T

Vanima palladia (A. Butler, 1867)  Butler’s Satyr  S Mexico to Brazil, Trinidad; ref.370  iN  M T S L

Cyllopsis  R. Felder, 1869  

Cyllopsis gemma (Hübner, 1808)  Eastern Gemmed-Satyr    iN

Cyllopsis gemma gemma (Hübner, 1808)  S IL to E TX, E to WV, S to FL  M S L1 L2 L3

Cyllopsis gemma freemani (D. Stallings & J. Turner, 1947)  S TX to Guatemala & Belize  M T1 T2 S L

Cyllopsis pertepida (Dyar, 1912)  Canyonland Gemmed-Satyr    iN

Cyllopsis pertepida dorothea (Nabokov, 1943)  SE NV, S & C UT, S & C CO, far W OK, N & C NM, N & C AZ  M S L

Cyllopsis pertepida maniola (Nabokov, 1942)  SE AZ, SW NM, far NW Mexico (N SON, CHIH)  M T S L

Cyllopsis pertepida avicula (Nabokov, 1942)  W TX, NE Mexico  M T S I Ha

Cyllopsis pertepida intermedia L. Miller, 1974  NW Mexico (W-central Chihuahua)  M T S

Cyllopsis pertepida pertepida (Dyar, 1912)  W to S-central Mexico  M T S L

Cyllopsis pyracmon (A. Butler, 1867)  Nabokov’s Satyr    iN

Cyllopsis pyracmon pyracmon (A. Butler, 1867)  C & S Mexico to Guatemala  M T S1 S2 L

Cyllopsis pyracmon henshawi (W. H. Edwards, 1876)  SE AZ, SW NM, NW Mexico  M T S L I

Cyllopsis brocki Grishin, 2023  ref.146

Cyllopsis argentella (A. Butler & H. Druce, 1872)  Ochre-patched Gemmed-Satyr  Costa Rica & W Panama  iN  M T S L

Cyllopsis caballeroi Beutelspacher, 1982  Cowboy Gemmed-Satyr  W Mexico (Nayarit, Jalisco)  iN  M T S

Cyllopsis clinas (Godman & Salvin, 1889)  Falcate Gemmed-Satyr  SW Mexico (Guerrero)  M T S

Cyllopsis diazi L. Miller, 1974  Diaz's Gemmed-Satyr  W Mexico, [Guatemala?]  iN  M T L Ha

Cyllopsis dospassosi L. Miller, 1974  dos Passos' Gemmed-Satyr  NE Mexico  iN  M T L

Cyllopsis emilia I. Chacón & Nishida, 2002  White Gemmed-Satyr  Costa Rica (Cerro de la Muerte, Villa Mills)  M S

Cyllopsis guatemalena L. Miller, 1974  Guatemalan Gemmed-Satyr  S Mexico (Chiapas) to Guatemala  iN  M T S L

Cyllopsis hedemanni R. Felder, 1869  Stub-tailed Gemmed-Satyr    iN

Cyllopsis hedemanni hedemanni R. Felder, 1869  [= vetones], E & S Mexico to W Panama  M T S L

Cyllopsis hedemanni tamaulipensis L. Miller, 1974  NE Mexico  M T S L

Cyllopsis hedemanni vetones (Godman & Salvin, 1878)  TL: Costa Rica  M T

Cyllopsis hilaria (Godman, 1901)  Two-toned Gemmed-Satyr  E Mexico to W Panama  iN  M T S L

Cyllopsis jacquelineae L. Miller, 1974  Jacqueline's Gemmed-Satyr  SW Mexico (Guerrero, Oaxaca)  iN  M T L

Cyllopsis nayarit (R. Chermock, 1947)  Nayarit Gemmed-Satyr  E & W Mexico  iN  M S L

Cyllopsis nelsoni (Godman & Salvin, 1881)  Yellow-striped Gemmed-Satyr  Guatemala (Cerro de Zunil)  iN  M T TI L

Cyllopsis pallens L. Miller, 1974  Pallid Gemmed-Satyr  W Mexico to Guatemala  iN  M T S L

Cyllopsis parvimaculata L. Miller, 1974  Weak-marked Gemmed-Satyr  S-central Mexico (TL: Tenancingo, Mexico State)  M T

Cyllopsis pephredo (Godman, 1901)  Big-spiked Gemmed-Satyr  SE Mexico to C Panama  iN  M T S

Cyllopsis perplexa L. Miller, 1974  Perplexing Gemmed-Satyr  S & SW Mexico  iN  M T L

Cyllopsis philodice (Godman & Salvin, 1878)  Pale-banded Gemmed-Satyr  Costa Rica & W Panama  iN  M T S L

Cyllopsis pseudopephredo (R. Chermock, 1947)  Chermock's Gemmed-Satyr  S-central Mexico (Distrito Federal)  M T

Cyllopsis rogersi (Godman & Salvin, 1878)  Costa Rican Gemmed-Satyr  Costa Rica & Panama  iN  M T S L

Cyllopsis schausi L. Miller, 1974  Schaus’ Gemmed-Satyr  W Mexico to Guatemala  M T S

Cyllopsis steinhauserorum L. Miller, 1974  Steinhausers' Gemmed-Satyr  W Mexico to El Salvador  iN  M T

Cyllopsis suivalenoides L. Miller, 1974  Big-eyed Gemmed-Satyr  W Mexico to Nicaragua  iN  M T S L

Cyllopsis suivalens (Dyar, 1914)  Dyar’s Gemmed-Satyr    iN

Cyllopsis suivalens suivalens (Dyar, 1914)  E Mexico (Puebla, Veracruz)  M T S L

Cyllopsis suivalens escalantei L. Miller, 1974  S Mexico to Guatemala  M T S L

Cyllopsis tomemmeli Warren & Nakahara, 2018  ref.299

Cyllopsis whiteorum L. Miller & J. Maza, 1984  Dark Gemmed-Satyr  S Mexico (N Oaxaca)  iN  M T L

Cyllopsis wellingi L. Miller, 1978  Belize Gemmed-Satyr  endemic to the Maya Mountains of Belize  M T S

Cyllopsis windi L. Miller, 1974  Wind's Gemmed-Satyr  [Possibly a form of C. nayarit], E & W Mexico  iN  M T S L

Erichthodes  Forster, 1964  

Erichthodes antonina (C. Felder & R. Felder, 1867)    iN  M T S

Erichthodes jovita (C. Felder & R. Felder, 1867)    iN  M T

Erichthodes julia (Weymer, 1911)    iN  M T

Erichthodes eremita Lamas, Willmott & Radford, 2018  ref.315

Stephenympha  Viloria, 2022  

Stephenympha arius (Weymer, 1911)  TL: Bolivia  M T

Stephenympha pauliana Viloria, 2022  TL: Venezuela

Stephenympha eriphule (A. Butler, 1867)    iN  M T

Atlanteuptychia  Freitas, Barbosa & O. Mielke, 2013  

Atlanteuptychia ernestina (Weymer, 1911)  ref.212  iN  M T

Euptychia  Hübner, 1818  

Euptychia alacristata Neild, Nakahara & Fratello, 2014  Amazon basin & S of the Guianas; ref.267

Euptychia aquila Fratello, Nakahara & Brévignon, 2015  ref.206

Euptychia atlantica Nakahara & Freitas, 2017  ref.259

Euptychia attenboroughi Neild, Nakahara, Fratello & Le Crom, 2015  ref.268

Euptychia audacia Brévignon, Fratello & Nakahara, 2015  ref.206

Euptychia boulleti (Le Cerf, 1919)  ref.210  M T

Euptychia cesarense Pulido, Andrade, C. Peña & Lamas, 2011  TL: Colombia

Euptychia cesarense cesarense Pulido, Andrade, C. Peña & Lamas, 2011  ref.279

Euptychia cesarense obtusa Nakahara, 2015  ref.252

Euptychia cesarense viloriai Andrade, Pulido, C. Peña & Lamas, 2011  ref.279  M S

Euptychia efraini Ríos, 2019  TL: Colombia

Euptychia enyita Nakahara, Lamas & Willmott, 2015  ref.252  iN

Euptychia enyo A. Butler, 1867    iN  M T

Euptychia favonius Nakahara, G. Vega & Willmott, 2016  ref.257

Euptychia fernandae Nakahara & Willmott, 2015  ref.252

Euptychia fetna A. Butler, 1870  Orange-patched Satyr  W & S Mexico to Guatemala  iN  M T S L

Euptychia granatina Nakahara, LeCrom & J. Hall, 2015  ref.252  iN

Euptychia hannemanni Forster, 1964    iN  M T

Euptychia insolata A. Butler & H. Druce, 1872  Insolata Satyr  Costa Rica & Panama  iN  M T S L

Euptychia jesia A. Butler, 1869  Jesia Satyr  S Mexico to Colombia  iN  M T S L

Euptychia juanjoi LeCrom, Nakahara & Lamas, 2015  ref.251

Euptychia lacandona A. Warren & Nakahara, 2015  ref.250

Euptychia marceli Brévignon, 2005  ref.174  iN

Euptychia marceli marceli Brévignon, 2005  ref.174

Euptychia marceli divisa M. Benmesbah, Costa, Attal & Viloria, 2021  TL: Venezuela

Euptychia meta Weymer, 1911    iN  M T

Euptychia mollina (Hübner, [1813])  Mollina Satyr  E Mexico to Colombia & Venezuela  iN

Euptychia mollina mollina (Hübner, [1813])  E Mexico to Colombia & Venezuela  M T S L

Euptychia mollina suzannae Brévignon, 2005  ref.174

Euptychia mollis Staudinger, 1876  Mollis Satyr  [Belize?] to Panama  iN  M T S

Euptychia neblina A. Warren & Nakahara, 2015  ref.250

Euptychia neildi Brévignon, 2005  ref.174

Euptychia padroni Nakahara, Lamas & Willmott, 2015  ref.252  M S

Euptychia pegasus Nakahara & J. Hall, 2015  ref.252  iN

Euptychia picea A. Butler, 1867    iN  M T S

Euptychia pillaca Nakahara & Willmott, 2015  ref.252

Euptychia roraima Nakahara, Fratello & Harvey, 2014  Guyana; ref.249

Euptychia rubrofasciata L. Miller & J. Miller, 1988  Red-webbed Satyr  NW Mexico  iN  M T S L

Euptychia rufocincta Weymer, 1911    iN  M T S

Euptychia sarah M. Benmesbah & Viloria, 2019  TL: Venezuela

Euptychia sophiae Zacca, Nakahara, Dolibaina & F. Dias, 2015  ref.268

Euptychia truncata Nakahara & J. Hall, 2015  ref.252

Euptychia westwoodi A. Butler, 1867  Westwood’s Satyr  S Mexico to Colombia & Venezuela  iN

Euptychia westwoodi westwoodi A. Butler, 1867  S Mexico to Colombia & Venezuela  M T S L

Euptychia westwoodi muli Brévignon, 2015  ref.174

Euptychia woroina Viloria & Benmesbah, 2020  ref.193

Euptychia [n. sp.#1] Lamas, MS  TL: Peru

Euptychia [n. sp.#2] Lamas, MS  TL: Peru

Euptychia [n. sp.#3] Lamas, MS  TL: Peru

Euptychia [n. sp.#4] Lamas, MS  TL: Peru

Euptychia [n. sp.#5] Lamas, MS  TL: Peru

Euptychia [n. sp.#6] Lamas, MS  TL: Peru

Euptychia [n. sp.#7] Lamas, MS  TL: Peru

Euptychia [n. sp.#8] Lamas, MS  TL: Peru

Euptychia [n. sp.#9] Lamas, MS  TL: Peru

Euptychia [n. sp.#10] Lamas, MS  TL: Peru

Euptychia [n. sp.#11] Brévignon, MS  TL: French Guiana

Euptychia [n. sp.#12] Brévignon, MS  TL: French Guiana

Euptychia [n. sp.#13] Lamas, MS  TL: Peru

Euptychia [n. sp.#14] Neild, MS  TL: Venezuela

Euptychia [n. sp.#15] Neild, MS  TL: Venezuela

Euptychoides  Forster, 1964  

Euptychoides albofasciata (Hewitson, 1869)    iN  M T

Euptychoides inani (Staudinger, [1886])  ref.201  iN  M T

Euptychoides laccine (C. Felder & R. Felder, 1867)    iN  M T S

Euptychoides pseudosaturnus Forster, 1964    iN  M T

Cisandina  Nakahara & Espeland, 2022  

Cisandina castanya Lamas & Nakahara, 2022  TL: Peru

Cisandina esmeralda Nakahara & E. Barbosa, 2022  TL: Brazil

Cisandina fida (Weymer, 1911)  ref.265  M T

Cisandina lea (Cramer, 1777)    iN

Cisandina philippa (A. Butler, 1867)  TL: Brazil (Am)  M T S

Cisandina sanmarcos (Nakahara & Lamas, 2018)  ref.265

Cisandina trinitensis (Brévignon & M. Benmesbah, 2011)  ref.179

Optimandes  Marín, Nakahara & Willmott, 2019  

Optimandes eugenia (C. Felder & R. Felder, 1867)    iN

Optimandes eugenia eugenia (C. Felder & R. Felder, 1867)  ref.316  M T

Optimandes eugenia transversa (Weymer, 1911)  ref.316  M T

Optimandes mocha Willmott, J. Hall & Lamas, 2019  ref.316  iN

Graphita  Nakahara, Marín & Barbosa, 2016  

Graphita griphe (C. Felder & R. Felder, 1867)  ref.254  iN  M T L

Godartiana  Forster, 1964  

Godartiana astronesthes Lamas & Nakahara, 2018  ref.261  iN

Godartiana byses (Godart, [1824])    iN  M T

Godartiana amadoi M. Paluch, Zacca & Freitas, 2017  ref.364

Godartiana muscosa (A. Butler, 1870)    iN  M T

Godartiana luederwaldti (Spitz, 1931)  ref.365  iN  M T

Godartiana armilla (A. Butler, 1867)  ref.365  iN  M T

Godartiana [n. sp.] Lamas, MS  Peru

Hermeuptychia  Forster, 1964  

Hermeuptychia sosybius (Fabricius, 1793)  Carolina Satyr  C TX & E OK, E to FL, N to S OH & VA  iN  M T OD TI S1 S2 S3 S4 IL Ma L1 L2 L3 L4 I G Txt

Hermeuptychia hermybius Grishin, 2014  South Texas Satyr  S TX & NE Mexico; ref.201,187  iN  M T1 T2 T3 OD S IL L I G

Hermeuptychia occidentalis Grishin, 2021  West Mexico Satyr  SW Mexico (Jal, Mor, Gue)

Hermeuptychia intricata Grishin, 2014  Intricate Satyr  SE US coastal plains to (possibly) Costa Rica; ref.201,187  iN  M T OD S IL Ma L Ha G

Hermeuptychia clara Nakahara, D. Tan, Lamas & Willmott, 2016  ref.201,256  iN

Hermeuptychia cucullina (Weymer, 1911)  Cucullina Satyr    iN  M T OD

Hermeuptychia gisella (Hayward, 1957)  Gisella Satyr  differences (genitalia, mitDNA) from H. cucullina support species status ...  iN  M T OD S G

Hermeuptychia atalanta (A. Butler, 1867)  Atalanta Satyr    iN  M T OI OD S G

Hermeuptychia lupita (Reakirt, [1867])  Lupita’s Satyr  SE Mexico  iN

Hermeuptychia hermes (Fabricius, 1775)  Hermes Satyr  probably in South America only  iN  M T OD S

Hermeuptychia canthe (Hübner, [1811])    iN  M T OD

Hermeuptychia sinuosa Grishin, 2021  Wavy Stayr  MX to Guatemala; ref.20  M T

Hermeuptychia maimoune (A. Butler, 1870)  Maimoune Satyr  ref.201  iN  M T OD S

Hermeuptychia pimpla (C. Felder & R. Felder, 1862)  Pimpla Satyr  ref.201  iN  M T OD S

Hermeuptychia harmonia (A. Butler, 1867)  Harmonia Satyr  [= calixta], Costa Rica to W Ecuador  iN  M T OD S L

Hermeuptychia fallax (C. Felder & R. Felder, 1862)  Fallax Satyr    iN

Hermeuptychia fallax fallax (C. Felder & R. Felder, 1862)  TL: [Peru]  M T OD S

Hermeuptychia fallax marinha Anken, 1994  TL: Brazil (RJ)  M T OI OD G

Hermeuptychia undulata (A. Butler, 1867)  ref.371  iN  M T OD S

Hermeuptychia [n. sp.#1] Lamas, MS  TL: Colombia

Hermeuptychia [n. sp.#2] Lamas, MS  TL: Peru

Hermeuptychia [n. sp.#3] Lamas, MS  TL: Peru

Hermeuptychia [n. sp.#4] Lamas, MS  TL: Peru

Huberonympha  Viloria & Costa, 2016  

Huberonympha neildi Viloria, Costa, Fratello & Nakahara, 2016  TL: Venezuela

Inbio  Nakahara & Espeland, 2015  

Inbio hilara (C. Felder & R. Felder, 1867)  Tawny-cornered Satyr  S Mexico to Colombia; ref.253  iN  M T S L

Magneuptychia  Forster, 1964  

Magneuptychia libye (Linnaeus, 1767)  Blue-gray Satyr  SE Mexico to Amazon Basin  iN  M T S L

Magneuptychia lethra (Möschler, 1883)    iN  M T S

Modica  Zacca, Casagrande & Willmott, [2023]  

Modica fugitiva (Lamas, [1997])    iN  M S L

Modica kamel (Benmesbah & Zacca, 2018)  ref.166

Modica maripa (Brévignon, 2005)  TL: French Guiana

Modica myncea (Cramer, 1780)  ref.201  iN  M T S

Modica confusa (Staudinger, 1887)  Confused Satyr  SE Mexico to Brazil; ref.201  iN  M T S L

Deltaya  Willmott, Nakahara & Espeland, [2023]  

Deltaya andrei (Zacca, Casagrande & O. Mielke, 2017)  ref.201,366  iN

Deltaya ocypete (Fabricius, 1776)  ref.201  iN  M T

Deltaya pallema (Schaus, 1902)  ref.201  iN  M T S

Deltaya louisammour (Benmesbah & Zacca, 2018)  ref.201,166

Deltaya opima (Weymer, 1911)  ref.201  iN  M T

Occulta  Nakahara & Willmott, [2023]  

Occulta ocnus (A. Butler, 1867)  ref.201  iN  M T S

Xenovena  Marín & Nakahara, [2023]  

Xenovena murrayae (Brévignon, 2005)  ref.174  iN

Trico  Nakahara & Espeland, [2023]  

Trico tricolor (Hewitson, 1850)    iN

Trico tricolor tricolor (Hewitson, 1850)  TL: [Brazil (Pa)]  M T S

Trico tricolor fulgora (A. Butler, 1869)  TL: Peru  M T

Scriptor  Nakahara & Espeland, 2020  

Scriptor sphenophorus Lamas & Nakahara, 2020    iN

Modestia  Viloria & M. Benmesbah, 2021  ref.167

Modestia agnata (Schaus, 1913)  Agnata Satyr  Costa Rica (Atlantic slope); ref.201  iN  M T S

Modestia analis (Godman, 1905)  ref.201  iN  M T S

Modestia drymo (Schaus, 1913)  Drymo Satyr  Costa Rica (Atlantic slope); ref.201  M T S

Modestia gomezi (Singer, DeVries & P. Ehrlich, 1983)  Gomez' Satyr  Costa Rica; ref.167  iN  M T S L

Modestia harpyia (C. Felder & R. Felder, 1867)  ref.201  iN

Modestia harpyia harpyia (C. Felder & R. Felder, 1867)  TL: Surinam  M T

Modestia harpyia batesii (A. Butler, 1867)  TL: Brazil (Pa)  M T S

Modestia modesta (A. Butler, 1867)  ref.167  iN  M T S

Modestia remypignoux M. Benmesbah & Viloria, 2021  ref.167

Modestia remypignoux remypignoux M. Benmesbah & Viloria, 2021  ref.167

Modestia remypignoux shueyi M. Benmesbah & Viloria, 2021  ref.167

Modestia sylvina (C. Felder & R. Felder, 1867)  ref.201  iN  M T

Colombeia  Viloria, Andrade & Le Crom, 2019  

Colombeia hotchkissi (Dyar, 1913)  ref.201  M T

Colombeia mycalesis (Röber, 1927)  ref.156  iN  M T

Colombeia nossis (Hewitson, 1862)  ref.201  iN  M T

Omacha  Andrade, Viloria, Henao & Le Crom, 2019  

Omacha pax (Huertas, Lamas, G. Fagua & Willmott, 2016)  ref.156  iN

Megeuptychia  Forster, 1964  

Megeuptychia antonoe (Cramer, 1775)  Cramer's Satyr  S Mexico to Amazon Basin  iN  M T S L

Megeuptychia monopunctata Willmott & J. Hall, 1995    iN  M T S

Megeuptychia souadae (M. Benmesbah, 2015)  ref.165  iN

Megisto  Hübner, [1819]  

Megisto cymela (Cramer, 1777)  Little Wood-Satyr    iN

Megisto cymela cymela (Cramer, 1777)  TL: South Africa  M T OI S L1 L2 L3 L4 L5 Ha G

Megisto cymela viola (Maynard, 1891)  TL: USA (Fl)  M TI S L

Llorenteana  Viloria & Luis-Martínez, [2019]  

Llorenteana pellonia (Godman, 1901)  Plateau Satyr  NW Mexico; ref.296  M T S

Neonympha  Hübner, 1818  

Neonympha areolatus (J. E. Smith, 1797)  Georgia Satyr    iN

Neonympha areolatus septentrionalis W. Davis, 1924  TL: USA (NJ)  M S

Neonympha areolatus areolatus (J. E. Smith, 1797)  [= helicta]; ref.134  M T S L1 L2 Ha

Neonympha areolatus dadeensis Gatrelle, 1999  TL: USA (Fl)  M T

Neonympha mitchellii French, 1889  Mitchell's Satyr    iN

Neonympha mitchellii mitchellii French, 1889  TL: USA (Mi)  M S1 S2 L

Neonympha mitchellii francisci Parshall & Kral, 1989  TL: USA (NC)  M T S L Ha

Moneuptychia  Forster, 1964  

Moneuptychia castrensis (Schaus, 1902)    iN  M T

Moneuptychia giffordi Freitas, Emery & O. Mielke, 2010  ref.209  iN

Moneuptychia itapeva Freitas, 2007  ref.208  iN

Moneuptychia melchiades (A. Butler, 1877)  TL: Argentina  M T

Moneuptychia montana Freitas, 2015  ref.214

Moneuptychia pervagata Freitas, R. Siewert & O. Mielke, 2015  ref.214  iN

Moneuptychia romanina (Bryk, 1953)    iN  M T

Moneuptychia soter (A. Butler, 1877)    iN  M T

Moneuptychia vitellina Freitas & E. Barbosa, 2015  ref.214  iN

Moneuptychia viviana (Romieux, 1927)  ref.164  M T

Moneuptychia wahlbergi Freitas, E. Barbosa, R. Siewert & O. Mielke, 2015  ref.214  iN

Paramacera  A. Butler, 1868  

Paramacera allyni L. Miller, 1972  Arizona Pine-Satyr  SE Arizona, far NW Mexico; species status supported by differences in and ...; ref.16  M T S IL L I G

Paramacera xicaque (Reakirt, [1867])  Mexican Pine-Satyr  E & central Mexico  iN  M T S IL L G

Paramacera rubrosuffusa L. Miller, 1972  Mexican Pine-Satyr  S Oaxaca (Sierra Madre del Sur); ref.16  M T S L

Paramacera copiosa L. Miller, 1972  Guerrero Pine-Satyr  N Oaxaca, Guerrero  M T S IL G

Paramacera chinanteca L. Miller, 1972  Oaxacan Pine-Satyr  N Oaxaca  iN  M T S IL L G

Pareuptychia  Forster, 1964  

Pareuptychia binocula (A. Butler, 1869)    iN  M T S

Pareuptychia difficilis Forster, 1964  TL: Bolivia  M T

Pareuptychia hervei Brévignon, 2005  ref.174

Pareuptychia hesionides Forster, 1964    iN

Pareuptychia hesionides hesionides Forster, 1964  TL: Bolivia  M T S

Pareuptychia hesionides deviae Brévignon, 2005  ref.174

Pareuptychia lydia (Cramer, 1777)    iN  M T

Pareuptychia metaleuca (Boisduval, 1870)  White-banded Satyr  E & S Mexico to Panama  iN

Pareuptychia metaleuca metaleuca (Boisduval, 1870)  TL: Guatemala  M T S L1 L2

Pareuptychia metaleuca [n. ssp.#1] Lamas, MS  TL: Colombia

Pareuptychia metaleuca [n. ssp.#2] Lamas, MS  TL: Peru

Pareuptychia metaleuca [n. ssp.#3] Lamas, MS  TL: Peru

Pareuptychia metaleuca [n. ssp.#4] Lamas, MS  TL: French Guiana

Pareuptychia milleri Nakahara, M. Marín & Neild, 2016  ref.255

Pareuptychia ocirrhoe (Fabricius, 1776)    iN

Pareuptychia ocirrhoe Northern segregate ,  [= hesione], E & S Mexico to Ecuador  M S L1 L2

Pareuptychia ocirrhoe ocirrhoe (Fabricius, 1776)  TL: Surinam  M T

Pareuptychia ocirrhoe interjecta (R.F. d'Almeida, 1952)  TL: Brazil (RJ)  M T

Pareuptychia ocirrhoe [n. ssp.#1] Lamas, MS  TL: Mexico

Pareuptychia ocirrhoe [n. ssp.#2] Lamas, MS  TL: Peru

Pareuptychia ocirrhoe [n. ssp.#3] Lamas, MS  TL: Trinidad

Pareuptychia summandosa (Gosse, 1880)    iN  M T

Pareuptychia [n. sp.] Lamas, MS  Peru

Paryphthimoides  Forster, 1964  

Paryphthimoides poltys (Prittwitz, 1865)    iN

Paryphthimoides poltys poltys (Prittwitz, 1865)  TL: Brazil (RJ)  M T OD

Paryphthimoides poltys numilia (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Paryphthimoides vestigiata (A. Butler, 1867)    iN  M T

Paryphthimoides fridae Zacca, Casagrande, Huertas & Nakahara, 2020  ref.369

Paryphthimoides jorupe Zacca, Casagrande, Checa & Willmott, 2020  ref.369

Paryphthimoides terrestris (A. Butler, 1867)  Cryptic Satyr    iN

Paryphthimoides terrestris terrestris (A. Butler, 1867)  E Mexico to Brazil  M T S L

Paryphthimoides terrestris araguaianus Zacca, Casagrande & O. Mielke, 2020  ref.369

Paryphthimoides terrestris grevei Zacca, Casagrande & O. Mielke, 2020  ref.369

Paryphthimoides terrestris muyrakytan Zacca, Casagrande & O. Mielke, 2020  ref.369

Paryphthimoides touloulou (M. Benmesbah, 2015)  ref.369

Paryphthimoides brixius (Godart, [1824])    iN

Paryphthimoides brixius brixius (Godart, [1824])  ref.369  M S

Paryphthimoides brixius madre Zacca, O. Mielke, Casagrande & Lamas, 2020  ref.369

Paryphthimoides brixius rouranensis Brévignon, 2020  ref.369

Paryphthimoides brixiola (A. Butler, 1867)  TL: Brazil (Pa)  M T

Paryphthimoides flavofascia (Zacca & R. Siewert, 2014)  ref.369  iN

Paryphthimoides pseudoconfusa (Singer, DeVries & P. Ehrlich, 1983)  Gold-stained Satyr  E Mexico to Panama; ref.201  iN  M T S L

Paryphthimoides sheba (Brévignon & M. Benmesbah, 2011)  ref.201,179

Emeryus  Zacca, Casagrande & O. Mielke, 2020  

Emeryus argulus (Godart, [1824])    iN

Emeryus argulus argulus (Godart, [1824])  ref.368  M T

Emeryus argulus magnum Zacca, Casagrande & O. Mielke, 2019  ref.368

Emeryus difficilis (Forster, 1964)  ref.368  iN  M T

Emeryus numeria (C. Felder & R. Felder, 1867)  ref.368  iN  M T

Pharneuptychia  Forster, 1964  

Pharneuptychia haywardiana Lamas, 2010  TL: Bolivia  M T

Pharneuptychia estoraquensis Henao & Meneses, [2018]  TL: Colombia

Pharneuptychia innocentia (C. Felder & R. Felder, 1867)    iN  M T

Pharneuptychia phares (Godart, [1824])    iN  M T

Pharneuptychia pharnabazos (Bryk, 1953)    iN  M T

Pharneuptychia pharnaces (Weymer, 1911)  TL: Brazil

Pindis  R. Felder, 1869  

Pindis squamistriga R. Felder, 1869  Variable Satyr  E & W Mexico to Guatemala  iN  M T S L I

Pseudeuptychia  Forster, 1964  

Pseudeuptychia hemileuca (Staudinger, [1886])  TL: Colombia  M T

Pseudeuptychia languida (A. Butler, 1871)    iN

Pseudeuptychia languida languida (A. Butler, 1871)  TL: Colombia  M T

Pseudeuptychia languida austrina Nakahara & Lamas, 2018  ref.261

Pseudeuptychia cuzquenya Nakahara & Lamas, 2018  ref.261

Pseudeuptychia callichloris (A. Butler, 1867)  ref.201  iN  M T

Pseudeuptychia herseis (Godart, [1824])  ref.201  iN  M T

Pseudeuptychia rectilinea (Brévignon, Rosant, Lamas & Willmott, 2019)  ref.201,180  iN

Pseudeuptychia marica (Weymer, 1911)  ref.201  iN  M T S

Satyrotaygetis  Forster, 1964  

Satyrotaygetis satyrina (H. Bates, 1865)  Wide-bordered Satyr  [= gigas (Butler)], S Mexico to Panama; ref.201  iN  M T S L1 L2

Satyrotaygetis iris (C. Felder & R. Felder, 1867)  ref.201  iN  M T S

Satyrotaygetis tiessa (Hewitson, 1869)  Tiessa Satyr  Nicaragua to W Ecuador  iN  M T S L

Splendeuptychia  Forster, 1964  

Splendeuptychia ambra (Weymer, [1911])    iN

Splendeuptychia ashna (Hewitson, 1869)    iN  M T

Splendeuptychia purusana (Aurivillius, 1929)    iN  M T

Splendeuptychia tupinamba A. Freitas, Huertas & Rosa, 2021  ref.201  iN

Splendeuptychia gera (Hewitson, 1850)  ref.201  iN

Splendeuptychia gera gera (Hewitson, 1850)  TL: Brazil  M T

Splendeuptychia gera nortia (Hewitson, 1862)  TL: Brazil  M T S

Splendeuptychia nobilis (Weymer, 1911)  ref.201

Splendeuptychia moderata (Weymer, 1911)  ref.201  iN  M T

Splendeuptychia metagera (A. Butler, 1867)  ref.201  M T

Splendeuptychia [n. sp.#1] Lamas, MS  TL: Peru

Splendeuptychia [n. sp.#2] Lamas, MS  TL: Peru

Splendeuptychia [n. sp.#3] Lamas, MS  TL: Colombia

Argentaria  Huertas & Willmott, 2023  

Argentaria argyropsacas (Bryk, 1953)  ref.201  iN  M T

Argentaria clementia (A. Butler, 1877)  ref.201  iN  M T S

Argentaria clorimena (Stoll, 1790)  ref.201  iN  M T

Argentaria cosmophila (Hübner, 1823)  ref.201  M T

Argentaria hygina (A. Butler, 1877)  ref.201  iN  M T

Argentaria itonis (Hewitson, 1862)  ref.201  iN  M T

Argentaria jadea (Brévignon & M. Benmesbah, 2011)  ref.201,179  iN

Argentaria kendalli (L. Miller, 1976)  Kendall's Satyr  NE Mexico; ref.201  iN  M T S L

Argentaria libitina (A. Butler, 1870)  ref.201  iN  M T

Argentaria pagyris (Godart, [1824])  ref.201  iN  M T

Argentaria quadrina (A. Butler, 1869)  ref.201  iN  M T

Argentaria salvini (A. Butler, 1867)  Salvin's Satyr  Panama; ref.201  iN  M T S

Argentaria telesphora (A. Butler, 1877)  ref.201  M T

Argentaria zischkai (Forster, 1964)  ref.201  M T

Argentaria [n. sp.#1] Lamas, MS  TL: Peru

Argentaria [n. sp.#2] Lamas, MS  TL: Peru

Argentaria [n. sp.#3] Lamas, MS  TL: Peru

Argentaria [n. sp.#4] Lamas, MS  TL: Peru

Argentaria [n. sp.#5] Lamas, MS  TL: Peru

Argentaria [n. sp.#6] Lamas, MS  TL: Peru

Argentaria [n. sp.#7] Lamas, MS  TL: Peru

Argentaria [n. sp.#8] Lamas, MS  TL: ?

Argentaria [n. sp.#9] Lamas, MS  TL: Brazil (SP)

Argentaria [n. sp.#10] Lamas, MS  TL: Brazil (MG)

Argentaria [n. sp.#11] Lamas, MS  TL: Peru

Argentaria [n. sp.#12] Lamas, MS  TL: Peru

Argentaria [n. sp.#13] Lamas, MS  TL: Peru

Argentaria [n. sp.#14] Lamas, MS  TL: Venezuela

Argentaria [n. sp.#15] Lamas, MS  TL: Peru

Argentaria [n. sp.#16] Lamas, MS  TL: Brazil

Argentaria [n. sp.#17] Lamas, MS  TL: Brazil (ES)

Argentaria [n. sp.#18] Lamas, MS  TL: Peru

Saurona  Huertas & Willmott, [2023]  

Saurona aurigera (Weymer, [1911])  ref.201  iN  M T

Saurona triangula (Aurivillius, 1929)  ref.201  iN  M T

Saurona [n. sp.] Lamas, MS  Peru

Nhambikuara  Freitas, Barbosa & Zacca, 2018  

Nhambikuara mima (A. Butler, 1867)  ref.216  iN  M T

Nhambikuara cerradensis Freitas, Barbosa & Zacca, 2018  ref.216  iN

Nhambikuara ackeryi (Huertas, C. Rios & LeCrom, 2009)  ref.201  iN

Nhambikuara doxes (Godart, [1824])  ref.201  iN  M T

Nhambikuara furina (Hewitson, 1862)  ref.201  iN  M T

Nhambikuara junonia (A. Butler, 1867)  ref.201  iN  M T

Nhambikuara latia (A. Butler, 1867)  ref.201  M T

Nhambikuara mercedes (Huertas, 2011)  ref.201,226

Nhambikuara toynei (Willmott & J. Hall, 1995)  ref.201  iN  M T

Stegosatyrus  Zacca, O. Mielke & Pyrcz, 2013  

Stegosatyrus hemiclara Pyrcz, Boyer & Zacca, 2013  ref.363

Stegosatyrus imbrialis (Weeks, 1901)  ref.363  iN  M T

Stegosatyrus ocelloides (Schaus, 1902)  ref.363  iN  M T

Stegosatyrus periphas (Godart, [1824])  ref.363  iN  M T

Stevenaria  Viloria, Costa, Neild & Nakahara, 2016  

Stevenaria divergens (A. Butler, 1867)  TL: Brazil (Am)  M T

Stevenaria nakaharai Viloria, Costa, Fratello & Neild, 2016  ref.189

Stevenaria yutajensis Viloria & Costa, 2016  ref.189

Taydebis  A. Freitas, 2003  

Taydebis clarissa (A. Freitas & O. Mielke, 2011)  ref.371  iN

Taydebis guria Zacca, Casagrande & O. Mielke, 2021  TL: Brazil

Taydebis melobosis (Capronnier, 1874)  ref.371  M T

Sepona  Freitas & E. Barbosa, 2016  

Sepona punctata (Weymer, 1911)  ref.215  iN  M T

Harjesia  Forster, 1964  

Harjesia argentata Nakahara, Zacca & Lamas, 2018  ref.261

Harjesia blanda (Möschler, 1877)    iN  M T S

Harjesia obscura (A. Butler, 1867)    iN  M T S L

Harjesia [n. sp.#1] Lamas, MS  TL: Peru

Harjesia [n. sp.#2] Lamas, MS  TL: Peru

Harjesia [n. sp.#3] Lamas, MS  TL: Peru

Harjesia [n. sp.#4] Lamas, MS  TL: Peru

Parataygetis  Forster, 1964  

Parataygetis albinotata (A. Butler, 1867)    iN  M T S

Parataygetis lineata (Godman & Salvin, 1880)  White-lined Satyr  Costa Rica to Colombia  iN  M T S L

Posttaygetis  Forster, 1964  

Posttaygetis penelea (Cramer, 1777)  Yellow-spiked Satyr  Costa Rica to Brazil  iN  M T S L

Forsterinaria  R. Gray, 1973  

Forsterinaria itatiaia C. Peña & Lamas, 2005  ref.276  iN

Forsterinaria quantius (Godart, [1824])    iN  M T

Forsterinaria necys (Godart, [1824])    iN  M T

Forsterinaria pronophila (A. Butler, 1867)  ref.243  iN  M T

Forsterinaria neonympha (C. Felder & R. Felder, 1867)  White-dotted Satyr  [= umbracea; = polyphemus; = cyclops], S Mexico to W Ecuador  iN  M T S L

Forsterinaria stella (Hayward, 1957)    iN  M T S L

Forsterinaria pyrczi C. Peña & Lamas, 2005  ref.276  iN

Forsterinaria proxima (Hayward, 1957)    iN  M T S

Forsterinaria pseudinornata (Forster, 1964)  TL: Ecuador  M T

Forsterinaria punctata C. Peña & Lamas, 2005  ref.276

Forsterinaria anophthalma (C. Felder & R. Felder, 1867)    iN  M T

Forsterinaria enjuerma C. Peña & Lamas, 2005  ref.276  iN

Forsterinaria pichita C. Peña & Lamas, 2005  ref.276  iN  M S

Forsterinaria guaniloi C. Peña & Lamas, 2005    iN

Forsterinaria pilosa C. Peña & Lamas, 2005  ref.276

Forsterinaria falcata C. Peña & Lamas, 2005  ref.276

Forsterinaria rotunda C. Peña & Lamas, 2005  ref.276

Forsterinaria emo A. Zubek, T. Pyrcz & Boyer, 2014  ref.374

Forsterinaria hannieri A. Zubek, T. Pyrcz & Boyer, 2011  ref.376

Forsterinaria difficilis (Forster, 1964)  TL: Bolivia  M T

Forsterinaria rustica (A. Butler, 1868)    iN

Forsterinaria rustica rustica (A. Butler, 1868)  TL: Bolivia  M T S

Forsterinaria rustica villarresi (Dognin, 1887)  TL: Ecuador  M T S

Forsterinaria rustica glendita C. Peña & Lamas, 2005  TL: Peru

Forsterinaria antje C. Peña & Lamas, 2005  ref.276  M S

Forsterinaria inornata (C. Felder & R. Felder, 1867)    iN  M T S

Forsterinaria pallida C. Peña & Lamas, 2005    iN

Forsterinaria pallida aurita C. Peña & Lamas, 2005  TL: Ecuador  M L

Forsterinaria pallida coipa C. Peña & Lamas, 2005  TL: Peru  M S

Forsterinaria pallida pallida C. Peña & Lamas, 2005  TL: Peru

Forsterinaria boliviana (Godman, 1905)    iN  M T S

Forsterinaria chaniorum Viloria & Costa, 2019  TL: Venezuela

Orotaygetis  Nakahara & Zacca, 2018  

Orotaygetis surui Nakahara, Zacca & Lamas, 2018  ref.261

Pseudodebis  Forster, 1964  

Pseudodebis celia (Cramer, 1779)  Celia Satyr  Nicaragua to Brazil & Bolivia; ref.243  iN  M T S L

Pseudodebis darrenthroopi Nakahara & Willmott, 2022  TL: Costa Rica

Pseudodebis nakamurai Nakahara & Willmott, 2021  TL: Ecuador

Pseudodebis pieti Nakahara & Willmott, 2021  TL: Ecuador

Pseudodebis tigrillo Nakahara & Willmott, 2022  TL: Ecuador

Pseudodebis puritana (Weeks, 1902)    iN

Pseudodebis puritana puritana (Weeks, 1902)  ref.243  M T S

Pseudodebis puritana [n. ssp.] L. Miller, MS  Brazil (Pr)  M T

Pseudodebis marpessa (Hewitson, 1862)    iN  M T S

Pseudodebis dubiosa Forster, 1964    iN  M T

Pseudodebis euptychidia (A. Butler, 1868)    iN  M T

Pseudodebis valentina (Cramer, 1779)    iN  M T S

Pseudodebis vrazi (Kheil, 1896)  ref.261  M T

Pseudodebis zimri (A. Butler, 1869)  Butler's Satyr  SE Mexico to Colombia  iN  M T S L

Taguaiba  A. Freitas, Zacca & R. Siewert, [2023]  

Taguaiba rectifascia (Weymer, 1907)  ref.201  iN  M T

Taguaiba ypthima (Hübner, [1821])  ref.201  iN  M T

Taguaiba drogoni (Siewert, Zacca, Dias & Freitas, 2013)  Drogon's Satyr  ref.201

Taguaiba servius (Weymer, 1910)  ref.201

Taguaiba fulginia (R.F. d'Almeida, 1922)  ref.201  M T

Taygetina  Forster, 1964  

Taygetina banghaasi (Weymer, 1910)  Bang-Haas' Satyr  Costa Rica & Bolivia  iN  M T S L

Taygetina accacioi Nakahara & Freitas, 2019  ref.263

Taygetina brocki Lamas & Nakahara, 2019  ref.263

Taygetina oreba (A. Butler, 1870)  Oreba Satyr  Costa Rica to Peru; ref.243  iN  M T S

Taygetina gulnare (A. Butler, 1870)    iN  M T S L

Taygetina weymeri (Draudt, 1912)  Mexican Satyr  W Mexico; ref.243  iN  M T S L

Taygetina kerea (A. Butler, 1869)  Kerea Satyr  E & W Mexico to Paraguay & Argentina; ref.243  iN  M T S L

Taygetina peribaea (Godman & Salvin, 1880)  ref.243  M T S

Taygetis  Hübner, [1819]  

Taygetis acuta Weymer, 1910    iN  M T

Taygetis angulosa Weymer, 1907    iN  M T S

Taygetis asterie Weymer, 1910    iN

Taygetis chiquitana Forster, 1964    iN  M T

Taygetis chrysogone E. Doubleday, [1849]    iN  M T S

Taygetis cleopatra C. Felder & R. Felder, 1867    iN  M T S

Taygetis echo (Cramer, 1775)    iN

Taygetis echo echo (Cramer, 1775)  TL: Surinam  M T

Taygetis echo koepckei Forster, 1964  TL: Peru  M T

Taygetis echo velutina Staudinger, [1887]  TL: Venezuela  M T

Taygetis echo [n. ssp.] L. Miller, MS  Trinidad

Taygetis elegia Weymer, 1910    iN  M T

Taygetis [n. sp.#1] L. Miller, MS  TL: Peru

Taygetis godmani Weymer, 1910    iN  M T

Taygetis inambari L. Miller & Lamas, 1999    iN  M T

Taygetis inconspicua Draudt, 1931  Inconspicuous Satyr  SE Mexico, Guatemala  iN  M T

Taygetis kharisma Brévignon & M. Benmesbah, 2011  ref.179

Taygetis laches Fabricius, 1793    iN

Taygetis laches laches (Fabricius, 1793)  TL: Guyana  M T

Taygetis laches marginata Staudinger, [1887]  TL: Brazil (SP)  M T

Taygetis laches [n. ssp.] L. Miller, MS  Peru

Taygetis [n. sp.#2] L. Miller, MS  TL: Peru

Taygetis larua C. Felder & R. Felder, 1867  Laura's Satyr  Panama to Bolivia  iN  M T S L

Taygetis leuctra A. Butler, 1870  Leuctra Satyr  Chiapas to Panama (at least)  iN  M T S L

Taygetis mermeria (Cramer, 1776)    iN

Taygetis mermeria mermeria (Cramer, 1776)  TL: Surinam  M T S L

Taygetis mermeria excavata A. Butler, 1868  E Mexico to Panama  M T S L

Taygetis mermeria griseomarginata L. Miller, 1978  W Mexico  M T S

Taygetis nympha A. Butler, 1868    iN  M T

Taygetis oyapock Brévignon, 2007  ref.175  iN

Taygetis rufomarginata Staudinger, 1888  Rufous-margined Satyr  [S Mexico?], Honduras, Costa Rica & Panama  iN

Taygetis rufomarginata rufomarginata Staudinger, 1888  [S Mexico?], Honduras, Costa Rica & Panama  M T S L

Taygetis rufomarginata tapulunmi Brévignon, 2005  TL: French Guiana

Taygetis salvini Staudinger, [1887]    iN  M T

Taygetis sosis Hopffer, 1874  Sosis Satyr  [= xenana godmani], [S Mexico] to Brazil & Peru  iN  M S L

Taygetis sylvia H. Bates, 1866  Sylvia's Satyr  Panama to Bolivia  iN  M T S L

Taygetis thamyra (Cramer, 1779)  Thamyra Satyr  E Mexico to S Brazil  iN  M T S L

Taygetis tripunctata Weymer, 1907    iN  M T S

Taygetis uncinata Weymer, 1907  Hook-lined Satyr  W & S Mexico, Guatemala  iN  M T S L

Taygetis uzza A. Butler, 1869  Uzza Satyr  [= salvini], [S Mexico?], Guatemala to Colombia  iN  M T OI S L

Taygetis virgilia (Cramer, 1776)  Stub-tailed Satyr  E & W Mexico to Brazil  iN  M T S L

Taygetis zippora A. Butler, 1869  TL: French Guiana  M T

Yphthimoides  Forster, 1964  

Yphthimoides angularis (A. Butler, 1867)    iN  M T

Yphthimoides bella E. Barbosa & Freitas, 2015  ref.162

Yphthimoides blanquita E. Barbosa, M. Marín & Freitas, 2016  ref.163  iN

Yphthimoides borasta (Schaus, 1902)    iN  M T S

Yphthimoides celmis (Godart, [1824])    iN

Yphthimoides acmenis (Hübner, 1823)    iN  M T

Yphthimoides cipoensis Freitas, 2004  ref.207  iN

Yphthimoides gabriela E. Barbosa, Freitas & M. Paluch, 2015  ref.162

Yphthimoides iserhardi Freitas & E. Barbosa, 2015  ref.162

Yphthimoides kinyoni Lamas & Nakahara, 2021  TL: Peru

Yphthimoides leguialimai (Dyar, 1913)    iN  M T

Yphthimoides manasses (C. Felder & R. Felder, 1867)  TL: Brazil (Ba)  M T

Yphthimoides nareta E. Barbosa & Freitas, 2016  ref.163

Yphthimoides ochracea (A. Butler, 1867)    iN  M T L

Yphthimoides ordinaria Freitas, L. Kaminski & O. Mielke, 2012  ref.211  iN

Yphthimoides pacta (Weymer, 1911)    iN  M T

Yphthimoides patricia (Hayward, 1957)    iN

Yphthimoides renata (Stoll, 1780)  Renata Satyr  S Mexico to W Ecuador  iN  M T S L1 L2

Yphthimoides saltuensis (Hayward, 1962)  TL: ?  M T

Yphthimoides straminea (A. Butler, 1867)    iN  M T

Yphthimoides yphthima (C. Felder & R. Felder, 1867)    iN  M T

Yphthimoides inornata (Hayward, 1962)  ref.277  iN  M T

Yphthimoides [n. sp.#1] Lamas, MS  TL: Peru

Yphthimoides [n. sp.#2] Lamas, MS  TL: Peru

Malaveria  Viloria & M. Benmesbah, 2021  ref.167

Malaveria affinis (A. Butler, 1867)  argyrospila A.G. Butler 1867 0 Euptychia Brazil (Am) S 3550 156 ...; ref.167  iN  M T

Malaveria alcinoe (C. Felder & R. Felder, 1867)  Simple Satyr  S Mexico to Bolivia; ref.167  iN  M T S L

Malaveria argyrospila (A. Butler, 1867)  ref.201  M T

Malaveria ballofi M. Benmesbah & Viloria, 2021  ref.167

Malaveria bottoi M. Benmesbah & Viloria, 2021  ref.167

Malaveria duponti M. Benmesbah & Murienne, 2021  ref.167

Malaveria erigone (A. Butler, 1867)  ref.201  M T

Malaveria grimon (Godart, [1824])  ref.201  iN

Malaveria guenzeli (Anken, 1994)  ref.201

Malaveria iseai (Viloria, [2022])  ref.201

Malaveria maepius (Godart, [1824])    iN  M T

Malaveria mimas (Godman, 1905)  ref.167  iN  M T S

Malaveria mimula (Hayward, 1954)    iN  M T

Malaveria mythra (Weymer, 1911)    iN  M T

Malaveria nebulosa (A. Butler, 1867)  ref.167  iN  M T

Malaveria neomaenas (Hayward, 1967)  ref.167  M T

Malaveria rodriguezi M. Benmesbah & Viloria, 2021  ref.167  iN

Koutalina  Viloria & M. Benmesbah, 2021  ref.167

Koutalina pamela (Hayward, 1957)  ref.167  iN

Zischkaia  Forster, 1964  

Zischkaia amalda (Weymer, 1911)  TL: Bolivia  M T

Zischkaia arctoa Nakahara, 2019  ref.262

Zischkaia chullachaki Nakahara & Zacca, 2019  ref.262  iN

Zischkaia baku Zacca, Dolibaina & Dias, 2019  ref.262  iN

Zischkaia arenisca Nakahara, Willmott & J. Hall, 2019  ref.262  iN

Zischkaia argyrosflecha Nakahara, L. Miller & Huertas, 2019  ref.262

Zischkaia pacarus (Godart, [1824])    iN  M T

Zischkaia abanico Nakahara & Petit, 2019  ref.262

Zischkaia saundersii (A. Butler, 1867)    iN  M T

Zischkaia josti Nakahara & Kleckner, 2019  ref.262

Zischkaia mielkeorum Dolibaina, Dias & Zacca, 2019  ref.262

Zischkaia warreni Dias, Zacca & Dolibaina, 2019  ref.262  iN

Amphidecta  A. Butler, 1867  

Amphidecta calliomma (C. Felder & R. Felder, 1862)  Calliomma Satyr  Colombia, Panama  iN  M T S L

Amphidecta clio (Weymer, 1911)  ref.201  iN  M T

Amphidecta pignerator A. Butler, 1867  Stub-tailed Satyr    iN

Amphidecta pignerator pignerator A. Butler, 1867  SE Mexico (CHIS) to Amazon Basin  M T S L

Amphidecta pignerator simplicia Weymer, 1910  TL: Brazil (MG)  M S

Amphidecta reynoldsi Sharpe, 1890    iN

Amphidecta reynoldsi reynoldsi Sharpe, 1890  TL: Brazil (Go)  M T S

Amphidecta reynoldsi [n. ssp.#1] Lamas, MS  TL: Brazil (MG/SC)  M T

Amphidecta reynoldsi [n. ssp.#2] Lamas, MS  TL: Colombia

Genera-unknown1  , []  not assigned to a genus yet

Genera-unknown1 insignis (A. Butler, 1867)  TL: [Brazil (Am)]  M T

Genera-unknown1 ordinata (Weymer, 1911)    iN  M T

  Tribe Satyrini, Subtribe Coenonymphina

Oressinoma  E. Doubleday, [1849]  

Oressinoma sorata Salvin & Godman, 1868    iN

Oressinoma sorata sorata Salvin & Godman, 1868  TL: Bolivia  M T S

Oressinoma sorata [n. ssp.] Lamas, MS  Peru

Oressinoma sorina Florczyk, Fåhraeus & Pyrcz, 2021  TL: Peru

Oressinoma typhla E. Doubleday, [1849]    iN

Oressinoma typhla Northern segregate ,  Costa Rica & Panama  M S L

Oressinoma typhla typhla E. Doubleday, [1849]  TL: Colombia  M T

Oressinoma typhla boliviana Forster, 1964  TL: Bolivia  M T

Oressinoma typhla [n. ssp.#1] Lamas, MS  TL: Costa Rica

Oressinoma typhla [n. ssp.#2] Lamas, MS  TL: Peru

Oressinoma typhla [n. ssp.#3] Lamas, MS  TL: Peru

Coenonympha  Hübner, [1819]  

Coenonympha haydenii (W. H. Edwards, 1872)  Hayden's Ringlet    iN  M S L1 L2 Ha

Coenonympha tullia (Müller, 1764)  Large Heath  ref.11  iN

Coenonympha tullia mixturata Alpheraky, 1897  this name may only apply to Siberian populations: existence of mixturata in Ameri ...  M S

Coenonympha tullia yukonensis W. Holland, 1900  USA (Ak) & Can (YT); ref.11  M T S1 S2 L

Coenonympha tullia kodiak (W. H. Edwards, 1869)  USA (Ak); ref.11  M S1 S2 IL

Coenonympha california Westwood, 1851  Common Ringlet  ref.11  iN

Coenonympha california nipisiquit McDunnough, 1939  salt marshes in NB & QC (Canada). Layberry et al. (1998), Webster (1999), and Sei ...  M T S

Coenonympha california mcisaaci dos Passos, 1935  TL: Canada (NL)  M T S

Coenonympha california inornata W. H. Edwards, 1861  TL: Canada (MB)  M T S L Ha

Coenonympha california benjamini McDunnough, 1928  TL: Canada (AB)  M S L Ha

Coenonympha california mackenziei D. Davenport, 1936  TL: Canada (NT)  M T S

Coenonympha california insulana McDunnough, 1928  TL: Canada (BC)  M S L

Coenonympha california yontocket Porter & Mattoon, 1989  TL: USA (Ca)  M T1 T2 S

Coenonympha california eryngii H. Edwards, 1877  TL: USA (Ca)  M T S L Ha

Coenonympha california california Westwood, 1851  TL: USA (Ca)  M T S L1 L2 I Ha

Coenonympha california columbiana McDunnough, 1928  TL: Canada (BC)  M S L

Coenonympha california eunomia (Dornfeld, 1967)  TL: USA (Or)  M T S L Ha

Coenonympha california ampelos W. H. Edwards, 1871  TL: USA (Ca/Or)  M T S1 S2 L I Ha

Coenonympha california mono Burdick, 1942  TL: USA (Ca)  M S L

Coenonympha california elko W. H. Edwards, 1881  TL: USA (Nv)  M S1 S2

Coenonympha california ochracea W. H. Edwards, 1861  TL: USA ([Co])  M T S L Ha

Coenonympha california pseudobrenda Austin & R. Gray, 1998  TL: USA (Nv)  M S1 S2 L

Coenonympha california furcae W. Barnes & Benjamin, 1926  TL: USA (Az)  M T S

Coenonympha california subfusca W. Barnes & Benjamin, 1926  TL: USA (Az)  M T S L Ha

  Tribe Satyrini, Subtribe Erebiina

Cercyonis  Scudder, 1875  

Cercyonis pegala (Fabricius, 1775)  Common Wood-Nymph    iN

Cercyonis pegala pegala (Fabricius, 1775)  TL: [USA (SC)]  M T S L1 L2

Cercyonis pegala alope (Fabricius, 1793)  (= maritima, = ochracea, = agawamensis); ref.141  M S L

Cercyonis pegala texana (W. H. Edwards, 1880)  TL: USA (Tx)  M T S L I

Cercyonis pegala Coahuila, low-elevation segregate ,   M S L Ha

Cercyonis pegala nephele (W. Kirby, 1837)  (= olympus, = borealis); ref.141  M S1 S2 S3 S4 L1 L2 I Ha

Cercyonis pegala ino G. Hall, 1924  TL: Canada (Ab)  M S

Cercyonis pegala incana (W. H. Edwards, 1880)  TL: USA (Wa)  M S

Cercyonis pegala boopis (Behr, 1864)  TL: USA (Ca)  M T S L

Cercyonis pegala ariane (Boisduval, 1852)  TL: USA (Ca)  M S L1 L2 Ha

Cercyonis pegala australosierra J. Emmel, T. Emmel & K. Davenport, 2008  TL: USA (Ca)  M T I

Cercyonis pegala Little Deschutes River segregate ,  Little Deschutes River & upper Deschutes R., Deschutes Co., OR  M S1 S2 L Ha

Cercyonis pegala Silver Lake segregate ,  vic. Silver Lake, Lake Co., OR  M S L1 L2 L3 Ha

Cercyonis pegala Crump Lake segregate ,  vic. Crump Lake, Lake Co., OR  M S1 S2

Cercyonis pegala Ravendale segregate ,   M S

Cercyonis pegala gabbii (W. H. Edwards, 1870)  TL: USA (Or)  M S L

Cercyonis pegala carsonensis Austin, 1992  TL: USA (Nv)  M T1 T2

Cercyonis pegala walkerensis Austin, 1992  TL: USA (Ca)  M T1 T2 S

Cercyonis pegala wheeleri (W. H. Edwards, 1873)  TL: USA (Ca)  M T

Cercyonis pegala paucilineatus Austin, 1992  TL: USA (Nv)  M T1 T2

Cercyonis pegala pluvialis Austin, 1992  TL: USA (Nv)  M T L

Cercyonis pegala paludum Austin, 1992  TL: USA (Nv)  M S L I Ha

Cercyonis pegala stephensi (W. G. Wright, 1905)  TL: USA (Ca)  M T S1 S2 L Ha

Cercyonis pegala utahensis Austin, 1992  TL: USA (Ut)  M T1 T2 S1 S2

Cercyonis meadii (W. H. Edwards, 1872)  Mead’s Wood-Nymph    iN

Cercyonis meadii meadii (W. H. Edwards, 1872)  TL: USA (Co)  M T S L1 L2 Ha

Cercyonis meadii alamosa T. Emmel & J. Emmel, 1969  TL: USA (Co)  M S L1 L2 I H Ha

Cercyonis meadii melania (Wind, 1946)  Requires confirmation in NE Chihuahua  M T S L

Cercyonis meadii mexicana (R. Chermock, 1949)  SE AZ, Sonora, W Chihuahua, etc.; ref.20  M T S

Cercyonis sthenele (Boisduval, 1852)  Great Basin Wood-Nymph    iN

Cercyonis sthenele masoni Cross, 1937  TL: USA (Co)  M S L

Cercyonis sthenele damei W. Barnes & Benjamin, 1926  Grand Canyon area; ref.20

Cercyonis sthenele paulus (W. H. Edwards, 1879)  TL: USA (Nv)  M S L1 L2

Cercyonis sthenele W OR Cascades segregate ,   M S

Cercyonis sthenele sineocellata Austin & J. Emmel, 1998  TL: USA (Or)  M T S L1 L2 Ha

Cercyonis sthenele behrii (F. Grinnell, 1905)  TL: USA (Ca)  M S L

Cercyonis sthenele sthenele (Boisduval, 1852)  TL: USA (Ca)  M T1 T2 TI S IL

Cercyonis hypoleuca Hawks & J. Emmel, 1998  Santa Cruz Is., CA, USA; ref.20

Cercyonis silvestris (W. H. Edwards, 1861)  Great Basin Wood-Nymph  LT not coonspecific with sthenele; ref.16

Cercyonis silvestris incognita J. Emmel, T. Emmel & Mattoon, 2012  N CA; ref.16  iN  M S L

Cercyonis silvestris silvestris (W. H. Edwards, 1861)  ref.16  M T S1 S2 L Ha

Cercyonis oetus (Boisduval, 1869)  Small Wood-Nymph    iN

Cercyonis oetus charon (W. H. Edwards, 1872)  TL: USA (Co)  M T S L1 L2 L3 L4 Ha

Cercyonis oetus phocus (W. H. Edwards, 1874)  TL: Canada (BC)  M T S

Cercyonis oetus oetus (Boisduval, 1869)  TL: USA (Ca)  M T S L

Cercyonis oetus pallescens T. Emmel & J. Emmel, 1971  TL: USA (Nv)  M S Ha

Cercyonis oetus alkalorum Austin, 1998  TL: USA (Nv)  M T1 T2

Erebia  Dalman, 1816  

Erebia magdalena Strecker, 1880  Magdalena Alpine    iN

Erebia magdalena hilchie Kemal & Koçak, 2007  N Rocky Mountains; W-central ALTA; E-central BC  M T S

Erebia magdalena magdalena Strecker, 1880  S Rocky Mountains; S-central MT, NW WY, N UT, CO, N-central NM  M T S L I Ha

Erebia mackinleyensis Gunder, 1932  Mt. McKinley Alpine  N, C & E AK, W YT, N BC   iN  M S L Ha

Erebia fasciata A. Butler, 1868  Banded Alpine    iN

Erebia fasciata avinoffi W. Holland, 1930  NW AK & E Russia  M S

Erebia fasciata fasciata A. Butler, 1868  Nunavut to C AK  M S L Ha G

Erebia discoidalis (W. Kirby, 1837)  Red-disked Alpine  Canada & NC USA (N MT, N ND, N MN, N WI )  iN  M S G

Erebia rossii (J. Curtis, 1835)  Ross's Alpine    iN

Erebia rossii kuskoquima W. Holland, 1931  coastal SW AK (Yukon & Kuskoquim River drainages)  M S

Erebia rossii gabrieli dos Passos, 1949  vic. Denali National Park, AK  M S L Ha

Erebia rossii rossii (J. Curtis, 1835)  most of range  M S G

Erebia rossii ornata Leussler, 1935  vic. Churchill, Manitoba  M T S

Erebia disa (Thunberg, 1791)  Disa Alpine    iN

Erebia disa steckeri W. Holland, 1930  N AK to W YK; tundra and forest habitats  M S L G

Erebia disa subarctica McDunnough, 1937  Richardson Mts. & MacKenzie Delta E to Nunavut  M S

Erebia mancinus E. Doubleday, [1849]  Taiga Alpine  taiga; Labrador to Alaska; S in Mts. To Banff, ALTA  iN  M S

Erebia lafontainei Troubridge & Philip, 1983  Lafontaine's Alpine  tundra AK, E YT & NW NT  iN  M OI S IL L Ha G Txt

Erebia youngi W. Holland, 1900  Young's Alpine    iN

Erebia youngi rileyi dos Passos, 1947  mostly tundra, W AK coast & vic. Denali National Park, AK  M S IL G Txt

Erebia youngi youngi W. Holland, 1900  mostly tundra in C AK to W YK  M S IL L Ha G Txt

Erebia youngi herscheli Leussler, 1935  mostly tundra, coastal N AK & N YK  M T S IL G Txt

Erebia occulta Roos & Kimmich, 1983  Eskimo Alpine    iN

Erebia occulta occulta Roos & Kimmich, 1983  small scree areas in AK & NW YT  M OI S IL L G Txt

Erebia pawloskii Ménétriés, 1859  Theano Alpine    iN

Erebia pawloskii alaskensis W. Holland, 1900  AK, YK, N BC  M S L Ha

Erebia pawloskii canadensis B. Warren, 1931  vic. Churchill, Manitoba  M S

Erebia pawloskii ethela W. H. Edwards, 1891  higher elevations in SC MT, NW WY & NC CO  M S L Ha G

Erebia pawloskii demmia B. Warren, 1936  San Juan Range in SW CO  M S L

Erebia epipsodea A. Butler, 1868  Common Alpine    iN

Erebia epipsodea remingtoni P. Ehrlich, 1952  SE AK, YT & NW BC  M S L

Erebia epipsodea sineocellata Skinner, 1889  NE BC, AB (except extreme SW), S SK, SW MB, N ID & N MT  M S G

Erebia epipsodea epipsodea A. Butler, 1868  S BC, SW AB, NW USA  M S L1 L2 Ha G

Erebia epipsodea brucei Elwes, 1889  moist meadows in S Rocky Mountains; SE WY, E UT, CO & N NM  M T S L Ha

Erebia vidleri Elwes, 1898  Vidler's Alpine  W BC & NW WA  iN  M OD S L Ha G

Erebia callias W. H. Edwards, 1871  Colorado Alpine    iN

Erebia callias callias W. H. Edwards, 1871  alpine meadows of SE MT, W WY, NE UT & C CO  M S L1 L2 Ha

  Tribe Satyrini, Subtribe Satyrina

Oeneis  Hübner, [1819]  

Oeneis ridingsii (W. H. Edwards, 1865)  Ridings' Satyr  ref.10  iN

Oeneis ridingsii minimus (Austin, 1986)  S ALTA E to SW Manitoba, N MT, W ND  M T S

Oeneis ridingsii wyomingo (Scott, 1998)  N CO, WY, MT, UT; flies in late summer  M S1 S2

Oeneis ridingsii ridingsii (W. H. Edwards, 1865)  C MT to N MN, W NE  M S1 S2 L1 L2 Ha

Oeneis ridingsii curicata (M. Fisher, Scott & Garhart, 2008)  upper Gunnison River Valley, Gunnison Co., CO  M T L Ha

Oeneis ridingsii coloalbiterra (Garhart & M. Fisher, 2008)  Roan Cliffs, Garfield Co., CO  M T

Oeneis ridingsii neomexicanus (Austin, 1986)  S-central AZ & NM  M T S L Ha

Oeneis ridingsii stretchii (W. H. Edwards, 1870)  W WY to W CO, W to SE OR, NV, N AZ  M S L Ha

Oeneis ridingsii pallidus (Austin, 1986)  Sierra Nevada (CA-NV), N to C Oregon  M S L1 L2 H

Oeneis carmen (A. Warren, Austin, Llorente, Luis & Vargas, 2008)  Joboni Satyr  northern Coah to southern NL; ref.108,10  iN  M T S L H

Oeneis pseudochazaroides (Durden, 2008)  TL: Mexico

Oeneis jutta (Hübner, [1806])  Jutta Arctic    iN

Oeneis jutta terraenovae dos Passos, 1935  Newfoundland  M T S

Oeneis jutta balder (Guérin-Méneville, [1832])  SE Canada & NE US W to SE Manitoba  M T OI S1 S2

Oeneis jutta ascerta Masters & Sorensen, 1968  N WI, MN, MI, adjacent parts of Ontario  M T S L

Oeneis jutta ridingiana F. Chermock & R. Chermock, 1940  SW Manitoba & SASK  M S

Oeneis jutta harperi dos Passos, [1977]  N Manitoba & E NWT  M T S

Oeneis jutta leussleri Bryant, 1935  W NWT  M T S

Oeneis jutta alaskensis W. Holland, 1900  AK, E to YK & N BC  M S1 S2 L I Ha

Oeneis jutta chermocki Wyatt, 1965  W ALTA, S BC  M S

Oeneis jutta reducta McDunnough, 1929  S Rocky Mountains (far E ID, W MT, WY, NE UT, NW CO)  M S L I1 I2 Ha

Oeneis melissa (Fabricius, 1775)  Melissa Arctic    iN

Oeneis melissa semidea (Say, 1828)  White Mountains, New Hampshire  M T S

Oeneis melissa melissa (Fabricius, 1775)  coastal Newfoundland & Labrador  M T S

Oeneis melissa semplei W. Holland, 1930  interior Labrador, N Quebec, margins of Hudson Bay  M S

Oeneis melissa assimilis A. Butler, 1868  Nunavut & NWT  M S

Oeneis melissa gibsoni W. Holland, 1930  W & N YK, AK  M S L

Oeneis melissa atlinensis C. Guppy & J. Shepard, 2001  NW BC, SW YK  M S

Oeneis melissa beanii Elwes & J. Edwards, 1893  S & C BC, SW & W-central ALTA, to N-central WA, MT, WY & NE UT  M S L Ha

Oeneis melissa lucilla W. Barnes & McDunnough, 1918  S Rocky Mountains (CO, N NM)  M S L1 L2 Ha

Oeneis polixenes (Fabricius, 1775)  Polixenes Arctic    iN

Oeneis polixenes katahdin (Newcomb, 1901)  Mt. Katahdin, Maine  M T S

Oeneis polixenes polixenes (Fabricius, 1775)  E Canadian arctic; Quebec to Labrador  M S

Oeneis polixenes subhyalina (J. Curtis, 1835)  arctic islands and coasts; Nunavut & NWT  M S

Oeneis polixenes beringianus Kurentsov, 1970  Churchill, Manitoba, W to AK, S to S ALTA; E Russia  M T S1 S2 L

Oeneis polixenes woodi Troubridge & Parshall, 1988  arctic coast of AK & YK  M S

Oeneis polixenes yukonensis A. Gibson, 1920  SW YK  M S

Oeneis polixenes brucei (W. H. Edwards, 1891)  S Rocky Mountains (NW WY, C & N CO, N NM)  M S L Ha

Oeneis philipi Troubridge, 1988  Philip's Arctic  C AK, YK, N BC  iN  M S

Oeneis alpina Kurentsov, 1970  Sentinel Arctic    iN

Oeneis alpina excubitor Troubridge, Philip, Scott & J. Shepard, 1982  far NW NWT, YK, N AK  M S G

Oeneis bore (Esper, 1789)  White-veined Arctic    iN

Oeneis bore taygete Geyer, [1830]  Labrador & N Quebec  M T S

Oeneis bore gaspeensis dos Passos, 1949  montane Gaspé Peninsula, Quebec  M S

Oeneis bore hanburyi Watkins, 1928  N-central Canadian arctic, from NE Manitoba to YK & E-central AK  M S L Ha

Oeneis bore mckinleyensis dos Passos, 1949  vic. Denali National Park, AK  M S L Ha

Oeneis bore fordi dos Passos, 1949  W AK, Russian Far East  M S

Oeneis bore edwardsi dos Passos, 1949  S Rocky Mountains (S-central MT, NW WY, NE UT, SW CO)  M S L Ha

Oeneis alberta Elwes, 1893  Alberta Arctic    iN

Oeneis alberta alberta Elwes, 1893  Canada, N-central US  M T S

Oeneis alberta ojibwe Scott & Kondla, 2006  SE Manitoba  M S

Oeneis alberta oslari Skinner, 1911  CO  M S1 S2 L Ha

Oeneis alberta capulinensis F. Brown, 1970  NE NM & SE CO  M S1 S2

Oeneis alberta daura (Strecker, 1894)  AZ: Graham & Coconino Cos.  M S L1 L2 Ha

Oeneis tanana A. Warren & Nakahara, 2016    iN  M S

Oeneis chryxus (E. Doubleday, [1849])  Chryxus Arctic    iN

Oeneis chryxus strigulosa McDunnough, 1934  N WI, N MI, S & E Ontario, SW Quebec  M S L

Oeneis chryxus calais (Scudder, 1865)  W Ontario, W to SASK  M S

Oeneis chryxus caryi Dyar, 1904  boreal ALTA, BC, NWT, YK, far E AK  M T S

Oeneis chryxus altacordillera Scott, 2006  high Canadian Zone in Rocky Mts. of Colorado  M S1 S2 L Ha

Oeneis chryxus chryxus (E. Doubleday, [1849])  Rocky Mountains from ALTA to NM  M OI S1 S2 Ab HS L Ha

Oeneis chryxus socorro R. Holland, 2010  NM: Socorro Co., Mt. Withington  M OI

Oeneis chryxus valerata Burdick, 1958  Olympic Mts., NW WA & Vancouver I.  M T S L Ha

Oeneis chryxus stanislaus Hovanitz, 1937  Sierra Nevada, CA-NV  M T S L1 L2 Ha

Oeneis chryxus ivallda (Mead, 1878)  Sierra Nevada, CA  M S L

Oeneis nevadensis (C. Felder & R. Felder, 1867)  Great Arctic    iN

Oeneis nevadensis gigas A. Butler, 1868  BC to NW OR (coastal)  M S L

Oeneis nevadensis nevadensis (C. Felder & R. Felder, 1867)  BC to N CA (inland)  M T S L I Ha

Oeneis nevadensis iduna (W. H. Edwards, 1874)  CA: Mendocino-Sonoma Cos.  M S

Oeneis macounii (W. H. Edwards, 1885)  Macoun's Arctic    iN  M S

Oeneis uhleri (Reakirt, 1866)  Uhler's Arctic    iN

Oeneis uhleri cairnesi A. Gibson, 1920  NE AK, YK, far NW NWT  M T OI OD S L Ha G

Oeneis uhleri nahanni Dyar, 1904  Mackenzie Mountains, NWT  M T

Oeneis uhleri varuna (W. H. Edwards, 1882)  S-central Canada and N-central US  M S L G

Oeneis uhleri reinthali F. Brown, 1953  CO: W slopes  M T S L1 L2 Ha G

Oeneis uhleri uhleri (Reakirt, 1866)  CO: E slopes, E WY  M S1 S2 L Ha G

Oeneis uhleri SW Utah segregate ,  Tushar Mts. of Beaver-Piute Cos., Utah  M S1 S2

  Subfamily Nymphalidae-incertae-sedis

Apanthesis  , []  a fossil genus

Apanthesis leuce Scudder, 1889  Fossil from Colorado

Lithodryas  , []  a fossil genus

Lithodryas styx (Scudder, 1889)  Fossil from Colorado

Nymphalites  , []  a fossil genus

Nymphalites obscurum Scudder, 1889  Fossil from Colorado

Nymphalites scudderi Beutenmüller & T. Cockerell, 1908  Fossil from Colorado

Family RIODINIDAE Grote, 1895 (1827) – METALMARKS

  Subfamily Euselasiinae

  Tribe Nemeobiini

Styx  Staudinger, 1876  

Styx infernalis Staudinger, 1876    iN  M T S

Corrachia  Schaus, 1913  

Corrachia leucoplaga Schaus, 1913  Costa Rican Metalmark  Costa Rica  iN  M T S

  Tribe Euselasiini

Methone  E. Doubleday, 1847  

Methone cecilia (Cramer, 1777)  ref.12  iN

Methone cecilia caduca Stichel, 1919  Costa Rica  M T S

Methone cecilia chrysomela (A. Butler, 1872)  Costa Rica to Colombia  M T S

Methone cecilia columbana Stichel, 1926  TL: Colombia  M T

Methone cecilia eurotias Stichel, 1919  TL: Ecuador  M T S

Methone cecilia cecilia (Cramer, 1777)  TL: Bengala  M T S

Methone cecilia magnarea (Seitz, 1913)  TL: Brazil (Am); Peru; Bolivia  M T

Methone hecamede (Hewitson, 1870)  ref.12  iN

Methone hecamede hecamede (Hewitson, 1870)  TL: Ecuador  M T S

Methone hecamede avicula (Stichel, 1909)  TL: Ecuador  M T

Methone hecamede haematites (Stichel, 1909)  TL: ?  M T

Methone hecamede hecate (Stichel, 1919)  TL: Colombia  M T

Methone noctula (Westwood, 1851)  White-rayed Metalmark  SE Mexico to Colombia and Venezuela; ref.12  iN  M T S1 S2 L I1 I2

Methone portentosa (Stichel, 1927)  Black Euselasia  Costa Rica  M T

Methone eucrates (Hewitson, 1872)    iN

Methone eucrates leucorrhoa (Godman & Salvin, 1878)  SE Mexico to Venezuela  M T S L

Methone eucrates eucrates (Hewitson, 1872)  TL: Ecuador  M T S

Methone eucrates eriphyla (Stichel, 1919)  TL: Ecuador  M T

Methone orion (Le Cerf, 1958)  TL: Colombia  M OI OD

Methone tenebrosa (Constantino, G. Rodríguez & Salazar, 2011)    iN  M T

Methone opimia (Stichel, 1919)  TL: Bolivia  M T S

Methone baucis (Stichel, 1919)    iN

Methone baucis baucis (Stichel, 1919)  TL: Peru  M T S

Methone baucis aethiops (Rebillard, 1958)  TL: Colombia  M OI

Methone pontasis (Callaghan, Llorente & Luis, 2007)  Pointed Euselasia  SE Mexico (VER, CHIS)  M T S

Methone hypophaea (Godman & Salvin, 1878)  Panamanian Euselasia  S Mexico to Panama  M T S

Methone dolichos (Staudinger, [1887])  TL: Peru  M T S

Methone euploea (Hewitson, [1855])    iN  M T S HS V

Methone eucerus (Hewitson, 1872)    iN  M T S

Methone eubule (R. Felder, 1869)  Dusky Euselasia    iN

Methone eubule W Mexican segregate (, )  W Mexico  M S

Methone eubule eubule (R. Felder, 1869)  [= mexicana], E Mexico to Panama  M T S1 S2 L I

Methone onorata (Hewitson, 1869)  Whitewashed Euselasia  Costa Rica to Ecuador  iN  M T TI S

Methone authe (Godman, 1903)    iN

Methone authe authe (Godman, 1903)  TL: Brazil (MT)  M T

Methone authe copiosa (Stichel, 1919)  TL: Peru; Bolivia  M T

Methone authe ocalea (H. Druce, 1904)  TL: Bolivia  M T S

Eurylasia  Grishin, 2021  

Eurylasia effima (Hewitson, 1869)    iN  M T S

Eurylasia euryone (Hewitson, 1856)    iN  M T S

Eurylasia thusnelda (Möschler, 1883)    iN  M T

Eurylasia eunaeus (Hewitson, [1855])  TL: Brazil (Am)  M T S

Eurylasia inini (Brévignon, 1996)  TL: French Guiana

Myselasia  Grishin, 2021  

Myselasia eulione (Hewitson, 1856)    iN  M T

Myselasia ella (Seitz, 1916)    iN

Myselasia ella terrea (Stichel, 1924)  TL: Brazil (Pa)  M T

Myselasia ella ella (Seitz, 1916)  TL: Bolivia  M T S

Myselasia mapatayna (J. Hall & Willmott, 1998)  TL: Ecuador  M T OI

Myselasia nauca (J. Hall & Willmott, 1998)  TL: Ecuador  M T OI

Myselasia hygenius (Stoll, 1790)    iN

Myselasia hygenius hygenius (Stoll, 1790)  TL: Surinam  M OI S

Myselasia hygenius occulta (Stichel, 1919)  TL: Brazil (RS)  M T

Myselasia eustola (Stichel, 1919)    iN

Myselasia eustola eustola (Stichel, 1919)  TL: Bolivia; Peru  M T S

Myselasia eustola wacapu (Brévignon, 1996)  TL: French Guiana

Myselasia cafusa (H. Bates, 1868)    iN  M T S

Myselasia janigena (Stichel, 1919)    iN  M T S

Myselasia alcmena (H. Druce, 1878)    iN  M T S V

Myselasia cyanofusa (J. Hall & Willmott, 1998)    iN  M T OI S

Myselasia gloori (Choimet, 2011)  TL: Costa Rica

Myselasia pullata (Stichel, 1927)  TL: Brazil (Pa)  M T S

Myselasia licinia (Godman, 1903)  TL: Guyana  M T

Myselasia mys (Herrich-Schäffer, [1853])  Variable Euselasia    iN

Myselasia mys mys (Herrich-Schäffer, [1853])  Belize to Amazonas, Guianas  M T S V

Myselasia mys cytis (Stichel, 1919)  TL: Brazil (MT)  M T S

Myselasia mys lara (Stichel, 1919)  TL: Brazil (ES)  M T S

Myselasia rhodon (Seitz, 1913)  TL: [Brazil (Pa)]  M T S

Myselasia pseudomys (Callaghan, 1999)    iN  M T OI S

Myselasia saulina (Brévignon, 1996)  TL: French Guiana

Myselasia crinon (Stichel, 1919)    iN  M T S

Myselasia marica (Stichel, 1919)    iN  M T S

Myselasia eberti (Callaghan, 1999)  TL: Brazil (SP)  M T OI

Myselasia athena (Hewitson, 1869)  TL: Ecuador  M T S

Myselasia procula (Godman & Salvin, 1885)  Orange-costa Euselasia  SE Mexico to Colombia  iN  M T S

Myselasia hieronymi (Salvin & Godman, 1868)  Red-rayed Euselasia    iN

Myselasia hieronymi SW Mexican segregate (, )  SW Mexico  M S

Myselasia hieronymi hieronymi (Salvin & Godman, 1868)  E Mexico to Nicaragua  M T S L Ha

Myselasia hieronymi bianala (J. Hall & Willmott, 1998)  Costa Rica to W Ecuador  M T OI S L

Myselasia inconspicua (Godman & Salvin, 1878)  Inconspicuous Euselasia  SE Mexico to Costa Rica and Panama  iN  M T S

Myselasia cucuta (Schaus, 1902)    iN  M T

Myselasia gradata (Stichel, 1927)  TL: Venezuela

Myselasia gradata gonzaloi (Salazar & Henao, 2019)  TL: Colombia  M OI S

Myselasia gradata gradata (Stichel, 1927)  TL: Venezuela  M T

Myselasia jigginsi (J. Hall & Willmott, 1998)  TL: Ecuador  M T OI S

Myselasia sergia (Godman & Salvin, 1885)  Sergia Euselasia  SE Mexico (VER, CHIS) to Panama  iN  M T S

Myselasia mystica (Schaus, 1913)  Mystical Euselasia  Nicaragua to Panama  iN

Myselasia mystica mystica (Schaus, 1913)  Nicaragua to Panama  M T S1 S2 L I1 I2

Myselasia mystica caucana (Constantino, Salazar & G. Rodríguez, 2019)  TL: Colombia  M OI

Myselasia mystica pseudomazaca (Salazar & J. Vargas, 2019)  TL: Colombia  M OI

Myselasia pance (Callaghan, 1999)    iN  M T OI

Myselasia leucon (Schaus, 1913)  Rusty-patched Euselasia  SE Mexico (OAX) to Panama  iN  M T S

Myselasia pusilla (R. Felder, 1869)  Pearly Euselasia    iN

Myselasia pusilla pusilla (R. Felder, 1869)  E Mexico to El Salvador  M T S L

Myselasia pusilla mazai (Beutelspacher, 1975)  Chiapas  M T S L

Myselasia matuta (Schaus, 1913)  White-ventered Euselasia  Costa Rica  M T S

Myselasia julia (H. Druce, 1878)  Amazonian  iN  M T S

Erythia  Hübner, [1819]  

Erythia teleclus (Stoll, 1787)    iN

Erythia teleclus maculata (Constantino & Salazar, 2019)  TL: Colombia  M OI

Erythia teleclus teleclus (Stoll, 1787)  TL: Surinam  M T OI S

Erythia archelaus (Seitz, 1916)    iN

Erythia archelaus archelaus (Seitz, 1916)  TL: Bolivia  M T

Erythia archelaus ecuadorensis (Lathy, 1926)  TL: Ecuador  M T

Erythia catoleuce Hübner, 1823  TL: America  M OI S HS V

Erythia thucydides (Fabricius, 1793)    iN

Erythia thucydides thucydides (Fabricius, 1793)  TL: Indiis  M T S

Erythia thucydides truncata (Callaghan, 2001)  TL: Brazil (DF)  M T

Erythia labdacus (Stoll, 1780)    iN

Erythia labdacus reducta (Lathy, 1926)  Costa Rica to Colombia  M T S

Erythia labdacus labdacus (Stoll, 1780)  TL: Surinam  M T OI

Erythia midas (Fabricius, 1775)    iN

Erythia midas midas (Fabricius, 1775)  TL: Indiis  M OI IL

Erythia midas ater (Seitz, 1913)  TL: [Colombia; Bolivia]  M T S

Erythia midas crotopiades (Stichel, 1919)  Costa Rica to Colombia  M T S L

Erythia midas crotopina (Seitz, 1916)  TL: [Peru]  M T S

Erythia midas erosina (Brévignon, 1995)  TL: French Guiana  M S

Erythia midas pelusia (Stichel, 1919)  TL: Peru  M T S

Erythia kartopus (Stichel, 1919)    iN  M T S

Erythia euhemerus (Hewitson, 1856)  TL: Brazil (Pa)  M T S

Erythia mutator (Seitz, 1916)  TL: [Peru]

Erythia mutator huilensis (G. Rodríguez & Salazar, 2019)  TL: Colombia  M OI

Erythia mutator mutator (Seitz, 1916)  TL: [Peru]  M T S

Erythia micaela (Schaus, 1902)  TL: Peru  M T

Erythia venezolana (Seitz, 1913)    iN

Erythia venezolana venezolana (Seitz, 1913)  TL: Venezuela  M T

Erythia venezolana jauffreti (Gallard, 2014)  TL: Brazil  M OI

Erythia psammathe (Seitz, 1916)  TL: [French Guiana]

Erythia psammathe hypocala (Le Cerf, 1958)  TL: Colombia  M T S

Erythia psammathe psammathe (Seitz, 1916)  TL: [French Guiana]  M T

Erythia gordios (Stichel, 1919)    iN  M T S

Erythia rhodogyne (Godman, 1903)    iN

Erythia rhodogyne rhodogyne (Godman, 1903)  TL: Colombia  M T S

Erythia rhodogyne patella (Stichel, 1927)  Costa Rica and Panama  M T S1 S2 I

Erythia fayneli (Gallard, 2006)  French Guiana; ref.154  M T

Erythia aurantiaca (Salvin & Godman, 1868)  Fiery Euselasia    iN

Erythia aurantiaca aurum (Callaghan, Llorente & Luis, 2007)  W Mexico  M T S

Erythia aurantiaca aurantiaca (Salvin & Godman, 1868)  E Mexico to Panama  M T S L

Erythia aurantiaca [n. ssp.] (Callaghan & Lamas, MS)  Colombia  M T

Erythia aurantiaca marginata (Lathy, 1926)  TL: Venezuela  M T

Erythia borrosa Grishin, 2023  Panama; ref.151

Erythia cheles (Godman & Salvin, 1889)  Cheles Euselasia  Costa Rica and Panama  iN  M T S1 S2 S3 I

Erythia paracheles Grishin, 2023  Panama; ref.151

Erythia palla (J. Hall & Willmott, 1998)  TL: Ecuador  M T OI

Pelolasia  Grishin, 2021  

Pelolasia mazaca (Hewitson, 1860)    iN  M T S

Pelolasia aurantia (A. Butler & H. Druce, 1872)  Bright-centered Euselasia  Costa Rica to Colombia  iN  M T S L

Pelolasia chrysippe (H. Bates, 1866)  Golden Euselasia  SE Mexico to Colombia  iN  M T S1 S2 L I

Pelolasia candaria (H. Druce, 1904)    iN  M T S

Pelolasia bettina (Hewitson, 1869)  Black-topped Euselasia  Nicaragua to Ecuador  iN  M T S L

Pelolasia cataleuca (R. Felder, 1869)  Black-edged Euselasia  SE Mexico, Belize, Honduras  iN  M T S L

Pelolasia leucophryna (Schaus, 1913)  Glowing Euselasia  Honduras, Costa Rica; ref.151  M T S

Pelolasia eusepus (Hewitson, [1853])  Eusepus Euselasia  Panama to S Brazil  iN  M T S L

Pelolasia eumenes (Hewitson, [1853])    iN  M T S

Pelolasia eupatra (Seitz, 1916)    iN  M T S

Pelolasia artos (Herrich-Schäffer, [1853])  TL: Surinam  M T S

Pelolasia ignitus (Stichel, 1924)    iN  M T S

Pelolasia rubrocilia (Lathy, 1926)  TL: French Guiana  M T

Pelolasia mirania (H. Bates, 1868)    iN  M T S

Pelolasia nytua (J. Hall & Willmott, 2009)  ref.225  M OI

Pelolasia eumedia (Hewitson, [1853])    iN

Pelolasia eumedia eumedia (Hewitson, [1853])  TL: Brazil (Pa)  M T S

Pelolasia eumedia ophias (Herrich-Schäffer, [1853])  TL: Surinam  M T S

Pelolasia pelor (Hewitson, [1853])    iN  M T S

Pelolasia pellonia (Stichel, 1919)  Red-rimmed Euselasia    iN

Pelolasia pellonia pellonia (Stichel, 1919)  Costa Rica, Peru  M T S I

Pelolasia pellonia azurea (Callaghan, 1999)  TL: Brazil (RJ)  M T OI

Pelolasia euboea (Hewitson, [1853])    iN

Pelolasia euboea euboea (Hewitson, [1853])  TL: Brazil (Pa)  M T S

Pelolasia euboea pasoula (Callaghan & Lamas, 2004)  TL: French Guiana  M S

Pelolasia seitzi (Lathy, 1926)  TL: Peru  M T S

Pelolasia amphidecta (Godman & Salvin, 1878)  Glossed Euselasia  Costa Rica and Panama  iN  M T S1 S2 L

Pelolasia melaphaea (Hübner, 1823)    iN

Pelolasia melaphaea melaphaea (Hübner, 1823)  TL: Surinam  M T S

Pelolasia melaphaea condensa (Stichel, 1927)  TL: Bolivia  M T S

Pelolasia argentea (Hewitson, 1871)  Orange-spotted Euselasia  Chiapas to Panama  M T S

Pelolasia fervida (A. Butler, 1874)    iN

Pelolasia fervida fervida (A. Butler, 1874)  TL: Venezuela  M T S

Pelolasia fervida fervidina (Stichel, 1919)  TL: Brazil (ES)  M T

Pelolasia hahneli (Staudinger, [1887])    iN  M T S

Marmessus  Hübner, [1819]  

Marmessus gyda (Hewitson, 1860)    iN

Marmessus gyda gydina (Stichel, 1919)  Costa Rica and Panama  M T S

Marmessus gyda gyda (Hewitson, 1860)  TL: Brazil  M T

Marmessus gyda sticheli (G. Rodríguez & Salazar, 2019)  TL: Colombia  M OI

Marmessus opalescens (Hewitson, [1855])    iN

Marmessus opalescens nielseni (Salazar & J. Vargas, 2019)  TL: Colombia  M OI

Marmessus opalescens opalescens (Hewitson, [1855])  TL: Brazil  M T

Marmessus opalescens interrupta (Lathy, 1924)  TL: French Guiana  M T

Marmessus opalescens joiceyi (Lathy, 1924)  TL: Ecuador  M T S

Marmessus opalescens opigena (Stichel, 1919)  TL: Bolivia  M T S

Marmessus opalina (Westwood, 1851)  TL: Brazil (Am)  M T S

Marmessus praeclara (Hewitson, 1869)    iN  M T S

Marmessus violacea (Lathy, 1924)  TL: Colombia  M T S

Marmessus fournierae (Lathy, 1924)  TL: Brazil (Am)  M T S

Marmessus jocotoco (J. Hall & Willmott, 2009)  ref.225  M OI S

Marmessus gelanor (Stoll, 1780)    iN  M T OI S

Marmessus bilineata (Lathy, 1926)    iN  M T S

Marmessus erilis (Stichel, 1919)    iN  M T S

Marmessus murina (Stichel, 1925)    iN  M T V

Marmessus rava (Stichel, 1928)  TL: Bolivia

Marmessus rava albomarginata (Salazar & J. Vargas, 2019)  TL: Colombia  M OI

Marmessus rava rava (Stichel, 1928)  TL: Bolivia  M T

Marmessus amblypodia (Lathy, 1926)    iN

Marmessus amblypodia funebris (Salazar & Constantino, 2019)  TL: Colombia  M OI

Marmessus amblypodia amblypodia (Lathy, 1926)  TL: Peru  M T S

Marmessus parca (Stichel, 1919)  TL: Brazil (Am)  M T

Marmessus uria (Hewitson, [1853])    iN

Marmessus uria amapaena (J. Jauffret, 2011)  TL: Brazi

Marmessus uria uria (Hewitson, [1853])  TL: Brazil (Pa)  M T S

Marmessus uria angustifascia (Lathy, 1926)  TL: Brazil (MT)  M T S

Marmessus extensa (H. Bates, 1868)    iN  M T S

Marmessus eumithres (Stichel, 1919)  TL: Brazil (Am)  M T

Marmessus eubotes (Hewitson, 1856)    iN  M T S

Marmessus scotinosa (Stichel, 1930)    iN

Marmessus scotinosa scotinosa (Stichel, 1930)  TL: Brazil (Am)  M T S

Marmessus scotinosa matouryensis (Brévignon & Gallard, 1993)  TL: French Guiana

Marmessus misteriosa (Salazar & J. Vargas, 2019)  TL: Colombia  M OI S

Marmessus urites (Hewitson, [1853])    iN

Marmessus urites litoralis (Salazar & Constantino, 2019)  TL: Colombia  M OI

Marmessus urites urites (Hewitson, [1853])  TL: Brazil (Pa)  M T S

Marmessus urites eglawahe (Brévignon & Gallard, 1993)  TL: French Guiana  M S

Marmessus lycaeus (Staudinger, 1888)    iN  M T S

Marmessus lysimachus (Staudinger, 1888)    iN  M T S

Marmessus euromus (Hewitson, 1856)    iN  M T S

Marmessus lisias (Cramer, 1777)    iN  M T S

Marmessus dorina (Hewitson, 1860)    iN  M T S

Marmessus oaxacensis (Callaghan, Llorente & Luis, 2013)  Stub-tailed Euselasia  ["angulata"], SE Mexico to Brazil  iN  M T

Marmessus eurypus (Hewitson, 1856)    iN  M T S L

Marmessus angulata (H. Bates, 1868)    iN  M T S

Marmessus clesa (Hewitson, 1856)    iN  M T S

Marmessus descherrei (Gallard, 2014)  TL: French Guiana  M T OI

Marmessus utica (Hewitson, [1855])  TL: Brazil  M T S

Marmessus euphaes (Hewitson, [1855])    iN  M T S

Marmessus subargentea (Lathy, 1904)  Great Euselasia  Costa Rica to Colombia  M T S

Maculasia  Grishin, 2021  

Maculasia albomaculiga (Callaghan, 1999)    iN  M T OI S

Euselasia  Hübner, [1819]  

Euselasia andreae J. Hall, Willmott & R. Busby, 1998  TL: Ecuador  M T OI S

Euselasia illarina J. Hall, Willmott & R. Busby, 1998  TL: Ecuador  M T OI

Euselasia violetta (H. Bates, 1868)    iN  M T S

Euselasia fabia (Godman, 1903)    iN

Euselasia fabia fabia (Godman, 1903)  TL: Peru; Ecuador  M T S

Euselasia fabia marcida Stichel, 1919  TL: Ecuador  M T S

Euselasia arbas (Stoll, 1782)    iN  M T OI S

Euselasia attrita Seitz, 1916    iN  M T S

Euselasia rasonea (Schaus, 1902)    iN  M T S

Euselasia praecipua Stichel, 1924    iN  M T S

Euselasia euoras (Hewitson, [1855])  Purple-washed Euselasia  Costa Rica to Peru and Brazil  iN

Euselasia euoras awala Brévignon, 2008  TL: French Guiana  M T

Euselasia euoras euoras (Hewitson, [1855])  Costa Rica to Peru and Brazil  M T S L

Euselasia manoa Brévignon, 1996  TL: French Guiana  M T S

Euselasia waponaka Brévignon, 2008    iN

Euselasia waponaka uypiranga Brévignon, 2008  TL: [Brazil]

Euselasia waponaka waponaka Brévignon, 2008  TL: French Guiana  M T S

Euselasia respectus Brévignon, 2008  TL: French Guiana  M T

Euselasia rufomarginata J. Hall & Harvey, 2004  TL: Peru  M T OI

Euselasia michaeli J. Hall & Harvey, 2004    iN  M OI S

Euselasia tarinta (Schaus, 1902)  TL: Colombia  M T S

Euselasia regipennis (A. Butler & H. Druce, 1872)  Purple-topped Euselasia    iN

Euselasia regipennis regipennis (A. Butler & H. Druce, 1872)  SE Mexico to Panama  M T S L

Euselasia regipennis eupepla (Godman & Salvin, 1885)  Panama and Colombia  M T S

Euselasia regipennis eurysthenes (Hewitson, 1872)  TL: Ecuador  M T S

Euselasia eurymachus (Hewitson, 1872)    iN  M T OI

Euselasia corduena (Hewitson, 1874)    iN

Euselasia corduena corduena (Hewitson, 1874)  TL: Bolivia  M T S

Euselasia corduena anadema Stichel, 1927  Costa Rica to Colombia  M T S

Euselasia euriteus (Cramer, 1777)    iN  M T OI S

Euselasia eutaea (Hewitson, [1853])    iN

Euselasia eutaea eutaea (Hewitson, [1853])  TL: Brazil (Pa)  M T S

Euselasia eutaea lindana Möschler, 1883  TL: Surinam  M T S

Euselasia chinguala J. Hall & Willmott, 1995  TL: Ecuador  M T

Euselasia zena (Hewitson, 1860)    iN  M T1 T2 S

Euselasia erythraea (H. Bates, 1868)  TL: Brazil (Am)  M T S

Euselasia gelon (Stoll, 1787)  TL: Surinam  M OI S

Euselasia thaumata J. Hall & Willmott, 1998    iN  M T OI S

Euselasia hypocrita Stichel, 1919  TL: Bolivia  M T S

Euselasia toppini Sharpe, 1915    iN  M T S

Euselasia eucritus (Hewitson, [1853])    iN  M T

Euselasia calligramma (H. Bates, 1868)    iN  M T S

Euselasia phelina (H. Druce, 1878)    iN  M T

Euselasia pallantis Brévignon & Gallard, 1993    iN  M T TI

Euselasia arpi Gallard, 2013    iN  M T OI S

Euselasia charilis (H. Bates, 1868)  TL: Brazil (Am)  M T S HS

Euselasia pellos Stichel, 1919    iN  M T S

Euselasia zara (Westwood, 1851)    iN  M T S

Euselasia uzita (Hewitson, [1853])    iN  M T S

Euselasia orfita (Cramer, 1777)    iN  M T OI S

Euselasia clithra (H. Bates, 1868)    iN  M T S

Euselasia eutychus (Hewitson, 1856)    iN  M T S

Euselasia cyanira Callaghan, 1997  TL: Peru  M T

Euselasia cuprea Lathy, 1926    iN  M T S

Euselasia phedica (Boisduval, 1836)    iN  M T S

Euselasia euodias (Hewitson, 1856)    iN

Euselasia euodias euodias (Hewitson, 1856)  TL: Brazil (Am)  M T S V

Euselasia euodias talidiman Brévignon & Gallard, 1992  TL: French Guiana  M T S

Euselasia orba Stichel, 1919    iN

Euselasia orba orba Stichel, 1919  TL: Bengala  M OD TI S HS V

Euselasia orba spectralis Stichel, 1919  TL: Brazil (Pa/Am); Peru  M T S

Euselasia pillaca J. Hall & Willmott, 1998  TL: Ecuador  M T OI

Euselasia issoria (Hewitson, 1869)    iN  M T S

Euselasia perisama J. Hall & Lamas, 2001    iN  M T

Eugelasia  Grishin, 2021  

Eugelasia modesta (H. Bates, 1868)  Brazil; ref.151  M T

Eugelasia satyroides (Lathy, 1926)  ref.151  iN  M T S

Eugelasia eugeon (Hewitson, 1856)    iN

Eugelasia eugeon eugeon (Hewitson, 1856)  TL: Brazil (Pa)  M T S

Eugelasia eugeon gorgoniensis (Constantino & Salazar, 2019)  TL: Colombia  M OI

Eugelasia brevicauda (Lathy, 1926)    iN  M T S

  Subfamily Riodininae

  Tribe Eurybiini

Alesa  E. Doubleday, 1847  

Alesa prema (Godart, [1824])    iN  M T S

Alesa telephae (Boisduval, 1836)    iN  M T S

Alesa fournierae Lathy, 1958  TL: Brazil (Am)  M T

Alesa amesis (Cramer, 1777)    iN  M T S

Alesa lipara H. Bates, 1867  TL: Brazil (Pa)  M T S

Alesa hemiurga H. Bates, 1867  TL: Brazil (Am)  M T

Alesa thelydrias H. Bates, 1867  TL: Brazil (Pa)  M T

Alesa rothschildi (Seitz, 1913)  TL: Guyana  M T S

Alesa esmeralda Salazar & Constantino, 2007  TL: Colombia  M T

Alesa amethystina Gallard & Fernandez, 2015  TL: French Guiana  M T

Alesa beneluzi Gallard & Fernandez, 2015  TL: French Guiana

Alesa humilis Brévignon, 2015  TL: French Guiana

Alesa suzana J. Hall & Ahrenholz, 2010  TL: Ecuador

Eurybia  [Illiger], 1807  

Eurybia carolina Godart, [1824]    iN  M S

Eurybia pergaea (Geyer, 1832)    iN  M T S

Eurybia nicaeus (Fabricius, 1775)    iN

Eurybia nicaeus nicaeus (Fabricius, 1775)  TL: India  M T S

Eurybia nicaeus erythinosa Stichel, 1910  TL: Ecuador  M T

Eurybia cyclopia Stichel, 1910  Fire-bordered Eurybia    iN

Eurybia cyclopia cyclopia Stichel, 1910  Costa Rica to Ecuador  M T S L

Eurybia cyclopia montsineryensis Brévignon, 1997  TL: French Guiana

Eurybia molochina Stichel, 1910    iN

Eurybia molochina molochina Stichel, 1910  TL: Brazil (SC)  M T S

Eurybia molochina hyacinthina Stichel, 1910  TL: Brazil (ES)  M T S

Eurybia caerulescens H. Druce, 1904    iN  M T S

Eurybia franciscana C. Felder & R. Felder, 1862    iN  M T S

Eurybia unxia Godman & Salvin, 1885  Azure-winged Eurybia  [= fulgens], Costa Rica and Panama  iN  M T S L

Eurybia lycisca Westwood, 1851  Blue-winged Eurybia  SE Mexico to Ecuador  iN  M T S L1 L2

Eurybia dardus (Fabricius, 1787)    iN

Eurybia dardus dardus (Fabricius, 1787)  TL: Surinam  M T S

Eurybia dardus annulata Stichel, 1910  TL: Colombia; Ecuador; Peru  M T S

Eurybia misellivestis Stichel, 1910    iN  M T

Eurybia silaceana Stichel, 1924    iN  M T

Eurybia jemima Hewitson, 1869    iN

Eurybia jemima jemima Hewitson, 1869  TL: Ecuador  M T S

Eurybia jemima sinnaces H. Druce, 1904  TL: Peru  M T

Eurybia donna C. Felder & R. Felder, 1862    iN  M T S

Eurybia latifasciata (Hewitson, 1869)  TL: Ecuador  M T S

Eurybia patrona Weymer, 1875    iN

Eurybia patrona patrona Weymer, 1875  TL: Colombia  M T

Eurybia patrona persona Staudinger, 1876  Honduras to Panama  M T S L

Eurybia patrona promota Stichel, 1910  TL: Brazil (Am/"SC"); Peru  M T S

Eurybia rubeolata Stichel, 1910    iN

Eurybia rubeolata rubeolata Stichel, 1910  TL: Bolivia; Peru  M T S L

Eurybia rubeolata planaltensis Callaghan, 2001  TL: Brazil (DF)  M T

Eurybia juturna C. Felder & R. Felder, 1865    iN  M T S

Eurybia halimede (Hübner, [1807])    iN

Eurybia halimede halimede (Hübner, [1807])  TL: Brazil  M T S

Eurybia halimede passercula Stichel, 1915  TL: Brazil (MT)  M T

Eurybia albiseriata Weymer, 1890    iN

Eurybia albiseriata albiseriata Weymer, 1890  TL: Brazil (Pa)  M T S

Eurybia albiseriata stellifera Stichel, 1910  TL: Peru; Bolivia  M T S

Eurybia elvina Stichel, 1910  Blind Eurybia    iN

Eurybia elvina elvina Stichel, 1910  E & W Mexico to Panama  M T S1 S2 L I

Eurybia elvina emidiata Stichel, 1915  TL: Brazil (Am)  M T S

Eurybia elvina granulata Stichel, 1910  TL: Peru; Ecuador  M T

Eurybia gonzaga Dolibaina, Dias, O. Mielke & Casagrande, 2014    iN

Eurybia mestiza Salazar, Villalobos & Vargas, 2022  TL: Colombia

  Tribe Mesosemiini

Eunogyra  Westwood, 1851  

Eunogyra satyrus Westwood, 1851    iN

Eunogyra satyrus infernalis Salazar & Constantino, 2009  TL: Colombia

Eunogyra satyrus satyrus Westwood, 1851  TL: Brazil (Pa)  M T S

Eunogyra curupira H. Bates, 1868    iN  M T S

Teratophthalma  Stichel, 1909  

Teratophthalma axilla (H. Druce, 1904)    iN

Teratophthalma axilla axilla (H. Druce, 1904)  TL: Bolivia  M T S

Teratophthalma axilla lactifusa Stichel, 1910  TL: Ecuador  M T

Teratophthalma axilla vegeta Stichel, 1910  TL: Peru  M T

Teratophthalma maenades (Hewitson, 1858)    iN

Teratophthalma maenades maenades (Hewitson, 1858)  TL: Colombia  M T S

Teratophthalma maenades coronata Stichel, 1915  TL: Ecuador  M T L

Teratophthalma maenades marsena (Hewitson, 1869)  TL: Ecuador  M T

Teratophthalma maenades marsidia (Hewitson, 1869)  TL: Ecuador  M T

Teratophthalma maenades polyplusia Stichel, 1910  TL: Peru  M T S

Teratophthalma bacche (Seitz, 1916)    iN

Teratophthalma bacche bacche (Seitz, 1916)  TL: Colombia  M T

Teratophthalma bacche adulter Stichel, 1929  TL: Peru  M T

Teratophthalma monochroma Stichel, 1910    iN  M T S

Teratophthalma phelina (C. Felder & R. Felder, 1862)    iN

Teratophthalma phelina phelina (C. Felder & R. Felder, 1862)  TL: Colombia  M T S

Teratophthalma phelina analoga (Seitz, 1916)  TL: Colombia  M T

Teratophthalma phelina vicina Stichel, 1926  TL: Peru  M T

Ectosemia  Grishin, 2021  ref.12

Ectosemia eumene (Cramer, 1776)  ref.12  iN  M T S

Ectosemia attavus (J. Zikán, 1952)  Amazonian; ref.151  M T S

Ectosemia erinnya (Stichel, 1910)  ref.12,151  iN  M T S

Ectosemia decolorata (Lathy, 1932)  ref.12

Ectosemia decolorata decolorata (Lathy, 1932)  TL: Brazil (MT)  M T S

Ectosemia decolorata meyi (Brévignon, 1997)  TL: French Guiana

Ectosemia steli (Hewitson, 1858)  ref.12  iN  M T S

Ectosemia nesti (Hewitson, 1858)    iN  M T S

Ectosemia acuta (Hewitson, 1873)  ref.151  iN  M T S

Ectosemia eurythmia (Stichel, 1915)  ref.151  M T

Endosemia  Grishin, 2021  ref.12

Endosemia ulrica (Cramer, 1777)  ref.12  iN

Endosemia ulrica ulrica (Cramer, 1777)  TL: Surinam  M T S

Endosemia ulrica messala (Hewitson, 1870)  TL: Ecuador  M T

Endosemia macella (Hewitson, 1859)  ref.12  iN  M T S

Mesosemia  Hübner, [1819]  

Mesosemia misipsa Hewitson, 1859    iN

Mesosemia misipsa misipsa Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia misipsa carapa Brévignon, 2013  TL: French Guiana

Mesosemia metura Hewitson, [1873]    iN

Mesosemia metura metura Hewitson, [1873]  TL: ?  M S HS

Mesosemia metura polyglauca Stichel, 1910  TL: Peru  M T

Mesosemia teulem Brévignon, 1995  TL: French Guiana  M T S

Mesosemia esperanza Schaus, 1913  Esperanza Eyed-Metalmark  Costa Rica and Panama  iN  M T S

Mesosemia coelestis Godman & Salvin, 1885  Pale-blue Eyed-Metalmark  Costa Rica to Colombia  iN  M T S L

Mesosemia bahia Callaghan, 1999  TL: Brazil (MG)  M T

Mesosemia mesoba Hewitson, [1873]    iN  M T S

Mesosemia zikla Hewitson, 1869  TL: Ecuador  M T

Mesosemia dulcis Stichel, 1910    iN  M T S

Mesosemia eugenea Stichel, 1910  TL: Surinam

Mesosemia eugenea eugenea Stichel, 1910  TL: Surinam  M T

Mesosemia eugenea pythonica Stichel, 1923  TL: Brazil (Pa)  M T

Mesosemia naliniae Brévignon, 2015  TL: French Guiana  M T S

Mesosemia ephyne (Cramer, 1776)    iN  M T

Mesosemia jucunda Stichel, 1923    iN  M T

Mesosemia gneris Westwood, 1851    iN  M T S

Mesosemia albipuncta Schaus, 1913  White-spotted Eyed-Metalmark  Costa Rica and Panama  M T S I

Mesosemia cyanira Stichel, 1909    iN  M T S

Mesosemia ackeryi Brévignon, 1997  TL: French Guiana

Mesosemia thetys Godman & Salvin, 1885    iN  M T S

Mesosemia hesperina A. Butler, 1874  Hesperina Eyed-Metalmark    iN

Mesosemia hesperina hesperina A. Butler, 1874  Nicaragua to Venezuela  M T S L

Mesosemia hesperina tenuivittata Stichel, 1910  TL: Brazil (RJ)  M T

Mesosemia lycorias Stichel, 1915  TL: Brazil (Pa)  M T S

Mesosemia synnephis Stichel, 1909    iN  M T S

Mesosemia maeotis Hewitson, 1859  TL: Brazil (Am)

Mesosemia maeotis maeotis Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia maeotis amicula Stichel, 1915  TL: Brazil (Pa)  M T S

Mesosemia cippus Hewitson, 1859    iN  M T S

Mesosemia sylvina H. Bates, 1867  TL: Brazil (Pa)  M T

Mesosemia walteri Brévignon, 1998    iN

Mesosemia walteri walteri Brévignon, 1998  TL: French Guiana  M T

Mesosemia walteri serenitas Brévignon, 2013  TL: French Guiana  M T

Mesosemia kahuapayani J. Hall & Harvey, 2004    iN  M T

Mesosemia minutula Gallard, 1996  TL: French Guiana  M T

Mesosemia ibycus Hewitson, 1859    iN  M T S

Mesosemia minos Hewitson, 1859    iN

Mesosemia minos minos Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia minos guaramiranga P. Jauffret, J. Jauffret & Pessôa, 2008  TL: Brazil

Mesosemia minos orarius Brévignon, 2013  TL: French Guiana

Mesosemia minos modica Stichel, 1910  TL: Surinam  M T S

Mesosemia minos tetrophthalma Stichel, 1915  TL: Brazil (Pe)  M T S

Mesosemia mosera Hewitson, 1860  TL: Brazil (Pa)  M T S

Mesosemia melese Hewitson, 1860  TL: Brazil (Pa)  M T S

Mesosemia kwokii D'Abrera, 1994    iN  M T

Mesosemia myonia Hewitson, 1859  TL: Brazil (Pa)  M T S

Mesosemia methion Hewitson, 1860    iN

Mesosemia methion methion Hewitson, 1860  TL: Brazil  M S

Mesosemia methion maera Hewitson, [1873]  Trinidad  iN  M T

Mesosemia araeostyla Stichel, 1915  TL: Guyana  M T

Mesosemia luperca Stichel, 1910    iN  M T S

Mesosemia melaene Hewitson, 1859    iN

Mesosemia melaene melaene Hewitson, 1859  TL: Brazil (Pa)  M T S

Mesosemia melaene pinguilenta Stichel, 1915  TL: Guyana  M T

Mesosemia inconspicua Lathy, 1932  TL: French Guiana  M T S

Mesosemia nyctea (Hoffmansegg, 1818)    iN

Mesosemia nyctea nyctea (Hoffmansegg, 1818)  TL: Surinam  M T S

Mesosemia nyctea fluminensis J. Zikán, 1952  TL: Brazil (RJ)

Mesosemia nyctea lato Stichel, 1910  TL: Brazil (Ba)  M T S

Mesosemia scotina Stichel, 1909  TL: Surinam  M T S

Mesosemia lacernata Stichel, 1909    iN  M T S

Mesosemia melpia Hewitson, 1859    iN

Mesosemia melpia melpia Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia melpia vaporosa Stichel, 1910  TL: Brazil (Pa)  M T S

Mesosemia metope Hewitson, 1859    iN

Mesosemia metope metope Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia metope pruinosa Stichel, 1910  TL: Peru  M T

Mesosemia metope ungulata Stichel, 1915  TL: Brazil (Pa)  M T S

Mesosemia ahava Hewitson, 1869    iN

Mesosemia ahava ahava Hewitson, 1869  TL: Ecuador  M T S

Mesosemia ahava veleda Stichel, 1910  TL: Bolivia  M T

Mesosemia olivencia H. Bates, 1868    iN  M T S

Mesosemia zanoa Hewitson, 1869    iN

Mesosemia zanoa zanoa Hewitson, 1869  TL: Ecuador  M T S

Mesosemia zanoa orthia Stichel, 1915  TL: Colombia  M T S

Mesosemia anceps Stichel, 1915    iN  M T

Mesosemia philocles (Linnaeus, 1758)    iN

Mesosemia philocles philocles (Linnaeus, 1758)  TL: Indiis  M S

Mesosemia philocles egabella H. Bates, 1868  TL: Brazil (Am)  M T S

Mesosemia philocles jeziela A. Butler, 1869  TL: Colombia  M T S

Mesosemia philocles laetifica H. Bates, 1868  TL: Brazil (Pa)  M T

Mesosemia philocles limbata Stichel, 1915  TL: Ecuador?  M T

Mesosemia philocles sylvia H. Druce, 1878  TL: Bolivia  M T

Mesosemia philocles thyestes H. Druce, 1878  TL: Peru  M T S

Mesosemia machaera Hewitson, 1860    iN

Mesosemia machaera machaera Hewitson, 1860  TL: Brazil (Am)  M T S

Mesosemia machaera dealbata Stichel, 1910  TL: Colombia  M T

Mesosemia machaera salesiana J. Zikán, 1952  TL: Brazil (Am)  M T

Mesosemia modulata Stichel, 1910  TL: Peru  M T S

Mesosemia magete Hewitson, 1860    iN

Mesosemia magete humboldtiana G. Rodríguez, Salazar & Constantino, 2010  TL: Colombia

Mesosemia magete magete Hewitson, 1860  TL: Brazil (Pa)  M T

Mesosemia magete bersabana Stichel, 1910  TL: Surinam  M T S

Mesosemia thymetus (Cramer, 1777)    iN

Mesosemia thymetus thymetus (Cramer, 1777)  TL: Surinam  M T S

Mesosemia thymetus thymetina A. Butler, 1869  TL: Colombia  M T S

Mesosemia thymetus umbrosa Stichel, 1909  TL: Bolivia  M T S

Mesosemia quadralineata J. Hall & Harvey, 2004  TL: Peru  M T

Mesosemia sifia (Boisduval, 1836)  TL: Brazil  M T S

Mesosemia isshia A. Butler, 1869    iN  M T S

Mesosemia latizonata A. Butler, 1874    iN  M T S

Mesosemia impedita Stichel, 1909  TL: Brazil (Pa)  M T

Mesosemia zonalis Godman & Salvin, 1885  Whitened Eyed-Metalmark  Honduras to Colombia  iN

Mesosemia zonalis zonalis Godman & Salvin, 1885  Honduras to Colombia  M T S L

Mesosemia zonalis gorgoniensis Constantino, G. Rodríguez & Salazar, 2010  TL: Colombia

Mesosemia gertraudis Stichel, 1910  TL: Peru  M T S

Mesosemia myrmecias Stichel, 1910  TL: Peru  M T

Mesosemia hedwigis Stichel, 1910  TL: Bolivia; Peru  M T S

Mesosemia amarantus Stichel, 1910  TL: Peru  M T

Mesosemia nympharena Stichel, 1909    iN  M T S

Mesosemia naiadella Stichel, 1909    iN

Mesosemia naiadella naiadella Stichel, 1909  TL: Peru; Guyana  M T S

Mesosemia naiadella dryadella Stichel, 1909  TL: Ecuador  M T

Mesosemia naiadella oreas Stichel, 1915  TL: Guyana  M T

Mesosemia pardalis Callaghan, 2001    iN

Mesosemia nerine Stichel, 1909  TL: Bolivia  M T

Mesosemia tenebricosa Hewitson, 1877    iN

Mesosemia tenebricosa tenebricosa Hewitson, 1877  TL: Ecuador  M T

Mesosemia tenebricosa anica H. Druce, 1904  TL: Peru  M T S

Mesosemia tenebricosa serenata Stichel, 1923  TL: Brazil (Pa)  M T

Mesosemia sirenia Stichel, 1909    iN  M T

Mesosemia judicialis A. Butler, 1874    iN  M T S

Mesosemia odice (Godart, [1824])    iN  M T S

Mesosemia mathania Schaus, 1902    iN  M T

Mesosemia cymotaxis Stichel, 1910  TL: Brazil  M T

Mesosemia veneris A. Butler, 1871  TL: Brazil (MG)  M T

Mesosemia rhodia (Godart, [1824])    iN  M T S

Mesosemia moesia Hewitson, [1857]    iN  M T

Mesosemia friburgensis Schaus, 1902    iN  M T S

Mesosemia meeda Hewitson, 1858    iN  M T S

Mesosemia mayi Lathy, 1958  TL: Brazil (RJ)  M T

Mesosemia esmeralda Gallard & Brévignon, 1989  TL: French Guiana  M T

Mesosemia epidius Hewitson, 1859  TL: French Guiana  M T S

Mesosemia evias Stichel, 1923    iN  M T S

Mesosemia bella Sharpe, 1890  TL: Brazil (Go)  M T

Mesosemia phace Godman, 1903  TL: Guyana  M T S

Mesosemia thera Godman, 1903    iN  M T S

Mesosemia thyas Stichel, 1910  TL: Brazil (Pa); Surinam  M T S

Mesosemia orbona Godman, 1903    iN

Mesosemia orbona orbona Godman, 1903  TL: Surinam  M T S

Mesosemia orbona caballina Brévignon, 1997  TL: French Guiana

Mesosemia carderi H. Druce, 1904    iN  M T S

Mesosemia ceropia H. Druce, 1874  Spying Eyed-Metalmark  Costa Rica to Colombia  M T S

Mesosemia hypermegala Stichel, 1909  Hiding Eyed-Metalmark  Costa Rica to Colombia  iN  M T S L

Mesosemia antaerice Hewitson, 1859    iN  M T S

Mesosemia ozora Hewitson, 1869  TL: Ecuador  M T

Mesosemia menoetes Hewitson, 1859    iN

Mesosemia menoetes menoetes Hewitson, 1859  TL: Brazil (Am)  M T S

Mesosemia menoetes mennonia Stichel, 1909  TL: Peru; Brazil (Am)  M T

Mesosemia menoetes paetula Stichel, 1915  TL: Bolivia  M T

Mesosemia menoetes pulverulenta Brévignon, 1997  TL: French Guiana

Mesosemia putli Seitz, 1913  TL: Peru  M T

Mesosemia metuana (C. Felder & R. Felder, 1865)    iN

Mesosemia metuana metuana (C. Felder & R. Felder, 1865)  TL: Colombia  M T S

Mesosemia metuana vargasi Salazar, Rodríguez & Constantino, 2010  TL: Colombia  M T

Mesosemia metuana montezuma Salazar, Geale & G. Rodríguez, 2017  TL: Colombia

Mesosemia metuana glaucoma Stichel, 1909  TL: Bolivia  M T S

Mesosemia cordillerensis Salazar & Constantino, 1993    iN  M T

Mesosemia mehida Hewitson, 1869  TL: Ecuador  M T S

Mesosemia reba Hewitson, 1869    iN

Mesosemia reba reba Hewitson, 1869  TL: Ecuador  M T S

Mesosemia reba agnilata Dognin, 1893  TL: Ecuador  M T

Mesosemia zorea Hewitson, 1869    iN

Mesosemia zorea chocoensis Salazar, G. Rodríguez & Constantino, 2010  TL: Colombia

Mesosemia zorea zorea Hewitson, 1869  TL: Ecuador  M T S

Mesosemia zorea toparcha Stichel, 1910  TL: Peru  M T S

Mesosemia macrina (C. Felder & R. Felder, 1865)    iN

Mesosemia macrina macrina (C. Felder & R. Felder, 1865)  TL: Colombia  M T S

Mesosemia macrina henaoi Salazar, Constantino & Rodríguez, 2010  TL: Colombia  M T

Mesosemia macrina sticheli Constantino, Salazar & Rodríguez, 2010  TL: Colombia  M T

Mesosemia adida Hewitson, 1869  TL: Ecuador  M T S

Mesosemia praeculta Stichel, 1910    iN

Mesosemia praeculta colombiana Salazar, G. Rodríguez & Constantino, 2010  TL: Colombia

Mesosemia praeculta praeculta Stichel, 1910  TL: Bolivia  M T S

Mesosemia telegone (Boisduval, 1836)  Violet-washed Eyed-Metalmark    iN

Mesosemia telegone telegone (Boisduval, 1836)  SE Mexico to Colombia  M T S L

Mesosemia telegone amiana C. Felder & R. Felder, 1861  TL: Venezuela  M T

Mesosemia telegone paramba H. Druce, 1904  TL: Ecuador  M T

Mesosemia lamachus Hewitson, 1857  Purple-washed Eyed-Metalmark  [= tertica], E & W Mexico to Colombia  iN  M T S1 S2 L I

Mesosemia carissima H. Bates, 1866  Blue-patched Eyed-Metalmark  Nicaragua to Panama  iN  M T S1 S2 L I

Mesosemia grandis H. Druce, 1874  Giant Eyed-Metalmark  Costa Rica and Panama  iN  M T S1 S2 L I1 I2

Mesosemia gaudiolum H. Bates, 1865  Gaudy Eyed-Metalmark  SE Mexico to Costa Rica  iN  M T S

Mesosemia gemina J. Maza & R. G. Maza, 1980  Turquoise Eyed-Metalmark  SE Mexico  iN  M T S L

Mesosemia sibyllina Staudinger, [1887]    iN  M T S

Mesosemia mamilia Hewitson, 1870    iN  M T S

Mesosemia mevania Hewitson, [1857]    iN

Mesosemia mevania mevania Hewitson, [1857]  TL: Colombia  M T S

Mesosemia mevania magnessa Stichel, 1909  TL: Peru  M T

Mesosemia mevania mimallonis Stichel, 1909  TL: Ecuador  M T S

Mesosemia pacifica Stichel, 1926    iN  M T S

Mesosemia mancia Hewitson, 1870    iN  M T S

Mesosemia loruhama Hewitson, 1869    iN

Mesosemia loruhama loruhama Hewitson, 1869  TL: Ecuador  M T S

Mesosemia loruhama candara H. Druce, 1904  TL: Peru  M T S

Mesosemia loruhama sidonia Stichel, 1924  TL: Colombia  M T

Mesosemia asa Hewitson, 1869  Deep-blue Eyed-Metalmark    iN

Mesosemia asa asa Hewitson, 1869  Nicaragua to Panama  M T S L1 L2 I

Mesosemia asa asopis Stichel, 1909  Panama and Colombia  M T S

Mesosemia asa iphigenia Stichel, 1926  TL: Colombia  M T

Mesosemia asa mycene Hewitson, 1870  TL: Ecuador  M T S

Mesosemia harveyi DeVries & J. Hall, 1996  Harvey's Eyed-Metalmark  Costa Rica and Panama  M T S

Mesosemia messeis Hewitson, 1860    iN

Mesosemia messeis messeis Hewitson, 1860  TL: Brazil (Am)  M T S

Mesosemia messeis itoupensis Gallard & Fernandez, 2012  TL: French Guiana

Mesosemia messeis amona Hewitson, 1876  TL: Bolivia  M T S

Mesosemia lapillus Stichel, 1910    iN  M T S

Mesosemia porce Rodríguez, Constantino & Salazar, 2010  TL: Colombia  M T

Mesosemia portentosa Salazar, Constantino & Rodríguez, 2010    iN  M T

Mesosemia vemania Constantino, Salazar & Rodríguez, 2010    iN  M T

Mesosemia atachibakka Brévignon, 2013  TL: French Guiana

Mesosemia exilinea Gallard, 2015  TL: French Guiana  M T

Mesosemia firmitas Brévignon, 2013  TL: French Guiana  M T

Mesosemia litany Brévignon, 2019  TL: French Guiana

Mesosemia pakaraima Fratello, 2019  TL: Guyana  M S

Mesosemia patientia Brévignon, 2013  TL: French Guiana  M T

Mesosemia perspicacitas Brévignon, 2013  TL: French Guiana  M T

Mesosemia sagacitas Brévignon, 2013  TL: French Guiana  M T

Mesosemia tenacitas Brévignon, 2013  TL: French Guiana  M T

Mesosemia vigilantia Brévignon, 2013  TL: French Guiana  M T

Semomesia  Westwood, 1851  

Semomesia geminus (Fabricius, 1793)    iN  M S

Semomesia croesus (Fabricius, 1777)    iN

Semomesia croesus croesus (Fabricius, 1777)  TL: Surinam  M T S

Semomesia croesus lacrimosa Stichel, 1915  TL: Colombia  M T S

Semomesia croesus meana (Hewitson, 1858)  TL: Brazil (Am)  M T S

Semomesia croesus siccata Stichel, 1919  TL: Bolivia  M T S

Semomesia croesus trilineata (A. Butler, 1874)  TL: Brazil (Am)  M T S

Semomesia croesus undosa Stichel, 1919  TL: Ecuador  M T

Semomesia alyattes J. Zikán, 1952    iN

Semomesia alyattes alyattes J. Zikán, 1952  TL: Brazil (Am)  M T S

Semomesia alyattes fassli P. Jauffret & J. Jauffret, 2009  TL: Brazil

Semomesia cecilae Gallard, 1997  TL: French Guiana  M T

Semomesia capanea (Cramer, 1779)    iN

Semomesia capanea capanea (Cramer, 1779)  TL: Surinam  M T S

Semomesia capanea ionima Stichel, 1910  TL: Brazil (Am)  M T

Semomesia capanea sodalis Stichel, 1919  TL: Peru  M T

Semomesia marisa (Hewitson, 1858)    iN

Semomesia marisa marisa (Hewitson, 1858)  TL: Brazil (Am)  M T S

Semomesia marisa hewitsoni Stichel, 1915  TL: Brazil (Pa)  M T S

Semomesia macaris (Hewitson, 1859)  TL: Brazil (Am)  M T S

Semomesia tenella Stichel, 1910    iN

Semomesia tenella tenella Stichel, 1910  TL: Peru  M T S

Semomesia tenella aetherea Stichel, 1910  TL: Peru  M T

Semomesia tenella lucana Stichel, 1924  TL: Ecuador  M T

Semomesia tenella optima Stichel, 1915  TL: Brazil (Pa)  M T S

Semomesia wallacei P. Jauffret & J. Jauffret, 2009  TL: Brazil

Leucochimona  Stichel, 1909  

Leucochimona icare (Hübner, [1819])    iN

Leucochimona icare icare (Hübner, [1819])  TL: Surinam  M T S

Leucochimona icare matatha (Hewitson, 1873)  TL: [Brazil (MG)]  M T

Leucochimona icare nivea (Godman & Salvin, 1885)  TL: Bolivia  M T

Leucochimona icare polita Stichel, 1910  TL: Colombia  M T

Leucochimona icare subalbata (Seitz, 1913)  TL: Bolivia  M T S

Leucochimona aequatorialis (Seitz, 1913)    iN  M T S

Leucochimona vestalis (H. Bates, 1865)  Vestalis Metalmark    iN

Leucochimona vestalis vestalis (H. Bates, 1865)  SE Mexico and Guatemala  M T S L

Leucochimona vestalis leucogaea (Godman & Salvin, 1885)  Guatemala to Panama  M T S

Leucochimona lepida (Godman & Salvin, 1885)  Satyr Metalmark    iN

Leucochimona lepida lepida (Godman & Salvin, 1885)  Nicaragua to Panama  M T S1 S2 L I

Leucochimona lepida nivalis (Godman & Salvin, 1885)  SE Mexico to Guatemala  M T S L

Leucochimona iphias Stichel, 1909  Iphias Metalmark  Panama  iN  M T S1 S2 L I

Leucochimona lagora (Herrich-Schäffer, [1853])  Lagora Metalmark  Honduras to Guianas & Ecuador  iN  M T S HS L V

Leucochimona molina (Godman & Salvin, 1885)  Molina Metalmark  Nicaragua to Panama  iN  M T TI S I

Leucochimona hyphea (Cramer, 1776)    iN

Leucochimona hyphea hyphea (Cramer, 1776)  TL: West Indies  M T S

Leucochimona hyphea pallida (Lathy, 1932)  TL: Peru  M T

Leucochimona matisca (Hewitson, 1860)    iN  M T S

Leucochimona anophthalma (C. Felder & R. Felder, 1865)    iN  M T S

Leucochimona libna Brévignon, 2013  TL: French Guiana

Mesophthalma  Westwood, 1851  

Mesophthalma idotea (Westwood, 1851)    iN  M T S

Mesophthalma mirita (Herrich-Schäffer, [1853])  TL: Surinam  M T S

Perophthalma  Westwood, 1851  

Perophthalma tullius (Fabricius, 1787)  Tullius Metalmark  Costa Rica to Brazil  iN  M T S L

Perophthalma lasus (Westwood, 1851)  Lasus Metalmark  SE Mexico to Panama  iN  M T S L I

Perophthalma lasciva Stichel, 1929    iN  M T S

Hyphilaria  Hübner, [1819]  

Hyphilaria anthias (Hewitson, 1874)    iN  M T S L

Hyphilaria nicia Hübner, [1819]    iN  M S

Hyphilaria parthenis (Westwood, 1851)  Fire-banded Metalmark  C Panama to Bolivia & S Brazil, Guianas  iN  M T S L

Eucorna  Strand, 1932  monotypic genus; ref.12

Eucorna sanarita (Schaus, 1902)  S Brazil; ref.12  iN  M T S

Cremna  E. Doubleday, 1847  ref.12

Cremna alector (Geyer, 1837)  Brazil; ref.12  iN  M T S

Cremna pupillata Stichel, 1915  Brazil; ref.151  M T S

Cremna radiata (Godman & Salvin, 1886)  White-rayed Metalmark  Costa Rica to Colombia & W Ecuador; ref.12  M T S

Cremna dentata (Stichel, 1910)  Colombia; ref.151  M T

Cremna theata (Stichel, 1910)  Blue-banded Metalmark  Costa Rica to W Ecuador; ref.12  iN  M T S I H

Cremna actoris (Cramer, 1776)  Actoris Metalmark  far SE Peru, Bolivia, C & S Amazonia; ref.12  iN  M T S L

Cremna heteroea H. Bates, 1867  Blue-washed Metalmark  Guianas, Venezuela, Colombia to S Peru, Amazonas; ref.12  iN  M T S L

Cremna telarania Grishin, 2023  Bolivia; ref.151  M T

Cremna calitra Hewitson, 1869  Ecuador; ref.151  M T S

Napaea  Hübner, [1819]  ref.12

Napaea sylva (Möschler, 1877)  ref.12  iN  M T S

Napaea beltiana (H. Bates, 1867)  ref.12  iN  M T S

Napaea dramba (J. Hall, Robbins & Harvey, 2004)  Fossil from Dominican Republic; ref.125,12  M OI

Napaea danforthi A. Warren & Opler, 1999  Ancient Metalmark  NW Mexico (Sonora to Jalisco); ref.124,12  iN  M T S L

Napaea umbra (Boisduval, 1870)  Quilted Metalmark  E Mexico to Panama; ref.12  iN

Napaea umbra W Mexican segregate (, )  W Mexico to Chiapas  M S L

Napaea umbra veracruzana (RG. Maza & Attal, 2016)  Mexico; ref.84

Napaea umbra Chiapas segregate (, )  S-central Chiapas (very large adults)  M S

Napaea umbra umbra (Boisduval, 1870)  TL: Honduras  M S1 S2 L I

Napaea loxicha (RG. Maza & J. Maza, 2016)  Mexico; ref.84,12  iN  M S

Napaea maya (J. Maza & Lamas, 2016)  Mexico; ref.84,12  iN  M S

Napaea necaxa (RG. Maza & J. Maza, 2018)  Mexico; ref.85,12

Napaea totonaca (RG. Maza & J. Maza, 2016)  Mexico; ref.84,12  iN  M S

Napaea rufolimba J. Hall, 2005  Red-margined Metalmark  SE Mexico to W & C Ecuador; ref.86,12  iN  M T S

Napaea tumbesia J. Hall & Lamas, 2001  Peru; ref.12  M T S L

Napaea fratelloi J. Hall & Harvey, 2005  ref.86,12  M S

Napaea eucharila (H. Bates, 1867)  White-stitched Metalmark  [= picina], SE Mexico to Bolivia, Paraguay, S Brazil, Guianas, T ...; ref.12  iN  M T OD TI S1 S2 HS L I

Napaea frustatoria Brévignon, 2019  ref.87,12  M T

Napaea phryxe (C. Felder & R. Felder, 1865)  Brazil; ref.12  iN  M T S

Napaea agroeca Stichel, 1910  S Brazil; ref.12  iN  M T S

Napaea cebrenia (Hewitson, [1873])  ref.12  iN  M T S

Napaea zikani Stichel, 1923  ref.12  M T S

Napaea elisae (J. Zikán, 1952)  ref.12  M T S

Napaea joinvilea J. Hall & Harvey, 2005  ref.86,12  M T S

Napaea melampia (H. Bates, 1867)  ref.12  M T

Napaea mellosa J. Hall & Harvey, 2005  ref.86,12  iN  M T S

Napaea gynaecomorpha J. Hall, Harvey & Gallard, 2005  ref.86,12  iN  M T S

Napaea merula (Thieme, 1907)  ref.12  iN  M T S

Napaea thasus (Stoll, 1780)  Thasus Metalmark  [= subrutila], genus tentative; SE Mexico S Brazil & Paraguay, Guianas  iN  M T S L

Ithomiola  C. Felder & R. Felder, 1865  [includes Hermathena as a subgenus]; ref.12

Ithomiola eburna (J. Hall & Harvey, 2005)  White Metalmark  [= C. Am. records of H. candidata], Costa Rica to W Ecuador  iN  M S

Ithomiola candidata (Hewitson, 1874)    iN  M T S

Ithomiola oweni (Schaus, 1913)  Peppered Metalmark  SE Mexico to Costa Rica  iN  M T S L

Ithomiola calculosa J. Hall & Harvey, 2005  Calculosa Metalmark  Costa Rica to C Panama  M T S

Ithomiola tessera J. Hall, 2005  White-spotted Metalmark  SE Mexico to Costa Rica; ref.154  M S L

Ithomiola tesseroides Grishin, 2024  Panama; ref.154  M T

Ithomiola theages (Godman & Salvin, 1878)  White-spotted Metalmark  Costa Rica to NW Ecuador; ref.154  iN  M T TI S L

Ithomiola coladoris Grishin, 2024  Panama; ref.154  M T

Ithomiola cribralis (Stichel, 1915)  Cloud-spotted Metalmark  C Panama to W Ecuador  iN  M T S

Ithomiola neildi (J. Hall & Willmott, 1998)    iN  M T

Ithomiola bajotanos J. Hall, 2005    iN  M T

Ithomiola perutanos Grishin, 2024  Peru; ref.154  M T

Ithomiola tanos (Stichel, 1910)    iN  M T S

Ithomiola nepos (Fabricius, 1793)    iN  M S

Ithomiola orpheus (Westwood, 1851)    iN  M T S

Ithomiola callixena (Hewitson, 1870)    iN  M T S

Ithomiola buckleyi J. Hall & Willmott, 1998  TL: Ecuador  M T S

Ithomiola floralis C. Felder & R. Felder, 1865    iN

Ithomiola floralis floralis C. Felder & R. Felder, 1865  TL: Surinam  M T S

Ithomiola floralis cascella (Hewitson, 1870)  TL: Ecuador  M T

Ithomiola floralis celtilla (Hewitson, 1870)  E Panama to Bolivia  M T S

  Tribe Dianesiini

Dianesia  Harvey & Clench, 1980  

Dianesia carteri (W. Holland, 1902)  Caribbean Metalmark    iN

Dianesia carteri carteri (W. Holland, 1902)  Bahamas (New Providence, L. San Salvador, Andros, Exumas)  M T S

Dianesia carteri ramsdeni (Skinner, 1912)  Cuba  M T S L

  Tribe Symmachiini

Lucillella  Strand, 1932  

Lucillella camissa (Hewitson, 1870)    iN  M T

Lucillella suberra (Hewitson, 1877)  TL: Ecuador  M T

Lucillella asterra (Grose-Smith, 1898)    iN  M T S

Lucillella pomposa (Stichel, 1910)    iN  M T S

Lucillella arcoirisa J. Hall & Willmott, 2010  TL: Ecuador

Lucillella splendida J. Hall & Harvey, 2007  ref.222  M S

Lucillella aphrodita Rodríguez, Salazar & Constantino, 2011    iN  M T

Esthemopsis  C. Felder & R. Felder, 1865  

Esthemopsis clonia C. Felder & R. Felder, 1865  Clonia Metalmark  S Mexico to Colombia  iN  M T S

Esthemopsis alicia (H. Bates, 1865)  Alicia's Metalmark    iN

Esthemopsis alicia alicia (H. Bates, 1865)  S Mexico & Guatemala  M T S

Esthemopsis alicia saracena Grose-Smith, 1902  TL: Brazil  M T S

Esthemopsis macara (Grose-Smith, 1902)  TL: Colombia  M T

Esthemopsis jesse (A. Butler, 1870)    iN

Esthemopsis jesse jesse (A. Butler, 1870)  TL: Venezuela  M T

Esthemopsis jesse aeniacus Hewitson, 1876  TL: Bolivia  M T S

Esthemopsis colaxes Hewitson, 1870  Cloaxes Metalmark    iN

Esthemopsis colaxes colaxes Hewitson, 1870  Costa Rica to W Ecuador  M T S L

Esthemopsis colaxes guianensis (A. Hall, 1939)  TL: Guyana

Esthemopsis sericina (H. Bates, 1867)    iN  M T S

Esthemopsis crystallina Brévignon & Gallard, 1992  TL: French Guiana  M T S

Esthemopsis pherephatte (Godart, [1824])  Godart's Metalmark  [= caeruleata], S Mexico to Panama  iN  M T S I V

Esthemopsis teras (Stichel, 1910)    iN

Esthemopsis teras teras (Stichel, 1910)  TL: Brazil (SC/Am)  M T S

Esthemopsis teras thouvenoti Gallard, 2017  TL: French Guiana

Esthemopsis pallida Lathy, 1932    iN  M T

Esthemopsis talamanca J. Hall & Harvey, 2007  ref.222  M S

Esthemopsis chocoana Constantino, Salazar & Rodríguez, 2011  TL: Colombia  M T

Esthemopsis diamantina Dias, Dolibaina, O. Mielke & Casagrande, 2014  TL: Brazil

Esthemopsis sateri Dolibaina, Dias, O. Mielke & Casagrande, 2014  TL: Brazil

Mesene  E. Doubleday, 1847  

Mesene aeolia (H. Bates, 1868)  ref.12  M T S

Mesene celina (H. Bates, 1868)  ref.12  iN  M T S

Mesene argentea H. Bates, 1866  Silvery Metalmark  ref.12  iN

Mesene argentea argentea H. Bates, 1866  S Mexico to Panama  M T S L

Mesene argentea carnutes (Hewitson, 1875)  TL: Colombia  M T S

Mesene argentea flavibasis (Lathy, 1932)  TL: Colombia  M T S

Mesene phareus (Cramer, 1777)  Cell-barred Metalmark  S Mexico to Brazil & Peru  iN  M T S L

Mesene leucophrys H. Bates, 1868    iN  M T1 T2 S

Mesene ingrumaensis Callaghan & Salazar, 1999  TL: Colombia  M T

Mesene epaphus (Stoll, 1780)    iN

Mesene epaphus epaphus (Stoll, 1780)  TL: Surinam  M T S

Mesene epaphus pyrrha H. Bates, 1868  TL: Brazil (Am)  M T S

Mesene pyrippe Hewitson, 1874    iN

Mesene pyrippe pyrippe Hewitson, 1874  TL: Brazil (ES)  M T S

Mesene pyrippe sanguilenta Stichel, 1910  TL: Brazil (SC/RS)  M T

Mesene paraena H. Bates, 1868    iN  M T S

Mesene hyale C. Felder & R. Felder, 1865  TL: Colombia  M T

Mesene mygdon Schaus, 1913  Red Metalmark  Costa Rica to Brazil  iN  M T S L

Mesene croceella H. Bates, 1865  Guatemalan Metalmark  E & S Mexico to Costa Rica, [Panama?]  iN  M T S L

Mesene cyneas (Hewitson, 1874)  TL: Bolivia  M T S

Mesene bomilcar (Stoll, 1790)  TL: Surinam

Mesene citrinella J. Hall & Willmott, 1995  TL: Ecuador  M T

Mesene nepticula Möschler, 1877  TL: Surinam

Mesene nepticula nepticula Möschler, 1877  TL: Surinam  M T

Mesene nepticula stigmosa Stichel, 1910  TL: Brazil (Pa); Ecuador  M T

Mesene bigemmis Stichel, 1925    iN  M T

Mesene simplex H. Bates, 1868    iN  M T S

Mesene leucogyna J. Hall & Lamas, 2007    iN

Mesene leucogyna leucogyna J. Hall & Lamas, 2007  TL: Peru  M S

Mesene leucogyna notia J. Hall & Lamas, 2007  TL: Peru

Mesene celetes H. Bates, 1868    iN  M T S

Mesene monostigma (Erichson, [1849])    iN

Mesene monostigma monostigma (Erichson, [1849])  TL: Guyana  M T S

Mesene monostigma discolor Stichel, 1929  TL: Bolivia  M T

Mesene philonis Hewitson, 1874    iN  M T S

Mesene babosa J. Hall & Willmott, 1995  TL: Ecuador  M T

Mesene veleda Stichel, 1923  TL: Brazil (Pa)

Mesene veleda veleda Stichel, 1923  TL: Brazil (Pa)  M T

Mesene veleda herteli J. Zikán, 1952  TL: Brazil (Am)  M T

Mesene nola Herrich-Schäffer, [1853]    iN  M T S

Mesene eupteryx H. Bates, 1868    iN  M T

Mesene patawa Brévignon, 1995  TL: French Guiana

Mesene sardonyx Stichel, 1910  TL: Bolivia  M T

Mesene jimena Callaghan, Llorente & Luis, 2011  Zebra-tipped Metalmark  SW Mexico  M S

Mesene margaretta (A. White, 1843)  Zebra-tipped Metalmark    iN

Mesene margaretta El Salvador segregate ,  El Salvador  M S1 S2

Mesene margaretta margaretta (A. White, 1843)  E Mexico to Colombia  M T S1 S2 L

Mesene margaretta anartia J. Hall & Lamas, 2007  TL: Peru

Mesene silaris Godman & Salvin, 1878  Yellow Metalmark  S Mexico to Peru, Venezuela, Guianas  iN  M T S L V

Mesene leucopus Godman & Salvin, 1886  White-legged Metalmark  S Mexico & Guatemala  iN  M T S L

Mesene gracielae Dolibaina, Dias, O. Mielke & Casagrande, 2016  TL: Brazil

Mesenopsis  Godman & Salvin, 1886  

Mesenopsis bryaxis (Hewitson, 1870)  Yellow-centered Metalmark  Nicaragua to Colombia  iN  M T S

Mesenopsis melanochlora (Godman & Salvin, 1878)  Orange-striped Metalmark  Nicaragua to Bolivia  iN  M T S I

Mesenopsis briseis Godman & Salvin, 1886    iN  M T S

Mesenopsis pulchella Godman, 1903  TL: Brazil (Am)  M T

Mesenopsis albivitta (Lathy, 1904)  TL: Brazil (ES)  M T S

Mesenopsis jordana Dias, Dolibaina, O. Mielke & Casagrande, 2013    iN  M S

Mesenopsis tricolor (Hewitson, 1867)  Tricolored Metalmark  [= hademanni], S Mexico to Colombia  iN  M T S1 S2 L I1 I2

Xynias  Hewitson, 1874  

Xynias lithosina (H. Bates, 1868)    iN

Xynias lithosina lithosina (H. Bates, 1868)  TL: Brazil (Am)  M T S

Xynias lithosina christalla Grose-Smith, 1902  TL: Ecuador  M T

Xynias lithosina cynosema Hewitson, 1874  TL: Bolivia  M T S

Xynias lithosina potaronus Kaye, 1919  TL: Guyana  M T S

Xynias lilacina Lathy, 1932  TL: Peru  M T

Xenandra  C. Felder & R. Felder, 1865  

Xenandra helius (Cramer, 1779)  Red-striped Metalmark    iN

Xenandra helius helius (Cramer, 1779)  Costa Rica to Colombia  M T S

Xenandra helius cruentata Stichel, 1909  TL: Peru  M T

Xenandra nigrivenata Schaus, 1913  Black-veined Metalmark  Costa Rica  M T S

Xenandra vulcanalis Stichel, 1910    iN  M T

Xenandra desora Schaus, 1928  Red-patched Metalmark  Costa Rica & Panama  iN  M T S

Xenandra poliotactis (Stichel, 1910)  TL: Peru  M T S

Xenandra pelopia (H. Druce, 1890)    iN  M T S

Xenandra agria (Hewitson, [1853])    iN  M T S

Xenandra ahrenholzi J. Hall & Willmott, 2007  ref.224  M OI

Symmachia  Hübner, [1819]  

Symmachia championi Godman & Salvin, 1886  Red-costa Metalmark  S Mexico & Guatemala  iN  M T S L

Symmachia belti Godman & Salvin, 1886  TL: Nicaragua  M T

Symmachia probetor (Stoll, 1782)  Red-costa Metalmark  Nicaragua to Colombia  iN  M T S L I

Symmachia praxila Westwood, 1851    iN  M T

Symmachia falcistriga Stichel, 1910    iN  M T S

Symmachia meyi Brévignon, 1998  TL: French Guiana

Symmachia menetas (Drury, 1782)    iN

Symmachia menetas menetas (Drury, 1782)  TL: Brazil

Symmachia menetas eurina Schaus, 1902  TL: Brazil (Pr)  M T S

Symmachia maeonius Staudinger, 1888  TL: Brazil (Am); Peru  M T

Symmachia rubina H. Bates, 1866  Rubina Metalmark    iN

Symmachia rubina rubina H. Bates, 1866  S Mexico to Colombia  M T S1 S2 L I

Symmachia rubina separata Lathy, 1932  TL: Bolivia; Brazil (MT)  M T

Symmachia technema Stichel, 1910  TL: Surinam; Trinidad  M T S

Symmachia suevia Hewitson, 1877  TL: Ecuador  M T S

Symmachia leena Hewitson, 1870  Rusted Metalmark    iN

Symmachia leena leena Hewitson, 1870  Nicaragua to Panama  M T S

Symmachia leena harmodius Godman & Salvin, 1886  TL: Colombia  M T

Symmachia leena punctata A. Butler, 1877  TL: Brazil (Am)  M T

Symmachia leena harveyi Brévignon, 1998  TL: French Guiana

Symmachia multesima Stichel, 1910    iN  M T S

Symmachia aconia Hewitson, 1876  TL: Brazil  M T

Symmachia calliste Hewitson, 1867  TL: Brazil (Pa)  M T S

Symmachia pardalia Stichel, 1924    iN  M T S

Symmachia tigrina Hewitson, 1867  TL: Brazil (Pa)  M T

Symmachia accusatrix Westwood, 1851  Accused Metalmark  Mexico to Colombia, Ecuador, Brazil  iN  M T S L

Symmachia arcuata Hewitson, 1867  TL: Brazil (Am)  M T

Symmachia threissa Hewitson, 1870  White-notched Metalmark    iN

Symmachia threissa threissa Hewitson, 1870  Nicaragua to Colombia  M T S

Symmachia threissa seducta Brévignon, 1993  TL: French Guiana

Symmachia hetaerina Hewitson, 1867  TL: Brazil (Pa)

Symmachia hetaerina hetaerina Hewitson, 1867  TL: Brazil (Pa)  M T

Symmachia hetaerina hesione Stichel, 1910  TL: Brazil (Am); Peru  M T

Symmachia jugurtha Staudinger, [1887]  TL: Colombia  M T S

Symmachia miron Grose-Smith, 1898    iN

Symmachia miron miron Grose-Smith, 1898  TL: Ecuador  M T

Symmachia miron pulchellita Brévignon & Gallard, 1992  TL: French Guiana  M T S

Symmachia titiana Hewitson, 1870    iN

Symmachia titiana occidentalis Salazar, G. Rodríguez & Constantino, 2011  TL: Colombia

Symmachia titiana titiana Hewitson, 1870  TL: Ecuador  M T S

Symmachia aureus Attal, 2009  TL: Ecuador

Symmachia aureus aureus Attal, 2009  TL: Ecuador  M T S

Symmachia aureus mallqui Attal & Büche, 2015  TL: Peru  M S

Symmachia aureus niger Attal & Büche, 2013  TL: Peru

Symmachia aurigera (Weeks, 1902)  TL: Colombia  M T S

Symmachia juratrix Westwood, 1851  TL: Brazil (Pa)  M T S

Symmachia leopardinum (C. Felder & R. Felder, 1865)  TL: Brazil (Ba)  M T S

Symmachia hippea Herrich-Schäffer, [1853]  TL: Surinam  M T S

Symmachia batesi (Staudinger, [1887])    iN  M T S

Symmachia norina Hewitson, 1867  TL: Brazil (Pa)  M T S

Symmachia nemesis Le Cerf, 1958    iN

Symmachia fassli J. Hall & Willmott, 1995  TL: Ecuador  M T S

Symmachia triangularis (Thieme, 1907)    iN  M T

Symmachia calligrapha Hewitson, 1867  TL: Brazil (Pa)  M T S

Symmachia eraste (H. Bates, 1868)  TL: Brazil (Am)  M T S

Symmachia calderoni J. Hall & Lamas, 2001  TL: Peru  M T S

Symmachia basilissa (H. Bates, 1868)    iN

Symmachia basilissa basilissa (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Symmachia basilissa brevignoni Gallard, 2008  TL: French Guiana

Symmachia basilissa pujarnii (P. Jauffret & Martins, 2006)  TL: Brazil

Symmachia basilissa paracatuensis Callaghan, 2001  TL: Brazil (MG)  M T

Symmachia splendida (Salazar & Constantino, 1993)  TL: Colombia  M T

Symmachia hazelana J. Hall & Willmott, 1996  TL: Ecuador  M T

Symmachia almeidai (J. Zikán, 1946)  TL: Brazil (RJ)  M T

Symmachia sepyra (Hewitson, 1877)  TL: Ecuador  M T

Symmachia busbyi J. Hall & Willmott, 2007  ref.224  M OI S

Symmachia emeralda J. Hall & Willmott, 2007  ref.224  M OI

Symmachia pena J. Hall & Lamas, 2007    iN  M S

Symmachia mielkei (J. Hall & Furtado, 1999)  TL: Brazil (MT)

Symmachia atlantica Dias, Dolibaina & O. Mielke, 2020    iN

Symmachia divisora Dolibaina & Leite, 2012  TL: Brazil

Symmachia estellina Gallard, 2008    iN  M T

Symmachia poirieri Gallard, 2009  TL: French Guiana  M T

Symmachia rosanti Gallard, 2008  TL: French Guiana  M T

Symmachia uirassu Dias, Dolibaina & Casagrande, 2020  TL: Brazil

Stichelia  J. Zikán, 1949  

Stichelia bocchoris (Hewitson, 1876)    iN  M T S

Stichelia cuneifascia (J. Zikán, 1946)    iN  M T

Stichelia dukinfieldia (Schaus, 1902)    iN  M T S

Stichelia pelotensis Biezanko, O. Mielke & Wedderhoff, 1979    iN  M T

Stichelia catarinensis Dias, Dolibaina, O. Mielke & Casagrande, 2013  TL: Brazil

Tigria  Grishin, 2021  

Tigria xypete (Hewitson, 1870)  Tiger-striped Metalmark  Nicaragua to Panama; ref.12  iN  M T S1 S2 I

Tigria rubrica (Stichel, 1929)  Colombia & Ecuador; ref.12  M T S

Tigria phaedra (H. Bates, 1868)  ref.12

Tigria phaedra phaedra (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Tigria phaedra asclepia (Hewitson, 1870)  TL: Ecuador  M T S

Tigria elinas (Rebillard, 1958)  ref.12  M T S

Asymma  Grishin, 2021  

Asymma pardalis (Hewitson, 1867)  ref.12  iN  M T S

Asymma hippodice (Godman, 1903)  ref.12  M T S

Asymma arion (C. Felder & R. Felder, 1865)  ref.12  iN  M T

Asymma virgatula (Stichel, 1910)  Golden-red Metalmark  S Mexico (Chiapas) to W Ecuador; ref.12

Asymma virgatula virgatula (Stichel, 1910)  TL: Colombia  M T S

Asymma virgatula urichi (Kaye, 1925)  TL: Trinidad  M T S

Asymma virgaurea (Stichel, 1910)  ref.12  iN

Asymma virgaurea virgaurea (Stichel, 1910)  TL: Colombia  M T

Asymma virgaurea caucaensis (Salazar & Constantino, 2000)  TL: Colombia  M T S

Asymma virgaurea souadae (Benmesbah & Costa, 2020)  TL: Venezuela  M T

Asymma stigmosissima (Stichel, 1910)  ref.12

Asymma stigmosissima eddyi (Gallard, 2008)  TL: French Guiana  M T S

Asymma stigmosissima stigmosissima (Stichel, 1910)  TL: Bolivia  M T S

Asymma satana (J. Hall & Harvey, 2007)  ref.222,12  M S

Asymma giffordi (P. Jauffret & J. Jauffret, 2010)  ref.12

Pirascca  J. Hall & Willmott, 1996  

Pirascca apolecta (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Pirascca histrica (Stichel, 1910)  TL: Panama  M T S I1 I2

Pirascca arbuscula (Möschler, 1883)  Arbuscula Metalmark  TL: Surinam

Pirascca arbuscula arbuscula (Möschler, 1883)  Panama to Guianas  M T

Pirascca arbuscula mandosa (H. Druce, 1904)  TL: Brazil (RJ)  M T S

Pirascca crocostigma (H. Bates, 1868)  TL: Brazil (Am)  M T S

Pirascca hanneri Gallard, Viloria, Fratello & Costa, 2017  ref.190  M T S

Pirascca iasis (Godman, 1903)    iN  M T S

Pirascca phoenicura (Godman & Salvin, 1886)  Dingy Metalmark  Nicaragua to Colombia  iN  M T S

Pirascca pluto (Stichel, 1911)    iN  M T S

Pirascca polemistes J. Hall & Willmott, 1996  TL: Ecuador  M T S

Pirascca sagaris (Cramer, 1775)    iN

Pirascca sagaris sagaris (Cramer, 1775)  TL: Surinam  M T S Txt

Pirascca sagaris amazonica (Stichel, 1925)  TL: Brazil (Pa)  M T

Pirascca sagaris phrygiana (Stichel, 1916)  TL: Brazil (SP)  M T

Pirascca sagaris satnius (Dalman, 1823)  TL: Brazil  M T S

Pirascca sticheli (Lathy, 1932)  TL: Ecuador

Pirascca sticheli sticheli (Lathy, 1932)  TL: Ecuador  M T

Pirascca sticheli kawensis Brévignon, 1998  TL: French Guiana

Pirascca suapure (Weeks, 1906)  TL: Venezuela

Pirascca suapure suapure (Weeks, 1906)  TL: Venezuela  M T

Pirascca suapure vittata (Stichel, 1910)  TL: Brazil (Pa)  M T

Pirascca tyriotes (Godman & Salvin, 1878)  Golden-banded Metalmark  S Mexico to Brazil, Trinidad  iN  M T S1 S2 I1 I2

Pirascca patriciae J. Hall & Willmott, 2007  ref.224  M OI

Pirascca mimica Gallard & Fernandez, 2012  TL: French Guiana

Pirascca pedias (Godman, 1903)    iN  M T S

Pterographium  Stichel, 1910  

Pterographium sicora (Hewitson, 1875)  TL: Brazil (ES)  M T S

Pterographium semiota (H. Bates, 1868)  TL: Brazil (Pa)

Pterographium semiota remedios (Vargas & Salazar, 2018)  TL: Colombia

Pterographium semiota semiota (H. Bates, 1868)  TL: Brazil (Pa)  M T S L

Pterographium elegans (Schaus, 1920)  Fire-banded Metalmark  S Mexico (Chiapas) to Panama  iN  M T S

Pterographium thyatira (Hewitson, [1853])    iN  M T S

Pterographium inaria (Westwood, 1851)    iN  M T S

Pterographium interrupta (Lathy, 1932)  TL: French Guiana  M T S

Pterographium hypochloris (H. Bates, 1868)  ref.12  iN  M S

Phaenochitonia  Stichel, 1910  

Phaenochitonia florus (Fabricius, 1793)  ref.12  iN  M T S

Phaenochitonia pyrsodes (H. Bates, 1868)  TL: Brazil (Am)  M T S

Phaenochitonia cingulus (Stoll, 1790)    iN  M T S

Phaenochitonia fuliginea (H. Bates, 1868)    iN  M T S

Phaenochitonia gallardi J. Hall & Willmott, 1996  TL: Venezuela  M T

Phaenochitonia pseudodebilis J. Hall & Willmott, 1996  TL: Brazil (MT)  M T S

Phaenochitonia ignipicta Schaus, 1913  Ignipicta Metalmark  Costa Rica & Panama  M T S

Phaenochitonia tale (Hübner, [1819])  TL: Surinam

Phaenochitonia irroratum (Godman, 1903)  ref.12  M T S

Phaenochitonia exigua (H. Bates, 1868)  ref.12  M T S

Phaenochitonia apachita (J. Hall & Willmott, 1996)  ref.12  M T S

Phaenochitonia clarissa (Sharpe, 1890)  ref.12  M T S

Phaenochitonia debilis (H. Bates, 1868)  ref.12  M T

Phaenochitonia eanes (Godman, 1903)  ref.12  M T S

Phaenochitonia ignicauda (Godman & Salvin, 1878)  Red-stained Metalmark  ref.12  iN  M T S

Phaenochitonia sophistes (H. Bates, 1868)  ref.12  iN

Phaenochitonia sophistes sophistes (H. Bates, 1868)  TL: Brazil (Am/Pa)  M T S

Phaenochitonia sophistes baba (Brévignon, 2013)  TL: French Guiana

Argyrogrammana  Strand, 1932  

Argyrogrammana holosticta (Godman & Salvin, 1878)  Dotted Metalmark  S Mexico to Peru  M T S L

Argyrogrammana stilbe (Godart, [1824])    iN  M T S

Argyrogrammana sublimis Brévignon & Gallard, 1995    iN  M S

Argyrogrammana venilia (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Argyrogrammana crocea (Godman & Salvin, 1878)  Crocea Metalmark  Costa Rica & Panama  iN  M T S V

Argyrogrammana occidentalis (Godman & Salvin, 1886)  TL: Colombia  M T

Argyrogrammana janzeni J. Hall, 2023  TL: Costa Rica  M T

Argyrogrammana eura J. Hall, 2023  TL: French Guiana  M T

Argyrogrammana placibilis (Stichel, 1910)    iN  M T S

Argyrogrammana barine (Staudinger, [1887])  Blue-studded Metalmark  Costa Rica to Colombia  iN  M T S

Argyrogrammana amalfreda (Staudinger, [1887])    iN  M T

Argyrogrammana alstonii (Smart, 1979)    iN  M T

Argyrogrammana nurtia (Stichel, 1911)  TL: Bolivia; Peru

Argyrogrammana nurtia nurtia (Stichel, 1911)  TL: Bolivia; Peru  M T

Argyrogrammana nurtia ludibunda Brévignon & Gallard, 1995  TL: French Guiana

Argyrogrammana physis (Stichel, 1911)    iN  M T

Argyrogrammana phyton (Stichel, 1911)  TL: Brazil (Am)  M T S

Argyrogrammana praestigiosa (Stichel, 1929)    iN  M T

Argyrogrammana astuta Grishin, 2023  ref.146  M T

Argyrogrammana pulchra (Talbot, 1929)  TL: Colombia  M T

Argyrogrammana sticheli (Talbot, 1929)    iN  M T

Argyrogrammana caesarion Lathy, 1958    iN  M T S

Argyrogrammana trochilia (Westwood, 1851)    iN  M T S

Argyrogrammana leptographia (Stichel, 1911)  Blue-stained Metalmark  Costa Rica to Colombia  iN

Argyrogrammana leptographia leptographia (Stichel, 1911)  Costa Rica to Colombia  M T S

Argyrogrammana leptographia magdalenae Constantino, G. Rodríguez & Salazar, 2012  TL: Colombia  M T L

Argyrogrammana rameli (Stichel, 1930)    iN  M S

Argyrogrammana saphirina (Staudinger, [1887])  TL: Colombia  M T S

Argyrogrammana glaucopis (H. Bates, 1868)    iN

Argyrogrammana glaucopis glaucopis (H. Bates, 1868)  TL: Brazil (Am)  M T S V

Argyrogrammana glaucopis virgata Brévignon & Gallard, 1995  TL: French Guiana

Argyrogrammana caerulea J. Hall, 2023  TL: Peru  M T

Argyrogrammana subota (Hewitson, 1877)  TL: Ecuador  M T

Argyrogrammana sebastiani Brévignon, 1995  TL: French Guiana  M T

Argyrogrammana denisi Gallard, 1995  TL: French Guiana  M T

Argyrogrammana chicomendesi Gallard, 1995  TL: French Guiana  M T

Argyrogrammana natalita J. Hall & Willmott, 1995    iN  M T S

Argyrogrammana cosnipata J. Hall, 2023  TL: Peru  M S

Argyrogrammana pacsa J. Hall & Willmott, 1998  TL: Ecuador  M T

Argyrogrammana caelestina J. Hall & Willmott, 1995  TL: Ecuador  M T

Argyrogrammana aparamilla J. Hall & Willmott, 1995  TL: Ecuador  M T

Argyrogrammana celata J. Hall & Willmott, 1995  TL: Ecuador  M T

Argyrogrammana bonita J. Hall & Willmott, 1995    iN  M T S

Argyrogrammana johannismarci Brévignon, 1995    iN  M T S

Argyrogrammana pastaza J. Hall & Willmott, 1996    iN  M T S

Argyrogrammana talboti Brévignon & Gallard, 1998    iN

Argyrogrammana talboti talboti Brévignon & Gallard, 1998  TL: French Guiana  M T

Argyrogrammana talboti naranjilla J. Hall & Furtado, 1999  TL: Brazil (Am)  M T

Argyrogrammana brevignoni Dolibaina & Dias, 2015  TL: Brazil

Argyrogrammana danieli P. Jauffret & Martins, 2006  TL: Brazil

Argyrogrammana francozi Gallard, 2017  TL: French Guiana

Argyrogrammana gallardi Dolibaina & Dias, 2015  TL: Brazil

Argyrogrammana halli Dolibaina & Dias, 2015  TL: Brazil

Argyrogrammana saulensis Gallard, 2008  TL: French Guiana

Argyrogrammana saulensis saulensis Gallard, 2008  TL: French Guiana  M T

Argyrogrammana saulensis tunari Gallard, 2008  TL: Bolivia

Argyrogrammana willmotti Dolibaina & Dias, 2015  TL: Brazil

Argyrogrammana janiceae Ahrenholz & J. Hall, 2023  TL: Ecuador  M S

  Tribe Helicopini

Helicopis  Fabricius, 1807  

Helicopis gnidus (Fabricius, 1787)    iN

Helicopis gnidus gnidus (Fabricius, 1787)  TL: Surinam  M S

Helicopis gnidus beaulieui Le Moult, 1939  TL: ?  M T

Helicopis gnidus galatea Stichel, 1919  TL: Brazil (Pa)  M T1 T2

Helicopis gnidus interrupta Le Moult, 1939  TL: Peru  M T

Helicopis gnidus medialis Schaus & T. Cockerell, 1923  TL: Colombia  M T

Helicopis gnidus nigrobasalis Aurivillius, 1929  TL: Brazil (Am)  M T

Helicopis gnidus obidonus Le Moult, 1939  TL: Brazil (Pa)  M T

Helicopis cupido (Linnaeus, 1758)    iN

Helicopis cupido cupido (Linnaeus, 1758)  TL: America  M T1 T2 S HS

Helicopis cupido belemensis Le Moult, 1940  TL: Brazil (Pa)  M T

Helicopis cupido erotica Seitz, 1913  TL: Brazil (Pa)  M T1 T2 T3 T4 S

Helicopis cupido gradiva Stichel, 1919  TL: Brazil (Am)  M T

Helicopis cupido incerta Meier-Ramel, 1928  TL: Brazil (Am)  M T1 T2 T3 T4 T5 L

Helicopis cupido lindeni Grote, 1874  TL: Brazil (Pa)  M T

Helicopis cupido nigra N. Riley, 1919  TL: Brazil (Am)  M T

Helicopis cupido pseudolindeni Le Moult, 1940  TL: Brazil (Ap)  M T

Helicopis cupido trinitatis Seitz, 1913  TL: Trinidad  M T S

Helicopis endymiaena (Hübner, [1819])    iN

Helicopis endymiaena endymiaena (Hübner, [1819])  TL: Surinam  M T S

Helicopis endymiaena elegans Kaye, 1904  TL: Trinidad  M T

Sarota  Westwood, 1851  

Sarota subtessellata (Schaus, 1913)  Regal Sarota  Costa Rica to W Colombia  iN  M T S

Sarota chrysus (Stoll, 1782)  Stoll's Sarota  SE Mexico to Bolivia & Brazil, Guianas, Trinidad  iN  M T S L

Sarota neglecta Stichel, 1910  Neglected Sarota  Costa Rica to W Ecuador & NW Venezuela  iN  M T S L

Sarota chocoensis J. Hall, 1998  Choco Sarota  C Panama to W Ecuador  iN  M T S

Sarota turrialbensis (Schaus, 1913)  Turrialba Sarota  Turrialba, Costa Rica  M T S

Sarota craspediodonta (Dyar, 1918)  Veracruzan Sarota  S Mexico & Guatemala  M T S

Sarota spicata (Staudinger, 1888)    iN  M T S

Sarota willmotti J. Hall, 1998    iN  M T S

Sarota acanthoides (Herrich-Schäffer, [1853])    iN  M T S

Sarota harveyi J. Hall, 1998  TL: Brazil (MT)  M T S

Sarota psaros Godman & Salvin, 1886  Guatemalan Sarota    iN

Sarota psaros psaros Godman & Salvin, 1886  S Mexico to Panama, Guianas  M T S L

Sarota psaros albidisca (Lathy, 1932)  TL: Peru  M T

Sarota psaros punctinegris Gallard, 2017  TL: French Guiana

Sarota psaros psaronius (Stichel, 1911)  TL: Brazil (ES)  M T

Sarota chloropunctata J. Hall, 1998  TL: Ecuador  M T

Sarota estrada Schaus, 1928  Schaus' Sarota    iN

Sarota estrada estrada Schaus, 1928  S Mexico to Costa Rica  M T S L

Sarota estrada sabanilla J. Hall, 1998  TL: Ecuador  M T S

Sarota gamelia Godman & Salvin, 1886  Panamanian Sarota    iN

Sarota gamelia gamelia Godman & Salvin, 1886  S Mexico to W Ecuador  M T S

Sarota gamelia alba J. Hall, 1998  TL: Peru  M T

Sarota acantus (Stoll, 1782)  Simple Sarota  Honduras to W Ecuador, Venezuela, Peru, Guianas, Trinidad  iN  M T S L I

Sarota miranda Brévignon, 1998    iN  M S

Sarota gyas (Cramer, 1775)  Guyanan Sarota  [Mexico to Panama?], Colombia to Peru, Brazil, Guianas  iN  M T S L

Sarota myrtea Godman & Salvin, 1886  Godman's Sarota  E Mexico to W Ecuador, Venezuela to Argentina  iN  M T S L

Sarota flavicincta (Lathy, 1932)  TL: Peru  M T S

Sarota completa J. Hall, 1998    iN  M T S

Sarota karishmae Brévignon, 2016  TL: French Guiana

Ourocnemis  Bethune-Baker, 1887  

Ourocnemis archytas (Stoll, 1787)  False Anteros  Panama, Surinam, E Ecuador, C Brazil, Paraguay  iN  M OI S L

Ourocnemis axiochus (Hewitson, 1867)  TL: Brazil  M T OI S

Ourocnemis boulleti Le Cerf, 1911    iN  M T

Ourocnemis carausius (Westwood, 1851)  Carousing Anteros  ref.12  iN

Ourocnemis carausius carausius (Westwood, 1851)  E & W Mexico to Colombia  M T S1 S2 L I

Ourocnemis carausius yurakchupa (J. Hall & Willmott, 1998)  TL: Ecuador  M T

Ourocnemis principalis (Hopffer, 1874)  ref.12  iN  M T S

Ourocnemis renaldus (Stoll, 1790)  ref.12  iN

Ourocnemis renaldus renaldus (Stoll, 1790)  TL: Surinam  M S

Ourocnemis renaldus indigator (Stichel, 1911)  Nicaragua to Panama  M T S1 S2 L I

Ourocnemis renaldus notius (Stichel, 1911)  TL: Brazil (SC)  M T

Ourocnemis aerosus (Stichel, 1924)  ref.12  iN  M T S

Anteros  Hübner, [1819]  

Anteros allectus Westwood, 1851  Rounded Anteros  [= cumulatus], Costa Rica to W Ecuador  iN

Anteros allectus allectus Westwood, 1851  [= cumulatus], Costa Rica to W Ecuador  M T S L I

Anteros allectus magnificus Brévignon, 2015  TL: French Guiana

Anteros roratus Godman & Salvin, 1886  Elegant Anteros  S Mexico to Panama  iN  M T S I

Anteros chrysoprasta Hewitson, 1867    iN  M T S

Anteros zikani Siewert, Dias & Dolibaina, 2014  TL: Brazil

Anteros otho Westwood, 1851  TL: Brazil (Pa)  M T S

Anteros micon H. Druce, 1875  Black-bellied Anteros  S Mexico to Panama  iN  M T S1 S2 L I1 I2

Anteros formosus (Cramer, 1777)    iN  M T S

Anteros aliceae Dias & Siewert, 2014    iN  M S

Anteros theleia Stichel, 1910  TL: [Peru]  M T

Anteros ethani Dias, Dolibaina & O. Mielke, 2014  TL: Brazil

Anteros aurigans Gallard & Brévignon, 1989    iN  M T

Anteros lectabilis Stichel, 1909    iN  M T

Anteros gentilis (Rebillard, 1958)  TL: Peru

Anteros nubosus J. Hall & Willmott, 1995    iN  M T S

Anteros cruentatus Stichel, 1911    iN  M T S

Anteros acheus (Stoll, 1781)    iN

Anteros acheus acheus (Stoll, 1781)  TL: Surinam  M S

Anteros acheus ampyx (Drury, 1782)  TL: Brazil (RJ)

Anteros acheus troas Stichel, 1909  TL: Bolivia  M T S

Anteros kupris Hewitson, 1875  Giant Anteros    iN

Anteros kupris antioquensis G. Rodríguez, Constantino & Salazar, 2010  TL: Colombia  M T

Anteros kupris kupris Hewitson, 1875  Costa Rica to Ecuador  M T S L

Anteros kupris aureocultus Stichel, 1909  TL: Bolivia  M T S

Anteros bracteata Hewitson, 1867    iN

Anteros bracteata chocoensis Constantino, Salazar & G. Rodríguez, 2010  TL: Colombia

Anteros bracteata bracteata Hewitson, 1867  TL: Brazil (Am)  M T S

Anteros milae Brévignon, 2015  TL: French Guiana

Anteros pardalis Salazar, Constantino & G. Rodríguez, 2010  TL: Colombia

  Tribe Callistiumini

Callistium  Stichel, 1911  

Callistium cleadas (Hewitson, 1866)  Brazil; ref.12  iN  M T S

  Tribe Calydnini

Echydna  J. Hall, 2002  

Echydna chaseba (Hewitson, 1854)    iN  M T S

Echydna punctata (C. Felder & R. Felder, 1861)    iN  M T S

Echydna bojeto P. Jauffret & J. Jauffret, 2009  TL: Brazil

Calydna  E. Doubleday, 1847  

Calydna calamisa Hewitson, 1854    iN  M T S

Calydna caieta Hewitson, 1854    iN  M T S

Calydna charila Hewitson, 1854    iN  M T S

Calydna venusta Godman & Salvin, 1886  Venusta Metalmark    iN

Calydna venusta venusta Godman & Salvin, 1886  E & W Mexico to Panama  M T S

Calydna venusta morio Stichel, 1929  TL: Brazil (MT)  M T

Calydna stolata Brévignon, 1998  TL: French Guiana  M S

Calydna lusca (Geyer, [1835])    iN  M S

Calydna fissilisima J. Hall, 2002  TL: Brazil (RJ)  M T S

Calydna thersander (Stoll, 1780)    iN  M T S

Calydna jeannea J. Hall, 2002    iN  M S

Calydna nicolayi J. Hall, 2002    iN  M S

Calydna cea Hewitson, 1859    iN  M T S

Calydna sturnula (Geyer, 1837)  Sturnula Metalmark  [= hegias, = sinuata], E & W Mexico to Panama  iN  M T S1 S2 L I

Calydna micra H. Bates, 1868    iN  M T S

Calydna carneia Hewitson, 1859    iN  M T S

Calydna candace Hewitson, 1859  TL: Brazil (Am)  M T

Calydna hiria (Godart, [1824])    iN  M T S

Calydna catana Hewitson, 1859    iN  M T S

Calydna cabira Hewitson, 1854  TL: Brazil (Am)  M T S

Calydna arius (Cramer, 1775)  TL: Surinam

Calydna isala (Godart, [1824])  TL: S. Amer.

Calydna pichita J. Hall & Lamas, 2004  TL: Peru  M T

Echenais  Hübner, [1819]  

Echenais thelephus (Cramer, 1775)    iN

Echenais thelephus thelephus (Cramer, 1775)  TL: West Indies  M T S

Echenais thelephus meridionalis (Lathy, 1932)  TL: Brazil (MT)  M T

Echenais mycea (Hewitson, 1865)    iN

Echenais mycea mycea (Hewitson, 1865)  TL: Colombia  M T

Echenais mycea glaucosmia (Hewitson, 1870)  TL: Ecuador  M T S

Echenais aenetus (Hewitson, 1874)    iN  M T S

  Tribe Befrostiini

Befrostia  Grishin, 2019  

Befrostia elegia (Stichel, 1929)    iN  M T

Befrostia lalannei (Gallard, 2008)  TL: French Guiana  M T

  Tribe Sertaniini

Xanthosa  Grishin, 2019  

Xanthosa xanthosa (Stichel, 1910)  TL: ?  M T

Sertania  Callaghan & Kaminski, 2017  

Sertania guttata (Stichel, 1910)    iN  M T S

Sertania jaibensis (Callaghan & Soares, 2001)  TL: Brazil (MG)  M T

Sertania lambedor (P. Jauffret, J. Jauffret & Pessôa, 2008)    iN

  Tribe Riodinini

Baeotis  Hübner, [1819]  

Baeotis hisbon (Cramer, 1775)    iN  M T S

Baeotis bacaenis Hewitson, 1874    iN  M T S

Baeotis elegantula Hopffer, 1874    iN  M T S L

Baeotis bacaenita Schaus, 1902    iN  M T S L

Baeotis creusis Hewitson, 1874    iN  M T S L

Baeotis staudingeri D'Abrera, 1994    iN  M T S

Baeotis nesaea Godman & Salvin, 1889  Nesaea Metalmark  Costa Rica to Peru  iN  M T S L

Baeotis melanis Hübner, [1829]    iN

Baeotis euprepes (H. Bates, 1868)    iN

Baeotis euprepes euprepes (H. Bates, 1868)  TL: Brazil (Pa)  M T

Baeotis euprepes obscurior Brévignon, 1998  TL: French Guiana

Baeotis euprepes orthotaenia Seitz, 1916  TL: Bolivia  M T S

Baeotis prima (H. Bates, 1868)    iN

Baeotis prima prima (H. Bates, 1868)  TL: Brazil (Pa/Am)  M T S

Baeotis prima nigricans Brévignon, 1998  TL: French Guiana

Baeotis felix Hewitson, 1874    iN

Baeotis felix felix Hewitson, 1874  TL: Bolivia  M T S

Baeotis felix felicissima Thieme, 1907  TL: Ecuador  M T

Baeotis zonata R. Felder, 1869  Bumblebee Metalmark    iN

Baeotis zonata zonata R. Felder, 1869  E & W Mexico to Venezuela  M T S L

Baeotis zonata dryades Dognin, 1891  TL: Ecuador  M T S

Baeotis choroniensis Lichy, 1946  TL: Venezuela  M T S

Baeotis capreolus Stichel, 1910  TL: Colombia

Baeotis capreolus capreolus Stichel, 1910  TL: Colombia  M T

Baeotis capreolus robini Brévignon, 1995  TL: French Guiana  M T S

Baeotis johannae Sharpe, 1890    iN

Baeotis johannae johannae Sharpe, 1890  TL: Brazil (Go)  M T S

Baeotis johannae cearaica Seitz, 1916  TL: Brazil (Ce)  M T

Baeotis barce Hewitson, 1875  Barce Metalmark    iN

Baeotis barce barce Hewitson, 1875  S Mexico to Honduras  M T OI

Baeotis barce corentyna Brévignon, 1998  TL: Guyana

Baeotis sulphurea (R. Felder, 1869)  Sulphur Metalmark  E Mexico to Guatemala  iN  M T S L

Baeotis macularia (Boisduval, 1870)  Sulphur Metalmark  S Mexico (Chiapas) to Colombia  iN  M T S L

Baeotis kadenii (C. Felder & R. Felder, 1861)    iN  M T S

Baeotis attali J. Hall & Willmott, 1998    iN  M T

Baeotis cephissa (Hewitson, 1875)    iN  M T

Baeotis vicosa P. Jauffret, J. Jauffret & Pessôa, 2008  TL: Brazil

Baeotis johnstoni (Dannatt, 1904)  ref.12  iN  M T

Baeotis banghaasi (Seitz, 1917)  ref.12  iN  M T

Baeotis seitzi (Salazar, Constantino & G. Rodríguez, 2011)  ref.12  iN  M T

Dachetola  J. Hall, 2001  

Dachetola azora (Godart, [1824])    iN  M T S

Dachetola virido (Lathy, 1958)    iN

Dachetola virido klagesi P. Jauffret & J. Jauffret, 2008  TL: Brazil

Dachetola virido virido (Lathy, 1958)  TL: Bolivia  M T

Dachetola caligata (Stichel, 1911)  TL: Colombia  M T S

Dachetola pione (H. Bates, 1868)  TL: Brazil (Pa)  M T

Cariomothis  Stichel, 1910  

Cariomothis erythromelas (Sepp, [1840])    iN

Cariomothis erythromelas erythromelas (Sepp, [1840])  TL: Surinam  M S

Cariomothis erythromelas erythraea Stichel, 1910  TL: Brazil (ES)  M T

Cariomothis erythromelas fulvus Lathy, 1932  TL: Brazil (MT)  M T S

Cariomothis erythromelas suffusa Lathy, 1932  TL: Ecuador  M T

Cariomothis poeciloptera (Godman & Salvin, 1878)  Fiery Metalmark    iN

Cariomothis poeciloptera poeciloptera (Godman & Salvin, 1878)  Costa Rica & Panama  M T TI S L

Cariomothis poeciloptera melusina (Staudinger, 1888)  Panama & Colombia  M T S

Cariomothis erotylus Stichel, 1910    iN  M T

Cariomothis chia (Hübner, 1823)    iN  M S

Inkana  Grishin, 2021  

Inkana incoides (Schaus, 1902)  ref.12  iN  M T S

Inkana cutisca (J. Hall & Willmott, 1998)  TL: Ecuador  M T S

Pheles  Herrich-Schäffer, [1853]  

Pheles heliconides Herrich-Schäffer, [1853]    iN

Pheles heliconides heliconides Herrich-Schäffer, [1853]  TL: Surinam  M T S

Pheles heliconides rufotincta H. Bates, 1868  TL: Brazil (Am)  M T S

Pheles melanchroia (C. Felder & R. Felder, 1865)  Melancholy Metalmark  Mexico & Guatemala  iN  M T S

Pheles strigosus (Staudinger, 1876)  Strigosa Metalmark    iN

Pheles strigosus strigosus (Staudinger, 1876)  S Mexico to Colombia  M T OI OD S L

Pheles strigosus tristis (Schaus, 1902)  TL: Peru  M T

Pheles incerta Staudinger, [1887]  Incerta Metalmark  Panama to Peru  iN  M T S L

Pheles eulesca (Dyar, 1909)  Dyar's Metalmark  S Mexico to Panama  M T S

Pheles ochracea (Stichel, 1910)  Ochracea Metalmark  Panama to Venezuela  M T S

Pheles atricolor (A. Butler, 1871)    iN

Pheles atricolor atricolor (A. Butler, 1871)  TL: Brazil  M T S

Pheles atricolor malmanoury (Brévignon, 1998)  TL: French Guiana

Pheles obscura C. Mielke, Dolibaina & Dias, 2015  TL: Brazil  M OI

Pheles plaumanni Dolibaina & Dias, 2015    iN

Syrmatia  Hübner, [1819]  

Syrmatia nyx (Hübner, [1817])  Spotted Tailwing  Costa Rica to Brazil, Trinidad  iN  M T S

Syrmatia astraea Staudinger, 1888  TL: Colombia  M T

Syrmatia lamia H. Bates, 1868    iN  M T S

Syrmatia aethiops Staudinger, 1888  White-tipped Tailwing  Costa Rica to Brazil  M T S

Parcella  Stichel, 1910  

Parcella amarynthina (C. Felder & R. Felder, 1865)  Orange-banded Metalmark  Panama to Argentina  iN  M T S L

Notheme  Westwood, 1851  

Notheme erota (Cramer, 1780)    iN

Notheme erota erota (Cramer, 1780)  TL: Surinam  M T S

Notheme erota agathon (C. Felder & R. Felder, 1865)  TL: Brazil (Ba)  M T

Notheme erota angellus Stichel, 1910  TL: Paraguay  M T

Notheme erota diadema Stichel, 1910  S Mexico to Bolivia  M T S L

Notheme erota hemicosmeta Seitz, 1917  TL: Brazil (SP)  M T

Riodina  Westwood, 1851  

Riodina lysippus (Linnaeus, 1758)    iN

Riodina lysippus lysippus (Linnaeus, 1758)  TL: America  M T S

Riodina lysippus erratica Seitz, 1913  TL: [Brazil]  M T S

Riodina lysippus lysias Stichel, 1910  TL: Ecuador; Bolivia  M T S

Riodina lycisca (Hewitson, [1853])    iN

Riodina lycisca lycisca (Hewitson, [1853])  TL: Brazil (MG)  M T S

Riodina lycisca lysistratus Burmeister, 1878  TL: Argentina  M T S

Riodina lycisca pelta (Schaus, 1902)  TL: Brazil (RJ)  M T S

Riodina lysippoides Berg, 1882    iN  M T S

Siseme  Westwood, 1851  

Siseme alectryo Westwood, 1851    iN

Siseme alectryo alectryo Westwood, 1851  TL: Colombia  M T S

Siseme alectryo lucilius Hopffer, 1874  TL: Peru; Bolivia  M T

Siseme alectryo spectanda Stichel, 1909  TL: Ecuador  M T

Siseme alectryo tantilla Thieme, 1907  TL: Colombia  M T

Siseme peculiaris H. Druce, 1904  TL: Peru  M T

Siseme militaris (Hopffer, 1874)    iN  M T S

Siseme aristoteles (Latreille, [1809])    iN

Siseme aristoteles aristoteles (Latreille, [1809])  TL: Colombia  M T

Siseme aristoteles leucodesma Seitz, 1916  TL: Colombia  M T

Siseme aristoteles minerva C. Felder & R. Felder, 1865  TL: Colombia  M T S

Siseme aristoteles ochrotaenia Seitz, 1916  TL: Ecuador

Siseme aristoteles sprucei H. Bates, 1868  TL: Ecuador  M T S

Siseme pallas (Latreille, [1809])    iN

Siseme pallas pallas (Latreille, [1809])  TL: Colombia  M T S

Siseme pallas defasciata Lathy, 1932  TL: Peru  M T

Siseme pallas xanthogramma H. Bates, 1868  TL: Peru  M T

Siseme atrytone Thieme, 1907    iN  M T L

Siseme neurodes (C. Felder & R. Felder, 1861)    iN

Siseme neurodes neurodes (C. Felder & R. Felder, 1861)  TL: Venezuela; Ecuador; Peru  M T

Siseme neurodes caudalis H. Bates, 1868  TL: Peru  M T S L

Siseme pseudopallas Weymer, 1890    iN

Siseme pseudopallas pseudopallas Weymer, 1890  TL: Peru  M T S

Siseme pseudopallas hellotis Thieme, 1907  TL: Peru  M T

Monethe  Westwood, 1851  

Monethe alphonsus (Fabricius, 1793)    iN  M S

Monethe albertus C. Felder & R. Felder, 1862    iN

Monethe albertus albertus C. Felder & R. Felder, 1862  TL: [Peru]  M T S

Monethe albertus rudolphus Godman & Salvin, 1885  Honduras to Colombia  M T S L

Themone  Westwood, 1851  

Themone pais (Hübner, [1820])    iN

Themone pais pais (Hübner, [1820])  TL: ?  M T S V

Themone pais bambino Bryk, 1953  TL: Brazil (Am)  M T S

Themone pais carveri (Weeks, 1906)  TL: Venezuela  M T

Themone pais eutacta Stichel, 1910  TL: Surinam  M T

Themone pais paidion Stichel, 1910  TL: Peru  M T

Themone pais storthynga Stichel, 1910  TL: Peru  M T S

Themone pais sublimata Stichel, 1925  TL: Brazil (Am)  M T

Themone pais trivittata Lathy, 1904  TL: Peru  M T

Paraphthonia  Stichel, 1910  

Paraphthonia molione (Godman, 1903)  TL: Peru  M T

Paraphthonia cteatus Seitz, 1917    iN  M T

Paraphthonia diadocis (Stichel, 1910)  [formerly Lyropteryx]; ref.151  iN  M T S

Brachyglenis  C. Felder & R. Felder, 1862  

Brachyglenis esthema C. Felder & R. Felder, 1862  White-rayed Metalmark    iN

Brachyglenis esthema esthema C. Felder & R. Felder, 1862  Costa Rica & Colombia  M T S

Brachyglenis esthema ecuadorensis Lathy, 1932  TL: Ecuador  M T

Brachyglenis dodone (Godman & Salvin, 1886)  Orange-bellied Metalmark  S Mexico to Panama  M T S

Brachyglenis dinora (H. Bates, 1866)  Dinora Metalmark  Nicaragua to Colombia  iN  M T S I

Brachyglenis drymo (Godman & Salvin, 1886)    iN  M T S

Isapis  E. Doubleday, 1847  

Isapis agyrtus (Cramer, 1777)    iN

Isapis agyrtus agyrtus (Cramer, 1777)  TL: Surinam  M T

Isapis agyrtus abydus Stichel, 1929  TL: Brazil (SP)  M T

Isapis agyrtus falcis Weymer, 1890  TL: Colombia  M T S

Isapis agyrtus hera Godman & Salvin, 1886  S Mexico to Panama  M T S L

Isapis agyrtus mithrophorus Seitz, 1913  TL: Surinam  M T S

Isapis agyrtus praxinus (Stichel, 1909)  TL: Venezuela  M T

Isapis agyrtus sestus (Stichel, 1909)  TL: Brazil (Pa/Am); Peru  M T S

Isapis poecila (H. Bates, 1868)  [formerly Themone]; ref.151  iN  M T S

Isapis inornata (Lathy, 1958)  [not poecila]; ref.155  M T

Isapis pulcherrima (Herrich-Schäffer, [1853])  [formerly Themone]; ref.151  iN  M T

Melanis  Hübner, [1819]  

Melanis smithiae (Westwood, 1851)    iN

Melanis smithiae smithiae (Westwood, 1851)  TL: Brazil ([Pe])  M T S

Melanis smithiae xarifa (Hewitson, [1853])  TL: Venezuela  M T

Melanis alena (Hewitson, 1870)    iN  M T S

Melanis cinaron (C. Felder & R. Felder, 1861)    iN  M T S

Melanis pixe (Boisduval, 1836)  Pixie    iN

Melanis pixe pixe (Boisduval, 1836)  [= sexpunctata], S TX, E & W Mexico to Guatemala  M T S L1 L2 L3 I

Melanis pixe sanguinea (Stichel, 1910)  Honduras to Colombia  M T S1 S2 L I

Melanis pixe corvina (Stichel, 1910)  TL: Colombia  M T

Melanis pixe pseudoalena Salazar & Vargas, 2018  TL: Colombia

Melanis gynaeceas (Godman & Salvin, 1889)  Forgotten Metalmark  E & S Mexico (TAMPS, PUE, VER, GRO)  iN  M T S L Ha

Melanis unxia (Hewitson, [1853])    iN

Melanis unxia unxia (Hewitson, [1853])  TL: Brazil (RJ)  M T S

Melanis unxia ludmila (J. Zikán, 1952)  TL: Brazil (MG)  M T

Melanis yeda (J. Zikán, 1952)  TL: [Brazil (MG)]  M T S

Melanis vidali (Dognin, 1891)    iN  M T

Melanis passiena (Hewitson, 1870)    iN  M T S

Melanis volusia (Hewitson, [1853])    iN  M T S

Melanis herminae (J. Zikán, 1952)    iN  M T

Melanis electron (Fabricius, 1793)    iN

Melanis electron electron (Fabricius, 1793)  TL: French Guiana  M T

Melanis electron auriferax (Stichel, 1910)  TL: Brazil (SP)  M T

Melanis electron epijarbas (Staudinger, 1888)  TL: Brazil (Pa)  M T

Melanis electron herellus (Snellen, 1887)  TL: Curaçao  M T

Melanis electron melantho (Ménétriés, 1855)  Belize & Guatemala to Panama  M S

Melanis electron pronostriga (Stichel, 1910)  TL: Brazil (Pa); Colombia  M T S

Melanis electron rabuscula (Stichel, 1910)  TL: Brazil (Am); Peru  M T S

Melanis hillapana (Röber, 1904)    iN

Melanis hillapana hillapana (Röber, 1904)  TL: Peru  M T

Melanis hillapana corinna (J. Zikán, 1952)  TL: Brazil (MG)  M T

Melanis hillapana cratippa (Seitz, 1913)  TL: Brazil (Ma/MT)  M T S

Melanis melandra Hübner, [1819]    iN  M T S

Melanis boyi (Stichel, 1923)  TL: Brazil (Pa)  M T

Melanis opites (Hewitson, 1875)  TL: Brazil  M T S

Melanis leucophlegma (Stichel, 1910)    iN  M T S L

Melanis aegates (Hewitson, 1874)    iN

Melanis aegates aegates (Hewitson, 1874)  TL: Bolivia  M T

Melanis aegates albugo (Stichel, 1910)  TL: Paraguay  M T S

Melanis aegates araguaya (Seitz, 1913)  TL: Brazil (Go)  M T

Melanis aegates cretiplaga (Stichel, 1910)  TL: Argentina  M T S

Melanis aegates lilybaeus (Stichel, 1926)  TL: Brazil (MG)  M T

Melanis aegates limbata (Stichel, 1925)  TL: Brazil (MG)  M T

Melanis aegates melliplaga (Stichel, 1909)  TL: French Guiana; Surinam  M T S

Melanis cephise (Ménétriés, 1855)  White-rayed Metalmark  W Mexico to Costa Rica  iN  M S L

Melanis acroleuca (R. Felder, 1869)  White-tipped Metalmark    iN

Melanis acroleuca huasteca J. White & A. White, 1989  E Mexico  M S L

Melanis acroleuca acroleuca (R. Felder, 1869)  W Mexico  M T S L

Melanis lycea Hübner, 1823    iN  M T S

Melanis lioba (J. Zikán, 1952)    iN  M T

Melanis xenia (Hewitson, [1853])    iN

Melanis xenia xenia (Hewitson, [1853])  TL: Brazil (MG)  M T S

Melanis xenia ambryllis (Hewitson, 1874)  TL: Bolivia  M T S

Melanis marathon (C. Felder & R. Felder, 1865)    iN

Melanis marathon marathon (C. Felder & R. Felder, 1865)  TL: Colombia  M T S

Melanis marathon assimulata (Stichel, 1910)  TL: Colombia  M T

Melanis marathon charon (A. Butler, 1874)  TL: Brazil (RJ)  M T

Melanis marathon stenotaenia (Röber, 1904)  TL: Peru  M T

Melanis seleukia (Stichel, 1910)    iN  M T S IL Txt

Melanis cercopes (Hewitson, 1874)    iN  M T S

Melanis cratia (Hewitson, 1870)  TL: ?  M T S

Melanis hodia (A. Butler, 1870)    iN  M T S

Melanis caatingensis (Callaghan & Nobre, 2014)  TL: Brazil

Melanis dulcis Callaghan & Costa, 2017  TL: Venezuela  M T S

Melanis weissi Brévignon, 2019  TL: Brazil

Lyropteryx  Westwood, 1851  [includes Necyria as a subgenus]; ref.151

Lyropteryx apollonia Westwood, 1851    iN

Lyropteryx apollonia apollonia Westwood, 1851  TL: Brazil (Am)  M T S

Lyropteryx apollonia diana Stichel, 1910  TL: Colombia  M T

Lyropteryx apollonia sparsa Stichel, 1924  TL: Brazil (MT)  M T

Lyropteryx lyra Saunders, 1859    iN

Lyropteryx lyra lyra Saunders, 1859  TL: Colombia  M T S

Lyropteryx lyra cleadas H. Druce, 1875  SE Mexico to Panama  M T S I

Lyropteryx terpsichore Westwood, 1851    iN

Lyropteryx terpsichore terpsichore Westwood, 1851  TL: Brazil  M S L

Lyropteryx terpsichore zygaena Stichel, 1910  TL: Paraguay  M T S

Lyropteryx melaniae (Stichel, 1930)  [formerly Melanis]; ref.151  iN  M T

Lyropteryx bellona (Westwood, 1851)    iN

Lyropteryx bellona bellona (Westwood, 1851)  TL: [Bolivia]  M T S L

Lyropteryx bellona ahrenholzi (Lamas, 2003)  TL: Peru  M T

Lyropteryx bellona gerhardi (Mengel, 1902)  TL: Peru  M T

Lyropteryx bellona juturna (Hewitson, 1869)  TL: Colombia  M T S

Lyropteryx bellona manco (Saunders, 1859)  TL: Colombia  M T S

Lyropteryx bellona saundersii (Hewitson, 1854)  TL: Colombia  M T

Lyropteryx bellona westwoodi (Hopffer, 1874)  TL: Peru  M T S

Lyropteryx bellona whitelyiana (H. Druce, 1874)  TL: Peru  M T S

Lyropteryx bellona zaneta (Hewitson, 1875)  TL: Ecuador  M T S

Lyropteryx ingaretha (Hewitson, 1872)  Orange-spotted Metalmark  Nicaragua & Costa Rica  M T OI S

Lyropteryx duellona (Westwood, 1851)    iN

Lyropteryx duellona duellona (Westwood, 1851)  TL: Ecuador  M T S

Lyropteryx duellona beltiana (Hewitson, 1870)  Nicaragua to Panama  M T TI S L I

Lyropteryx duellona diva (Staudinger, [1887])  TL: Colombia  M T

Lyropteryx larunda (Godman & Salvin, 1885)  Larunda Metalmark  Chiapas & Guatemala  iN  M S

Ancyluris  Hübner, [1819]  

Ancyluris meliboeus (Fabricius, 1777)    iN

Ancyluris meliboeus meliboeus (Fabricius, 1777)  TL: Surinam  M T S

Ancyluris meliboeus eudaemon Stichel, 1910  TL: Peru; Colombia; Bolivia  M T S

Ancyluris meliboeus julia (Saunders, 1850)  TL: Brazil (Pa)  M T S

Ancyluris miniola (H. Bates, 1868)    iN  M T S

Ancyluris rubrofilum Stichel, 1909    iN  M T

Ancyluris etias (Saunders, 1859)    iN

Ancyluris etias etias (Saunders, 1859)  TL: Peru  M T S

Ancyluris etias gracilis Stichel, 1910  TL: Surinam  M T HS V

Ancyluris etias mendita (H. Druce, 1904)  TL: Bolivia  M T

Ancyluris melior Stichel, 1910  TL: Brazil (Am); Peru  M T S

Ancyluris pomposa Stichel, 1910  TL: Brazil (Am)  M T

Ancyluris paramba D'Abrera, 1994    iN  M T S

Ancyluris paetula Stichel, 1916  TL: Peru  M T

Ancyluris tedea (Cramer, 1777)    iN

Ancyluris tedea tedea (Cramer, 1777)  TL: Surinam  M S

Ancyluris tedea silvicultrix Stichel, 1909  TL: Peru; Bolivia  M T S

Ancyluris aulestes (Cramer, 1777)    iN

Ancyluris aulestes aulestes (Cramer, 1777)  TL: Surinam  M T S

Ancyluris aulestes eryxo (Saunders, 1859)  TL: Peru  M T S

Ancyluris aulestes jocularis Stichel, 1909  TL: Colombia  M T S

Ancyluris aulestes callaghani Lamas, Constantino & Salazar, 2013  E. foothills of the Cordillera Oriental, Colombia, 400-800 m  M T

Ancyluris aulestes pandama (Saunders, 1850)  TL: Brazil (Ba)  M T

Ancyluris colubra (Saunders, 1859)    iN  M T S

Ancyluris mira (Hewitson, 1874)    iN

Ancyluris mira mira (Hewitson, 1874)  TL: Bolivia  M T

Ancyluris mira toussaintensis Brévignon, 2019  TL: French Guiana

Ancyluris mira furia Stichel, 1925  TL: Ecuador; Colombia  M T

Ancyluris mira thaumasia Stichel, 1910  TL: Bolivia  M T S

Ancyluris jurgensenii (Saunders, 1850)  Costa-spotted Metalmark    iN

Ancyluris jurgensenii jurgensenii (Saunders, 1850)  S Mexico to Panama  M T S1 S2 L I

Ancyluris jurgensenii atahualpa (Saunders, 1859)  TL: Colombia  M T S

Ancyluris aristodorus (Morisse, 1838)    iN  M T

Ancyluris formosissima (Hewitson, 1870)    iN

Ancyluris formosissima formosissima (Hewitson, 1870)  TL: Ecuador  M T S

Ancyluris formosissima venerabilis Stichel, 1916  TL: Peru  M T

Ancyluris inca (Saunders, 1850)  Inca Metalmark    iN

Ancyluris inca inca (Saunders, 1850)  S Mexico to Panama  M T S1 S2 L I1 I2

Ancyluris inca cacica (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Ancyluris inca formosa (Hewitson, 1870)  TL: Ecuador  M T

Ancyluris inca caucaensis C. Rodríguez, G. Rodríguez & Constantino, 2013  rio Cauca in Colombia  M T

Ancyluris inca huascar (Saunders, 1859)  TL: Colombia  M T S

Ancyluris inca miranda (Hewitson, 1874)  TL: Bolivia  M T S

Ancyluris inca pulchra (Hewitson, 1870)  TL: Ecuador  M T

Ancyluris lais (Boisduval, 1870)  TL: Colombia  M T

Cyrenia  Westwood, 1851  

Cyrenia martia Westwood, 1851  Red-spotted Metalmark    iN

Cyrenia martia martia Westwood, 1851  Amazonas  M T S

Cyrenia martia androgyne Stichel, 1910  TL: Bolivia  M T

Cyrenia martia pyrippe Godman & Salvin, 1878  Costa Rica & Panama  M T TI

Cyrenia martia themis Le Cerf, 1958  TL: Colombia  M T

Rhetus  Swainson, [1829]  

Rhetus belphegor (Westwood, 1851)    iN  M T S

Rhetus arcius (Linnaeus, 1763)    iN

Rhetus arcius arcius (Linnaeus, 1763)  TL: ["Asia"]  M T

Rhetus arcius amycus Stichel, 1909  TL: Brazil (SC)  M T

Rhetus arcius angustifascia Lathy, 1932  TL: Ecuador  M T

Rhetus arcius beutelspacheri Llorente, 1988  W Mexico  M S L

Rhetus arcius castigatus Stichel, 1909  Guatemala to Costa Rica  M T S L I1 I2

Rhetus arcius huana (Saunders, 1859)  TL: Brazil (Am)  M T S

Rhetus arcius pacificus Stichel, 1924  TL: Colombia  M T

Rhetus arcius thia (Morisse, 1838)  E Mexico  M T S L

Rhetus periander (Cramer, 1777)    iN

Rhetus periander periander (Cramer, 1777)  TL: Surinam  M T S

Rhetus periander arthuriana (Sharpe, 1890)  TL: Brazil (Go)  M T

Rhetus periander eleusinus Stichel, 1910  TL: Brazil (SP/SC)  M T

Rhetus periander laonome (Morisse, 1838)  TL: Brazil; Colombia  M S

Rhetus periander naevianus Stichel, 1910  S Mexico to Panama  M T S L I

Rhetus dysonii (Saunders, 1850)    iN

Rhetus dysonii dysonii (Saunders, 1850)  TL: Venezuela  M T S

Rhetus dysonii caligosus Stichel, 1929  Costa Rica to Colombia  M T S L

Rhetus dysonii psecas (Saunders, 1850)  TL: Bolivia  M T S

Ithomeis  H. Bates, 1862  

Ithomeis eulema Hewitson, 1870    iN

Ithomeis eulema imitatrix Godman & Salvin, 1878  Costa Rica & Panama  M T S L

Ithomeis eulema caucana Salazar & Vargas, 2018  TL: Colombia

Ithomeis eulema eulema Hewitson, 1870  TL: Colombia  M T

Ithomeis eulema serena (Stichel, 1910)  TL: Colombia?  M T S

Ithomeis aurantiaca H. Bates, 1862    iN

Ithomeis aurantiaca aurantiaca H. Bates, 1862  TL: Brazil (Am)  M T

Ithomeis aurantiaca astrea (C. Felder & R. Felder, 1862)  TL: Venezuela  M T S

Ithomeis aurantiaca bernardi Brévignon, 1998  TL: French Guiana  M T

Ithomeis aurantiaca corena (C. Felder & R. Felder, 1862)  TL: Colombia  M T

Ithomeis aurantiaca delecta Stichel, 1910  TL: Brazil (SC)  M T

Ithomeis aurantiaca heliconina H. Bates, 1862  TL: Brazil (Am)  M T S

Ithomeis aurantiaca lauronia Schaus, 1902  TL: Peru  M T S

Ithomeis aurantiaca mimica H. Bates, 1862  TL: Brazil (Am)  M T S

Ithomeis aurantiaca satellites H. Bates, 1862  TL: Brazil (Pa)  M T S

Ithomeis aurantiaca stalachtina H. Bates, 1862  TL: Brazil (Am)  M T

Ithomeis aurantiaca coenoides (Hewitson, 1870)  Ecuador; ref.12  iN  M T

Ithomeis aurantiaca ecuadorica Strand, 1911  TL: Ecuador  M T

Ithomeis aurantiaca trochois (Hewitson, 1877)  TL: Ecuador  M T

Chorinea  G. Gray, 1832  

Chorinea octauius (Fabricius, 1787)    iN

Chorinea octauius octauius (Fabricius, 1787)  TL: Indiis  M T S

Chorinea octauius orchestris Stichel, 1910  TL: Peru  M T

Chorinea amazon (Saunders, 1859)    iN

Chorinea amazon amazon (Saunders, 1859)  TL: Brazil (Am)  M T S

Chorinea amazon antoniana Brévignon, 1998  TL: French Guiana

Chorinea bogota (Saunders, 1859)  Northern Chorinea  Guatemala to Colombia  iN  M T S L

Chorinea heliconides (Swainson, [1833])    iN  M T S

Chorinea licursis (Fabricius, 1775)    iN  M T S

Chorinea sylphina (H. Bates, 1868)    iN  M T S L

Chorinea gratiosa Stichel, 1910    iN  M T S

Chorinea batesii (Saunders, 1859)    iN

Chorinea batesii batesii (Saunders, 1859)  TL: Brazil (Am)  M T S

Chorinea batesii regina Brévignon, 1998  TL: French Guiana

Panara  E. Doubleday, 1847  

Panara phereclus (Linnaeus, 1758)    iN  M T S

Panara jarbas (Drury, 1782)    iN

Panara jarbas jarbas (Drury, 1782)  TL: Brazil  M S

Panara jarbas episatnius Prittwitz, 1865  TL: Brazil (RJ)  M T

Panara jarbas thymele Stichel, 1909  TL: Brazil (SP/RJ)  M T

Panara aureizona A. Butler, 1874    iN  M T S

Panara soana Hewitson, 1875    iN

Panara soana soana Hewitson, 1875  TL: Brazil  M T S

Panara soana bacana Callaghan, 1997  TL: Brazil (MG)  M T

Panara soana ruschii Callaghan, 1997  TL: Brazil (ES)  M T

Panara ovifera Seitz, 1913    iN  M T S

Amarynthis  Hübner, [1819]  

Amarynthis meneria (Cramer, 1776)    iN  M T S L

Chalodeta  Stichel, 1910  

Chalodeta theodora (C. Felder & R. Felder, 1862)    iN  M T S L

Chalodeta lypera (H. Bates, 1868)  Green-stitched Metalmark  Guatemala to Brazil  iN  M T S

Chalodeta panurga Stichel, 1910    iN  M T

Chalodeta pescada J. Hall & Willmott, 1998    iN  M T S

Chalodeta chelonis (Hewitson, 1866)  TL: Brazil (RJ)  M T S

Chalodeta chlosine J. Hall, 2002    iN  M T S

Chalodeta chitinosa J. Hall, 2002    iN  M T S

Chalodeta chaonitis (Hewitson, 1866)  Orange-stitched Metalmark  Mexico to Brazil, Trinidad  iN  M T S L

Crocozona  C. Felder & R. Felder, 1865  

Crocozona pheretima C. Felder & R. Felder, 1865    iN  M T S

Crocozona fasciata (Hopffer, 1874)    iN

Crocozona fasciata fasciata (Hopffer, 1874)  TL: Peru  M T S

Crocozona fasciata clusia (Hewitson, 1875)  TL: Bolivia  M T L

Crocozona coecias (Hewitson, 1866)    iN

Crocozona coecias coecias (Hewitson, 1866)  TL: Brazil (Am)  M T S

Crocozona coecias arcuata (Godman, 1903)  TL: Colombia  M T S

Crocozona croceifasciata J. Zikán, 1952    iN  M T

Calephelis  Grote & Robinson, 1869  

Calephelis borealis (Grote & Robinson, 1866)  Northern Metalmark  E OK, MO, IN to VA, local in New England  iN  M S L Ha G

Calephelis muticum McAlpine, 1937  Swamp Metalmark  N AR, MO, KY?, WO to MI, OH  iN  M T S L Ha G

Calephelis virginiensis (Guérin-Méneville, [1832])  Little Metalmark  E TX to FL, N to VA  iN  M T S L1 L2 I H G

Calephelis laverna (Godman & Salvin, 1886)  Laverna Calephelis    iN

Calephelis laverna laverna (Godman & Salvin, 1886)  S Mexico to Brazil  M T S I G

Calephelis laverna parva Austin, 1993  Osa Peninsula, SW Costa Rica  M T1 T2

Calephelis laverna trinidadensis McAlpine, 1971  TL: Trinidad  M T S

Calephelis nilus (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Calephelis argyrodines (H. Bates, 1866)  Argyrodines Calephelis  S Mexico to Costa Rica  iN  M T S G

Calephelis nemesis (W. H. Edwards, 1871)  Fatal Calephelis    iN

Calephelis nemesis nemesis (W. H. Edwards, 1871)  SE AZ, S NM, NW Mexico  M T S HS L1 L2 I G

Calephelis nemesis australis (W. H. Edwards, 1877)  S TX, NE Mexico  M T S L1 L2 L3 I1 I2 I3 H G

Calephelis nemesis dammersi McAlpine, 1971  [= californica], S CA, Baja California  M T1 T2 S L G

Calephelis bajaensis McAlpine, 1971  Baja Calephelis  Baja California Sur  iN  M T S L G

Calephelis exiguus Austin, 1993  Exigua Calephelis  Costa Rica  M T1 T2

Calephelis costaricicola Strand, 1916  Costa Rican Calephelis  S Mexico to Panama  M T S HS G

Calephelis perditalis W. Barnes & McDunnough, 1918  Rounded Calephelis    iN

Calephelis perditalis perditalis W. Barnes & McDunnough, 1918  S TX to SE Mexico  M T S L1 L2 I G

Calephelis perditalis donahuei McAlpine, 1971  vic. Jacala, Hidalgo, Mexico  M T S G

Calephelis mexicana McAlpine, 1971  Mexican Calephelis  E & W Mexico  iN  M T S L G

Calephelis sixola McAlpine, 1971  Greater Calephelis  Costa Rica  iN  M T S G

Calephelis wrighti W. Holland, 1930  Wright’s Calephelis  Baja California, lower Colorado River drainage  iN  M T S1 S2 L1 L2 H G

Calephelis fulmen Stichel, 1910  Fulmen Calephelis  E & W Mexico to Panama  iN  M T S G

Calephelis rawsoni McAlpine, 1939  Rawson’s Calephelis  W & S TX, NE Mexico  iN  M T S L H G

Calephelis freemani McAlpine, 1971  Freeman's Calephelis  Davis Mts., Jeff Davis Co., TX  M T G

Calephelis arizonensis McAlpine, 1971  Arizona Calephelis  SE AZ, SW NM, NW Mexico  iN  M T S L1 L2 L3 I H G

Calephelis sinaloensis McAlpine, 1971  Sinaloan Calephelis    iN

Calephelis sinaloensis sinaloensis McAlpine, 1971  NW Mexico (S Sinaloa)  M T S G

Calephelis sinaloensis nuevoleon McAlpine, 1971  NE & N-central Mexico (Nuevo Leon to Durango)  M T S G

Calephelis dreisbachi McAlpine, 1971  Dreisbach's Calephelis  NW Mexico (Sinaloa-Nayarit), [SE Arizona?]  M T G

Calephelis stallingsi McAlpine, 1971  Stallings' Calephelis  E Mexico to Guatemala  M T G

Calephelis matheri McAlpine, 1971  Mather’s Calephelis  W-central Mexico  M T S L G

Calephelis huasteca McAlpine, 1971  Huastecan Calephelis  E Mexico  iN  M T S G

Calephelis montezuma McAlpine, 1971  Montezuma’s Calephelis  E & W Mexico  M T S G

Calephelis acapulcoensis McAlpine, 1971  Acapulco Calephelis  SW Mexico  M T S G

Calephelis azteca McAlpine, 1971  Aztec Calephelis  E Mexico  iN  M T S G

Calephelis yucatana McAlpine, 1971  Yucatan Calephelis  Yucatan Peninsula and vicinity  M T S G

Calephelis maya McAlpine, 1971  Mayan Calephelis  Yucatan Peninsula and vicinity  M T S G

Calephelis yautepequensis R. G. Maza & Turrent, 1977  Morelos Calephelis  S-central Mexico  M T

Calephelis wellingi McAlpine, 1971  Welling's Calephelis  TL: Mexico (Tab)

Calephelis wellingi wellingi McAlpine, 1971  SE Mexico to Belize  M T S G

Calephelis wellingi baleuensis McAlpine, 1971  Alta Verapaz, Guatemala  M T G

Calephelis sacapulas McAlpine, 1971  Sacapulas Calephelis  Guatemala, [S Mexico?]  iN  M T G

Calephelis clenchi McAlpine, 1971  Clench's Calephelis  Guatemala to El Salvador  iN  M T S G

Calephelis browni McAlpine, 1971  Brown's Calephelis  S Mexico to Panama  iN  M T S G

Calephelis sodalis Austin, 1993  Sodalis Calephelis  Costa Rica, [Panama?]  M T1 T2

Calephelis schausi McAlpine, 1971  Schaus' Calephelis  Honduras to Panama  iN  M T S G

Calephelis guatemala McAlpine, 1971  Guatemalan Calephelis  Guatemala  M T OI G

Calephelis inca McAlpine, 1971  Inca Calephelis  Costa Rica to Brazil & Trinidad  iN  M T S G

Calephelis tapuyo McAlpine, 1971  Tapuyo Calephelis  possibly Costa Rica  M T G

Calephelis aymaran McAlpine, 1971    iN  M T

Calephelis distinctus Austin, 2008  Iridistinct Calephelis  Brazil

Calephelis canus Austin, 2008  Distogray Calephelis  Brazil

Calephelis brevis Austin, 2008  Transtishort Calephelis  Brazil

Calephelis rotundum Austin, 2008  Allround Calephelis  Brazil

Calephelis indistinctus Austin, 2008  Indistinct Calephelis  Brazil

Calephelis braziliensis McAlpine, 1971    iN  M T

Calephelis burgeri McAlpine, 1971  TL: Colombia  M T

Calephelis tikal Austin, 1993  Tikal Calephelis  SE Mexico (Yucatan Peninsula) to Guatemala  iN  M T S

Calephelis candiope (H. Druce, 1904)  Candiope Calephelis  Costa Rica to Colombia  M T S

Calephelis iris (Staudinger, 1876)  Iris Calephelis  Guatemala to Colombia  iN  M T S L I G

Calephelis velutina (Godman & Salvin, 1878)  Dark Calephelis  S Mexico to Colombia  iN  M T S L G

Amphiselenis  Staudinger, 1888  

Amphiselenis chama (Staudinger, [1887])    iN  M T S

Charis  Hübner, [1819]  

Charis anius (Cramer, 1776)  Anius Metalmark  Honduras to Brazil  iN  M S L

Chadia  Grishin, 2021  

Chadia cadytis (Hewitson, 1866)  Brazil; ref.12  iN  M T S

Detritivora  J. Hall & Harvey, 2002  

Detritivora major (Lathy, 1932)  TL: Ecuador  M T

Detritivora matic (Harvey & J. Hall, 2002)    iN  M T S

Detritivora ma (Harvey & J. Hall, 2002)    iN  M T S

Detritivora cleonus (Stoll, 1782)    iN  M T

Detritivora rocana (Harvey & J. Hall, 2002)  TL: Brazil (Ap)  M T

Detritivora negro (Harvey & J. Hall, 2002)  TL: Venezuela  M T

Detritivora ariquemes (Harvey & J. Hall, 2002)  TL: Brazil (Rd)  M T

Detritivora manu (Harvey & J. Hall, 2002)    iN  M T S

Detritivora humaita (Harvey & J. Hall, 2002)  TL: Brazil (Am)

Detritivora iquitos (Harvey & J. Hall, 2002)  TL: Peru  M T S

Detritivora tefe (Harvey & J. Hall, 2002)  TL: Brazil (Am)  M T S

Detritivora palcazu (Harvey & J. Hall, 2002)    iN  M T S

Detritivora cacaulandia (Harvey & J. Hall, 2002)  TL: Brazil (Rd)  M T

Detritivora manicore (Harvey & J. Hall, 2002)  TL: Brazil (Am)  M T

Detritivora ipiranga (Harvey & J. Hall, 2002)  TL: Brazil (Am)  M T

Detritivora cuiaba (Harvey & J. Hall, 2002)  TL: Brazil (MT)  M T

Detritivora maues (Harvey & J. Hall, 2002)  TL: Brazil (Am)  M T

Detritivora tapajos (Harvey & J. Hall, 2002)  TL: Brazil (Pa)  M T

Detritivora santarem (Harvey & J. Hall, 2002)    iN  M T

Detritivora breves (Harvey & J. Hall, 2002)  TL: Brazil (Pa)  M T

Detritivora caryatis (Hewitson, 1866)    iN  M T S

Detritivora brasilia (Harvey & J. Hall, 2002)    iN

Detritivora nicolayi (J. Hall & Harvey, 2001)  Nicolay's Metalmark  Costa Rica & Panama  iN  M T

Detritivora hermodora (C. Felder & R. Felder, 1861)  Hermodora Metalmark  Costa Rica to Venezuela  iN  M T S L

Detritivora gynaea (Godart, [1824])  Gynaea Metalmark  Costa Rica to Brazil  iN  M T S

Detritivora gallardi (J. Hall & Harvey, 2001)  TL: French Guiana  M T

Detritivora barnesi (J. Hall & Harvey, 2001)  Barnes' Metalmark  SE Mexico to W Ecuador  iN  M T S L1 L2

Detritivora callaghani (J. Hall & Harvey, 2001)  TL: Colombia  M T

Detritivora zama (H. Bates, 1868)    iN  M T S

Detritivora mapaou Gallard & Fernandez, 2017  TL: French Guiana

Putridivora  Grishin, 2021  

Putridivora argyrea (H. Bates, 1868)  ref.12  M T S

Putridivora smalli (J. Hall & Harvey, 2001)  Small's Metalmark  ref.12

Putridivora smalli smalli (J. Hall & Harvey, 2001)  E Panama to W Ecuador  M T

Putridivora smalli harveyi (Gallard, 2008)  TL: French Guiana  M T

Lasaia  H. Bates, 1868  

Lasaia callaina Clench, 1972  Shining-blue Lasaia  E & W Mexico to N Costa Rica; ref.154  M T S1 S2 L I1 I2

Lasaia agesilas (Latreille, [1809])  Shining-blue Lasaia  S Costa Rica to Bolivia & Brazil  iN  M T S L

Lasaia aerugo Clench, 1972  TL: Peru  M T

Lasaia maritima J. Hall & Lamas, 2001  TL: Peru  M T

Lasaia arsis Staudinger, [1887]    iN  M T S

Lasaia meris (Stoll, 1781)  Variegated Lasaia  S Mexico to Brazil  iN  M S

Lasaia pseudomeris Clench, 1972  Clench's Lasaia  Costa Rica to Venezuela & Peru  iN  M T S L

Lasaia pallida Grishin, 2024  Colombia & Venezuela; ref.154  M T

Lasaia peninsularis Clench, 1972  Blue Lasaia  S TX, E Mexico; ref.151  M T S L1 L2 L3 L4

Lasaia sula Staudinger, 1888  Blue Lasaia  W Mexico to Costa Rica; ref.151  iN  M T S L

Lasaia maria Clench, 1972  Blue-gray Lasaia    iN

Lasaia maria anna Clench, 1972  E Mexico  M T S L

Lasaia maria maria Clench, 1972  W Mexico to Guatemala, Honduras  M T S L

Lasaia sessilis Schaus, 1890  Gray Lasaia  E & S Mexico to Costa Rica  iN

Lasaia sessilis sessilis Schaus, 1890  E & S Mexico to Costa Rica  iN  M T S L

Lasaia sessilis oaxacensis Grishin, 2024  Mexico: Oax; ref.154

Lasaia moeros Staudinger, 1888    iN  M T S

Lasaia kennethi Weeks, 1901  TL: Bolivia  M T

Lasaia oileus Godman, 1903  Oily Lasaia  Belize to Paraguay, French Guiana, Trinidad  iN  M T S

Exoplisia  Godman & Salvin, 1886  

Exoplisia hypochalybe (C. Felder & R. Felder, 1861)  Many-banded Metalmark    iN

Exoplisia hypochalybe hypochalybe (C. Felder & R. Felder, 1861)  S Mexico to Colombia  M T S

Exoplisia hypochalybe muscolor (Weeks, 1901)  TL: Bolivia  M T

Exoplisia cadmeis (Hewitson, 1866)  Cadmeis Metalmark  Honduras to Peru  iN  M T S L

Exoplisia myrtis (H. Druce, 1904)    iN  M T S

Exoplisia azuleja Callaghan, Llorente & Luis, 2007  Occidental Metalmark  W Mexico  iN  M S

Exoplisia aphanis (Stichel, 1910)  SE S. Amer.; ref.12  iN  M T

Metacharis  A. Butler, 1867  

Metacharis ptolomaeus (Fabricius, 1793)    iN  M S

Metacharis lucius (Fabricius, 1793)    iN  M T S

Metacharis regalis A. Butler, 1867    iN

Metacharis regalis regalis A. Butler, 1867  TL: Brazil (Am)  M T S

Metacharis regalis indissimilis Weeks, 1901  TL: Colombia  M T S

Metacharis cuparina H. Bates, 1868  TL: Brazil (Pa)  M T S

Metacharis nigrella H. Bates, 1868  TL: Brazil (Pa)  M T

Metacharis victrix (Hewitson, 1870)  Victrix Metalmark  Nicaragua to W Ecuador  iN  M T S1 S2 L I

Metacharis umbrata J. Hall, 2005    iN  M T S I

Metacharis syloes Hewitson, 1877  Confused Metalmark  N Venezuela to Bolivia  iN  M T S

Metacharis xanthocraspedum Stichel, 1910    iN  M T

Metacharis fergusi J. Hall, 2005  ref.219  iN  M S

Metacharis smalli J. Hall, 2005  Small's Metalmark  Panama; ref.219  M T S G

Astraeodes  Staudinger, [1887]  

Astraeodes areuta (Westwood, 1851)    iN  M T S

Cartea  W. F. Kirby, 1871  

Cartea vitula (Hewitson, [1853])    iN

Cartea vitula vitula (Hewitson, [1853])  TL: Brazil (Am)  M T

Cartea vitula tapajona (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Cartea vitula trailii A. Butler, 1877  TL: Brazil (Am)  M T S

Cartea ucayala Thieme, 1907    iN  M T

Cartea chiribiquetensis Salazar & Constantino, 2007  TL: Colombia

Oco  Grishin, 2021  

Oco ocellata (Hewitson, 1867)  Venezuela; ref.12  M T S

Chamaelimnas  C. Felder & R. Felder, 1865  

Chamaelimnas tircis C. Felder & R. Felder, 1865    iN

Chamaelimnas tircis tircis C. Felder & R. Felder, 1865  TL: Brazil (Ba)  M T

Chamaelimnas tircis iaeris H. Bates, 1868  TL: Brazil (Am)  M T S

Chamaelimnas tircis pansa Godman, 1903  TL: Brazil (MT)  M T S

Chamaelimnas briola H. Bates, 1868    iN

Chamaelimnas briola briola H. Bates, 1868  TL: Peru  M T S

Chamaelimnas briola doryphora Stichel, 1910  TL: Brazil (RJ)  M T S

Chamaelimnas briola meridionalis Lathy, 1932  TL: Paraguay  M T

Chamaelimnas briola urbana Stichel, 1916  TL: Peru  M T S

Chamaelimnas cydonia Stichel, 1910  Cydonia Metalmark  [= villagomes], Panama, [Costa Rica?]  M T S

Chamaelimnas cercides Hewitson, 1871    iN  M T

Chamaelimnas joviana Schaus, 1902    iN  M T S

Chamaelimnas splendens Grose-Smith, 1902  TL: Bolivia  M T

Barbicornis  Godart, [1824]  

Barbicornis basilis Godart, [1824]    iN

Barbicornis basilis basilis Godart, [1824]  TL: Brazil  M T S

Barbicornis basilis acroleuca Berg, 1896  TL: Paraguay  M T S

Barbicornis basilis bahiana Azzará, 1979  TL: Brazil (Ba)  M T

Barbicornis basilis ephippium Thieme, 1907  TL: Brazil (RS)  M T

Barbicornis basilis marginata Seitz, 1913  TL: Brazil (Ba)  M T

Barbicornis basilis mona Westwood, 1851  TL: Brazil (ES)  M T

Barbicornis basilis paraopeba Azzará, 1979  TL: Brazil (MG)  M T

Barbicornis basilis tucumana Thieme, 1907  TL: Argentina  M T S

Seco  J. Hall & Harvey, 2002  

Seco calagutis (Hewitson, 1871)    iN  M T S L

Caria  Hübner, 1823  

Caria ino Godman & Salvin, 1886  Red-bordered Metalmark    iN

Caria ino ino Godman & Salvin, 1886  W Mexico  M T S L

Caria ino melicerta Schaus, 1890  E Mexico, S TX  M T S Ma L1 L2 L3 I H

Caria domitianus (Fabricius, 1793)    iN

Caria domitianus domitianus (Fabricius, 1793)  TL: Guadeloupe  M T S

Caria domitianus galbula (C. Felder & R. Felder, 1861)  TL: Venezuela  M T

Caria domitianus vejento Clench, 1967  SE Mexico to Panama  M T S L

Caria rhacotis (Godman & Salvin, 1878)  Honduran Metalmark  S Mexico to Peru  iN  M T S1 S2 L I

Caria stillaticia Dyar, 1912  Gold-headed Metalmark  [= rabatta], SW Mexico  iN  M T S L Ha

Caria melino Dyar, 1912  Melino Metalmark  SE Mexico  M T S L

Caria trochilus Erichson, [1849]    iN

Caria trochilus trochilus Erichson, [1849]  TL: Guyana  M T

Caria trochilus arete (C. Felder & R. Felder, 1861)  TL: [Peru]  M T S

Caria castalia (Ménétriés, 1855)    iN  M S L

Caria marsyas Godman, 1903    iN  M T S

Caria tabrenthia Schaus, 1902    iN  M T

Caria mantinea (C. Felder & R. Felder, 1861)    iN

Caria mantinea mantinea (C. Felder & R. Felder, 1861)  TL: [Peru]  M T S

Caria mantinea lampeto Godman & Salvin, 1886  S Mexico to Bolivia  M T S1 S2 L I

Caria plutargus (Fabricius, 1793)    iN

Caria plutargus plutargus (Fabricius, 1793)  TL: Indiis  M T S

Caria plutargus amazonica (H. Bates, 1868)  TL: Brazil (Am)  M T S

Caria [n. sp.] Callaghan & Lamas, MS  Ecuador  M T

Caria chrysame (Hewitson, 1874)    iN

Caria chrysame chrysame (Hewitson, 1874)  TL: Bolivia  M T S

Caria chrysame psittacus (Hopffer, 1874)  TL: Peru  M T S

Caria sponsa (Staudinger, [1887])    iN  M T S

  Tribe Emesidini

Curvie  Grishin, 2019  

Curvie emesia (Hewitson, 1867)  Curve-winged Emesis  E & W Mexico to Yucatan  iN  M T S1 S2 S3 L1 L2 I1 I2 H

Apodemia  C. Felder & R. Felder, 1865  

Apodemia mormo (C. Felder & R. Felder, 1859)  Mormon Metalmark    iN

Apodemia mormo mormo (C. Felder & R. Felder, 1859)  Columbia Basin, NW Great Basin to E edge Cascades, Siskiyous  M T S1 S2 L1 L2 H Ha

Apodemia mormo parva Austin, 1998  NE NV, W & N UT, SE ID, W WY, NE CO  M T S L

Apodemia mormo autumnalis Austin, 1998  N AZ, N NM, SE CA, SW UT, S NV; possibly E to SW CO  M T S L I

Apodemia mormo SE Arizona segregate ,  SE Arizona  M S L I Ha

Apodemia mormo cythera (W. H. Edwards, 1873)  widespread in S CA, univoltine in fall  M T S1 S2 L

Apodemia mormo tuolumnensis Opler & Powell, 1962  C & S Sierra Nevada, at least  M T S

Apodemia mormo langei J. A. Comstock, 1939  vic. Antioch, Contra Costa Co., CA  M T S L

Apodemia mormo C & S CA Coast Ranges segregate ,  CA San Fran. Bay area S to Baywood Park area, SLO Co.  M S L1 L2 H Ha

Apodemia mormo Cottonwood Pass segregate ,  [nr. langei], Ortega Grade to Cottonwood Pass, SLO Co., CA  M S L1 L2 L3 L4 H Ha

Apodemia virgulti (Behr, 1865)  Behr's Metalmark    iN

Apodemia virgulti peninsularis J. Emmel, T. Emmel & Pratt, 1998  San Jacinto, Palomar & Laguna Mts., Riv., S.D. cos., CA; univoltine  M T S L I Ha

Apodemia virgulti virgulti (Behr, 1865)  SW CA, Baja California (N); widespread, multivoltine  M T S1 S2 L1 L2 L3 L4 I H Ha

Apodemia virgulti arenaria J. Emmel, T. Emmel & Pratt, 1998  El Segundo, Los Angeles Co., CA  M T

Apodemia virgulti nigrescens J. Emmel & T. Emmel, 1998  vic. Colton, San Bernardino Co., CA  M T S

Apodemia virgulti pratti J. Emmel & T. Emmel, 1998  Holcomb Valley, San Bernardino Mts., San Bernardino Co., CA  M T

Apodemia virgulti dialeuca Opler & Powell, 1962  Baja California (Norte)  M T S

Apodemia virgulti dialeucoides J. Emmel, T. Emmel & Pratt, 1998  San Bernardino Mts.; Sugarloaf Mt. E to Onyx Peak; univoltine  M T S

Apodemia virgulti mojavelimbus J. Emmel, T. Emmel & Pratt, 1998  SW edge Mojave Desert & San Gabriel Mts.; univoltine (spring)  M T S L Ha

Apodemia virgulti davenporti J. Emmel, T. Emmel & Pratt, 1998  E slope S. Nevada (Olancha to Walker Pass); Tehachapis, Piutes  M T S L

Apodemia mejicanus (Behr, 1865)  Mexican Metalmark    iN

Apodemia mejicanus mejicanus (Behr, 1865)  NW Mexico, SE AZ  M S1 S2 L1 L2 I

Apodemia mejicanus maxima (Weeks, 1891)  Baja California  M T TI S

Apodemia mejicanus pueblo Scott, 1998  E NM, S-central CO, N CO, SE WY, E MT, W ND  M S1 S2 L1 L2 H

Apodemia mejicanus deserti W. Barnes & McDunnough, 1918  Colorado Desert  M T S L1 L2 I H Ha

Apodemia mejicanus S San Luis Valley segregate ,  S San Luis Valley, CO-NM  M S L1 L2 H

Apodemia duryi (W. H. Edwards, 1882)  Dury's Metalmark  S-central NM, W TX, N-central Mexico to QRO  iN  M T S L

Apodemia multiplaga Schaus, 1902  Narrow-winged Metalmark  S TX (rare stray), E & W Mexico to NW Costa Rica  iN  M T S L

Apodemia palmerii (W. H. Edwards, 1870)  Palmer’s Metalmark    iN

Apodemia palmerii palmerii (W. H. Edwards, 1870)  Baja California (N), S CA, S NV, SW UT, W AZ  M T S IL L

Apodemia palmerii arizona Austin, [1989]  C AZ to W TX, NW Mexico (SON, SIN, CHIH)  M T S IL L1 L2 I

Apodemia palmerii australis Austin, [1989]  C Mexico (DGO to MICH & HGO)  M T S IL L

Apodemia murphyi Austin, [1989]  Murphy’s Metalmark  Baja California  iN  M T S IL L

Apodemia hepburni Godman & Salvin, 1886  Hepburn’s Metalmark    iN

Apodemia hepburni remota Austin, 1991  Baja California Sur  M T S L

Apodemia hepburni hepburni Godman & Salvin, 1886  NW Mexico, uncommon in SE AZ, W TX  M T S L

Apodemia walkeri Godman & Salvin, 1886  Walker’s Metalmark  S TX (rare stray), E & W Mexico to NW Costa Rica  iN  M T S L1 L2

Apodemia selvatica J. Maza & RG. Maza, 2017  TL: Mexico (Chia)

Apodemia hypoglauca (Godman & Salvin, 1878)  Falcate Metalmark    iN

Apodemia hypoglauca hypoglauca (Godman & Salvin, 1878)  E & W Mexico, Guatemala  M T S L

Apodemia hypoglauca wellingi Ferris, 1985  Yucatan Peninsula & vicinity  M T S

Apodemia gerres (Thieme, 1907)  Colombia; ref.11  iN  M T S

Apodemia calvus (Staudinger, [1887])  Peru; ref.11  iN  M T S

Apodemia nais (W. H. Edwards, 1877)  Nais Metalmark  N-central CO to SW NM, C & SE AZ, NW Mexico (CHIH, SON, DGO)  iN  M T S L1 L2 I Ha

Apodemia chisosensis H. Freeman, 1964  Chisos Metalmark  SW TX, Hypothetical for N-central Mexico  iN  M T S L

Apodemia ares (W. H. Edwards, 1882)  Ares Emesis  SE AZ, SW NM, NW Mexico  iN  M T S L I

Apodemia zela (A. Butler, 1870)  Zela Emesis  W & C Mexico to Guatemala  iN  M T S L

Apodemia cleis (W. H. Edwards, 1882)  Zela Emesis  S AZ, SW NM, NW Mexico; ref.16  M T S L1 L2 I

Apodemia arnacis (Stichel, 1928)  Miahuatlan Emesis  SW Mexico (Sierra Madre del Sur, Oaxaca)  iN  M T S L Ha

Emesis  Fabricius, 1807  

Emesis cereus (Linnaeus, 1767)    iN

Emesis cereus cereus (Linnaeus, 1767)  TL: Indiis  M T S HS

Emesis cereus cronina Schaus, 1928  Paraguay  iN  M T S

Emesis malik Callaghan, Costa, Trujano-Ortega & M. Benmesbah, 2021  Venezuela  M T

Emesis neemias Hewitson, 1872    iN

Emesis neemias cayennensis Gallard, 2017  TL: French Guiana

Emesis neemias neemias Hewitson, 1872  TL: Brazil  M T S

Emesis orichalceus Stichel, 1916    iN  M T S

Emesis aerigera (Stichel, 1910)  TL: Brazil (SP)  M T S

Emesis lacrines Hewitson, 1870  Striated Emesis  Nicaragua to Panama  iN  M T S1 S2 L I

Emesis fatimella Westwood, 1851    iN

Emesis fatimella fatimella Westwood, 1851  TL: Surinam; Brazil (Am)  M T S

Emesis fatimella nobilata Stichel, 1910  S Mexico to Colombia, Guianas, Trinidad  M T S L

Emesis furor A. Butler & H. Druce, 1872  Great Emesis  E & W Mexico to Costa Rica  M T S1 S2 S3 L I1 I2 I3 I4

Emesis mandana (Cramer, 1780)  Great Emesis  S Costa Rica to Brazil  iN  M T S1 S2 L

Emesis russula Stichel, 1910    iN  M T S

Emesis melancholica Stichel, 1916  TL: Brazil (ES)  M T S

Emesis ocypore (Geyer, 1837)    iN

Emesis ocypore ocypore (Geyer, 1837)  TL: Africa  M T S

Emesis ocypore aethalia H. Bates, 1868  S Mexico to Colombia  M T S1 S2 L I

Emesis ocypore zelotes Hewitson, 1872  TL: Brazil  M T S

Emesis angularis Hewitson, 1870    iN  M T S L

Emesis sinuatus Hewitson, 1877  TL: Ecuador  M T

Emesis heterochroa Hopffer, 1874    iN  M T S

Emesis tenedia C. Felder & R. Felder, 1861  Falcate Emesis  S TX (as stray), E & W Mexico to Brazil  iN

Emesis tenedia tenedia C. Felder & R. Felder, 1861  S TX (as stray), E & W Mexico to Brazil  M T S1 S2 S3 L

Emesis tenedia guyanensis Gallard, 2017  TL: French Guiana

Emesis [n. sp.] N/A, MS  Hook-tipped Emesis  Chiapas  M S

Emesis toltec Reakirt, 1866  Toltec Emesis  Mexico  M S IL

Emesis saturata Godman & Salvin, 1886  Oaxacan Emesis  S Mexico  iN  M T S L

Emesis cypria C. Felder & R. Felder, 1861    iN

Emesis cypria cypria C. Felder & R. Felder, 1861  TL: Venezuela  M T S

Emesis cypria capnodis Stichel, 1911  TL: Venezuela  M T

Emesis cypria cilix Hewitson, 1870  TL: Ecuador  M T S

Emesis cypria guppyi Kaye, 1904  TL: Trinidad; Colombia  M T S

Emesis cypria paphia R. Felder, 1869  S Mexico to Colombia  M T S1 S2 L I

Emesis eleanorae Gallardo & Grishin, 2021  Honduras; ref.117

Emesis tegula Godman & Salvin, 1886  Tegula Emesis  E & W Mexico to Colombia  iN  M T S1 S2 L I

Emesis lupina Godman & Salvin, 1886  Lupina Emesis  E & W Mexico to Venezuela  iN  M T OI S

Emesis phyciodoides (W. Barnes & Benjamin, 1924)  Crescent Metalmark  SE AZ (extirpated), NW Mexico (SON, CHIH)  iN  M T S L

Emesis planeca (RG. Maza & J. Maza, 2017)  ref.2

Emesis poeas Godman, 1901  Thorn-scrub Emesis  W Mexico  iN  M T S L

Emesis brimo Godman & Salvin, 1889  Brimo Emesis    iN

Emesis brimo vimena Schaus, 1928  S Mexico to Panama  M T

Emesis brimo brimo Godman & Salvin, 1889  Panama to Colombia  M T S1 S2 I

Emesis brimo insulicola Vargas & Salazar, 2018  TL: Colombia

Emesis progne (Godman, 1903)    iN  M T S

Emesis temesa (Hewitson, 1870)    iN

Emesis temesa temesa (Hewitson, 1870)  TL: Ecuador  M T S

Emesis temesa peruviana (Lathy, 1904)  TL: Peru  M T S

Emesis satema (Schaus, 1902)  TL: Brazil (RJ)  M T S

Emesis liodes Godman & Salvin, 1886  Liodes Emesis  Yucatan Peninsula & vicinity  iN  M T S L

Emesis aurimna (Boisduval, 1870)  White-spotted Emesis  E & S Mexico to Colombia  iN  M T S1 S2 L I1 I2

Emesis glaucescens Talbot, 1929  White-frosted Emesis  Colombia  iN  M T S

Emesis lucinda (Cramer, 1775)  White-patched Emesis  Costa Rica to N Brazil & Guianas  iN  M T S1 S2 L I

Emesis fastidiosa Ménétriés, 1855    iN  M T S

Emesis condigna Stichel, 1925    iN  M T S

Emesis castigata Stichel, 1910    iN  M T S

Emesis opaca Stichel, 1910  TL: French Guiana  M T S

Emesis eurydice Godman, 1903    iN  M T S

Emesis spreta H. Bates, 1868    iN  M T S

Emesis vulpina Godman & Salvin, 1886  Pale Emesis  E & W Mexico to Guatemala  iN  M T S1 S2 L

Emesis diogenia Prittwitz, 1865    iN  M T S

Emesis heteroclita Stichel, 1929    iN

Emesis heteroclita heteroclita Stichel, 1929  TL: Peru  M T S

Emesis heteroclita vicaria Le Cerf, 1958  TL: Amazon  M T

Emesis heteroclita adelpha Le Cerf, 1958  Bolivia  iN  M T S

  Tribe Nymphidiini

Zabuella  Stichel, 1911  

Zabuella tenellus (Burmeister, 1878)  Argentina & Paraguay  iN  M T S L

Zabuella castanea (Prittwitz, 1865)  SE Brazil  iN  M T S

Zabuella paucipuncta (Spitz, 1930)  SE Brazil  iN  M T S

Zabuella maculosa (H. Bates, 1868)  Mexico to Brazil  iN  M T S

Zabuella tinea (H. Bates, 1868)  NW Mexico to S Brazil & Argentina  iN  M T S L

Zabuella argiella (H. Bates, 1868)  SE Brazil  iN  M T S

Catocyclotis  Stichel, 1911  

Catocyclotis wiltoni J. Hall, 2018  TL: Ecuador  M T

Catocyclotis malca (Schaus, 1902)    iN  M T S

Catocyclotis sejuncta (Stichel, 1910)    iN  M T S

Catocyclotis marmoreus J. Hall, 2018  TL: Brazil(MG)  M T

Catocyclotis leucarpis (Stichel, 1925)    iN  M T

Catocyclotis aemulius (Fabricius, 1793)    iN  M T S

Catocyclotis adelina (A. Butler, 1872)  Adelina Metalmark  Costa Rica to W Ecuador  iN  M T S L

Catocyclotis densemaculata (Hewitson, 1870)  Tan Metalmark  Nicaragua to Peru  iN  M T S L

Catocyclotis elpinice (Godman, 1903)  TL: Colombia  M T S

Catocyclotis zerna (Hewitson, 1872)    iN  M T S

Catocyclotis glauca (Godman & Salvin, 1886)  Glauca Metalmark  Costa Rica to Venezuela  M T S L

Catocyclotis curulis (Hewitson, 1874)    iN  M T S

Nymphidium  Fabricius, 1807  

Nymphidium scurrilis Stichel, 1929  TL: Peru  M T S

Nymphidium nealces Hewitson, 1871  TL: French Guiana  M T S

Nymphidium omois Hewitson, 1865    iN  M T S

Nymphidium callaghani Brévignon, 1999  TL: French Guiana  M S

Nymphidium ariari Callaghan, 1988  TL: Colombia  M T

Nymphidium lenocinium Schaus, 1913  Lenocinium Metalmark  SE Mexico (CHIS) to Colombia  M T S

Nymphidium pseudominuta J. Hall, 2018  TL: Ecuador  M T S

Nymphidium azanoides A. Butler, 1867    iN

Nymphidium azanoides azanoides A. Butler, 1867  TL: Brazil (Pa)  M T S L

Nymphidium azanoides amazonensis Callaghan, 1986  TL: Brazil (Am)  M T S L

Nymphidium fulminans H. Bates, 1868    iN

Nymphidium fulminans fulminans H. Bates, 1868  TL: Brazil (Pa)  M T S

Nymphidium fulminans sardinata Callaghan, 1985  Colombia  M T

Nymphidium guyanensis Gallard & Brévignon, 1989  TL: French Guiana  M T S

Nymphidium olinda H. Bates, 1865  Olinda Metalmark  Panama to Venezuela & Brazil  iN  M T S

Nymphidium mantus (Cramer, 1775)  Mantus Metalmark  Costa Rica to Venezuela & Brazil  iN  M T S L

Nymphidium manicorensis Callaghan, 1985  TL: Brazil (Am)  M T S

Nymphidium minuta H. Druce, 1904    iN  M T

Nymphidium baeotia Hewitson, [1853]    iN  M T S L

Nymphidium robiginosum Stichel, 1929  TL: Brazil (Am)  M T S

Nymphidium aurum Callaghan, 1985  TL: Brazil (Am)  M T S

Nymphidium menalcus (Stoll, 1782)    iN  M T S

Nymphidium ninias Hewitson, 1865    iN  M T S

Nymphidium medusa H. Druce, 1904    iN

Nymphidium medusa medusa H. Druce, 1904  TL: Peru  M T S

Nymphidium medusa semiramis Stichel, 1924  TL: Colombia  M T S

Nymphidium leucosia (Hübner, [1806])    iN  M T S

Nymphidium chione H. Bates, 1867    iN  M T S L

Nymphidium cachrus (Fabricius, 1787)    iN  M T S L V

Nymphidium chimborazium H. Bates, 1868    iN  M T S

Nymphidium haematostictum Godman & Salvin, 1878  Blood-spot Metalmark  Costa Rica & Panama  iN  M T S L

Nymphidium acherois (Boisduval, 1836)    iN  M T S

Nymphidium balbinus Staudinger, [1887]    iN  M T S

Nymphidium onaeum Hewitson, 1869  Hewitson's Metalmark  S Mexico to Panama  iN  M T S1 S2 I

Nymphidium lisimon (Stoll, 1790)    iN  M T S L

Nymphidium velatum Stichel, 1914  TL: Peru  M T S

Nymphidium plinthobaphis Stichel, 1910    iN  M T S L

Nymphidium carmentis Stichel, 1910    iN  M T S

Nymphidium caricae (Linnaeus, 1758)    iN

Nymphidium caricae caricae (Linnaeus, 1758)  TL: America  M T S V

Nymphidium caricae erubescens Stichel, 1929  TL: Venezuela  M T S

Nymphidium caricae trinidadi Callaghan, 1999  Trinidad  iN  M T S

Nymphidium colleti Gallard, 2008  TL: French Guiana  M T

Nymphidium hermieri Gallard, 2008  TL: French Guiana  M T

Minotauros  J. Hall, 2007  

Minotauros lampros (H. Bates, 1868)  TL: Brazil (Am)  M T S

Minotauros charessa (Stichel, 1910)  TL: Brazil (Am)  M T

Minotauros confusa Salazar, Nielsen & Vargas, 2018  TL: Colombia

Parvospila  J. Hall, 2018  

Parvospila lucianus (Fabricius, 1793)  Lucianus Metalmark  Costa Rica to Venezuela, Guianas, Trinidad  iN  M T S L

Parvospila lucetia (Hübner, 1821)    iN  M T S

Parvospila emylius (Cramer, 1775)    iN  M T S

Parvospila asteria (Stichel, 1911)  Asteria Metalmark  Panama to Colombia  M T S

Parvospila cilissa (Hewitson, 1863)  Cilissa Metalmark  Honduras to Colombia  iN  M T S L

Periplacis  Geyer, 1837  

Periplacis felsina (Hewitson, 1863)    iN  M T S

Periplacis laobotas (Hewitson, 1875)  Three-spotted Metalmark  Nicaragua to Ecuador  iN  M T OI S

Periplacis apotheta (H. Bates, 1868)    iN  M T S

Periplacis coruscans (A. Butler, 1867)  TL: Brazil (Pa)  M T S

Periplacis menander (Stoll, 1780)    iN  M T S L

Periplacis nitida (A. Butler, 1867)    iN  M T S

Periplacis glaucoma Geyer, 1837    iN  M T S

Periplacis splendida (A. Butler, 1867)  TL: Brazil (Pa)  M T S L

Periplacis pretus (Cramer, 1777)    iN  M T S1 S2 I

Periplacis homerus J. Hall, 2018  TL: Colombia  M T

Periplacis aldasi (J. Hall & Willmott, 1995)  TL: Ecuador  M T S

Periplacis cicuta (Hewitson, 1863)  TL: Brazil (Am)  M T S

Periplacis clotho (Stichel, 1911)  TL: Peru  M T S

Periplacis hebrus (Cramer, 1775)    iN  M T S

Periplacis thalassicus (Brévignon & Gallard, 1992)  TL: French Guiana  M T

Periplacis perisama J. Hall & Ahrenholz, 2020  TL: Ecuador

Livendula  J. Hall, 2007  

Livendula balista (Hewitson, 1863)  TL: Brazil (Am)  M T S

Livendula epixanthe (Stichel, 1911)  TL: Brazil (Am)  M T S

Livendula aminias (Hewitson, 1863)    iN  M T S

Livendula leucophaea (Hübner, [1821])    iN  M T

Livendula jasonhalli (Brévignon & Gallard, 1999)    iN

Livendula leucocyana (Geyer, 1837)    iN  M S

Livendula aristus (Stoll, 1790)    iN  M T S L

Livendula pauxilla (Stichel, 1911)    iN  M T S

Livendula huebneri (A. Butler, 1867)    iN  M T S L

Livendula violacea (A. Butler, 1867)    iN  M T S

Livendula amasis (Hewitson, 1870)    iN  M T S

Adelotypa  W. Warren, 1895  

Adelotypa bolena (A. Butler, 1867)    iN  M T S

Adelotypa borsippa (Hewitson, 1863)    iN  M T S

Calociasma  Stichel, 1910  

Calociasma nycteus (Godman & Salvin, 1886)  Brown-and-white Metalmark  SE Mexico to Panama  M T S

Calociasma laius (Godman & Salvin, 1886)  Laius Metalmark  Guatemala to El Salvador  iN  M T S

Calociasma ictericum (Godman & Salvin, 1878)  Icterica Metalmark  Costa Rica & Panama  M T OI S

Joiceya  Talbot, 1928  

Joiceya praeclarus Talbot, 1928  TL: Brazil (MT)  M T

Joiceya fannia (Godman, 1903)  TL: Guyana  M T S

Joiceya satyroides (Lathy, 1932)  TL: Guyana  M T

Ahrenholzia  J. Hall, 2018  

Ahrenholzia pimpilala J. Hall, 2018  TL: Ecuador  M T

Pandemos  Hübner, [1819]  

Pandemos pasiphae (Cramer, 1775)    iN  M T S

Zelotaea  H. Bates, 1868  

Zelotaea cindra (Staudinger, [1887])  TL: Brazil (Am)  M T

Zelotaea acuta (Brévignon & Gallard, 1999)  TL: French Guiana

Zelotaea costalis (H. Bates, 1868)  TL: Brazil (Am)

Zelotaea costalis costalis (H. Bates, 1868)  TL: Brazil (Am)  M T S

Zelotaea costalis gwiyanensis (Brévignon, 1999)  TL: French Guiana

Zelotaea grosnyi (Le Cerf, 1958)  TL: Brazil (Pa)  M S

Zelotaea brunescens P. Jauffret & J. Jauffret, 2007  TL: Brazil (Pa)

Zelotaea suffusca Brévignon & Gallard, 1993  TL: French Guiana

Zelotaea lya Lathy, 1958  TL: Amazon?  M S

Zelotaea alba Gallard & Brévignon, 1989  TL: French Guiana  M T S

Zelotaea spectra J. Hall, 2018  TL: Ecuador  M T

Zelotaea phasma H. Bates, 1868    iN

Zelotaea phasma phasma H. Bates, 1868    iN  M T S

Zelotaea phasma serena Brévignon, 2021  TL: French Guiana

Zelotaea anushkae Brévignon, 2021  TL: French Guiana

Zelotaea eidothea A. Butler, 1873  TL: Brazil (RJ)  M T S

Dysmathia  H. Bates, 1868  

Dysmathia portia H. Bates, 1868  TL: Brazil (Pa)  M T S

Pseudolivendula  J. Hall, 2018  

Pseudolivendula byzeres (Hewitson, 1872)  TL: Brazil (Pa)  M T OI TI S

Pseudolivendula candace (H. Druce, 1904)  TL: Brazil (RJ)  M T

Pseudolivendula wia (Brévignon & Gallard, 1992)  TL: French Guiana  M T

Pseudolivendula hemileuca (H. Bates, 1868)    iN

Pseudolivendula hemileuca hemileuca (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Pseudolivendula hemileuca sagittaria (Brévignon, 2012)  TL: French Guiana  M T

Pseudolivendula simplaris (Stichel, 1911)  TL: Brazil (Am)  M T

Pseudolivendula subtilis (Brévignon, 2012)  TL: French Guiana  M T

Pseudolivendula temperans (Brévignon, 2012)  TL: French Guiana  M T

Pseudolivendula tenuis Brévignon, 2019  TL: French Guiana  M T

Diminutiva  J. Hall, 2018  

Diminutiva glaucina (Stichel, 1928)  TL: Brazil (Am)  M T S

Rodinia  Westwood, 1851  

Rodinia calphurnia (Saunders, 1850)    iN  M T S

Rodinia justinia Gallard, 2017  TL: Peru  M S

Hypophylla  Boisduval, 1836  

Hypophylla lasthenes (Hewitson, 1870)  TL: Nicaragua

Hypophylla lasthenes lasthenes (Hewitson, 1870)  TL: Nicaragua  M T S

Hypophylla lasthenes idae Callaghan, 2001  TL: Colombia  M T

Hypophylla zeurippa Boisduval, 1836  Zeurippa Metalmark  W Mexico to Panama  iN  M T S1 S2 Ha

Hypophylla florus (Staudinger, [1887])  TL: Colombia  M T S

Hypophylla sudias (Hewitson, [1858])  White-banded Metalmark  S Mexico to Panama  iN  M T S L

Hypophylla argenissa (Stoll, 1790)  Deep-blue Metalmark  Costa Rica to Colombia  iN  M T S

Argyraspila  J. Hall, 2018  

Argyraspila maeonoides (Godman, 1903)  TL: Guyana

Argyraspila maeonoides maeon (Godman, 1903)  TL: Colombia  M T

Argyraspila maeonoides maeonoides (Godman, 1903)  TL: Guyana  M T S

Argyraspila antonii (Brévignon, 1995)  TL: French Guiana  M S

Argyraspila urichi (S. May, 1972)  TL: Trinidad  M T

Argyraspila pirene (Godman, 1903)    iN  M T S

Argyraspila caecina (C. Felder & R. Felder, 1865)    iN  M T

Argyraspila napoensis (J. Hall & Harvey, 2004)  TL: Peru  M T S

Argyraspila boyi J. Hall, 2018  TL: Brazil (Am)

Argyraspila thara (Hewitson, 1858)    iN  M T S

Argyraspila alma J. Hall, 2018  TL: Ecuador  M T S

Argyraspila rhesa (Hewitson, 1858)    iN  M T S

Argyraspila tavakiliani (Brévignon & Gallard, 1995)  TL: French Guiana  M S

Argyraspila gyges (Stichel, 1911)    iN  M T S

Argyraspila cayapona (Stichel, 1916)  TL: Brazil (MT)  M T S

Argyraspila zeanger (Stoll, 1790)    iN  M S

Argyraspila cuprea (A. Butler, 1867)  TL: Brazil (Pa)  M T S

Argyraspila eupolemia (Godart, [1824])  nomen dubium

Calospila  Geyer, 1832  

Calospila rhodope (Hewitson, 1853)    iN

Calospila rhodope rhodope (Hewitson, 1853)  TL: Brazil (Am)  M T S V

Calospila rhodope amphis (Hewitson, 1870)  TL: Ecuador  M T S

Calospila parthaon (Dalman, 1823)    iN

Calospila parthaon pelarge (Godman & Salvin, 1878)  S Mexico to W Ecuador  iN  M T S

Calospila parthaon parthaon (Dalman, 1823)  TL: Brazil  M T S

Calospila furvolinea J. Hall, 2018    iN  M T

Calospila ignotaterra J. Hall, 2020  TL: Unknown

Calliona  H. Bates, 1868  

Calliona siaka (Hewitson, [1858])    iN  M T S

Calliona irene (Westwood, 1851)  TL: Brazil (Pa)  M T S

Calliona latona (Hewitson, 1853)    iN  M T S

Calliona delia (Stichel, 1911)  TL: Venezuela  M T S

Sanguinea  J. Hall, 2018  

Sanguinea alcmaeon (Hewitson, 1876)  Blue-rayed Metalmark  Costa Rica to W Ecuador  iN  M T S

Sanguinea cleomedes (Hewitson, 1870)  Cleomedes Metalmark  Nicaragua to Panama  iN  M T S

Sanguinea heliodora (Staudinger, [1887])    iN  M T S

Sanguinea luceres (Hewitson, 1870)  TL: Ecuador  M T S

Sanguinea cruentata (A. Butler, 1867)  TL: Brazil (Am)

Sanguinea cruentata cruentata (A. Butler, 1867)  TL: Brazil (Am)  M T

Sanguinea cruentata cumulata J. Hall, 2018  TL: Peru  M T S

Sanguinea cruentata helice (Godman, 1903)  TL: Brazil (Am)  M T

Catagrammina  H. Bates, 1868  

Catagrammina hedonisma J. Hall, 2018  TL: Peru  M T

Catagrammina tapaja (Saunders, 1859)    iN

Catagrammina tapaja tapaja (Saunders, 1859)  TL: Brazil (Pa)  M T S

Catagrammina tapaja augea J. Hall, 2018  TL: Peru  M T

Catagrammina tapaja eumaea J. Hall, 2018  TL: Peru  M T

Setabis  Westwood, 1851  

Setabis flammula (H. Bates, 1868)    iN  M T S

Setabis disparilis (H. Bates, 1868)    iN

Setabis disparilis disparilis (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Setabis disparilis notios J. Hall, 2018  TL: Brazil (Ro)  M T

Setabis myrtis (Westwood, 1851)    iN  M T S

Setabis gelasine (H. Bates, 1868)  TL: Brazil (Am)  M T S

Setabis butleri (H. Bates, 1868)  TL: Brazil (Pa)  M T S

Setabis pythioides (A. Butler, 1867)    iN  M T S L

Setabis lagus (Cramer, 1777)    iN

Setabis lagus lagus (Cramer, 1777)  TL: Surinam  M T S IL L Txt

Setabis lagus jansoni (A. Butler, 1870)  S Mexico to Panama  M T S L

Setabis buckleyi (Grose-Smith, 1898)    iN  M T S

Setabis epitus (Cramer, 1780)    iN  M T S

Setabis velutina (A. Butler, 1867)    iN  M T S

Setabis phaedon (Godman, 1903)    iN  M T S

Setabis plagiaria (Grose-Smith, 1902)    iN  M T S

Setabis serica (Westwood, 1851)  TL: Brazil (Pa)

Setabis serica serica (Westwood, 1851)  TL: Brazil (Pa)  M T S L

Setabis serica extensa (Lathy, 1932)  TL: Peru  M T S

Setabis brunneus J. Hall, 2018  TL: Peru

Aricoris  Westwood, 1851  

Aricoris constantius (Fabricius, 1793)  Brazil  iN  M S

Aricoris emeryi Callaghan, Lemes & Kaminski, 2020  Brazil  iN

Aricoris tutana (Godart, [1824])  Brazil  iN  M T S

Aricoris monotona Stichel, 1910  S Brazil  iN  M T S

Aricoris terias (Godman, 1903)  Paraguay  iN  M T S

Aricoris middletoni (Sharpe, 1890)  Brazil  iN  M T S

Aricoris colchis (C. Felder & R. Felder, 1865)  S Brazil  M T S

Aricoris arenarum (Schneider, 1937)  Uruguay; ref.142  iN  M S V

Aricoris notialis (Stichel, 1910)  Argentina  iN  M T S

Aricoris schneideri Siewert, Dolibaina, O. Mielke & Casagrande, 2014  Brazil, Argentina, Uruguay  iN

Aricoris cinericia (Stichel, 1910)  TL: Argentina  M T S

Aricoris chilensis (C. Felder & R. Felder, 1865)  Bolivia, Argentina, Uruguay  iN  M T S

Aricoris erostratus (Westwood, 1851)  Erostratus Metalmark  Panama to Venezuela  iN  M T S L

Aricoris domina (H. Bates, 1865)  Domina Metalmark  Costa Rica & Panama  M T S L

Aricoris aurinia (Hewitson, 1863)  Brazil  iN  M T

Aricoris incana (Stichel, 1910)  Peru to Argentina  iN  M T S

Aricoris gauchoana (Stichel, 1910)  [= montana] Argentina, Uruguay; ref.142  iN  M T

Aricoris indistincta (Lathy, 1932)  Argentina  iN  M T S

Aricoris caracensis (Callaghan, 2001)  Brazil  iN  M T S

Aricoris campestris (H. Bates, 1868)  Brazil  iN  M T S

Aricoris zachaeus (Fabricius, 1798)  French Guiana, a nomen dubium: is it epulus?  M T

Aricoris epulus (Cramer, 1775)  the Guianas & Brazil  iN  M T S

Aricoris propitia (Stichel, 1910)  Brazil  iN  M T S

Aricoris signata (Stichel, 1910)  S Brazil  iN  M T S

Aricoris hubrichi (Stichel, 1926)  Argentina, Uruguay  iN  M T

Aricoris [n. sp.#1] J. Hall & Harvey, MS  TL: Brazil (Pr)

Aricoris [n. sp.#2] J. Hall & Harvey, MS  TL: Bolivia

Ariconias  J. Hall & Harvey, 2002  

Ariconias albinus (C. Felder & R. Felder, 1861)  Albinus Metalmark  Panama to Venezuela  iN  M T S L

Ariconias glaphyra (Westwood, 1851)    iN  M T S L

Lemonias  Hübner, [1807]  

Lemonias zygia Hübner, [1807]    iN

Lemonias zygia zygia Hübner, [1807]  TL: Brazil  M T S

Lemonias zygia chea Hewitson, 1863  TL: Brazil (Am)  M T S

Lemonias egaensis (A. Butler, 1867)    iN  M T S

Lemonias stalachtioides (A. Butler, 1867)    iN  M T S

Lemonias sontella (Schaus, 1902)  TL: Brazil (Pr)  M T

Lemonias caliginea (A. Butler, 1867)  Butler's Metalmark  S Mexico to Guatemala  iN  M T S

Lemonias ochracea (Mengel, 1902)    iN  M T S

Lemonias theodora (Godman, 1903)  TL: Brazil (MT)  M T

Lemonias albofasciata (Godman, 1903)    iN  M T S

Thisbe  Hübner, [1819]  

Thisbe irenea (Stoll, 1780)  Irenia Metalmark  S Mexico to Brazil  iN  M T S1 S2 L I

Thisbe rupestre Callaghan, 2001  TL: Brazil (Go)  M T

Thisbe molela (Hewitson, 1865)    iN  M T S

Thisbe lycorias (Hewitson, [1853])  Banner Metalmark  E & W Mexico to Colombia & Peru  iN  M T S L I

Thisbe hyalina (A. Butler, 1867)    iN  M T S

Thisbe incubus (J. Hall, Lamas & Willmott, 2001)    iN  M T S

Thisbe ucubis (Hewitson, 1870)  Ucubis Metalmark  Costa Rica to Colombia  iN  M T S

Thisbe silvestre Kaminski, Iserhard & Freitas, 2015    iN

Juditha  Hemming, 1964  

Juditha odites (Cramer, 1775)    iN

Juditha odites odites (Cramer, 1775)  TL: Surinam  M T S

Juditha odites praeclarum (H. Bates, 1866)  Costa Rica to W Ecuador  M T S L

Juditha pulcherrima (A. Butler, 1867)    iN

Juditha pulcherrima pulcherrima (A. Butler, 1867)  TL: Peru  M T S

Juditha pulcherrima comparata (Stichel, 1911)  TL: Peru  M T

Juditha inambari J. Hall & Harvey, 2001    iN  M T

Juditha dorilis (H. Bates, 1866)  Dorilis Metalmark  Nicaragua to W Ecuador  M T S

Juditha naza J. Hall & Harvey, 2001  TL: Colombia  M T S

Juditha azan (Westwood, 1851)    iN

Juditha azan azan (Westwood, 1851)  TL: Brazil (Pa)  M T S

Juditha azan completa (Lathy, 1904)  TL: Peru  M T S

Juditha azan majorina Brévignon & Gallard, 1998  TL: Surinam  M T

Juditha molpe (Hübner, [1808])    iN  M T S

Juditha caucana (Stichel, 1911)  Molpe Metalmark  [ = records of molpe from region], SE Mexico to W Ecuador]  iN  M T S1 S2 L1 L2 I

Synargis  Hübner, [1819]  

Synargis ethelinda (Hewitson, 1870)    iN  M T

Synargis nymphidioides (A. Butler, 1872)  Greater Metalmark    iN

Synargis nymphidioides septentrionalis Callaghan, Llorente & Luis, 2007  S Mexico (S Oaxaca to C Veracruz)  M T S L

Synargis nymphidioides praedictum Llorente, Luis & Arellano, 2013  TL: Mexico

Synargis nymphidioides nymphidioides (A. Butler, 1872)  S Mexico (Chiapas) to Panama, Pacific slope  M T S L

Synargis palaeste (Hewitson, 1870)  Palaeste Metalmark  Costa Rica to Colombia  M T S L

Synargis soranus (Stoll, 1781)  [= orestessa]; ref.151  iN  M T OI S

Synargis arche (Hewitson, 1865)  [formerly orestessa]; ref.151  iN  M T S

Synargis abaris (Cramer, 1776)    iN  M T S

Synargis tytia (Cramer, 1777)    iN  M T S

Synargis calyce (C. Felder & R. Felder, 1862)    iN  M T S

Synargis mycone (Hewitson, 1865)  Rusty Metalmark  W Mexico to Nicaragua  iN  M T S1 S2 L I1 I2

Synargis galena (H. Bates, 1868)    iN  M T S

Synargis gela (Hewitson, [1853])    iN  M T S

Synargis agle (Hewitson, [1853])    iN  M T S

Synargis pittheus (Hoffmansegg, 1818)    iN  M T S

Synargis dirca (Stichel, 1911)  Dirca Metalmark  Panama  M T S

Synargis ochra (H. Bates, 1868)  Ochra Metalmark  S Mexico to Colombia, Peru  iN  M T S L

Synargis sylvarum (H. Bates, 1867)  TL: Brazil (Pa)  M T

Synargis chaonia (Hewitson, [1853])  TL: Brazil (Am)  M T S

Synargis regina Grishin, 2024  Peru; ref.154  M T

Synargis reginella Grishin, 2024  Lower Amazonian region; ref.154  M T

Synargis regulus (Fabricius, 1793)  South America  iN  M T S

Synargis attilius (Stichel, 1925)  SE Brazil; ref.154  M T S

Synargis flavicauda Grishin, 2024  Peru; ref.154  M T

Synargis tenebritorna Grishin, 2024  E Brazil; ref.154  M T

Synargis latidifa Grishin, 2024  Lower Amazonian region; ref.154  M T

Synargis gorpa Dolibaina, Dias, O. Mielke & Casagrande, 2013  TL: Brazil  M OI

Synargis ochrophlegma (Stichel, 1911)    iN  M T

Synargis bifasciata (Mengel, 1902)    iN  M T S HS V

Synargis perrupta (Seitz, 1932)  TL: Brazil (Go)  M T

Synargis axenus (Hewitson, 1876)    iN  M T S

Synargis fandanga Dolibaina, Dias, O. Mielke & Casagrande, 2013    iN  M T

Synargis rasqueada Dolibaina, Dias, O. Mielke & Casagrande, 2013    iN  M T

Synargis fenestrella (Lathy, 1932)    iN  M T S

Synargis paulistina (Stichel, 1910)    iN  M T S V

Synargis victrix (Rebel, 1901)    iN  M T S

Synargis phliasus (Clerck, 1764)    iN

Synargis phliasus velabrum (Godman & Salvin, 1878)  Honduras to Colombia  M T S

Synargis phliasus phliasus (Clerck, 1764)  TL: ?  M T S

Synargis phliasus maravalica (Seitz, 1913)  TL: Trinidad  M T

Synargis anapis (Godart, [1824])  Brazil; help please, if you know this species

Thenpea  J. Hall, 2018  

Thenpea penthea (Cramer, 1777)    iN  M T S

Annulata  J. Hall, 2018  

Annulata annulifera (Godman, 1903)    iN  M T S

Calicosama  J. Hall & Harvey, 2001  

Calicosama lilina (A. Butler, 1870)  White-posted Metalmark  S Mexico to Panama, Trinidad  iN  M T S L

Calicosama robbinsi J. Hall & Harvey, 2001  TL: Panama  M T S

Behemothia  J. Hall, 2000  

Behemothia godmanii (Dewitz, 1877)  Godman's Metalmark  E & W Mexico to Costa Rica  iN  M T S L

Archaeonympha  J. Hall, 1998  

Archaeonympha smalli J. Hall & Harvey, 1998  Panamanian Metalmark  Nicaragua to C & E Panama  iN  M T S L

Archaeonympha drepana (H. Bates, 1868)  TL: Brazil (Am)  M T

Archaeonympha urichi (Vane-Wright, 1994)  Vane-Wright's Metalmark  Trinidad, Panama  M S

Theope  E. Doubleday, 1847  

Theope nycteis (Westwood, 1851)  Eyed Theope  Costa Rica to Argentina, Guianas, Trinidad  iN  M T S L

Theope brevignoni (Gallard, 1996)  TL: French Guiana  M T S

Theope fayneli Gallard, 2002    iN  M T S

Theope philotes (Westwood, 1851)    iN  M T S

Theope tetrastigma H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope orphana (Stichel, 1911)  TL: Brazil (Am)  M T S

Theope hypoleuca H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope simplicia H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope azurea H. Bates, 1868  TL: Brazil (Am)  M T S

Theope archimedes (Fabricius, 1793)    iN

Theope archimedes archimedes (Fabricius, 1793)  TL: Indiis  M T S

Theope archimedes zyzyxoxyx D'Abrera, 1994  Costa Rica to Colombia & Venezuela  M T S

Theope matuta Godman & Salvin, 1897  Giant Theope  TL: Costa Rica

Theope matuta matuta Godman & Salvin, 1897  Costa Rica & Panama  M T S

Theope matuta heureka J. Hall, 1999  TL: Colombia  M T

Theope pseudopedias J. Hall, 1999  Hall's Theope  S Mexico to Costa Rica  M T S

Theope pedias Herrich-Schäffer, [1853]  Yellow-bottomed Theope  S Mexico to Brazil, Guianas, Trinidad  iN  M T S

Theope devriesi J. Hall & Willmott, 1996  DeVries’ Theope  S Mexico (Chiapas) to W Ecuador  iN  M T S I

Theope pepo Willmott & J. Hall, 1994  TL: Ecuador  M T S

Theope eudocia Westwood, 1851  Orange Theope  Honduras to Brazil, Guianas, Trinidad  iN  M T S1 S2 I

Theope acosma Stichel, 1910  Tawny Theope  Costa Rica to Brazil, Guiana  iN  M T S

Theope excelsa H. Bates, 1868  Elegant Theope  Panama to Amazonas, Guianas  M T S

Theope kingi J. Hall & Willmott, 1996  King's Theope  Panama  M T S

Theope barea Godman & Salvin, 1878  Panamanian Theope  SW Mexico, Nicaragua to Brazil, French Guiana  iN  M T S

Theope mundula Stichel, 1926  Stichel's Theope  Honduras to Brazil, French Guiana  M T S

Theope zafaran Brévignon, 2011  TL: French Guiana  M S

Theope aureonitens H. Bates, 1868  TL: Brazil (Am)  M T S

Theope galionicus Gallard & Brévignon, 1989  TL: French Guiana

Theope dabrerae J. Hall & Willmott, 1996  TL: Ecuador  M T

Theope sericea H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope lampropteryx H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope eurygonina H. Bates, 1868    iN  M T S

Theope speciosa Godman & Salvin, 1897  Glazed Theope  Costa Rica to W Ecuador  iN  M T S

Theope pieridoides C. Felder & R. Felder, 1865  White Theope  S Mexico to Brazil, French Guiana, Trinidad  iN  M T S

Theope cratylus Godman & Salvin, 1886  Scallop-patched Theope  S Mexico to Colombia  iN  M T S

Theope sisemina Seitz, 1920    iN

Theope sisemina sisemina Seitz, 1920  TL: Colombia  M T S

Theope sisemina tabacona J. Hall & Willmott, 1996  TL: Peru  M T

Theope antanitis (Hewitson, 1874)  TL: Bolivia  M T

Theope iani Willmott & J. Hall, 1994  TL: Ecuador  M T S

Theope villai Beutelspacher, 1981  West-Mexican Theope  W Mexico (GRO & MEX)  iN  M T S L Ha

Theope virgilius (Fabricius, 1793)  Blue-based Theope  E & W Mexico to Brazil, Guianas, Trinidad, Netherlands Antilles  iN  M T S1 S2 S3 L I1 I2

Theope eupolis Schaus, 1890  Veracruzan Theope  [= diores], W & S Mexico to Panama  iN  M T S

Theope nobilis H. Bates, 1868  TL: Brazil (Pa)  M T

Theope sanjuani D'Abrera, 1994  TL: Colombia  M T S

Theope publius C. Felder & R. Felder, 1861  Shaded Theope    iN

Theope publius publius C. Felder & R. Felder, 1861  Nicaragua to Peru & Venezuela  M T S L I

Theope publius incompositus J. Hall, 1999  W Mexico & Yucatan Peninsula to El Salvador  M T S L Ha

Theope syngenes H. Bates, 1868    iN  M T S

Theope terambus (Godart, [1824])    iN  M S

Theope bacenis Schaus, 1890  Curve-lined Theope  [= mania], E & W Mexico to Colombia  iN  M T S L

Theope basilea H. Bates, 1866  Straight-lined Theope  Costa Rica to W Ecuador  iN  M T S L

Theope comosa Stichel, 1911  TL: Peru  M T S

Theope zostera H. Bates, 1868  TL: Brazil (Am)  M T S

Theope nodosus J. Hall, 1999  TL: Ecuador  M T

Theope phaeo Prittwitz, 1865  Falcate Theope  S Mexico (CHIS) to Brazil, Guianas, Trinidad  iN  M T S L I

Theope theritas Hewitson, 1860  TL: Brazil (Am)  M T

Theope thootes Hewitson, 1860    iN  M T S

Theope apheles H. Bates, 1868  TL: Brazil (Am)  M T S

Theope sobrina H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope janus H. Bates, 1867  TL: Brazil (Ma)  M T S

Theope guillaumei Gallard, 1996    iN

Theope guillaumei guillaumei Gallard, 1996  TL: French Guiana  M T

Theope guillaumei cecropia DeVries & J. Hall, 1996  Costa Rica  M T L

Theope discus H. Bates, 1868  TL: Brazil (Pa)  M T S

Theope thestias Hewitson, 1860    iN  M T S

Theope decorata Godman & Salvin, 1878  Painted Theope  Nicaragua to Brazil, French Guiana  iN  M T S

Theope lycaenina H. Bates, 1868  Gray Theope  Costa Rica to Brazil, Guianas, Trinidad  iN  M T S L

Theope leucanthe H. Bates, 1868  Faded Theope  Panama to Peru, Brazil, French Guiana, Trinidad, Venezuela  iN  M T S

Theope amicitiae J. Hall, Gallard & Brévignon, 1998  TL: French Guiana  M T

Theope christiani J. Hall & Willmott, 1999  TL: Guyana  M T

Theope methemona H. Bates, 1868  TL: Brazil (Pa)  M T

Theope sticheli J. Hall, 1998  TL: Brazil (Pa)  M T

Theope busbyi J. Hall, 1998  TL: Ecuador  M T

Theope turneri J. Hall & Austin, 1997  TL: Brazil (Rd)  M T

Theope batesi J. Hall, 1998  TL: Brazil (Am)  M T S

Theope atima H. Bates, 1868  TL: Brazil (Am)  M T

Theope foliorum H. Bates, 1868  Plain Theope  Panama to Peru, Brazil & Paraguay, Guianas, Trinidad  iN  M T S

Theope wallacei J. Hall, 1998  Wallace's Theope  Panama to Peru, French Guiana  M T S

Theope pakitza J. Hall & Harvey, 1998  TL: Peru  M T

Theope tetrastigmoides J. Hall, 2008  ref.220

Theope harrisi J. Hall, 2008  ref.220

Theope euselasina J. Hall, 2008  ref.220

Theope johannispetreus Brévignon, 2010  TL: French Guiana

Theope martinae Brévignon, 2010  TL: French Guiana

Theope minialba Gallard, 2006  TL: French Guiana  M T

Theope palambala Gallard, 2009  TL: French Guiana  M T

Theope rochambellus Brévignon, 2010  TL: French Guiana  M T

Theope shaphir Brévignon, 2009  TL: French Guiana

Pseudotinea  J. Hall & Callaghan, 2003  

Pseudotinea volcanicus (Callaghan & Salazar, 1997)  TL: Colombia  M T

Pseudotinea eiselei Callaghan & J. Hall, 2003  TL: Argentina  M T

Pseudotinea gagarini Callaghan & J. Hall, 2003    iN  M T

Pseudotinea hemis (Schaus, 1927)    iN  M T V

Pseudotinea caprina (Hewitson, 1859)  TL: Brazil  M T

Petrocerus  Callaghan, 1979  

Petrocerus catiena (Hewitson, 1875)  Brazil; ref.12  iN  M T

Pachythone  H. Bates, 1868  

Pachythone erebia H. Bates, 1868  TL: Brazil (Am)

Pachythone erebia thalassina (Vargas & Salazar, 2018)  TL: Colombia

Pachythone erebia erebia H. Bates, 1868  TL: Brazil (Am)  M T S

Pachythone robusta Lathy, 1932  TL: Brazil (MT)  M T

Pachythone pasicles Hewitson, [1873]    iN  M T

Pachythone palades Hewitson, [1873]  TL: Brazil (ES)  M T S

Pachythone philonis Hewitson, [1873]  TL: Colombia  M T S

Pachythone distigma H. Bates, 1868  TL: Brazil (Pa)  M T S

Pachythone analuciae J. Hall, Furtado & DeVries, 1999  TL: Brazil (MT)  M T S

Pachythone lateritia H. Bates, 1868    iN

Pachythone lateritia lateritia H. Bates, 1868  TL: Brazil (Pa/Am)  M T S

Pachythone lateritia coccineata (Kaye, 1904)  TL: Trinidad  M T

Pachythone xanthe H. Bates, 1868    iN

Pachythone xanthe xanthe H. Bates, 1868  TL: Brazil (Am)  M T S

Pachythone xanthe mimula H. Bates, 1868  TL: Brazil (Pa)  M T

Pachythone xanthe thaumaria Stichel, 1911  TL: Guyana  M T

Pachythone gigas Godman & Salvin, 1878  Gigas Metalmark    iN

Pachythone gigas nigriciliata Schaus, 1913  S Mexico to Panama, Atlantic slope  M T S

Pachythone gigas gigas Godman & Salvin, 1878  Costa Rica & Panama, Pacific slope  M T S

Pachythone gigas ignifer Stichel, 1911  TL: Colombia  M T

Pachythone obstinata (J. Hall & Willmott, 1995)  Ecuador; ref.12  M T

Pachythone watkinsi (D'Abrera, 1994)  Peru; ref.12

Pachythone aenigmatica (Rodríguez, Salazar & Constantino, 2011)  Colombia; ref.12  M T

Pachythone bicolor (Godman & Salvin, 1886)  Yellowed Metalmark  Panama, [Costa Rica?]  iN  M T OI

Pachythone conspersa Stichel, 1926    iN  M T S

Pachythone sumare Callaghan, 1999  TL: Brazil (RJ)  M T

Pachythone rubigo (H. Bates, 1868)  TL: Brazil (Pa)  M T

Pachythone crepusculi Gallard & Fernandez, 2017  TL: French Guiana

Pachythone strati (Kaye, 1925)  TL: Trinidad  M T S

Pachythone velazquezi (Beutelspacher, 1976)  Brown-tipped Metalmark  SW Mexico to Guatemala  M T S Ha

Pachythone lencates (Hewitson, 1875)  Lencates Metalmark  Costa Rica to Brazil, Trinidad; ref.11  M T S

Pachythone flocculus (Brévignon & Gallard, 1993)  ref.11  M T

Pachythone floccus (Brévignon, 2013)  ref.11

Pachythone heberti (P. Jauffret & J. Jauffret, 2007)  ref.11

Pachythone marajoara (P. Jauffret & J. Jauffret, 2007)  ref.11

Pseudonymphidia  Callaghan, 1985  

Pseudonymphidia agave (Godman & Salvin, 1886)  Agave Metalmark    iN

Pseudonymphidia agave agave (Godman & Salvin, 1886)  S Mexico to Panama  M T S

Pseudonymphidia agave leucogonia (Stichel, 1911)  TL: Colombia  M T S

Pseudonymphidia clearista (A. Butler, 1871)  Clearista Metalmark  W Mexico to Panama  iN  M T S

Minstrellus  J. Hall, 2007  

Minstrellus grandis (Callaghan, 1999)  base of E Andes: Peru, W Brazil & Bolivia  iN  M T S

Minstrellus nivosa (Stichel, 1929)  base of E Andes in N Peru  M T

Minstrellus emphatica (Stichel, 1911)  base of E Andes: S Colombia to Bolivia, wet lowland forest <1000m  M T S

Minstrellus leucotopus (Stichel, 1911)  E Andes: S Peru to E Bolivia, lower montane forest 1000-1500 m  M T

Protonymphidia  J. Hall, 2000  

Protonymphidia senta (Hewitson, 1853)    iN  M T S

Stalachtis  Hübner, 1818  

Stalachtis calliope (Linnaeus, 1758)    iN

Stalachtis calliope calliope (Linnaeus, 1758)  TL: Indiis  M T S

Stalachtis calliope bicoler Staudinger, [1887]  TL: Peru  M T

Stalachtis calliope voltumna Stichel, 1911  TL: Ecuador  M T S

Stalachtis eugenia (Cramer, 1777)    iN

Stalachtis eugenia eugenia (Cramer, 1777)  TL: Surinam  M T S

Stalachtis eugenia terpsichore Brévignon & Gallard, 1999  TL: French Guiana  M T

Stalachtis magdalena (Westwood, [1851])    iN

Stalachtis magdalena magdalena (Westwood, [1851])  TL: Colombia  M T S

Stalachtis magdalena cleove Staudinger, 1888  Panama to Colombia  M T S L

Stalachtis phlegia (Cramer, 1779)    iN

Stalachtis phlegia phlegia (Cramer, 1779)  TL: Surinam  M T S

Stalachtis phlegia nocticoelum Seitz, 1917  TL: Brazil (Pa)  M T

Stalachtis phlegia phlegetontia (Perty, 1833)  TL: Brazil  M T

Stalachtis phlegia susanna (Fabricius, 1787)    iN  M T S

Stalachtis phlegia venezolana Seitz, 1917  TL: Venezuela  M T

Stalachtis halloweeni J. Hall, 2006  TL: Guyana

Stalachtis halloweeni miranda Costa & Fratello, 2019  TL: Venezuela

Stalachtis halloweeni halloweeni J. Hall, 2006  TL: Guyana  M S L

Stalachtis euterpe (Linnaeus, 1758)    iN

Stalachtis euterpe euterpe (Linnaeus, 1758)  TL: America  M S HS V

Stalachtis euterpe adelpha Staudinger, 1888  TL: Brazil (Am)  M T

Stalachtis euterpe latefasciata Staudinger, 1888  TL: [Peru]  M T S

Stalachtis phaedusa Hübner, [1818]    iN

Stalachtis phaedusa phaedusa Hübner, [1818]  TL: Brazil  M T S

Stalachtis phaedusa duvalii (Perty, 1833)  TL: Brazil  M T

Stalachtis phaedusa exul Seitz, 1917  TL: French Guiana  M T

Stalachtis phaedusa phaloe Staudinger, [1887]  TL: Peru  M T

Stalachtis phaedusa trangeri Schaus, 1928  TL: Colombia  M T

Stalachtis phaedusa zephyritis (Dalman, 1823)  TL: ?  M T S

Stalachtis lineata (Guérin-Méneville, [1844])    iN  M T S

  Subfamily Riodinidae-incertae-sedis

Riodinella  Durden & Rose, 1978  a fossil genus

Riodinella nympha Durden & Rose, 1978  Fossil from USA: CO, questionably a riodinid  M OI

Lithopsyche  A. Butler, 1889  a fossil genus

Lithopsyche antiqua A. Butler, 1889  Fossil from England, questionably a riodinid  M T

Genus-unknown1   ,  

Genus-unknown1 misenes (Cramer, 1777)  "Surinam", identity unclear, could be a blue  M OI

Genus-unknown2   ,  

Genus-unknown2 spondiae (Shaw, 1808)  "Surinam", identity unclear, probably a hairstreak  M OI OD

Genus-unknown3   ,  

Genus-unknown3 ethemon (Cramer, 1775)  "West Indies", identity unclear, was placed in Euselasia, but probably a hairstreak  M OI OD

Family LYCAENIDAE [Leach], [1815] – GOSSAMERWINGS

  Subfamily Miletinae

  Tribe Spalgini

Feniseca  Grote, 1869  

Feniseca tarquinius (Fabricius, 1793)  Harvester    iN

Feniseca tarquinius tarquinius (Fabricius, 1793)  TL: Indiis  M S L1 L2 L3 I Ha

Feniseca tarquinius novascotiae McDunnough, 1935  TL: Canada  M S

  Subfamily Lycaeninae

Lycaena  Fabricius, 1807  

Lycaena cupreus (W. H. Edwards, 1870)  Lustrous Copper    iN

Lycaena cupreus lapidicola J. Emmel & Pratt, 1998  CA Sierra Nevada from Nevada to Tulare-Inyo Cos.; similar in White Mts.  M T S L H

Lycaena cupreus cupreus (W. H. Edwards, 1870)  [= artemisia], N CA, C-E OR, ID, NE NV, MT, NW WY, N UT  M T S L1 L2 Ha

Lycaena cupreus henryae (Cadbury, 1937)  BC, ALTA, N WA, NW MT  M S L

Lycaena cupreus snowi (W. H. Edwards, [1881])  S Rocky Mts. of SE WY, CO, N NM; high Uinta and La Sal Mts., UT  M S L1 L2 Ha

Lycaena hypophlaeas (Linnaeus, 1761)  American Copper  [formerly phlaeas, restricted to the Old World]; ref.16  iN

Lycaena hypophlaeas hypophlaeas (Boisduval, 1852)  TL: USA (Ma)  M S Ab L1 L2 H

Lycaena hypophlaeas feildeni (M'Lachlan, 1878)  coastal tundra in Nunavut, NWT, YK, AK  M S L I

Lycaena hypophlaeas arethusa (Wolley-Dod, 1907)  TL: not specified  M S

Lycaena hypophlaeas weberi Kohler, 2007  TL: not specified  M S

Lycaena hypophlaeas arctodon Ferris, 1974  TL: not specified  M T S L Ha

Lycaena hypophlaeas shields Kemal & Koçak, 2007  [alpestris is preoccupied]  M T S

Tharsalea  Scudder, 1876  

Tharsalea arota (Boisduval, 1852)  Tailed Copper  ref.11  iN

Tharsalea arota arota (Boisduval, 1852)  TL: USA (Ca)  M T TI S L I

Tharsalea arota C CA interior Coast Range segregate (, )  M S L Ha

Tharsalea arota Willamette Valley segregate (, )  Willamette Valley, W Oregon  M S

Tharsalea arota virginiensis (W. H. Edwards, 1870)  TL: USA (Nv)  M T S L1 L2 H

Tharsalea arota nubila J. A. Comstock, 1926  TL: USA (Ca)  M T S

Tharsalea arota schellbachi (Tilden, 1955)  TL: USA (Az)  M S L1 L2 Ha

Tharsalea hermes (W. H. Edwards, 1870)  Hermes Copper  ref.11  iN  M T S L1 L2 L3 L4 H

Tharsalea dione (Scudder, 1868)  Gray Copper  ref.11  iN  M S L Ha

Tharsalea editha (Mead, 1878)  Edith's Copper  ref.11  iN

Tharsalea editha editha (Mead, 1878)  TL: USA (Ca)  M T S L1 L2 L3 Ha

Tharsalea editha pseudonexa (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S

Tharsalea editha obscuramaculata (Austin, 1989)  TL: not specified  M T S L Ha

Tharsalea editha NE Oregon segregate (, )  [nr. editha], NE Oregon  M S

Tharsalea editha vurali (Koçak, 1984)  TL: not specified  M S L1 L2 L3 Ha

Tharsalea xanthoides (Boisduval, 1852)  Great Copper  ref.11  iN

Tharsalea xanthoides nigromaculata (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S1 S2 L

Tharsalea xanthoides xanthoides (Boisduval, 1852)  TL: USA (Ca)  M T S L1 L2

Tharsalea xanthoides obsolescens (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S Ha

Tharsalea gorgon (Boisduval, 1852)  Gorgon Copper  ref.11  iN

Tharsalea gorgon gorgon (Boisduval, 1852)  TL: USA (Ca)  M T S Ab L1 L2 I Ha

Tharsalea gorgon micropunctata (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S L Ha

Tharsalea gorgon jacquelineae (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S Ab L1 L2 I Ha

Tharsalea gorgon dorothea (J. Emmel & Pratt, 1998)  TL: USA (Or)  M T S L1 L2 Ha

Tharsalea rubidus (Behr, 1866)  Ruddy Copper  ref.11  iN

Tharsalea rubidus monachensis (K. Johnson & Balogh, 1977)  TL: USA (Ca)  M S L1 L2

Tharsalea rubidus incana (Austin, 1998)  TL: USA (Nv)  M T

Tharsalea rubidus rubidus (Behr, 1866)  [= duofascies]  M S1 S2 L

Tharsalea rubidus duofacies (K. Johnson & Balogh, 1977)  TL: USA (Id)  M T

Tharsalea rubidus perkinsorum (K. Johnson & Balogh, 1977)  TL: USA (Or)  M T S L

Tharsalea rubidus sirius (W. H. Edwards, 1871)  TL: USA (Co)  M T S L1 L2 L3 Ha

Tharsalea rubidus longi (K. Johnson & Balogh, 1977)  TL: USA (Ne)  M T S

Tharsalea rubidus ferrisi (K. Johnson & Balogh, 1977)  TL: USA (Az)  M T S L1 L2 I

Tharsalea heteronea (Boisduval, 1852)  Blue Copper  ref.11  iN

Tharsalea heteronea heteronea (Boisduval, 1852)  TL: USA (Ca)  M T TI S L

Tharsalea heteronea northi (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S I

Tharsalea heteronea submaculata (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S L Ha

Tharsalea heteronea clara (H. Edwards, [1880])  TL: USA (Ca)  M T S L

Tharsalea heteronea austin (Kemal & Koçak, 2007)  [rutila is preoccupied]  M T S L1 L2

Tharsalea heteronea rava (Austin, 1998)  TL: USA (Nv)  M T S L Ha

Tharsalea heteronea klotsi (W. D. Field, 1936)  TL: not specified  M S L1 L2 Ha

Tharsalea heteronea gravenotata (Klots, 1930)  TL: USA (Co)  M S L1 L2 Ha

Tharsalea heteronea E base Cascades- boldly spotted segregate (, )  [= nr. gravenotata], E base Cascades far N CA, OR, far S WA  M S L1 L2 L3 Ha

Tharsalea hyllus (Cramer, 1775)  Bronze Copper  ref.11  iN  M T S L1 L2 L3 I1 I2 Ha

Tharsalea epixanthe (Boisduval & Le Conte, [1835])  Bog Copper  ref.11  iN

Tharsalea epixanthe epixanthe (Boisduval & Le Conte, [1835])  TL: USA (NJ)  M S L H

Tharsalea epixanthe phaedrus (G. Hall, 1924)  TL: Canada (NS)  M S

Tharsalea epixanthe michiganensis (Rawson, 1948)  TL: USA (Mi)  M S L

Tharsalea dorcas (W. Kirby, 1837)  Dorcas Copper  ref.11  iN

Tharsalea dorcas claytoni (A. E. Brower, 1940)  TL: USA (Me)  M S L Ha

Tharsalea dorcas michuron (Scott, 2006)  TL: USA (Mi)  M S L

Tharsalea dorcas dorcas (W. Kirby, 1837)  TL: not specified  M S L

Tharsalea dorcas arcticus (Ferris, 1977)  TL: Canada (Yt)  M T S L

Tharsalea dorcas florus (W. H. Edwards, [1884])  TL: not specified  M S L Ha

Tharsalea dorcas megaloceras (Ferris, 1977)  TL: USA (Wy)  M T S

Tharsalea dorcas caldera (Scott & Kondla, 2017)  TL: USA (WY)

Tharsalea dorcas sangremar (Scott, 2008)  TL: USA (NM)  M T

Tharsalea dospassosi (McDunnough, 1940)  Dos Passos' Copper  ref.11  iN  M S

Tharsalea helloides (Boisduval, 1852)  Purplish Copper  [= castro]; ref.11  iN  M T S L1 L2 L3 L4 L5 L6 I Ha1 Ha2

Tharsalea nivalis (Boisduval, 1869)  Lilac-bordered Copper  ref.11  iN

Tharsalea nivalis nivalis (Boisduval, 1869)  TL: USA (Ca)  M S Ab L1 L2

Tharsalea nivalis W Cascadian segregate (, )  [nr. bichroma]  M S L1 L2 L3 Ha

Tharsalea nivalis bichroma (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S L I Ha

Tharsalea nivalis warnermontana (J. Emmel & Pratt, 1998)  TL: USA (Ca)  M T S L1 L2 Ha

Tharsalea nivalis NE Oregon segregate (, )  [nr. warnermontana]  M S L Ha

Tharsalea nivalis praetexta (Austin, 1998)  TL: USA (Nv)  M T S L Ha

Tharsalea nivalis browni (dos Passos, 1938)  TL: USA (Id)  M S L1 L2 Ha

Tharsalea mariposa (Reakirt, 1866)  Mariposa Copper  ref.11  iN

Tharsalea mariposa mariposa (Reakirt, 1866)  TL: USA (Ca)  M S L1 L2 Ha

Tharsalea mariposa ximena (Pyle & Hammond, 2018)  TL: USA, CA  M S L Ha

Tharsalea mariposa marya (Pyle & Hammond, 2018)  W Siskiyous, Josephine Co., OR

Tharsalea mariposa cascadia (Pyle & Hammond, 2018)  TL: USA, OR

Tharsalea mariposa thea (Pyle & Hammond, 2018)  TL: USA, OR

Tharsalea mariposa penroseae (W. D. Field, 1938)  TL: USA (Wy)  M T S L1 L2 L3 Ha

Tharsalea mariposa sperlingi (Pyle & Hammond, 2018)  TL: Canada

Tharsalea mariposa iridescens (Pyle & Hammond, 2018)  TL: Canada

Tharsalea mariposa charlottensis (W. Holland, 1930)  TL: not specified  M S

Tharsalea mariposa junia (Pyle & Hammond, 2018)  TL: USA, WA

Tharsalea mariposa candia (Pyle & Hammond, 2018)  TL: USA, OR

Tharsalea mariposa makah (Pyle & Hammond, 2018)  TL: USA, WA

Iophanus  Draudt, 1920  

Iophanus pyrrhias (Godman & Salvin, 1887)  Guatemalan Copper  Chiapas and Guatemala  iN  M T S L

  Subfamily Theclinae

  Tribe Theclini

Hypaurotis  Scudder, 1876  includes Habrodais; ref.11,129

Hypaurotis crysalus (W. H. Edwards, 1873)  Colorado Hairstreak  ref.11  iN

Hypaurotis crysalus crysalus (W. H. Edwards, 1873)  C & S CO, NM, far W TX, AZ, SON  M T S L1 L2 L3 I Ha

Hypaurotis crysalus intermedia Austin, 1998  SE NV, SW UT  M T S

Hypaurotis crysalus citima (H. Edwards, 1881)  N UT; NW CO pops. intermediate towards nominate  M T S I Ha

Hypaurotis crysalus Mexican segregate ,  NE Mexico (COAH, NL, DGO), Oaxaca  M S L

Hypaurotis grunus (Boisduval, 1852)  Golden Hairstreak  ref.11  iN

Hypaurotis grunus lorquini (W. D. Field, 1938)  SW ORE (Siskiyous) to C CA Coast Ranges  M T S L1 L2 L3 I H Ha

Hypaurotis grunus grunus (Boisduval, 1852)  S CA & Sierra Nevada  M T S L

Hypaurotis grunus herri (W. D. Field, 1938)  Cascade Mountains, far S WA (Skamania Co.), OR, N OR Coast Range  M T S L1 L2 H Ha

Hypaurotis poodiae (J. Brown & Faulkner, 1982)  Baja Hairstreak  Baja California (Sierra San Pedro Martir)

Hypaurotis poodiae poodiae (J. Brown & Faulkner, 1982)  Baja California (Sierra San Pedro Martir)  M T S

Hypaurotis poodiae browni (Hsu, 2005)  TL: Mexico  M T

  Tribe Eumaeini

Eumaeus  Hübner, [1819]  

Eumaeus childrenae (G. Gray, 1832)  Great Cycadian  [= debora], E & S Mexico to Honduras  iN  M T S L I H

Eumaeus godartii (Boisduval, 1870)  White-tipped Cycadian  Nicaragua to W Ecuador  iN  M T S1 S2 L I1 I2

Eumaeus minyas (Hübner, [1809])  Minyas Cycadian  Colombia to Peru & C Brazil  iN  M T

Eumaeus toxea (Godart, [1824])  Mexican Cycadian  E & W Mexico to Nicaragua (S TX as very rare stray)  iN  M T S L1 L2

Eumaeus atala (Poey, 1832)  Atala  S FL, Bahamas, Cuba, Isla de Juventud  iN  M T S L I

Eumaeus toxana (Boisduval, 1870)  Toxana  Venezuela to Bolivia  iN  M T

Theorema  Hewitson, 1865  

Theorema sapho (Staudinger, 1888)  Banded Cycadian  Panama to Ecuador  iN  M T S HS V

Theorema eumenia Hewitson, 1865  Tailed Cycadian  SE Mexico to Ecuador  iN  M T S1 S2 I

Theorema pyrczia Bálint, 2010  Pyrcz's Tailed Cycadian  Venezuela  M T

Theorema dysmenia Draudt, 1919  Dark Cycadian  Colombia, ?Peru  iN  M T S

Paiwarria  Kaye, 1904  

Paiwarria antinous (C. Felder & R. Felder, 1865)  Felders' Hairstreak  S Mexico (CHIS) to Colombia  iN  M T S

Paiwarria telemus (Cramer, 1775)  Telemus Hairstreak  Colombia to S Brazil  iN  M S

Paiwarria venulius (Cramer, 1779)  Venulius Hairstreak  Colombia to S Brazil  iN  M T S

Paiwarria aphaca (Hewitson, 1867)  Aphaca Hairstreak  C to S Brazil and Paraguay  iN  M T S

Paiwarria episcopalis (Fassl, 1912)  Episcopal Hairstreak  Colombia  M T

Paiwarria chuchuvia J. Hall & Willmott, 2005  Chuchuvia Hairstreak  Ecuador; ref.223  iN  M T

Paiwarria umbratus (Geyer, 1837)  Thick-tailed Hairstreak  E & W Mexico to Costa Rica  iN  M T S L I

Mithras  Hübner, [1819]  

Mithras elis (Cramer, 1779)  Elis Mithras  E Ecuador & Peru to Surinam; ref.181  iN  M T

Mithras orobia (Hewitson, 1867)  Orobia Mithras  Ecuador to Bolivia & C Brazil; ref.181  iN  M T S

Mithras colombiensis (K. Johnson & Constantino, 1997)  Colombian Mithras  Colombia & S. Brazil; ref.181  iN  M T S

Mithras oroanna (Bálint, 2002)  Oroanna Mithras  E Ecuador to Peru; ref.181  iN  M T

Mithras orobiana (Hewitson, 1867)  Orobiana Mithras  Amazonian region of Brazil; ref.181  M T

Mithras orocana (H. Druce, 1912)  Orocana Mithras  Colombia to Peru; ref.181  iN  M T S

Mithras nautes (Cramer, 1779)  Nautes Mithras  Venezuela to Peru & C Brazil  iN  M T S

Mithras catrea (Hewitson, 1874)  Catrea Mithras  S Brazil; ref.181  iN  M T S

Mithras apuya (Busby & Robbins, 2017)  ref.181

Mithras azul (Busby, Robbins & Faynel, 2017)  ref.181  iN

Mithras vossoroca (Bálint & Moser, 2001)  Vossoroca Mithras  S. Brazil; ref.181  iN  M T

Mithras cosmo (Busby, Robbins & Faynel, 2017)  ref.181  iN

Mithras emma (Busby & Robbins, 2017)  ref.181  iN

Mithras grande (Busby, Robbins & Moser, 2017)  ref.181

Mithras honor (Busby, Robbins & J. Hall, 2017)  ref.181  iN

Mithras lilyana (Busby & Robbins, 2017)  ref.181

Mithras noemi (Busby & Robbins, 2017)  ref.181

Mithras sine (Busby & Robbins, 2017)  ref.181

Mithras transvesta Robbins & Busby, 2017  ref.181

Dabreras  Bálint, [2008]  

Dabreras teucria (Hewitson, 1868)  Double-patched Hairstreak  Venezuela to Peru & C Brazil  iN  M T S

Brangas  Hübner, [1819]  

Brangas moserorum Bálint & Faynel, 2008  Mosers' Brangas  Paraguay & S Brazil  iN  M T

Brangas neora (Hewitson, 1867)  Widespread Brangas  E & W Mexico to S Brazil  iN  M T S L I

Brangas rita (Goodson, 1945)  Brown Brangas  Panama & Colombia  iN  M T

Brangas dydimaon (Cramer, 1777)  Twin Brangas  Colombia, Trinidad to Bolivia & C Brazil  M T OI

Brangas coccineifrons (Godman & Salvin, 1887)  Black-veined Brangas  SE Mexico to Colombia  iN  M T S

Brangas torfrida (Hewitson, 1867)  Torfrida Brangas  C & S Brazil  iN  M S

Brangas carthaea (Hewitson, 1868)  Green-spotted Brangas  E Mexico to Ecuador  iN  M T S L

Brangas [n. sp.#1] Robbins, MS  TL: Costa Rica

Brangas getus (Fabricius, 1787)  Bright Brangas  SE Mexico to S Brazil, Trinidad  iN  M T S L

Brangas caranus (Stoll, 1780)  Caranus Brangas  Venezuela to S Brazil  iN  M S

Brangas [n. sp.#2] Robbins, MS  TL: Venezuela

Brangas rambutorum Bálint & Faynel, 2008  Rambuteau's Brangas  Colombia & Venezuela  iN  M T

Brangas neildonatus Bálint & Faynel, 2008  Neild's Brangas  Venezuela  M T

Brangas silumena (Hewitson, 1867)  Silumena Brangas  Paraguay & S Brazil  iN  M T

Brangas felderi (Goodson, 1945)  Felders' Brangas  Colombia & Ecuador  iN  M T

Brangas [n. sp.#3] Robbins, MS  TL: not specified

Brangas polonus Bálint, 2008  Pyrcz's Brangas  Peru & Bolivia  M T

Brangas insolitus Bálint & Faynel, 2008  Isolated Brangas  Ecuador  iN  M T L

Thaeides  K. Johnson, Kruse & Kroenlein, 1997  

Thaeides theia (Hewitson, 1870)  Brown-barred Hairstreak  SE Mexico to Bolivia & SE Brazil  iN  M T S

Thaeides muela (Dyar, 1913)  Muela Hairstreak  Ecuador to Chile  iN  M T

Thaeides goleta (Hewitson, 1877)  Goleta Hairstreak  Colombia  M T

Thaeides xavieri (Le Crom & K. Johnson, 1997)  Xavier's Hairstreak  Colombia  M T

Thaeides pyrczi (K. Johnson, Le Crom & Constantino, 1997)  Pyrcz's Hairstreak  Venezuela  M T

Enos  K. Johnson, Kruse & Kroenlein, 1997  

Enos falerina (Hewitson, 1867)  Falerina Hairstreak  S Mexico (CHIS) to S Brazil  iN  M T S L

Enos thara (Hewitson, 1867)  Thara Hairstreak  SE Mexico to S Brazil  iN  M T S L

Enos myrtea (Hewitson, 1867)  Myrtea Hairstreak  Colombia to C Brazil  iN  M T S

Enos mazurka (Hewitson, 1867)  Mazurka Hairstreak  Brazil  M T S

Enos polka Lamas & Robbins, 2009  Spotted Hairstreak  Ecuador to C Brazil  iN  M T S

Lamasina  Robbins, 2002  

Lamasina draudti (Lathy, 1926)  Draudt's Hairstreak  [= godmani], Guatemala to Colombia  iN  M T S L

Lamasina lathyi (Bálint, 2005)  Lathy's Hairstreak  Ecuador, Peru  iN  M T S

Lamasina ganimedes (Cramer, 1775)  Ganimedes Hairstreak  Amazonas & Guianas  iN  M T OI S HS

Airamanna  Bálint, 2009  

Airamanna columbia (Bálint, 2005)  Colombian Hairstreak  Colombia  iN  M T S

Airamanna rhaptissima (K. Johnson, 1991)  Rhaptissima Hairstreak  Colombia to Peru  iN  M T S

Airamanna rhapsodia (Bálint, 2005)  Rhapsodia Hairstreak  Bolivia  M T

Evenus  Hübner, [1819]  

Evenus regalis (Cramer, 1775)  Regal Hairstreak  E & W Mexico to S Brazil  iN  M S1 S2 L I1 I2

Evenus coronata (Hewitson, 1865)  Crowned Hairstreak  S Mexico (CHIS) to Ecuador  iN  M T S

Evenus felix Neild & Bálint, 2014  Felix's Hairstreak  Venezuela to Peru  iN  M T

Evenus gabriela (Cramer, 1775)  Gabriela Hairstreak  Colombia to C Brazil  iN  M OI S

Evenus batesii (Hewitson, 1865)  Bates’ Hairstreak  [= teresina], SE Mexico to S Brazil  iN  M T S L

Evenus candidus (H. Druce, 1907)  Candid Hairstreak  Panama to Amazonas  iN  M T S

Evenus sponsa (Möschler, 1877)  Sponsa Hairstreak  Surinam and Guianas  iN  M T OI S

Evenus sumptuosa (H. Druce, 1907)  Sumptuosa Hairstreak  Ecuador to S Brazil  iN  M T HS

Evenus tagyra (Hewitson, 1865)  Tagyra Hairstreak  Peru to C Brazil  iN  M T S

Evenus floralia (H. Druce, 1907)  Floral Hairstreak  Peru to C Brazil  iN  M T

Evenus temathea (Hewitson, 1865)  Timid Hairstreak  Panama to Guianas  M T S

Evenus satyroides (Hewitson, 1865)  Satyr Hairstreak  Colombia to S Brazil  iN  M T S

Evenus latreillii (Hewitson, 1865)  Latreille's Hairstreak  Paraguay & S Brazil  iN  M T S

Atlides  Hübner, [1819]  

Atlides halesus (Cramer, 1777)  Great Blue Hairstreak    iN

Atlides halesus halesus (Cramer, 1777)  SE USA to New England  M T S1 S2 L1 L2 I

Atlides halesus corcorani Clench, 1942  W OR, CA, SW USA, NW, NE & E Mexico  M T S L I Ha

Atlides halesus cynara (Godman & Salvin, 1887)  C Mexico to Oaxaca  M T S Ab

Atlides gaumeri (Godman, 1901)  White-tipped Hairstreak  E & W Mexico to Panama  iN  M T S L

Atlides [n. sp.#1] Robbins, MS  TL: Peru

Atlides dahnersi Bálint, Constantino & K. Johnson, 2003  Dahners' Hairstreak  W Colombia  M T

Atlides polybe (Linnaeus, 1763)  Black-veined Hairstreak  E & W Mexico to S Brazil & N Argentina  iN  M S L I

Atlides inachus (Cramer, 1775)  Spying Hairstreak  [= carpophora], SE Mexico to C Brazil  iN  M T S

Atlides [n. sp.#2] Robbins, MS  TL: Costa Rica

Atlides cosa (Hewitson, 1867)  Cosa Hairstreak  S Brazil  iN  M T

Atlides misma D'Abrera, 1995  Misma Hairstreak  C & S Brazil, Paraguay  iN  M T

Atlides atys (Cramer, 1779)  Atys Hairstreak  [= scamander], Panama to S Brazil  iN  M S L

Atlides halljasoni Bálint, Kertész & Wojtusiak, 2006  Jason Hall's Hairstreak  Ecuador  M T

Atlides bacis (Godman & Salvin, 1887)  Bacis Hairstreak  Belize to Peru & C Brazil  iN  M T S

Atlides carpasia (Hewitson, 1868)  Jeweled Hairstreak  E & W Mexico to Panama  iN  M T S Ma L

Atlides browni Constantino, Salazar & K. Johnson, 1993  Brown's Hairstreak  Colombia  M T

Atlides rustan (Stoll, 1790)  Rustan Hairstreak  SE Mexico to S Brazil  iN  M T S L

Atlides polama (Schaus, 1902)  Polama Hairstreak  C & S Brazil  iN  M T

Atlides havila (Hewitson, 1865)  Havila Hairstreak  Colombia to Peru  iN  M T

Atlides zava (Hewitson, 1878)  Zava Hairstreak  S Brazil  iN  M T

Atlides thargelia (Burmeister, 1878)  Thargelia Hairstreak  Bolivia to Argentina  iN  M T

Atlides [n. sp.#3] Robbins, MS  TL: Brazil (Ba)

Arcas  Swainson, 1832  

Arcas imperialis (Cramer, 1775)  Imperial Arcas  SE Mexico to Guianas, Bolivia, Argentina & S Brazil  iN  M T OI S L

Arcas ducalis (Westwood, 1852)  Ducal Arcas  S Brazil (ES to RS)  iN  M T S

Arcas cypria (Geyer, 1837)  Mexican Arcas  E & W Mexico to Colombia & Venezuela  iN  M T OI S1 S2 L I Ha

Arcas gozmanyi (Bálint, 2006)  Gozmány’s Arcas  Costa Rica to W Colombia & W Ecuador  iN  M T S

Arcas splendor (H. Druce, 1907)  Splendid Arcas  Andean Region of Colombia and E Ecuador  iN  M T OI

Arcas alleluia Bálint, 2002  Bare Arcas  Andean Region of Peru  M T OI S

Arcas jivaro Nicolay, 1971  Hidden Arcas  E slope of the Andes in Ecuador to Bolivia  M T G

Arcas delphia Nicolay, 1971  Nicolay's Arcas  E slope of Costa Rica to W Colombia & W Ecuador  iN  M T S

Arcas tuneta (Hewitson, 1865)  Frosted Arcas  E Colombia, E Ecuador, E Peru, Bolivia to Guianas and W & S Brazil  iN

Arcas tuneta tuneta (Hewitson, 1865)  E Colombia, E Ecuador, E Peru, Bolivia to W Brazil & Guianas  M T HS G

Arcas tuneta arcadia Bálint, 2002  S Brazil  M T

Pseudolycaena  Wallengren, 1858  

Pseudolycaena damo (H. Druce, 1875)  Sky-blue Hairstreak  E & W Mexico to Colombia, W Ecuador & Venezuela  iN  M T S L1 L2 I1 I2

Pseudolycaena marsyas (Linnaeus, 1758)  Marsyas Hairstreak  ?Nicaragua and Trinidad-Tobago to S Brazil & Argentina  iN  M T S L

Pseudolycaena cybele (Godman & Salvin, 1896)  St. Vincent Hairstreak  St. Vincent  M T S L

Lucilda  D'Abrera & Bálint, 2001  

Lucilda danaus (C. Felder & R. Felder, 1865)  ref.242  iN  M T

Lucilda margaritacea (Draudt, 1919)  ref.242  iN  M T

Aveexcrenota  Salazar & K. Johnson, 1997  

Aveexcrenota anna (H. Druce, 1907)  TL: Colombia  M T S

Theritas  Hübner, 1818  

Theritas mavors Hübner, 1818  Deep-green Hairstreak  SE Mexico to Guianas  iN  M T S1 S2 L I

Theritas triquetra (Hewitson, 1865)    iN  M T S

Theritas paupera (C. Felder & R. Felder, 1865)    iN  M T

Theritas drucei (Lathy, 1926)    iN  M T

Theritas harrietta (Weeks, 1901)    iN  M T

Theritas [n. sp.#1] Robbins, MS  TL: Peru

Theritas adamsi (H. Druce, 1909)  TL: Peru  M T

Theritas deniva (Hewitson, 1874)    iN  M T V

Theritas [n. sp.#2] Robbins, MS  TL: Brazil (SC)

Theritas [n. sp.#3] Robbins, MS  TL: Peru

Theritas crines (H. Druce, 1907)  Crines Hairstreak  Panama to Colombia  iN  M T S

Denivia  K. Johnson, 1992  

Denivia acontius (Goodson, 1945)  ref.242  iN  M T

Denivia arene (Goodson, 1945)  ref.242  iN  M T

Denivia augustinula (Goodson, 1945)  Augustinula Hairstreak  E & S Mexico to Panama; ref.242  iN  M T S L

Denivia augustula (W. F. Kirby, 1877)  Augustula Hairstreak  Nicaragua & Costa Rica; ref.242  iN  M T S I

Denivia chaluma (Schaus, 1902)  ref.242  iN  M T

Denivia curitabaensis (K. Johnson, 1992)  ref.242  iN  M T

Denivia hemon (Cramer, 1775)  Pale-clubbed Hairstreak  SE Mexico to S Brazil; ref.242  iN  M T S1 S2 L1 L2 I

Denivia lisus (Stoll, 1790)  Lisus Hairstreak  [= hisbon], SE Mexico to S Brazil; ref.242  iN  M T S L I

Denivia monica (Hewitson, 1867)  ref.242  iN  M T

Denivia phegeus (Hewitson, 1865)  ref.242  iN  M T S

Denivia theocritus (Fabricius, 1793)  Pearly Hairstreak  E & S Mexico to Colombia; ref.242  iN  M T S L

Denivia silma (Martins, Faynel & Robbins, 2016)  Martins et al. (2019, Zootaxa, 4563: 119-134); ref.242  iN

Denivia viresco (H. Druce, 1907)  ref.242  iN  M T S

Denivia espiritosanto Bálint & Moser, 2007  S Brazil  iN

Denivia boliboyerus (Bálint & Wojtusiak, 2006)  TL: Bolivia

Johnsonita  Salazar & Constantino, 1995  

Johnsonita auda (Hewitson, 1867)  Auda Hairstreak  Panama to Venezuela  iN  M T S

Johnsonita [n. sp.#1] Robbins, MS  TL: Ecuador

Johnsonita [n. sp.#2] Robbins, MS  TL: Ecuador

Johnsonita chlamydem (H. Druce, 1907)    iN  M T S

Johnsonita pardoa (D'Abrera, 1995)    iN  M T

Johnsonita [n. sp.#3] Robbins, MS  TL: Ecuador

Johnsonita [n. sp.#4] Robbins, MS  TL: Peru

Johnsonita [n. sp.#5] Robbins, MS  TL: Bolivia

Johnsonita catadupa (Hewitson, 1869)  TL: Ecuador  M T

Johnsonita [n. sp.#6] Robbins, MS  TL: Ecuador

Johnsonita [n. sp.#7] Robbins, MS  TL: Peru

Johnsonita johnsoni Salazar & Constantino, 1995  TL: Colombia  M T

Johnsonita [n. sp.#8] Robbins, MS  TL: Ecuador

Johnsonita [n. sp.#9] Robbins, MS  TL: Ecuador

Johnsonita assula (Draudt, 1919)  TL: Colombia  M T

Johnsonita carpia Bálint, Boyer & Pyrcz, 2021  TL: Ecuador

Johnsonita iacinta Bálint, Boyer & Pyrcz, 2021  TL: Peru

Johnsonita ianusca Bálint, Lorenc-Brudecka & Pyrcz, 2021  TL: Ecuador

Johnsonita johnbanksi Bálint, 2003  TL: Peru

Johnsonita oxalida Bálint, Boyer & Pyrcz, 2021  TL: Peru

Johnsonita subcunicula Bálint, Cerdeña & Pyrcz, 2021  TL: Peru

Johnsonita turquisca Bálint, Boyer & Pyrcz, 2021  TL: Ecuador

Johnsonita zubkova Bálint, Boyer & Lorenc-Brudecka, 2021  TL: Peru

Brevianta  K. Johnson, Kruse & Kroenlein, 1997  

Brevianta busa (Godman & Salvin, 1887)  White-scalloped Hairstreak  E & W Mexico to Costa Rica  iN  M T S L

Brevianta perpenna (Godman & Salvin, 1887)  Perpenna Hairstreak  Panama  M T OI

Brevianta undulata (Hewitson, 1867)    iN  M T S

Brevianta undulella (Strand, 1918)    iN  M T

Brevianta ematheon (Cramer, 1777)    iN  M S

Brevianta bathoryon Bálint & Wojtusiak, 2011  Bathory Hairstreak  Colombia & Ecuador  M T

Brevianta magnifica Salazar & Constantino, 2010  TL: Colombia  M T

Brevianta celelata (Hewitson, 1874)    iN  M T

Brevianta tolmides (C. Felder & R. Felder, 1865)  White-frosted Hairstreak  Nicaragua to Colombia  iN  M T S

Brevianta hyas (Godman & Salvin, 1887)  Hyas Hairstreak  C Mexico to Costa Rica  iN  M T S L

Brevianta saphonota (Constantino, Salazar & K. Johnson, 1993)  Colombia & Ecuador  M T S

Ianusanta  Bálint, 2011  

Ianusanta ianusi Bálint, 2011  Peru  M T

Micandra  Staudinger, 1888  

Micandra platyptera (C. Felder & R. Felder, 1865)  Platyptera Hairstreak  Costa Rica, Venezuela to Bolivia  iN  M T S L

Micandra ion (H. Druce, 1890)  TL: Colombia  M T

Micandra extrema (Draudt, 1919)    iN  M T

Micandra comae (H. Druce, 1907)    iN  M T

Micandra aegides (C. Felder & R. Felder, 1865)    iN  M T S

Micandra [n. sp.] Robbins, MS  Peru

Micandra tongida Clench, 1971  Tongida Hairstreak  C Mexico  iN  M T S L

Micandra cyda (Godman & Salvin, 1887)  Cyda Hairstreak  [= furina], S Mexico to Costa Rica  iN  M T S L

Micandra dignota (Draudt, 1919)    iN  M T

Micandra sylvana (Jörgensen, 1934)    iN  M T S

Micandra stephanieae Prieto, 2011  TL: not specified

Rhamma  K. Johnson, 1992  

Rhamma commodus (C. Felder & R. Felder, 1865)    iN  M T

Rhamma familiaris (K. Johnson, 1991)    iN  M T L

Rhamma oxida (Hewitson, 1870)    iN  M T

Rhamma inexpectata K. Johnson, 1992  TL: Peru  M T

Rhamma [n. sp.] Robbins, MS  Ecuador

Rhamma tyrrius (H. Druce, 1907)  TL: Ecuador  M T

Rhamma amethystina (Hayward, 1949)  TL: Argentina  M T

Rhamma adriana (K. Johnson, 1992)    iN  M T

Rhamma adunca (Draudt, 1919)  TL: Colombia  M T

Rhamma livida Torres & K. Johnson, 1997  TL: Colombia  M T

Rhamma aurugo (Draudt, 1919)    iN  M T

Rhamma shapiroi (K. Johnson, 1992)    iN  M T

Rhamma mishma (Hewitson, 1878)    iN  M T

Rhamma comstocki K. Johnson, 1992    iN  M T

Rhamma hybla (H. Druce, 1907)    iN  M T

Rhamma catamarca K. Johnson, 1992  TL: Argentina  M T

Rhamma arria (Hewitson, 1870)    iN  M T

Rhamma nigrasarotina K. Johnson, 1992  TL: Peru  M T

Rhamma tarma K. Johnson, 1992  TL: Peru  M T

Rhamma confusa (Jörgensen, 1934)  TL: Argentina

Rhamma lapazensis (K. Johnson, 1992)  TL: Bolivia  M T

Rhamma brunea (K. Johnson, 1992)  TL: Argentina  M T

Rhamma coquimbiensis (K. Johnson, 1992)  TL: Chile

Rhamma bilix (Draudt, 1919)  TL: Colombia  M T

Rhamma anosma (Draudt, 1919)    iN  M T

Rhamma magenta K. Johnson, 1992  TL: Peru  M T

Rhamma tolimensis (K. Johnson, 1992)  TL: Colombia  M T

Rhamma dawkinsi Prieto & Lorenc-Brudecka, 2017  TL: Colombia

Phothecla  Robbins, 2004  

Phothecla photismos (H. Druce, 1907)  TL: Ecuador  M T

Phothecla margarita (Draudt, 1920)  TL: Bolivia  M T

Phothecla thespia (Hewitson, 1870)  TL: Ecuador  M T

Salazaria  D'Abrera & Bálint, 2001  

Salazaria sala (Hewitson, 1867)    iN  M T S

Salazaria elizabetha (Salazar, Vélez & K. Johnson, 1997)  TL: Colombia  M T

Salazaria [n. sp.#1] Robbins, MS  TL: Ecuador

Salazaria [n. sp.#2] Robbins, MS  TL: Ecuador

Salazaria [n. sp.#3] Robbins, MS  TL: Ecuador

Salazaria [n. sp.#4] Robbins, MS  TL: Ecuador

Salazaria [n. sp.#5] Robbins, MS  TL: Peru

Timaeta  K. Johnson, Kruse & Kroenlein, 1997  

Timaeta timaeus (C. Felder & R. Felder, 1865)    iN  M T S

Timaeta christina Robbins & R. Busby, 2008    iN  M T

Timaeta molinopampa (Bálint & Wojtusiak, 2000)  TL: Peru

Timaeta werneri (Salazar, Vélez, Cardona & K. Johnson, 1997)    iN  M T

Timaeta matthewi R. Busby & Robbins , 2008  TL: Ecuador

Timaeta walteri R. Busby & Robbins , 2008    iN

Timaeta trochus (H. Druce, 1907)    iN  M T

Timaeta roberti R. Busby & Robbins , 2008  TL: Ecuador  M S

Timaeta aepea (Hewitson, 1874)  TL: Ecuador  M T

Timaeta cospata Robbins & R. Busby, 2008  TL: Peru

Timaeta eronos (H. Druce, 1890)  TL: Colombia  M T

Timaeta pilosa Robbins & R. Busby, 2008    iN  M S

Timaeta gabriela R. Busby & Robbins , 2008  TL: Ecuador

Timaeta romero Robbins & R. Busby, 2008  TL: Venezuela

Timaeta balzabamba (Goodson, 1945)  TL: Venezuela; Ecuador  M T S

Timaeta boyeropampa (Bálint & Wojtusiak, 2010)  TL: Peru

Temecla  Robbins, 2004  

Temecla tema (Hewitson, 1867)    iN  M T

Temecla paron (Godman & Salvin, 1887)  Paron Hairstreak  E Mexico to Panama  iN  M T S

Temecla [n. sp.#1] Robbins, MS  TL: Venezuela

Temecla [n. sp.#2] Robbins, MS  TL: Bolivia

Temecla [n. sp.#3] Robbins, MS  TL: Peru

Temecla heraclides (Godman & Salvin, 1887)  Heraclides Hairstreak  S Mexico (CHIS) to ...  M T OI

Temecla sergius (Godman & Salvin, 1887)  TL: Venezuela  M T

Temecla bennetti (Dyar, 1913)  TL: Peru  M T

Temecla peona (Hewitson, 1874)  TL: ?  M T

Ipidecla  Dyar, 1916  

Ipidecla miadora Dyar, 1916  West-Mexican Ipidecla  W Mexico (SIN to GRO)  iN  M T S L

Ipidecla schausi (Godman & Salvin, 1887)  Schaus’ Ipidecla  E Mexico to Paraguay  iN  M T S L

Ipidecla crepundia (H. Druce, 1909)  Whitened Ipidecla  Peru to Trinidad  iN  M T S L

Penaincisalia  K. Johnson, 1990  

Penaincisalia loxurina (C. Felder & R. Felder, 1865)    iN  M T S

Penaincisalia alcacera (Bálint, Attal, Boyer & Wojtusiak, 2004)  TL: Ecuador

Penaincisalia santamarta (Bálint & Wojtusiak, 2004)  TL: Colombia  M T

Penaincisalia chachapoya (Bálint & Wojtusiak, 2003)  Chachapoya Penelfin  TL: Peru

Penaincisalia tegulina (Bálint & Boyer, 2008)  Tegulina Penelfin    iN  M T

Penaincisalia atymna (Hewitson, 1870)    iN  M T S

Penaincisalia felizitas Bálint, 2004  Felizitas Penelfin  TL: Peru  M T

Penaincisalia sigsiga (Bálint, Le Crom & Wojtusiak, 2004)  Sigsig Penelfin  TL: Ecuador

Penaincisalia cillutincarae (Draudt, 1919)    iN  M T

Penaincisalia [n. sp.#1] Robbins, MS  TL: Peru

Penaincisalia browni (K. Johnson, 1992)    iN  M T S

Penaincisalia regala (Le Crom & K. Johnson, 1997)  TL: Colombia  M T

Penaincisalia elisabeth Prieto, 2010  Elisabeth's Penelfin  TL: Colombia  M T G

Penaincisalia saraha (K. Johnson, 1992)  TL: Ecuador  M T S G

Penaincisalia vittata (K. Johnson, 1992)  TL: Ecuador  M T S

Penaincisalia cuiva Prieto & Rodriguez, 2007  Cuiva Penelfin  TL: Colombia

Penaincisalia magnifica (K. Johnson, 1992)  TL: Colombia  M T

Penaincisalia purpurea (K. Johnson, 1992)    iN  M T

Penaincisalia amazona (Bálint & Wojtusiak, 2003)  TL: Peru

Penaincisalia caeruleanota J. Hall & Willmott, 2005  ref.221

Penaincisalia amatamba Prieto, Boyer, Le Crom & Bálint, 2007  Cuiva Penelfin  TL: Ecuador

Penaincisalia balzapamba (K. Johnson, 1992)  TL: Ecuador  M T

Penaincisalia ismaeli R. Busby & J. Hall, 2005  ref.221

Penaincisalia dominiquae (Bálint, Boyer & Prieto, 2007)  Dominique's Penelfin  TL: Peru  M T

Penaincisalia duviolsi (Bálint, Boyer & Wojtusiak, 2006)  Duviols' Penelfin  TL: Peru  M T

Penaincisalia oxapampa (Bálint, Boyer & Wojtusiak, 2006)  Oxapampa Penelfin  TL: Peru  M S

Penaincisalia molipampa (Bálint, Boyer & Wojtusiak, 2006)  Molinopampa Penelfin  TL: Peru

Penaincisalia falcatamba (Bálint, Boyer & Wojtusiak, 2006)  Sicklemargined Penelfin  TL: Peru

Penaincisalia swarthea (Le Crom & K. Johnson, 1997)    iN  M T

Penaincisalia bimediana K. Johnson, 1990  TL: Ecuador  M T S

Penaincisalia cyanomediana (K. Johnson, 1992)  TL: Ecuador  M T

Penaincisalia [n. sp.#2] Robbins, MS  TL: Ecuador

Penaincisalia contraloxurina (K. Johnson, 1992)  TL: Colombia  M T

Penaincisalia amatista (Dognin, 1895)    iN  M T S

Penaincisalia [n. sp.#3] Robbins, MS  TL: Peru

Penaincisalia albalineata (K. Johnson, 1992)    iN  M T

Penaincisalia [n. sp.#4] Robbins, MS  TL: Peru

Penaincisalia [n. sp.#5] Robbins, MS  TL: Peru

Penaincisalia [n. sp.#6] Robbins, MS  TL: Ecuador

Penaincisalia penai K. Johnson, 1990    iN  M T S

Penaincisalia downeyi K. Johnson, 1990  TL: Colombia  M T

Penaincisalia rawlinsi K. Johnson, 1990  TL: Peru  M T

Penaincisalia culminicola (Staudinger, 1894)    iN  M T

Penaincisalia biophot Bálint & Wojtusiak, 2008  Biophot Penelfin  TL: Peru  M T

Penaincisalia caudata K. Johnson, 1990  TL: Peru  M T

Penaincisalia alatus (H. Druce, 1907)    iN  M T

Penaincisalia [n. sp.#7] Lamas & Robbins, MS  TL: Peru

Penaincisalia [n. sp.#8] Lamas & Robbins, MS  TL: Peru

Penaincisalia [n. sp.#9] Lamas & Robbins, MS  TL: Peru

Penaincisalia [n. sp.#10] Lamas & Robbins, MS  TL: Peru

Penaincisalia [n. sp.#11] Lamas & Robbins, MS  TL: Peru

Penaincisalia [n. sp.#12] Lamas & Robbins, MS  TL: Bolivia

Penaincisalia aurulenta K. Johnson, 1990  TL: Peru  M T

Penaincisalia perezi Bálint, 2001    iN

Penaincisalia descimoni K. Johnson, 1990  TL: Peru  M T

Penaincisalia lamasi Bálint, 2001    iN

Penaincisalia [n. sp.#13] Lamas & Robbins, MS  TL: Peru

Penaincisalia juliae J. Hall & Willmott, 2005  ref.221  iN

Penaincisalia andreae R. Busby & J. Hall, 2005  ref.221

Penaincisalia libertada J. Hall, 2005  ref.221  iN

Penaincisalia alina Bálint, 2019  TL: Peru

Penaincisalia celosia Prieto, 2016  TL: Colombia

Penaincisalia fortunatas (Bálint, Boyer & Pyrcz, 2023)  TL: Peru

Lathecla  Robbins, 2004  

Lathecla mitzi Robbins, MS  Mitzi's Hairstreak  S Brazil (Rio de Janeiro and Paraná); ref.288

Lathecla latagus (Godman & Salvin, 1887)  Latagus Hairstreak  SE Mexico to W Ecuador  iN  M T OI

Lathecla mimula (Draudt, 1920)  Mimula Hairstreak  lowland forest of South America  iN  M T

Lathecla fernandezi Robbins & Busby, 2015  Fernandez's Hairstreak  Venezuela & Ecuador; ref.288

Lathecla vichai Robbins & Busby, 2015  Vichai's Hairstreak  Guatemala to Panama; ref.288

Lathecla carolyna Busby, 2015  Carolyn's Hairstreak  Ecuador; ref.288  iN

Lathecla winnie Robbins & Busby, 2015  Winnie's Hairstreak  Costa Rica (maybe Panama); ref.288  iN

Podanotum  Torres & K. Johnson, 1996  

Podanotum metallicus Torres & K. Johnson, 1996  Metallic Podelfin    iN  M T

Podanotum clarissimus J. Hall, Willmott & K. Johnson, 1996  Bright Podelfin  TL: Ecuador  M T

Podanotum melanissimum Bálint & Wojtusiak, 2006  Melanic Podelfin  TL: Ecuador  M T

Podanotum glorissimum Bálint & Wojtusiak, 2002  Glorious Podelfin  TL: Peru

Podanotum andrewneildi Bálint, 2001  Andrew Neild's Podelfin  TL: Venezuela  M T

Podanotum paramosa (Constantino & Salazar, 1998)  Paramosa Podelfin  TL: Colombia  M T

Podanotum vanewrighti (K. Johnson & Adams, 1993)  Vane-Wrighti's Podelfin    iN  M T

Podanotum aquaphilum Bálint & Attal, 2007  Hydrophilic Podelfin  TL: Ecuador  M T

Podanotum magnificum Bálint & Attal, 2007  Magnificent Podelfin  TL: Ecuador  M T

Podanotum [n. sp.#1] Robbins, MS  TL: Ecuador

Podanotum [n. sp.#2] Robbins, MS  TL: Ecuador

Podanotum salaeides (Draudt, 1919)  Salaeides Podelfin    iN  M T S

Podanotum pajaten Farfán, Cerdeña & Bálint, 2021  TL: Peru

Podanotum pulsar Prieto & Rodríguez, 2012  TL: Colombia

Busbiina  Robbins, 2004  

Busbiina bosora (Hewitson, 1870)    iN  M T

Thereus  Hübner, [1819]  

Thereus cithonius (Godart, [1824])  Pale-lobed Hairstreak  E & W Mexico to Argentina  iN  M T S L

Thereus brocki Robbins, Heredia & Busby, 2015  ref.290

Thereus lomalarga Robbins, Heredia & Busby, 2015  ref.290  iN

Thereus wojtusiaki Bálint, 2006    iN  M T

Thereus gabathana (Strand, 1918)  TL: Colombia  M T

Thereus molena (E. Jones, 1912)  TL: Brazil (Pr)  M T

Thereus [n. sp.#1] Robbins, MS  TL: Brazil (Ba)

Thereus guianivaga (K. Johnson, 1989)  TL: Venezuela  M T

Thereus [n. sp.#2] Robbins, MS  TL: Guatemala

Thereus [n. sp.#3] Robbins, MS  TL: Panama

Thereus lutzi (Huntington, 1932)  Lutz's Hairstreak  Panama  M T S

Thereus caltha (H. Druce, 1907)  TL: Brazil (Pa)  M T

Thereus enenia (Hewitson, 1867)  Enenia Hairstreak  Guianas to C Brazil  iN  M T

Thereus [n. sp.#4] Robbins, MS  Nicaragua to Panama

Thereus oppia (Godman & Salvin, 1887)  Oppia Hairstreak  Mexico to Nicaragua  iN  M T S L

Thereus [n. sp.#5] Robbins, MS  TL: Panama

Thereus orasus (Godman & Salvin, 1887)  Crimson-spot Hairstreak  Mexico to Panama  iN  M T S L

Thereus ortalus (Godman & Salvin, 1887)  Ortalus Hairstreak  Mexico to S Brazil  iN  M T S L I Ha

Thereus [n. sp.#6] Robbins, MS  TL: Ecuador

Thereus praxis (Godman & Salvin, 1887)  Praxis Hairstreak  Costa Rica to N Brazil  iN  M T S

Thereus pedusa (Hewitson, 1867)  Pedusa Hairstreak  Panama to Amazonas  iN  M T S

Thereus lausus (Cramer, 1779)  Lausus Hairstreak  S Mexico (CHIS) to Guianas, Amazonas  iN  M S L I

Thereus timoclea (Hewitson, 1870)  Timoclea Hairstreak  Panama to Ecuador  M T S

Thereus ismarus (Cramer, 1777)  TL: Surinam  M T

Thereus phalanthus (Stoll, 1780)  TL: Surinam  M T

Thereus genena (Hewitson, 1867)    iN  M T

Thereus tiasa (Hewitson, 1869)    iN  M T

Thereus [n. sp.#7] Robbins, MS  TL: Brazil (Am)

Thereus endera (Hewitson, 1867)    iN  M T S

Thereus columbicola (Strand, 1916)  TL: Colombia  M T

Thereus eryssus (Herbst, 1800)  TL: Surinam  M S

Thereus illex (Schaus, 1902)  TL: Colombia  M T

Thereus [n. sp.#8] Robbins, MS  TL: Peru

Thereus pseudarcula (Giacomelli, 1914)    iN  M T

Rekoa  Kaye, 1904  

Rekoa meton (Cramer, 1779)  Meton Hairstreak  E & W Mexico to Bolivia, Paraguay, S Brazil, Venezuela to Guianas, Trinidad  iN  M T S L

Rekoa malina (Hewitson, 1867)    iN  M T

Rekoa palegon (Cramer, 1780)  Gold-bordered Hairstreak  S TX, E & W Mexico to N Argentina, Venezuela to Guianas, Trinidad, Tobago  iN  M T S L1 L2 I

Rekoa zebina (Hewitson, 1869)  Zebina Hairstreak  E & W Mexico to NW Costa Rica  iN  M T S

Rekoa bourkei (Kaye, [1925])  Jamaican Hairstreak  Jamaica  M S

Rekoa abeja (K. Johnson & Matusik, 1988)  Hispaniolan Hairstreak  Hispaniola  M T OI G

Rekoa marius (Lucas, 1857)  Marius Hairstreak  S TX, E & W Mexico to Bolivia, Paraguay, SE Brazil, Venezuela, Trinidad  iN  M T S L1 L2 I1 I2

Rekoa stagira (Hewitson, 1867)  Smudged Hairstreak  E & W Mexico to Paraguay, SE Brazil, Guianas, Trinidad, Venezuela  iN  M T S L

Arawacus  Kaye, 1904  

Arawacus togarna (Hewitson, 1867)  Togarna Hairstreak  SE Mexico to Amazonas  iN  M T S L1 L2 I

Arawacus lincoides (Draudt, 1917)    iN  M T S L

Arawacus aetolus (Sulzer, 1776)    iN  M T

Arawacus separata (Lathy, 1926)    iN  M T

Arawacus aethesa (Hewitson, 1867)  TL: Brazil (Ba)  M T

Arawacus sito (Boisduval, 1836)  Fine-lined Hairstreak  E & W Mexico to Costa Rica  iN  M T S1 S2 L I

Arawacus leucogyna (C. Felder & R. Felder, 1865)  Leucogyna Hairstreak  Belize to Venezuela & Peru  iN  M T S L

Arawacus meliboeus (Fabricius, 1793)    iN  M T

Arawacus jada (Hewitson, 1867)  Creamy Hairstreak  S AZ (as vagrant), E & W Mexico to Costa Rica  iN  M T S L I

Arawacus ellida (Hewitson, 1867)    iN  M T

Arawacus hypocrita (Schaus, 1913)  Pallid Hairstreak  E & S Mexico to Costa Rica  iN  M T S L

Arawacus dolylas (Cramer, 1777)  Dolylas Hairstreak  Panama to Surinam & S Brazil  iN  M T S L

Arawacus dumenilii (Godart, [1824])  Dumenil's Hairstreak  Honduras, Panama to Guianas, Trinidad  iN  M T S

Arawacus euptychia (Draudt, 1920)    iN

Arawacus tadita (Hewitson, 1877)    iN  M T

Arawacus binangula (Schaus, 1902)    iN  M T

Arawacus tarania (Hewitson, 1868)    iN  M T

Contrafacia  K. Johnson, 1989  

Contrafacia bassania (Hewitson, 1868)  White-etched Hairstreak  Mexico to Colombia  iN  M T S L

Contrafacia marmoris (H. Druce, 1907)    iN  M T

Contrafacia ahola (Hewitson, 1867)  Half-Blue Hairstreak  Mexico to Venezuela  iN  M T S L

Contrafacia imma (Prittwitz, 1865)  Imma Hairstreak  Mexico to S Brazil  iN  M T S L

Contrafacia francis (Weeks, 1901)    iN  M T

Contrafacia muattina (Schaus, 1902)    iN  M T

Contrafacia catharina (Draudt, 1920)    iN  M T

Contrafacia [n. sp.] Robbins, MS  Peru

Kolana  Robbins, 2004  

Kolana ligurina (Hewitson, 1874)  Ligurina Hairstreak  E & W Mexico to Brazil  iN  M T S1 S2 L

Kolana [n. sp.] Robbins, MS  Brazil (SC)

Kolana buccina (H. Druce, 1907)    iN  M T

Kolana lyde (Godman & Salvin, 1887)  Lyde Hairstreak  E & W Mexico to Venezuela  iN  M T S L

Kolana ergina (Hewitson, 1867)    iN  M T

Kolana chlamys (H. Druce, 1907)    iN  M T

Satyrium  Scudder, 1876  

Satyrium alcestis (W. H. Edwards, 1871)  Soapberry Hairstreak  ref.10  iN

Satyrium alcestis alcestis (W. H. Edwards, 1871)  TL: USA (Tx)  M T S L I

Satyrium alcestis oslari (Dyar, 1904)  TL: USA (Az)  M T S L I Ha

Satyrium fuliginosa (W. H. Edwards, 1861)  Western Sooty Hairstreak    iN

Satyrium fuliginosa fuliginosa (W. H. Edwards, 1861)  TL: USA (Ca)  M T TI S

Satyrium fuliginosa tildeni Mattoon & Austin, 1998  TL: USA (Ca)  M T1 T2 S L Ha

Satyrium fuliginosa albolineatum Mattoon & Austin, 1998  TL: USA (Ca)  M S

Satyrium fuliginosa E Klamath drainage segregate ,  CSNM, Jackson Co., OR, to vic Weed, Siskiyou Co. CA  M S L

Satyrium semiluna Klots, 1930  Sagebrush Sooty Hairstreak    iN

Satyrium semiluna semiluna Klots, 1930  TL: USA (Wy)  M S L I Ha

Satyrium semiluna maculadistinctum Mattoon & Austin, 1998  TL: USA (Nv)  M T S L1 L2

Satyrium semiluna Columbia Basin segregate ,  Columbia Basin, S BC to N OR  M S L1 L2 Ha

Satyrium semiluna Pine Mtn. segregate ,  Pine Mountain, Deschutes Co., OR; similar in Ochocos  M S L Ha

Satyrium semiluna C OR Cascades segregate ,  Cascades of C OR (e.g., Three Creeks Meadow, Deschutes Co.)  M S L1 L2 Ha

Satyrium semiluna Mt. Hood segregate ,  Mt. Hood, Clackamas / Hood River Cos., OR  M S L1 L2 Ha

Satyrium behrii (W. H. Edwards, 1870)  Behr's Hairstreak    iN

Satyrium behrii behrii (W. H. Edwards, 1870)  S CA, N through Sierra Nevada & Cascades to C WA  M S L1 L2 I H

Satyrium behrii columbia (McDunnough, 1944)  [often considered a synonym of S. b. behrii], Chelan Co. WA, northward  M S

Satyrium behrii northern intermontane segregate ,  NE OR, S ID, NE WY, N UT, NW CO  M S1 S2 L Ha

Satyrium behrii southern intermontane segregate ,  SE NV to C UT, S to C AZ  M S L

Satyrium behrii crossi (W. D. Field, 1938)  SE WY, E & S CO, N NM, far SE UT, far NE AZ  M T S L Ha

Satyrium acadica (W. H. Edwards, 1862)  Acadian Hairstreak    iN

Satyrium acadica acadica (W. H. Edwards, 1862)  TL: Canada (On)  M S L

Satyrium acadica montanensis (F. Watson & W. Comstock, 1920)  TL: USA (Mt)  M T S L Ha

Satyrium californica (W. H. Edwards, 1862)  California Hairstreak    iN

Satyrium californica californica (W. H. Edwards, 1862)  TL: USA (Ca)  M T S L1 L2 I Ha

Satyrium californica cygnus (W. H. Edwards, 1871)  TL: USA (Nv)  M T S L

Satyrium californica obscurafacies Austin, 1998  [= brashor], SE BC to NE NV  M T S1 S2 L1 L2 Ha

Satyrium californica wapiti M. Fisher, 2006  TL: USA (Co)  M T

Satyrium californica CO-WY Front Range segregate ,  Front Range of CO & SE WY  M S

Satyrium sylvinus (Boisduval, 1852)  Sylvan Hairstreak  ref.16  iN

Satyrium sylvinus nootka M. Fisher, 1998  TL: Canada (BC)  M T S L Ha

Satyrium sylvinus C Columbia Basin segregate ,  C Columbia Basin, WA  M S

Satyrium sylvinus sylvinus (Boisduval, 1852)  TL: USA (Ca)  M T TI S L Ha

Satyrium dryope (W. H. Edwards, 1870)  Dryope Hairstreak  ref.16

Satyrium dryope putnami (H. Edwards, 1877)  TL: USA (Ut)  M T S1 S2 L Ha

Satyrium dryope megapallidum Austin, 1998  TL: USA (Nv)  M T S L Ha

Satyrium dryope itys (W. H. Edwards, 1882)  TL: USA (Az)  M T S L

Satyrium dryope desertorum (F. Grinnell, 1917)  TL: USA (Ca)  M T S L I

Satyrium dryope dryope (W. H. Edwards, 1870)  TL: USA ([Ca])  M T S L

Satyrium titus (Fabricius, 1793)  Coral Hairstreak    iN

Satyrium titus titus (Fabricius, 1793)  TL: USA (Ga)  M S L1 L2

Satyrium titus winteri (Gatrelle, 2004)  TL: USA (Ma)  M T S1 S2 L

Satyrium titus campus (Gatrelle, 2004)  TL: USA (Ia)  M T S L I

Satyrium titus watsoni (W. Barnes & Benjamin, 1926)  TL: USA (Tx)  M T S L

Satyrium titus immaculosus (W. Comstock, 1913)  TL: USA (Ut)  M S L Ha

Satyrium titus carrizozo R. Holland, 2010  TL: USA (NM)

Satyrium titus occidentalis (Austin & J. Emmel, 1998)  TL: USA (Nv)  M T S L

Satyrium edwardsii (Grote & Robinson, 1867)  Edwards' Hairstreak    iN

Satyrium edwardsii edwardsii (Grote & Robinson, 1867)  TL: Canada (On)  M S L1 L2 I Ha

Satyrium edwardsii meridionale Gatrelle, 2001  TL: USA (SC)  M T1 T2 L

Satyrium calanus (Hübner, [1809])  Banded Hairstreak    iN

Satyrium calanus calanus (Hübner, [1809])  TL: ?  M S1 S2 L

Satyrium calanus falacer (Godart, [1824])  TL: USA (Pa)  M T S1 S2 L1 L2 L3 Ha

Satyrium calanus godarti (W. D. Field, 1938)  TL: USA (Co)  M T S L1 L2 Ha

Satyrium calanus albidus Scott, 1981  TL: USA (Co)  M T S1 S2

Satyrium caryaevorus (McDunnough, 1942)  Hickory Hairstreak    iN  M S L F

Satyrium kingi (Klots & Clench, 1952)  King's Hairstreak    iN  M S L1 L2 H

Satyrium liparops (Le Conte, 1833)  Striped Hairstreak    iN

Satyrium liparops floridensis Gatrelle, 2001  TL: USA (Fl)  M T1 T2 S L H

Satyrium liparops liparops (Le Conte, 1833)  TL: USA (Ga)  M T S L

Satyrium liparops strigosa (T. Harris, 1862)  TL: USA (Ma)  M S L Ha

Satyrium liparops fletcheri (Michener & dos Passos, 1942)  TL: Canada (Mb)  M T S

Satyrium liparops aliparops (Michener & dos Passos, 1942)  TL: USA (Co)  M S L

Satyrium auretorum (Boisduval, 1852)  Gold-hunter’s Hairstreak    iN

Satyrium auretorum auretorum (Boisduval, 1852)  TL: USA (Ca)  M T S L

Satyrium auretorum spadix (H. Edwards, 1881)  TL: USA (Ca)  M T S L H

Satyrium auretorum fumosum J. Emmel & Mattoni, 1990  TL: USA (Ca)  M T

Satyrium tetra (W. H. Edwards, 1870)  Mountain Mahogany Hairstreak  Baja California (N) to N CA  iN

Satyrium tetra Northern segregate ,  SE Oregon, NE CA, N W NV  M S

Satyrium tetra tetra (W. H. Edwards, 1870)  TL: USA (Ca)  M T S L1 L2 I Ha

Satyrium saepium (Boisduval, 1852)  Hedgerow Hairstreak    iN

Satyrium saepium saepium (Boisduval, 1852)  TL: USA (Ca)  M T TI S L Ha

Satyrium saepium fulvescens (H. Edwards, 1877)  TL: USA (Ca)  M T

Satyrium saepium chalcis (W. H. Edwards, 1869)  [= fulvescens]  M T S1 S2 L I

Satyrium saepium rubrotenebrosum J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S

Satyrium saepium caliginosum J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L1 L2 Ha

Satyrium saepium chlorophora (F. Watson & W. Comstock, 1920)  TL: USA (Ca)  M T L

Satyrium saepium subaridum J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L Ha

Satyrium saepium obscurofuscum Austin, 1998  TL: USA (Nv)  M T

Satyrium saepium latalinea Austin & Savage, 1998  TL: USA (Ut)  M T1 T2 S

Satyrium saepium provo (F. Watson & W. Comstock, 1920)  TL: USA (Ut)  M T S L Ha

Satyrium favonius (J. E. Smith, 1797)  Oak Hairstreak    iN

Satyrium favonius ontario (W. H. Edwards, 1868)  TL: Canada (On)  M S L Ha

Satyrium favonius autolycus (W. H. Edwards, 1871)  TL: USA (Tx)  M T S L1 L2 I

Satyrium favonius violae (D. Stallings & J. Turner, 1947)  TL: USA (NM)  M T S

Satyrium favonius favonius (J. E. Smith, 1797)  TL: USA (Ga)  M S1 S2 S3 Ab L Ha

Satyrium ilavia (Beutenmüller, 1899)  Ilavia Hairstreak    iN  M T S L1 L2 I Ha

Satyrium polingi (W. Barnes & Benjamin, 1926)  Poling’s Hairstreak    iN

Satyrium polingi organensis (Ferris, 1980)  TL: USA (NM)  M T

Satyrium polingi polingi (W. Barnes & Benjamin, 1926)  TL: USA (Tx)  M T S L Ha

Ocaria  Clench, 1970  

Ocaria clenchi (K. Johnson, 1992)  Clench's Hairstreak  Mexico, El Salvador  iN  M T S L

Ocaria [n. sp.#1] Robbins, MS  TL: Panama

Ocaria aholiba (Hewitson, 1867)  Colombian Hairstreak  Costa Rica to Venezuela  iN  M T S

Ocaria [n. sp.#2] Robbins, MS  TL: Brazil (SP)

Ocaria clepsydra (H. Druce, 1907)    iN  M T

Ocaria arpoxais (Godman & Salvin, 1887)  Blue-spotted Hairstreak  Mexico to Panama  M T S

Ocaria cinerea (Lathy, 1936)    iN  M T

Ocaria [n. sp.#3] Robbins, MS  TL: Guyana

Ocaria sadiei (Weeks, 1901)    iN  M T

Ocaria petelina (Hewitson, 1877)  Petelina Hairstreak  SE Mexico to Panama  M T S L

Ocaria elvira (K. Johnson, 1992)    iN  M T

Ocaria calesia (Hewitson, 1870)    iN  M T S

Ocaria elongata (Hewitson, 1870)    iN  M T

Ocaria thales (Fabricius, 1793)  Thales Blackstreak  Mexico to S Brazil and Argentina  iN  M T S L

Ocaria arcula (H. Druce, 1907)    iN  M T

Ocaria ocrisia (Hewitson, 1868)  Black Hairstreak  Mexico to S Brazil and Argentina  iN  M T S L1 L2 L3 I

Ocaria faurei Bálint, Attal & Costa, 2018  ref.191

Ocaria moseranki (D'Abrera, 2022)  TL: Brazil

Chlorostrymon  Clench, 1961  

Chlorostrymon simaethis (Drury, 1773)  Silver-banded Hairstreak    iN

Chlorostrymon simaethis sarita (Skinner, 1895)  Mexico to Panama  M T S L1 L2 L3 L4 L5 I H

Chlorostrymon simaethis simaethis (Drury, 1773)  S Florida, (most) Greater & Lesser Antilles, in S. Amer. to Argentina  M S L I

Chlorostrymon simaethis rosario Nicolay, 1980  TL: Ecuador  M T

Chlorostrymon simaethis jago (W. Comstock & Huntington, 1943)  Jamaica  M T S

Chlorostrymon kuscheli (Ureta, 1949)  TL: Chile  M T

Chlorostrymon maesites (Herrich-Schäffer, 1865)  Amethyst Hairstreak  S Florida, Greater & (most) Lesser Antilles  iN  M T S L

Chlorostrymon clenchi (W. Comstock & Huntington, 1943)  Dominican Hairstreak  Dominica  M T S

Chlorostrymon lalitae Brévignon, 2000  ref.178

Chlorostrymon telea (Hewitson, 1868)  Telea Hairstreak  S TX (stray) to S Brazil  iN  M T S L I

Chlorostrymon orbis K. Johnson & D. Smith, 1993  Orbis Hairstreak  Jamaica  M T OI

Magnastigma  Nicolay, 1977  

Magnastigma hirsuta (Prittwitz, 1865)  Hirsuta Hairstreak  Costa Rica to Brazil  iN  M T S

Magnastigma julia Nicolay, 1977  TL: Brazil (DF)  M T S

Magnastigma elsa (Hewitson, 1877)  Elsa Hairstreak  Mexico to Panama  iN  M T S L

Cyanophrys  Clench, 1961  

Cyanophrys goodsoni (Clench, 1946)  Goodson’s Greenstreak  S TX to Costa Rica  iN  M T S L H

Cyanophrys argentinensis (Clench, 1946)    iN  M T

Cyanophrys bertha (E. Jones, 1912)    iN  M T

Cyanophrys acaste (Prittwitz, 1865)    iN  M T

Cyanophrys amyntor (Cramer, 1775)  Amyntor Greenstreak  S Mexico to S Brazil  iN  M T S L

Cyanophrys fusius (Godman & Salvin, 1887)  Brown Greenstreak  E & W Mexico to Colombia  iN  M T S I

Cyanophrys herodotus (Fabricius, 1793)  Tropical Greenstreak  S TX, E & W Mexico to Argentina  iN  M T S L1 L2 H

Cyanophrys miserabilis (Clench, 1946)  Clench’s Greenstreak  S TX to Costa Rica  iN  M T S L

Cyanophrys velezi K. Johnson & Kruse, 1997  TL: Colombia  M T

Cyanophrys crethona (Hewitson, 1874)  Jamaican Greenstreak  Jamaica  M T S

Cyanophrys hartii T. Turner & J. Miller, 1992  Dallas Mountain Greenstreak  Jamaica  M T

Cyanophrys roraimiensis K. Johnson & D. Smith, 1993  TL: Brazil (Rr)  M T

Cyanophrys longula (Hewitson, 1868)  Mountain Greenstreak  E & W Mexico to Venezuela; one possible record for SE AZ  iN  M T S L1 L2 I Ha

Cyanophrys pseudolongula (Clench, 1944)    iN  M T

Cyanophrys agricolor (A. Butler & H. Druce, 1872)  Stained Greenstreak  E & W Mexico to Panama  iN  M T S L

Cyanophrys banosensis (Clench, 1944)    iN  M T

Cyanophrys remus (Hewitson, 1868)    iN  M T

Callophrys  Billberg, 1820  

Callophrys affinis (W. H. Edwards, 1862)  Western Green Hairstreak    iN

Callophrys affinis washingtonia Clench, 1944  TL: USA (Wa)  M T S L Ha

Callophrys affinis affinis (W. H. Edwards, 1862)  TL: USA (Wy)  M T S1 S2 L I1 I2 Ha

Callophrys affinis homoperplexa W. Barnes & Benjamin, 1923  TL: USA (Co)  M T S L Ha

Callophrys affinis albipalpus Gorelick, 2005  TL: USA (NM)  M T

Callophrys affinis apama (W. H. Edwards, 1882)  TL: USA (Az)  M T S L1 L2

Callophrys affinis chapmani Gorelick, 2005  Sonora & Durango  M T S

Callophrys dumetorum (Boisduval, 1852)  Lotus Hairstreak    iN

Callophrys dumetorum oregonensis Gorelick, [1969]  TL: USA (Wa)  M T S L Ha

Callophrys dumetorum dumetorum (Boisduval, 1852)  TL: USA (Ca)  M T S1 S2 L1 L2 I Ha

Callophrys dumetorum perplexa W. Barnes & Benjamin, 1923  TL: USA (Ca)  M T

Callophrys dumetorum superperplexa J. Emmel, T. Emmel & Mattoon, 1998  [possibly conspecific with C. sheridanii]  M T S L I

Callophrys viridis (W. H. Edwards, 1862)  Coastal' Green Hairstreak  [possibly conspecific with C. sheridanii]  iN  M T S1 S2 HS L I

Callophrys sheridanii (W. H. Edwards, 1877)  Sheridan's Hairstreak    iN

Callophrys sheridanii pseudodumetorum J. Emmel, T. Emmel & Mattoon, 1998  [= nr. dumetorum of Warren], W OR, NW CA Siskiyous low elev.  M T S L I H

Callophrys sheridanii newcomeri Clench, 1963  TL: USA (Wa)  M S1 S2 L

Callophrys sheridanii lemberti Tilden, 1963  TL: not specified  M T S L Ha

Callophrys sheridanii interrupta Austin, 1998  TL: USA (Nv)  M T S Ha

Callophrys sheridanii comstocki Henne, 1940  TL: USA (Ca)  M T S L I Ha

Callophrys sheridanii paradoxa Scott, 1986  TL: USA (Ut)  M T1 T2 S1 S2

Callophrys sheridanii neoperplexa W. Barnes & Benjamin, 1923  TL: USA (Ut)  M T S1 S2 I

Callophrys sheridanii W CO montane segregate ,   M S

Callophrys sheridanii sheridanii (W. H. Edwards, 1877)  TL: USA (Wy)  M S L

Callophrys sheridanii sacramento Scott, 2006  TL: USA (NM)  M S I

Callophrys gryneus (Hübner, [1819])  Juniper Hairstreak    iN

Callophrys gryneus sweadneri (F. Chermock, 1945)  TL: USA (Fl)  M T S L

Callophrys gryneus smilacis (Boisduval & Le Conte, [1835])  TL: USA (Ga)  M T S

Callophrys gryneus gryneus (Hübner, [1819])  TL: USA (Va)  M T S L1 L2

Callophrys gryneus castalis (W. H. Edwards, 1871)  TL: USA (Tx)  M T S1 S2 Ab L1 L2

Callophrys gryneus SE Coahuila segregate ,  SE Coahuila  M S

Callophrys gryneus siva (W. H. Edwards, 1874)  [= clenchi, turkingtoni]  M T S L1 L2 L3 L4 I Ha

Callophrys gryneus chalcosiva Clench, 1981  TL: USA (Ut)  M T1 T2 S1 S2 L1 L2 I1 I2 Ha

Callophrys gryneus Columbia Basin segregate ,  [nr. chalcosiva]  M S L Ha

Callophrys gryneus S Sierra Nevada on Juniperus segregate ,  S Sierra Nevada (Tulare Co.); on Juniperus; formerly "barryi"; similar in SB Mts  M S L

Callophrys gryneus Guadalupe Island segregate ,  nelsoni  M S

Callophrys gryneus nelsoni (Boisduval, 1869)  TL: USA (Ca)  M T TI S IL L1 L2

Callophrys gryneus plicataria K. Johnson, 1976  TL: Canada  M T1 T2 S L Ha

Callophrys gryneus rosneri K. Johnson, 1976  TL: Canada  M T S

Callophrys gryneus byrnei K. Johnson, 1976  TL: USA (Id)  M T

Callophrys gryneus juniperaria (J. A. Comstock, 1925)  TL: USA (Ca)  M T S L

Callophrys gryneus mansfieldi (Tilden, 1951)  TL: USA (Ca)  M T S L

Callophrys loki (Skinner, 1907)  Loki Hairstreak  Baja California Norte  iN

Callophrys loki loki (Skinner, 1907)  Baja California Norte  M T S1 S2 S3 L Ha

Callophrys loki thornei (J. Brown, 1983)  TL: USA (Ca)  M T1 T2 S L1 L2 I H

Callophrys loki cedrosensis (J. Brown & Faulkner, 1989)  TL: Mexico (BCN)  M T S L I Ha

Callophrys muiri (H. Edwards, 1881)  Muir's Hairstreak    iN  M T S1 S2 L

Callophrys hesseli (Rawson & Ziegler, 1950)  Hessel's Hairstreak    iN

Callophrys hesseli hesseli (Rawson & Ziegler, 1950)  TL: USA (NJ)  M T S1 S2

Callophrys hesseli angulata (Gatrelle, 2001)  TL: USA (SC)  M T S L H

Callophrys xami (Reakirt, [1867])  Xami Hairstreak    iN

Callophrys xami xami (Reakirt, [1867])  S AZ, W & C Mexico to Oaxaca  M T S L1 L2 I H

Callophrys xami texami Clench, 1981  S TX, far NE Mexico  M T S L I H

Callophrys scaphia Clench, 1981  Chiapas Hairstreak  Chiapas and Guatemala  iN  M T S L

Callophrys mcfarlandi P. Ehrlich & Clench, 1960  Sandia Hairstreak    iN  M T S L1 L2 I F Ha

Callophrys dospassosi Clench, 1981  Dos Passos' Hairstreak    iN

Callophrys dospassosi dospassosi Clench, 1981  E Mexico  M T S L

Callophrys dospassosi searsi Clench, 1981  W Mexico  M S L I

Callophrys estela Clench, 1981  Estela Hairstreak  TL: Mexico (Dur)  M S

Callophrys spinetorum (Hewitson, 1867)  Thicket Hairstreak    iN

Callophrys spinetorum spinetorum (Hewitson, 1867)  TL: USA (Ca)  M T S L1 L2 Ha

Callophrys spinetorum millerorum Clench, 1981  TL: Mexico (Hid)  M T S L

Callophrys johnsoni (Skinner, 1904)  Johnson's Hairstreak    iN  M S L I H Ha

Callophrys guatemalena Clench, 1981  Guatemalan Hairstreak    iN  M T S L

Callophrys augustinus (Westwood, 1852)  Brown Elfin    iN

Callophrys augustinus croesioides (Scudder, 1876)  TL: Canada  M T S L1 L2 LAb Ha

Callophrys augustinus helenae (dos Passos, 1943)  TL: Canada  M T S

Callophrys augustinus augustinus (Westwood, 1852)  TL: Canada  M T S L

Callophrys augustinus concava (Austin, 1998)  TL: USA (Nv)  M T S L Ha

Callophrys augustinus iroides (Boisduval, 1852)  Baja California Norte  M T S L1 L2 I Ha

Callophrys augustinus annetteae (dos Passos, 1943)  far NW Mexico  M T S L I

Callophrys fotis (Strecker, [1878])  Desert Elfin    iN

Callophrys fotis fotis (Strecker, [1878])  TL: USA (Az)  M S L I H

Callophrys fotis mojavensis (Austin, 1998)  TL: USA (Nv)  M T S L Ha

Callophrys mossii (H. Edwards, 1881)  Moss' Elfin    iN

Callophrys mossii schryveri (Cross, 1937)  TL: USA (Co)  M S L H

Callophrys mossii NE Cascadian segregate ,  NE Cascades (N OR, WA), E slope  M S Ha

Callophrys mossii mossii (H. Edwards, 1881)  TL: not specified  M S L Ha

Callophrys mossii W Oregon segregate ,  [nr. mossii]  M S L H

Callophrys mossii marinensis (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S I

Callophrys mossii bayensis R. Brown, 1969  TL: USA (Ca)  M T S

Callophrys mossii doudoroffi (dos Passos, 1940)  TL: USA (Ca)  M S L F Ha

Callophrys mossii hidakupa (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Callophrys mossii windi (Clench, 1943)  TL: USA (Ca)  M T S L I H

Callophrys mossii Siskiyou segregate ,  Siskiyou Mts., SW OR, Mts.of NW CA  M S

Callophrys mossii Mt. Diablo segregate ,  Mt. Diablo & vic., Contra Costa Co., CA  M S

Callophrys polios (Cook & F. Watson, 1907)  Hoary Elfin    iN

Callophrys polios polios (Cook & F. Watson, 1907)  TL: USA (NJ)  M T S1 S2 L Ha

Callophrys polios obscurus (Ferris & M. Fisher, 1973)  TL: USA (Co)  M T S L H

Callophrys polios NW montane segregate ,   M S

Callophrys polios Puget Trough segregate ,  Puget Trough, NW Washington  M S L

Callophrys polios C Oregon coastal segregate ,  C Oregon coast (Lincoln Co.)  M S

Callophrys polios maritima (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S L

Callophrys irus (Godart, [1824])  Frosted Elfin    iN

Callophrys irus irus (Godart, [1824])  TL: USA (NJ)  M T S1 S2 L Ha

Callophrys irus arsace (Boisduval & Le Conte, [1835])  TL: not specified  M T S1 S2 L1 L2 I H

Callophrys irus hadros (Cook & F. Watson, 1909)  TL: USA (Tx)  M T S1 S2 L1 L2 I1 I2 I3 H

Callophrys henrici (Grote & Robinson, 1867)  Henry's Elfin    iN

Callophrys henrici henrici (Grote & Robinson, 1867)  TL: USA (Pa)  M T S1 S2 L Ha

Callophrys henrici viridissima (Pavulaan, 1998)  TL: USA (NC)  M T S L

Callophrys henrici yahwehus Gatrelle, 1999  TL: USA (SC)  M T1 T2 S L

Callophrys henrici margaretae (dos Passos, 1943)  TL: USA (Fl)  M T S1 S2 L

Callophrys henrici turneri (Clench, 1943)  TL: USA (Ks)  M T S L1 L2 F

Callophrys henrici solatus (Cook & F. Watson, 1909)  TL: USA (Tx)  M T S L I

Callophrys lanoraieensis (Sheppard, 1934)  Bog Elfin    iN  M S L

Callophrys niphon (Hübner, [1819])  Eastern Pine Elfin    iN

Callophrys niphon niphon (Hübner, [1819])  TL: USA (Fl)  M S L F

Callophrys niphon clarki (T. Freeman, 1938)  TL: not specified  M T S1 S2 Ab L Ha

Callophrys eryphon (Boisduval, 1852)  Western Pine Elfin    iN

Callophrys eryphon northeastern segregate ,   M S

Callophrys eryphon purpurascens (Austin & J. Emmel, 1998)  TL: USA (Ca)  M T S L I

Callophrys eryphon sheltonensis (F. Chermock & Frechin, [1949])  TL: USA (Wa)  M T

Callophrys eryphon eryphon (Boisduval, 1852)  TL: USA (Ca)  M T S1 S2 L I1 I2 Ha

Callophrys eryphon pallescens (Austin, 1998)  TL: USA (Nv)  M T S L

Callophrys eryphon fusca (Austin, 1998)  TL: USA (Az)  M T S L1 L2 L3

Bistonina  Robbins, 2004  

Bistonina erema (Hewitson, 1867)  Erema Hairstreak  Guatemala, Panama  M T V

Bistonina biston (Möschler, 1877)    iN  M S

Bistonina bactriana (Hewitson, 1868)    iN  M T

Bistonina [n. sp.#1] Robbins, MS  TL: Costa Rica

Bistonina feretria (Hewitson, 1878)  TL: ?  M T S

Bistonina [n. sp.#2] Robbins, MS  TL: Costa Rica

Bistonina [n. sp.#3] Robbins, MS  TL: Peru

Bistonina mantica (H. Druce, 1907)    iN  M T L

Bistonina olbia (Hewitson, 1867)  TL: Brazil  M T

Megathecla  Robbins, 2002  

Megathecla corentini Faynel, 2009  TL: French Guiana

Megathecla gigantea (Hewitson, 1867)  TL: Brazil (Pa)  M T

Cupathecla  Bálint, 2006  

Cupathecla cupentus (Stoll, 1781)  Cupentus Hairstreak  SE Mexico to Guianas, N Brazil  iN  M T S L

Thestius  Hübner, [1819]  

Thestius pholeus (Cramer, 1777)  Dotted Thestius  Venezuela to N Brazil  iN  M T OI OD S IL Txt

Thestius selina (Hewitson, 1869)  Brazilian Thestius  Brazil  iN  M T OI S Txt

Thestius meridionalis (Draudt, 1920)  Common Thestius  Colombia to Venezuela and Bolivia   iN  M T OI OD S IL

Thestius epopea (Hewitson, 1870)  Epopea Hairstreak  Nicaragua to Peru  iN  M T S

Thestius [n. sp.#1] Robbins, MS  TL: Peru

Thestius [n. sp.#2] Robbins, MS  TL: Ecuador

Thestius azaria (Hewitson, 1867)  Azaria Hairstreak  S Brazil  iN  M T

Thestius [n. sp.#3] Robbins, MS  TL: Ecuador

Thestius lycabas (Cramer, 1777)  Lycabas Hairstreak  Peru to Brazil  iN  M T

Thestius [n. sp.#4] Robbins, MS  TL: Panama

Allosmaitia  Clench, [1964]  

Allosmaitia coelebs (Herrich-Schäffer, 1862)  Cuban Hairstreak  Cuba  M T S

Allosmaitia fidena (Hewitson, 1867)  Fidena Hairstreak  Hispaniola and Puerto Rico  iN  M T S

Allosmaitia piplea (Godman & Salvin, 1896)  Piplea Hairstreak  S Lesser Antilles (St. Vincent, Grenada, St. Lucia, Dominica)  M T S

Allosmaitia strophius (Godart, [1824])  Strophius Hairstreak  S TX, E & W Mexico to S Brazil  iN  M T S L

Allosmaitia myrtusa (Hewitson, 1867)  Myrtusa Hairstreak  Panama to Bolivia  iN  M T S

Laothus  K. Johnson, Kruse & Kroenlein, 1997  

Laothus gibberosa (Hewitson, 1867)    iN  M T S

Laothus numen (H. Druce, 1907)  TL: Guyana  M T

Laothus phydela (Hewitson, 1867)    iN  M T

Laothus erybathis (Hewitson, 1867)  Erybathis Hairstreak  E & W Mexico to Colombia  iN  M T S L I

Laothus oceia (Godman & Salvin, 1887)  Oceia Hairstreak  [= laothoe], SE Mexico to Colombia  iN  M T S L

Laothus viridicans (C. Felder & R. Felder, 1865)    iN  M T

Laothus barajo (Reakirt, [1867])  Barajo Hairstreak  SE Mexico to Panama  iN  M T S L1 L2 I

Laothus cockaynei (Goodson, 1945)  TL: Venezuela  M T S

Janthecla  Robbins & Venables, 1991  

Janthecla rocena (Hewitson, 1867)  Rocena Hairstreak  MX to S Brazil  iN  M T S

Janthecla adrienne Robbins, 2009  TL: Peru  M T

Janthecla malvina (Hewitson, 1867)    iN  M T

Janthecla janthodonia (Dyar, 1918)  Janthodonia Hairstreak  S MX, Guatemala, Belize  iN  M T S L

Janthecla cydonia (H. Druce, 1890)  Cydonia Hairstreak  Costa Rica to Colombia  iN  M T S L

Janthecla leea Venables & Robbins, 1991    iN  M T S

Janthecla armilla (H. Druce, 1907)  TL: Brazil (RJ)  M T

Janthecla halli R. Busby & Robbins, 2009  TL: Ecuador

Janthecla janthina (Hewitson, 1867)  Janthina Hairstreak  MX to Venezuela  iN  M T S L

Janthecla sista (Hewitson, 1867)    iN  M T S

Janthecla aurora (H. Druce, 1907)    iN  M T S

Janthecla flosculus (H. Druce, 1907)    iN  M T

Lamprospilus  Geyer, 1832  

Lamprospilus ayawi Bálint, Fratello, Costa & Benmesbah, 2019  ref.192

Lamprospilus genius Geyer, 1832    iN  M T

Lamprospilus japola (E. Jones, 1912)    iN  M T

Lamprospilus [n. sp.#1] Robbins, MS  TL: Guyana

Lamprospilus decorata Lathy, 1926    iN

Lamprospilus decorata decorata Lathy, 1926    iN  M T

Lamprospilus decorata valluna Prieto & Faynel, 2023  TL: Colombia

Lamprospilus nicetus (C. Felder & R. Felder, 1865)    iN  M T

Lamprospilus [n. sp.#2] Robbins, MS  TL: Panama

Lamprospilus draudti Lathy, 1932    iN  M T

Lamprospilus occidentalis K. Johnson & Salazar, 2004    iN

Lamprospilus collucia (Hewitson, 1877)  Two-toned Groundstreak  Mexico to Ecuador  iN  M T S L

Lamprospilus orcidia (Hewitson, 1874)    iN  M T L

Lamprospilus [n. sp.#3] Robbins, MS  TL: Ecuador

Lamprospilus aunus (Cramer, 1775)    iN  M T

Lamprospilus [n. sp.#4] Robbins, MS  TL: Ecuador

Lamprospilus coelicolor (A. Butler & H. Druce, 1872)  Central American Groundstreak  Mexico to Bolivia  iN  M T S

Lamprospilus bicolor Faynel & Prieto, 2023  TL: Colombia

Lamprospilus bicolor bicolor Faynel & Prieto, 2023  TL: Colombia

Lamprospilus bicolor mirador Faynel & Prieto, 2023  TL: Peru

Lamprospilus prieta (Bálint, 2022)  TL: Colombia

Lamprospilus stegmaier Prieto & Faynel, 2023  TL: Colombia

Badecla  Robbins & Duarte, 2010  

Badecla badaca (Hewitson, 1868)    iN  M T1 T2

Badecla picentia (Hewitson, 1868)    iN  M T

Badecla quadramacula (Austin & K. Johnson, 1997)    iN  M T

Badecla lanckena (Schaus, 1902)    iN  M T

Badecla argentinensis (K. Johnson & Kroenlein, 1993)    iN  M T

Badecla clarissa (Draudt, 1920)    iN  M T

Arzecla  Robbins & Duarte, 2010  

Arzecla arza (Hewitson, 1874)  Single-banded Groundstreak  Mexico to S Brazil  iN  M T S L

Arzecla [n. sp.#1] Robbins, MS  TL: Ecuador

Arzecla tarpa (Godman & Salvin, 1887)  Double-banded Groundstreak  Mexico to Panama  iN  M T S

Arzecla canacha (Hewitson, 1877)    iN  M T

Arzecla [n. sp.#2] Robbins, MS  TL: Ecuador

Arzecla calatia (Hewitson, 1873)  Nicaraguan Groundstreak  Mexico, Guatemala, Nicaragua  iN  M T OI

Arzecla tucumanensis (K. Johnson & Kroenlein, 1993)    iN  M T

Arzecla sethon (Godman & Salvin, 1887)  Large Groundstreak  Mexico to Panama  iN  M T S Ab L

Arzecla nubilum (H. Druce, 1907)    iN  M T

Arzecla paralus (Godman & Salvin, 1887)  Paralus Groundstreak  Guatemala to Venezuela  iN  M T OI

Arzecla taminella (Schaus, 1902)  Taminella Groundstreak  Mexico to S Brazil  iN  M T S L

Arzecla albolineata (Lathy, 1936)    iN  M T

Arzecla straelena Bálint, 2019  Colombia  iN

Arzecla straboris Bálint, 2019  TL: Colombia

Arumecla  Robbins & Duarte, 2004  

Arumecla aruma (Hewitson, 1877)    iN  M T

Arumecla galliena (Hewitson, 1877)  Red-based Groundstreak  Mexico to Ecuador  iN  M T S L

Arumecla [n. sp.#1] Robbins, MS  TL: Peru

Arumecla [n. sp.#2] Robbins, MS  TL: Ecuador

Arumecla nisaee (Godman & Salvin, 1887)  Nisaee Groundstreak  Costa Rica, Panama  iN  M T OI

Arumecla [n. sp.#3] Robbins, MS  TL: Peru

Arumecla [n. sp.#4] Robbins, MS  TL: Ecuador

Arumecla [n. sp.#5] Robbins, MS  TL: Ecuador

Arumecla [n. sp.#6] Robbins, MS  TL: Panama

Arumecla [n. sp.#7] Robbins, MS  TL: Peru

Arumecla [n. sp.#8] Robbins, MS  TL: Ecuador

Arumecla [n. sp.#9] Robbins, MS  TL: Ecuador

Arumecla [n. sp.#10] Robbins, MS  TL: Peru

Arumecla [n. sp.#11] Robbins, MS  TL: Peru

Arumecla [n. sp.#12] Robbins, MS  TL: Panama

Arumecla netesca (Draudt, 1920)  TL: Guyana; Peru  M T

Camissecla  Robbins & Duarte, 2004  

Camissecla charichlorus (A. Butler & H. Druce, 1872)  Charichlorus Groundstreak  Honduras to Colombia  iN  M T S

Camissecla [n. sp.#1] Robbins, MS  TL: Ecuador

Camissecla simasca (Draudt, 1920)  TL: Colombia  M T

Camissecla gedrosia (Hewitson, 1868)  TL: Brazil (Pa)  M T

Camissecla camissa (Hewitson, 1870)    iN  M T L

Camissecla cleocha (Hewitson, 1870)    iN  M T

Camissecla [n. sp.#2] Robbins, MS  TL: Ecuador

Camissecla [n. sp.#3] Robbins, MS  TL: Bolivia

Camissecla pactya (Hewitson, 1874)    iN  M T

Camissecla vespasianus (A. Butler & H. Druce, 1872)  Vespasianus Groundstreak  Mexico to Costa Rica  iN  M T S

Camissecla melma Schaus, 1913  Melma Groundstreak  Costa Rica and Panama  iN  M T S

Camissecla saphronotis (K. Johnson & Kroenlein, 1993)    iN  M T

Camissecla vesper (H. Druce, 1909)    iN  M T S L

Camissecla verbenaca (H. Druce, 1907)    iN  M T

Camissecla [n. sp.#4] Robbins, MS  TL: Brazil (SC)

Electrostrymon  Clench, 1961  

Electrostrymon endymion (Fabricius, 1775)    iN  M T

Electrostrymon constantinoi (K. Johnson & Kroenlein, 1993)  TL: Colombia  M T

Electrostrymon hugon (Godart, [1824])  Ruddy Hairstreak  [= sangala, = cyphara] S TX to Venezuela  iN  M T S L I

Electrostrymon joya (Dognin, 1895)  Muted Hairstreak  [= canus] S TX to Peru  iN  M T S L

Electrostrymon dominicana (Lathy, 1904)  Dominican Hairstreak  Dominica  iN

Electrostrymon dominicana dominicana (Lathy, 1904)  Dominica  M T S

Electrostrymon dominicana karukera Brévignon, 2000  TL: Guadeloupe

Electrostrymon pan (Drury, 1773)  Pan Hairstreak  Jamaica  iN  M T S L

Electrostrymon picoloro Prieto & Robbins, 2015  ref.278

Electrostrymon angelia (Hewitson, 1874)  Fulvous Hairstreak    iN

Electrostrymon angelia angelia (Hewitson, 1874)  S Florida and Cuba  M T S L

Electrostrymon angelia dowi (Clench, 1941)  Bahamas & Turks and Caicos Islands  M T S L

Electrostrymon angelia pantoni (W. Comstock & Huntington, 1943)  Jamaica  M T S

Electrostrymon angelia boyeri (W. Comstock & Huntington, 1943)  Hispaniola to St. Croix  M T S

Electrostrymon angerona (Godman & Salvin, 1896)  Angerona Hairstreak  Lesser Antilles  iN  M T S

Pendantus  K. Johnson & Kroenlein, 1993  

Pendantus denarius (A. Butler & H. Druce, 1872)  Coppery Groundstreak  E Mexico to Guatemala; ref.10  iN  M T S L

Pendantus guzanta (Schaus, 1902)  Orange-crescent Groundstreak  W and S TX to Nicaragua; ref.10  iN  M T S L1 L2 I Ha

Pendantus thurman (Thompson & Robbins, 2016)  Panama; ref.10  iN

Pendantus perisus (H. Druce, 1907)  ref.10  iN  M T

Rubroserrata  K. Johnson & Kroenlein, 1993  

Rubroserrata mathewi (Hewitson, 1874)  Mathew’s Groundstreak  Mexico to Panama  iN  M T S1 S2 L I

Rubroserrata ecbatana (Hewitson, 1868)    iN  M T S

Ziegleria  K. Johnson, 1993  

Ziegleria hesperitis (A. Butler & H. Druce, 1872)  Hesperitis Groundstreak  Mexico to S Brazil  iN  M T S L

Ziegleria hoffmani K. Johnson, 1993  Hoffmann’s Groundstreak  Mexico to Costa Rica  iN  M T S I

Ziegleria ceromia (Hewitson, 1877)  Ceromia Groundstreak  Mexico to Venezuela  iN  M T S

Ziegleria hernandezi (K. Johnson & Kroenlein, 1993)  TL: Trinidad  M T

Kisutam  K. Johnson & Kroenlein, 1993  

Kisutam micandriana (K. Johnson, 1992)  Micandriana Groundstreak  Mexico to Nicaragua; ref.294  M T S

Kisutam rosemary Thompson & Robbins, 2015  Rosemary's Groundstreak  Costa Rica and Panama; ref.294

Kisutam syllis (Godman & Salvin, 1887)  Sky-blue Groundstreak  S TX (rare stray), E & W Mexico to S Brazil; ref.294  iN  M T S L

Calycopis  Scudder, 1876  

Calycopis tamos (Godman & Salvin, 1887)  Tamos Hairstreak  [possibly a synonym of C. matho]  iN  M T S

Calycopis matho (Godman & Salvin, 1887)  TL: Guyana  M T

Calycopis clarina (Hewitson, 1874)  White-striped Groundstreak    iN  M T S L Ha

Calycopis cos (H. Druce, 1907)    iN  M T

Calycopis atnius (Herrich-Schäffer, [1853])  Atnius Groundstreak    iN  M T S L

Calycopis mimas (Godman & Salvin, 1887)  Mimas Groundstreak    iN  M T S

Calycopis [n. sp.#1] Robbins, MS  TL: Brazil (Ma)

Calycopis petaurister (H. Druce, 1907)    iN  M T

Calycopis calor (H. Druce, 1907)    iN  M T

Calycopis [n. sp.#2] Robbins, MS  TL: Panama

Calycopis johnsoni (Salazar, 2000)    iN  M T

Calycopis demonassa (Hewitson, 1868)  Shining Groundstreak    iN  M T S L

Calycopis buphonia (Hewitson, 1868)  Buphonia Groundstreak  Costa Rica to N Brazil  iN  M T S

Calycopis calus (Godart, [1824])  Calus Groundstreak    iN  M S L

Calycopis wolfii (K. Johnson, 1993)    iN  M T L

Calycopis caesaries (H. Druce, 1907)  Caesaries Groundstreak  Belize to N Brazil  iN  M T OI S L

Calycopis sullivani Robbins & Duarte, 2005  Sullivan's Hairstreak  Atlantic coast of Panama and Costa Rica  iN  M T S

Calycopis cicero Robbins & Duarte, 2005    iN

Calycopis ledaea (Hewitson, 1868)  TL: Brazil (Am)  M T

Calycopis cerata (Hewitson, 1877)  Cerata Groundstreak    iN  M T S L

Calycopis cecrops (Fabricius, 1793)  Red-banded Hairstreak  E USA, New Providence Island, Bahamas  iN  M T S Ma L1 L2 L3 L4

Calycopis isobeon (A. Butler & H. Druce, 1872)  Dusky-blue Groundstreak    iN  M T S1 S2 Ab L1 L2 I Txt

Calycopis quintana (K. Johnson, 1991)  Yucatan Groundstreak  Yucatan Peninsula of Mexico and N Belize  iN  M T S

Calycopis bactra (Hewitson, 1877)  Hewitson's Groundstreak    iN  M T S

Calycopis lerbela W. D. Field, 1967    iN  M T

Calycopis caulonia (Hewitson, 1877)    iN  M T

Calycopis [n. sp.#3] Robbins, MS  TL: Brazil (RJ)

Calycopis nicolayi W. D. Field, 1967    iN  M T

Calycopis [n. sp.#4] Robbins, MS  TL: Peru

Calycopis bellera (Hewitson, 1877)  Bellera Groundstreak  Panama, Surinam, Brazil  iN  M T OI S L

Calycopis xeneta (Hewitson, 1877)  Brilliant Groundstreak    iN  M T S L

Calycopis centoripa (Hewitson, 1868)    iN  M T S

Calycopis thama (Hewitson, 1877)  Thama Hairstreak  Costa Rica to Colombia  iN  M T S

Calycopis torqueor (H. Druce, 1907)    iN  M T

Calycopis vibulena (Hewitson, 1877)    iN  M T

Calycopis origo (Godman & Salvin, 1887)  Origo Groundstreak  [= susanna]  iN  M T S L

Calycopis drusilla W. D. Field, 1967  Drusilla Groundstreak    iN  M T S L

Calycopis vitruvia (Hewitson, 1877)    iN  M T

Calycopis partunda (Hewitson, 1877)    iN  M T

Calycopis indigo (H. Druce, 1907)    iN  M T

Calycopis anfracta (H. Druce, 1907)  TL: Peru  M T

Calycopis [n. sp.#5] Robbins, MS  TL: Ecuador

Calycopis anastasia W. D. Field, 1967  TL: Bolivia  M T

Calycopis fractunda W. D. Field, 1967  TL: Peru  M T

Calycopis anthora (Hewitson, 1877)    iN  M T

Calycopis spadectis (K. Johnson & Kroenlein, 1993)  TL: Ecuador  M S

Calycopis malta (Schaus, 1902)    iN  M T L

Calycopis gentilla (Schaus, 1902)    iN  M T

Calycopis trebula (Hewitson, 1868)  Trebula Groundstreak    iN  M T S L

Calycopis puppius (Godman & Salvin, 1887)    iN  M T

Calycopis [n. sp.#6] Robbins, MS  TL: Brazil (RJ)

Calycopis meleager (H. Druce, 1907)  TL: Surinam  M T S

Calycopis pisis (Godman & Salvin, 1887)  Pisis Groundstreak    iN  M T S

Calycopis [n. sp.#7] Robbins, MS  TL: Peru

Calycopis talama (Schaus, 1902)    iN  M T

Calycopis tifla (W. D. Field, 1967)  TL: Peru  M T

Calycopis [n. sp.#8] Robbins, MS  TL: Peru

Calycopis janeirica (C. Felder, 1862)    iN  M T

Calycopis vidulus (H. Druce, 1907)    iN  M T

Calycopis cyanus (Draudt, 1920)  TL: Bolivia  M T

Calycopis [n. sp.#9] Robbins, MS  TL: Peru

Calycopis orcillula (Strand, 1916)  Strand's Groundstreak    iN  M T S

Calycopis mirna Robbins & Duarte, 2002  TL: Argentina  M T

Calycopis orcilla (Hewitson, 1874)    iN  M T S

Calycopis blora (W. D. Field, 1967)  TL: French Guiana  M T

Calycopis naka (W. D. Field, 1967)  TL: Peru  M T

Calycopis barza (W. D. Field, 1967)  TL: Peru  M T

Calycopis gizela (Hewitson, 1877)    iN  M T

Calycopis suda (Draudt, 1920)    iN  M S

Calycopis boliviensis (K. Johnson, 1991)  TL: Bolivia  M T

Calycopis cinniana (Hewitson, 1877)    iN  M T

Calycopis plumans (H. Druce, 1907)  TL: Brazil (MT)  M T

Calycopis hosmeri (Weeks, 1906)  TL: Venezuela  M T

Calycopis [n. sp.#10] Robbins, MS  TL: Peru

Calycopis vesulus (Stoll, 1781)  TL: Surinam  M T

Calycopis cissusa (Hewitson, 1877)    iN  M T S

Strymon  Hübner, 1818  

Strymon melinus (Hübner, [1813])  Gray Hairstreak    iN

Strymon melinus melinus (Hübner, [1813])  TL: [USA (Ga)]  M T S L1 L2

Strymon melinus humuli (T. Harris, 1841)  TL: USA (Ma)  M T S L

Strymon melinus franki W. D. Field, 1938  TL: USA (Ks)  M T S Ab L1 L2 L3 L4 L5 I H

Strymon melinus setonia McDunnough, 1927  TL: Canada  M S L Ha

Strymon melinus atrofasciata McDunnough, 1921  TL: Canada  M S L Ha

Strymon melinus pudica (H. Edwards, 1877)  Baja California  M S1 S2 L1 L2 I Ha

Strymon melinus clarionensis (Heid, 1933)  [Apparently a species-level taxon], Clarion I., Colima, Mexico  M T S

Strymon melinus caldasensis Salazar, Vélez & K. Johnson, 1997  TL: Colombia  M T

Strymon avalona (W. G. Wright, 1905)  Avalon Scrub-Hairstreak  Santa Catalina Island, S CA  iN  M T S L Ha

Strymon daraba (Hewitson, 1867)    iN  M T

Strymon sabinus (C. Felder & R. Felder, 1865)    iN  M T

Strymon rufofusca (Hewitson, 1877)  Red-crescent Scrub-Hairstreak    iN

Strymon rufofusca rufofusca (Hewitson, 1877)  S TX, E & W Mexico to Paraguay, Lesser Antilles  M T S L1 L2

Strymon rufofusca bruneifimbria Brévignon, 2020  ref.178

Strymon auyana Bálint, Benmesbah & Viloria, 2018  ref.191

Strymon cyanofusca K. Johnson, Eisele & MacPherson, 1990  TL: Argentina  M T

Strymon lamasi Nicolay & Robbins, 2005  ref.270  iN  M T S

Strymon albata (C. Felder & R. Felder, 1865)  White Scrub-Hairsreak  [= sedecia] S TX to Venezuela  iN  M T S L I H

Strymon alea (Godman & Salvin, 1887)  Lacey’s Scrub-Hairstreak  [= laceyi] TX to Venezuela  iN  M T S L I H

Strymon bebrycia (Hewitson, 1868)  Red-lined Scrub-Hairstreak  SE AZ to S TX to Costa Rica  iN  M T S L H

Strymon yojoa (Reakirt, [1867])  Yojoa Scrub-Hairstreak  SE AZ, S TX to S Brazil  iN  M T S L1 L2 L3

Strymon tegaea (Hewitson, 1868)    iN  M T

Strymon ohausi (Spitz, 1933)    iN  M T

Strymon sylea (Hewitson, 1867)  TL: Brazil (Pa)  M T

Strymon mulucha (Hewitson, 1867)  Mottled Scrub-Hairstreak  Mexico to S Brazil  iN  M T S L

Strymon cestri (Reakirt, [1867])  Tailless Scrub-Hairstreak  SE AZ, S TX to S Brazil  iN  M T S L

Strymon davara (Hewitson, 1868)    iN  M T

Strymon crambusa (Hewitson, 1874)    iN  M T

Strymon astiocha (Prittwitz, 1865)  Gray-spotted Scrub-Hairstreak  Mexico to S Brazil  iN  M T S L

Strymon martialis (Herrich-Schäffer, 1865)  Martial Scrub-Hairstreak  S Florida, Bahamas, Cuba, Jamaica, Caymans  iN  M T S1 S2 L I H

Strymon christophei (W. Comstock & Huntington, 1943)  Hispaniolan Scrub-hairstreak  Hispaniola  M T S

Strymon michelle Nicolay & Robbins, 2005  ref.270  M T S

Strymon acis (Drury, 1773)  Caribbean Scrub-Hairstreak    iN

Strymon acis bartrami (W. Comstock & Huntington, 1943)  S Florida  M T S L Ha

Strymon acis casasi (W. Comstock & Huntington, 1943)  Cuba, Caymans  M T S

Strymon acis armouri Clench, 1943  Great Bahama Bank (Andros to Long), possibly Crooked I.  M T S

Strymon acis Great Inagua segregate ,  Great Inagua Island, Bahamas  M S

Strymon acis leucosticha Clench, 1992  Middle Caicos Island (at least)  M T

Strymon acis gossei (W. Comstock & Huntington, 1943)  Jamaica  M T S L

Strymon acis petioni (W. Comstock & Huntington, 1943)  Hispaniola  M T S

Strymon acis mars (Fabricius, 1776)  Puerto Rico (include. Mona & Culebran Is.) and Virgin Islands  M S L

Strymon acis acis (Drury, 1773)  Lesser Antilles (St. Martin to Dominica)  M T S

Strymon bazochii (Godart, [1824])  Lantana Scrub-Hairstreak    iN

Strymon bazochii bazochii (Godart, [1824])  S TX, E & W Mexico to Argentina  M T S L1 L2 I1 I2 H

Strymon bazochii gundlachianus M. Bates, 1935  Cuba, Jamaica, Hispaniola  M T S

Strymon [n. sp.] Nicolay & Robbins, MS  Venezuela

Strymon istapa (Reakirt, [1867])  Mallow Scrub-Hairstreak    iN

Strymon istapa clenchi Austin & J. Emmel, 1998  Baja California  M T S L

Strymon istapa istapa (Reakirt, [1867])  S TX, E & W Mexico to Argentina  M S Ab Ma L1 L2 L3 L4 I1 I2 H

Strymon istapa modesta (Maynard, 1873)  S Florida  M S L

Strymon istapa cybira (Hewitson, 1874)  Bahamas, Greater Antilles, (Dry Tortugas?)  M T S L

Strymon istapa arecibo (W. Comstock & Huntington, 1943)  Puerto Rico  M T S L

Strymon istapa clarionica (Vázquez, 1958)  Clarion Island; Relationship to S. i. clenchi requires study  M T

Strymon istapa socorroica (Vázquez, 1958)  [Apparently a species-level taxon], Socorro I., Colima, Mexico  M T

Strymon bubastus (Stoll, 1780)    iN

Strymon bubastus ponce (W. Comstock & Huntington, 1943)  Mona I., Puerto Rico, Virgin Is., Lesser Antilles S to Grenada  M T S L

Strymon bubastus bubastus (Stoll, 1780)  TL: Cape of Good Hope  M T

Strymon columella (Fabricius, 1793)  Columella Scrub-Hairstreak  Culebran Is., Virgin Is., St. Kitts to Antigua  iN  M T S

Strymon limenia (Hewitson, 1868)  Limenia Scrub-Hairstreak  S Florida, Greater Antilles, St. Thomas  iN  M T S

Strymon toussainti (W. Comstock & Huntington, 1943)  Toussaint's Scrub-Hairstreak  Hispaniola, E Cuba  iN  M T1 T2 T3 S

Strymon eurytulus (Hübner, [1819])    iN  M T

Strymon flavaria (Ureta, 1956)  TL: Chile  M T

Strymon colombiana (K. Johnson, L. Miller & Herrera, 1992)    iN  M T

Strymon bicolor (Philippi, 1859)    iN  M T

Strymon heodes (H. Druce, 1909)  TL: Peru  M T

Strymon wagenknechti (Ureta, 1947)    iN  M T

Strymon nivea (K. Johnson, L. Miller & Herrera, 1992)    iN  M T

Strymon rana (Schaus, 1902)    iN  M T

Strymon oribata (Weymer, 1890)    iN  M T

Strymon serapio (Godman & Salvin, 1887)  Bromeliad Scrub-Hairstreak  S TX (stray) to S Brazil  iN  M T1 T2 S L I

Strymon glorissima K. Johnson & Salazar, 1993    iN  M T

Strymon solitario Grishin & Durden, 2012  Big Bend Scrub-Hairstreak  TX Big Bend and vicinity & possibly NE MX; ref.61  iN  M T L I F Ha

Strymon jacqueline Nicolay & Robbins, 2005  ref.270  M T

Strymon gabatha (Hewitson, 1870)  Great Scrub-Hairstreak  Mexico to Peru  iN  M T OI S1 S2

Strymon monopeteinus A. Schwartz & J. Miller, 1985  Shining Scrub-Hairstreak  Hispaniola  M T S

Strymon oreala (Hewitson, 1868)    iN  M T

Strymon dindus (Fabricius, 1793)  TL: India

Strymon lucena (Hewitson, 1868)    iN  M T

Strymon cardus (Hewitson, 1874)    iN  M T

Strymon azuba (Hewitson, 1874)    iN  M T

Strymon eremica (Hayward, 1949)    iN  M T

Strymon ahrenholzi Nicolay & Robbins, 2005  ref.270  M S

Strymon veterator (H. Druce, 1907)  TL: Paraguay  M T

Strymon giffordi Nicolay & Robbins, 2005  ref.270  iN

Strymon megarus (Godart, [1824])  Megarus Scrub-Hairstreak  Mexico to S Brazil  iN  M T1 T2 OI S1 S2 L I G

Strymon ziba (Hewitson, 1868)  Ziba Scrub-Hairstreak  Mexico to S Brazil and Argentina  iN  M T S1 S2 L1 L2 G

Strymon anthracaetus Salazar, Vélez & Johnson, 1997  TL: Colombia

Strymon cryptodes Bálint, Cerdeña, Boyer & Pyrcz, 2019  TL: Peru

Tmolus  Hübner, [1819]  

Tmolus echion (Linnaeus, 1767)    iN

Tmolus echion echiolus (Draudt, 1920)  Hawaii, Mexico to S Brazil  M T S L

Tmolus echion echion (Linnaeus, 1767)  TL: West Indies  M T

Tmolus [n. sp.#1] Robbins, MS  TL: Panama

Tmolus crolinus A. Butler & H. Druce, 1872  Crolinus Hairstreak  Nicaragua to Venezuela  iN  M T S L

Tmolus venustus (H. Druce, 1907)    iN  M T S

Tmolus cydrara (Hewitson, 1868)  Cydrara Hairstreak  Mexico to S Brazil  iN  M T S L

Tmolus [n. sp.#2] Robbins, MS  TL: Peru

Tmolus [n. sp.#3] Robbins, MS  TL: Costa Rica

Tmolus ufentina (Hewitson, 1868)    iN  M T

Tmolus [n. sp.#4] Robbins, MS  TL: Ecuador

Tmolus [n. sp.#5] Robbins, MS  TL: Brazil (SC)

Tmolus [n. sp.#6] Robbins, MS  TL: Guatemala

Tmolus [n. sp.#7] Robbins, MS  TL: Peru

Tmolus [n. sp.#8] Robbins, MS  TL: Ecuador

Tmolus [n. sp.#9] Robbins, MS  TL: Peru

Tmolus mutina (Hewitson, 1867)  Mutina Midistreak  Costa Rica to Guianas & N Brazil  iN  M T S L

Nicolaea  K. Johnson, 1993  

Nicolaea cauter (H. Druce, 1907)    iN  M T

Nicolaea schausa (E. Jones, 1912)    iN  M T

Nicolaea dolium (H. Druce, 1907)  Dolium Hairstreak  Mexico to S Brazil  iN  M T S

Nicolaea ceglusa (Hewitson, 1868)  TL: Brazil  M T

Nicolaea bagrada (Hewitson, 1868)    iN  M T

Nicolaea xorema (Schaus, 1902)    iN  M T

Nicolaea [n. sp.#1] Robbins, MS  TL: Brazil (SP)

Nicolaea [n. sp.#2] Robbins, MS  TL: Brazil (Pr)

Nicolaea [n. sp.#3] Robbins, MS  TL: Ecuador

Nicolaea fabulla (Hewitson, 1868)    iN  M T

Nicolaea [n. sp.#4] Robbins, MS  TL: Mexico (Oax)

Nicolaea demilineata (Lathy, 1936)  TL: Paraguay  M T

Nicolaea [n. sp.#5] Robbins, MS  TL: Brazil (RJ)

Nicolaea [n. sp.#6] Robbins, MS  TL: Brazil (MT)

Nicolaea obelus (H. Druce, 1907)    iN  M T S

Nicolaea castinotus (K. Johnson & Le Crom, 1997)    iN  M T

Nicolaea [n. sp.#7] Robbins, MS  TL: Brazil (MT)

Nicolaea salki (Faynel & Robbins, 2014)  FG, Brazil (BA) & Peru; ref.204  iN

Nicolaea cupa (H. Druce, 1907)    iN  M T S

Nicolaea petilla (Hewitson, 1868)    iN  M T

Nicolaea velina (Hewitson, 1868)  Garnet-patched Hairstreak  [= phobe] Mexico to Brazil  iN  M T S L

Nicolaea torris (H. Druce, 1907)    iN  M T S

Nicolaea besidia (Hewitson, 1868)    iN  M T

Nicolaea socia (Hewitson, 1868)    iN  M T

Nicolaea [n. sp.#8] Robbins, MS  TL: Peru

Nicolaea viceta (Hewitson, 1868)  Nicaragua to S Brazil  M T

Nicolaea opalia (Hewitson, 1868)    iN  M T

Nicolaea opaliana (Hayward, 1967)  TL: Argentina

Nicolaea ophia (Hewitson, 1868)  Ophia Hairstreak  Nicaragua to S Brazil  iN  M T S I

Nicolaea [n. sp.#9] Robbins, MS  TL: Brazil (MT)

Nicolaea heraldica (Dyar, 1914)  Heraldica Hairstreak  Mexico to C Brazil  iN  M T S L

Nicolaea [n. sp.#10] Robbins, MS  TL: Panama

Nicolaea pyxis (K. Johnson, 1993)    iN  M T

Nicolaea umuarama (K. Johnson, 1993)    iN  M T

Nicolaea [n. sp.#11] Robbins, MS  TL: Ecuador

Nicolaea [n. sp.#12] Robbins, MS  TL: Ecuador

Nicolaea laconia (Hewitson, 1868)    iN  M T S

Nicolaea lemuria (Hewitson, 1868)    iN  M T

Nicolaea munditia (H. Druce, 1907)    iN  M T

Nicolaea [n. sp.#13] Robbins, MS  TL: Panama

Ministrymon  Clench, 1961  

Ministrymon leda (W. H. Edwards, 1882)  Leda Ministreak  SW USA to C Mexico  iN  M T S L1 L2 I Ha

Ministrymon clytie (W. H. Edwards, 1877)  Clytie Ministreak  S TX (?S AZ) to Costa Rica  iN  M T S L1 L2

Ministrymon albimimicus (K. Johnson, 1986)    iN  M T

Ministrymon megacles (Stoll, 1780)    iN  M T S

Ministrymon arola (Hewitson, 1868)  Arola Ministreak  [= coronta?] Mexico to Costa Rica  iN  M T S1 S2 L

Ministrymon coronta (Hewitson, 1874)  Central America  iN  M T

Ministrymon santans (Dyar, 1926)  Central America  iN  M T

Ministrymon [n. sp.#1] Robbins, MS  TL: Costa Rica

Ministrymon zilda (Hewitson, 1873)  Square-spotted Ministreak  Mexico to S Brazil  iN  M T S L

Ministrymon ollantaitamba (K. Johnson, L. Miller & Herrera, 1992)  TL: Peru  M T

Ministrymon sanguinalis (Burmeister, 1878)    iN  M T

Ministrymon gamma (H. Druce, 1909)    iN  M T

Ministrymon inoa (Godman & Salvin, 1887)  Veracruzan Ministreak  Mexico to Venezuela  iN  M T S

Ministrymon ligia (Hewitson, 1877)    iN  M T

Ministrymon phrutus (Geyer, 1832)  Purple-webbed Ministreak  Mexico to Brazil  iN  M T S L

Ministrymon cruenta (Gosse, 1880)    iN  M T L

Ministrymon arthuri Bálint, K. Johnson & Austin, [1999]    iN  M T S L

Ministrymon azia (Hewitson, 1873)  Gray Ministreak  [= guacanagari], SE AZ & S TX (strays) to Argentina, Antilles  iN  M T S L

Ministrymon janevicroy Glassberg, 2013  Pebbly Ministreak  Texas to Costa Rica & Curacao and Isla Margarita (Venezuela)  iN  M T S L G

Ministrymon cleon (Fabricius, 1775)  Cleon Ministreak  Costa Rica to S Brazil  iN  M T S

Ministrymon [n. sp.#2] Robbins, MS  TL: Guatemala

Ministrymon [n. sp.#3] Robbins, MS  TL: Mexico (Yuc)

Ministrymon fostera (Schaus, 1902)    iN  M T

Ministrymon una (Hewitson, 1873)    iN

Ministrymon una scopas (Godman & Salvin, 1887)  W & S Mexico at least to Costa Rica  M T S L

Ministrymon una una (Hewitson, 1873)  TL: Venezuela; Brazil (RJ)  M T S

Exorbaetta  K. Johnson, Austin, Le Crom & Salazar, 1997  

Exorbaetta metanira (Hewitson, 1867)    iN  M T S

Exorbaetta [n. sp.] Robbins, MS  Peru

Gargina  Robbins, 2004  

Gargina gargophia (Hewitson, 1877)  Gargophia Hairstreak  Mexico to Brazil  iN  M T S L

Gargina thyesta (Hewitson, 1869)  Thyesta Hairstreak  Costa Rica to Brazil  iN  M T S I

Gargina caninius (H. Druce, 1907)  Caninus Hairstreak  Mexico to S Brazil  iN  M T S

Gargina gnosia (Hewitson, 1868)  Gnosia Hairstreak  Mexico to Argentina  iN  M T S L

Gargina [n. sp.] Robbins, MS  Brazil (Pa)

Gargina emessa (Hewitson, 1867)  Emessa Hairstreak  Nicaragua to Brazil  iN  M T S

Gargina humber (Schaus, 1902)  TL: Venezuela  M T

Gargina thoria (Hewitson, 1869)  Thoria Hairstreak  Mexico to Venezuela  iN  M T S

Gargina panchaea (Hewitson, 1869)    iN  M T

Siderus  Kaye, 1904  

Siderus leucophaeus (Hübner, [1813])  White-spotted Hairstreak  Guatemala to Brazil  iN  M T S

Siderus [n. sp.#1] Robbins, MS  TL: Colombia

Siderus parvinotus Kaye, 1904  TL: Trinidad  M T

Siderus [n. sp.#2] Robbins, MS  TL: Peru

Siderus [n. sp.#3] Robbins, MS  TL: Ecuador

Siderus [n. sp.#4] Robbins, MS  TL: Ecuador

Siderus [n. sp.#5] Robbins, MS  TL: Honduras

Siderus guapila (Schaus, 1913)  Schaus' Hairstreak  Costa Rica  M T

Siderus [n. sp.#6] Robbins, MS  TL: Panama

Siderus [n. sp.#7] Robbins, MS  TL: Peru

Siderus [n. sp.#8] Robbins, MS  TL: Panama

Siderus eliatha (Hewitson, 1867)    iN  M T

Siderus [n. sp.#9] Robbins, MS  TL: Peru

Siderus [n. sp.#10] Robbins, MS  TL: Panama

Siderus bouvieri (Lathy, 1936)  TL: Ecuador  M T

Siderus athymbra (Hewitson, 1867)    iN  M T S

Siderus [n. sp.#11] Robbins, MS  TL: Colombia

Siderus [n. sp.#12] Robbins, MS  TL: Peru

Siderus [n. sp.#13] Robbins, MS  TL: Ecuador

Siderus [n. sp.#14] Robbins, MS  TL: Brazil

Siderus philinna (Hewitson, 1868)  Bold-spotted Hairstreak  MX to S Brazil  iN  M T1 T2 S I

Siderus giapor (Schaus, 1902)    iN  M T

Siderus nouraguensis Faynel & Robbins, 2014  ref.204  iN

Theclopsis  Godman & Salvin, 1887  

Theclopsis demea (Hewitson, 1874)  Demea Hairstreak  Nicaragua to Ecuador  iN  M T S

Theclopsis lydus (Hübner, [1819])  [= savaria]  iN  M T S L

Theclopsis murex (H. Druce, 1907)    iN  M T

Theclopsis leos (Schaus, 1913)  Leos Hairstreak  Honduras, Costa Rica  M T S I

Theclopsis epidius (Godman & Salvin, 1887)  Epidius Hairstreak  Panama to Colombia  iN  M T OI S

Theclopsis gargara (Hewitson, 1868)  Gargara Hairstreak  Colombia to S Brazil  iN  M T S L

Theclopsis mycon (Godman & Salvin, 1887)  Mycon Hairstreak  MX to Venezuela  iN  M T S1 S2 L I

Theclopsis [n. sp.#1] Robbins, MS  TL: Ecuador

Theclopsis [n. sp.#2] Robbins, MS  TL: Peru

Theclopsis aurina Robbins, 2002    iN  M T

Theclopsis sanmartina Bálint, 2022  TL: Peru

Grishinata  Robbins & Busby, 2022  

Grishinata penny Busby, J. Hall & Robbins, 2022  ref.204

Ostrinotes  K. Johnson, Austin, Le Crom & Salazar, 1997  

Ostrinotes purpuriticus (H. Druce, 1907)  Cryptic Hairstreak  SE Mexico to Colombia  iN  M T S

Ostrinotes silva (Faynel & Robbins, 2014)  Costa Rica to Bolivia & Brazil; ref.204  iN

Ostrinotes [n. sp.#1] Robbins, MS  TL: Brazil (Pa)

Ostrinotes empusa (Hewitson, 1867)  Empusa Hairstreak  Panama to Argentina  iN  M T S

Ostrinotes [n. sp.#2] Robbins, MS  TL: Peru

Ostrinotes halciones (A. Butler & H. Druce, 1872)  Halciones Hairstreak  MX to Brazil  iN  M T S L

Ostrinotes [n. sp.#3] Robbins, MS  TL: Panama

Ostrinotes [n. sp.#4] Robbins, MS  TL: Ecuador

Ostrinotes [n. sp.#5] Robbins, MS  TL: Peru

Ostrinotes tympania (Hewitson, 1869)    iN  M T

Ostrinotes tarena (Hewitson, 1874)    iN  M T

Ostrinotes gentiana (H. Druce, 1907)    iN  M T S

Ostrinotes sospes (Draudt, 1920)  TL: Colombia  M T

Ostrinotes [n. sp.#6] Robbins, MS  TL: Peru

Ostrinotes keila (Hewitson, 1869)  Keila Hairstreak  MX to Colombia  iN  M T S L

Ostrinotes sophocles (Fabricius, 1793)    iN

Ostrinotes [n. sp.#7] Robbins, MS  TL: Brazil (Go)

Strephonota  K. Johnson, Austin, Le Crom & Salazar, 1997  

Strephonota tephraeus (Geyer, 1837)  Pearly-gray Hairstreak  S Texas to Brazil  iN  M T S L

Strephonota sphinx (Fabricius, 1775)  Sphinx Hairstreak  Panama to Brazil  iN  M T S

Strephonota azurinus (A. Butler & H. Druce, 1872)  Azurinus Hairstreak  Costa Rica & Panama  iN  M T S

Strephonota [n. sp.#1] Robbins, MS  TL: Panama

Strephonota [n. sp.#2] Robbins, MS  TL: Ecuador

Strephonota [n. sp.#3] Robbins, MS  TL: Brazil (RJ)

Strephonota [n. sp.#4] Robbins, MS  TL: Brazil (SC)

Strephonota jactator (H. Druce, 1907)    iN  M T

Strephonota [n. sp.#5] Robbins, MS  TL: Peru

Strephonota [n. sp.#6] Robbins, MS  TL: Ecuador

Strephonota cyllarissus (Herbst, 1800)    iN  M T S

Strephonota bicolorata Faynel, 2003  TL: French Guiana  M T

Strephonota [n. sp.#7] Robbins, MS  TL: Peru

Strephonota pulchritudo (H. Druce, 1907)  TL: Amazons  M T

Strephonota [n. sp.#8] Robbins, MS  TL: Ecuador

Strephonota porphyritis (H. Druce, 1907)    iN  M T S

Strephonota [n. sp.#9] Robbins, MS  TL: Brazil (Rd)

Strephonota purpurantes (H. Druce, 1907)  TL: Peru  M T

Strephonota [n. sp.#10] Robbins, MS  TL: Peru

Strephonota foyi (Schaus, 1902)    iN  M T

Strephonota [n. sp.#11] Robbins, MS  TL: Ecuador

Strephonota [n. sp.#12] Robbins, MS  TL: Peru

Strephonota falsistrephon Faynel & Brévignon, 2003    iN  M T

Strephonota syedra (Hewitson, 1867)  Syedra Hairstreak  Mexico to Brazil  iN  M T S

Strephonota adela (Staudinger, 1888)    iN  M T S

Strephonota strephon (Fabricius, 1775)    iN  M T

Strephonota [n. sp.#13] Robbins, MS  TL: Peru

Strephonota parvipuncta (Lathy, 1926)    iN  M T

Strephonota [n. sp.#14] Robbins, MS  TL: Ecuador

Strephonota [n. sp.#15] Robbins, MS  TL: Peru

Strephonota tyriam (H. Druce, 1907)    iN  M T

Strephonota elika (Hewitson, 1867)    iN  M T

Strephonota [n. sp.#16] Robbins, MS  TL: Ecuador

Strephonota agrippa (Fabricius, 1793)    iN  M T

Strephonota trebonia (Hewitson, 1870)    iN  M T S

Strephonota [n. sp.#17] Robbins, MS  TL: Peru

Strephonota [n. sp.#18] Robbins, MS  TL: Peru

Strephonota perola (Hewitson, 1867)    iN  M T

Strephonota carteia (Hewitson, 1870)    iN  M T S

Strephonota ambrax (Westwood, 1852)  Ambrax Hairstreak  Mexico to Brazil  iN  M T S L

Strephonota malvania (Hewitson, 1867)  Malvania Hairstreak  Costa Rica & Panama  iN  M T S

Strephonota [n. sp.#19] Robbins, MS  TL: Peru

Strephonota acameda (Hewitson, 1867)    iN  M T

Strephonota ericeta (Hewitson, 1867)  Ericeta Hairstreak  Guatemala to Colombia  iN  M T S L

Strephonota buechei Faynel & Moser, 2011  TL: not specified

Strephonota caeruleus Faynel & Moser, 2011    iN

Strephonota berardi Faynel & Moser, 2011    iN  M S

Strephonota hightoni (Bálint, 2014)  TL: Venezuela

Panthiades  Hübner, [1819]  

Panthiades bitias (Cramer, 1777)  Bitias Hairstreak  Mexico to Brazil  iN  M T S1 S2 L1 L2 I1 I2

Panthiades hebraeus (Hewitson, 1867)    iN  M T

Panthiades aeolus (Fabricius, 1775)    iN  M T S

Panthiades boreas (C. Felder & R. Felder, 1865)    iN  M T

Panthiades ochus (Godman & Salvin, 1887)  Ochus Hairstreak  Mexico to Ecuador  iN  M T S L

Panthiades paphlagon (C. Felder & R. Felder, 1865)    iN  M T

Panthiades bathildis (C. Felder & R. Felder, 1865)  Zebra-striped Hairstreak  Mexico to N Brazil  iN  M T S L1 L2

Panthiades phaleros (Linnaeus, 1767)  Phaleros Hairstreak  SE Mexico to S Brazil  iN  M S HS L

Oenomaus  Hübner, [1819]  

Oenomaus ortygnus (Cramer, 1779)  Aquamarine Hairstreak  Mexico to Brazil  iN  M T S L

Oenomaus gwenish Robbins & Faynel, 2012  ref.203  M T G

Oenomaus mancha R. Busby & Faynel, 2012  ref.203  M T G

Oenomaus atesa (Hewitson, 1867)  Atesa Hairstreak  S Mexico to Brazil  iN  M T S

Oenomaus lea Faynel & Robbins, 2012  ref.203  iN  M T G

Oenomaus myrteana R. Busby, Robbins & Faynel, 2012  ref.203  iN  M T G

Oenomaus geba (Hewitson, 1877)  TL: ?  M T

Oenomaus brulei Faynel, 2008  TL: French Guiana  M T

Oenomaus griseus Faynel & Moser, 2008  TL: Brazil (DF)  M T

Oenomaus jauffreti Faynel & Moser, 2008    iN  M T

Oenomaus magnus Faynel & Moser, 2008  TL: Brazil (Am)  M T

Oenomaus poirieri Faynel, 2008  TL: French Guiana  M T

Oenomaus nigra Faynel & Moser, 2008  TL: Peru  M T

Oenomaus atena (Hewitson, 1867)  Atena Hairstreak  Costa Rica to Amazonas  iN  M T S L

Oenomaus morroensis Faynel & Moser, 2008    iN  M T

Oenomaus cortica (D'Abrera, 1995)    iN  M T

Oenomaus ambiguus Faynel, 2008  TL: French Guiana  M T

Oenomaus druceus Faynel & Moser, 2008    iN  M T

Oenomaus gaia Faynel, 2008  TL: French Guiana  M T

Oenomaus andi R. Busby & Faynel, 2012  ref.203  iN  M T G

Oenomaus moseri Robbins & Faynel, 2012  ref.203  iN  M T G

Oenomaus mentirosa Faynel & Robbins, 2012  ref.203  M T G

Oenomaus cyanovenata (D'Abrera, 1995)  Veined Hairstreak  Panama to Guianas  iN  M T S

Oenomaus floreus (H. Druce, 1907)    iN  M T

Oenomaus isabellae Faynel, 2006  TL: French Guiana  M T

Oenomaus taua Faynel & Moser, 2008  TL: Brazil (Pa)  M T

Oenomaus curiosa Faynel & Moser, 2008  TL: Peru  M T S

Oenomaus melleus (H. Druce, 1907)  Melleus Hairstreak  Nicaragua to Brazil  M T S I

Porthecla  Robbins, 2004  

Porthecla forasteira Faynel & Moser, 2011  ref.202  M T

Porthecla ravus (H. Druce, 1907)    iN  M T S

Porthecla annette Faynel & Robbins, 2011  Annette's Hairstreak  Panama to Amazonas; ref.202  M T S

Porthecla porthura (H. Druce, 1907)  Porthura Hairstreak  Honduras to W Ecuador  iN  M T S L

Porthecla minyia (Hewitson, 1867)    iN  M T

Porthecla gemma (H. Druce, 1907)    iN  M T

Porthecla johanna Faynel & Robbins, 2011  ref.202  M T

Porthecla peruensis Faynel & Moser, 2011  ref.202  M T

Porthecla prietoi Faynel & R. Busby, 2011  ref.202  M T

Porthecla willmotti R. Busby, Faynel & Moser, 2011  ref.202  M T

Porthecla barba (H. Druce, 1907)    iN  M T S

Porthecla dinus (Hewitson, 1867)  TL: Brazil  M T S

Thepytus  Robbins, 2004  

Thepytus epytus (Godman & Salvin, 1887)  Epytus Hairstreak  Panama to Ecuador  M T OI

Thepytus thyrea (Hewitson, 1867)    iN  M T

Thepytus arindela (Hewitson, 1874)  Arindela Hairstreak  S Mexico to Panama  M T S

Thepytus jennifer R. Busby & Robbins, 2010  TL: Ecuador

Thepytus beatrizae (Bálint & Dahners, 2006)  TL: Colombia  M T

Thepytus nancyana R. Busby & Robbins, 2010  ref.289

Thepytus carmen Robbins & Duarte, 2010  ref.289

Thepytus echelta (Hewitson, 1867)  Velvet Hairstreak  S Mexico to Brazil  iN  M T OI S

Parrhasius  Hübner, [1819]  

Parrhasius polibetes (Stoll, 1781)  Black-spot Hairstreak  E & W Mexico to Uruguay & Argentina  iN  M T S1 S2 L

Parrhasius orgia (Hewitson, 1867)  Variable Hairstreak  E & W Mexico to S Brazil & Argentina  iN  M T S1 S2 L

Parrhasius m-album (Boisduval & Le Conte, [1833])  White-M Hairstreak  S. Ontario CAN, E US from NH & WI to TX  iN  M T S L1 L2 I Ha

Parrhasius moctezuma (Clench, 1971)  Mexican-M Hairstreak  Mexico to Nicaragua  iN  M T S L

Parrhasius urraca Nicolay, 1979  Panamanian-M Hairstreak  Costa Rica & Panama  iN  M T S

Parrhasius selika (Hewitson, 1874)  Brazilian-M Hairstreak  Colombia to Brazil  iN  M T S

Michaelus  Nicolay, 1979  

Michaelus phoenissa (Hewitson, 1867)  Two-banded Hairstreak  SE Mexico to N Brazil & Peru  iN  M T S L I

Michaelus jebus (Godart, [1824])  Variegated Hairstreak  E & W Mexico to S Brazil & Argentina  iN  M T S L

Michaelus thordesa (Hewitson, 1867)  Thordesa Hairstreak  SE Mexico to Bolivia & SE Brazil  iN  M T S L

Michaelus hecate (Godman & Salvin, 1887)  Hecate Hairstreak  E & W Mexico to Costa Rica  iN  M T S

Michaelus ira (Hewitson, 1867)  Ira Hairstreak  E & W Mexico to SE Brazil & Argentina  iN  M T S1 S2 L

Michaelus joseph Robbins, 2010  Joseph Hairstreak  Costa Rica to W Amazon (Peru) & Brazil  iN  M T S

Ignata  K. Johnson, 1992  

Ignata mulsus (H. Druce, 1907)    iN  M T

Ignata elana (Hewitson, 1874)    iN  M T

Ignata gadira (Hewitson, 1867)  Gadira Hairstreak  Mexico to Panama  iN  M T S L

Ignata caldas Robbins, 2010  Caldas Hairstreak  Mexico to SE Peru in the Amazon Basin  iN  M T S L

Ignata brasiliensis (Talbot, 1928)    iN  M T

Ignata norax (Godman & Salvin, 1887)  Norax Hairstreak  S Mexico to Brazil  iN  M T S

Ignata levis (H. Druce, 1907)    iN  M T

Olynthus  Hübner, [1819]  

Olynthus narbal (Stoll, 1790)  Narbal Hairstreak  Panama to Peru, Guianas  iN  M T S L

Olynthus ophelia (Hewitson, 1867)    iN  M T

Olynthus avoca (Hewitson, 1867)  TL: Brazil  M T S

Olynthus [n. sp.#1] Robbins, MS  TL: Venezuela

Olynthus obsoleta (Lathy, 1926)  TL: Brazil  M T

Olynthus fulvoventris Austin & K. Johnson, 1998  TL: Brazil (Rd)  M T

Olynthus [n. sp.#2] Robbins, MS  TL: Brazil (MG)

Olynthus nitor (H. Druce, 1907)  TL: Brazil (Am)  M T

Olynthus negrus Austin & K. Johnson, 1998  TL: Brazil (Rd)  M T

Olynthus fancia (E. Jones, 1912)    iN  M T

Olynthus hypsea (Godman & Salvin, 1887)  Hypsea Hairstreak  Panama to Venezuela  M T OI

Olynthus [n. sp.#3] Robbins, MS  TL: Honduras

Olynthus [n. sp.#4] Robbins, MS  TL: Brazil (Pa)

Olynthus punctum (Herrich-Schäffer, [1853])  Punctum Hairstreak  [= stigmatos], Panama to Guianas & C Brazil  iN  M T S

Olynthus essus (Herrich-Schäffer, [1853])    iN  M T

Olynthus lorea (Möschler, 1883)  TL: Surinam  M T

Olynthus ostia (Hewitson, 1867)  TL: Brazil (RJ)  M T S

Olynthus [n. sp.#5] Robbins, MS  TL: Venezuela

Olynthus [n. sp.#6] Robbins, MS  TL: Ecuador

Olynthus porphyreticus (H. Druce, 1907)  TL: Peru  M T OI S

Hypostrymon  Clench, 1961  

Hypostrymon critola (Hewitson, 1874)  Sonoran Hairstreak  S AZ, BCS, W Mexico to GRO  iN  M T S L I

Hypostrymon [n. sp.#1] Robbins, MS  TL: Costa Rica

Hypostrymon renidens (Draudt, 1920)    iN  M T

Hypostrymon [n. sp.#2] Robbins, MS  TL: Venezuela

Hypostrymon aepeona (Draudt, 1920)  TL: Peru  M T

Hypostrymon [n. sp.#3] Robbins, MS  TL: Ecuador

Hypostrymon asa (Hewitson, 1868)  Asa Hairstreak  Guatemala to Brazil  iN  M T S L

Hypostrymon [n. sp.#4] Robbins, MS  TL: Peru

Hypostrymon [n. sp.#5] Robbins, MS  TL: Brazil (ES)

Marachina  Robbins, 2004  

Marachina maraches (H. Druce, 1912)    iN  M T S

Marachina [n. sp.#1] Robbins, MS  TL: Ecuador

Marachina [n. sp.#2] Robbins, MS  TL: Peru

Marachina fidelia (Hewitson, 1874)  TL: Venezuela  M T

Marachina peonida (Draudt, 1919)  TL: Colombia  M T

Apuecla  Robbins, 2004  

Apuecla maeonis (Godman & Salvin, 1887)  Maeonis Hairstreak  Mexico to Panama  iN  M T S L

Apuecla picus (H. Druce, 1907)  TL: Guyana  M T

Apuecla upupa (H. Druce, 1907)  Upupa Hairstreak  SE Mexico to Ecuador and Venezuela  iN  M T S

Nesiostrymon  Clench, [1964]  

Nesiostrymon calchinia (Hewitson, 1868)  Calachinia Hairstreak  Mexico to Argentina  iN  M T1 T2 S L

Nesiostrymon hyccara (Hewitson, 1868)  TL: Brazil  M T

Nesiostrymon celida (Lucas, 1857)  Caribbean Hairstreak    iN

Nesiostrymon celida celida (Lucas, 1857)  Cuba & Isla de Juventud  M T S

Nesiostrymon celida baorucoensis K. Johnson, 1991  SW Dominican Republic  M T

Nesiostrymon celida aibonito (W. Comstock & Huntington, 1943)  N Hispaniola & Puerto Rico  M T S

Nesiostrymon shoumatoffi (W. Comstock & Huntington, 1943)  Shoumatoff's Hairstreak  Jamaica  M T S

Nesiostrymon celona (Hewitson, 1874)  Celona Hairstreak  Mexico to Brazil  iN  M T S

Nesiostrymon dodava (Hewitson, 1877)  Dodava Hairstreak  Mexico to Venezuela  M T S

Nesiostrymon endela (Hewitson, 1874)    iN  M T

Nesiostrymon tristis (Lathy, 1926)  TL: Brazil (MT)  M T

Balintus  D'Abrera, 2001  

Balintus tityrus (C. Felder & R. Felder, 1865)    iN  M T

Aubergina  K. Johnson, 1991  

Aubergina vanessoides (Prittwitz, 1865)  Vanessoides Hairstreak  Colombia to Brazil  iN  M T S L

Aubergina paetus (Godman & Salvin, 1887)  Paetus Hairstreak  Mexico to Colombia  iN  M T S L

Aubergina hicetas (Godman & Salvin, 1887)  Hicetas Hairstreak  Mexico to Colombia  iN  M T S

Aubergina alda (Hewitson, 1868)    iN  M T

Aubergina [n. sp.#1] Robbins, MS  TL: Panama

Aubergina [n. sp.#2] Robbins, MS  TL: Peru

Aubergina hesychia (Godman & Salvin, 1887)  Hesychia Hairstreak  Costa Rica to Brazil  iN  M T OI

Terenthina  Robbins, 2004  

Terenthina terentia (Hewitson, 1868)  Terentia Hairstreak  Costa Rica to Brazil  iN  M T S L

Terenthina bradyae (D'Abrera, 1995)    iN  M T

Iaspis  Kaye, 1904  

Iaspis verania (Hewitson, 1868)    iN  M T

Iaspis ornata Austin & K. Johnson, 1996  TL: Brazil (Rd)  M T

Iaspis castitas (H. Druce, 1907)    iN  M T

Iaspis exiguus (H. Druce, 1907)    iN  M T

Iaspis talayra (Hewitson, 1868)    iN  M T

Iaspis andersoni Robbins, 2010  Anderson's Hairstreak  Mexico to N Colombia & Isla Coiba  iN  M T S1 S2 L I

Iaspis castimonia (H. Druce, 1907)  Castimonia Hairstreak  Mexico to Panama  iN  M T OI S L

Iaspis thabena (Hewitson, 1868)    iN  M T

Iaspis beera (Hewitson, 1870)    iN  M T

Iaspis grandis Austin & K. Johnson, 1996  TL: Brazil (Rd)  M T

Iaspis temesa (Hewitson, 1868)  Temesa Hairstreak  SE Mexico to S Brazil, Guianas  iN  M T S L

Celmia  K. Johnson, 1991  

Celmia celmus (Cramer, 1775)  Celmus Hairstreak  Mexico to Brazil and Paraguay  iN  M T S L

Celmia [n. sp.] Robbins, MS  Ecuador

Celmia uzza (Hewitson, 1873)    iN  M T

Celmia color (H. Druce, 1907)    iN  M T S

Celmia mecrida (Hewitson, 1867)  TL: Brazil  M T S L

Celmia anastomosis (Draudt, [1918])    iN  M T

Celmia conoveria (Schaus, 1902)  Conoveira Hairstreak  Mexico to Brazil  iN  M T S L

Dicya  K. Johnson, 1991  

Dicya dicaea (Hewitson, 1874)  Dicaea Hairstreak  Mexico to Brazil  iN  M T S

Dicya carnica (Hewitson, 1873)  Carnica Hairstreak  Mexico to Argentina  iN  M T S L

Dicya iambe (Godman & Salvin, 1887)  Iambe Hairstreak  Costa Rica to Peru  iN  M T OI S

Dicya eumorpha (Hayward, 1949)    iN  M T

Dicya lucagus (Godman & Salvin, 1887)  Lucagus Hairstreak  Mexico  iN  M T S

Dicya lollia (Godman & Salvin, 1887)  Lollia Hairstreak  Panama  iN  M T OI

Trichonis  Hewitson, 1865  

Trichonis hyacinthus (Cramer, 1775)    iN

Trichonis immaculata Lathy, 1930    iN  M T

Erora  Scudder, 1872  

Erora laeta (W. H. Edwards, 1862)  Early Hairstreak  SE CAN and NE US, from ME to N GA  iN  M S L

Erora quaderna (Hewitson, 1868)  Arizona Hairstreak  [= sanfordi], C & SE AZ, SW NM, W TX, E & W Mexico to Oaxaca  iN

Erora quaderna quaderna (Hewitson, 1868)  TL: Mexico  M T S L1 L2 L3 I

Erora quaderna caudata L. Miller, 1980  Chiapas  M T S

Erora subflorens (Schaus, 1913)  Emerald Hairstreak  Mexico to Venezuela  iN  M T S L

Erora biblia (Hewitson, 1868)    iN  M T S

Erora [n. sp.#1] Robbins, MS  TL: Brazil (Pa)

Erora nitetis (Godman & Salvin, 1887)  Nitetis Hairstreak  Mexico to Panama  iN  M T S L

Erora [n. sp.#2] Robbins, MS  TL: Panama

Erora [n. sp.#3] Robbins, MS  TL: Ecuador

Erora [n. sp.#4] Robbins, MS  TL: Peru

Erora aura (Godman & Salvin, 1887)  Aura Hairstreak  Mexico to W Colombia  iN  M T S

Erora tella (Schaus, 1902)    iN  M T

Erora phrosine (H. Druce, 1909)    iN  M T S

Erora carla (Schaus, 1902)  Carla Hairstreak  Mexico to Brazil  iN  M T S L

Erora gabina (Godman & Salvin, 1887)  Gabina Hairstreak  Mexico to Brazil  iN  M T S L

Erora opisena (H. Druce, 1912)  Opisena Hairstreak  Mexico to Venezuela  iN  M T S L

Erora [n. sp.#5] Robbins, MS  TL: Venezuela

Erora [n. sp.#6] Robbins, MS  TL: Brazil (SC)

Erora campa (E. Jones, 1912)    iN  M T

Erora [n. sp.#7] Robbins, MS  TL: Brazil (RJ)

Erora [n. sp.#8] Robbins, MS  TL: Peru

Erora [n. sp.#9] Robbins, MS  TL: Peru

Erora [n. sp.#10] Robbins, MS  TL: Peru

Erora badeta (Hewitson, 1873)  Badeta Hairstreak  Costa Rica to Brazil  iN  M T OI S I

Erora muridosca (Dyar, 1918)  Muridosca Hairstreak  Mexico, El Salvador  M T S L

Erora lampetia (Godman & Salvin, 1887)  Lampetia Hairstreak  Costa Rica to Brazil  M T S

Erora [n. sp.#11] Robbins, MS  TL: Ecuador

Erora [n. sp.#12] Robbins, MS  TL: Panama

Erora facuna (Hewitson, 1877)  TL: ?  M T

Erora senta (Draudt, 1920)  TL: Colombia  M T

Erora caespes (H. Druce, 1907)    iN  M T

Erora melba (Hewitson, 1877)    iN  M T

Erora nana (C. Felder & R. Felder, 1865)  TL: Colombia  M T

Erora lorina (Hewitson, 1874)  TL: ?  M T

Erora [n. sp.#13] Robbins, MS  TL: Brazil (RJ)

Erora aguaca (Draudt, 1920)  TL: Colombia

Semonina  Robbins, 2004  

Semonina ares (Godman & Salvin, 1887)  Ares Hairstreak  Panama to W Ecuador  M T OI

Semonina semones (Godman & Salvin, 1887)  Semones Hairstreak  Mexico to Brazil  iN  M T S

Semonina [n. sp.#1] Robbins, MS  TL: Panama

Semonina [n. sp.#2] Robbins, MS  TL: Brazil (MG)

Chalybs  Hübner, [1819]  

Chalybs janias (Cramer, 1779)  Janias Greenstreak  Mexico to Brazil  iN  M T S L

Chalybs chloris (Hewitson, 1877)    iN  M T S

Chalybs hassan (Stoll, 1790)  Hassan Greenstreak  Mexico to Brazil  iN  M T S L

Chalybs lineata (Lathy, 1936)    iN  M T

Chalybs mitaraka Faynel, 2019  TL: French Guiana

Symbiopsis  Nicolay, 1971  

Symbiopsis tanais (Godman & Salvin, 1887)  Tanais Hairstreak  Costa Rica to W Ecuador  iN  M T S

Symbiopsis [n. sp.#1] Robbins, MS  TL: Mexico (Col)

Symbiopsis pencilatus K. Johnson & Le Crom, 1997    iN  M T

Symbiopsis perulera Robbins, 2004    iN  M T

Symbiopsis [n. sp.#2] Robbins, MS  TL: Ecuador

Symbiopsis [n. sp.#3] Robbins, MS  TL: Colombia

Symbiopsis aprica (Möschler, 1883)    iN  M T S

Symbiopsis panamensis (Draudt, 1920)  Panama Hairstreak  Costa Rica, Panama  iN  M T S

Symbiopsis morpho Nicolay, 1971  TL: Ecuador  M T

Symbiopsis rickmani (Schaus, 1902)  Rickman's Hairstreak  Nicaragua to Colombia  iN  M T S

Symbiopsis pupilla (Draudt, 1920)    iN  M T

Symbiopsis pentas Nicolay, 1971    iN  M T

Symbiopsis [n. sp.#4] Robbins, MS  TL: Brazil (Pb)

Symbiopsis [n. sp.#5] Robbins, MS  TL: Brazil (ES)

Symbiopsis strenua (Hewitson, 1877)    iN  M T

Symbiopsis lenitas (H. Druce, 1907)    iN  M T

Symbiopsis nivepunctata (H. Druce, 1907)  TL: Guyana; Surinam  M T

  Subfamily Polyommatinae

  Tribe Polyommatini, Subtribe Celastrinina

Udara  Toxopeus, 1928  

Udara blackburni (Tuely, 1878)  Hawaiian Blue  Endemic to Hawaii  iN  M S L1 L2

Celastrina  Tutt, 1906  

Celastrina lucia (W. Kirby, 1837)  Lucia Azure    iN

Celastrina lucia lucia (W. Kirby, 1837)  TL: Canada (Sk)  M S1 S2 Ab L

Celastrina lucia lumarco Scott, 2006  TL: USA (Co)  M S L1 L2 L3 I Ha

Celastrina asheri LaBar, Pelham & Kondla, 2022  Asher's Azure    iN  M S1 S2 S3 L

Celastrina echo (W. H. Edwards, 1864)  Western Azure    iN

Celastrina echo echo (W. H. Edwards, 1864)  nigrescens Fletcher, 1903 is a homonym; ref.132  M T S1 S2 Ab L1 L2 I Ha

Celastrina echo sidara (Clench, 1944)  TL: USA (Co)  M S L I Ha

Celastrina echo cinerea (W. H. Edwards, 1883)  TL: USA (Az)  M T S L1 L2 I

Celastrina echo BCS segregate ,   M IL

Celastrina echo gozora (Boisduval, 1870)  TL: Honduras; Mexico  M T S L Ha

Celastrina ladon (Cramer, 1780)  Spring Azure    iN  M T L1 L2 F

Celastrina idella D. Wright & Pavulaan, 1999  Holly Azure    iN  M T S L

Celastrina serotina Pavulaan & D. Wright, 2005  Cherry Gall Azure    iN  M T S Ab L I H

Celastrina neglecta (W. H. Edwards, 1862)  Summer Azure    iN  M S1 S2 Ab L1 L2 L3 L4 L5 L6 I H

Celastrina humulus Scott & D. Wright, 1998  Hops Azure    iN  M S H

Celastrina neglectamajor Opler & Krizek, 1984  Appalachian Azure    iN  M T S L I

Celastrina nigra (W. Forbes, 1960)  Dusky Azure    iN  M S L Ha

  Tribe Polyommatini, Subtribe Zizeeriina

Zizina  Chapman, 1910  

Zizina otis (Fabricius, 1787)  Lesser Grass Blue    iN

Zizina otis otis (Fabricius, 1787)  Introduced to the Hawaiian Islands  M S L

  Tribe Polyommatini, Subtribe Lampidina

Lampides  Hübner, [1819]  

Lampides boeticus (Linnaeus, 1767)  Pea Blue  Introduced to Hawaii  iN  M T S L I

  Tribe Polyommatini, Subtribe Scolitantidina

Glaucopsyche  Scudder, 1872  

Glaucopsyche lygdamus (E. Doubleday, 1841)  Silvery Blue    iN

Glaucopsyche lygdamus lygdamus (E. Doubleday, 1841)  costal plains of eastern USA, only a couple of specimens are known  M S IL

Glaucopsyche lygdamus nittanyensis F. Chermock, 1945  differs in size, maculation and habitat preference  M T S1 S2 L1 L2 H Txt

Glaucopsyche lygdamus mildredae F. Chermock, 1945  TL: Canada (NS)  M T1 T2

Glaucopsyche lygdamus couperi Grote, 1873  TL: Canada (Qc)  M S L

Glaucopsyche lygdamus afra (W. H. Edwards, [1884])  TL: Canada (Mb)  M S

Glaucopsyche lygdamus jacki D. Stallings & Turner, 1947  TL: USA (Ks)  M T1 T2 S

Glaucopsyche lygdamus oro (Scudder, 1876)  TL: USA (Co)  M S L

Glaucopsyche lygdamus erico R. Holland, 2011  TL: USA (NM)  M OI

Glaucopsyche lygdamus ruidoso R. Holland, 2011  TL: USA (NM)  M OI

Glaucopsyche lygdamus arizonensis McDunnough, 1936  Hypothetical for Sonora, requires confirmation  M S L I

Glaucopsyche lygdamus minipunctum Austin, 1998  TL: USA (Nv)  M T S

Glaucopsyche lygdamus deserticola Austin & J. Emmel, 1998  TL: USA (Nv)  M T1 T2 S

Glaucopsyche lygdamus australis F. Grinnell, 1917  Baja California Norte  M T S L1 L2 I Ha

Glaucopsyche lygdamus maritima (Weeks, 1902)  Baja California Sur  M T OI

Glaucopsyche lygdamus palosverdesensis E. Perkins & J. Emmel, 1977  TL: USA (Ca)  M T1 T2 L

Glaucopsyche lygdamus sabulosa J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T1 T2 L Ha

Glaucopsyche lygdamus xerces (Boisduval, 1852)  TL: USA (Ca)  M T TI S1 S2 S3

Glaucopsyche lygdamus pseudoxerces J. Emmel & T. Emmel, 1998  TL: USA (Ca)  M T

Glaucopsyche lygdamus incognitus Tilden, 1974  TL: USA (Ca)  M T S L I Ha

Glaucopsyche lygdamus orcus (W. H. Edwards, 1869)  TL: [USA (Nv)]  M T S L Ha

Glaucopsyche lygdamus Sierra Nevada, high elevations segregate ,   M S L

Glaucopsyche lygdamus columbia (Skinner, 1917)  TL: USA (Wa)  M S L Ha

Glaucopsyche piasus (Boisduval, 1852)  Arrowhead Blue    iN

Glaucopsyche piasus umbrosa J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L I Ha

Glaucopsyche piasus sagittigera (C. Felder & R. Felder, 1865)  TL: [USA (Ca)]  M T S L I

Glaucopsyche piasus gabrielina J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S

Glaucopsyche piasus excubita J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L1 L2 I H

Glaucopsyche piasus piasus (Boisduval, 1852)  TL: USA (Ca)  M T TI S L

Glaucopsyche piasus toxeuma F. Brown, 1971  TL: Canada (BC)  M S L1 L2 Ha

Glaucopsyche piasus nevada F. Brown, 1975  TL: USA (Nv)  M S L Ha

Glaucopsyche piasus daunia (W. H. Edwards, 1871)  TL: USA (Co)  M S L

Philotes  Scudder, 1876  

Philotes sonorensis (C. Felder & R. Felder, 1865)  Sonoran Blue    iN

Philotes sonorensis sonorensis (C. Felder & R. Felder, 1865)  TL: [USA (Ca)]  M T S Ab L1 L2 L3 I H Ha Txt

Philotes sonorensis extinctis Mattoni, 1991  TL: USA (Ca)  M S1 S2

Philotes sonorensis Sierra Nevada segregate ,  Sierra Nevada Mts., CA  M S L I H

Euphilotes  Mattoni, [1978]  (includes Philotiella); ref.10

Euphilotes speciosa (H. Edwards, 1877)  Small Blue  ref.10  iN

Euphilotes speciosa speciosa (H. Edwards, 1877)  TL: USA (Ca)  M T S L1 L2 L3 I1 I2 H1 H2 Ha

Euphilotes speciosa septentrionalis (Austin, 1998)  TL: USA (Nv)  M T S I H

Euphilotes speciosa bohartorum (Tilden, [1969])  TL: USA (Ca)  M T S H

Euphilotes speciosa purisima (Priestaf & J. Emmel, 1998)  TL: USA (Ca)  M T S L1 L2 F Ha

Euphilotes leona (Hammond & McCorkle, 2000)  Leona's Blue  USA (Or); ref.10  iN  M T S Ab L1 L2 I Ha

Euphilotes battoides (Behr, 1867)  Square-spotted Blue    iN

Euphilotes battoides high-elevation, C & S Cascades segregate ,  [on Eriogonum marifolium], C & S Cascades, OR, far N CA  M S L1 L2 L3 H Ha

Euphilotes battoides battoides (Behr, 1867)  TL: USA (Ca)  M T S Ha

Euphilotes battoides argocyanea Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes battoides mazourka Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes battoides panamintensis Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes battoides fusimaculata Austin, 1998  TL: USA (Nv)  M T1 T2

Euphilotes battoides anasazi Scott, 1998  TL: USA (NM)  M T

Euphilotes allyni (Shields, 1975)  El Segundo Blue  El Segundo, Los Angeles County, CA; ref.130,131  iN  M T S1 S2 L

Euphilotes glaucon (W. H. Edwards, 1871)  Glaucon Blue    iN

Euphilotes glaucon glaucon (W. H. Edwards, 1871)  TL: USA (Nv)  M S L H

Euphilotes glaucon oregonensis (W. Barnes & McDunnough, 1917)  TL: USA (Or)  M T S L H

Euphilotes glaucon australoglaucon Pratt & J. Emmel, 1998  TL: USA (Ca)  M S L

Euphilotes glaucon comstocki (Shields, 1975)  TL: USA (Ca)  M T S

Euphilotes glaucon intermedia (W. Barnes & McDunnough, 1917)  TL: USA (Ca)  M T S L Ha

Euphilotes glaucon Siskiyou segregate ,  [nr. intermedia]  M S

Euphilotes glaucon W Cascadian segregate ,  W Cascades, Lane Co., OR  M S

Euphilotes glaucon N Cascades segregate ,  Mt. Hood, OR, Mt. Adams, WA, N into WA Cascades  M S L1 L2 Ha

Euphilotes centralis (W. Barnes & McDunnough, 1917)  Central Blue    iN

Euphilotes centralis centralis (W. Barnes & McDunnough, 1917)  TL: USA (Co)  M S L I H

Euphilotes centralis hadrochilus Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes bernardino (W. Barnes & McDunnough, 1916)  Bernardino Blue    iN

Euphilotes bernardino bernardino (W. Barnes & McDunnough, 1916)  TL: USA (Ca)  M T S L1 L2 L3 Ha

Euphilotes bernardino inyomontana Pratt & J. Emmel, 1998  TL: USA (Ca)  M S L

Euphilotes bernardino minuta Austin, 1998  TL: USA (Nv)  M T

Euphilotes bernardino martini (Mattoni, 1954)  Hypothetical for Sonora  M T S L F Ha

Euphilotes bernardino garthi Mattoni, 1989  TL: Mexico (BCN)  M T S L Ha

Euphilotes ellisii (Shields, 1975)  Ellis' Blue    iN

Euphilotes ellisii ellisii (Shields, 1975)  TL: USA (Co)  M T S L I

Euphilotes ellisii basinensis Austin, 1998  TL: USA (Nv)  M T

Euphilotes ellisii euromojavensis Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes ellisii avawatz Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes heracleoides Kohler & A. Warren, 2021  Heracleoides Blue    iN

Euphilotes baueri (Shields, 1975)  Bauer's Blue    iN

Euphilotes baueri borealis Kohler, 2021  TL: USA (Mt)

Euphilotes baueri shoshone Kohler, 2021  TL: USA (Id)

Euphilotes baueri vernalis Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes baueri baueri (Shields, 1975)  TL: USA (Ca)  M T S L H Ha

Euphilotes baueri orientis Austin, 1998  TL: USA (Nv)  M T S

Euphilotes baueri far N Great Basin, northward segregate ,   M S Ha

Euphilotes oakleyi Kohler, 2021  Oakley's Blue  TL: USA (Mt)

Euphilotes oakleyi oakleyi Kohler, 2021  TL: USA (Mt)

Euphilotes oakleyi madisonensis Kohler, 2021  TL: USA (Mt)

Euphilotes mojave (F. Watson & W. Comstock, 1920)  Mojave Blue    iN

Euphilotes mojave mojave (F. Watson & W. Comstock, 1920)  TL: USA (Ca)  M S L Ha

Euphilotes mojave virginensis Austin, 1998  TL: USA (Nv)  M T L I H

Euphilotes stanfordorum Opler & A. Warren, 2009  Stanfords' Blue  W Colorado, far NW New Mexico, possibly E Utah; on E. ovalifolium  iN  M T S I

Euphilotes enoptes (Boisduval, 1852)  Dotted Blue    iN

Euphilotes enoptes enoptes (Boisduval, 1852)  TL: USA (Ca)  M T TI S L I H Ha

Euphilotes enoptes aridorum Austin, 1998  TL: USA (Nv)  M T Ha

Euphilotes enoptes bayensis (Langston, 1964)  TL: USA (Ca)  M T S L

Euphilotes enoptes N OR Coast Range segregate ,  Polk Co., OR  M S

Euphilotes enoptes Siskiyou segregate ,  Siskiyou Mts., NW CA, SW OR  M S L H Ha

Euphilotes enoptes Cascades, low elevations segregate ,  W ORE Cascades below 4000', also SE WA Cascades  M S L H

Euphilotes enoptes OR Cascades, high elevations segregate ,  ORE Cascades above 4000'  M S L1 L2 Ha

Euphilotes enoptes tildeni (Langston, 1964)  TL: USA (Ca)  M T S

Euphilotes enoptes arenacola Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes enoptes smithi (Mattoni, 1954)  TL: USA (Ca)  M S L1 L2 H

Euphilotes enoptes dammersi (J. A. Comstock & Henne, 1933)  TL: USA (Ca)  M T S1 S2 L I

Euphilotes enoptes cryptorufes Pratt & J. Emmel, 1998  TL: USA (Ca)

Euphilotes enoptes opacapulla Austin, 1998  TL: USA (Nv)  M T

Euphilotes enoptes primavera Austin, 1998  TL: USA (Nv)  M T

Euphilotes enoptes langstoni (Shields, 1975)  TL: USA (Ca)  M T S L Ha

Euphilotes rita (W. Barnes & McDunnough, 1916)  Rita’s Blue    iN

Euphilotes rita rita (W. Barnes & McDunnough, 1916)  TL: USA (Az)  M T S L

Euphilotes rita four corners area segregate ,  NW New Mexico, S UT, N AZ  M S

Euphilotes rita coloradensis (Mattoni, [1966])  TL: USA (Co)  M T S L H

Euphilotes rita montanensis Kohler, 2021  TL: USA (Mt)

Euphilotes columbiae (Mattoni, 1954)  Columbian Blue    iN

Euphilotes columbiae columbiae (Mattoni, 1954)  TL: USA (Wa)  M S L1 L2 L3 I H

Euphilotes columbiae C Oregon segregate ,  Ochoco Mts, also W Jefferson Co., OR  M S L1 L2 I H Ha

Euphilotes ancilla (W. Barnes & McDunnough, 1918)  Ancilla Blue    iN

Euphilotes ancilla montosa Kohler, 2021  TL: USA (Mt)

Euphilotes ancilla campestris Kohler, 2021  TL: USA, MT

Euphilotes ancilla barnesi Opler & M. Fisher, 2009  TL: USA (Co)  M T S L1 L2 Ha

Euphilotes ancilla ancilla (W. Barnes & McDunnough, 1918)  TL: USA (Ut)  M T S L I H

Euphilotes ancilla SE OR segregate ,   M S L1 L2 Ha

Euphilotes ancilla giulianii Pratt & J. Emmel, 1998  TL: USA (Nv)

Euphilotes ancilla gilvatunica Austin, 1998  TL: USA (Nv)  M T

Euphilotes ancilla shieldsi Austin, 1998  TL: USA (Nv)  M T L

Euphilotes ancilla purpura Austin, 1998  N Spring Mts., Clark Co., NV, on E. umbellatum var. juniporinum  M T S

Euphilotes ancilla cryptica Austin & Boyd, 2008  TL: USA (Nv)

Euphilotes ancilla pseudointermedia Pratt & J. Emmel, 1998  TL: USA (Ca)  M S

Euphilotes pallescens (Tilden & Downey, 1955)  Pallid Blue    iN

Euphilotes pallescens emmeli (Shields, 1975)  TL: USA (Ut)  M T S L I H

Euphilotes pallescens pallescens (Tilden & Downey, 1955)  TL: USA (Ut)  M T S

Euphilotes pallescens mattonii (Shields, 1975)  TL: USA (Nv)  M T S

Euphilotes pallescens arenamontana Austin, 1998  TL: USA (Nv)  M T S Ha

Euphilotes pallescens ricei Austin, 1998  TL: USA (Nv)  M T

Euphilotes pallescens calneva J. Emmel & T. Emmel, 1998  TL: USA (Ca)  M S1 S2

Euphilotes pallescens confusa Pratt & J. Emmel, 1998  TL: USA (Ca)  M S L

Euphilotes pallescens elvirae (Mattoni, [1966])  TL: USA (Ca)  M T S

Euphilotes spaldingi (W. Barnes & McDunnough, 1917)  Spalding's Blue    iN

Euphilotes spaldingi spaldingi (W. Barnes & McDunnough, 1917)  TL: USA (Ut)  M S L1 L2 I H

Euphilotes spaldingi pinjuna Scott, 1981  TL: USA (Co)  M T S L H

  Tribe Polyommatini, Subtribe Brephidiina

Zizula  Chapman, 1910  

Zizula cyna (W. H. Edwards, 1881)  Cyna Blue    iN  M T S L I

Brephidium  Scudder, 1876  [includes Oraidium as a subgenus]; ref.154

Brephidium exilis (Boisduval, 1852)  Western Pygmy-Blue    iN

Brephidium exilis exilis (Boisduval, 1852)  TL: USA (Ca)  M T S L1 L2 L3 I H

Brephidium exilis yucateca Clench, 1970  Yucatan Peninsula  M T S

Brephidium exilis thompsoni Carpenter & H. Lewis, 1943  Cayman Islands  M T L

Brephidium exilis isophthalma (Herrich-Schäffer, 1862)  Bahamas, Cuba, Jamaica, Hispaniola  M T S L

Brephidium pseudofea (Morrison, 1873)  Eastern Pygmy-Blue    iN

Brephidium pseudofea insularus Pavulaan & Gatrelle, 1999  TL: USA (SC)  M T S

Brephidium pseudofea pseudofea (Morrison, 1873)  TL: USA (Fl)  M T S L1 L2 L3 Ha

  Tribe Polyommatini, Subtribe Leptotina

Leptotes  Scudder, 1876  

Leptotes cassius (Cramer, 1775)    iN

Leptotes cassius cassioides (Boisduval, 1870)  [= chadwicki], Windward and some Leeward Islands  M T S L

Leptotes cassius catilina (Fabricius, 1793)  Culebra, Virgin Is. to St. Martin  M T S

Leptotes cassius theonus (Lucas, 1857)  FL, Bahamas, Greater Antilles, Caymans  M T S L I

Leptotes cassius cassidula (Boisduval, 1870)  [= striata], S TX, E & W Mexico to Panama  M T S Ab L1 L2 H

Leptotes cassius surbaja Scott, 2017  Mexico: Baja California Sur  M S L

Leptotes cassius cassius (Cramer, 1775)  TL: Surinam  M T

Leptotes marina (Reakirt, 1868)  Marine Blue    iN  M T S L1 L2 L3 L4 I1 I2 H

Leptotes perkinsae Kaye, 1931  Perkins' Blue  Jamaica  iN  M T S

Leptotes hedgesi A. Schwartz & K. Johnson, 1992  Hedges' Blue  Cuba  M OI G

Leptotes idealus K. Johnson & Matusik, 1988  Ideal Blue  SW Hispaniola  M T S

Leptotes bathyllos Tessmann, 1928    iN  M T

Leptotes delalande Bálint & K. Johnson, 1995    iN  M T

Leptotes lamasi Bálint & K. Johnson, 1995    iN  M T

Leptotes trigemmatus (A. Butler, 1881)    iN  M T

Leptotes parrhasioides (Wallengren, 1860)    iN  M T

Leptotes andicola (Godman & Salvin, 1891)    iN  M T

Leptotes callanga (Dyar, 1913)    iN  M T S

Leptotes numerius (Stoll, 1790)  unclear if belongs here or from Suriname

  Tribe Polyommatini, Subtribe Everina

Elkalyce  Bálint & K. Johnson, [1996]  

Elkalyce cogina (Schaus, 1902)    iN  M T

Cupido  Schrank, 1801  

Cupido comyntas (Godart, [1824])  Eastern Tailed-Blue    iN

Cupido comyntas comyntas (Godart, [1824])  SE Canada, E USA to E CO, N & NE TX  M T S1 S2 Ab L1 L2 L3 L4 L5

Cupido comyntas texana (F. Chermock, 1945)  C & W TX, S NM, SE AZ, S to Ecuador  M T S L1 L2 I

Cupido comyntas sissona (W. G. Wright, 1905)  [= pacnowe], Baja to W WA + NE OR, E WA, N ID, SE BC  M T S L1 L2 Ha

Cupido amyntula (Boisduval, 1852)  Western Tailed-Blue    iN

Cupido amyntula montanorum (Austin, 1998)  TL: USA (Nv)  M S L I Ha

Cupido amyntula amyntula (Boisduval, 1852)  Baja California Norte  M T S L1 L2 I1 I2 Ha

Cupido amyntula nesiotes (J. Emmel & T. Emmel, 1998)  TL: USA (Ca)  M T1 T2

Cupido amyntula herrii (F. Grinnell, 1901)  TL: USA (Az)  M T S L1 L2

Cupido amyntula valeriae (Clench, 1944)  TL: USA (SD)  M T S Ha

Cupido amyntula albrighti (Clench, 1944)  TL: USA (Mo)  M T S1 S2

  Tribe Polyommatini, Subtribe Polyommatina

Pseudochrysops  Nabokov, 1945  

Pseudochrysops bornoi (W. Comstock & Huntington, 1943)  Antillean Blue    iN

Pseudochrysops bornoi bornoi (W. Comstock & Huntington, 1943)  Hispaniola  M T S

Pseudochrysops bornoi escobioi A. Schwartz, 1987  Puerto Rico  M T S

Pseudochrysops bornoi yateritas D. Smith & L. Hernández, 1992  SE Cuba  M T OI

Itylos  Draudt, 1921  [includes Paralycaeides as a subgenus]; ref.146

Itylos inconspicua Draudt, 1921  TL: Peru  M T

Itylos shade (Bálint, 1993)  TL: Peru  M T

Itylos vapa (Staudinger, 1894)    iN  M T

Itylos mashenka (Bálint, 1993)  TL: Peru  M T

Itylos fumosus (Balletto, 1993)  TL: Peru  M T

Itylos mira Bálint & Lamas, [1999]  TL: Peru  M T

Itylos pasco Bálint & Lamas, 1994    iN  M T

Itylos pnin Bálint, 1993    iN  M T

Itylos titicaca (Weymer, 1890)    iN  M T

Itylos ardisensis (Bálint & Lamas, [1997])  TL: Peru  M T

Itylos bella (Bálint & Lamas, [1997])  TL: Peru  M T

Itylos pacis Draudt, 1921    iN  M T

Itylos cobaltana (Bálint & Lamas, 1994)  TL: Peru  M T

Itylos nodo (Bálint & K. Johnson, 1995)    iN  M T

Itylos moza (Staudinger, 1894)    iN  M T

Itylos lolita (Bálint, 1993)    iN  M T L

Itylos pelorias (Weymer, 1890)    iN  M T

Itylos sigal (Benyamini, Bálint & K. Johnson, 1995)  TL: Chile  M T

Itylos colca (Bálint & Lamas, [1997])  TL: Peru  M T

Itylos gradoslamasi (Bálint, 1997)  TL: Peru  M T

Itylos huascarana (Bálint & Lamas, 1994)  TL: Peru  M T

Itylos koa (H. Druce, 1876)    iN  M T

Itylos tintarrona (Bálint & K. Johnson, 1995)  TL: Peru  M T

Itylos lea (Benyamini, Bálint & K. Johnson, 1995)  TL: Bolivia  M T

Itylos ludicra (Weymer, 1890)  TL: Chile  M T

Itylos malvasa (Bálint & Pyrcz, [2000])  TL: Colombia

Itylos vokoban (Bálint & C. Johnson, 1995)    iN  M T

Itylos [n. sp.] Lamas, MS  Peru

Pseudolucia  Nabokov, 1945  

Pseudolucia andina (Calvert, 1893)  TL: Chile  M T

Pseudolucia annamaria Bálint & K. Johnson, 1993  TL: Chile  M T

Pseudolucia arauco Bálint, Benyamini & K. Johnson, 2001    iN

Pseudolucia argentina (Balletto, 1993)    iN  M T

Pseudolucia asafi Benyamini, Bálint & K. Johnson, 1995  TL: Chile  M T

Pseudolucia aureliana Bálint & K. Johnson, 1993  TL: Chile  M T

Pseudolucia avishai Benyamini, Bálint & K. Johnson, 1995  TL: Chile  M T

Pseudolucia benyamini Bálint & K. Johnson, 1995    iN  M T

Pseudolucia charlotte Bálint & K. Johnson, 1993    iN  M T

Pseudolucia chilensis (Blanchard, 1852)    iN  M T

Pseudolucia clarea Bálint & K. Johnson, 1993  TL: Chile  M T

Pseudolucia collina (Philippi, 1859)    iN  M T

Pseudolucia dubi Bálint, 2001    iN

Pseudolucia grata (Köhler, 1934)    iN  M T

Pseudolucia hazearum Bálint & K. Johnson, 1993  TL: Chile  M T

Pseudolucia henyah Bálint, Benyamini & K. Johnson, 2001  TL: Chile  M T

Pseudolucia humbert Bálint & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia jujuyensis Bálint, Eisele & K. Johnson, 2000  TL: Argentina

Pseudolucia kechico Bálint, Benyamini & K. Johnson, 2001  TL: Chile

Pseudolucia kinbote Bálint & K. Johnson, 1993  TL: Chile

Pseudolucia lanin Bálint & K. Johnson, 1993    iN  M T

Pseudolucia magellana Benyamini, Bálint & K. Johnson, 1995  TL: Chile

Pseudolucia neuqueniensis Bálint & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia oligocyanea (Ureta, 1956)  TL: Chile  M T

Pseudolucia oraria Bálint & Benyamini, 2001    iN

Pseudolucia parana Bálint, 1993    iN  M T

Pseudolucia patago (Mabille, 1889)    iN  M T

Pseudolucia penai Bálint & K. Johnson, 1993  TL: Chile  M T

Pseudolucia plumbea (A. Butler, 1881)  TL: Chile  M T

Pseudolucia scintilla (Balletto, 1993)  TL: Chile  M T

Pseudolucia shapiroi Bálint & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia sibylla (W. F. Kirby, 1871)  TL: Chile  M T

Pseudolucia talia Bálint, Benyamini & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia tamara Bálint & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia ugartei Bálint & Benyamini, 2001  TL: Chile

Pseudolucia vera Bálint & K. Johnson, 1993    iN  M T

Pseudolucia whitakeri Bálint & K. Johnson, 1995  TL: Argentina  M T

Pseudolucia zina Benyamini, Bálint & K. Johnson, 1995    iN  M T

Pseudolucia barrigai Benyamini & Bálint, 2011  TL: not specified

Pseudolucia faundezi Benyamini & Bálint, 2011  TL: not specified

Pseudolucia johnsoni Benyamini & Bálint, 2011  TL: not specified

Pseudolucia luzmaria Benyamini & Bálint, 2011  TL: not specified

Pseudolucia munozae Benyamini & Bálint, 2011  TL: not specified

Pseudolucia zoellneri Benyamini & Bálint, 2011  TL: not specified

Pseudolucia sigal Benyamini & Bálint, 2011  TL: not specified

Pseudolucia valentina Benyamini & Bálint, 2011    iN

Pseudolucia atutia Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia calingasta Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia kieslingi Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia maricunga Bálint & Benyamini, 2013  TL: Chile

Pseudolucia mercedaria Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia ofir Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia pilgrama Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia sosneada Benyamini & Bálint, 2016  TL: Argentina

Pseudolucia ulibarrii Benyamini & Bálint, 2016  TL: Argentina

Nabokovia  Hemming, 1960  [includes Eldoradina as a subgenus]; ref.146

Nabokovia cyanea Balletto, 1993  TL: Peru  M T

Nabokovia sylphis (Draudt, 1921)  TL: Peru  M T

Nabokovia cuzquenha Bálint & Lamas, [1997]    iN  M T

Nabokovia faga (Dognin, 1895)    iN

Nabokovia faga faga (Dognin, 1895)  TL: Ecuador  M T

Nabokovia faga excisicosta (Dyar, 1913)  TL: Peru  M T

Nabokovia ada Bálint & K. Johnson, 1994  TL: Chile

Hemiargus  Hübner, 1818  

Hemiargus martha (Dognin, 1887)    iN  M T

Hemiargus [n. sp.] Lamas, MS  Peru

Hemiargus huntingtoni (Rindge & W. Comstock, 1953)    iN

Hemiargus huntingtoni hannoides Clench, 1965  Yucatan Peninsula to Panama  M T S

Hemiargus huntingtoni continentalis Clench, 1965  TL: Colombia  M T

Hemiargus huntingtoni huntingtoni (Rindge & W. Comstock, 1953)  TL: Trinidad  M T

Hemiargus ceraunus (Fabricius, 1793)  Ceraunus Blue    iN

Hemiargus ceraunus gyas (W. H. Edwards, 1871)  SW USA, NW Mexico  M T S L I H

Hemiargus ceraunus astenidas (Lucas, 1857)  [= zachaeina], C & S TX, E & S Mexico to Panama  M T S Ab HS L1 L2 I

Hemiargus ceraunus antibubastus Hübner, [1818]  Florida (vagrant N), Grand Bahama I.  M T S L1 L2

Hemiargus ceraunus filenus (Poey, 1832)  Cuba, Isla de Juventud, Caymans, Grand Turk, Gr. Inagua, etc.  M T S L

Hemiargus ceraunus ceraunus (Fabricius, 1793)  Jamaica, Hispaniola, some Bahama Islands (New Prov., etc.)  M T S L

Hemiargus hanno (Stoll, 1790)  Hanno Blue    iN

Hemiargus hanno watsoni W. Comstock & Huntington, 1943  Mona, Puerto Rico, S through Lesser Antilles to Grenada  M T S L

Hemiargus hanno bogotana Draudt, 1921  Panama and Colombia  M T

Hemiargus hanno hanno (Stoll, 1790)  range uncertain, mainly S America; ceraunus is not conspecific  M T S L

Hemiargus ramon (Dognin, 1887)    iN  M T

Echinargus  Nabokov, 1945  

Echinargus isola (Reakirt, [1867])  Reakirt’s Blue  [= alce],  iN  M T S L1 L2 L3 L4 L5 I Ha

Cyclargus  Nabokov, 1945  

Cyclargus thomasi (Clench, 1941)  Miami Blue    iN

Cyclargus thomasi thomasi (Clench, 1941)  Great Bahama Bank; ref.244  M T S L

Cyclargus thomasi bahamensis (Clench, [1943])  Crooked I., Acklins I., Plana Cays, Ragged I.; ref.244  M T S

Cyclargus thomasi clenchi (L. Miller, Simon & Harvey, 1992)  Mayaguana, Gr. & Little Iguana, Turks & Caicos banks; ref.244  M T S

Cyclargus thomasi noeli (W. Comstock & Huntington, 1943)  Hispaniola and satellite islands; ref.244  M T S L

Cyclargus thomasi bethunebakeri (W. Comstock & Huntington, 1943)  S Florida, Bimini Islands; ref.244  M T S L

Cyclargus woodruffi (W. Comstock & Huntington, 1943)  Miami Blue  [= oualiri] Mona P. Rico, Culebra and vic., Virgin Islands to L ...; ref.244  iN  M T S

Cyclargus ammon (Lucas, 1857)  Nickerbean Blue  FL Keys, Cuba, Isla de Juventud; ref.244  iN  M T S L

Cyclargus dominica (Möschler, 1886)  Jamaican Blue  Jamaica; ref.244  iN  M T S L

Icaricia  Nabokov, [1945]  

Icaricia emigdionis (F. Grinnell, 1905)  San Emigdio Blue  USA (Ca); ref.20  iN  M S L1 L2 L3 I

Icaricia shasta (W. H. Edwards, 1862)  Shasta Blue    iN

Icaricia shasta shasta (W. H. Edwards, 1862)  TL: USA (Ca)  M T S L1 L2 Ha

Icaricia shasta calchas (Behr, 1867)  TL: USA (Ca)  M T

Icaricia shasta NW fringe Great Basin segregate (, )  White Mts. CA/NV to Warner Mts. CA/OR, E to Harney Co., OR  M S L

Icaricia shasta pallidissima Austin, 1998  TL: USA (Nv)  M T1 T2 S L Ha

Icaricia shasta minnehaha (Scudder, 1874)  TL: USA (ND)  M T S L

Icaricia shasta charlestonensis (Austin, 1980)  TL: USA (Nv)  M S

Icaricia shasta pitkinensis (Ferris, 1976)  [= platazul]  M T S L1 L2 Ha

Icaricia neurona (Skinner, 1902)  Veined Blue    iN  M S1 S2 L1 L2 I

Icaricia dedeckera J. Emmel, T. Emmel & Mattoon, 1998  Lupine Blue  ref.ref. Davenport18  M T1 T2 S

Icaricia chlorina (Skinner, 1902)  Chlorina Blue  ref.ref. Davenport18

Icaricia chlorina chlorina (Skinner, 1902)  TL: USA (Ca)  M T S L Ha

Icaricia chlorina argentata J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S

Icaricia monticola (Clémence, 1909)  Monticola Blue  Baja California Norte; ref.ref. Davenport18  M T S1 S2 L

Icaricia lupini (Boisduval, 1869)  Lupine Blue    iN

Icaricia lupini Cascades, high elevations complex segregate (, )  C Cascades of OR to S WA, on Eriogonum pyrolifolium  M S1 S2 L1 L2 H Ha

Icaricia lupini spangelatus (Burdick, 1942)  TL: USA (Wa)  M S Ab

Icaricia lupini SE Oregon segregate (, )  M L

Icaricia lupini W Columbia Basin segregate (, )  M S L I Ha1 Ha2

Icaricia lupini SE Cascades, OR, NE CA segregate (, )  M S L1 L2 Ha

Icaricia lupini Siskiyous, W OR Cascades segregate (, )  M S Ha

Icaricia lupini N OR Coast Range segregate (, )  Polk Co., OR  M S

Icaricia lupini lutzi (dos Passos, 1938)  TL: USA (Id)  M S L I Ha

Icaricia lupini lupini (Boisduval, 1869)  TL: USA (Ca)  M T S L

Icaricia lupini alpicola J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L

Icaricia lupini goodpasturei Austin, 1998  TL: USA (Nv)  M T S L

Icaricia lupini texanus (Goodpasture, 1973)  TL: USA (Az)  M T S L1 L2 I Ha

Icaricia acmon (Westwood, [1851])  Acmon Blue  WA, OR, NV, CA & Baja California  iN  M T S1 S2 L1 L2 L3 L4 I Ha

Icaricia cotundra (Scott & M. Fisher, 2006)  Cotundra Blue    iN  M T S L Ha

Icaricia icarioides (Boisduval, 1852)  Boisduval's Blue    iN

Icaricia icarioides evius (Boisduval, 1869)  Baja California Norte  M T S L1 L2 H

Icaricia icarioides santana J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T1 T2 Ha

Icaricia icarioides moroensis (Sternitzky, 1930)  TL: USA (Ca)  M T S L1 L2 H Ha

Icaricia icarioides atascadero J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T1 T2 L

Icaricia icarioides pardalis (Behr, 1867)  TL: USA (Ca)  M T S1 S2 L

Icaricia icarioides missionensis (Hovanitz, 1937)  TL: USA (Ca)  M T S L

Icaricia icarioides pheres (Boisduval, 1852)  TL: USA (Ca)  M S1 S2

Icaricia icarioides parapheres J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S

Icaricia icarioides fenderi (Macy, 1931)  TL: USA (Or)  M T L I

Icaricia icarioides Mary's Peak, Benton Co., W OR segregate (, )  M S Ha

Icaricia icarioides blackmorei (W. Barnes & McDunnough, 1919)  TL: Canada (BC)  M T L I Ha

Icaricia icarioides montis (Blackmore, 1923)  TL: Canada (BC)  M S L

Icaricia icarioides N Cascades, OR, high elevations segregate (, )  M S L Ha

Icaricia icarioides S Cascades, OR, high elevations segregate (, )  M S L Ha

Icaricia icarioides helios (W. H. Edwards, 1871)  TL: USA (Ca)  M T S

Icaricia icarioides icarioides (Boisduval, 1852)  TL: USA (Ca)  M T S L1 L2 Ha

Icaricia icarioides eosierra J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L

Icaricia icarioides albihalos J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S L1 L2 L3 H

Icaricia icarioides inyo J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T1 T2

Icaricia icarioides argusmontana J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T S

Icaricia icarioides panamintina J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Ca)  M T1 T2

Icaricia icarioides austinorum J. Emmel, T. Emmel & Mattoon, 1998  TL: USA (Nv)  M T S

Icaricia icarioides fulla (W. H. Edwards, 1870)  TL: USA (Ca)  M T S L Ha

Icaricia icarioides pembina (W. H. Edwards, 1862)  TL: USA (Mt)  M S Ab L1 L2 I Ha

Icaricia icarioides lycea (W. H. Edwards, 1864)  TL: USA (Co)  M S Ab L I

Icaricia icarioides nigrafem (R. Holland, 2011)  TL: USA (NM)  M OI

Icaricia icarioides sacre (R. Holland, 2011)  TL: USA (NM)  M OI

Icaricia icarioides buchholzi (dos Passos, 1938)  Chihuahua  M S L

Icaricia saepiolus (Boisduval, 1852)  Greenish Blue    iN

Icaricia saepiolus amica (W. H. Edwards, 1863)  TL: Canada (NT)  M S L

Icaricia saepiolus insulanus (Blackmore, 1920)  TL: Canada (BC)  M S

Icaricia saepiolus littoralis (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T1 T2 T3 S

Icaricia saepiolus saepiolus (Boisduval, 1852)  TL: USA (Ca)  M T TI S

Icaricia saepiolus rufescens (Boisduval, 1869)  TL: USA (Ca)  M S L I Ha

Icaricia saepiolus aehaja (Behr, 1867)  TL: USA (Ca)  M T S L1 L2 L3 I Ha

Icaricia saepiolus albomontanus (Austin & J. Emmel, 1998)  TL: USA (Ca)  M T1 T2 S L

Icaricia saepiolus aureolus (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Icaricia saepiolus hilda (J. Grinnell & F. Grinnell, 1907)  Hypothetical for Baja California Norte  M T S L

Icaricia saepiolus maculosus (Austin, 1998)  TL: USA (Nv/Ut)  M T S L

Icaricia saepiolus gertschi (dos Passos, 1938)  TL: USA (Ut)  M S Ab L

Icaricia saepiolus whitmeri (F. Brown, 1951)  TL: USA (Co)  M T S Ab L1 L2 Ha

Agriades  Hübner, [1819]  

Agriades optilete (Knoch, 1781)  Cranberry Blue    iN

Agriades optilete yukona (W. Holland, 1900)  TL: USA (Ak)  M S L Ha

Agriades glandon (de Prunner, 1798)  Arctic Blue    iN

Agriades glandon labrador Schmidt, Scott & Kondla, 2006  TL: Canada (NL)  M S

Agriades glandon franklinii (J. Curtis, 1835)  TL: Canada (Nu)  M S

Agriades glandon lacustris (T. Freeman, 1939)  [= manitoba]  M T S

Agriades glandon bryanti (Leussler, 1935)  TL: Canada (NT)  M S L Ha

Agriades glandon megalo (McDunnough, 1927)  TL: Canada (BC)  M T S L

Agriades glandon rustica (W. H. Edwards, 1865)  TL: USA (Co)  M S L1 L2 I1 I2 Ha

Agriades glandon punctatus Austin, 1998  TL: USA (Az)  M T S L1 L2 I H

Agriades glandon cassiope J. Emmel & T. Emmel, 1998  TL: USA (Ca)  M T S L H

Agriades glandon kelsoni J. Emmel & T. Emmel, 1998  TL: USA (Ca)  M T S

Agriades podarce (C. Felder & R. Felder, 1865)  Sierra Blue    iN

Agriades podarce cilla (Behr, 1867)  TL: USA (Ca)  M T S L1 L2 L3 H

Agriades podarce podarce (C. Felder & R. Felder, 1865)  TL: USA (Ca)  M S

Agriades podarce klamathensis J. Emmel & T. Emmel, 1998  TL: USA (Ca)  M T S L1 L2 Ha

Plebejus  Kluk, 1780  [includes Arianna as a subgenus]; ref.154

Plebejus idas (Linnaeus, 1761)  Northern Blue    iN

Plebejus idas empetri (T. Freeman, 1938)  TL: Canada (NS)  M T S

Plebejus idas aster (W. H. Edwards, 1882)  TL: Canada (NL)  M S1 S2

Plebejus idas scudderii (W. H. Edwards, 1861)  boreal Canada (except far W)  M S

Plebejus idas nabokovi (Masters, 1972)  TL: USA (Mn)  M S

Plebejus idas alaskensis (F. Chermock, 1945)  Alaska, W YK  M T1 T2 S L Ha

Plebejus idas ferniensis (F. Chermock, 1945)  TL: Canada (BC)  M T S

Plebejus idas atrapraetextus (W. D. Field, 1939)  TL: USA (Id)  M T1 T2 S

Plebejus idas NE Oregon segregate ,  [nr. atrapraetextus]  M L1 L2 L3 Ha

Plebejus idas longinus (Nabokov, 1949)  TL: USA (Wy)  M S L Ha

Plebejus idas sublivens (Nabokov, 1949)  montane SW Colorado  M S

Plebejus fridayi (F. Chermock, 1945)  Friday's Blue    iN

Plebejus fridayi fridayi (F. Chermock, 1945)  TL: USA (Ca)  M T S L

Plebejus fridayi White Mountains segregate ,   M L

Plebejus anna (W. H. Edwards, 1861)  Anna's Blue    iN

Plebejus anna anna (W. H. Edwards, 1861)  TL: USA (Ca)  M T S L1 L2 L3 L4

Plebejus anna ricei (Cross, 1937)  TL: USA (Or)  M T S L1 L2 L3 I Ha

Plebejus anna lotis (Lintner, 1878)  TL: USA (Ca)  M S

Plebejus anna azureus (J. Emmel, T. Emmel & Mattoon, 1998)  TL: USA (Ca)  M T S

Plebejus anna vancouverensis (C. Guppy & J. Shepard, 2001)  TL: Canada (BC)  M T S

Plebejus anna benwarner Scott, 2006  TL: USA (Ca)  M S L1 L2 Ha

Plebejus melissa (W. H. Edwards, 1873)  Melissa Blue    iN

Plebejus melissa melissa (W. H. Edwards, 1873)  TL: USA (Co)  M T S Ab L1 L2 L3 L4 I

Plebejus melissa pseudosamuelis (Nabokov, 1949)  TL: USA (Co)  M T S L Ha

Plebejus melissa annetta (W. H. Edwards, 1882)  montane N UT  M S

Plebejus melissa alateres (Austin, 1998)  TL: USA (Nv)  M T S

Plebejus melissa inyoensis (Nabokov, 1949)  TL: USA (Ca)  M S1 S2 L1 L2 L3 Ha

Plebejus melissa paradoxa (F. Chermock, 1945)  Baja California Norte  M T S L

Plebejus melissa mexicana (Clench, 1965)  N Sonora, N Chihuahua  M T S

Plebejus samuelis (Nabokov, 1944)  Karner Blue    iN  M S1 S2 L1 L2 I Ha

Polyommatus  Latreille, 1804  

Polyommatus icarus (Rottemburg, 1775)  Common Blue  Introduced to SE Canada  iN

Polyommatus icarus icarus (Rottemburg, 1775)  TL: Germany  M S

  Tribe Polyommatini, Subtribe Undetermined

Genera-unknown1  , []  

Genera-unknown1 planuma (K. Johnson, 1992)  Argentina; nomen dubium

Family HESPERIIDAE Latreille, 1809 – SKIPPERS

  Subfamily Eudaminae

  Tribe Entheini

Udranomia  A. Butler, 1870  

Udranomia eurus (Mabille & Boullet, 1919)  Eurus Skipper  Costa Rica to Venezuela & Amazon basin  iN  M S I

Udranomia tomdaleyi Burns, 2017  Costa Rica; ref.21

Udranomia sallydaleyae Burns, 2017  Costa Rica; ref.21

Udranomia kikkawai (Weeks, 1906)  Nervous Skipper  E & W Mexico to Brazil; ref.21  iN  M T TI S1 S2 S3 L I1 I2

Udranomia orcinus (C. Felder & R. Felder, 1867)  Orcinus Skipper  SE Mexico to Bolivia  iN  M T S1 S2 IL I G

Udranomia spitzi (Hayward, 1942)  Brazil  iN  M S

Drephalys  E. Watson, 1893  

Drephalys oria Evans, 1952  Oria Skipper  Mexico to Honduras  iN  M T S G

Drephalys oriander (Hewitson, 1867)  Oriander Skipper  Mexico, Costa Rica to Guianas and N Brazil  iN  M T S1 S2 IL G

Drephalys talboti (Le Cerf, 1922)  TL: French Guiana  M T S G

Drephalys dumeril (Latreille, [1824])  Dumeril Skipper  Mexico to Ecuador and Brazil  iN  M T OI OD S1 S2 IL G

Drephalys croceus Austin, 1995  TL: Brazil (Rd)  M T OI G

Drephalys tortus Austin, 1995  TL: Brazil (Rd)  M T OI G

Drephalys alcmon (Cramer, 1780)  Alcmon Skipper  Guatemala to Guianas and Brazil  iN  M T S1 S2 S3 IL L I G

Drephalys mourei O. Mielke, 1968  SE & S Brazil  iN  M T IL

Drephalys olvina Evans, 1952  TL: Guyana  M S

Drephalys kidonoi Burns, 2000  Kidono's Skipper  W Costa Rica  M T S1 S2 I G

Drephalys helixus (Hewitson, 1877)  Helixus Skipper  Panama  iN  M T OD S IL G

Drephalys phoenice (Hewitson, 1867)  TL: Brazil (Am)  M T S IL

Drephalys heraclides E. Bell, 1942  TL: Peru  M T S

Drephalys miersi O. Mielke, 1968  TL: Brazil (SC)  M T S IL

Drephalys phoenicoides (Mabille & Boullet, 1919)  Curious Skipper  Costa Rica to Guianas and NE Brazil  iN  M S

Drephalys dracarys Madruga, Siewert, O. Mielke & Dolibaina, 2018  Brazil; ref.35

Drephalys electrinus Siewert, Madruga, O. Mielke & Dolibaina, 2018  Brazil; ref.35

Drephalys citrinus Madruga, Siewert, O. Mielke & Casagrande, 2018  Brazil; ref.35

Drephalys eous (Hewitson, 1867)  TL: Brazil (Pa)  M T S

Drephalys diovalis Grishin, 2023  Ecuador; ref.153

Drephalys opifex Evans, 1952  TL: Surinam  M OD

Drephalys olva Evans, 1952  TL: Brazil (Pa)  M T OD

Phanus  Hübner, [1819]  

Phanus australis L. Miller, 1965    iN  M T S

Phanus vitreus (Stoll, 1781)  Widespread Phanus  Oaxaca and Chiapas to southern Brazil  iN  M T S1 S2 L I G

Phanus ecitonorum Austin, 1993  Army-ant Phanus  Costa Rica to southern Brazil  iN  M T S L

Phanus confusis Austin, 1993  Confusing Phanus  eastern Mexico to El Salvador  iN  M T S

Phanus albiapicalis Austin, 1993  White-tipped Phanus  E & W Mexico to Panama, W Ecuador and Peru  iN  M T S L

Phanus rilma Evans, 1952  West-Mexican Phanus  Pacific coast, Nayarit to Guatemala  iN  M T S G

Phanus grandis Austin, 1993  Grand Phanus  Nicaragua, Costa Rica, Panama, Colombia, Venezuela  iN  M T S L

Phanus obscurior Kaye, 1925  Dark Phanus    iN

Phanus obscurior obscurior Kaye, 1925  SE Mexico to Colombia and Trinidad  M T S1 S2 I G

Phanus obscurior prestoni L. Miller, 1965  TL: Brazil (Am)  M T S

Phanus marshalli (W. F. Kirby, 1880)  Common Phanus  E & W Mexico to Bolivia  iN  M S1 S2 L I G

Hyalothyrus  Mabille, 1878  

Hyalothyrus infernalis (Möschler, 1877)    iN  M T S

Hyalothyrus infa Evans, 1952  Colombia; ref.7  M T S

Hyalothyrus leucomelas (Geyer, 1832)  Leucomelas Skipper  Panama to Brazil and Peru  iN  M S

Hyalothyrus nitocris (Stoll, 1782)    iN  M S

Hyalothyrus mimicus Mabille & Boullet, 1919    iN  M S

Hyalothyrus neleus (Linnaeus, 1758)    iN

Hyalothyrus neleus pemphigargyra (Mabille, 1888)  SE Mexico to Colombia and Venezuela  M T S L I G

Hyalothyrus neleus neleus (Linnaeus, 1758)  TL: Indiis  M T S L

Hyalothyrus neda Evans, 1952  Bolivia; ref.7  M T S

Entheus  Hübner, [1819]  

Entheus eumelus (Cramer, 1777)    iN  M T OI OD S IL G

Entheus ninyas H. Druce, 1912  TL: Bolivia  M T OI OD S

Entheus eunyas Austin, O. Mielke & Steinhauser, 1997    iN  M T OI S G

Entheus lemna (A. Butler, 1870)  TL: ?  M T OD TI S IL Txt

Entheus gentius (Cramer, 1777)    iN  M T OI OD TI S1 S2 S3 HS IL G Txt

Entheus bombus Austin, O. Mielke & Steinhauser, 1997    iN  M T OI S G

Entheus aureolus Austin, O. Mielke & Steinhauser, 1997    iN  M T OI S1 S2 S3 S4 S5 S6 IL G

Entheus huertasae Grishin, 2013  Blanca's Entheus  Panama & Colombia; ref.70  M T IL G

Entheus telemus Mabille, 1898  TL: Brazil  M OD S IL G

Entheus latebrosus Austin, 1997    iN  M T OI S

Entheus priassus (Linnaeus, 1758)    iN

Entheus priassus priassus (Linnaeus, 1758)  TL: Indiis  M OI OD TI S HS IL G Txt

Entheus priassus pralina Evans, 1952  TL: Brazil (ES)  M T IL G

Entheus aureanota Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M T OI S G

Entheus curvus Austin, 1997  TL: Peru  M T OI S G

Entheus matho Godman & Salvin, 1879  Giant Entheus    iN

Entheus matho matho Godman & Salvin, 1879  SE Mexico to Costa Rica  M T OD S IL L G Txt

Entheus matho latifascius M. Hering, 1925  TL: Colombia  M OI OD S

Entheus matho marmato Salazar & Vargas, [2017]  Colombia; ref.91

Entheus matho aequatorius Mabille & Boullet, 1919  Costa Rica to Ecuador  M OD S IL G

Entheus matho dius Mabille, 1898  Panama Guianas, Peru, and Bolivia  M T OD TI S IL G

Entheus crux Steinhauser, 1989  Mexican Entheus  SE Mexico to Nicaragua  iN  M T OD S G

Entheus warreni Grishin, 2012  Warren's Entheus  Ecuador; ref.63  M T S G

Augiades  Hübner, [1819]  

Augiades crinisus (Cramer, 1780)  Crinisus Skipper  Costa Rica to Bolivia and Brazil  iN  M T S L

Augiades vespasius (Fabricius, 1793)  [= epimethea]; ref.38  iN

Augiades vespasius bicolor (Mabille & Boullet, 1919)  Costa Rica to Bolivia, Guianas and N Brazil  M T S

Augiades vespasius vespasius (Fabricius, 1793)  TL: Indiis  M T

Tarsoctenus  E. Watson, 1893  

Tarsoctenus praecia (Hewitson, 1857)    iN

Tarsoctenus praecia praecia (Hewitson, 1857)  TL: Brazil (Pa)  M T OI S

Tarsoctenus praecia rufibasis Boullet, 1910  TL: French Guiana  M T IL

Tarsoctenus praecia plutia (Hewitson, 1857)  TL: Brazil (Am)  M T OI S L

Tarsoctenus praecia luna Evans, 1952  TL: Bolivia  M T

Tarsoctenus gaudialis (Hewitson, 1876)  Gaudy Skipper  Nicaragua to Colombia; ref.16  iN  M T OI S IL L G

Tarsoctenus corytus (Cramer, 1777)    iN

Tarsoctenus corytus corytus (Cramer, 1777)  TL: Surinam  M T OI OD S

Tarsoctenus corytus corba Evans, 1952  TL: Peru  M T S

Tarsoctenus papias (Hewitson, 1857)    iN  M T OI S L

  Tribe Phocidini

Phocides  Hübner, [1819]  

Phocides lilea (Reakirt, [1867])  Guava Skipper  S Texas to Costa Rica; ref.11  iN  M T S1 S2 L1 L2 I1 I2 G

Phocides polybius (Fabricius, 1793)  Guava Skipper    iN

Phocides polybius polybius (Fabricius, 1793)  [= imbreus] Panama, Colombia, Venezuela, Guianas, Trinidad; ref.11  M T S IL L G

Phocides polybius phanias (Burmeister, 1880)  TL: Argentina  M S L

Phocides charon (C. Felder & R. Felder, 1859)    iN  M T S IL

Phocides petroleum Siewert, Leviski, O. Mielke & Casagrande, 2018  Bolivia (La Paz); ref.34

Phocides distans (Herrich-Schäffer, 1869)    iN

Phocides distans licinus (Möschler, 1879)  Honduras, Costa Rica, Panama  M T S L G

Phocides distans distans (Herrich-Schäffer, 1869)  TL: ?  M T S IL

Phocides distans silva Evans, 1952  TL: Peru  M T S

Phocides perillus (Mabille, 1888)    iN  M T OI S

Phocides metrodorus E. Bell, 1932  Metrodorus Skipper    iN

Phocides metrodorus nigrescens E. Bell, 1938  Costa Rica, Colombia, Venezuela, Ecuador, Peru; species?  M T S1 S2 I1 I2

Phocides metrodorus metrodorus E. Bell, 1932  Panama to Bolivia and Brazil  M T S G

Phocides metrodorus metron Evans, 1952  TL: Paraguay  M T OD G

Phocides vulcanides Röber, 1925    iN  M T S

Phocides novalis Evans, 1952    iN  M T S G

Phocides thermus (Mabille, 1883)  Thermus Skipper    iN

Phocides thermus thermus (Mabille, 1883)  SE Mexico to Venezuela and Ecuador  M T S G

Phocides thermus bellina Evans, 1952  TL: Ecuador  M T S

Phocides thermus valgus (Mabille, 1883)  TL: French Guiana  M T S IL

Phocides partia Evans, 1952  Peru & Bolivia  M T G

Phocides padrona Evans, 1952    iN  M T S G

Phocides belus Godman & Salvin, 1893  Belus Skipper  S Texas (one-time stray) to Costa Rica  iN  M T S1 S2 L I

Phocides pigmalion (Cramer, 1779)  Pigmalion Skipper    iN

Phocides pigmalion pigmalion (Cramer, 1779)  TL: Surinam  M T OI IL G

Phocides pigmalion hewitsonius (Mabille, 1883)  TL: Brazil (Am)  M T TI IL

Phocides batabano (Lucas, 1857)  Mangrove Skipper  Cuba, I. de Juventud, Andros, Little Cayman; ref.12

Phocides batabano okeechobee (Worthington, 1881)  S Florida; mangroves  M T S L1 L2 I G

Phocides batabano batabanoides (W. Holland, 1902)  most Bahamian Islands  M T S

Phocides batabano batabano (Lucas, 1857)  Cuba, I. de Juventud, Andros, Little Cayman  M T S G

Phocides bicolora (Boddaert, 1783)  'Hispaniolan' Mangrove Skipper  Hispaniola; ref.12  M T S

Phocides johnsoni E. Bell, 1947    iN  M T S

Phocides urania (Westwood, 1852)  Urania Skipper  Mexico, Guatemala; usually in montane forest  iN  M T S L G

Phocides vida (A. Butler, 1872)  Vida Skipper  Costa Rica; ref.13  M T S

Phocides pialia (Hewitson, 1857)    iN

Phocides pialia maximus (Mabille, 1888)  TL: Brazil  M T OI S

Phocides pialia intermedia O. Mielke, 1992  TL: Brazil (MG)  M OI S

Phocides pialia pialia (Hewitson, 1857)  TL: Brazil  M T OI S G

Phocides oreides (Hewitson, [1875])    iN

Phocides oreides colombiana E. Bell, 1938  TL: Colombia  M T

Phocides oreides oreides (Hewitson, [1875])  TL: Peru  M T OI S IL

Phocides yokhara (A. Butler, 1870)    iN

Phocides yokhara yokhara (A. Butler, 1870)  TL: Peru  M T S L

Phocides yokhara inca Le Cerf, 1922  TL: Peru  M T

Phocides yokhara dryas Le Cerf, 1922  TL: Peru  M T

Phocides yokhara charonotis (Hewitson, 1874)  TL: Bolivia  M T TI

Phocides lincea (Herrich-Schäffer, 1869)  TL: ?  M S

Phocides perkinsi (Kaye, 1931)  Perkins' Skipper  Jamaica; ref.77  M T S

Nascus  E. Watson, 1893  [includes Pseudonascus, Praxa & Bron as subgenera]; ref.13

Nascus paulliniae (Sepp, [1842])  Least Scarlet-eye  S Mexico to Bolivia and S Brazil; ref.13  iN  M T OI S1 S2 I1 I2 G

Nascus prax Evans, 1952  Brazil (Pa); ref.13  iN  M T

Nascus broteas (Cramer, 1780)  Broteas Scarlet-eye  [= solon] E Mexico to Argentina, S Brazil and Paraguay; ref.13,146  iN  M T S1 S2 I1 I2 G

Nascus corilla Evans, 1952  Corilla Scarlet-eye  S Mexico to Colombia and Venezuela  M T S1 S2 I1 I2

Nascus lux Grishin, 2023  Luminous Scarlet-eye  [= Evans' "solon solon"]; ref.146  iN  M S L

Nascus phintias Schaus, 1913  Schaus’ Scarlet-eye  SE Mexico to French Guiana and Peru; ref.13  iN  M T S1 S2 IL I

Nascus phocus (Cramer, 1777)  Common Scarlet-eye  E & W Mexico to S Brazil, Paraguay and Argentina; ref.13  iN  M T S1 S2 IL L I1 I2 G

Aurina  Evans, 1937  

Aurina dida Evans, 1937  French Guiana; ref.7  M T OI OD S G Txt

Aurina azines (Hewitson, 1867)    iN  M T OI S L

Emmelus  O. Mielke & Casagrande, 2016  

Emmelus purpurascens (Mabille & Boullet, 1912)  Peru, Brazil; ref.119,7  iN  M OD S Co G Txt

Porphyrogenes  E. Watson, 1893  [includes Ocyba as subgenus]; ref.7

Porphyrogenes stupa Evans, 1952  TL: ?  M T

Porphyrogenes omphale (A. Butler, 1871)  Blue Scarlet-eye  Costa Rica, Panama, Venezuela, Guianas to Bolivia  iN  M T OI S1 S2 L G

Porphyrogenes passalus (Herrich-Schäffer, 1869)  Passalus Scarlet-eye  Panama, Colombia, Bolivia, Brazil (at least)  M T S

Porphyrogenes speciosus Austin & O. Mielke, 2008  Speciosus Scarlet-eye  Costa Rica  M T G

Porphyrogenes convexus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes sparus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes spadix Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes peterwegei Burns, 2010  Peter's Scarlet-eye  NW Costa Rica  M T1 T2 T3 S I1 I2 H G

Porphyrogenes sororcula (Mabille & Boullet, 1912)  TL: French Guiana  M T OI S

Porphyrogenes specularis Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes probus (Möschler, 1877)    iN  M T IL

Porphyrogenes vulpecula (Plötz, 1882)    iN  M T S G

Porphyrogenes zohra (Möschler, 1879)  Zohra Scarlet-eye  Honduras, Nicaragua, Venezuela, Guianas, N Brazil (Para)  M T S G

Porphyrogenes virgatus (Mabille, 1888)  Virgatus Scarlet-eye  Honduras, Guyana, Peru  M T OI

Porphyrogenes eudemus (Mabille, 1888)  Eudemus Scarlet-eye  Panama, Peru  iN  M T OI TI S G

Porphyrogenes splendidus Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes simulator Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes sula (R. Williams & E. Bell, 1940)  Sula Scarlet-eye  Honduras, possibly Costa Rica  iN  M T S I G

Porphyrogenes spina Austin & O. Mielke, 2008  Spina Scarlet-eye  Panama  M T G

Porphyrogenes sporta Austin & O. Mielke, 2008  TL: Colombia  M OI

Porphyrogenes stresa Evans, 1952  TL: Peru  M T

Porphyrogenes boliva Evans, 1952  TL: Venezuela  M T

Porphyrogenes spoda Evans, 1952  Spoda Scarlet-eye  Panama  M T G

Porphyrogenes ferruginea (Plötz, 1883)  TL: Brazil (Ba)  M T S

Porphyrogenes despecta (A. Butler, 1870)  TL: Brazil (Pa)  M T S

Porphyrogenes pausias (Hewitson, 1867)  TL: Brazil (Am)  M T S

Porphyrogenes spanda Evans, 1952  TL: Brazil (Pa)  M T S

Porphyrogenes sparta Evans, 1952  TL: Brazil (Pa)  M T

Porphyrogenes glavia Evans, 1952  Glavia Scarlet-eye  Panama  M T G

Porphyrogenes unassociated_unnamed_female_#1 ,  TL: not specified  M S IL

Porphyrogenes unassociated_unnamed_female_#2 ,  TL: not specified  M IL

Porphyrogenes unassociated_unnamed_female_#3 ,  Mystery Scarlet-eye #3  Panama  M S

Porphyrogenes unassociated_unnamed_female_#4 ,  Mystery Scarlet-eye #4  Panama  M S

Porphyrogenes unassociated_unnamed_female_#5 ,  Mystery Scarlet-eye #5  Costa Rica  M S

Porphyrogenes unassociated_unnamed_females ,  Mystery Female Scarlet-eyes  Porphyrogenes females whose identity remains a mystery  M S HS

Porphyrogenes unassociated_unnamed_male ,  Mystery Scarlet-eye #6  Costa Rica  M S G

Porphyrogenes calathana (Hewitson, 1868)  Yellow-rimmed Scarlet-eye  ref.7  iN

Porphyrogenes calathana calanus (Godman & Salvin, 1894)  Mexico to Costa Rica  M T OI S1 S2 IL L I G

Porphyrogenes calathana calathana (Hewitson, 1868)  Panama and Colombia  M T S IL

Porphyrogenes calathana compusa (Hewitson, 1868)  TL: Brazil (Am)  M T

Nicephellus  Austin, 2008  

Nicephellus nicephorus (Hewitson, 1876)  Two-spotted Scarlet-eye  SE Mexico to Guianas, N & S Brazil, NE Peru  iN  M T S IL I

Salatis  Evans, 1952  

Salatis canalis (Skinner, 1920)  Canal Scarlet-eye  Costa Rica to Ecuador  iN  M T S1 S2 I1 I2

Salatis salatis (Stoll, 1782)  Variable Scarlet-eye  Nicaragua, Panama to Guianas, Peru and S Brazil  iN  M T S IL L G

Salatis flavomarginatus (Sepp, [1851])  very long apiculus, no specimen known, Suriname  M OI I

Salantoia  Grishin, 2019  ref.7

Salantoia eriopis (Hewitson, 1867)  Brazil; ref.7  iN  M T S

Salantoia metallica Grishin, 2022  Guyana; ref.13  M T

Salantoia gildo (Mabille, 1888)  Brazil (Am); ref.13  M T OI

Ornilius  Grishin, 2022  ref.13

Ornilius rotundus Grishin, 2022  [= Evans' "cebrenus"]; ref.13  M T

Adina  Grishin, 2022  ref.13

Adina adrastor (Mabille & Boullet, 1912)  [= dinka] S. Amer.; ref.13,140  M T OI

Sarmientoia  Berg, 1897  

Sarmientoia phaselis (Hewitson, 1867)  TL: Brazil  M T S IL G

Sarmientoia faustinus (Burmeister, 1878)    iN  M T S IL G

Sarmientoia haywardi O. Mielke, 1967    iN  M OI S IL G

Sarmientoia similis O. Mielke, 1967    iN  M OI S IL G

Sarmientoia almeidai O. Mielke, 1967  TL: Brazil (MT)  M OI S G

Sarmientoia browni O. Mielke, 1967  TL: Brazil (DF)  M OI S G

Fulvatis  Grishin, 2022  ref.13

Fulvatis fulvius (Plötz, 1882)  Brazil (Pa); ref.13  M T S HS

Fulvatis scyrus (E. Bell, 1934)  Peru; ref.13  M T

Bungalotis  E. Watson, 1893  

Bungalotis erythus (Cramer, 1775)  Spotted Scarlet-eye  SE Mexico to Guianas, Brazil and Bolivia  iN  M S1 S2 I1 I2 G

Bungalotis corentinus (Plötz, 1882)  [= diophorus, = gagarini]; ref.13,140  iN  M T OI S1 S2 IL I1 I2 G

Bungalotis quadratum (Sepp, [1845])  Pallid Scarlet-eye    iN

Bungalotis quadratum quadratum (Sepp, [1845])  SE Mexico to Guianas  M T OI S1 S2 L I1 I2

Bungalotis quadratum barba Evans, 1952  TL: Brazil (Pa)  M T

Bungalotis lactos Evans, 1952  Lactos Scarlet-eye  TL: Peru  M T G

Bungalotis clusia Evans, 1952  Clusia Scarlet-eye  TL: Brazil (Pa)  M T

Bungalotis sipa de Jong, 1983  Sipa Scarlet-eye  Surinam  M T G

Bungalotis borax Evans, 1952  Borax Scarlet-eye  TL: French Guiana  M T S G

Bungalotis milleri H. Freeman, 1977  Miller's Scarlet-eye  SE Mexico to Costa Rica  iN  M T OD G

Bungalotis astylos (Cramer, 1780)  Dark-cheeked Scarlet-eye  SE Mexico to Guianas, Peru and S Brazil  iN  M T S1 S2 L I1 I2 G

Bungalotis aureus Austin, 2008  Golden Scarlet-Eye  TL: Ecuador  M OI G

Bungalotis midas (Cramer, 1775)  White-cheeked Scarlet-eye  SE Mexico to S Brazil and Paraguay  iN  M S1 S2 IL L I1 I2 G

Dyscophellus  Godman & Salvin, 1893  

Dyscophellus porcius (C. Felder & R. Felder, 1862)  Fiery Scarlet-eye  S Mexico to Paraguay and S Brazil  iN  M T S IL L G

Dyscophellus doriscus (Hewitson, 1867)  S Brazil; ref.13  M T S HS

Dyscophellus sebaldus (Stoll, 1781)  Sebaldus Scarlet-eye  Costa Rica to Guianas and N Brazil  iN  M OI S IL

Dyscophellus damias (Plötz, 1882)  [= erythras]; ref.13  iN  M T OI S

Dyscophellus mielkei Austin, 2008  TL: Brazil (Rd)  M OI

Dyscophellus ramon Evans, 1952  Plain Scarlet-eye  SE Mexico to Panama  iN  M T S1 S2 HS L I1 I2 G

Dyscophellus diaphorus (Mabille & Boullet, 1912)  Diaphorus Scarlet-eye  [part of Evans' ramusis] Panama to Guianas and N Brazil  M T OI

Dyscophellus ramusis (Stoll, 1781)  Subtle Scarlet-eye  in S. Amer. to S Brazil  iN

Dyscophellus ramusis astrapaeus (Hewitson, 1876)  Amazonian region  M T S HS

Dyscophellus ramusis ramusis (Stoll, 1781)  Colombia to Bolivia  M S IL

Dyscophellus australis Grishin, 2022  Southern Scarlet-eye  [= Evans' "damias"] S Brazil & Paraguay; ref.13  iN  M S L

Dyscophellus porsena (E. Bell, 1934)    iN  M T

Dyscophellus basialbus Grishin, 2022  White-base Scarlet-eye  [= Evans' "diaphorus"]; ref.13  iN  M T S

Euriphellus  Austin, 2008  

Euriphellus lama (Evans, 1952)  Big-spotted Scarlet-eye  SE Mexico to Honduras; ref.7  M T S

Euriphellus phraxanor (Hewitson, 1876)  Big-spotted Scarlet-eye  Nicaragua to Ecuador; ref.7  iN  M T S1 S2 L I G

Euriphellus panador Grishin, 2023  Ecuador; ref.153

Euriphellus panamicus Grishin, 2023  Panama; ref.153

Euriphellus mena (Evans, 1952)  Ecuador; ref.7  M T

Euriphellus marian (Evans, 1952)  Ecuador; ref.7  M T S

Euriphellus colombiensis Grishin, 2023  Colombia; ref.151  M T

Euriphellus ecuadoricus Grishin, 2023  Ecuador; ref.151  M T

Euriphellus cebrenus (Cramer, 1777)  [Evans' "cebrenus" is Ornilius rotundus] Suriname; ref.13  M TI S

Euriphellus euribates (Stoll, 1782)  Euribates Scarlet-eye  Costa Rica to Brazil and Bolivia, may be a synonym of cebrenus; ref.13  iN  M T OI S G

Euriphellus polygius (Latreille, [1824])  S Brazil; ref.7  M T IL

Phareas  Westwood, 1852  

Phareas burnsi Grishin, 2013  Burns' Skipper  Costa Rica, Panama, ?Colombia; ref.74  iN  M T1 T2 S I

Phareas coeleste Westwood, 1852  Magnificent Skipper  Colombia to Bolivia and S Brazil  iN  M T OI OD S1 S2 S3 IL L G

  Tribe Eudamini, Subtribe Eudamina

Cecropterus  Herrich-Schäffer, 1869  includes Thorybes as a subgenus; ref.7,126

Cecropterus acanthopoda (O. Mielke, 1977)  Brazil; ref.7  M T OI G

Cecropterus rinta (Evans, 1952)  Argentina; ref.7  M T S

Cecropterus zarex (Hübner, 1818)  Sharp Banded-Skipper  E & S Mexico to Argentina  iN  M T S1 S2 HS L I G

Cecropterus longipennis Plötz, 1882  Spike Banded-Skipper  E & S Mexico to S Brazil  iN  M T S L I G

Cecropterus evenus (Ménétriés, 1855)    iN  M S IL

Cecropterus lyciades (Geyer, 1832)  Hoary Edge  E USA, (Veracruz)  iN  M S1 S2 HS L1 L2 L3 L4 L5 I G

Cecropterus casica (Herrich-Schäffer, 1869)  Desert Cloudywing  SW USA (SE AZ to C TX), Mexico  iN  M T S L I H G

Cecropterus tehuacana (Draudt, 1922)  Tehuacan Cloudywing  S TX (rare stray), E Mexico to Guatemala  iN  M T OD S L G

Cecropterus confusis (E. Bell, 1923)  Confused Cloudywing  SE USA  iN  M T1 T2 S1 S2 S3 L1 L2 L3 I G

Cecropterus bathyllus (J. E. Smith, 1797)  Southern Cloudywing  E USA  iN  M OI TI S1 S2 S3 HS L1 L2 L3 L4 L5 L6 G

Cecropterus diversus (E. Bell, 1927)  Western Cloudywing  C & N Sierra Nevada and Siskiyou Mts., CA, far SW OR  iN  M S L1 L2 L3 I G

Cecropterus nevada (Scudder, 1872)  Sierra Nevada Cloudywing  ref.11  iN

Cecropterus nevada aemilea (Skinner, 1893)  Cascade Mts. N CA to C OR; ref.11  M T S L1 L2 Ha

Cecropterus nevada blanca (Scott, 1981)  White Mts., CA-NV; ref.11  M S L1 L2

Cecropterus nevada nevada (Scudder, 1872)  CA-NV Sierra Nevada; ref.11  M S L1 L2 Ha G

Cecropterus dobra (Evans, 1952)  Dobra Cloudywing  S Rocky Mts. (S WY-ID to NM, AZ) to NW Mexico; ref.11  iN  M T S1 S2 L1 L2 Ha

Cecropterus mexicana (Herrich-Schäffer, 1869)  Mexican Cloudywing  ref.11  iN

Cecropterus mexicana mexicana (Herrich-Schäffer, 1869)  C & W Mexico  M T S L Ha G

Cecropterus mexicana ducia (Evans, 1952)  TL: Panama  M OD

Cecropterus floridianus Grishin, 2023  Florida Cloudywing  ref.146  M S L

Cecropterus pylades (Scudder, 1870)  Northern Cloudywing  Canada, E USA  iN  M T S1 S2 L1 L2 L3 L4 L5 L6 L7 L8 L9 L10 L11 L12 I G

Cecropterus indistinctus (Austin & J. Emmel, 1998)  Indistinct Cloudywing  Baja California (N) to Washington, Cascades (& Ochocos) west  M T S L1 L2 Ha

Cecropterus rockiensis Grishin, 2023  Rocky Mountains Cloudywing  NE OR, S ID, MT, WY, W SD, W NE, N & C UT, CO, NM, AZ; ref.146  M T S L1 L2

Cecropterus albosuffusa (H. Freeman, 1943)  Whitewashed Cloudywing  W Texas, NE Mexico

Cecropterus albosuffusa albosuffusa (H. Freeman, 1943)  W Texas, NE Mexico  M T S1 S2

Cecropterus albosuffusa southwestern segregate (, )  S AZ, S NM, W Mexico to Oaxaca  M S L1 L2 L3 L4 H

Cecropterus albosuffusa BCS segregate (, )  Baja California Sur  M S

Cecropterus oaxacensis Grishin, 2023  Oaxacan Cloudywing  ref.146  M T

Cecropterus drusius (W. H. Edwards, [1884])  Drusius Cloudywing, White-fringed Cloudywing  SE AZ, SW NM, W TX, MX  iN  M T S L I Ha

Cecropterus cincta Plötz, 1882  Chisos Banded-Skipper  W Texas to Guatemala  iN  M T S L I Ha G

Cecropterus vectilucis (A. Butler, 1872)  Central American Banded-Skipper  Chiapas to Panama  iN  M T OD S L G

Cecropterus pseudocellus (Coolidge & Clémence, [1910])  Sonoran Banded-Skipper  Mexico; extirpated from S AZ  iN  M T S L Ha G

Cecropterus palliolum (H. Druce, 1908)  Ochre-stained Flasher  Costa Rica  M T OI S

Cecropterus egregius (A. Butler, 1870)  Small-spotted Flasher    iN

Cecropterus egregius egregius (A. Butler, 1870)  S TX (rare stray) to Colombia and Venezuela  M T S L Ha G

Cecropterus egregius coxeyi (R. Williams, 1931)  TL: Ecuador  M T

Cecropterus virescens (Mabille, 1877)  Green-washed Longtail  Panama to S Brazil and Paraguay  iN  M T S IL L G

Cecropterus viridissimus Grishin, 2023  Ecuador; ref.153

Cecropterus dorantes (Stoll, 1790)  Dorantes Longtail    iN

Cecropterus dorantes calafia (R. Williams, 1926)  Baja California Sur  M T S L

Cecropterus dorantes dorantes (Stoll, 1790)  continental; Florida, S AZ, S NM, TX to Argentina  M T S HS IL L1 L2 L3 I1 I2 H G

Cecropterus dorantes cramptoni (W. Comstock, 1944)  Hispaniola, Puerto Rico, Mona, Culebra, Virgin Is., Gr. Cayman  M T S

Cecropterus dorantes santiago (Lucas, 1857)  Cuba, N Andros, Mangrove Cay, Grand Bahama  M T S IL

Cecropterus dorantes galapagensis (F. Williams, 1911)  TL: Galapagos  M T

Cecropterus obscurus (Hewitson, 1867)  Dark Longtail  Lesser Antilles; St. Martin S to Grenadines, Barbados  iN  M T S L

Cecropterus markwalkeri Grishin, 2023  Walker's Cloudywing  SE AZ as stray, W Mexico; ref.20  M T S L I

Cecropterus albociliatus (Mabille, 1877)  White-edged Cloudywing    iN

Cecropterus albociliatus albociliatus (Mabille, 1877)  [= toxeus] S TX (?) as stray, Mexico to Costa Rica; ref.20  M T S1 S2 L I G

Cecropterus albociliatus leucophrys (Mabille, 1898)  Panama and Colombia  M T S

Cecropterus albociliatus nocera (Plötz, 1882)  TL: Colombia  M T OI

Cecropterus coyote (Skinner, 1892)  Coyote Cloudywing  [former toxeus] S Texas & E Mexico (at least); ref.20  iN  M S L G

Cecropterus nigrociliata (Mabille & Boullet, 1912)  Dark-edged Cloudywing  [former toxeus] SW Mexico to Panama; ref.20  M T S1 S2 I

Cecropterus jalapus (Plötz, 1881)  Jalapus Cloudywing  S Texas (rare stray) to Colombia  iN  M T S1 S2 L I1 I2 G

Cecropterus athesis (Hewitson, 1867)  Athesis Cloudywing  Panama, Colombia, Venezuela to Tobago  iN  M T S IL L G

Cecropterus phalaecus (Godman & Salvin, 1893)  Yellow-edged Flasher  E & W Mexico to Colombia  iN  M T S L G

Cecropterus reductus (N. Riley, 1919)    iN  M T S

Cecropterus doryssus (Swainson, 1831)  White-tailed Longtail    iN

Cecropterus doryssus chales (Godman & Salvin, 1893)  W Mexico  M T S IL L G

Cecropterus doryssus E Mexican segregate (, )  ["d. doryssus"], S Texas to El Salvador  M S L1 L2

Cecropterus doryssus doryssus (Swainson, 1831)  Chiapas to S Brazil & Bolivia  M T OI OD S IL L1 L2 I1 I2

Cecropterus doryssus albicuspis (Herrich-Schäffer, 1869)  TL: ?  M T IL

Cecropterus albimargo (Mabille, 1875)  White-edged Longtail  NE Mexico to Venezuela and Ecuador  iN  M T S1 S2 IL L I

Cecropterus takuta (Evans, 1952)  Amazonian; ref.7  iN  M T S

Cecropterus rica (Evans, 1952)  SE S. Amer.; ref.7  iN  M T

Cecropterus dariensis Grishin, 2023  Panama; ref.153

Cecropterus trebia (Möschler, 1879)  White-capped Longtail  El Salvador to Colombia and Trinidad  M T S IL G

Cecropterus carmelita (Herrich-Schäffer, 1869)  Carmelita Longtail  Panama to Brazil  iN  M T S IL

Cecropterus barra (Evans, 1952)  Brazil (Ba); ref.7  M T OD G

Spicauda  Grishin, 2019  

Spicauda cindra (Evans, 1952)    iN  M T S

Spicauda zagorus (Plötz, 1880)    iN  M T IL

Spicauda atelis Grishin, 2023  Atelis Longtail  S Texas to Costa Rica; ref.19  M S1 S2 L1 L2 I G

Spicauda teleus (Hübner, 1821)  Teleus Longtail  Panama to Argentina; ref.19  iN  M T S Ab IL L G

Spicauda zalanthus (Plötz, 1881)  S Brazil; ref.19  M T

Spicauda ambiguus (de Jong, 1983)    iN  M T

Spicauda tanna (Evans, 1952)  Tanna Longtail  S Texas (as stray) to Ecuador, Guianas, Trinidad  iN  M T S L G

Spicauda procne (Plötz, 1881)  Brown Longtail  S Texas, SE AZ/SW NM (as stray), to Argentina; ref.7  iN  M S L1 L2 L3 I G

Spicauda simplicius (Stoll, 1790)  Plain Longtail  S Texas (as stray) to Argentina  iN  M T S IL L1 L2 I1 I2 G

Urbanus  Hübner, [1807]  

Urbanus esmeraldus (A. Butler, 1877)  Esmeralda Longtail  (S TX as stray), E & W Mexico to Peru & Guianas, Trinidad  iN  M T S1 S2 IL L I G

Urbanus esma Evans, 1952  Esma Longtail  Panama to Brazil, Trinidad  iN  M T S

Urbanus prodicus E. Bell, 1956  Montane Longtail  E & W Mexico to Costa Rica; often in cloud forest  iN  M T S L

Urbanus elmina Evans, 1952  Andean Longtail  (Costa Rica?), Colombia, Ecuador  iN  M T S

Urbanus evona Evans, 1952  Turquoise Longtail  E & W Mexico to Colombia  iN  M T S1 S2 L I G

Urbanus esta Evans, 1952  Esta Longtail  E & W Mexico to Ecuador and Brazil  iN  M T S1 S2 L I

Urbanus viridis H. Freeman, 1970  Rare Longtail  SE Mexico (VER)  M T

Urbanus velinus (Plötz, 1881)  Velinus Skipper  (SE Mexico?). Guatemala to Brazil and Trinidad  iN  M T S IL

Urbanus proteus (Linnaeus, 1758)  Long-tailed Skipper    iN

Urbanus proteus proteus (Linnaeus, 1758)  continental; E USA, S CA, S AZ, S NM, TX to Argentina  M S1 S2 HS IL L1 L2 I1 I2 I3 H G

Urbanus proteus domingo (Scudder, 1872)  Bahamas and Antilles; ref.151  M T S

Urbanus cubanus Grishin, 2023  Cuban Longtail  W Cuba; ref.151  M T

Urbanus magnus Steinhauser, 1981  TL: Ecuador  M T

Urbanus rickardi Grishin, 2023  Rickard's Longtail  S TX to possibly Panama; ref.19  M T S L

Urbanus pronus Evans, 1952  Pronus Longtail  Colombia to Argentina  iN  M T S

Urbanus pronta Evans, 1952  Spot-banded Longtail  E & W Mexico to Argentina  iN  M T S1 S2 L I1 I2

Urbanus parvus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus longicaudus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus villus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus huancavillcas (R. Williams, 1926)  TL: Ecuador  M T S

Urbanus belli (Hayward, 1935)  Bell's Longtail  Argentina  iN  M TI IL G

Urbanus alva Evans, 1952  Alva Longtail  [= ehakernae] Mexico to Costa Rica; ref.82,10,19  iN  M T S1 S2 L I1 I2 G

Urbanus oplerorum Grishin, 2023  the Oplers' Longtail  S TX & MX; ref.19  M T

Urbanus bernikerni Burns, 2014  Costa Rica; ref.82  iN

Urbanus segnestami Burns, 2014  Costa Rica; ref.82  iN

Urbanus viterboana (Ehrmann, 1907)  Bluish Longtail  E & W Mexico to Colombia and Ecuador  iN  M T S1 S2 L I G

Urbanus dubius Steinhauser, 1981    iN  M T

Urbanus megalurus (Mabille, 1877)  Long-tailed Flasher  E & W Mexico to Colombia; ref.7  iN  M T S L G

Urbanus tucuti (R. Williams, 1927)  Tucuti Flasher  SE Mexico to Ecuador; ref.7  iN  M T OI S L

Urbanus mericuti Grishin, 2023  S America to Peru & Bolivia; ref.153

Telegonus  Hübner, [1819]  

Telegonus brevicauda (Plötz, 1886)  Short-tailed Flasher  [= talthybius], SE Mexico to Costa Rica  iN  M T S1 S2 L I

Telegonus chalco (Hübner, 1823)  Great White Longtail  Costa Rica to S Brazil and Paraguay  iN  M S IL L G

Telegonus tsongae Grishin, 2023  Qian's Flasher  S TX & MX; ref.19  M T L1 L2 I H

Telegonus misitra (Plötz, 1881)  Mexico; ref.19  M T

Telegonus catemacoensis (H. Freeman, 1967)  SE Mexico; ref.7  iN  M T Txt

Telegonus azul (Reakirt, [1867])  Two-barred Flasher, Flashing Astraptes  This is complex of continental species-level taxa; ref.7,19  iN  M T OD S L

Telegonus fulgerator (Walch, 1775)  S America; ref.18  iN  M T OI IL G

Telegonus fulminator (Sepp, [1841])  Suriname; ref.19  M T

Telegonus fulgor Hayward, 1939    iN  M TI S IL

Telegonus naxos (Hewitson, 1867)    iN  M T S

Telegonus christyi Sharpe, 1898  Antillean Flasher  Hispaniola; ref.78  M T S

Telegonus xagua (Lucas, 1857)  Antillean Flasher    iN

Telegonus xagua harveyi (Clench, Steinhauser & J. Miller, 2017)  Bahamas; ref.78

Telegonus xagua xagua (Lucas, 1857)  Cuba, Andros  M T S

Telegonus talus (Cramer, 1777)  Green Flasher  E & S Mexico to Argentina; Jamaica, Cuba, Hispaniola, etc.  iN  M T S L G

Telegonus pastus Grishin, 2023  SE Mexico to W Colombia; ref.153

Telegonus apastus (Cramer, 1777)  Broad-banded Flasher    iN

Telegonus apastus apastus (Cramer, 1777)  E Colombia to Guianas and N Brazil  M S L I G

Telegonus apastus pusa (Evans, 1952)  TL: Brazil (Ba)  M T

Telegonus elorus (Hewitson, 1867)    iN  M T S IL L

Telegonus alardus (Stoll, 1790)  Frosted Flasher    iN

Telegonus alardus latia (Evans, 1952)  S TX (rare stray), E Mexico to Colombia  M T S1 S2 L I Ha

Telegonus alardus aquila (Evans, 1952)  TL: Colombia  M T

Telegonus alardus alardus (Stoll, 1790)  TL: Surinam  M T OD S IL L G

Telegonus habana (Lucas, 1857)  Frosty Flasher  Cuba, Isla de Juventud  iN  M S

Telegonus heriul Mabille & Boullet, 1912  Frosty Flasher  Hispaniola; ref.7  M T S

Telegonus alector (C. Felder & R. Felder, 1867)    iN

Telegonus alector hopfferi (Plötz, 1881)  [= gilberti], S TX (stray), E & W Mexico to NW Brazil and Bolivia  M T S L

Telegonus alector alector (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Telegonus cretatus Hayward, 1939  Cretatus Flasher    iN

Telegonus cretatus cretatus Hayward, 1939  [= alfius], Nicaragua, Colombia to Bolivia  M T G

Telegonus cretatus adoba (Evans, 1952)  TL: Brazil (ES)  M T

Telegonus creteus (Cramer, 1780)  Whitened Flasher    iN

Telegonus creteus crana (Evans, 1952)  SE Mexico to Panama  M T S

Telegonus creteus crilla (Evans, 1952)  TL: Ecuador  M T

Telegonus creteus cyprus (Evans, 1952)  TL: Bolivia  M T

Telegonus creteus creteus (Cramer, 1780)  Panama to Guianas and Bolivia  M S IL G

Telegonus creteus siges Mabille, 1903  TL: Brazil  M T

Telegonus bifascia (Herrich-Schäffer, 1869)  Green-headed Flasher  SE Mexico to Panama  M T S

Telegonus latimargo (Herrich-Schäffer, 1869)  White-edged Flasher  Panama, Colombia, Ecuador  iN  M T OI S

Telegonus tinda (Evans, 1952)  TL: Brazil (Pa)  M T

Telegonus chiriquensis Staudinger, 1876  Chiriqui Flasher    iN

Telegonus chiriquensis chiriquensis Staudinger, 1876  E Mexico to Venezuela and Ecuador  M T S IL L

Telegonus chiriquensis erana (Evans, 1952)  TL: Ecuador  M T

Telegonus chiriquensis meretrix (Hewitson, 1876)  TL: Ecuador  M T

Telegonus chiriquensis oenander (Hewitson, 1876)  TL: Brazil (Pa)  M IL G

Telegonus weymeri (Plötz, 1882)  Weymer’s Flasher    iN  M S HS

Telegonus cretellus (Herrich-Schäffer, 1869)  Jamaican Flasher  [= jaira] Jamaica; ref.13  iN  M T S L

Telegonus cassander (Fabricius, 1793)  Cuban Flasher  Cuba and Isla de Juventud  iN  M T S G

Telegonus galesus Mabille, 1888  Peru; ref.13  iN  M T OI

Telegonus subflavus Grishin, 2022  Ecuador; ref.13  M T

Telegonus cassius (Evans, 1952)  Dull Flasher  Costa Rica to Venezuela; ref.7,13  M T S L G

Telegonus anausis Godman & Salvin, 1896  'Caribbean' Yellow-tipped Flasher  ref.11  iN

Telegonus anausis annetta (Evans, 1952)  S Texas to Guiana and Peru  M T S1 S2 L1 L2 I G

Telegonus anausis anoma (Evans, 1952)  TL: Trinidad  M T

Telegonus anausis aniza (Evans, 1952)  TL: Peru  M T

Telegonus anausis anausis Godman & Salvin, 1896  Greater Antilles, Virgin Is., Montserrat to Grenada  M T S

Telegonus anaphus (Cramer, 1777)  ref.11  iN  M T IL

Telegonus hyster (Dyar, 1916)  Hyster Skipper  Mexico, El Salvador  iN  M T S L G

Telegonus cellus (Boisduval & Le Conte, [1837])  Golden Banded-Skipper  E US to SE AZ, S to El Salv.  iN  M T OI OD S1 S2 S3 S4 IL L1 L2 L3 I Ha G Txt

Telegonus siermadror (Burns, 1984)  East-Mexican Banded-Skipper  NE Mexico; cloud forest  iN  M T L G

Autochton  Hübner, 1823  

Autochton sulfureolus (Mabille, 1883)    iN  M S G

Autochton reflexus (Mabille & Boullet, 1912)  TL: Brazil (SC)  M T S

Autochton neis (Geyer, 1832)  Broad Banded-Skipper  E & W Mexico to S Brazil and Paraguay  iN  M T S Ab L G

Autochton dora Grishin, 2023  Ecuador; ref.153

Autochton integrifascia (Mabille, 1891)    iN  M S L

Autochton itylus Hübner, 1823    iN  M OI S L

Autochton bipunctatus (Gmelin, [1790])  Gmelin's Banded-Skipper  SE Mexico to Bolivia and Brazil  iN  M T S L I

Autochton potrillo (Lucas, 1857)  Potrillo Skipper  Jamaica, Cuba, Hispaniola  iN  M T G

Autochton caballo Grishin, 2023  Caballo Skipper  SE AZ (stray), S TX to Costa Rica; ref.19  M S1 S2 L1 L2 I G

Autochton reducta (Mabille & Boullet, 1919)  Reduced-saddle Skipper  Panama, Colombia, Venezuela, Trinidad; ref.7,19  M S

Autochton oryx (C. Felder & R. Felder, 1862)  ref.7  iN  M T OI S

Spathilepia  A. Butler, 1870  

Spathilepia clonius (Cramer, 1775)  Falcate Skipper  S TX (sporadic) to N Argentina  iN  M T S1 S2 IL L I G

Astraptes  Hübner, [1819]  

Astraptes erycina (Plötz, 1881)  Brazil  iN  M S IL G

Astraptes mabillei Steinhauser, 1989  TL: Bolivia  M T S

Astraptes halesius (Hewitson, 1877)  Amazonian  iN  M T S

Astraptes centralis Grishin, 2023  Nicaragua to Panama; ref.153

Astraptes aulus (Plötz, 1881)  Green-tinged Flasher  in S. Am. to Peru and Argentina  iN  M T OI S G

Astraptes enotrus (Stoll, 1781)  White-spotted Flasher  E & W Mexico to S Brazil, Paraguay, Bolivia  iN  M T S IL L G

Astraptes janeira (Schaus, 1902)  Schaus' Flasher    iN  M T S L

Narcosius  Steinhauser, 1986  

Narcosius colossus (Herrich-Schäffer, 1869)  Colossal Flasher  SE Mexico to Colombia and Venezuela  iN  M T S1 S2 I1 I2 G

Narcosius granadensis (Möschler, 1879)  Venezuela; ref.7  M T S G

Narcosius dosula (Evans, 1952)    iN  M T

Narcosius hercules (E. Bell, 1956)  TL: Bolivia  M T

Narcosius steinhauseri Austin, 1996  TL: Brazil (Rd)  M T OI G

Narcosius narcosius (Stoll, 1790)    iN  M T S G

Narcosius aulina (Evans, 1952)  TL: French Guiana  M T

Narcosius helen (Evans, 1952)  Steely Flasher  E & W Mexico to Panama; ref.7  M T S I G

Narcosius parisi (R. Williams, 1927)    iN  M T IL G

Narcosius odysseus Austin, 1996  TL: Brazil (Rd)  M T OI G Txt

Narcosius mura (R. Williams, 1927)  TL: Brazil (Am)  M T S IL G Txt

Narcosius pseudomura Austin, 1996  TL: Brazil (Rd)  M T OI G Txt

Narcosius nazaraeus Steinhauser, 1986  Nazareus Flasher  SE Mexico to Guianas, Brazil, Peru, Ecuador and Bolivia  iN  M T S L G

Narcosius samson (Evans, 1952)  Samson Flasher  SE Mexico to Brazil and Bolivia  iN  M T S1 S2 I1 I2 G

Proteides  Hübner, [1819]  

Proteides mercurius (Fabricius, 1787)  Mercurial Skipper    iN

Proteides mercurius mercurius (Fabricius, 1787)  continental, S Texas to Argentina, SE Arizona as rare stray  M OI S1 S2 IL L I1 I2 G

Proteides mercurius sanantonio (Lucas, 1857)  Cuba, [one-time stray to Florida]  M S L

Proteides mercurius sanchesi E. Bell & W. Comstock, 1948  Hispaniola  M T S

Proteides mercurius jamaicensis Skinner, 1920  Jamaica  M T S

Proteides mercurius pedro (Dewitz, 1877)  Puerto Rico  M S

Proteides mercurius vincenti E. Bell & W. Comstock, 1948  St. Vincent  M T S

Proteides mercurius angasi Godman & Salvin, 1884  Dominica, St. Lucia, Tobago  M T S

Proteides mercurius grenadensis Enrico & Pinchon, 1969  Grenada  M S

Proteides maysi (Lucas, 1857)  Mays' Skipper  Cuba and Isla de Juventud  iN  M T S

Epargyreus  Hübner, [1819]  

Epargyreus zestos (Geyer, 1832)  Zestos Skipper    iN

Epargyreus zestos zestos (Geyer, 1832)  Florida, Bahamas, Puerto Rico, Virgin Islands to St. Vincent  M T S L G

Epargyreus zestos inaguarum Clench & Bjorndal, 1980  Great Inagua, Crooked I., Turks and Caicos Islands  M T S

Epargyreus clarus (Cramer, 1775)  Silver-spotted Skipper  ref.11,16  iN

Epargyreus clarus californicus MacNeill, 1975  BC to Baja California Norte; Sierra Nevada-Cascades, westward  M S L I H

Epargyreus clarus Rocky Mountain segregate ,  NE OR, E WA, ID, MT, WY, C & N UT, CO, N NM, Black Hills  M S

Epargyreus clarus clarus (Cramer, 1775)  E USA and Canada to far NE Mexico (hypothetical)  M S HS L1 L2 L3 L4 L5 I H G

Epargyreus huachuca Dixon, 1955  'Arizona' Silver-spotted Skipper  ref.11,16

Epargyreus huachuca profugus Austin, 1998  S NV, N AZ, possibly to central AZ-NM  M T L

Epargyreus huachuca huachuca Dixon, 1955  S AZ, S NM, far NW Mexico  M T S L I H

Epargyreus antaeus (Hewitson, 1867)  Jamaican Silverdrop  Jamaica  iN  M T S L G

Epargyreus spanna Evans, 1952  Hispaniolan Silverdrop  Hispaniola  iN  M T S G

Epargyreus orizaba Scudder, 1872  Round-spotted Silverdrop  E Mexico to Guatemala; ref.7  M T S L G

Epargyreus dicta Evans, 1952  Bolivia; ref.7  iN  M T

Epargyreus socus (Hübner, [1825])    iN

Epargyreus socus sinus Evans, 1952  TL: Brazil (Pa)  M T S

Epargyreus socus socus (Hübner, [1825])  TL: ?  M G

Epargyreus pseudexadeus Westwood, 1852  Brazil; ref.7  M T

Epargyreus windi H. Freeman, 1969  Wind’s Silverdrop  W Mexico  iN  M T S L G

Epargyreus fractigutta Grishin, 2023  Broken Silverdrop  S TX [CA? as stray] to Panama; ref.19  M T S IL L

Epargyreus cruza Evans, 1952  Bruised Silverdrop  E & W Mexico to Panama; ref.7,19  iN  M T L G

Epargyreus exadeus (Cramer, 1779)    iN  M S IL G

Epargyreus nutra Evans, 1952    iN  M T G

Epargyreus aspina Evans, 1952  Spineless Silverdrop  E & W Mexico to Venezuela and Ecuador  iN  M T L G

Epargyreus spina Evans, 1952  Spined Silverdrop    iN

Epargyreus spina spina Evans, 1952  E & W Mexico to Brazil and Bolivia  M T S G

Epargyreus spina verruga Evans, 1952  TL: Peru  M T

Epargyreus spinta Evans, 1952    iN  M T S

Epargyreus spinosa Evans, 1952  Suffused Silverdrop  W Mexico to Panama  iN  M T S G

Epargyreus clavicornis (Herrich-Schäffer, 1869)  [= chota]; ref.19  iN  M T IL G

Epargyreus gaumeri Godman & Salvin, 1893  Small-spotted Silverdrop  SE Mexico (?) to Honduras; ref.19

Epargyreus gaumeri gaumeri Godman & Salvin, 1893  SE Mexico (?) to Honduras; ref.19  M T OI G

Epargyreus gaumeri tenda Evans, 1955  SE Mexico to Venezuela and Ecuador; ref.19  M T S L G

Epargyreus brodkorbi H. Freeman, 1969  Barely-spotted Silverdrop  Chiapas  iN  M T S L G

Epargyreus deleoni H. Freeman, 1977  Long-spotted Silverdrop  Yucatan Peninsula (Camp, QRoo, Yuc Mexico) and Belize  iN  M T S G

Epargyreus tmolis (Burmeister, 1875)    iN  M S IL G

Epargyreus barisses (Hewitson, 1874)    iN

Epargyreus barisses barisses (Hewitson, 1874)  TL: Bolivia  M T S IL G

Epargyreus barisses argentina Mabille, 1903  TL: ?  M T S G

Epargyreus enispe (Hewitson, 1867)  TL: S. Amer.

Epargyreus enispe enispe (Hewitson, 1867)  TL: S. Amer.  M T S IL G

Epargyreus enispe elta Evans, 1952  TL: Bolivia  M T

Chioides  Lindsey, 1921  

Chioides albofasciatus (Hewitson, 1867)  White-striped Longtail  S TX, S NM, SE AZ, to Costa Rica; Baja California Sur  iN  M T S IL L1 L2 L3 I H G

Chioides catillus (Cramer, 1779)    iN

Chioides catillus albius Evans, 1952  Costa Rica and Panama  M T S1 S2 L I G

Chioides catillus catillus (Cramer, 1779)  TL: Surinam  M S1 S2 IL L

Chioides catillus jethira (A. Butler, 1870)  TL: Peru  M T S IL G

Chioides churchi E. Bell & W. Comstock, 1948  Jamaican Longtail  Jamaica  iN  M T S

Chioides cinereus (Mabille & Vuillot, 1891)  TL: Brazil  M T S IL

Chioides concinnus (Mabille, 1877)  Peru  iN  M T OI S G

Chioides vintra Evans, 1952  St. Vincent Longtail  St. Vincent, Grenadines, Grenada  M T S

Chioides iverna Evans, 1952    iN  M T S

Chioides zilpa (A. Butler, 1872)  Zilpa Longtail  S AZ-NM (stray) & S TX, S to Venezuela and Ecuador  iN  M T OD S1 S2 IL L I1 I2 G

Chioides ixion (Plötz, 1880)  Hispaniolan Longtail  Hispaniola  iN  M T S

Chioides marmorosa (Herrich-Schäffer, 1865)  Cuban Longtail  Cuba  M T S

  Tribe Eudamini, Subtribe Loboclina

Aguna  R. Williams, 1927  

Aguna malia Evans, 1952  Venezuela; ref.20  M T G

Aguna megaeles (Mabille, 1888)    iN  M T OI S IL

Aguna mcguirei Grishin, 2023  McGuire's Aguna  S TX to Panama; ref.19  M S L I

Aguna metophis (Latreille, [1824])  Long-tailed Aguna  C Am. to Brazil; ref.19  iN  M T IL

Aguna camagura (R. Williams, 1926)    iN  M T S

Aguna asander (Hewitson, 1867)  Gold-spotted Aguna    iN

Aguna asander asander (Hewitson, 1867)  SE AZ & S TX (as stray) to Argentina  M T S L I G

Aguna asander haitensis (Mabille & Boullet, 1912)  Cuba and Hispaniola  M T S

Aguna asander jasper Evans, 1952  Jamaica  M T S L

Aguna albistria (Plötz, 1880)    iN

Aguna albistria albistria (Plötz, 1880)  TL: Brazil ([Pa])  M T TI S IL L

Aguna albistria leucogramma (Mabille, 1888)  SE Mexico to Colombia and Venezuela (Surinam?)  M T OI S

Aguna claxon Evans, 1952  Emerald Aguna  Cuba, S Texas to Colombia and Surinam  iN  M T S L I G

Aguna claxonica Grishin, 2023  Ecuador; ref.153

Aguna prasinus Siewert, Leviski, O. Mielke & Casagrande, 2015  Panama; ref.32  M OI OD

Aguna clina Evans, 1952    iN  M T S

Aguna latifascia Austin & O. Mielke, 1998    iN  M T S

Aguna coelus (Stoll, 1781)  Stoll's Aguna  Costa Rica and Panama to Peru and central Brazil  iN  M T S

Aguna cirrus Evans, 1952  TL: Brazil (SP)  M T S G

Aguna coeloides Austin & O. Mielke, 1998  Austin's Aguna  SE Mexico to southern Brazil  iN  M T S1 S2 HS I1 I2

Aguna nicolayi Austin & O. Mielke, 1998    iN  M T S

Aguna venezuelae O. Mielke, 1971  TL: Venezuela  M T S

Aguna latimacula Austin & O. Mielke, 1998  TL: Brazil (MG)  M T

Aguna aurunce (Hewitson, 1867)    iN

Aguna aurunce aurunce (Hewitson, 1867)  TL: Brazil (Am)  M T S IL

Aguna aurunce hypozonius (Plötz, 1880)  SE Mexico to Venezuela  M S L I

Aguna glaphyrus (Mabille, 1888)    iN  M T S

Aguna esmeralda Grishin, 2023  Ecuador; ref.153

Aguna penicillata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna spicata Austin & O. Mielke, 1998    iN  M T S

Aguna longicauda Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T OI

Aguna ganna (Möschler, 1879)  Ganna Aguna  Costa Rica to Colombia and Venezuela  iN  M T S IL

Aguna panama Austin & O. Mielke, 1998  Panamanian Aguna  Honduras to Venezuela  iN  M T1 T2 S I

Aguna spatulata Austin & O. Mielke, 1998    iN  M T S

Aguna similis Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna lata Grishin, 2023  Guyana; ref.153

Aguna mesodentata Austin & O. Mielke, 1998    iN  M T S

Aguna squamalba Austin & O. Mielke, 1998    iN  M T S

Aguna parva Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Zeutus  Grishin, 2019  

Zeutus zeutus (Möschler, 1879)  Zeutus Banded-Skipper  SE Mexico to Guianas and Bolivia; ref.7  iN  M T S1 S2 L I G

Lobocla  Moore, 1884  the only Old World genus of Eudaminae

Lobocla liliana (Atkinson, 1871)  Asian species, shown here for comparison  iN

Lobocla liliana ignatius (Plötz, 1882)  Asian species, shown here for comparison  M T S

Lobocla liliana liliana (Atkinson, 1871)  Asian species, shown here for comparison  M S G

Lobocla aborica (Tytler, 1915)  Asian species, shown here for comparison

Lobocla aborica aborica (Tytler, 1915)  Asian species, shown here for comparison  M T G

Lobocla aborica zesta Evans, 1952  Asian species, shown here for comparison  M T

Lobocla aborica tonka Evans, 1952  Asian species, shown here for comparison  M T

Lobocla quadripunctata Fan & Wang, 2004  Asian species, shown here for comparison  M T S G

Lobocla disparalis Murayama, 1995  Asian species, shown here for comparison  M OI OD

Lobocla bifasciatus (Bremer & Grey, 1853)  Asian species, shown here for comparison  iN  M T S G

Lobocla contractus (Leech, 1894)  Asian species, shown here for comparison  M T S G

Lobocla germanus (Oberthür, 1886)  Asian species, shown here for comparison  iN  M T S G

Lobocla nepos (Oberthür, 1886)  Asian species, shown here for comparison  iN

Lobocla nepos nepos (Oberthür, 1886)  Asian species, shown here for comparison  M T S G

Lobocla nepos phyllis (Hemming, 1933)  Asian species, shown here for comparison  M S

Lobocla proximus (Leech, 1891)  Asian species, shown here for comparison  iN  M T S G

Lobocla simplex (Leech, 1891)  Asian species, shown here for comparison  iN  M T S G

Lobotractus  Grishin, 2019  

Lobotractus mysie (Dyar, 1904)  Mysie Skipper  SE AZ & N Son.; ref.113,114,115,12  M T S L I H F Ha

Lobotractus valeriana (Plötz, 1881)  Valeriana Skipper  N. Mexico to Oaxaca; ref.113,114,115,7,12  iN  M T S L G

Lobotractus uvydixa (Dyar, 1914)  Variable Mottled-Skipper  S Mexico: Guerrero to Chiapas  iN

Lobotractus uvydixa dark segregate (, )  W Mexico: Sonora to Michoacan  M S L I

Lobotractus uvydixa uvydixa (Dyar, 1914)  TL: Mexico (Gue)  M T S L Ha

Lobotractus cyda (Godman, 1901)  Cyda Mottled-Skipper  Chiapas to Nicaragua  iN  M T OI S IL

Codatractus  Lindsey, 1921  

Codatractus cyledis (Dyar, 1912)  Cyledis Mottled-Skipper  S Mexico  iN  M T S L Ha G

Codatractus bryaxis (Hewitson, 1867)  Golden Mottled-Skipper  N Mexico to Honduras  iN  M T S IL L G

Codatractus imalena (A. Butler, 1872)  Bluish Mottled-Skipper  Costa Rica to northern Brazil  iN  M T OI S1 S2 IL L I1 I2

Codatractus arizonensis (Skinner, 1905)  Arizona Mottled-Skipper  S AZ, S NM, W TX to Oaxaca  iN  M T S L1 L2 I H G

Codatractus sallyae A. Warren, 1995  Sally’s Mottled-Skipper  W Mexico, Yucatan Peninsula, Guatemala; ref.22  iN  M T S L

Codatractus melon (Godman & Salvin, 1893)  Melon Mottled-Skipper  N Mexico to Costa Rica  iN  M T S1 S2 IL L I

Codatractus carlos Evans, 1952  Carlos' Mottled-Skipper  C Mexico to Colombia (at least)  iN  M T S L G

Codatractus rowena Evans, 1952  Venezuela; ref.7  iN

Codatractus rowena rowena Evans, 1952  TL: Venezuela  M T G

Codatractus rowena arguta Evans, 1952  TL: Trinidad  M T

Codatractus alcaeus (Hewitson, 1867)  White-crescent Mottled-Skipper  S Texas to Costa Rica  iN  M T S IL L I G

Codatractus apulia Evans, 1952  White-crescent Mottled-Skipper  Panama (?) to W Ecuador; ref.7  M T S G

Codatractus yucatanus H. Freeman, 1977  Yucatan Mottled-Skipper  Yucatan Peninsula  iN  M T

Codatractus aminias (Hewitson, 1867)    iN  M T IL L

Zestusa  Lindsey, 1925  

Zestusa dorus (W. H. Edwards, 1882)  Short-tailed Skipper (Northern Zestusa)  far S CO-UT, AZ, NM, W TX, N Mexico  iN  M T S L1 L2 L3 I Ha G

Zestusa elwesi (Godman & Salvin, 1893)  Mexican Zestusa  Durango to Oaxaca  iN  M T S L G

Zestusa staudingeri (Mabille, 1888)  Southern Zestusa  Chiapas to El Salvador  iN  M T OI S L

Zestusa levona Steinhauser, 1972  Levona's Zestusa  Chiapas to El Salvador  iN  M T

Ridens  Evans, 1952  

Ridens harpagus (C. Felder & R. Felder, 1867)    iN  M T S IL

Ridens philistus (Hopffer, 1874)    iN

Ridens philistus philistus (Hopffer, 1874)  TL: Peru  M T S

Ridens philistus philia Evans, 1952  TL: Colombia  M T S

Ridens allyni H. Freeman, 1979  Allyn’s Ridens  SE Mexico, Guatemala  iN  M T1 T2 G

Ridens mephitis (Hewitson, 1876)  Hewitson's Ridens  SE Mexico to Peru and Bolivia  iN  M T S1 S2 I1 I2

Ridens fulminans (Herrich-Schäffer, 1869)    iN  M T S

Ridens angulinea Grishin, 2023  S Peru & Bolivia; ref.153  M T

Ridens fulima Evans, 1952  S Brazil  iN  M T S L

Ridens pacasa (R. Williams, 1927)  Peru & Bolivia  iN  M T S

Ridens bidens Austin, 1998  TL: Brazil (Rd)  M OI

Ridens conservationinternationalis Burns & Grishin, 2023  TL: Costa Rica

Ridens bridgmani (Weeks, 1902)  TL: Bolivia  M T S

Ridens miltas (Godman & Salvin, 1893)  Mexican Ridens  E Mexico  M T S IL G

Ridens mercedes Steinhauser, 1983  White-tailed Ridens  SW Mexico  M T S G

Ridens crison (Godman & Salvin, 1893)  Many-spotted Ridens  SE Mexico, Guatemala  iN  M T S IL G

Ridens howarthi Steinhauser, 1974  Many-spotted Ridens  El Salvador  M T G

Ridens cachinnans (Godman, 1901)  Many-spotted Ridens  Costa Rica, Panama  M T OI S IL I G

Ridens biolleyi (Mabille, 1900)  Biolley's Ridens  Costa Rica, Panama  M T S1 S2 I1 I2 I3

Ridens ridens (Hewitson, 1876)  Frosted Skipper  Costa Rica & Panama  M T S G

Ridens fieldi Steinhauser, 1974  Field's Ridens  Guatemala (Volcan Sta. Maria)  M T G

Ridens toddi Steinhauser, 1974  Todd’s Skipper  S Mexico (CHIS) to El Salvador  M T S G

Ridens panche (R. Williams, 1927)  Panche Ridens  Costa Rica to Colombia and Ecuador  iN  M T S1 S2 I1 I2

Ridens tristis (Draudt, [1922])    iN  M T S G

Ridens telegonoides (Mabille & Boullet, 1912)  TL: Colombia  M T OI

Ridens nora Evans, 1952  Colombia, Ecuador & Peru  iN  M T S

Venada  Evans, 1952  

Venada [n. sp.] N/A, N/A  Cryptic Scarlet-eye  ["advena"], Mexico, Guatemala, (Nicaragua?)  M S

Venada nevada Burns, 2005  Nevada Scarlet-eye  Costa Rica  M T OD S1 S2 I1 I2 G

Venada advena (Mabille, 1889)  Advena Scarlet-eye  Panama and Colombia, at least  iN  M T OI

Venada daneva Burns, 2005  Daneva Scarlet-eye  Costa Rica  iN  M T OD S1 S2 L I1 I2 G

Venada lamella Burns, 2013  Lamella Scarlet-eye  Mexico, Guatemala, (Nicaragua?), Costa Rica, Panama  M T S1 S2 G

Venada cacao Burns, 2005  Cacao Scarlet-eye  Costa Rica and Panama  iN  M T OD S I G

Venada naranja Burns, 2005  Orange Scarlet-eye  Costa Rica  iN  M T OD S1 S2 I1 I2 G

  Tribe Eudamini, Subtribe Cephisina

Cephise  Evans, 1952  

Cephise impunctus Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise cephise (Herrich-Schäffer, 1869)  TL: ?  M T S IL

Cephise nuspesez Burns, 1996  Burns' Scarlet-eye  E Mexico to Costa Rica  M T S1 S2 I1 I2

Cephise malesedis Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise maculatus Austin & O. Mielke, 2000    iN  M T

Cephise burnsi Austin & O. Mielke, 2000  TL: Brazil (ES)  M T OI

Cephise mexicanus Austin & O. Mielke, 2000  Mexican Scarlet-eye  SLP, OAX (Mexico)  M T

Cephise glarus (Mabille, 1888)  TL: Brazil (Pa)  M T OI S

Cephise aelius (Plötz, 1881)  Longtailed Scarlet-eye  E & S Mexico to Venezuela and Colombia  iN  M T S1 S2 IL L I1 I2 G

Cephise callias (Mabille, 1888)  TL: [Venezuela]  M T OI S IL

Cephise procerus (Plötz, 1880)  TL: Brazil (Pa)  M T IL

Cephise guatemalaensis (H. Freeman, 1977)  Guatemalan Scarlet-eye  Guatemala, Chiapas, Quintana Roo  M T

  Tribe Eudamini, Subtribe Telemiadina

Ectomis  Mabille, 1878  phenotypicaly diverse but genetically compact genus; ref.7

Ectomis gyges (Evans, 1952)    iN  M T S

Ectomis hirtius (A. Butler, 1870)    iN  M T IL

Ectomis roma (Evans, 1952)    iN  M T

Ectomis asine (Hewitson, 1867)  Asine Longtail  N Mexico to Panama  iN  M T S1 S2 IL L I1 I2

Ectomis mexicanus (H. Freeman, 1969)  Mexican Longtail  SE AZ, S TX (as stray), N Mexico to Costa Rica  iN  M T S1 S2 L

Ectomis octomaculata (Sepp, [1844])  Eight-spotted Longtail  (SE AZ as stray), S Texas to Argentina  iN  M T OI S1 S2 IL L I G

Ectomis maizae (Hellebuyck, 1998)  Melanic Longtail  Nicaragua  M T

Ectomis ceculus (Herrich-Schäffer, 1869)    iN  M T IL

Ectomis caunus (Herrich-Schäffer, 1869)  Four-spotted Longtail  E Mexico to S Brazil and Paraguay  iN  M T S1 S2 L I G

Ectomis auginus (Hewitson, 1867)  Auginus Longtail  SE Mexico to S Brazil  iN  M T S IL L I

Ectomis metallescens (Mabille, 1888)  Metallescens Skipper  SE Mexico to Peru and northern Brazil  iN  M T S L G

Ectomis kanshul (Shuey, 1991)  Kanshul Longtail  S Mexico, Belize, NE Costa Rica  iN  M S L I1 I2

Ectomis eudoxus (Stoll, 1781)    iN  M OI IL L

Ectomis minvanes (R. Williams, 1926)  TL: Brazil (MT)  M T G

Ectomis labriaris (A. Butler, 1877)    iN  M T S IL

Ectomis cythna (Hewitson, 1878)    iN

Ectomis cythna cythna (Hewitson, 1878)  TL: ?  M T S L

Ectomis cythna ega Evans, 1953  TL: Brazil (Am)  M T OD

Ectomis speculum (Austin, 2008)  Hidden Mirror Skipper  French Guiana to Brazil  M OI OD S G

Ectomis teutas (Hewitson, 1876)  TL: Brazil (Am)  M T S

Ectomis pervivax (Hübner, [1819])  Pervivax Skipper  Costa Rica to Bolivia, Peru and N Brazil  iN  M T OI S1 S2 HS I

Ectomis bahiana (Herrich-Schäffer, 1869)    iN  M T OI S

Ectomis orphne (Plötz, 1881)  TL: Brazil (RJ)  M T TI S

Ectomis otriades (Hewitson, 1867)  TL: Brazil (Am)  M T S

Ectomis epicincea (A. Butler & H. Druce, 1872)  Stub-tailed Skipper  SE Mexico to Costa Rica  iN  M T S G

Ectomis perniciosus (Herrich-Schäffer, 1869)  Perniciosus Skipper  Panama to Bolivia and C Brazil  iN  M T S L

Ectomis cuminaensis (R.F. d'Almeida, 1976)  TL: Brazil (Pa)  M T

Ectomis orpheus (Plötz, 1881)    iN  M T TI

Ectomis perna (Evans, 1952)  TL: Colombia  M T

Ectomis albovenae (E. Bell, 1932)    iN  M T

Telemiades  Hübner, [1819]  

Telemiades litanicus (Hewitson, 1876)  TL: Brazil (Am)  M T S

Telemiades amphion (Geyer, 1832)    iN  M T S

Telemiades belli Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades fides E. Bell, 1949  Small Telemiades  E & W Mexico to Colombia, Ecuador and upper Amazon  iN  M T S1 S2 L I1 I2

Telemiades marpesus (Hewitson, 1876)    iN  M IL G

Telemiades misitheus Mabille, 1888  TL: Peru  M T OI S L

Telemiades pekahia (Hewitson, 1868)  TL: Venezuela  M T Txt

Telemiades avitus (Stoll, 1781)  Yellow-spotted Telemiades  SE Mexico to N Amazon  iN  M T S L I G

Telemiades xantho Hayward, 1939  TL: Argentina

Telemiades austini Siewert, O. Mielke & Casagrande, 2020  ref.36  iN

Telemiades laogonus (Hewitson, 1876)    iN  M T

Telemiades vansa Evans, 1953  TL: Guyana  M T OD

Telemiades penidas (Hewitson, 1867)  Penidas Telemiades  Costa Rica to S Brazil  iN  M T S IL L

Telemiades pallidus Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades antiope (Plötz, 1882)  Plötz's Telemiades  Nicaragua, Costa Rica, Colombia, S Brazil  iN  M T OI OD S1 S2 L I1 I2

Telemiades atlantiope Siewert, O. Mielke & Casagrande, 2020  ref.36  iN

Telemiades lamasi Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades corbulo (Stoll, 1781)  Corbulo Skipper  Guianas, Ecuador, Peru, Bolivia, upper and lower Amazon  iN  M T S IL L G

Telemiades quammeni Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades insulsus (Draudt, 1922)  TL: Brazil  M T G

Telemiades oiclus (Mabille, 1889)  Oiclus Skipper  [= cora], Honduras to Panama  M T OI S I1 I2

Telemiades choricus (Schaus, 1902)  Mexican Telemiades  W & S Mexico, Belize  iN  M T S L G

Telemiades epicalus Hübner, [1819]  Hübner's Telemiades  Panama to S Brazil  iN  M T S L G

Telemiades sila Evans, 1953  Sila Telemiades  Nicaragua to Colombia, Venezuela and Ecuador  iN  M T S

Telemiades dawkinsi Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades warreni Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades mielkei Siewert & Casagrande, 2020  ref.36

Telemiades nicki Siewert, O. Mielke & Casagrande, 2021  ref.37

Telemiades centrites (Hewitson, 1870)    iN  M T S Txt

Telemiades esmeraldas Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades contra Evans, 1953  Ecuador; ref.7  M T OD

Telemiades gallius (Mabille, 1888)  Centrites Skipper  Costa Rica and Panama, mid-elevation rain forest  iN  M T OI S L I

Telemiades chrysorrhoea (Godman & Salvin, 1893)  Chrysorrhoea Skipper  Costa Rica and Panama, cloud forest  M T S1 S2 L I1 I2

Telemiades megallus Mabille, 1888  Orange Telemiades  SE Mexico (?) to Costa Rica  M T OI S1 S2 I1 I2 G

Telemiades squanda Evans, 1953    iN  M T S IL G

Telemiades moa Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades nicomedes (Möschler, 1879)  Dark Telemiades  SE Mexico to Guianas and upper Amazon  iN  M T S

Telemiades trenda Evans, 1953  TL: Surinam  M T OD

Telemiades pseudotrenda Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades amazonica Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades meris (Plötz, 1886)  Meris Telemiades  Panama & Colombia  iN  M T S1 S2 I1 I2

Telemiades brazus E. Bell, 1949  Brazil; ref.7  M T TI

Telemiades cryptus Siewert, O. Mielke & Casagrande, 2020  ref.36

Telemiades delalande (Latreille, [1824])  Delalande Skipper  Guatemala to Venezuela, Peru, upper Amazon  iN  M T OI S L

Polygonus  Hübner, [1825]  

Polygonus arizonensis (Skinner, 1911)  North America Polygon  SW USA (mostly as strays) & S TX to Central America; ref.20  M T S L I1 I2 G

Polygonus pallida Röber, 1925  Hammock Skipper  Central America to Argentina; ref.20  M T S L

Polygonus histrio Röber, 1925  Florida and Cuba, some Bahamian islands; ref.20  M S L I

Polygonus leo (Gmelin, [1790])  [= lividus, = ishmael], Hispaniola, Puerto Rico, SE to St. Lucia  iN  M T S IL L G

Polygonus hagar Evans, 1952  Jamaica; ref.20  M T S

Polygonus punctus E. Bell & W. Comstock, 1948  Dotted Polygon  Lesser Antilles; St. Martin S to Grenadines; ref.19  M T S

Polygonus pardus Grishin, 2023  Spotted Polygon  S TX to Panama; ref.19  M S1 S2 HS L I

Polygonus savigny (Latreille, [1824])  Manuel’s Skipper  [= manueli], Panama to Argentina; ref.19  iN  M T S G

  Tribe Oileidini, Subtribe Oileidina

Oileides  Hübner, [1825]  

Oileides fenestratus (Gmelin, [1790])  TL: [French Guiana]  M T S

Oileides guyanensis (Mabille & Boullet, 1912)  ref.7  M T S

Oileides vulpinus Hübner, [1825]    iN  M OI S

Flattoides  Grishin, 2019  

Flattoides amazonensis (E. Bell, 1947)  ref.5

Flattoides amazonensis amazonensis (E. Bell, 1947)  TL: Colombia  M T S G

Flattoides amazonensis renta (Evans, 1952)  TL: Peru  M T G

  Tribe Oileidini, Subtribe Typhedanina

Typhedanus  A. Butler, 1870  includes several species formerly in Cogia; ref.7

Typhedanus umber (Herrich-Schäffer, 1869)  TL: ?  M T S

Typhedanus ampyx (Godman & Salvin, 1893)  Gold-tufted Skipper  W & NE Mexico to Costa Rica  iN  M T S1 S2 L I G

Typhedanus salas H. Freeman, 1977  Salas Skipper  S Mexico & Belize  iN  M T S L

Typhedanus cajeta (Herrich-Schäffer, 1869)  Yellow-haired Skipper  [not Cogia]; ref.7  iN

Typhedanus cajeta cajeta (Herrich-Schäffer, 1869)  SE Mexico to Nicaragua  M T S L G

Typhedanus cajeta eluina (Godman & Salvin, 1894)  W Mexico to Costa Rica  M T S1 S2 L I

Typhedanus aventinus (Godman & Salvin, 1894)  Trimmed Skipper  W Mexico, Yucatan; ref.7  iN  M T S1 S2 IL L1 L2 Ha

Typhedanus mala (Evans, 1953)  Bad Gogia  Guatemala; ref.7  M T HS

Typhedanus buena (A. Warren, Dolibaina & Hernández-Mejía, 2015)  Good Gogia  S Mexico; ref.23,7  iN  M S

Oechydrus  E. Watson, 1893  

Oechydrus chersis (Herrich-Schäffer, 1869)    iN

Oechydrus chersis chersis (Herrich-Schäffer, 1869)  TL: ?  M T S

Oechydrus chersis ochrilinea (Schaus, 1902)  TL: Peru  M T S

Oechydrus chersis rufus Evans, 1953  TL: Brazil (RJ)  M T OD S

Oechydrus evelinda (A. Butler, 1870)  S Brazil; ref.7  M T TI S G

Cogia  A. Butler, 1870  includes several species formerly in Typhedanus; ref.7

Cogia crameri (McHenry, 1960)  ref.7  iN  M T S IL L

Cogia undulatus (Hewitson, 1867)  Mottled Longtail  [not Typhedanus] S Texas (as stray) to Argentina; ref.7  iN  M T S1 S2 HS IL L I G

Cogia galbula (Plötz, 1880)  ref.7  iN  M T S IL

Cogia optica (Evans, 1952)  ref.7  iN  M T S

Cogia goya (Evans, 1952)  Brazil; ref.7  M T S

Cogia stylites (Herrich-Schäffer, 1869)  ref.7  iN  M T S IL

Cogia aziris (Hewitson, 1867)  Brazil; ref.7  iN  M T S

Cogia cursinoi (O. Mielke, 1979)  Brazil; ref.7  M T OI G

Cogia eliasi (O. Mielke, 1979)  Brazil; ref.7  M T OI G

Cogia hassan A. Butler, 1870    iN

Cogia hassan hassan A. Butler, 1870  TL: Brazil (Pa)  M T S

Cogia hassan evansi E. Bell, 1937  TL: Bolivia  M T S

Cogia abdul Hayward, 1947    iN  M TI G

Cogia troilus Mabille, 1898  TL: Bolivia  M T S

Cogia punctilia Plötz, 1882    iN  M T S

Cogia cerradicola (O. Mielke, 1967)    iN  M T OI S G

Cogia outis (Skinner, 1894)  Outis Skipper  NE Mexico (CHIH, COAH), Texas, S-C OK to far SW MO  iN  M T S L1 L2 I H Ha G

Cogia hippalus (W. H. Edwards, 1882)  Acacia Skipper    iN

Cogia hippalus peninsularis L. Miller & MacNeill, 1969  Baja California Sur  M T S

Cogia hippalus hippalus (W. H. Edwards, 1882)  AZ, SW NM, W & S TX, W Mexico  M T S L I H G

Cogia hiska Evans, 1953  'East-Mexican' Acacia Skipper  E Mexico to Costa Rica; ref.146

Cogia hiska hiska Evans, 1953  E Mexico to Costa Rica; ref.146  M T S1 S2 L I G

Cogia hiska hester Evans, 1953  Colombia & Venezuela; ref.146  M T OD

Cogia azila Evans, 1953  TL: Bolivia  M T OD

Cogia moschus (W. H. Edwards, 1882)  Gold-costa Skipper  SE AZ, S NM to N MX; ref.146  M S L I H G

Cogia caicus (Herrich-Schäffer, 1869)  Mexican Gold-costa Skipper  N MX to Oax; ref.146  iN  M T S L G

Cogia chiagua Grishin, 2023  Southern Gold-costa Skipper  Chiapas to Costa Rica; ref.146  M T S

Cogia calchas (Herrich-Schäffer, 1869)  Mimosa Skipper  S Texas to N Argentina  iN  M T S1 S2 L1 L2 I G

Cogia grandis N. Riley, 1921    iN  M T S

Cogia elaites (Hewitson, 1867)  TL: Brazil (MG)  M T

Nerula  Mabille, 1888  

Nerula fibrena (Hewitson, 1877)  TL: Brazil (Am)  M T OI S

Nerula tuba Evans, 1953  TL: Brazil (Pa)  M T OD

Marela  Mabille, 1903  

Marela tamyroides (C. Felder & R. Felder, 1867)    iN  M T S L

Marela tamyris Mabille, 1903    iN  M T TI S L

Marela tamba Evans, 1953  Peru; ref.7  M T S

  Subfamily Pyrrhopyginae

  Tribe Pyrrhopygini

Azonax  Godman & Salvin, 1893  

Azonax typhaon (Hewitson, 1877)  Typhaon Skipper  SE Mexico (OAX), Nicaragua  M T S L

Azonax guiana O. Mielke, Brockmann & C. Mielke, 2022  French Guiana; ref.30  M S

Azonax urupa O. Mielke, Brockmann & C. Mielke, 2022  Brazil (Rd); ref.30

Zonia  Evans, 1951  

Zonia zonia Evans, 1951  Zonia Skipper    iN

Zonia zonia panamensis Nicolay, 1975  Belize, Panama  M T OI S G

Zonia zonia zonia Evans, 1951  N Brazil (Am)  M T TI

Zonia zonia diabo O. Mielke & Casagrande, 1998  SE Brazil (SP)  M T

Zonia zonia eberti O. Mielke, C. Mielke & Casagrande, 2022  ref.30

Granila  Mabille, 1903  

Granila paseas (Hewitson, 1857)  Paseas Skipper  SE Brazil, Paraguay  iN  M T OI S IL

Aspitha  Evans, 1951  

Aspitha leander (Boullet, 1912)  Leander Firetip  SE Mexico (OAX), Panama, Colombia  iN  M T S IL G Txt

Aspitha aspitha (Hewitson, 1866)  Aspitha Firetip    iN

Aspitha aspitha parima (Plötz, 1886)  Trinidad, French Guiana, Surinam, N Brazil  M TI

Aspitha aspitha aspitha (Hewitson, 1866)  N Brazil (Pa, Am)  M T TI S

Aspitha aspitha rufescens (N. Riley, 1919)  Brazil (Rd)  M T S

Aspitha teffa Evans, 1951  ref.30  M T

Aspitha bassleri (E. Bell, 1940)  Bassler's Firetip  Peru  M T S

Aspitha agenoria (Hewitson, 1876)  Red-lobed Firetip    iN

Aspitha agenoria cruor (H. Druce, 1908)  Peru  M T

Aspitha agenoria agenoria (Hewitson, 1876)  Colombia, Peru, Bolivia  M T S IL

Myscelus  Hübner, [1819]  

Myscelus nobilis (Cramer, 1777)  Noble Myscelus  Surinam, Venezuela, Peru, Brazil, Bolivia  iN  M T S

Myscelus illustris Mabille, 1903  ref.30  M T HS IL

Myscelus amystis (Hewitson, 1867)  Widespread Myscelus    iN

Myscelus amystis hages Godman & Salvin, 1893  E & W Mexico to Panama  M T S1 S2 HS L I Ha G

Myscelus amystis muttra Evans, 1951  Colombia (Valle del Cauca)  M T

Myscelus amystis amystis (Hewitson, 1867)  Trinidad, Venezuela, Colombia  M T TI S

Myscelus amystis distinctus Röber, 1925  Ecuador  M S G

Myscelus amystis mysus Evans, 1951  Peru, Bolivia  M T S

Myscelus epigona Herrich-Schäffer, 1869  S Brazil, Paraguay, Argentina; ref.30  M T TI S HS IL

Myscelus phoronis (Hewitson, 1867)  Phoronis Myscelus    iN

Myscelus phoronis phoronis (Hewitson, 1867)  Guatemala, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia  M T TI S IL L

Myscelus phoronis caucanus Staudinger, 1888  Colombia  M T S

Agara  Mabille & Boullet, 1908  

Agara belti (Godman & Salvin, 1879)  Belt's Myscelus  SE Mexico (OAX, VER, CHIS) to Venezuela, Colombia & Ecuador  iN

Agara belti belti (Godman & Salvin, 1879)  SE Mexico (OAX, VER, CHIS) to Venezuela, Colombia & Ecuador  M T TI S I1 I2 I3

Agara belti caucaensis (Constantino, Salazar & Vargas, 2013)  W Colombia; ref.58

Agara perissodora (Dyar, 1914)  Dyar's Myscelus  S Mexico (GRO, OAX, VER) to Colombia & Venezuela  iN  M T S L I G

Agara pegasus (Mabille, 1903)  Pegasus Myscelus  French Guiana, Venezuela, Ecuador  M T TI S

Agara draudti (N. Riley, 1926)  Draudt's Myscelus  Peru, Bolivia  M T S

Agara epimachia (Herrich-Schäffer, 1869)  Red Myscelus    iN

Agara epimachia verina (Evans, 1951)  Ecuador  M T S

Agara epimachia epimachia (Herrich-Schäffer, 1869)  Venezuela, Colombia, Ecuador, Peru, Bolivia, S Brazil  M S IL

Agara epimachia nevada O. Mielke & Casagrande, 2022  ref.30

Agara epimachia hermieri O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Agara epimachia edix (Evans, 1951)  SE Brazil, Paraguay  M T S

Agara santhilarius (Latreille, [1824])  Hilarious Myscelus  SE Brazil  iN  M T S IL

Agara assaricus (Cramer, 1779)  Fiery Myscelus    iN

Agara assaricus assaricus (Cramer, 1779)  French Guiana, Surinam, N Brazil  M T S G

Agara assaricus mapirica Strand, 1921  S Venezuela, Peru, Brazil (Rd)  M T S IL

Agara michaeli (Nicolay, 1975)  Fiery Myscelus  SE Mexico (OAX, CHIS), Guatemala, Costa Rica, Panama  M T OI S1 S2 L I G

Agara pardalina (C. Felder & R. Felder, 1867)  Flaming Myscelus    iN

Agara pardalina kesselringi (O. Mielke & Casagrande, 2011)  Peru  M T OI

Agara pardalina guarea (Evans, 1951)  N Brazil (Para)  M T G

Agara pardalina pardalina (C. Felder & R. Felder, 1867)  Colombia, Ecuador, Peru, Bolivia, Brazil  M T S IL

Agara pardalina yacutinga (O. Mielke & Casagrande, 2011)  Argentina & Paraguay  M T OI

Agara gallardi O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Passova  Evans, 1951  

Passova nigrocephala (E. Bell, 1934)  Black-headed Firetip  Colombia  iN  M T S

Passova polemon (Hopffer, 1874)  Polemon Firetip  S Brazil  iN  M T S IL L

Passova ganymedes (E. Bell, 1931)  Mimetic Firetip    iN

Passova ganymedes ganymedes (E. Bell, 1931)  Colombia  M T S

Passova ganymedes gulia Evans, 1951  Ecuador  M T

Passova ganymedes gelina Evans, 1951  Peru  M T S

Passova gellias (Godman & Salvin, 1893)  Gellias Firetip  Belize, Honduras, Costa Rica, Panama & Venezuela  iN  M T S1 S2 IL I1 I2

Passova glacia Evans, 1951  Shining Firetip  Surinam, French Guiana  iN  M T S

Passova gazera (Hewitson, 1866)  Gazera Firetip  Brazil (Amazonas)  M T TI S IL

Passova passova (Hewitson, 1866)  Passova Firetip    iN

Passova passova gortyna (Hewitson, 1866)  N Brazil  M T TI S G

Passova passova passova (Hewitson, 1866)  French Guiana  M T S IL

Passova passova rudex Evans, 1951  N Brazil, Peru  M T S

Passova passova styx (Möschler, 1879)  Colombia, Venezuela, Bolivia  M T S

Passova passova practa Evans, 1951  S Brazil, Paraguay  M T S

Passova passova stacer Evans, 1951  Surinam, Guyana  M T

Passova passova diamantino O. Mielke, C. Mielke & Casagrande, 2022  ref.30

Passova greta Evans, 1951  Great Firetip  Peru, Bolivia  iN  M T S

Passova vilna Evans, 1951  Vilna Firetip    iN

Passova vilna vilna Evans, 1951  Bolivia  M T

Passova vilna vanta Evans, 1951  Bolivia  M T

Pyrrhopyge  Hübner, [1819]  

Pyrrhopyge zenodorus Godman & Salvin, 1893  Red-headed Firetip  SE Mexico to Costa Rica  iN  M T S L I1 I2 G

Pyrrhopyge evansi E. Bell, 1947  Evans' Firetip  TL: Colombia

Pyrrhopyge evansi evansi E. Bell, 1947  Costa Rica, Panama, Colombia, Venezuela (Bolivar), French Guiana, [Ecuador?]  M T S G

Pyrrhopyge evansi borburata Orellana, [2010]  Venezuela (N slope Cordillera de la Costa, Carabobo)

Pyrrhopyge phidias (Linnaeus, 1758)  Original Firetip    iN

Pyrrhopyge phidias hyperici Hübner, 1823  Brazil (Para)  M T S IL

Pyrrhopyge phidias latifasciata A. Butler, 1873  Colombia  M T S IL

Pyrrhopyge phidias aldo O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge phidias bixae (Linnaeus, 1758)  Surinam, French Guiana, N Brazil to Matto Grosso  M S

Pyrrhopyge phidias rusca Evans, 1951  Ecuador, Peru, SW Brazil  M T

Pyrrhopyge phidias leucoloma Erschoff, 1875  Peru, Bolivia  M S

Pyrrhopyge phidias garata Hewitson, 1866  Guyana, Surinam, [N Venezuela?]  M T S IL

Pyrrhopyge phidias phidias (Linnaeus, 1758)  Trinidad, Surinam, Guyana, Venezuela, Colombia, Peru, Bolivia, Paraguay, Brazil  M T S IL

Pyrrhopyge guianae E. Bell, 1932  Trinidad, Guyana, French Guiana  M T

Pyrrhopyge aziza Hewitson, 1866  Aziza Firetip    iN

Pyrrhopyge aziza aziza Hewitson, 1866  Colombia  M T TI S

Pyrrhopyge aziza araethyrea Hewitson, 1870  Ecuador  M T TI S IL

Pyrrhopyge aziza troja Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge attis E. Bell, 1931  Bolivia

Pyrrhopyge attis lexos Evans, 1951  Guyana, Colombia, Ecuador, Peru  M T S

Pyrrhopyge attis arbor Evans, 1951  Colombia, Venezuela  M T S

Pyrrhopyge attis cardus Mabille, 1891  N Brazil  M T S

Pyrrhopyge attis attis E. Bell, 1931  Bolivia  M T

Pyrrhopyge attis miersi O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge attis subnubilus Hayward, 1935  Argentina

Pyrrhopyge pusca Evans, 1951  Pusca Firetip  Peru, Bolivia  M T S

Pyrrhopyge tingo O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge rafael O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge iquitos O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge sabrina O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge draudti E. Bell, 1931  TL: Bolivia

Pyrrhopyge draudti bruno O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge draudti lina E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge draudti cintra Evans, 1951  French Guiana, Surinam, Guyana, N Brazil, NE Peru  M T S

Pyrrhopyge draudti draudti E. Bell, 1931  Venezuela, Colombia, Ecuador, Peru, Bolivia (Andean)  M T S

Pyrrhopyge proculus Hopffer, 1874  Proculus Firetip    iN

Pyrrhopyge proculus proculus Hopffer, 1874  Trinidad, Guyana, Venezuela (E Cordillera de la Costa), Colombia, N Brazil  M T IL

Pyrrhopyge proculus ponicia Evans, 1951  French Guiana, Surinam, N Brazil  M T S

Pyrrhopyge infantilis H. Druce, 1908  Infant Firetip    iN

Pyrrhopyge infantilis infantilis H. Druce, 1908  Peru, Bolivia  M T S IL

Pyrrhopyge infantilis agala Evans, 1951  Bolivia  M T S

Pyrrhopyge thericles Mabille, 1891  Impostor Firetip    iN

Pyrrhopyge thericles orientis E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge thericles thericles Mabille, 1891  NE Brazil  M T S

Pyrrhopyge thericles pseudophidias E. Bell, 1931  Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru, N Brazil  M T S

Pyrrhopyge thericles raymondi Orellana, [2010]  Venezuela (Cordillera de la Costa, Carabobo, Aragua, Guarico, Miranda)

Pyrrhopyge thericles fola Evans, 1951  Colombia, SW Venezuela  M T

Pyrrhopyge thericles rileyi E. Bell, 1931  Ecuador, Bolivia  M T

Pyrrhopyge thericles ronda Evans, 1953  Trinidad  M T

Pyrrhopyge amythaon E. Bell, 1931  Amythaon Firetip  TL: Brazil (Pe)

Pyrrhopyge amythaon gradens Evans, 1951  N Brazil (Para)  M T S

Pyrrhopyge amythaon peron E. Bell, 1947  Venezuela (Bolivar), Colombia, NE Peru  M T S

Pyrrhopyge amythaon podina Evans, 1951  Peru, Bolivia  M T

Pyrrhopyge amythaon amythaon E. Bell, 1931  Brazil (Pernambuco)  M T

Pyrrhopyge josephina Draudt, 1921  Peru, Bolivia

Pyrrhopyge josephina polka Evans, 1951  French Guiana  M T S

Pyrrhopyge josephina josephina Draudt, 1921  Peru, Bolivia  M T S IL

Pyrrhopyge josephina orino Evans, 1951  Colombia, Venezuela (S of Orinoco), Guyana  M T S

Pyrrhopyge josephina pollio Evans, 1951  Peru  M T

Pyrrhopyge divisor O. Mielke & Casagrande, 2022  ref.30

Pyrrhopyge mina E. Bell, 1931  TL: Bolivia

Pyrrhopyge mina mina E. Bell, 1931  TL: Bolivia  M T

Pyrrhopyge mina ganus E. Bell, 1947  Guyana, Venezuela (Cordillera de la Costa), [Colombia, Ecuador, Peru, Brazil?]  M T

Pyrrhopyge mina previsto O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge sergius Hopffer, 1874  Sergius Firetip    iN

Pyrrhopyge sergius selina Evans, 1951  Brazil (Para)  M T S

Pyrrhopyge sergius andros Evans, 1951  Colombia, Venezuela (Amazonas)  M T

Pyrrhopyge sergius andronicus E. Bell, 1931  Ecuador, Peru, Bolivia  M T S

Pyrrhopyge sergius semana Evans, 1951  Surinam, French Guiana  M T S

Pyrrhopyge sergius sergius Hopffer, 1874  Peru  M T S IL

Pyrrhopyge michael O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge jola O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge caribe Orellana, [2010]  Caribbean Firetip  TL: Venezuela

Pyrrhopyge caribe caribe Orellana, [2010]  Venezuela (Aragua, Carabobo, Dtto. Capital, Lara, Trujillo, Vargas)

Pyrrhopyge caribe camachoi Orellana, [2010]  Venezuela (Gran Sabana)

Pyrrhopyge eduardo O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge erazoae Orellana, [2010]  Erazo's Firetip  Venezuela (Bolivar)  M S

Pyrrhopyge charybdis Westwood, 1852  Charybdis Firetip    iN

Pyrrhopyge charybdis semita Evans, 1951  Bolivia, S Brazil  M T S

Pyrrhopyge charybdis charybdis Westwood, 1852  S Brazil  M T OI S IL L

Pyrrhopyge cressoni E. Bell, 1932  Cresson's Firetip  Ecuador, Bolivia  M T S

Pyrrhopyge pelota Plötz, 1879  Pelota Firetip  Bolivia, Paraguay, S Brazil  iN  M T S

Pyrrhopyge amyclas (Cramer, 1779)  Yellow-edged Firetip    iN

Pyrrhopyge amyclas amyclas (Cramer, 1779)  Trinidad, Venezuela, Guyana, Surinam, French Guiana, Brazil  M S IL

Pyrrhopyge amyclas denticulata Herrich-Schäffer, 1869  Ecuador, Peru, Brazil  M S L

Pyrrhopyge phylleia Hewitson, 1874  Orange-edged Firetip  S Peru, Bolivia  iN  M T TI S IL L G

Pyrrhopyge arax Evans, 1951  Arax Firetip  SE Peru, Bolivia  iN  M T S L

Pyrrhopyge caranavi O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge haemon Godman & Salvin, 1893  Tawny-rimmed Firetip  Costa Rica  M T S

Pyrrhopyge papius Hopffer, 1874  Papius Firetip    iN

Pyrrhopyge papius pasca Evans, 1951  Colombia  M T S

Pyrrhopyge papius papius Hopffer, 1874  Venezuela, Colombia, Ecuador, Peru (Andean)  M T S

Pyrrhopyge frona Evans, 1951  Inca Firetip  Peru  iN  M T S

Pyrrhopyge melanomerus Mabille & Boullet, 1908  Melanomerus Firetip  TL: Bolivia

Pyrrhopyge melanomerus patma Evans, 1951  S Ecuador, NE Peru  M T

Pyrrhopyge melanomerus melanomerus Mabille & Boullet, 1908  Peru, Bolivia  M T L

Pyrrhopyge afluente O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge decipiens Mabille, 1903  Red-spotted Firetip  [Ecuador?], Peru, Bolivia  iN  M T S

Pyrrhopyge placeta Evans, 1951  Placeta Firetip  Brazil (ES)  M T

Pyrrhopyge sanpedro O. Mielke & Lamas, 2022  ref.30

Pyrrhopyge hadassa Hewitson, 1866  Hadassa Firetip  Colombia, Ecuador, Peru  iN  M T TI S IL

Pyrrhopyge olivacea Röber, 1925  TL: Peru  M T

Pyrrhopyge hanga Evans, 1951  Peru  M T

Pyrrhopyge halma Evans, 1951  Peru, Bolivia  M T S L

Pyrrhopyge pseudohadassa Mabille & Boullet, 1908  Peru  M T S L

Pyrrhopyge cosnipata O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge terra Evans, 1951  Terra Firetip  Bolivia  M T S

Pyrrhopyge telassina Staudinger, 1888  Peru  iN  M T S HS

Pyrrhopyge telassa Hewitson, 1866  Telassa Firetip  Ecuador  iN  M T TI S IL

Pyrrhopyge phaeax Hopffer, 1874  Peru  M T S IL L

Pyrrhopyge croceimargo Mabille & Boullet, 1908  S Peru, Bolivia  M T S IL L

Pyrrhopyge leticia O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge mendax Mabille & Boullet, 1908  TL: Peru  M T S

Pyrrhopyge shiva Evans, 1951  SE Peru, Bolivia  M T S L

Pyrrhopyge martena Hewitson, [1869]  Martena Firetip  Ecuador  iN  M T OI S

Pyrrhopyge schausi E. Bell, 1931  Schaus' Firetip  Colombia, Ecuador, N Peru  M T S

Pyrrhopyge sadia Evans, 1951  Sadia Firetip  Ecuador, Peru  iN  M T S

Pyrrhopyge creona H. Druce, 1874  Creona Firetip  Peru, Brazil (Am)  M T S IL

Pyrrhopyge kelita Hewitson, [1869]  Streaked Firetip  SE Peru, Bolivia  iN  M T OI IL L

Pyrrhopyge crista Evans, 1951  Crista Firetip  Bolivia  M T

Pyrrhopyge crida Hewitson, 1871  White-banded Firetip  SE Mexico to Colombia and Ecuador  iN  M T TI S1 S2 IL I1 I2 G

Pyrrhopyge mopsus (E. Bell, 1931)  Mopsus Firetip  Peru

Pyrrhopyge mopsus mopsus (E. Bell, 1931)  Peru  M T S

Pyrrhopyge mopsus elisa O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge sarpedon (E. Bell, 1931)  Sarpedon Firetip  Peru (Amazonas)  M T

Pyrrhopyge arinas (Cramer, 1777)  Arinas Firetip  TL: Surinam

Pyrrhopyge arinas arinas (Cramer, 1777)  French Guiana, Surinam, N Brazil, NE Peru  M T TI S IL

Pyrrhopyge arinas erica O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge arinas temenos (E. Bell, 1931)  Peru  M T S

Pyrrhopyge cacaulandia O. Mielke, Brockmann & C. Mielke, 2022  ref.30

Pyrrhopyge creusae (E. Bell, 1931)  Creusae Firetip  French Guiana  M T

Pyrrhopyge steyermarki Orellana, Costa & Grishin, 2021  Venezuela; ref.83

Pyrrhopyge tatei (E. Bell, 1932)  Tate's Firetip  Venezuela  M T

Pyrrhopyge boulleti Le Cerf, 1922  Boullet's Firetip  Colombia, Venezuela (Tachira)  iN  M S

Guyanna  Grishin, 2022  

Guyanna tetricus (E. Bell, 1931)  Tetricus Skipper  Guyana, Peru; ref.17  iN  M T

Gunayan  O. Mielke, 2002  

Gunayan rhacia (Hewitson, [1875])  Rhacia Skipper  S Brazil  iN  M T OI IL

Gunayan rubricollis (Sepp, [1841])  Red-collared Skipper  French Guiana, Peru, Bolivia, Brazil  iN

Gunayan rubricollis rubricollis (Sepp, [1841])  French Guiana, Peru, Bolivia, Brazil  M T TI S IL L

Gunayan rubricollis flemingi (E. Bell, 1947)  TL: Venezuela  M T

Gunayan timaeus (E. Bell, 1931)  Timaeus Skipper  Colombia, Peru  M T S

Yanguna  E. Watson, 1893  

Yanguna cometes (Cramer, 1779)  Comet Skipper    iN

Yanguna cometes cometes (Cramer, 1779)  French Guiana, N Brazil (Para)  M S

Yanguna cometes staudingeri (Plötz, 1879)  Surinam, Venezuela (Bolivar), S Colombia (upper Putumayo), Ecuador, N Peru  M S IL L

Yanguna cometes angeli Orellana, [2010]  Venezuela (Tachira)

Yanguna cometes cometides Mabille & Boullet, 1908  Peru, Bolivia  M S IL

Yanguna thelersa (Hewitson, 1866)  Thelersa Skipper  French Guiana, N Brazil, Ecuador, Peru  iN  M T S

Yanguna blossomiae R. Williams, 1931  TL: Ecuador

Yanguna blossomiae blossomiae R. Williams, 1931  TL: Ecuador  M T OI

Yanguna blossomiae galion O. Mielke, Brockmann & C. Mielke, 2022  French Guiana; ref.30

Yanguna moa O. Mielke, Casagrande & Brockmann, 2022  Peru; ref.30

Yanguna moa moa O. Mielke, Casagrande & Brockmann, 2022  Peru; ref.30

Yanguna moa guiana O. Mielke, Casagrande & Brockmann, 2022  French Guiana; ref.30

Yanguna cosyra (H. Druce, 1875)  Cosyra Skipper  Guatemala to Colombia and Ecuador  iN  M T S1 S2 IL I1 I2 I3 G

Yanguna spatiosa (Hewitson, 1870)  Spatiosa Skipper    iN

Yanguna spatiosa spatiosa (Hewitson, 1870)  W Colombia, Ecuador  M T TI S IL L

Yanguna spatiosa julian O. Mielke, Brockmann & C. Mielke, 2022  Colombia; ref.30

Yanguna aspilos Mabille & Boullet, 1908  Bolivia

Yanguna aspilos mabillei H. Druce, 1909  Peru  M T S L

Yanguna aspilos lara O. Mielke, Brockmann & C. Mielke, 2022  Peru; ref.30

Yanguna aspilos aspilos Mabille & Boullet, 1908  Bolivia  M T OI IL

Yanguna erebus (Plötz, 1879)  Erebus Skipper  Brazil (Ba)  M T

Apyrrothrix  Lindsey, 1921  

Apyrrothrix arizonae (Godman & Salvin, 1893)  Araxes Skipper  SE AZ, SW NM, W TX (local), Son, Chih, NL, Tam, SLP; ref.30  iN  M T S L I G

Apyrrothrix araxes (Hewitson, 1867)  Araxes Skipper    iN

Apyrrothrix araxes Sinaloa-Durango segregate ,  NW Mexico (SIN, DGO [probably NW NAY])  M S

Apyrrothrix araxes Nayarit segregate ,  W-central Mexico (SW NAY)  M S

Apyrrothrix araxes Guerrero-Michoacan segregate ,  SW Mexico (GRO, MICH)  M S

Apyrrothrix araxes araxes (Hewitson, 1867)  central Mexico (COL, JAL, DF, MOR, HGO, PUE, VER)  M T S IL L G

Apyrrothrix araxes cyrillus (Plötz, 1879)  S Mexico (OAX)  M T S L

Apyrrothrix mulleri E. Bell, 1934  Red-spotted Skipper  SE Mexico (CHIS, OAX, VER)  iN  M T S L G

Apyrrothrix hoffmanni (H. Freeman, 1977)  Hoffmann's Skipper  SE Mexico (OAX, TAB, CHIS), Belize  iN  M T OD S L G

Apyrrothrix erythrosticta (Godman & Salvin, 1879)  Spotted Skipper  SE Mexico to Panama  iN  M T OD S IL I G Txt

Apyrrothrix burnsi (O. Mielke, Brockmann & C. Mielke, 2022)  Dotted Skipper  SE Mexico to Costa Rica  M S1 S2 S3 HS L I1 I2

Apyrrothrix maculosa (Hewitson, 1866)  Maculosa Skipper  Panama, Colombia, Ecuador, Venezuela (Zulia)  iN  M T S IL L

Apyrrothrix cossea (H. Druce, 1875)  Cossea Skipper  Colombia  M T S

Apyrrothrix sangaris (Skinner, 1921)  Sangaris Skipper  Colombia  M T S

Apyrrothrix chalybea (Scudder, 1872)  Orange-rimmed Skipper    iN

Apyrrothrix chalybea chalybea (Scudder, 1872)  C Mexico interior (central JAL, GTO, QRO)  M S IL L G

Apyrrothrix chalybea chloris (Evans, 1951)  W and S Mexico, Belize, Guatemala, Honduras  M T S IL L

Apyrrothrix hygieia (C. Felder & R. Felder, 1867)  Colombia, N Ecuador; ref.8  iN  M T S IL

Apyrrothrix zereda (Hewitson, 1866)  Zereda Skipper    iN

Apyrrothrix zereda zereda (Hewitson, 1866)  Ecuador  M T IL

Apyrrothrix zereda rufipectus (Godman & Salvin, 1879)  Colombia, C Ecuador  M T S

Apyrrothrix zereda zepha (Evans, 1951)  S Ecuador  M T

Apyrrothrix zereda insana (Staudinger, 1876)  Peru  M T S IL

Apyrrothrix rufinucha (Godman & Salvin, 1879)  Peru, Bolivia; ref.30  M T S IL L

Apyrrothrix aesculapus (Staudinger, 1876)  Fire-edged Skipper  Costa Rica, Panama, Colombia, Ecuador  iN  M T OD S1 S2 IL I1 I2 G

Creonpyge  O. Mielke, 2002  

Creonpyge creon (H. Druce, 1874)  Creon Skipper    iN

Creonpyge creon creon (H. Druce, 1874)  N Costa Rica to Cocle Prov., Panama, Atlantic & Pacific slopes  M T S1 S2 IL L I1 I2 G

Creonpyge creon lilliana (Nicolay & Small, 1969)  Cerro Campana, far E Cocle, far W Panama Prov., Panama  M T S

Creonpyge creon taylori (Nicolay & Small, 1981)  Fila Cruces, Puntarenas, SE Costa Rica  M T S

Jonaspyge  O. Mielke, 2002  

Jonaspyge jonas (C. Felder & R. Felder, 1859)  Scallop-edged Firetip  SE Mexico (HGO, OAX, PUE, VER, CHIS), Guatemala, Nicaragua  iN  M T OD S IL L G

Jonaspyge tzotzili (H. Freeman, 1969)  Freeman's Firetip  SE Mexico (CHIS, OAX)  M T OD S

Jonaspyge elizabethae Gallardo & Grishin, 2021  Honduras; ref.116

Ardaris  E. Watson, 1893  

Ardaris aerata (Godman & Salvin, 1879)  Aerata Skipper  Colombia  iN  M T S

Ardaris eximia (Hewitson, 1871)  Crystal-banded Skipper  Venezuela (Cordillera de Merida from Tachira to Trujillo & Lara)  iN  M T TI S IL

Ardaris hantra Evans, 1951  Amber-banded Skipper  Venezuela (Cordillera de Merida in Merida)  M T

Ardaris sela (Hewitson, 1866)  Sela Skipper    iN

Ardaris sela sela (Hewitson, 1866)  Colombia  M T OI S

Ardaris sela aequatorea (Röber, 1925)  Colombia, Ecuador, N Peru  M T S

Ardaris sela peruviana (Draudt, 1921)  Peru  M T S G

Ardaris periphema (Hewitson, [1875])  Peru, Bolivia  M T OI S L

Ardaris lomax (Evans, 1951)  Lomax Skipper  Peru  iN  M T S

Ardaris montra (Evans, 1951)  Montra Skipper  Peru  iN  M T S

Ardaris pityusa (Hewitson, 1857)  Pityusa Skipper  Ecuador, Peru  iN  M T OI S IL L

Ardaris porus (Plötz, 1879)  Porus Skipper    iN

Ardaris porus porus (Plötz, 1879)  Colombia  M T OI S HS IL

Ardaris porus mortis (Evans, 1951)  Peru  M T S

Ardaris minthe (Godman & Salvin, 1879)  Minthe Skipper  Colombia, Ecuador, N Peru  iN  M T S IL

Mysoria  E. Watson, 1893  

Mysoria cosinga (Hewitson, 1874)  Cosinga Firetip    iN

Mysoria cosinga catana (Evans, 1951)  Peru, Bolivia  M T S

Mysoria cosinga circe (Evans, 1951)  Bolivia  M T S

Mysoria cosinga cedra (Evans, 1951)  Bolivia  M T S G

Mysoria cosinga cosinga (Hewitson, 1874)  Peru, Bolivia  M T TI S HS IL

Mysoria xanthippe (Latreille, [1824])  Xanthippe Firetip    iN

Mysoria xanthippe xanthippe (Latreille, [1824])  S Brazil  M T OI IL

Mysoria xanthippe spixii (Plötz, 1879)  S Brazil  M T S HS IL

Mysoria damippe (Mabille & Boullet, 1908)  Damippe Firetip  S Brazil  iN  M T OI S IL

Mysoria pertyi (Plötz, 1879)  Perty's Firetip  S Brazil  iN  M IL

Mysoria curitiba (O. Mielke & Casagrande, 2002)  Curitiba Firetip  S Brazil  M OI S

Mysoria soza (Evans, 1951)  Soza Firetip  S Brazil (MG, RJ, Pr, SC)  iN  M T IL G

Mysoria oneka (Hewitson, 1866)  Oneka Firetip  [S Brazil]  M T IL

Mysoria catomelaena (Mabille & Boullet, 1908)  Cato Firetip  S Brazil  iN

Mysoria catomelaena catomelaena (Mabille & Boullet, 1908)  S Brazil  M T OI

Mysoria catomelaena pico (O. Mielke & Casagrande, 2022)  Brazil (Ba); ref.30

Mysoria antias (C. Felder & R. Felder, 1859)  Antias Firetip  S Brazil  iN  M T S IL

Mysoria sejanus (Hopffer, 1874)    iN

Mysoria sejanus stolli (O. Mielke, 2002)  Costa Rica, Colombia, Venezuela, French Guiana, Brazil, NE Peru  M T S IL L

Mysoria sejanus sejanus (Hopffer, 1874)  Ecuador, Peru  M T IL

Mysoria sejanus erythrostigma Röber, 1925  Bolivia  M T OD S

Mysoria ambigua (Mabille & Boullet, 1908)  Royal Firetip  SE Mexico (CHIS, TAB, VER) to Costa Rica  iN  M T S1 S2 L I Txt

Mysoria barcastus (Sepp, [1851])  Barcastus Firetip    iN

Mysoria barcastus venezuelae (Scudder, 1872)  Panama, Colombia, Venezuela, Tobago, [Guianas]  M T S HS IL L G

Mysoria barcastus alta Evans, 1951  Trinidad  M T

Mysoria barcastus barcastus (Sepp, [1851])  Guyana, Suriname, French Guiana  M T OI OD I

Mysoria barcastus pallens (Mabille, 1891)  S Venezuela (Amazonas, Bolivar), Colombia  M T

Mysoria barcastus antila Evans, 1951  N Brazil  M T

Mysoria barcastus barta Evans, 1951  Bolivia, Paraguay, S Brazil, Argentina  M T S L

Mysoria amra (Hewitson, 1871)  Blue-collared Firetip  W Mexico, Guatemala  iN  M T OI S L G

Mysoria affinis (Herrich-Schäffer, 1869)  Red-collared Firetip  [= wilsoni], W Mexico, Guatemala  iN  M T S IL L G

Mysoria galgala (Hewitson, 1866)  Red-banded Firetip  S Mexico (SW CHIS), Panama, Colombia and Venezuela  iN  M T S IL L G

Mimoniades  Hübner, 1823  

Mimoniades picia (Evans, 1951)  Colombia  M T S1 S2

Mimoniades pionia (Hewitson, 1857)  Pionia Skipper    iN

Mimoniades pionia sandra (Orellana, [2010])  Colombia, Venezuela (E slope Andes in Tachira, Barinas)  M S

Mimoniades pionia pionia (Hewitson, 1857)  Colombia, Venezuela (Cordillera de la Costa), disjunct in Bolivia, N Argentina  M T OI S IL L

Mimoniades pedro (O. Mielke & Casagrande, 2022)  Brazil (Ba); ref.30

Mimoniades ponina (Herrich-Schäffer, 1869)  Venezuela, Surinam

Mimoniades ponina ponina (Herrich-Schäffer, 1869)  Venezuela, Surinam  M S

Mimoniades ponina rogeri (Orellana, [2010])  Venezuela (Merida, Lara, at least)  M S

Mimoniades baroni (Godman & Salvin, 1895)  Baron's Skipper  Peru  M T S Txt

Mimoniades eupheme (Godman & Salvin, 1879)  Ecuador, Peru, Bolivia; ref.30  M T S L

Mimoniades versicolor (Latreille, [1824])  Versicolor Skipper  Brazil; ref.30  iN  M T S

Mimoniades montana J. Zikán, 1938  Quadricolor Skipper  SE Brazil  iN  M S

Mimoniades nurscia (Swainson, 1821)  Nursica Skipper    iN

Mimoniades nurscia malis (Godman & Salvin, 1879)  Colombia  M T S IL

Mimoniades nurscia nurscia (Swainson, 1821)  Colombia, Ecuador, N Peru  M S

Mimoniades nurscia amans Skinner, 1920  Colombia, Peru  M T S IL

Mimoniades ocyalus Hübner, 1823  Ocyalus Skipper  S Brazil  iN  M S L

Mimoniades fallax (Mabille, 1878)  Fallax Skipper    iN

Mimoniades fallax fiska (Evans, 1951)  Colombia: Cauca & Cundinamarca  M T

Mimoniades fallax fida (Evans, 1951)  Ecuador, Peru, Bolivia  M T S1 S2 L

Mimoniades fallax solaris (Hayward, 1942)  Colombia, Venezuela, Guyana, N Brazil, E Peru, Bolivia  M T S1 S2

Mimoniades fallax fallax (Mabille, 1878)  Guianas, N Brazil  M T OD S L Txt

Jemadia  E. Watson, 1893  

Jemadia menechmus (Mabille, 1878)  Menechmus Skipper    iN

Jemadia menechmus menechmus (Mabille, 1878)  Surinam, Venezuela, Colombia, Ecuador, Peru, Bolivia, N & W Brazil  M T OD S

Jemadia menechmus desousai Orellana, [2010]  Venezuela (Amazonas)  M OI

Jemadia scomber H. Druce, 1908  Mammoth Skipper  Peru  iN  M T S L

Jemadia pseudognetus (Mabille, 1878)  Dot-collared Skipper  SE Mexico to Venezuela and upper Amazon  iN

Jemadia pseudognetus pseudognetus (Mabille, 1878)  SE Mexico to Venezuela and upper Amazon  iN  M T OD S1 S2 L I1 I2 G Txt

Jemadia pseudognetus imitator (Mabille, 1891)  Colombia  M T S

Jemadia hospita (A. Butler, 1877)  Hospita Skipper    iN

Jemadia hospita hospita (A. Butler, 1877)  S Colombia (upper Putumayo), Ecuador, Peru, Bolivia, W Brazil  M T S IL

Jemadia hospita hephaestos (Plötz, 1879)  Surinam  M T S HS L

Jemadia demarmelsi Orellana, [2010]  Demarmels' Skipper  Venezuela (Bolivar)  M OI

Jemadia mabillei O. Mielke, Brockmann & C. Mielke, 2022  [= Evans' "sosia"] SW Venezuela (Tachira), Colombia, Ecuador, Pe ...; ref.30  iN  M T

Jemadia gnetus (Fabricius, 1781)  Gnetus Skipper  Surinam, Guyana, Venezuela, Coloimbia, Ecuador, Peru, Bolivia, S Brazil  iN

Jemadia gnetus gnetus (Fabricius, 1781)  Surinam, Guyana, Venezuela, Coloimbia, Ecuador, Peru, Bolivia, S Brazil  M T OI OD S1 S2 HS L

Jemadia gnetus brevipennis Schaus, 1902  Brazil (DF, SP, Pr), Paraguay  iN  M T OD Txt

Jemadia gnetus mineira (O. Mielke, C. Mielke & Casagrande, 2022)  Brazil (MG); ref.30

Jemadia suekentonmiller Grishin, 2014  Speciesex Skipper  Costa Rica & Panama; ref.72  M T1 T2 T3 Txt

Jemadia ovid Evans, 1951  Ovid Skipper  Colombia, Ecuador  iN  M T S G

Jemadia pater Evans, 1951  Pater Skipper  Panama, Colombia, Venezuela  iN

Jemadia pater pater Evans, 1951  Panama, Colombia, Venezuela  iN  M T S1 S2 G Txt

Jemadia pater ortizi Orellana, [2010]  Venezuela (SE Tachira), Colombia  M OI S

Jemadia albescens Röber, 1925  Ecuador, Peru, Bolivia  M OD S1 S2 Ma L1 L2 G Txt

Jemadia hewitsonii (Mabille, 1878)  French Guiana, Venezuela (Amazonas, Bolivar), N Brazil, Peru  iN

Jemadia hewitsonii hewitsonii (Mabille, 1878)  French Guiana, Venezuela (Amazonas, Bolivar), N Brazil, Peru  M T OI OD TI S1 S2 S3 S4 HS L G

Jemadia sosia (Mabille, 1878)  Sosia Skipper  Brazil; ref.30  M T OD S Txt

Nosphistia  Mabille & Boullet, 1908  

Nosphistia zonara (Hewitson, 1866)  Zonara Skipper  Colombia, Ecuador, N Brazil (Amazonas)  iN  M T OD TI S IL

Croniades  Mabille, 1903  

Croniades pieria (Hewitson, 1857)  Fire-banded Skipper  French Guiana, N Brazil; ref.30  iN  M T OI S IL

Croniades auraria H. Druce, 1908  Peru, Bolivia; ref.30  M T S

Protelbella  O. Mielke, 1995  

Protelbella ruficauda (Hayward, 1932)  Red-tailed Skipper  S Brazil, Argentina  iN  M T

Protelbella alburna (Mabille, 1891)  Alburna Skipper    iN

Protelbella alburna alburna (Mabille, 1891)  Ecuador, Peru, Bolivia  M T S IL

Protelbella alburna brasiliensis O. Mielke, 1995  French Guiana, N & W Brazil  M OI

Parelbella  O. Mielke, 1995  

Parelbella machaon (Westwood, 1852)  Yellow-banded Skipper    iN

Parelbella machaon machaon (Westwood, 1852)  SE Brazil  M T OI S IL

Parelbella machaon seabrai (O. Mielke, 1995)  SE Brazil (RJ)  M T OI

Parelbella nigra O. Mielke, Austin & A. Warren, 2008  Mielke’s Skipper  SE Mexico (OAX, VER); ref.95  iN  M T

Parelbella macleannani (Godman & Salvin, 1893)  Macleannan’s Skipper  SE Mexico (CHIS, VER) to NW Ecuador  iN  M T S1 S2 IL I1 I2 G

Parelbella ahira (Hewitson, 1866)  Ahira Skipper    iN

Parelbella ahira ahira (Hewitson, 1866)  French Guiana, Colombia, Peru, Bolivia, Paraguay, N & W Brazil  M T OI TI S IL L

Parelbella ahira extrema (Röber, 1925)  SE Brazil, Paraguay  M T OI S IL

Parelbella peruana O. Mielke, 1995  Peruana Skipper  Peru  M T OI S

Parelbella polyzona (Latreille, [1824])  Polyzona Skipper  SE Brazil  M T IL

Microceris  E. Watson, 1893  

Microceris merops (E. Bell, 1934)  Merops Skipper  French Guiana, Venezuela (Tachira), Colombia, Peru, Bolivia, Brazil  M T S IL

Microceris patrobas (Hewitson, 1857)  Patrobas Skipper    iN

Microceris patrobas mexicana (O. Mielke, 1995)  SE Mexico (QROO, TAB, CHIS, VER, OAX), Belize  M T OI S

Microceris patrobas patrobas (Hewitson, 1857)  Guatemala to Colombia  M T OI S IL I1 I2 G

Microceris patrobas evansi (O. Mielke, 1995)  Colombia, Venezuela (Tachira)  M T OI

Microceris patrobas melanina (Mabille & Boullet, 1908)  Peru, Bolivia  M T OI

Microceris patrobas tingo (O. Mielke, 1995)  Peru, W Brazil  M T OI L

Microceris patrobas amazonica (O. Mielke, 1995)  N Brazil (Amazonas)  M T OI

Microceris blanda (Evans, 1951)  Blanda Skipper  French Guiana, Colombia, Peru, Bolivia, Paraguay, Brazil  iN  M T TI S IL

Microceris lustra (Evans, 1951)  Lustra Skipper  SW Venezuela (Tachira), Colombia, N Ecuador  M T IL

Microceris azeta (Hewitson, 1866)  Azeta Skipper    iN

Microceris azeta azeta (Hewitson, 1866)  Venezuela (Bolivar), Colombia, Peru, Bolivia, Brazil  M T IL

Microceris azeta giffordi (O. Mielke, 1995)  S Brazil  M T OI

Microceris miodesmiata (Röber, 1925)  Rober's Skipper  SE Mexico (OAX), Panama, Colombia  iN  M OI S

Microceris madeira (O. Mielke, 1995)  Madeira Skipper  W Brazil, Peru  M T OI S

Microceris patroclus (Plötz, 1879)  Patroculus Skipper  TL: Peru

Microceris patroclus patroclus (Plötz, 1879)  Panama, Ecuador, Peru, Bolivia  M T TI S HS

Microceris patroclus acala (Evans, 1951)  SW Venezuela (Tachira), Colombia  M T S IL

Microceris bicuspis (de Jong, 1983)  Bicuspis Skipper  Surinam  M T IL

Microceris rondonia (O. Mielke, 1995)  Rondonian Skipper  W Brazil, Peru  M T OI S

Microceris etna (Evans, 1951)  Etna Skipper  Trinidad, Colombia, Peru, Bolivia, Brazil  iN  M T S IL

Microceris adonis (E. Bell, 1931)  Adonis Firetip  SE Brazil, S Paraguay  M T Ab IL

Microceris iphinous (Latreille, [1824])  Yellow-barred Skipper  SE Brazil  iN

Microceris iphinous iphinous (Latreille, [1824])  SE Brazil  M T OI Ab IL

Microceris iphinous mariae (E. Bell, 1931)  SE Brazil  M T IL

Microceris iphinous molinai (Hayward, 1941)  SE Brazil; ref.30  M S IL

Microceris luteizona (Mabille, 1877)  Zona Firetip  SE Brazil; ref.30  iN  M T OD TI S HS IL

Microceris hegesippe (Mabille & Boullet, 1908)  Hegesippe Firetip  SE Brazil  M T OI S IL

Microceris theseus (E. Bell, 1934)  Theseus Skipper  Colombia, Venezuela (Tachira), Ecuador, Peru, Bolivia, N Brazil  iN  M T S IL

Microceris scylla (Ménétriés, 1855)  Scylla Firetip  E & W Mexico to Costa Rica  iN  M S IL L I1 I2 G

Microceris dulcinea (Plötz, 1879)  Dulcinea Firetip  Costa Rica, Panama, Colombia, Venezuela, Ecuador  iN  M T S IL

Microceris intersecta (Herrich-Schäffer, 1869)  Variable Firetip    iN

Microceris intersecta intersecta (Herrich-Schäffer, 1869)  French Guiana, Surinam, S Venezuela, Colombia, Peru, Ecuador, Bolivia, Brazil  M T S HS IL

Microceris intersecta peter (Evans, 1951)  NE Brazil  M T S IL

Microceris intersecta paraensis (O. Mielke, 1995)  NE Brazil  M T OI

Microceris intersecta rufitegula (O. Mielke, 1995)  SE Brazil (coastal)  M T OI

Microceris intersecta strova (Evans, 1951)  Panama, C Venezuela  M T S IL

Microceris intersecta losca (Evans, 1951)  SE Brazil (interior)  M T IL

Microceris viriditas (Skinner, 1920)  Gold-rimmed Firetip  TL: Honduras

Microceris viriditas viriditas (Skinner, 1920)  Paraguay  M T IL

Microceris viriditas chia (Evans, 1951)  N Brazil  M T S IL

Microceris viriditas boliviana (O. Mielke, 1995)  Bolivia  M T OI

Microceris lamprus (Hopffer, 1874)  Lamprus Firetip    iN

Microceris lamprus lamprus (Hopffer, 1874)  SE Brazil, S Paraguay  M T S IL

Microceris lamprus albociliata (O. Mielke, 1995)  SE Brazil  M T OI S L

Microceris variicolor (Ménétriés, 1855)  Zigzag Firetip  C & S-central Brazil  iN  M S IL G

Oxynetra  C. Felder & R. Felder, 1862  [includes Olafia as a subgenus]; ref.8

Oxynetra roscius (Hopffer, 1874)  Olaf's Firetip    iN

Oxynetra roscius roscius (Hopffer, 1874)  SE Brazil (RS), Paraguay, Argentina  M T S

Oxynetra roscius iphimedia (Plötz, 1886)  SE Brazil (SP)  M T S IL

Oxynetra roscius koikei (O. Mielke, Brockmann & C. Mielke, 2022)  Brazil (RJ); ref.30

Oxynetra aureopecta A. Warren & Grishin, 2017  Orange-chested Firetip  Mexico: Hgo., Ver., Chis.; ref.97  M T

Oxynetra hopfferi Staudinger, 1888  Hopffer's Firetip  SE Mexico (VER), Costa Rica, Panama  M T OI OD TI S G

Oxynetra stangelandi Grishin & Burns, 2013  Stangeland's Firetip  currently known only from ACG in Costa Rica; ref.73  M T1 T2 L I1 I2

Oxynetra semihyalina C. Felder & R. Felder, 1862  Felder's Firetip  Colombia, Ecuador, Peru, Bolivia  iN  M T OD S L

Oxynetra confusa Staudinger, 1888  Confusing Firetip  Venezuela (E slope Andes), Ecuador, Peru, Bolivia, [French Guiana?]  iN

Oxynetra confusa franciscana Salazar & Vargas, [2017]  Panama; ref.91

Oxynetra confusa confusa Staudinger, 1888  Venezuela (E slope Andes), Ecuador, Peru, Bolivia, [French Guiana?]  M T S

  Tribe Jerini

Jera  Lindsey, 1925  ref.8

Jera tricuspidata (Mabille, 1902)  Ecuador; ref.8  M T TI S IL

  Subfamily Pyrginae

  Tribe Achlyodini, Subtribe Pythonidina

Livida  Grishin, 2019  

Livida assecla (Mabille, 1883)  Many-spotted Blue-Skipper  SE Mexico to upper Amazon (Peru); ref.5  M T S IL

Livida grandis (Mabille, 1878)  Brazil; ref.5  iN  M T S

Pythonides  Hübner, [1819]  

Pythonides lerina (Hewitson, 1868)    iN  M T S IL

Pythonides lera Grishin, 2023  Peru; ref.153

Pythonides jovianus (Stoll, 1782)  Variable Blue-Skipper    iN

Pythonides jovianus amaryllis Staudinger, 1876  SE Mexico to Ecuador and Peru  M T S IL L I

Pythonides jovianus jovianus (Stoll, 1782)  Guatemala, Colombia, Guianas, Trinidad, upper Amazon  M S IL L

Pythonides jovianus crameri (Mabille & Boullet, 1917)  TL: Colombia  M T S IL

Pythonides jovianus fabricii W. F. Kirby, 1871  TL: Indiis  M S IL

Pythonides latemarginatus Grishin, 2023  Panama; ref.153

Lirra  Grishin, 2023  ref.149

Lirra pteras (Godman & Salvin, 1895)  Narrow-winged Blue-Skipper  Chiapas to Colombia  iN  M T S IL I

Lirra limaea (Hewitson, 1868)    iN  M T S

Gindanes  Godman & Salvin, 1895  [includes many species formerly in Pythonides]; ref.149

Gindanes brontinus Godman & Salvin, 1895  Straight-edged Skipper  TL: Nicaragua

Gindanes brontinus brontinus Godman & Salvin, 1895  E & W Mexico to Costa Rica  M T S I G

Gindanes brontinus bronta Evans, 1953  TL: Peru  M T S

Gindanes brebisson (Latreille, [1824])    iN

Gindanes brebisson panaetius Godman & Salvin, 1895  SE Mexico to Panama  M T S

Gindanes brebisson brebna Evans, 1953  TL: Bolivia  M T S

Gindanes brebisson phagesia (Hewitson, 1868)  TL: Brazil (Pa)  M T S

Gindanes brebisson brebisson (Latreille, [1824])  TL: Brazil  M T S

Gindanes variegatus Grishin, 2023  Brazil; ref.153  M S

Gindanes homer (Evans, 1953)  TL: Brazil (MT)  M T S

Gindanes nides (O. Mielke & Casagrande, 2002)  TL: Brazil (ES)  M T

Gindanes kelso (Evans, 1953)  Guyana; ref.5  M T S

Gindanes maraca (O. Mielke & Casagrande, 1992)    iN  M S

Gindanes jenmorrisae (Shuey & Ramírez, 2022)  Ecuador & Peru; ref.92  M S

Gindanes tullia (Evans, 1953)  TL: Brazil (RJ)  M T S

Gindanes herennius (Geyer, [1838])  Herennius Blue-Skipper    iN

Gindanes herennius herennius (Geyer, [1838])  Panama to Peru and lower Amazon  M T S IL L

Gindanes herennius lusorius (Mabille, 1891)  TL: Brazil (RJ)  M T S

Gindanes proxenus (Godman & Salvin, 1895)  Proxenus Blue-Skipper  SE Mexico to Venezuela and Ecuador  iN  M T S IL L I G

Gindanes parallelus (Mabille, 1898)    iN  M T S

Gindanes braga (Evans, 1953)  TL: Guyana  M T

Gindanes bora Evans, 1953    iN  M T S

Gindanes hampa (Evans, 1953)  TL: Brazil (Pa)  M T S

Gindanes rosa (Steinhauser, 1989)  Steinhauser's Blue-Skipper  S Mexico (CHIS)  iN  M T S G

Gindanes neivai (Hayward, 1940)  TL: Ecuador  M S

Gindanes mundo (H. Freeman, 1979)  Freeman's Blue-Skipper  S Mexico (OAX)  M T S G

Gindanes eminus (E. Bell, 1934)    iN

Gindanes eminus pasha (Evans, 1953)  TL: Brazil (MT)  M T S

Gindanes eminus eminus (E. Bell, 1934)  TL: Peru  M T S

Quadrus  Lindsey, 1925  [includes Ouleus and Zera as subgenera]; ref.149

Quadrus ineptus (Draudt, 1922)    iN  M T OI S G

Quadrus francesius H. Freeman, 1969  Chiapan Blue-Skipper  S Mexico (Chiapas) to Costa Rica  M T S I

Quadrus jacobus (Plötz, 1884)  TL: Brazil (RJ)  M T S

Quadrus contubernalis (Mabille, 1883)  Striped Blue-Skipper    iN

Quadrus contubernalis anicius (Godman & Salvin, 1894)  SE Mexico, Guatemala  M T S IL L G

Quadrus contubernalis contubernalis (Mabille, 1883)  SE Mexico to upper Amazon  M T S1 S2 IL L I

Quadrus deyrollei (Mabille, 1877)    iN

Quadrus deyrollei deyrollei (Mabille, 1877)  TL: French Guiana  M T S IL

Quadrus deyrollei portulana (Plötz, 1884)  TL: S. Amer.  M S

Quadrus deyrollei porta Evans, 1953  TL: Brazil (Am)  M T S IL

Quadrus lancea (Hewitson, 1868)    iN  M S IL

Quadrus cerialis (Stoll, 1782)  Common Blue-Skipper  E & W Mexico to Argentina  iN  M OI S1 S2 IL L I G

Quadrus tros Evans, 1953    iN  M T

Quadrus fanda Evans, 1953  TL: Brazil (Pa)  M T S

Quadrus u-lucida (Plötz, 1884)    iN

Quadrus u-lucida u-lucida (Plötz, 1884)  TL: Colombia  M S IL

Quadrus u-lucida mimus (Mabille & Boullet, 1917)  TL: Brazil (RS); Paraguay  M T S

Quadrus u-lucida parabus O. Mielke, 1968  TL: Brazil (MG)  M T

Quadrus lugubris (R. Felder, 1869)  Tanned Blue-Skipper  E & W Mexico to Costa Rica; ref.149  iN  M T S1 S2 IL L I G

Quadrus ophia (A. Butler, 1870)  TL: Venezuela  M T S

Quadrus truncata (Hewitson, 1870)    iN  M T S IL

Quadrus salvina (Evans, 1953)  Salvin's Skipper  E Mexico to Costa Rica  iN  M T S L

Quadrus panna (Evans, 1953)  Panna Skipper  Guatemala to Colombia  iN  M T S L

Quadrus fridericus (Geyer, 1832)    iN

Quadrus fridericus hilarina (Mabille, 1903)  TL: Brazil (Pa)  M T S

Quadrus fridericus fridericus (Geyer, 1832)  TL: Surinam  M T1 T2 S

Quadrus fridericus sinepunctis (Kaye, 1904)  TL: Trinidad  M T S

Quadrus fridericus sheldoni (Cock, 2017)  TL: Tobago

Quadrus fridericus candangus (O. Mielke, 1968)  TL: Brazil (DF)  M T S

Quadrus fridericus riona (Evans, 1953)  TL: Brazil (RJ)  M T S IL G

Quadrus fatinitza (Plötz, 1884)  TL: Colombia  M TI

Quadrus juxta (E. Bell, 1934)    iN

Quadrus juxta dampa (Evans, 1953)  TL: French Guiana  M T

Quadrus juxta juxta (E. Bell, 1934)  TL: Peru  M T S L G

Quadrus eboneus E. Bell, 1947  Eboneus Skipper  SE Mexico; ref.149  M T

Quadrus negrus (Nicolay, 1980)  Negrus Skipper  Costa Rica and Panama  M T S G

Quadrus cristatus (Steinhauser, 1989)  [could be a synonym of pescada] Colombia; ref.149  M T

Quadrus pescada (E. Bell, 1956)  Ecuador; ref.149  M T

Quadrus dilla (Evans, 1953)  TL: Ecuador

Quadrus dilla baru (Nicolay, 1973)  Costa Rica and Panama  M T OI S G

Quadrus dilla dilla (Evans, 1953)  TL: Ecuador  M T

Quadrus accedens (Mabille, 1895)    iN

Quadrus accedens noctis Lindsey, 1925  TL: Peru  M S

Quadrus accedens finda (Evans, 1953)  TL: Brazil (ES)  M T S

Quadrus accedens accedens (Mabille, 1895)  TL: Brazil (SC)  M T S

Quadrus bubaris (Godman & Salvin, 1895)  Bubaris Skipper  SE Mexico, Guatemala  M T S

Quadrus calavius (Godman & Salvin, 1895)  Calavius Skipper  SE Mexico to Panama  iN  M T S

Quadrus candidus (Steinhauser, 1989)  TL: Colombia  M T

Quadrus zora (Evans, 1953)  Ecuador; ref.149  M S

Quadrus cyrna (Mabille, 1895)  Hidden-yellow Skipper  Chiapas to Panama  M T S I

Quadrus difficilis (Weeks, 1901)  Difficult Skipper  Chiapas, Guianas and upper Amazon  iN  M T OI S G

Quadrus zera (A. Butler, 1870)  Zera Skipper  Costa Rica to Venezuela, Ecuador and S Brazil  M T S IL L G

Quadrus teresa (Steinhauser, 1989)  TL: Brazil (ES)  M S HS

Quadrus belti (Godman & Salvin, 1894)  Belti Skipper  Chiapas to Nicaragua  M T OI S IL G

Quadrus nolckeni (Mabille, 1891)  Nolcken's Skipper  Panama to Bolivia (not in Mexico)  iN  M T S IL L

Quadrus phila (Godman & Salvin, 1894)  Phila Skipper  Panama to Peru and Bolivia  iN  M T OI S IL L

Quadrus hosta (Evans, 1953)  Hosta Skipper  Chiapas to Colombia  iN  M T S I

Quadrus hyacinthinus (Mabille, 1877)  Bruised Skipper  E & W Mexico to Colombia  iN  M T S IL L

Quadrus gellius (Mabille, 1903)  S Brazil & Argentina; ref.149  M S

Quadrus tetrastigma (Sepp, [1847])  Tetrastigma Skipper  Chiapas to Guianas, Trinidad, upper Amazon  iN  M T S IL G

Quadrus menedemus (Godman & Salvin, 1894)  ref.149  M T IL

Quadrus servius (Plötz, 1884)  [= erisichthon] Brazil; ref.149  M T TI S IL

Quadrus vivax Grishin, 2023  S Brazil; ref.149  M S

Haemactis  Mabille, 1903  

Haemactis sanguinalis (Westwood, 1852)  Ecuador; ref.118  iN  M T S

Haemactis albamarita Austin, 1994  Brazil (Rd); ref.118  M T S

Haemactis [n. sp.] N/A, N/A  Red-washed Skipper  [likely conspecific with pyrrhosphenus] Nicaragua and Costa Rica; ref.118  M S L

Haemactis pyrrhosphenus Lindsey, 1919  Colombia; ref.118  M S

Milanion  Godman & Salvin, 1895  [includes Atarnes and Eburuncus as subgenera]; ref.149

Milanion sallei (C. Felder & R. Felder, 1867)  Orange-spotted Skipper  E & W Mexico to Colombia  iN  M T S1 S2 L I G

Milanion hierax (Hopffer, 1874)  White-veined Tuskyhook  Peru; ref.64  M T S IL G

Milanion unifasciata (C. Felder & R. Felder, 1867)  Single-banded Tuskyhook  SE Mexico to Ecuador and Peru; ref.64  iN  M T S1 S2 IL L I G

Milanion virga Grishin, 2023  [= Evans' "hemestinus"] Tropical S. Am.; ref.153  M S

Milanion hemes (Cramer, 1777)  Hemes Skipper  (= clito); ref.153  iN

Milanion hemes hemes (Cramer, 1777)  (= clito) Costa Rica to Guianas and Trinidad; ref.153  M T S L Txt

Milanion hemes pemba Evans, 1953  TL: Peru  M T S

Milanion hemes memba Evans, 1953  TL: Brazil (MT)  M T S

Milanion latior Mabille & Boullet, 1917  Guatemala to Ecuador; ref.153  M T

Milanion marciana Godman & Salvin, 1895  Marciana Skipper  Panama  iN  M T S1 S2 I G

Milanion leucaspis (Mabille, 1878)    iN  M T S G

Milanion pilumnus Mabille & Boullet, 1917    iN

Milanion pilumnus albidior Mabille & Boullet, 1917  TL: Venezuela  M T S

Milanion pilumnus pilumnus Mabille & Boullet, 1917  (= hemestinus); ref.153  M T S G

Milanion pilta Evans, 1953  ref.153  M T S

Milanion alaricus (Plötz, 1884)    iN  M T S G

Milanion furvus Grishin, 2023  Panama; ref.153

Milanion laricus Grishin, 2023  Ecuador; ref.153

Milanion cramba Evans, 1953    iN  M T S

Paramimus  Hübner, [1819]  

Paramimus stigma (C. Felder & R. Felder, 1867)    iN

Paramimus stigma stanna Evans, 1953  Panama, (Colombia)  M T S L

Paramimus stigma stigma (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Paramimus scurra (Hübner, [1809])    iN

Paramimus scurra scurra (Hübner, [1809])  TL: ?  M T S G

Paramimus scurra herberti Godman & Salvin, 1896  TL: Brazil (MT)  M T S

Charidia  Mabille, 1903  

Charidia empolaeus (Westwood, 1852)  TL: ?  M T S

Charidia lucaria (Hewitson, 1868)    iN  M T S

Charidia ronda Grishin, 2023  Brazil (Ro); ref.153  M S

Charidia pilea Evans, 1953  Colombia; ref.153  M T S

Charidia pocus Evans, 1953  Peru; ref.153

Charidia pocus pocus Evans, 1953  TL: Peru  M T S

Charidia pocus mayo Evans, 1953  TL: Peru  M T

  Tribe Achlyodini, Subtribe Achlyodina

Cabirus  Hübner, [1819]  

Cabirus procas (Cramer, 1777)  The Guianas & N Brazil  iN  M OI OD S IL

Cabirus junta Evans, 1952  Peru, Bolivia, Brazil; ref.140  M T S1 S2 IL

Cabirus purda Evans, 1952  Ecuador & Peru; ref.140  M T S L

Pseudodrephalys  Burns, 1999  

Pseudodrephalys sohni Burns, 1999    iN  M T S

Pseudodrephalys tinas Grishin, 2023  NE Peru; ref.153  M T

Pseudodrephalys atinas (Mabille, 1888)  TL: [Peru]  M T OI S G

Pseudodrephalys hypargus (Mabille, 1891)  TL: Brazil (Am)  M T S

Pseudodrephalys argus Grishin, 2023  The Guianas; ref.153  M T

Eantis  Boisduval, 1836  

Eantis pallida (R. Felder, 1869)  Pale Sicklewing  [= selva], W Mexico & S Texas (as stray) to Bolivia; ref.5  iN  M T S L1 L2 I1 I2 G

Eantis tamenund (W. H. Edwards, 1871)  Northern Sicklewing  C & S Texas (further N as strays) to N Belize and Guatemala  iN  M T S L1 L2 L3 I1 I2 H G

Eantis thraso (Hübner, [1807])  Southern Sicklewing  SW Chiapas, S Belize, Guatemala to Argentina  iN  M S1 S2 IL L I1 I2 G

Eantis mithridates (Fabricius, 1793)  Jamaican Sicklewing  Jamaica  M T S L

Eantis minor W. Comstock, 1944  Lesser Sicklewing  Puerto Rico to Dominica and Guadeloupe  iN  M T S L

Eantis papinianus (Poey, 1832)  Cuban Sicklewing  [= sagra], Cuba, Hispaniola  iN  M T S L I

Eantis munroei E. Bell, 1956  Munroe's Sicklewing  Cuba  M T OD S G

Eantis minna Evans, 1953  TL: Brazil (Ba)  M T S HS G

Eantis later (Mabille, 1891)  TL: Peru  M T S

Eantis haber (Mabille, 1891)  TL: Peru  M T S

Eantis tosta (Evans, 1953)  Peru; ref.5  M T OD S G

Eantis pseudospeculum (Siewert, O. Mielke & Casagrande, 2015)  Brazil; ref.31

Aethilla  Hewitson, 1868  

Aethilla gigas (Mabille, 1877)  TL: Peru  M T S G

Aethilla echina Hewitson, 1870  Echina Skipper    iN

Aethilla echina echina Hewitson, 1870  SE Mexico to S Brazil  M T S IL L G

Aethilla echina coracina A. Butler, 1870  TL: Brazil (SP)  M T S

Aethilla melas Plötz, 1882  TL: Brazil (RJ)  M OD S G Txt

Aethilla eleusinia Hewitson, 1868    iN  M T S L G

Aethilla memmius A. Butler, 1870    iN  M T OI S G Txt

Aethilla epicra Hewitson, 1870  TL: Ecuador  M T S HS G

Aethilla lavochrea A. Butler, 1872  Yellow-rimmed Skipper  E & W Mexico to Colombia  iN  M T S1 S2 S3 L I

Aethilla chiapa H. Freeman, 1969  Highlands Skipper  Oaxaca, Chiapas, Guatemala  M T OD S G

Achlyodes  Hübner, [1819]  

Achlyodes busirus (Cramer, 1779)  Giant Sicklewing    iN

Achlyodes busirus heros Ehrmann, 1909  E & W Mexico to Bolivia  M T S1 S2 HS IL L I1 I2 Txt

Achlyodes busirus busirus (Cramer, 1779)  TL: Surinam  M OI OD S G

Achlyodes busirus negro (Kaye, 1921)  TL: Trinidad  M T S G

Achlyodes busirus rioja Evans, 1953  TL: Brazil (RJ)  M T S IL G

Achlyodes calvus Grishin, 2023  S Brazil & Paraguay; ref.153  M HS

Doberes  Godman & Salvin, 1895  

Doberes anticus (Plötz, 1884)  Dark Doberes  SE Mexico (VER, OAX, CHIS)  iN

Doberes anticus anticus (Plötz, 1884)  TL: Mexico  M S L G

Doberes anticus Pale segregate ,  S Mexico (Chiapas) to Costa Rica  M S I

Doberes sobrinus (Godman & Salvin, 1895)  West-Mexican Doberes  W Mexico (Guerrero)  M T S

Doberes hewitsonius (Reakirt, [1867])  Pale Doberes  S Mexico  iN  M T S L G

Spioniades  Hübner, [1819]  

Spioniades artemis Grishin, 2023  Costa Rica & Panama; ref.153

Spioniades artemides (Stoll, 1782)  Artemides Skipper  Panama to S Brazil  iN  M S L I G

Spioniades artemidoides Grishin, 2023  S Brazil; ref.153  M S

Spioniades abbreviata (Mabille, 1888)  White-tipped Skipper  Nicaragua to Colombia  iN  M T OI S1 S2 I

Spioniades anta Evans, 1953  S Peru & Bolivia; ref.153  M T S

Spioniades libethra (Hewitson, 1868)  TL: Brazil (Am)  M T OI S1 S2 S3

Eracon  Godman & Salvin, 1894  

Eracon sarahburnsae Grishin, 2014  Sarah's Skipper  Costa Rica & Panama; ref.75  M T1 T2 T3 T4 S I

Eracon clinias (Mabille, 1878)  Clinias Skipper  Costa Rica to upper and lower Amazon  iN  M T S1 S2 IL L G

Eracon pebana Evans, 1953  Pebana Skipper    iN  M T S IL G

Eracon asymmetrica Austin, 1997  Asymmetric Skipper  TL: Brazil (Rd)  M T OI G

Eracon biternata (Mabille, 1889)  Biternata Skipper  Costa Rica and Panama  M T OI TI S G

Eracon paulinus (Stoll, 1782)  Tear-drop Skipper  Costa Rica to Guiana and upper Amazon  iN  M T OI OD S1 S2 S3 S4 IL L I G Txt

Eracon lachesis (Dyar, 1918)  Lachesis Skipper  S Mexico to Panama  M T S1 S2 S3 L I

Mimia  Evans, 1953  

Mimia chiapaensis H. Freeman, 1969  Chiapan Skipper  CHIS  M T S I G

Mimia phidyle (Godman & Salvin, 1894)  Phidyle Skipper    iN

Mimia phidyle phidyle (Godman & Salvin, 1894)  Mexico (Chis) to Panama  M T S L

Mimia phidyle tatamaensis Salazar & Vargas, [2017]  Colombia; ref.91

Mimia pazana Evans, 1953  Bolivia; ref.149  M T S G

Xispia  Lindsey, 1925  

Xispia quadrata (Mabille, 1889)    iN  M T OI S G

Xispia satyrus (Jörgensen, 1935)    iN  M S IL

Myrinia  Evans, 1953  

Myrinia binoculus (Möschler, 1877)  TL: Surinam  M T S

Myrinia orieca Grishin, 2023  E Ecuador; ref.153

Myrinia raymundo H. Freeman, 1979  Raymundo's Skipper  SE Mexico (TAB, CHIS)  iN  M T S G

Myrinia aragua Grishin, 2023  Venezuela; ref.153

Myrinia laddeyi (E. Bell, 1942)    iN  M T

Myrinia maculosa Grishin, 2023  Guatemala; ref.153

Myrinia catua O. Mielke, 1968    iN  M T S

Myrinia myris (Mabille, 1898)  Myris Skipper  Campeche, Belize, Nicaragua to Guianas, Bolivia, S Brazil  iN  M T S

Myrinia santa Evans, 1953    iN

Myrinia santa monka Evans, 1953  TL: Brazil (Pa)  M T S

Myrinia santa santa Evans, 1953  TL: Brazil (ES)  M T

Myrinia manchada Grishin, 2023  The Guianas; ref.153  M T S

Grais  Godman & Salvin, 1894  

Grais juncta Evans, 1953  'Jamaican' Hermit Skipper  Jamaica; ref.77  M T S

Grais stigmaticus (Mabille, 1883)  Hermit Skipper  C & S TX (as stray), E & W Mexico to Argentina  iN  M T S1 S2 L I G

Morvina  Evans, 1953  

Morvina caecus (Austin, 2008)  Blind Skipper  Brazil (Rondonia)  M T S

Morvina falisca (Hewitson, 1878)    iN

Morvina falisca falisca (Hewitson, 1878)  TL: French Guiana  M T S

Morvina falisca falia Evans, 1953  TL: Peru  M T S

Morvina fissimacula (Mabille, 1878)    iN

Morvina fissimacula pelarge (Godman & Salvin, 1894)  Nicaragua to Panama  M T S I G

Morvina fissimacula lenia Evans, 1953  TL: Colombia  M T

Morvina fissimacula rema Evans, 1953  TL: Brazil (Pa)  M T S

Morvina fissimacula fissimacula (Mabille, 1878)  TL: Brazil  M T S

Morvina morvus (Plötz, 1884)    iN

Morvina morvus morvus (Plötz, 1884)  TL: Brazil  M S

Morvina morvus para Evans, 1953  TL: Brazil (Pa)  M T S

Morvina morvus cyclopa Evans, 1953  TL: Bolivia  M T S

  Tribe Carcharodini, Subtribe Cyclosemiina

Cyclosemia  Mabille, 1878  

Cyclosemia earina (Hewitson, 1878)  TL: Brazil (Pa)  M T S

Cyclosemia anastomosis Mabille, 1878  Northern Eyed-Skipper  E Mexico to Ecuador  iN  M T S1 S2 L I G

Cyclosemia lathaea (Hewitson, 1878)    iN  M T S

Cyclosemia leppa Evans, 1953  Leppa Eyed-Skipper  Guatemala to Bolivia  iN  M T S IL

Cyclosemia lyrcaea (Hewitson, 1878)  Lyrcaea Eyed-Skipper  Panama to SE Brazil  iN  M T S

Cyclosemia pedro R. Williams & E. Bell, 1940  Pedro Eyed-Skipper  Honduras to N Brazil ("Amazons")  iN  M T G

Cyclosemia caerulea Austin & DeVries, 2001  Caerulea Eyed-Skipper  Costa Rica  M T S

Cyclosemia subcaerulea Schaus, 1913  Subcaerulea Eyed-Skipper  Costa Rica to Ecuador  iN  M T S1 S2 I

Cyclosemia herennius (Stoll, 1782)    iN  M S

Cyclosemia elelea (Hewitson, 1878)  TL: French Guiana  M T S

  Tribe Carcharodini, Subtribe Ilianina

Cornuphallus  Austin, 1997  

Cornuphallus albafascia Austin, 1997  White-banded Hornydick    iN  M T OI G

Cornuphallus onoribo (Möschler, 1883)  Onoribo Hornydick  TL: Surinam  M T S

Cornuphallus problematica (R. Williams & E. Bell, 1940)  Tricky Hornydick  TL: Argentina  M T G

Iliana  E. Bell, 1937  ref.119,19

Iliana peruvianus (Mabille, 1883)  TL: [Peru]  M T S G

Iliana bolivianus O. Mielke & Casagrande, 2016  Bolivia; ref.119  M T

Iliana miersi O. Mielke & Casagrande, 2016  Brazil; ref.119  M T

Iliana remus E. Bell, 1937  TL: Peru  M T OD S G

Iliana sapasoa (Nicolay, 1973)    iN  M T OD S G

Iliana romulus E. Bell, 1937    iN  M T OD S G

Iliana nirtoni O. Mielke & Casagrande, 2016  S Brazil; ref.119

Iliana taurus (Evans, 1953)  TL: Peru  M T OD S G

Iliana gorgus (E. Bell, 1937)  Gorgus Skipper  Belize, Costa Rica; ref.149  M T OD S I G

Tiana  Grishin, 2019  

Tiana platypterus (Mabille, 1895)  Platypterus Skipper  SE Mexico to Panama; ref.7  M T S I G

Tiana niger (R. Williams & E. Bell, 1940)  Colombian Skipper  Costa Rica to Colombia; ref.7  M T OD S I1 I2 G

  Tribe Carcharodini, Subtribe Nisoniadina

Arteurotia  A. Butler & H. Druce, 1872  

Arteurotia artistella Grishin, 2023  Artistarred Skipper  S TX to Mexico; ref.19  M S L

Arteurotia tractipennis A. Butler & H. Druce, 1872  Starred Skipper    iN

Arteurotia tractipennis tractipennis A. Butler & H. Druce, 1872  [Mexico?] El Salvador to Surinam and Bolivia; ref.19  M T S IL I G

Arteurotia tractipennis contractipennis Mabille & Boullet, 1916  Venezuela; ref.19  M T

Conognathus  C. Felder & R. Felder, 1862  

Conognathus platon C. Felder & R. Felder, 1862    iN  M T OI S L

Ocella  Evans, 1953  

Ocella albata (Mabille, 1888)    iN  M T OI S

Ocella diophthalma (Plötz, 1884)  TL: ?  M S

Ocella monophthalma (Plötz, 1884)  Brazil; ref.16  iN  M T S

Sophista  Plötz, 1879  [includes Pachyneuria and Austinus as subgenera]; ref.149

Sophista heroica (Evans, 1953)  Colombia; ref.7  M T OD G

Sophista heros (Mabille & Boullet, 1917)  Guianas; ref.119  iN  M T OD S L G

Sophista obscura (Mabille, 1888)  TL: Peru  M T OI TI S

Sophista jaguar (Evans, 1953)  TL: Peru  M T S

Sophista milleri (Steinhauser, 1989)  TL: Peru  M T

Sophista lineatopunctata (Mabille & Boullet, 1917)    iN

Sophista lineatopunctata belema (Evans, 1953)  TL: Brazil (Pa)  M T

Sophista lineatopunctata lineatopunctata (Mabille & Boullet, 1917)  TL: Peru  M T S

Sophista damon (E. Bell, 1937)  TL: Brazil (Am)  M T S

Sophista duidae (E. Bell, 1932)    iN

Sophista duidae duidae (E. Bell, 1932)  TL: Venezuela  M T S

Sophista duidae pozuza (Evans, 1953)  TL: Peru  M T

Sophista inops (Mabille, 1877)    iN  M T TI S

Sophista helena (Hayward, 1939)  TL: Ecuador  M S

Sophista plinius Plötz, 1882  Brazil; ref.33  iN  M T S

Sophista aristoteles (Westwood, 1852)    iN  M T S IL L

Sophista latifasciata (Spitz, 1930)    iN  M S

Sophista matto Evans, 1953  Brazil; ref.33  M S

Sophista spectrum Siewert, Zacca, O. Mielke & Casagrande, 2016  Brazil; ref.33

Polyctor  Evans, 1953  

Polyctor cleta Evans, 1953  Cleta Tufted-Skipper  E & W Mexico to Costa Rica  iN  M T S1 S2 L I G

Polyctor polyctor (Prittwitz, 1868)  Polyctor Tufted-Skipper  Costa Rica to Argentina  iN  M T S1 S2 IL L I G

Polyctor dagua Evans, 1953  Colombia; ref.149  M T

Polyctor enops (Godman & Salvin, 1894)  Enops Tufted-Skipper  E & S Mexico to Costa Rica  iN  M T S1 S2 L I

Polyctor lamperus Grishin, 2023  Panama; ref.153

Polyctor fera (Weeks, 1901)    iN

Polyctor fera fera (Weeks, 1901)  TL: Bolivia  M T S

Polyctor fera falla Evans, 1953  TL: Venezuela  M T

Polyctor tensa Evans, 1953    iN  M T S

Viola  Evans, 1953  

Viola alicus (Schaus, 1902)    iN  M T S

Viola minor (Hayward, 1933)    iN  M S

Viola olla Evans, 1953  TL: Brazil (Am)  M T S

Viola violella (Mabille, 1898)    iN  M T S L

Viola egra Evans, 1953    iN  M T S

Viola kuma (E. Bell, 1942)  [= dagamba] Venezuela, Guyana; ref.20  M T

Viuria  Grishin, 2019  formerly in Pachyneuria, but sister to Viola; ref.5,148

Viuria licisca (Plötz, 1882)  Immaculate Tufted-Skipper  E & W Mexico to Colombia and Venezuela; ref.5,148  iN  M T S1 S2 L I G

Viuria lista (Evans, 1953)  French Guiana & Brazil (Ro); ref.5,148  M T S

Viuria herophile (Hayward, 1940)  Ecuador, Peru, Brazil (Ac); ref.5,148  M S

Viuria acadia Siewert, Lemes, O. Mielke & Casagrande, 2023  Ecuador & Brazil (Ac, Ro, Mt); ref.148

Viuria inanna Siewert, Lemes, O. Mielke & Casagrande, 2023  E & SE Brazil & Paraguay; ref.148

Pellicia  Herrich-Schäffer, 1870  [includes Mictris as a subgenus]; ref.149

Pellicia klugi R. Williams & E. Bell, 1939  TL: Peru  M T S

Pellicia santana R. Williams & E. Bell, 1939  TL: Peru  M T

Pellicia simulator R. Williams & E. Bell, 1939  TL: Peru  M T

Pellicia arina Evans, 1953  Glazed Tufted-Skipper  S TX to Panama  iN  M T S1 S2 L I G

Pellicia costimacula Herrich-Schäffer, 1870  Frosted Tufted-Skipper    iN

Pellicia costimacula costimacula Herrich-Schäffer, 1870  Panama to S Brazil and Paraguay  M T S G

Pellicia costimacula litoralis Biezanko & O. Mielke, 1973  TL: Brazil (RS)  M T S

Pellicia tyana Plötz, 1882    iN

Pellicia tyana toza Evans, 1953  TL: Colombia  M T

Pellicia tyana tyana Plötz, 1882  TL: S. Amer.  M T S

Pellicia vecina Schaus, 1902    iN

Pellicia vecina naja Steinhauser, 1989  TL: Peru  M T

Pellicia vecina vecina Schaus, 1902  TL: Brazil (RJ)  M T S

Pellicia najoides Hayward, 1933  TL: Argentina  M S

Pellicia angra Evans, 1953  Rare Tufted-Skipper  TL: Mexico (Ver)

Pellicia angra angra Evans, 1953  SE Mexico (VER) to Panama  M T S G

Pellicia angra axina Evans, 1953  TL: Ecuador  M T

Pellicia tonga Evans, 1953  Tonga Tufted-Skipper  Panama, Colombia, Venezuela  M T S

Pellicia trax Evans, 1953  TL: Ecuador  M T S

Pellicia theon Plötz, 1882  TL: S. Amer.  M T TI S

Pellicia nema R. Williams & E. Bell, 1939  ref.149  M T

Pellicia ranta Evans, 1953    iN

Pellicia ranta ranta Evans, 1953  TL: Brazil (RJ)  M T

Pellicia ranta rancida Evans, 1953  TL: Paraguay  M T S

Pellicia bilinea Mabille, 1889  Central America; ref.149  M T OI

Pellicia dimidiata Herrich-Schäffer, 1870  Morning Glory Tufted-Skipper    iN

Pellicia dimidiata dimidiata Herrich-Schäffer, 1870  W Mexico and S TX (as stray) to Guianas, Trinidad, up. Amazon  M T S1 S2 L I G

Pellicia dimidiata meno (Mabille, 1889)  TL: Panama  M T OI S

Pellicia dimidiata zamia Plötz, 1882  TL: S. Amer.  M T TI S

Pellicia hersilia Hayward, 1939    iN  M T S

Pellicia demetrius (Plötz, 1882)  TL: Brazil  M T TI

Pellicia alta Steinhauser, 2007  TL: Peru

Pellicia crispus Herrich-Schäffer, 1870    iN

Pellicia crispus caerula (Mabille, 1877)  CHIS to Ecuador  M T S1 S2 L I1 I2 G

Pellicia crispus crispus Herrich-Schäffer, 1870  TL: Venezuela  M T S

Nisoniades  Hübner, [1819]  

Nisoniades godma Evans, 1953  Godma Tufted-Skipper  E & S Mexico to Costa Rica  iN  M T S1 S2 L I G

Nisoniades panama Evans, 1953  Panamanian Tufted-Skipper  Panama to Ecuador  M T S

Nisoniades suprapanama Steinhauser, 1989  TL: Colombia  M T

Nisoniades benda Evans, 1953  TL: Ecuador  M T S

Nisoniades bessus (Möschler, 1877)    iN  M T S

Nisoniades brazia Evans, 1953    iN  M T S

Nisoniades hecale (Hayward, 1940)  TL: Ecuador  M S

Nisoniades cauca Evans, 1953    iN  M T

Nisoniades remo Evans, 1953  TL: Peru  M T

Nisoniades guianae (R. Williams & E. Bell, 1939)  TL: French Guiana  M T

Nisoniades montana (R. Williams & E. Bell, 1939)  TL: Venezuela  M T

Nisoniades coca Steinhauser, 1989  TL: Ecuador  M T

Nisoniades lata Steinhauser, 1989  TL: Ecuador  M T S

Nisoniades bromias (Godman & Salvin, 1894)  Purplish Tufted-Skipper  W Mexico & S Texas (as stray) to Panama; ref.11  iN  M T S1 S2 L I H G

Nisoniades lutum Grishin, 2023  W Mexico; ref.153

Nisoniades rubescens (Möschler, 1877)  Purplish Tufted-Skipper  Panama to lower Amazon and Bolivia; ref.11  iN  M T S G

Nisoniades mimas (Cramer, 1775)    iN  M T S

Nisoniades bipuncta (Schaus, 1902)    iN  M T S

Nisoniades inca (Lindsey, 1925)  [= brunneata]; ref.16  iN  M T

Nisoniades ephora (Herrich-Schäffer, 1870)  Ephora Tufted-Skipper  E & W Mexico to Guianas, Trinidad, Ecuador and Peru  iN  M T S L I G

Nisoniades torta Steinhauser, 1989  Torta Tufted-Skipper  Costa Rica to Panama  M T S G

Nisoniades borra (E. Bell, 1947)  TL: Colombia  M T

Nisoniades indistincta (R. Williams & E. Bell, 1939)  TL: Colombia  M T

Nisoniades laurentina (R. Williams & E. Bell, 1939)  Dark Tufted-Skipper  SE Mexico (VER) to Guianas and Trinidad  M T S

Nisoniades evansi Steinhauser, 1989  TL: Ecuador  M T S

Nisoniades supra Steinhauser, 1989  TL: Peru  M T

Nisoniades maura (Mabille & Boullet, 1917)    iN  M T S

Nisoniades hora (Hayward, 1939)    iN  M T

Nisoniades criton (Mabille, 1898)    iN  M T

Nisoniades hesperia (Hayward, 1939)  TL: Paraguay

Nisoniades castolus (Hewitson, 1878)  Castolus Tufted-Skipper  SE Mexico (VER) to S Brazil  iN  M T S1 S2 I G

Nisoniades haywardi (R. Williams & E. Bell, 1939)    iN  M T S

Nisoniades rimana (E. Bell, 1942)  TL: Venezuela  M T S

Nisoniades macarius (Herrich-Schäffer, 1870)  Macarius Tufted-Skipper  (doubtful for Mexico, Honduras), Panama to Argentina  iN  M T TI S G

Nisoniades nyctineme (A. Butler, 1877)  TL: Brazil (Pa)  M T

  Tribe Carcharodini, Subtribe Burcina

Burca  E. Bell & W. Comstock, 1948  

Burca braco (Herrich-Schäffer, 1865)  Braco Skipper    iN

Burca braco braco (Herrich-Schäffer, 1865)  Cuba  M T S L

Burca braco castigata Rindge, 1955  Bahamas; Great Bahama Bank  M T

Burca stillmanni E. Bell & W. Comstock, 1948  Stillman's Skipper  Hispaniola  M T S

Burca hispaniolae E. Bell & W. Comstock, 1948  Hispaniolan Skipper  Hispaniola  iN  M T S

Burca concolor (Herrich-Schäffer, 1865)  Concolorous Skipper    iN

Burca concolor concolor (Herrich-Schäffer, 1865)  Cuba  M T S

Burca concolor atrata Rindge, 1955  Bahamas; Great Bahama Bank  M T S

Burca cubensis (Skinner, 1913)  Cuban Skipper  Cuba  M T L

  Tribe Carcharodini, Subtribe Pholisorina

Gorgopas  Godman & Salvin, 1894  

Gorgopas trochilus (Hopffer, 1874)    iN  M T S

Gorgopas chlorocephala (Herrich-Schäffer, 1870)  Green-headed Sootywing    iN

Gorgopas chlorocephala chlorocephala (Herrich-Schäffer, 1870)  Nicaragua to Bolivia  M T S L G

Gorgopas chlorocephala sneiderni E. Bell, 1947  TL: Colombia  M T

Gorgopas gutta Evans, 1953    iN  M T S L

Gorgopas agylla (Mabille, 1898)  TL: Bolivia  M T S L

Gorgopas petale (Mabille, 1888)    iN  M T OI S

Gorgopas extensa (Mabille, 1891)  Peru; ref.13  iN  M T S L

Incisus  Grishin, 2019  

Incisus incisus (Mabille, 1878)  ref.5  iN  M T S IL L G

Incisus fasciatus (Hayward, 1933)  ref.5  iN  M S

Incisus angulata (E. Bell, 1937)  TL: Brazil (MG)  M T

Clytius  Grishin, 2019  

Clytius clytius (Godman & Salvin, 1897)  Mottled Sootywing  SE AZ & S TX (rare strays) to Honduras; ref.5  iN  M T S L G

Clytius shola (Evans, 1953)  Venezuela; ref.13  M T

Perus  Grishin, 2019  

Perus menuda (Weeks, 1902)  Bolivia; ref.5

Perus parvus (Steinhauser & Austin, 1993)  Parvus Sootywing  Costa Rica; ref.13  M T

Perus manx (Evans, 1953)  Colombia; ref.13  M T

Perus minor (Schaus, 1902)  Peru; ref.5  iN  M T S IL G

Perus coecatus (Mabille, 1891)  S Brazil; ref.5  iN  M T S

Perus mossi (Evans, 1953)  Peru; ref.5  iN  M T S

Perus cordillerae (Lindsey, 1925)  Peru; ref.5  iN  M S

Perus narycus (Mabille, 1889)  Peru; ref.13  iN  M T OI S IL

Bolla  Mabille, 1903  

Bolla cybele Evans, 1953  Veracruzan Sootywing  SE Mexico (VER)  M T IL

Bolla cylindus (Godman & Salvin, 1896)  Checkered Sootywing  SE Mexico (VER) to Costa Rica  iN  M T S G

Bolla sonda Evans, 1953  TL: Mexico (Ver)  M T

Bolla cyclops (Mabille, 1877)  Cyclops Sootywing  [= sonda], SE Mexico, Guatemala, Colombia  iN  M T S

Bolla aplica (E. Bell, 1937)  [not eusebius] W Mexico; ref.149  M T

Bolla sodalis Schaus, 1913  [not imbras]; ref.149  M T

Bolla solitaria Steinhauser, 1991  Solitary Sootywing  S Mexico (OAX)  iN  M T S G

Bolla saletas (Godman & Salvin, 1896)  Saletas Sootywing  Guatemala to El Salvador  M T S G

Bolla tepeca (E. Bell, 1942)  Grizzled Sootywing  C & S Mexico (mostly DF to OAX); ref.143  iN  M T S L

Bolla fenestra Steinhauser, 1991  Oaxacan Sootywing  S Mexico (OAX)  iN  M T S L Ha G

Bolla nigerrima Mabille & Boullet, 1917    iN  M T

Bolla catharina (E. Bell, 1937)    iN  M T S

Bolla morona (E. Bell, 1940)    iN

Bolla morona lorea (E. Bell, 1947)  TL: Colombia  M T

Bolla morona morona (E. Bell, 1940)  TL: Peru  M T S

Bolla cupreiceps (Mabille, 1891)  Copper-headed Sootywing  E & S Mexico to Bolivia and S Brazil  iN  M T S IL L G

Bolla imbras (Godman & Salvin, 1896)  Rounded Sootywing  [= pullata, = subapicatus], E & S Mexico to Colombia and Venezuela; ref.149  iN  M T S1 S2 L G

Bolla phylo Mabille, 1903    iN  M T

Bolla mancoi (Lindsey, 1925)  TL: Peru  M S

Bolla atahuallpai (Lindsey, 1925)  Atahuallpai Sootywing  Panama to Argentina  iN  M OI S IL G

Bolla evippe (Godman & Salvin, 1896)  Rough-tipped Sootywing  E & W Mexico, Guatemala, Nicaragua  iN  M T S G

Bolla orsines (Godman & Salvin, 1896)  Hook-tipped Sootywing  E & W Mexico to El Salvador  iN  M T S G

Bolla eusebius (Plötz, 1884)  Spatulate Sootywing  E & W Mexico to Costa Rica  iN  M T S

Bolla chilpancingo (E. Bell, 1937)  [= Evans' "subapicatus"] W & S Mexico, Guatemala; ref.149  M T

Bolla guerra Evans, 1953  Guerrero Sootywing  SW Mexico to Nicaragua  iN  M T S L

Bolla oriza Evans, 1953  Orizaba Sootywing  SE Mexico (VER, CHIS?)  M T S

Bolla brennus (Godman & Salvin, 1896)  Obscure Sootywing    iN

Bolla brennus brennus (Godman & Salvin, 1896)  S Texas (one-time stray), E Mexico to Panama  M T S G

Bolla brennus vexta Evans, 1953  TL: Colombia  M T

Bolla dorsolaciniae Steinhauser, 1989  TL: Colombia  M T

Bolla antha Evans, 1953  TL: Colombia  M T

Bolla giselus (Mabille, 1883)  TL: Colombia

Bolla giselus tornea Evans, 1953  TL: Colombia  M T

Bolla giselus giselus (Mabille, 1883)  TL: Colombia  M T S

Bolla boliviensis (E. Bell, 1937)  Bolivian Sootywing  Guatemala to Bolivia  iN  M T S IL G

Bolla zorilla (Plötz, 1886)  Zorilla Sootywing  SE Mexico (VER, OAX) to Bolivia  iN  M S G

Bolla vena Grishin, 2023  Venezuela; ref.153

Bolla madrea (R. Williams & E. Bell, 1940)  Peru; ref.149  M T

Bolla hazelae (Hayward, 1940)  Ecuador; ref.149  M T

Bolla evemerus (Godman & Salvin, 1896)  Evemerus Sootywing  Nicaragua to Panama; ref.149  M T S I G

Bolla chlora (Evans, 1953)  Bolivia; ref.149  iN  M T S

Bolla astra (R. Williams & E. Bell, 1940)  Peru; ref.149  iN  M T S

Bolla balsa (E. Bell, 1937)  Peru; ref.149  M T

Bolla tridentis (Steinhauser, 1989)  Colombia; ref.149  M T

Bolla esmeraldus (L. Miller, 1966)  Esmeraldus Sootywing  E Costa Rica; ref.149  M T G

Bolla chlorocephala (Latreille, [1824])  SE & S Brazil; ref.149  iN  M T S

Bolla incanus (E. Bell, 1932)  SE & S Brazil; ref.149  M T S

Staphylus  Godman & Salvin, 1896  [includes Hesperopsis as a subgenus]; ref.149

Staphylus cartagoa (R. Williams & E. Bell, 1940)  Cartagoa Sootywing  Costa Rica, El Salvador  M T G

Staphylus vincula Plötz, 1886  Mountain Sootywing  [= iguala, what was called "vincula" is opites] E & W MX to Venez.; ref.14  iN  M T TI S1 S2 L G

Staphylus opites (Godman & Salvin, 1896)  Opites Sootywing  [formerly called "vincula"]; ref.14  iN  M T

Staphylus ceos (W. H. Edwards, 1882)  Golden-headed Sootywing  far S CA, S NV (rare), S AZ, S NM, W TX (El Paso), to QRO  iN  M T S HS L1 L2 I G

Staphylus ecos Grishin, 2022  Texas Sootywing  W, C & S Texas and Mexico (Coah); ref.14  M T S L I1 I2

Staphylus vula Grishin, 2023  Mexico; ref.153

Staphylus vulga Grishin, 2023  Panama; ref.153

Staphylus vulgata (Möschler, 1879)  Golden-snouted Sootywing  E & W Mexico to Argentina  iN  M T S1 S2 IL L I G

Staphylus melaina (Hayward, 1947)    iN  M S IL G

Staphylus kayei Cock, 1996    iN  M T

Staphylus sambo Evans, 1953  TL: Brazil (Pa)  M T

Staphylus putumayo (E. Bell, 1937)    iN  M T S

Staphylus saxos Evans, 1953    iN  M T

Staphylus satrap Evans, 1953  ref.149  M T S

Staphylus tingo Steinhauser, 1989  TL: Peru  M T

Staphylus oeta (Plötz, 1884)  Oeta Sootywing  Costa Rica to Bolivia and Argentina  iN  M T TI S G

Staphylus punctiseparatus Hayward, 1933  TL: Argentina  M IL

Staphylus lizeri (Hayward, 1938)    iN

Staphylus lizeri album Steinhauser, 1989  TL: Brazil (MG)  M T

Staphylus lizeri lizeri (Hayward, 1938)  TL: Bolivia  M T S

Staphylus caribbea (R. Williams & E. Bell, 1940)  Panamanian Sootywing  Nicaragua to Ecuador  iN  M T S1 S2 I G

Staphylus buena (R. Williams & E. Bell, 1940)  TL: Bolivia  M T S

Staphylus melius Steinhauser, 1989  TL: Argentina  M T

Staphylus musculus (Burmeister, 1875)    iN  M T S IL L G

Staphylus imperspicua (Hayward, 1940)  TL: Ecuador  M S

Staphylus corumba (R. Williams & E. Bell, 1940)  TL: Brazil (MS)  M T S

Staphylus ascalon (Staudinger, 1876)    iN  M T TI S IL

Staphylus eryx Evans, 1953  TL: Brazil (Pa)  M T

Staphylus melangon (Mabille, 1883)    iN

Staphylus melangon epicaste Mabille, 1903  ref.151  M T S IL G

Staphylus melangon melangon (Mabille, 1883)  TL: S. Amer.  M T S

Staphylus melangon stanga Evans, 1953  TL: Ecuador  M T

Staphylus tucumanus (Plötz, 1884)    iN  M S

Staphylus perna Evans, 1953    iN  M T S

Staphylus insignis O. Mielke, 1980  TL: Brazil (RS)  M OI S G

Staphylus tyro (Mabille, 1878)  TL: Brazil; Venezuela  M T

Staphylus azteca (Scudder, 1872)  Aztec Sootywing  E & W Mexico to Costa Rica  iN  M T S1 S2 L I G

Staphylus floridus Grishin, 2022  Florida Sootywing  USA: FL; ref.14  M T L I

Staphylus hayhurstii (W. H. Edwards, 1870)  Hayhurst's Sootywing  S Ontario, E USA to far SE SD, NB, KS, to S-C TX  iN  M T OD S L1 L2 I H G

Staphylus mazans (Reakirt, [1867])  Mazans Sootywing  C TX to CHIS  iN  M T OD S L1 L2 G Txt

Staphylus unicornis Steinhauser & Austin, 1993  Unicorn Sootywing  Costa Rica  M T1 T2

Staphylus ascalaphus (Staudinger, 1876)  Central American Sootywing  SE Mexico to Bolivia, S Brazil  iN  M T OD S1 S2 L G

Staphylus yucatanus Grishin, 2023  Yucatan Peninsula; ref.153  M S HS

Staphylus lenis Steinhauser, 1989  Lenis Sootywing  Trinidad  iN  M T G

Staphylus perforata (Möschler, 1879)  Perforata Sootywing  Panama, Colombia, Peru  M T S G

Staphylus rotundalus Grishin, 2023  Ecuador; ref.153

Staphylus tierra Evans, 1953  West-Mexican Sootywing  W Mexico (SON to CHIS), El Salvador  iN  M T S L

Staphylus veytius H. Freeman, 1969  TL: Mexico (Chia)  M T

Staphylus huigra (R. Williams & E. Bell, 1940)  Huigra Sootywing  TL: Ecuador

Staphylus huigra huigra (R. Williams & E. Bell, 1940)  El Salvador, Ecuador to Bolivia  M T S G

Staphylus huigra canka Evans, 1953  TL: Peru  M T

Staphylus alpheus (W. H. Edwards, 1876)  Saltbush Sootywing    iN

Staphylus alpheus oricus (W. H. Edwards, 1879)  E & S CA, NV, SW UT, NW AZ  M T S L Ha

Staphylus alpheus alpheus (W. H. Edwards, 1876)  [= texana], S AZ, NM, CO, far S MT, TX (verification needed for NE Mexico)  M S1 S2 L1 L2 I1 I2 G

Staphylus gracielae (MacNeill, 1970)  MacNeill's Sootywing  lower Colorado River drainage, Wa Co., UT to SON + Baja  iN  M T S L I H

Staphylus libya (Scudder, 1878)  Mojave Sootywing    iN

Staphylus libya joaquina (J. Emmel, T. Emmel & Mattoon, 1998)  S San Joaquin Valley, CA  M T S

Staphylus libya libya (Scudder, 1878)  Mojave Desert, CA- S NV-AZ  M S L1 L2 H Ha G V

Staphylus libya Great Basin segregate (, )  SE OR, SW ID, most of NV, E CA  M S L H Ha

Staphylus libya confertiblanca (Scott, 1992)  [perhaps a synonym of lena], W CO, far NW NM, UT?, N AZ?  M T S I

Staphylus libya lena (W. H. Edwards, 1882)  W ND, E MT, WY, UT?  M S

Pholisora  Scudder, 1872  

Pholisora mejicanus (Reakirt, [1867])  Mexican Sootywing  S-C Colorado to Oaxaca  iN  M T S L G

Pholisora albicirrus Glassberg, 2023  White-tufted Sootywing  SE AZ & NW MX; ref.145,146  M S L

Pholisora catullus (Fabricius, 1793)  Common Sootywing  widespread in USA and S Canada, S to Col. and Hid....S to Oax.  iN  M S HS L I G

Pholisora crestar Scott & Davenport, 2017  Starred Sootywing  Tulare & Kern Cos, CA; ref.11

Pholisora litus (Dyar, 1912)  Many-spotted Sootywing  mostly W Mexico; ref.13  iN  M T S L

  Tribe Carcharodini, Subtribe Carcharodina

Noctuana  E. Bell, 1937  

Noctuana lactifera (A. Butler & H. Druce, 1872)  Cryptic Skipper    iN

Noctuana lactifera bipuncta (Plötz, 1884)  E & W Mexico to Nicaragua  M S L

Noctuana lactifera lactifera (A. Butler & H. Druce, 1872)  Costa Rica to Colombia  M T S L I G

Noctuana noctua (C. Felder & R. Felder, 1867)  Noctua Skipper  Panama to Ecuador and Peru  iN  M T S

Noctuana brunneofusca (Mabille & Boullet, 1917)    iN  M T

Noctuana haematospila (C. Felder & R. Felder, 1867)    iN  M T S

Noctuana stator (Godman, 1899)  Red-studded Skipper  W Mexico & S TX (one-time stray) to Guianas, Trinidad, Peru  iN  M T S L I G

Noctuana diurna (A. Butler, 1870)    iN  M T S

Windia  H. Freeman, 1969  

Windia windi H. Freeman, 1969  Wind’s Skipper  SE Arizona (one-time stray), NW Mexico to Costa Rica  iN  M T S L G

  Tribe Pyrgini

Pyrgus  Hübner, [1819]  

Pyrgus centaureae (Rambur, [1842])  Grizzled Skipper    iN

Pyrgus centaureae dzekh Gorbunov, 2007  AK; ref.12  M S

Pyrgus centaureae freija (B. Warren, 1924)  boreal N America  M T S Ha

Pyrgus centaureae loki Evans, 1953  Rocky Mountains  M T S1 S2 S3 L1 L2 I1 I2 Ha

Pyrgus centaureae wyandot (W. H. Edwards, 1863)  E & NE USA  M T S L1 L2 G

Pyrgus ruralis (Boisduval, 1852)  Two-banded Checkered-Skipper    iN

Pyrgus ruralis ruralis (Boisduval, 1852)  NW CO, UT, W WY, W MT, SW ALTA, S BC to C CA  M T S1 S2 S3 L1 L2 Ha G

Pyrgus ruralis lagunae Scott, 1981  Laguna Mts., San Diego Co., CA  M T1 T2 T3 S L Ha1 Ha2

Pyrgus xanthus W. H. Edwards, 1878  Mountain Checkered-Skipper  S UT, N & E AZ, NM, S-C & SW CO  iN  M T S IL L1 L2 I H G

Pyrgus scriptura (Boisduval, 1852)  Small Checkered-Skipper    iN

Pyrgus scriptura scriptura (Boisduval, 1852)  at least San Francisco Bay area, CA  M T S L G

Pyrgus scriptura apertorum Austin, 1998  most of range, ALTA to Zacatecas  M T S L I Ha

Chirgus  Grishin, 2019  

Chirgus barrosi (Ureta, 1956)  Chile  M S

Chirgus limbata (Erschoff, 1876)  Peru, Bolivia, Argentina & Chile  iN  M T S

Chirgus nigella (Weeks, 1902)  Peru, Bolivia; ref.7  M S L

Chirgus fides (Hayward, 1940)  Peru, Bolivia, Argentina & Chile  iN  M T S IL G

Chirgus bocchoris (Hewitson, 1874)  Peru, Bolivia, Argentina & Chile  iN

Chirgus bocchoris cuzcona (Draudt, 1923)  TL: Peru  M T S L G

Chirgus bocchoris bocchoris (Hewitson, 1874)  TL: Bolivia  M T S IL G

Chirgus bocchoris trisignatus (Mabille, 1875)  TL: Chile  M T S

Chirgus veturius (Plötz, 1884)  Brazil & Bolivia  iN  M T S

Burnsius  Grishin, 2019  ref.7

Burnsius notatus (Blanchard, 1852)  Chile  iN

Burnsius notatus notatus (Blanchard, 1852)  TL: Chile  M T S

Burnsius notatus valdiviana (Reed, 1877)  TL: Chile  M T

Burnsius crisia (Herrich-Schäffer, 1865)  Antillean Checkered-Skipper  Cuba and Hispaniola  iN  M T S G

Burnsius communis (Grote, 1872)  Common Checkered-Skipper  USA & S Canada, S to Oaxaca; ref.7,15  iN

Burnsius communis W Oregon segregate (, )  W Oregon, N CA Siskiyous  M S L Ha

Burnsius communis high-elevation Cascades-Sierra Nevada segregate (, )  high elevation C Cascades, OR, & N Sierra Nevada, CA  M L

Burnsius communis communis (Grote, 1872)  widespread in USA & S Canada, S to C Mexico  M T S1 S2 Ab L1 L2 L3 L4 I F Ha G

Burnsius communis albescens (Plötz, 1884)  S Mexico; ref.15  M T

Burnsius albezens Grishin, 2022  White Checkered-Skipper  [former albescens] S USA (CA to SC), S to Oaxaca; ref.15  iN  M T S L1 L2 I H G V

Burnsius burnsi Grishin, 2022  Burns' Checkered-Skipper  [N. Amer. "adepta"] Mexico to Panama; ref.15  M T S1 S2 Ab L G

Burnsius adepta (Plötz, 1884)  E Colombia; ref.15  iN

Burnsius adepta adepta (Plötz, 1884)  E Colombia; ref.15  M T S

Burnsius adepta inepta Grishin, 2022  W Colombia & Ecuador; ref.15  M T S

Burnsius orcynus Grishin, 2022  Curacao & Margarita Is. (Venezuela); ref.15  M T IL

Burnsius orcynoides (Giacomelli, 1928)    iN  M S IL L G

Burnsius chloe (Evans, 1942)  Ecuador & Peru; ref.15  iN  M T S

Burnsius titicaca (Reverdin, 1921)  Peru; ref.15  iN  M S

Burnsius philetas (W. H. Edwards, 1881)  Desert Checkered-Skipper  S AZ to S TX, S to Chiapas  iN  M T S L I H G

Burnsius oileus (Linnaeus, 1767)  Tropical Checkered-Skipper  SE AZ, SW NM, S TX to GA, S to CR, Gr. & L. Antilles  iN  M T S1 S2 HS Ma L1 L2 L3 L4 I G

Burnsius orcus (Stoll, 1780)  Orcus Checkered-Skipper  [= varus] Oax and Chis to Argentina, Trinidad to Dominica; ref.15  iN  M T S1 S2 IL Co L1 L2 G

Burnsius brenda (Evans, 1942)  Brenda Checkered-Skipper  Ecuador & Peru  iN  M T S Co G

Heliopetes  Billberg, 1820  [includes Leucoscirtes & Heliopyrgus as subgenera]; ref.7

Heliopetes americanus (Blanchard, 1852)  ref.7  iN

Heliopetes americanus americanus (Blanchard, 1852)  TL: Chile  M S

Heliopetes americanus bellatrix (Plötz, 1884)  Argentina; ref.15  M T S IL G

Heliopetes willi (Plötz, 1884)  Brazil, Argentina & Paraguay; ref.15  M S IL G

Heliopetes domicella (Erichson, [1849])  Erichson’s White-Skipper  ref.7  iN

Heliopetes domicella domicella (Erichson, [1849])  S CA & S NV to S TX, S to Venezuela  M T S L1 L2 I G

Heliopetes domicella margarita (E. Bell, 1937)  TL: Venezuela  M T S

Heliopetes sublinea Schaus, 1902  East-Mexican White-Skipper  S TX & E Mexico to Yucatan Peninsula; ref.7  iN  M T S L G

Heliopetes purgia Schaus, 1902  Brazil; ref.7  iN  M T S

Heliopetes omrina (A. Butler, 1870)  Stained White-Skipper  Panama to Argentina  iN  M T S IL L G

Heliopetes ericetorum (Boisduval, 1852)  Northern White-Skipper  Baja California (norte), CA, Columbia Basin, Great Basin, AZ  iN  M T S L1 L2 L3 I Ha G

Heliopetes laviana (Hewitson, 1868)  Laviana White-Skipper    iN

Heliopetes laviana laviana (Hewitson, 1868)  S AZ and Baja California, C & S TX, most of Mexico to Venez.  M T S L1 L2 L3 L4 L5 I H G

Heliopetes laviana leca (A. Butler, 1870)  TL: Venezuela  M T

Heliopetes libra Evans, 1944    iN  M T S IL L G

Heliopetes macaira (Reakirt, [1867])  Turk’s-cap White-Skipper  S TX, E & W Mexico to Panama  iN  M T S L I G

Heliopetes nivella (Mabille, 1883)    iN  M T S

Heliopetes orbigera (Mabille, 1888)  Brazil; ref.7  M T OI S

Heliopetes marginata Hayward, 1940    iN  M T

Heliopetes elonmuski Grishin, 2023  SpaceX White-Skipper  S TX to Costa Rica; ref.19  M T S L1 L2 I G

Heliopetes arsalte (Linnaeus, 1758)  Veined White-Skipper  Panama to Argentina; ref.19  iN  M T S IL G

Heliopetes lana Grishin, 2023  Mexico to Guatemala; ref.153  M S

Heliopetes acuta Grishin, 2024  Mexico: Oax; ref.154  M T

Heliopetes alana (Reakirt, 1868)  Alana White-Skipper  [= janice] Costa Rica to Argentina; ref.20  iN  M T S1 S2 IL L1 L2 G

Heliopetes chimbo Evans, 1953    iN  M T S

Heliopetes petrus (Hübner, [1819])    iN  M S

Heliopetes ochroleuca J. Zikán, 1938    iN  M T S IL L G

Heliopetes leucola (Hewitson, 1868)    iN  M T S

Celotes  Godman & Salvin, 1899  

Celotes sabinus Grishin, 2023  Arizona Streaky-Skipper  ref.146  M T S L I

Celotes nessus (W. H. Edwards, 1877)  Common Streaky-Skipper  far SE CA, S AZ, NM, S OK, TX, N Mexico  iN  M T S L I H G

Celotes spurcus A. Warren, Steinhauser, Hernandez-Mejia & Grishin, 2008  Mexican Streaky-Skipper  Central Mexico; ref.98  iN  M T L Ha G

Celotes limpia Burns, 1974  West-Texas Streaky-Skipper  W TX to Coahuila  iN  M T S L G

Systaspes  Weeks, 1901  

Systaspes funebris (R. Felder, 1869)  West-Mexican Spurwing  W & S Mexico; ref.5  iN  M T S L G

Systaspes corrosus (Mabille, 1878)  Small Spurwing  E Mexico to Panama; ref.5  iN  M T S L

Trina  Evans, 1953  

Trina geometrina (C. Felder & R. Felder, 1867)    iN

Trina geometrina geometrina (C. Felder & R. Felder, 1867)  TL: Venezuela; Colombia  M T S IL L

Trina geometrina phalaena (Mabille, 1898)  TL: Bolivia  M T S L

Antina  Grishin, 2022  ref.16

Antina minor (O. Mielke, 1980)  S Brazil; ref.16  iN  M T OI S G

Antigonus  Hübner, [1819]  

Antigonus nearchus (Latreille, [1817])  Large Spurwing  E & W Mexico to S Brazil and Paraguay  iN  M T S IL L I G

Antigonus erosus (Hübner, [1812])  Common Spurwing  W Mexico & S TX (rare stray) to S Brazil and Paraguay  iN  M T S1 S2 IL L1 L2 I G

Antigonus emorsa (R. Felder, 1869)  White Spurwing  W Mexico (SE AZ as rare stray, SON to OAX)  iN  M T S1 S2 L I G

Diaeus  Godman & Salvin, 1895  

Diaeus varna Evans, 1953  Camouflaged Skipper  E Mexico to Panama  iN  M T S L G

Diaeus lacaena (Hewitson, 1869)    iN  M T S L

Diaeus ambata Evans, 1953    iN  M T S

Diaeus variegata (Plötz, 1884)    iN  M T TI S G

Onenses  Godman & Salvin, 1895  

Onenses hyalophora (R. Felder, 1869)  Crystal-winged Skipper  E Mexico to Guatemala  iN  M S L G

Systasea  W. H. Edwards, 1877  

Systasea microsticta Dyar, 1923  Rare Spurwing  S Mexico (GRO, MOR, PUE, OAX)  iN  M T S L Ha

Systasea pulverulenta (R. Felder, 1869)  Texas Powdered-Skipper  S AZ, S NM, C & S TX to Guatemala  iN  M T S L1 L2 I H G

Systasea zampa (W. H. Edwards, 1876)  Arizona Powdered-Skipper  Baja, far S CA, S NV, S AZ, S NM, W TX, SON, CHIH, COAH  iN  M T1 T2 TI S L I H G

Zobera  H. Freeman, 1970  

Zobera albopunctata H. Freeman, 1970  Coliman Zobera  W Mexico (Colima)  M T S G

Zobera marginata H. Freeman, 1979  Western Zobera  W Mexico (NAY, MICH, OAX)  iN  M T S L G

Zobera oaxaquena Steinhauser, 1991  Oaxacan Zobera  S Mexico (OAX)  iN  M T1 T2 S L G

Xenophanes  Godman & Salvin, 1895  

Xenophanes ruatanensis Godman & Salvin, 1895  Glassy-winged Skipper  S Texas to Costa Rica; ref.16  M T S1 S2 L1 L2 I1 I2 G

Xenophanes tryxus (Stoll, 1780)  Classy-winged Skipper  Panama to Argentina; ref.16  iN  M T S IL G

Canesia  Grishin, 2019  ref.5

Canesia canescens (R. Felder, 1869)  Hoary Skipper  W Mexico & S TX to Panama; ref.5  iN  M T S1 S2 IL L I G

Canesia pallida (Röber, 1925)  S Brazil; ref.5,13  M T S IL

Canesia lilloi (Hayward, 1947)  Ecuador; ref.5  M TI IL G

Canesia leada (A. Butler, 1870)  Butler's Hoary-Skipper  Costa Rica to Bolivia and S Brazil; ref.5  iN  M T S HS IL G

Canesia recurva (Austin, 2000)  Brazil; ref.5  M T OI G

Canesia callipetes (Godman & Salvin, 1895)  Cloud-forest Hoary-Skipper  S Mexico and Guatemala; ref.5  iN  M T OI S L G

Canesia meridensis (Godman & Salvin, 1895)  Costa Rican Hoary-Skipper  Nicaragua to Colombia and Venezuela; ref.5  iN  M T OI S L G

Canesia ella Grishin, 2023  Beautiful Hoary-Skipper  Venezuela; ref.153

Carrhenes  Godman & Salvin, 1895  

Carrhenes fuscescens (Mabille, 1891)  Tanned Hoary-Skipper  E & W Mexico to Costa Rica  iN  M T S1 S2 L I G

Carrhenes conia Evans, 1953  the Guianas; ref.13  M T S G

Carrhenes chaeremon (Mabille, 1891)    iN  M T S IL G

Carrhenes calidius Godman & Salvin, 1895  Rain-forest Hoary-Skipper  SE Mexico to Bolivia  iN  M T S1 S2 I G

Carrhenes sinesinus Steinhauser, 1989  TL: Colombia  M T

Carrhenes bamba Evans, 1953    iN  M T S IL G

Carrhenes infuscescens Steinhauser, 1989  TL: Argentina  M T S

Carrhenes decens (A. Butler, 1874)  Peru; ref.13  iN  M T S L

Santa  Grishin, 2019  ref.5

Santa santes (E. Bell, 1940)  Peru and Bolivia; ref.5  iN  M T S

Santa trifasciatus (Lindsey, 1925)  Peru; ref.5  iN  M S HS

Santa palica (Mabille, 1888)  [= Paches era] Peru; ref.13  M T OI S

Paches  Godman & Salvin, 1895  [includes Tiges as a subgenus]; ref.13

Paches mutilatus (Hopffer, 1874)  Peru; ref.13  iN  M T S L1 L2 G

Paches liborius (Plötz, 1884)  Brazil (Ba); ref.13  iN

Paches liborius liborius (Plötz, 1884)  Brazil (Ba); ref.13  M T S IL L G

Paches liborius areta (Evans, 1953)  TL: Brazil (RS)  M T S

Paches exosa (A. Butler, 1877)  TL: Brazil (Am)  M T S L

Paches polla (Mabille, 1888)  Polla Blue-Skipper  E & W Mexico to Costa Rica  iN  M T OI S IL L

Paches gladiatus (A. Butler, 1870)    iN  M T S

Paches loxana Evans, 1953  Bolivia; ref.153  M T S

Paches loxeca Grishin, 2023  S Ecuador & N Peru; ref.153

Paches loxus (Westwood, 1852)  Glorious Blue-Skipper  Panama to Venezuela, Ecuador, N Brazil  iN  M T OI S IL L G

Paches gloriosus Röber, 1925  Glorious Blue-Skipper  [= zonula], SE Mexico to Costa Rica; ref.153  M T S1 S2 S3 L I

Plumbago  Evans, 1953  

Plumbago plumbago (Plötz, 1884)    iN  M T OI S L

Plumbago pulverea (Mabille, 1878)  TL: French Guiana  M T TI S

Plumbago serapion (Plötz, 1884)  TL: Brazil (RJ)  M T

Zopyrion  Godman & Salvin, 1896  [includes Timochreon as a subgenus]; ref.13

Zopyrion subvariegata Hayward, 1942    iN

Zopyrion subvariegata subvariegata Hayward, 1942  TL: Ecuador  M T S

Zopyrion subvariegata thyas Evans, 1953  TL: Peru  M T S

Zopyrion reticulata Hayward, 1942  TL: Brazil (Go)  M S

Zopyrion evenor Godman, 1901    iN

Zopyrion evenor tressa Evans, 1953  TL: Brazil (Pa)  M T

Zopyrion evenor thania Evans, 1953  TL: Brazil (Pe)  M T S

Zopyrion evenor evenor Godman, 1901  TL: Brazil (MS)  M T S

Zopyrion sandace Godman & Salvin, 1896  Sandy Skipper  E & W Mexico to Costa Rica  iN  M T S L G

Zopyrion satyrina (C. Felder & R. Felder, 1867)  Colombia & Venezuela; ref.13  iN  M T S

Zopyrion satyrus (C. Felder & R. Felder, 1867)  Satyr Skipper  ref.13  iN

Zopyrion satyrus satyrus (C. Felder & R. Felder, 1867)  Costa Rica to upper Amazon  M T S1 S2 L I G

Zopyrion satyrus forta (Evans, 1953)  TL: Brazil (Pa)  M T S

Zopyrion satyrus tampa (Evans, 1953)  TL: Brazil (MT)  M T S IL G

Zopyrion doria (Plötz, 1884)  Brazil; ref.13  iN  M S

Anisochoria  Mabille, 1877  

Anisochoria bacchus Evans, 1953  Northern Snout-Skipper  SE Mexico to El Salvador  iN  M T S L G

Anisochoria polysticta Mabille, 1877  Central American Snout-Skipper  Costa Rica to Guianas; ref.13  M T S I

Anisochoria pedaliodina (A. Butler, 1870)    iN  M T S IL L1 L2 G

Anisochoria extincta Hayward, 1933  Argentina; ref.13  M S IL G

Anisochoria sublimbata Mabille, 1883    iN  M T TI S IL L G

Anisochoria subpicta Schaus, 1902    iN  M T S

Anisochoria verda Evans, 1953  Ecuador; ref.13  M T S

Anisochoria minorella Mabille, 1898    iN  M T S L

Anisochoria quadrifenestrata (Bryk, 1953)  TL: Brazil (Ba)  M T S

Anisochoria superior Mabille, 1898  TL: Bolivia  M T S

Anisochoria vianna Evans, 1953  TL: Brazil (Go)  M T S

Bralus  Grishin, 2019  ref.5

Bralus nadia (Nicolay, 1980)  Ecuador; ref.5,13  iN  M T

Bralus albida (Mabille, 1888)  Peru; ref.5  M T OI S

Bralus alco (Evans, 1953)  Bolivia; ref.13  M T S

  Tribe Erynnini

Clito  Evans, 1953  

Clito jonkersi de Jong, 1983  Jonkers's Clito  TL: Surinam  M T OD

Clito palotschka Grishin, 2014  I-Clito  Ecuador; ref.71  M T

Clito litteroides Grishin, 2014  Subscripted Clito  Venezuela; ref.71  M T

Clito trinidadensis Cock, 1998  Trinidad Clito  Trinidad  M T OD

Clito littera (Mabille, 1877)  Scripted Clito  S Brazil; ref.71  M T OD S IL Txt

Clito nebulosa (Draudt, 1924)  Nebulous Clito  Bolivia; ref.71  M T OD G

Clito anda Evans, 1953  White-patched Clito  Amazonian; ref.71  M T S IL G

Clito bibulus (N. Riley, 1929)  Mottled Clito  Brazil  M T S G

Clito congruens Grishin, 2023  S & W Mexico to Panama; ref.153

Clito aberrans (Draudt, 1924)  Northern Clito  [= Evans' "clito"], Colombia to S Brazil  iN  M T OD S1 S2 S3 S4 IL L I1 I2 G

Clito sompa Evans, 1953  Brazil  iN  M T S G

Clito tuva Evans, 1953  Brazil  M T S G

Clito zelotes (Hewitson, 1873)  Hewitson’s Clito  SE Mexico (VER) to French Guiana and upper Amazon  iN  M T S IL L G

Clito mnemon (Schaus, 1913)  Horseshoe-spotted Skipper  Costa Rica  iN  M T S

Clito zenda Evans, 1953  TL: Surinam  M T S G

Clito clada (Evans, 1953)  TL: Brazil (Ma)  M T S IL G

Anastrus  Hübner, [1824]  

Anastrus meliboea (Godman & Salvin, 1894)  Scarce Anastrus    iN

Anastrus meliboea meliboea (Godman & Salvin, 1894)  (SE Mexico?), Costa Rica to Colombia  M T S L G

Anastrus meliboea bactra Evans, 1953  TL: Bolivia  M T S L

Anastrus neaeris (Möschler, 1879)  Brilliant Anastrus    iN

Anastrus neaeris neaeris (Möschler, 1879)  S Mexico to Venezuela and Ecuador  M T S1 S2 L I G

Anastrus neaeris narva Evans, 1953  Panama to Bolivia and E-C Brazil  M T S IL L G

Anastrus virens Austin, 1999    iN

Anastrus virens albopannus Austin, 2002  SE Mexico (OAX) to Panama  M T S L G

Anastrus virens virens Austin, 1999  TL: Brazil (Rd)  M T G

Anastrus chaqua Evans, 1953  Peru  M T S

Anastrus obscurus Hübner, [1824]    iN  M OI S

Anastrus ulpianus Poey, 1832  S Brazil  iN  M T S

Mylon  Godman & Salvin, 1894  

Mylon illineatus Mabille & Boullet, 1917  Illineatus Mylon    iN

Mylon illineatus illineatus Mabille & Boullet, 1917  Ecuador, Peru, Bolivia  M T S IL Co Ma L G

Mylon illineatus toxina Evans, 1953  Upper Amazonian region of Colombia, Ecuador, NE Peru and NW Brazil  M T OD Co L G

Mylon lassia (Hewitson, 1868)  Bold Mylon  E & W Mexico to Guianas and Trinidad  iN  M T S1 S2 IL L I G

Mylon zephus (A. Butler, 1870)  Zephus Mylon    iN

Mylon zephus zephus (A. Butler, 1870)  Colombia & Venezuela, hypothetical for Panama  M T S

Mylon zephus albodiscus Mabille & Boullet, 1917  Peru, Bolivia  M T L

Mylon orsa Evans, 1953  Orsa Mylon  Costa Rica and Panama  M T S IL G

Mylon mestor Evans, 1953  Mestor Mylon  Ecuador  iN  M T S IL G

Mylon salvia Evans, 1953  Evans’ Mylon  Mexico, Guatemala to Ecuador  iN  M T S1 S2 IL I G

Mylon ander Evans, 1953  Narrow-winged Mylon    iN

Mylon ander andrea Evans, 1953  SE Mexico to Colombia  M T OD S L

Mylon ander ander Evans, 1953  Colombia to Bolivia and S Brazil  M T OD S IL L G

Mylon maimon (Fabricius, 1775)  Common Mylon  [= menippus], E & W Mexico to Argentina  iN  M T S1 S2 S3 IL L1 L2 L3 I G Txt

Mylon cajus (Plötz, 1884)    iN

Mylon cajus hera Evans, 1953  (SE Mexico?), Guatemala to Panama  M T S IL L G

Mylon cajus cajus (Plötz, 1884)  Colombia to S Brazil  M T S L

Hoodus  Grishin, 2019  ref.5

Hoodus pelopidas (Fabricius, 1793)  Pale Mylon  SE AZ (one-time stray), E & W Mexico to Paraguay; ref.5  iN  M T S1 S2 IL Ma L1 L2 I G

Hoodus cristata (Austin, 2000)  Austin's Mylon  Campeche and Chiapas to Guianas and Brazil; ref.5  iN  M OI S IL G

Hoodus jason (Ehrmann, 1907)  Jason’s Mylon  (SE Mexico?), Guatemala to S Brazil and Paraguay; ref.5  iN  M TI S L G

Hoodus exstincta (Mabille & Boullet, 1917)  Forgotten Mylon  NW Brazil; ref.5  M IL L G

Hoodus simplex (Austin, 2000)  Unadorned Mylon  NW Brazil; ref.5  M T OI L G

Hoodus argonautarum (Austin, 2000)  Argonaut Mylon  NW Brazil; ref.5  iN  M T OI S G

Echelatus  Godman & Salvin, 1894  

Echelatus sempiternus (A. Butler & H. Druce, 1872)  Common Anastrus  ref.5  iN

Echelatus sempiternus sempiternus (A. Butler & H. Druce, 1872)  S TX (one-time stray), E & W Mexico to Venezuela and Ecuador  M T S L I1 I2 G

Echelatus sempiternus simplicior (Möschler, 1877)  Colombia to Brazil; ref.13  M T S HS L G

Echelatus sempiternus dilloni E. Bell & W. Comstock, 1948  Jamaica, Hispaniola, Cuba  M T S L

Tolius  Grishin, 2022  ref.13

Tolius luctuosus (Godman & Salvin, 1894)  West-Mexican Anastrus  [= Mexican "robigus"], W Mexico; ref.13  iN  M T S G

Tolius tolimus (Plötz, 1884)  Blurry Anastrus    iN

Tolius tolimus tolimus (Plötz, 1884)  E Mexico to Ecuador; ref.13  M S L G

Tolius tolimus robigus (Plötz, 1884)  S. Amer.; ref.13  M T TI S G

Anaxas  Grishin, 2019  ref.5

Anaxas petius (Möschler, 1877)  ref.5  iN

Anaxas petius peto (Evans, 1953)  Guatemala to Costa Rica  M T S G

Anaxas petius petius (Möschler, 1877)  TL: Surinam  M T S L G

Anaxas isidro (Grishin, 2012)  Isidro's Skipper  Costa Rica and Panama; ref.62,5  M T G

Anaxas obliqua (Plötz, 1884)  S. Amer.; ref.5  iN  M T S IL G

Potamanaxas  Lindsey, 1925  

Potamanaxas thoria (Hewitson, 1870)  Thoria Skipper  Nicaragua to Peru  iN  M T OI OD S1 S2 S3 L I G

Potamanaxas okroogly Grishin, 2013  Ovalpatch Skipper  E Colombia, Ecuador, Peru & Bolivia; ref.69  M T S HS IL L G

Potamanaxas cranda Evans, 1953  Cranda Skipper  Costa Rica & Panama  M T OD S1 S2 G

Potamanaxas louisghilli Grishin, 2015  Hill's Skipper  Costa Rica & Panama; ref.76  M T S

Potamanaxas [n. sp.] Grishin, MS  Potamanaxas louisghilli complex  Colombia to Peru  M S

Potamanaxas melicertes (Godman & Salvin, 1895)  Melicertes Skipper  Costa Rica to Ecuador  iN  M T OI OD S IL

Potamanaxas paphos Evans, 1953  Paphos Skipper  Costa Rica to Ecuador  iN  M T OD S1 S2 S3 G

Potamanaxas hirta (Weeks, 1901)  Hirta Skipper  Colombia to Bolivia  iN  M T OI OD TI S1 S2 S3 S4 L G Txt

Potamanaxas flavofasciata (Hewitson, 1870)  Yellowbanded Skipper    iN

Potamanaxas flavofasciata flavofasciata (Hewitson, 1870)  Ecuador, Peru  M T S1 S2 G

Potamanaxas flavofasciata pantra Evans, 1953  Bolivia  M T OD S IL G

Potamanaxas effusa (Draudt, 1922)  Effusa Skipper    iN

Potamanaxas effusa effusa (Draudt, 1922)  Colombia to Guianas and Peru  M OI OD S1 S2 G

Potamanaxas effusa confusa (Draudt, 1922)  Costa Rica, Panama & Colombia  M OI OD S

Potamanaxas xantholeuce (Mabille, 1888)  Xantholeuce Skipper  Panama  iN  M T OI TI S

Potamanaxas hermieri Grishin, 2013  Hermier's Skipper  Ecuador; ref.66  M T

Potamanaxas paralus (Godman & Salvin, 1895)  Paralus Skipper  Colombia to Bolivia  iN  M T OD S1 S2 IL L Txt

Potamanaxas latrea (Hewitson, 1875)  Latrea Skipper    iN

Potamanaxas latrea latrea (Hewitson, 1875)  Nicaragua to Colombia  M T OI OD TI S

Potamanaxas latrea caliadne (Godman & Salvin, 1895)  Costa Rica  M T OI OD S IL G

Potamanaxas latrea tusca Evans, 1953  Ecuador  M T OD S IL L G

Potamanaxas latrea tyndarus Evans, 1953  Peru  M T OD S G

Potamanaxas thestia (Hewitson, 1870)  Thestia Skipper  Ecuador, Peru, Bolivia  iN  M T S IL L G

Potamanaxas forum Evans, 1953  Forum Skipper  S Peru, Bolivia  M T OD S G

Potamanaxas lamasi Grishin, 2013  Lamas' Skipper  N & central Peru; ref.65  M T OD S G

Potamanaxas fuma Evans, 1953  Fuma Skipper  central Peru  M T OD G

Potamanaxas frenda Evans, 1953  Frenda Skipper  Peru  iN  M T OD S IL G

Potamanaxas bana E. Bell, 1956  Bana skipper  Ecuador  M T OI OD S L G

Potamanaxas serp Grishin, 2013  Sickle-clasp Skipper  Colombia; ref.67  M T

Potamanaxas quira E. Bell, 1956  Quira Skipper  Colombia  iN  M T OI OD S G

Potamanaxas trigga Evans, 1953  Trigga Skipper  Colombia, Ecuador, Peru  iN  M T OD S G

Potamanaxas laoma (Hewitson, 1870)  Laoma Skipper    iN

Potamanaxas laoma fumida (Draudt, 1922)  Colombia  M OI OD S

Potamanaxas laoma laoma (Hewitson, 1870)  Ecuador  M T S1 S2 G

Potamanaxas laoma violacea (Dognin, 1888)  Ecuador, Peru  M T OI OD S1 S2 S3 S4 S5 IL G

Potamanaxas laoma cosna Evans, 1953  Peru, Bolivia  M T OD S1 S2 G

Potamanaxas trex Evans, 1953  Trex Skipper  Colombia, Ecuador  M T OD S G

Potamanaxas tunga E. Bell, 1956  Tunga Skipper  Ecuador  M T OI OD G

Potamanaxas zagadka Grishin, 2013  Phylomystery Skipper  Ecuador; ref.68  M T

Potamanaxas andraemon (Mabille, 1898)  Andraemon Skipper  Colombia (West Andes)  iN  M T OD S L G

Potamanaxas perornatus Hayward, 1940  Perornatus Skipper  Colombia (East Andes), Ecuador, N Peru  M T OD S G

Potamanaxas tschotky Grishin, 2013  Crisppatch Skipper  Colombia & Ecuador; ref.69  iN  M T S L G

Festivia  Grishin, 2019  ref.5

Festivia cronion (C. Felder & R. Felder, 1867)  ref.5  iN  M T S IL L

Festivia festiva (Erichson, [1849])  ref.5  iN  M T S IL L

Festivia caerulans (Mabille & Boullet, 1917)  ref.5  iN  M T

Festivia grippa (Evans, 1953)  Grippa Skipper  Costa Rica to Ecuador and Bolivia; ref.5  iN  M T S L

Festivia peruvia Grishin, 2023  Peru; ref.153

Festivia jinna (Evans, 1953)  Colombia; ref.5  M T OD

Festivia adamantinus (Mabille, 1898)  Peru and Bolivia; ref.5  M T S IL

Sostrata  Godman & Salvin, 1895  

Sostrata nordica Evans, 1953  Blue-studded Skipper  S TX to Costa Rica  iN  M T S1 S2 L I

Sostrata bifasciata (Ménétriés, 1829)    iN

Sostrata bifasciata leucorrhoa Godman & Salvin, 1895  Costa Rica to Colombia  M T S L

Sostrata bifasciata adamas (Plötz, 1884)  TL: Brazil  M S

Sostrata bifasciata bifasciata (Ménétriés, 1829)  TL: Brazil (MG)  M T S

Sostrata pusilla Godman & Salvin, 1895  Pusilla Skipper    iN

Sostrata pusilla pusilla Godman & Salvin, 1895  Nicaragua to upper Amazon  M T S I

Sostrata pusilla pulsa Evans, 1953  TL: Ecuador  M T OD S

Gorgythion  Godman & Salvin, 1896  

Gorgythion vox Evans, 1953  Crab’s-claw Skipper  E Mexico to Costa Rica  iN  M T OD S L G

Gorgythion begga (Prittwitz, 1868)    iN

Gorgythion begga pyralina (Möschler, 1877)  S TX to Guianas and Bolivia  M OI OD S1 S2 L I1 I2 G

Gorgythion begga begga (Prittwitz, 1868)  ref.151  M T OD S IL L G

Gorgythion guyanus Grishin, 2023  Guyana; ref.151

Gorgythion marginata Schaus, 1902  Peru; ref.151  M T OD

Gorgythion plautia (Möschler, 1877)    iN  M T OI OD S L G

Gorgythion beggina Mabille, 1898  Beggina Skipper  Costa Rica to Argentina  iN

Gorgythion beggina escalophoides (Hayward, 1941)  Costa Rica to Argentina  M T OD TI S IL G Txt

Gorgythion beggina beggina Mabille, 1898  TL: Bolivia  M T OD S G

Gorgythion canda Evans, 1953  TL: Paraguay  M T OD S G

Gorgythion alcandra (Dognin, 1891)    iN  M T OD G Txt

Chiothion  Grishin, 2019  

Chiothion georgina (Reakirt, 1868)  White-patched Skipper  ref.5  iN

Chiothion georgina pelagica (Weeks, 1891)  Baja California  M T OI S L

Chiothion georgina georgina (Reakirt, 1868)  SE AZ & S TX, E & W Mexico to Costa Rica  M T S1 S2 L1 L2 I G

Chiothion asychis (Stoll, 1780)  ref.5  iN

Chiothion asychis simon (Evans, 1953)  Costa Rica to Venezuela, Trinidad  M T S1 S2 G

Chiothion asychis vincenta (Evans, 1953)  St. Vincent  M T S G

Chiothion asychis grenada (Evans, 1953)  St. Lucia, Grenadines, Grenada  M T S G

Chiothion asychis asychis (Stoll, 1780)  TL: Surinam  M T TI S IL G

Chiothion asychis zania (Evans, 1953)  TL: Peru  M T S G

Chiothion asychis autander (Mabille, 1891)  TL: Argentina  M T S IL L G

Chiothion khalili (N. Riley, 1934)  Brazil; ref.5  M T S G

Chiothion basigutta (Plötz, 1884)  ref.5  iN  M T S G

Theagenes  Godman & Salvin, 1896  

Theagenes aegides (Herrich-Schäffer, 1869)  White-centered Bent-Skipper  E & W Mexico to Panama  iN  M S L G

Theagenes albiplaga (C. Felder & R. Felder, 1867)    iN  M T S IL G

Theagenes dichrous (Mabille, 1878)  Brazil; ref.13  iN  M T S IL G

Helias  Fabricius, 1807  

Helias cama Evans, 1953  Squared Bent-Skipper  E Mexico to Colombia  iN  M T S1 S2 L I G

Helias godmani (Mabille & Boullet, 1917)  Godman's Bent-Skipper  Costa Rica to Ecuador  iN  M T S

Helias phalaenoides Fabricius, 1807  Fabricius' Bent-Skipper    iN

Helias phalaenoides phalaenoides Fabricius, 1807  Panama to Bolivia  M S L

Helias phalaenoides palpalis (Latreille, [1824])  TL: Brazil  M T S IL G

Ebrietas  Godman & Salvin, 1896  

Ebrietas osyris (Staudinger, 1876)  Yellow-patched Bent-Skipper  SE Mexico to Venezuela and Ecuador  iN  M T S1 S2 IL L I G

Ebrietas infanda (A. Butler, 1877)    iN  M T S L G

Ebrietas badia (Plötz, 1884)  Badia Bent-Skipper  Costa Rica to Bolivia  iN  M S G

Ebrietas anacreon (Staudinger, 1876)  Common Bent-Skipper    iN

Ebrietas anacreon anacreon (Staudinger, 1876)  E & W Mexico to Argentina  M T S1 S2 HS L I G

Ebrietas anacreon triseriata (Plötz, 1884)  TL: Venezuela

Ebrietas anacreon leucophaea Mabille & Boullet, 1917  TL: Colombia; Ecuador; Brazil  M T

Ebrietas sappho Steinhauser, 1974  Sappho Bent-Skipper  E Mexico to Costa Rica  iN  M T S G

Ebrietas elaudia (Plötz, 1884)  Elaudia Bent-Skipper    iN

Ebrietas elaudia livius Mabille, 1898  SE Mexico (TAB, CHIS) to Bolivia  M T S IL

Ebrietas elaudia elaudia (Plötz, 1884)  Panama, Colombia, Venezuela  M S

Ebrietas evanidus Mabille, 1898  Blurred Bent-Skipper  SE Mexico to Peru and upper Amazon  iN  M T S L G

Cycloglypha  Mabille, 1903  

Cycloglypha thrasibulus (Fabricius, 1793)  Widespread Bent-Skipper    iN

Cycloglypha thrasibulus thrasibulus (Fabricius, 1793)  E & W Mexico to S Brazil & Paraguay  M T S1 S2 IL L I G

Cycloglypha thrasibulus flinta Evans, 1953  TL: Ecuador  M T

Cycloglypha tisias (Godman & Salvin, 1896)  Tisias Bent-Skipper  SE Mexico (OAX) to Brazil  iN  M T OD S G

Cycloglypha corax Grishin, 2023  SE & S Brazil; ref.153

Cycloglypha caeruleonigra Mabille, 1903    iN  M T S G

Cycloglypha stellita J. Zikán, 1938    iN  M S

Cycloglypha enega (Möschler, 1877)  Enega Bent-Skipper  Nicaragua to Bolivia and S Brazil  iN  M T S L

Cycloglypha polax Evans, 1953    iN  M T S G

Camptopleura  Mabille, 1877  

Camptopleura janthinus (Capronnier, 1874)    iN  M T S G

Camptopleura oaxaca H. Freeman, 1969  Oaxacan Bent-Skipper  SE Mexico (Oaxaca)  iN  M T

Camptopleura auxo (Möschler, 1879)  Auxo Bent-Skipper  SE Mexico to S Brazil & Paraguay  iN  M T S1 S2 IL L I1 I2 G

Camptopleura theramenes Mabille, 1877  Mabille's Bent-Skipper  SE Mexico to Bolivia and Brazil  iN  M T S IL G

Camptopleura cincta Mabille & Boullet, 1917  Bolivia; ref.13  M T

Camptopleura termon (Hopffer, 1874)    iN  M T S G

Camptopleura orsus (Mabille, 1889)  [= impressus] Panama to Minas Gerais; ref.13  iN  M T OI S L G

Timochares  Godman & Salvin, 1896  

Timochares ruptifasciata (Plötz, 1884)  Brown-banded Skipper  SE AZ / SW NM, S TX to Chiapas and Belize  iN  M T S L1 L2 I1 I2 G

Timochares runia Evans, 1953  Jamaican-banded Skipper  Jamaica  M T S

Timochares trifasciata (Hewitson, 1868)  Many-banded Skipper    iN

Timochares trifasciata trifasciata (Hewitson, 1868)  E & W Mexico to S Brazil and Paraguay  M T S1 S2 HS L G

Timochares trifasciata sanda Evans, 1953  TL: Argentina  M T S G

Chiomara  Godman & Salvin, 1899  

Chiomara mithrax (Möschler, 1879)  Mithrax Duskywing  S AZ (rare stray) W & E Mexico to Argentina, Cuba  iN  M T S1 S2 IL L I G

Chiomara gundlachi (Skinner & Ramsden, 1924)  TL: Cuba  M T

Crenda  Grishin, 2019  ref.5

Crenda crenda (Evans, 1953)  Brazil; ref.5  M T S HS G

Neomorphuncus  Burns, 2019  

Neomorphuncus eugramma (Mabille, 1888)  Chiriqui Skipper  Costa Rica and Panama  iN  M T OI S1 S2 I

Ephyriades  Hübner, [1819]  

Ephyriades brunnea (Herrich-Schäffer, 1865)  'Cuban' Florida Duskywing    iN

Ephyriades brunnea floridensis E. Bell & W. Comstock, 1948  S Florida  M T S L I

Ephyriades brunnea brunnea (Herrich-Schäffer, 1865)  Cuba, Isla de Juventud, Bahamas  M T S L

Ephyriades jamaicensis (Möschler, 1879)  'Jamaican' Florida Duskywing  Jamaica; ref.13  M T S

Ephyriades zephodes (Hübner, [1825])  Zephodes Duskywing  Cuba, Hispaniola  iN  M T OI S G

Ephyriades arcas (Drury, 1773)  Caribbean Duskywing    iN

Ephyriades arcas philemon (Fabricius, 1775)  Greater Antilles  M T S IL L

Ephyriades arcas arcas (Drury, 1773)  northern Leeward Islands  M S

Ephyriades dominicensis E. Bell & W. Comstock, 1948  'Lesser Antillean' Florida Duskywing  Dominica and Guadeloupe; ref.7  M T S

Gesta  Evans, 1953  includes most species formerly in Erynnis; ref.10,127

Gesta telemachus (Burns, 1960)  Rocky Mountain Duskywing  E NV, UT, AZ, NM, CO, far S-C WY, W TX; ref.10  iN  M T S1 S2 L1 L2 L3 L4 L5 I G

Gesta juvenalis (Fabricius, 1793)  Juvenal's Duskywing  ref.10  iN

Gesta juvenalis juvenalis (Fabricius, 1793)  E USA, SE Canada, far NE Mexico (COAH)  M T S L1 L2 L3 L4 L5 L6 F G

Gesta juvenalis clitus (W. H. Edwards, 1883)  SE AZ, SW NM, W TX, E & W Mexico to Chiapas  M T S L I G

Gesta propertius (Scudder & Burgess, 1870)  Propertius Duskywing  Cascades-Sierra Nevada, westward, S BC to N Baja; ref.10  iN  M S1 S2 L1 L2 L3 Ha G

Gesta meridianus (E. Bell, 1927)  Meridian Duskywing  ref.10  iN

Gesta meridianus meridianus (E. Bell, 1927)  S NV, SW UT, AZ, NM, C & W TX, far N Mexico  M T S L I G

Gesta meridianus fieldi (Burns, 1964)  C Mexico  M T S Ha

Gesta horatius (Scudder & Burgess, 1870)  Horace's Duskywing  (far SE UT, far NE AZ), CO, NM, E USA, NE MX (Tam); ref.10  iN  M S1 S2 L1 L2 L3 L4 L5 I G

Gesta tristis (Boisduval, 1852)  Mournful Duskywing  ref.10  iN

Gesta tristis pattersoni (Burns, 1964)  Baja California Sur  M T S L

Gesta tristis tristis (Boisduval, 1852)  CA and Baja California Norte  M T S L Ha G

Gesta tristis tatius (W. H. Edwards, 1883)  AZ, S NM, W & S TX to Colombia  M T S1 S2 IL L1 L2 I Ha G

Gesta zarucco (Lucas, 1857)  Zarucco Duskywing  SE USA (OK-NC-FL), Cuba, Hispaniola; ref.10  iN  M S L1 L2 G

Gesta funeralis (Scudder & Burgess, 1870)  Funereal Duskywing  S CA, S AZ, S NM, TX S to Argentina; strays widely; ref.10  iN  M T S IL L1 L2 L3 L4 I Ha G

Gesta baptisiae (W. Forbes, 1936)  Wild Indigo Duskywing  E USA (far S Ontario) to N FL; ref.10  iN  M T S L1 L2 L3 L4 I H

Gesta lucilius (Scudder & Burgess, 1870)  Columbine Duskywing  NE USA, SE Canada; ref.10  iN  M S G

Gesta afranius (Lintner, 1878)  Afranius Duskywing  SE ALTA, S SASK to N Mexico; ref.10  iN

Gesta afranius BCN segregate (, )  SW CA and Baja California (norte)  M S

Gesta afranius afranius (Lintner, 1878)  TL: USA (Co)  M T S1 S2 L1 L2 L3 L4 I F G

Gesta afranius SE Mexican segregate (, )  SE Mexico (Puebla and Mexico States)  M S

Gesta persius (Scudder, 1863)  Persius Duskywing  ref.10  iN

Gesta persius persius (Scudder, 1863)  NE USA  M S L G

Gesta persius borealis (M. Cary, 1906)  most of W Canada, possibly S along Cascades (& W) to N CA  M T S L

Gesta persius avinoffi (W. Holland, 1930)  YT, Alaska  M T S

Gesta persius fredericki (H. Freeman, 1943)  Rocky Mountain region, USA, possibly S SASK, S ALTA  M T1 T2 T3 T4 S L1 L2 I1 I2 Ha

Gesta scudderi (Skinner, 1914)  Scudder’s Duskywing  SE AZ, SW NM, W TX to Guatemala; ref.10  iN  M T S L Ha G

Gesta pacuvius (Lintner, 1878)  Pacuvius Duskywing  ref.10  iN

Gesta pacuvius pacuvius (Lintner, 1878)  S UT, S CO, AZ, NM to Oaxaca  M S L I G

Gesta pacuvius lilius (Dyar, 1904)  NW CO, N UT, W WY, MT, ID, BC, WA, OR, W NV, N & E CA  M T S L I H

Gesta pacuvius pernigra (F. Grinnell, 1905)  San Francisco Bay area, C CA  M T S

Gesta pacuvius callidus (F. Grinnell, 1904)  S CA coast ranges to Baja California (norte)  M T S L H Ha

Gesta martialis (Scudder, [1870])  Mottled Duskywing  MN to NH, S to N FL, W to C TX, CO, N NM W NE, W SD; ref.10  iN  M S1 S2 L H G

Gesta invisus (A. Butler & H. Druce, 1872)  False Duskywing  C & S TX to Costa Rica  iN  M T S L I G

Gesta gesta (Herrich-Schäffer, 1863)  Impostor Duskywing  Greater Antilles, Costa Rica to Argentina  iN  M T S1 S2 IL L I G

Gesta inga Evans, 1953    iN  M T

Gesta austerus (Schaus, 1902)    iN  M T S IL G

Gesta heteropterus (Plötz, 1884)    iN  M S

Erynnis  Schrank, 1801  

Erynnis icelus (Scudder & Burgess, 1870)  Dreamy Duskywing  widespread in USA and Canada  iN  M S1 S2 L1 L2 L3 I Ha G

Erynnis lacustra (W. G. Wright, 1905)  'Lacustra' Sleepy Duskywing  Baja California Norte, S & C CA  M T S L Ha G

Erynnis burgessi (Skinner, 1914)  'Rocky Mountain' Sleepy Duskywing  S Rocky Mountain states, W TX, far SE CA, USA, far NW Mexico

Erynnis burgessi burgessi (Skinner, 1914)  S Rocky Mountain states, W TX, far SE CA, USA, far NW Mexico  M T S1 S2 L I Ha G

Erynnis burgessi muelleri (Draudt, 1923)  C & W Mexico  M T S L Ha G

Erynnis brizo (Boisduval & Le Conte, [1837])  Sleepy Duskywing  SE Canada, E USA, possibly far NE Mexico  iN  M S1 S2 L1 L2 L3 L4 F Ha G

Erynnis somnus (Lintner, 1881)  'Florida' Sleepy Duskywing  Peninsular Florida  M T S L

Erynnis mercurius (Dyar, 1926)  Mexican Duskywing  S-C Mexico (MEX, DF)  M T S

  Subfamily Tagiadinae

  Tribe Celaenorrhinini

Celaenorrhinus  Hübner, [1819]  

Celaenorrhinus aegiochus (Hewitson, 1876)  Flashing Flat  Costa Rica and Panama  iN  M T S L G

Celaenorrhinus savia (Evans, 1952)  TL: Peru  M T S IL

Celaenorrhinus orneates Austin, 1996  TL: Brazil (Rd)  M T OI S

Celaenorrhinus monartus (Plötz, 1884)  Dotted Flat  S Mexico to N Brazil  iN  M T S IL L G

Celaenorrhinus astrigera (A. Butler, 1877)    iN  M T S

Celaenorrhinus fritzgaertneri (Bailey, 1880)  Fritzgaertner’s Flat  SE AZ & S TX (as strays) to Costa Rica  iN  M T S1 S2 IL L I G

Celaenorrhinus stallingsi H. Freeman, 1946  Stallings’ Flat  S Texas (rare stray), E Mexico to Costa Rica  iN  M T S L I Ha

Celaenorrhinus shema (Hewitson, 1877)    iN

Celaenorrhinus shema shema (Hewitson, 1877)  TL: French Guiana  M T S IL

Celaenorrhinus shema ochra Evans, 1952  TL: Brazil (Am)  M T S

Celaenorrhinus shema vox Evans, 1952  TL: Peru  M T S

Celaenorrhinus shema mercedensis O. Mielke, 1992  TL: Peru  M T OI

Celaenorrhinus par Steinhauser & Austin, 1996  TL: Brazil (Rd)  M T OI

Celaenorrhinus disjunctus E. Bell, 1940  TL: Peru  M T S

Celaenorrhinus vagra Evans, 1952  TL: Peru  M T

Celaenorrhinus stola Evans, 1952  Stola Flat  E & W Mexico to Nicaragua  iN  M T S L

Celaenorrhinus approximatus R. Williams & E. Bell, 1940  Approximate Flat  Costa Rica and Panama  iN  M T S L I G

Celaenorrhinus bifurcus E. Bell, 1934  Bifurcate Flat  Costa Rica to Guianas, Trinidad, Ecuador  iN  M T S L

Celaenorrhinus similis Hayward, 1933    iN  M S IL G

Celaenorrhinus eligius (Stoll, 1781)  Eligius Flat    iN

Celaenorrhinus eligius eligius (Stoll, 1781)  Costa Rica to Guianas, Trinidad, Tobago, to Peru  M T S IL I G

Celaenorrhinus eligius punctiger (Burmeister, 1878)  TL: Argentina; Brazil (RJ)  M T S IL G

Celaenorrhinus songoensis Draudt, 1922    iN

Celaenorrhinus songoensis saroma Evans, 1952  TL: Colombia  M T S

Celaenorrhinus songoensis tonio Evans, 1952  TL: Colombia  M T

Celaenorrhinus songoensis songoensis Draudt, 1922  TL: Bolivia  M T OI S L G

Celaenorrhinus cynapes (Hewitson, 1870)  Small Flat    iN

Celaenorrhinus cynapes darius Evans, 1952  TL: Ecuador  M T S

Celaenorrhinus cynapes cynapes (Hewitson, 1870)  SE Mexico to Ecuador  M T G

Celaenorrhinus tritonae (Weeks, 1901)  TL: Bolivia  M TI S

Celaenorrhinus anchialus (Mabille, 1878)  TL: Brazil  M T S IL

Celaenorrhinus jao (Mabille, 1889)    iN  M T OI S

Celaenorrhinus autochton Steinhauser & Austin, 1996    iN  M T S

Celaenorrhinus suthina (Hewitson, 1877)    iN  M T OI S L

Celaenorrhinus syllius (C. Felder & R. Felder, 1862)    iN  M T S1 S2 S3 L

  Subfamily Heteropterinae

  Tribe Heteropterini

Carterocephalus  Lederer, 1853  

Carterocephalus mandan (W. H. Edwards, 1863)  Arctic Skipperling  SE SASK through Great Lakes region; ref.11  iN

Carterocephalus mandan mesapano (Scudder, 1868)  New England, NE Canadian Provinces  M S L G

Carterocephalus mandan mandan (W. H. Edwards, 1863)  SE SASK through Great Lakes region  M S L

Carterocephalus skada (W. H. Edwards, 1870)  Arctic Skipperling  Alaska to N CA; ref.11  iN

Carterocephalus skada S Rocky Mountain segregate ,  NW WY to ALTA  M S L Ha

Carterocephalus skada skada (W. H. Edwards, 1870)  Alaska to N CA  M T S L1 L2 Ha

Carterocephalus skada magnus Mattoon & Tilden, 1998  coastal Sonoma Co., CA  M T S

Dalla  Mabille, 1904  

Dalla calaon (Hewitson, 1877)  Calaon Skipperling  Ecuador  M T

Dalla thalia Evans, 1955  Thalia Skipperling  Ecuador  M T G

Dalla orsines (Hewitson, 1877)  Orsines Skipperling  Venezuela, Bolivia  M T S G

Dalla grovius (Mabille, 1898)  Grovius Skipperling    iN

Dalla grovius grovius (Mabille, 1898)  Ecuador (Loja), Peru  M T S

Dalla grovius amba Evans, 1955  Ecuador (Ambato)  M S

Dalla grovius floxa Evans, 1955  Peru (Amazonas, Chachapoyas)  M S

Dalla riza (Mabille, 1889)  Riza Skipperling  Peru, Bolivia  M T S

Dalla agathocles (C. Felder & R. Felder, 1867)  Agathocles Skipperling    iN

Dalla agathocles polydesma (Mabille, 1889)  Venezuela  M T S

Dalla agathocles agathocles (C. Felder & R. Felder, 1867)  Colombia (Bogota)  M S

Dalla agathocles lanna Evans, 1955  Colombia (Cauca), Ecuador, Peru  M S

Dalla agathocles lonia Evans, 1955  Peru (Huancabamba)  M S

Dalla lenda Evans, 1955  Lenda Skipperling  Ecuador (Zamora, Loja)  M S

Dalla lorda Evans, 1955  Lorda Skipperling  Ecuador (Zamora, Loja)  M T

Dalla costala Evans, 1955  Costala Skipperling    iN

Dalla costala ascha Evans, 1955  Peru (Huancabamba, Pozuzo, La Merced)  M S

Dalla costala zona Evans, 1955  Peru (Amazonas, Chachapoyas)  M S

Dalla costala costala Evans, 1955  Peru, Bolivia  M S

Dalla huanca Evans, 1955  Huancabamba Skipperling  Peru (Huancabamba)  M S

Dalla sepia Evans, 1955  Falcate Skipperling  Peru  M S

Dalla hesperioides (C. Felder & R. Felder, 1867)  Hesperioides Skipperling    iN

Dalla hesperioides hesperioides (C. Felder & R. Felder, 1867)  Colombia  M T S

Dalla hesperioides hister Evans, 1955  Peru (Limbani)  M T S

Dalla polycrates (C. Felder & R. Felder, 1867)  Polycrates Skipperling    iN

Dalla polycrates polycrates (C. Felder & R. Felder, 1867)  Colombia  M T

Dalla polycrates ambala Evans, 1955  Ecuador, Peru (Huancabamba)  M S

Dalla polycrates lania Evans, 1955  SE Peru (Aqualani), Bolivia  M S

Dalla superior Draudt, 1923  Superior Skipperling  Colombia, Ecuador  iN  M T S G

Dalla epiphaneus (C. Felder & R. Felder, 1867)  Epiphaneus Skipperling    iN

Dalla epiphaneus epiphaneus (C. Felder & R. Felder, 1867)  Colombia  M T S

Dalla epiphaneus gaujoni (Mabille, 1898)  S Ecuador, Peru  M T S

Dalla epiphaneus poya Evans, 1955  Peru (Amazonas)  M S

Dalla epiphaneus anca Evans, 1955  Peru (Huancabamba)  M S

Dalla epiphaneus limba Evans, 1955  Peru  M S

Dalla epiphaneus junga Evans, 1955  Bolivia  M T

Dalla cola E. Bell, 1959  Cola Skipperling  Colombia  M T

Dalla frontinia Evans, 1955  Frontinia Skipperling  TL: Colombia

Dalla frontinia venda Evans, 1955  Venezuela, [Peru ?]  M T S

Dalla frontinia frontinia Evans, 1955  Colombia, Ecuador  M S

Dalla frontinia vanca Evans, 1955  Peru  M S

Dalla taza Evans, 1955  Taza Skipperling  Colombia, Ecuador  M T S

Dalla pota E. Bell, 1959  Pota Skipperling  Ecuador  M T

Dalla bubobon (Dyar, 1921)  Bubobon Skipperling  S-C Mexico (COL, MICH, GRO)  iN  M T S

Dalla freemani A. Warren, 1997  Freeman's Skipperling  Guatemala; ref.100  M T G

Dalla austini A. Warren & R. G. Maza, 2011  Austin's Skipperling  S Mexico (Chiapas); ref.99  M T

Dalla amatlanensis A. Warren, Gott & Legal, 2020  ref.25

Dalla steinhauseri H. Freeman, 1991  Steinhauser's Skipperling  E Oaxaca  iN  M T L

Dalla pulchra (Godman, 1900)  Costa Rican Skipperling  Costa Rica  iN  M T S L

Dalla nubes Steinhauser, 1991  Chiapan Skipperling  S Mexico (OAX & CHIS)  M T S L

Dalla ligilla (Hewitson, 1877)  Ligilla Skipperling  W (JAL) & S Mexico to Costa Rica  iN  M T S L

Dalla dividuum (Dyar, 1913)  Dyar's Skipperling  W Mexico (SON to CHIS)  iN  M T S L Ha

Dalla dimidiatus (C. Felder & R. Felder, 1867)  Dimidiatus Skipperling    iN

Dalla dimidiatus dimidiatus (C. Felder & R. Felder, 1867)  Venezuela, Colombia, Ecuador  M T S

Dalla dimidiatus lilla Evans, 1955  S Ecuador, Peru, Bolivia  M S

Dalla dimidiatus pucer Evans, 1955  Peru (Huancabamba)  M S

Dalla cocha Evans, 1955  Cochabamba Skipperling  Peru, Bolivia  iN  M S L

Dalla miser Evans, 1955  Miser Skipperling  Colombia, Peru  iN  M S L

Dalla caicus (Hewitson, 1868)  Caicus Skipperling    iN

Dalla caicus caicus (Hewitson, 1868)  Venezuela  M T S

Dalla caicus inca Draudt, 1923  Colombia, Ecuador, Peru, Bolivia  M T S L G

Dalla hilina (A. Butler, 1870)  Hilina Skipperling  Colombia, Venezuela  M T

Dalla dognini (Mabille, 1889)  Dognin's Skipperling  Ecuador  iN  M T S L

Dalla scylla (Mabille, 1898)  Scylla Skipperling  Peru, Bolivia, N Argentina  iN  M T S1 S2

Dalla genes (Mabille, 1898)  Genes Skipperling    iN

Dalla genes saleca (Mabille, 1898)  Colombia  M T S

Dalla genes genes (Mabille, 1898)  S Ecuador  M T S

Dalla genes golia Evans, 1955  N Peru  M T S

Dalla genes nona Evans, 1955  S Peru, Bolivia  M T S

Dalla cyprius (Mabille, 1898)  Cyprius Skipperling    iN

Dalla cyprius quinka Evans, 1955  Peru  M S

Dalla cyprius cyprius (Mabille, 1898)  Bolivia  M T S

Dalla vinca Evans, 1955  Vinca Skipperling  TL: Peru  M T S

Dalla cypselus (C. Felder & R. Felder, 1867)  Cypselus Skipperling    iN

Dalla cypselus cypselus (C. Felder & R. Felder, 1867)  Venezuela, Colombia  M T S

Dalla cypselus evages (Hewitson, 1877)  Ecuador, Peru, Bolivia  M T S

Dalla dora E. Bell, 1947  Dora Skipperling  Ecuador  iN  M T S

Dalla spica Hayward, 1939  Spica Skipperling    iN

Dalla spica livia Evans, 1955  Peru, Bolivia  M T S

Dalla spica spica Hayward, 1939  Argentina  M S

Dalla quasca E. Bell, 1947  Quasca Skipperling    iN

Dalla quasca quasca E. Bell, 1947  Venezuela, Colombia  M T S

Dalla quasca equatoria E. Bell, 1947  Ecuador  M T S

Dalla pincha Steinhauser, 1991  Pinche Skipperling  Ecuador  M T S

Dalla gelus (Mabille, 1898)  Gelus Skipperling  Ecuador  iN  M T S HS

Dalla cupavia (Mabille, 1898)  Cupavia Skipperling    iN

Dalla cupavia elka Evans, 1955  N & C Peru  M S

Dalla cupavia cupavia (Mabille, 1898)  S Peru, Bolivia  M T S

Dalla frater (Mabille, 1878)  Frater Skipperling  Panama to Bolivia  iN  M T S1 S2 L

Dalla mesoxantha (Plötz, 1884)  Mesoxantha Skipperling  Venezuela, Colombia, Ecuador, Peru, Bolivia  iN  M T S

Dalla wardi Steinhauser, 2002  Ward's Skipperling  Colombia (Magdalena, 8 km E San Pedro, 2550m)  M T OD S G

Dalla merida Evans, 1955  Merida Skipperling  Venezuela, Colombia  M T S

Dalla xantha Steinhauser, 1991  Xantha Skipperling  Colombia  iN  M T L

Dalla geon (Mabille, 1898)  Geon Skipperling  Ecuador  M T S

Dalla octomaculata (Godman, 1900)  Eight-spotted Skipperling  Costa Rica  M S

Dalla oxaites (Hewitson, 1877)  Oxaites Skipperling  Peru, Bolivia  iN  M T S

Dalla eryonas (Hewitson, 1877)  Central American Skipperling  Costa Rica to Colombia  iN  M T S L

Dalla mentor Evans, 1955  Guerrero Skipperling  SW Mexico (Guerrero & Oaxaca)  M S

Dalla kemneri Steinhauser, 1991  Pelon Skipperling  SE Mexico (Oaxaca & Puebla)  iN  M T S L

Dalla lalage (Godman, 1900)  Godman’s Skipperling  SW Mexico (CHIS, GRO, OAX)  iN  M S L

Dalla lethaea (Schaus, 1913)  Schaus' Skipperling  S Mexico (CHIS, GRO, OAX, PUE) to Costa Rica  iN  M T OI S L

Dalla faula (Godman, 1900)  West-Mexican Skipperling  W Mexico (SON to MICH & PUE)  iN  M S L

Dalla lysis (Schaus, 1913)  Lysis Skipperling  Costa Rica  M OI

Dalla diraspes (Hewitson, 1877)  Widespread Skipperling  Costa Rica to S Brazil  iN  M T S L

Dalla ramirezi H. Freeman, 1969  Gold-rayed Skipperling  S Mexico (OAX, PUE, VER) to Guatemala  iN  M T S L Ha

Dalla jelskyi (Erschoff, 1875)  Jelsky's Skipperling    iN

Dalla jelskyi jelskyi (Erschoff, 1875)  Peru  M S

Dalla jelskyi aurosa J. Zikán, 1938  S Brazil  M S

Dalla xicca (Dyar, 1913)  Xicca Skipperling  TL: Peru

Dalla xicca xicca (Dyar, 1913)  Peru  M T S

Dalla xicca paza Evans, 1955  Bolivia  M T S

Dalla semiargentea (C. Felder & R. Felder, 1867)  Half-silvered Skipperling  Colombia (Cordillera Oriental)  M T S

Dalla superargentea Viloria, A. Warren & Austin, 2008  Silvered Skipperling  Serrania de Valledupar, Sierra de Perija, Colombia-Venezuela; ref.101  M T

Ladda  Grishin, 2019  ref.5

Ladda pedro (Steinhauser, 2002)  Pedro Skipperling  Colombia (Magdalena, 8 km E San Pedro, 2550m)  iN  M T OD G

Ladda connexa (Draudt, 1923)  Connexa Skipperling  Colombia, Ecuador, Peru  iN  M T S G Txt

Ladda disconnexa (Steinhauser, 2002)  Disconnexa Skipperling  Ecuador  iN  M T OD G

Ladda vista (Steinhauser, 2002)  Vista Skipperling  Ecuador  M T OD S G

Ladda ticidas (Mabille, 1898)  Ticidas Skipperling  Peru, Bolivia  iN  M T S

Ladda caenides (Hewitson, 1868)  Caenides Skipperling  Venezuela, Colombia, Ecuador  iN  M T S

Ladda bos (Steinhauser, 1991)  Bos Skipperling  Colombia (Cauca)  M T

Ladda pura (Steinhauser, 1991)  Pura Skipperling  Colombia (Cauca)  M T

Ladda mora (Evans, 1955)  Mora Skipperling  Ecuador, N Peru  iN  M T S G

Ladda carnis (Evans, 1955)  Carnis Skipperling    iN

Ladda carnis sondra (Evans, 1955)  Peru  M T S

Ladda carnis carnis (Evans, 1955)  S Peru, Bolivia  M S

Ladda monospila (Mabille, 1898)    iN  M T S

Ladda simplicis (Steinhauser, 1991)  Simple Skipperling  Colombia  M T S

Ladda seirocastnia (Draudt, 1923)  Seirocastnia Skipperling  Colombia, Ecuador  M T S G

Ladda celsus (Steinhauser, 2002)  Celsus Skipperling  Ecuador (Chimborazo)  M T OD G

Ladda pantha (Evans, 1955)  Pantha Skipperling  N Peru (Huancabamba)  M T

Ladda puracensis (Steinhauser, 1991)  Purace Skipperling  TL: Colombia

Ladda puracensis puracensis (Steinhauser, 1991)  Colombia  M T S

Ladda puracensis cotopa (Steinhauser, 1991)  Ecuador  M T

Ladda puracensis quindio (Steinhauser, 1991)  Colombia  M T

Ladda ochrolimbata (Draudt, 1923)  Ochro Skipperling  Peru  M T G

Ladda tona (Evans, 1955)  Tona Skipperling  Peru (Huancabamba)  M T

Ladda decca (Evans, 1955)  Decca Skipperling    iN

Ladda decca decca (Evans, 1955)  Colombia, Ecuador  M S

Ladda decca doppa (Evans, 1955)  Peru  M S

Ladda eburones (Hewitson, 1877)  Eburones Skipperling  ref.5  iN

Ladda eburones inornata (E. Bell, 1937)  Colombia, Ecuador  M T S

Ladda eburones ena (Evans, 1955)  Peru (Amazonas, Chachapoyas)  M T S

Ladda eburones eburones (Hewitson, 1877)  SE Peru, Bolivia  M T S

Ladda morva (Mabille, 1898)  Morva Skipperling  Peru, Bolivia  M T S

Ladda ibhara (A. Butler, 1870)  Ibhara Skipperling  Venezuela, Colombia, Ecuador, Peru, Bolivia  iN  M T S

Ladda plancus (Hopffer, 1874)  Plancus Skipperling  Peru, Bolivia  iN  M T S L

Ladda crithote (Hewitson, 1874)  Crithoe Skipperling  Colombia, Ecuador  iN  M T S

Ladda calima (Steinhauser, 1991)  Calima Skipperling  Colombia (Valle del Cauca)  iN  M T

Ladda rosea (Evans, 1955)  Rosy Skipperling  Colombia  M T S

Ladda granites (Mabille, 1898)  Granites Skipperling  TL: Ecuador

Ladda granites granites (Mabille, 1898)  Ecuador  M T

Ladda granites privata (Draudt, 1923)  Bolivia  M OI S

Ladda parma (Evans, 1955)  Parma Skipperling  Ecuador  M T S G

Ladda mars (Evans, 1955)  Mars Skipperling  Colombia  M T

Ladda cuadrada (Weeks, 1901)  Bolivian Skipperling  Bolivia  iN  M T OD TI Txt

Ladda quadristriga (Mabille, 1889)  Quadristriga Skipperling    iN

Ladda quadristriga quadristriga (Mabille, 1889)  Colombia, Ecuador  M T S G

Ladda quadristriga regia (Evans, 1955)  Peru (Amazonas, Chachapoyas)  M T

Ladda quadristriga rota (Evans, 1995)  SE Peru  M T S

Ladda quadristriga rusa (Evans, 1955)  Bolivia  M T S

Ladda rubia (Evans, 1955)  Rubia Skipperling  Peru (Huancabamba)  M T

Freemaniana  A. Warren, 2001  

Freemaniana rawlinsi (A. Warren, 2001)  Rawlins' Skipperling  Ecuador (Zamora); ref.102,103  M S

Dardarina  Evans, 1937  

Dardarina dardaris (Hewitson, 1877)  Dardaris Skipperling  NW Mexico (SON) to Costa Rica  iN  M T S L I Ha

Dardarina amadryas Hayward, 1942  Amadryas Skipperling  Brazil  iN  M S

Dardarina tessellatus (Hayward, 1934)  Argentine Skipperling  Argentina  iN  M S

Dardarina salta Hayward, 1937  Salta Skipperling  Argentina  iN  M S

Dardarina daridaeus (Godman, 1900)  Daridaeus Skipperling  S Brazil, Paraguay, Argentina  iN  M T S

Dardarina rana Evans, 1955  Rana Skipperling  S Brazil (Pr)  iN  M S

Dardarina castra Evans, 1955  Castra Skipperling  S Brazil (Pr)  M S

Dardarina umuarama O. Mielke, 1978  Umuarama Skipperling  S Brazil (SP)  iN  M T

Dardarina jonesi Evans, 1955  Jones' Skipperling  S Brazil (Pr)  M S

Dardarina angeloi O. Mielke & Casagrande, 2005  Angelo's Skipperling  SE & S Brazil (SP)

Dardarina angeloi angeloi O. Mielke & Casagrande, 2005  SE Brazil (SP)  M T

Dardarina angeloi machadoi O. Mielke & Casagrande, 2005  S Brazil (SC)  M T

Dardarina gaucha O. Mielke, 1980  Gaucha Skipperling  S Brazil (RS)  iN  M T

Dardarina para O. Mielke, 1968  Mielke's Skipperling  S Brazil (MG)  M T

Dardarina aspila O. Mielke, 1966  Aspila Skipperling  S Brazil (Pr)  iN  M T S

Piruna  Evans, 1955  

Piruna pirus (W. H. Edwards, 1878)  Russet Skipperling  SE ID, W & S WY, CO, UT, AZ, NM, (W TX?)  iN  M T S L I Ha G

Piruna cyclosticta (Dyar, 1920)  Plateau Skipperling  Mexican Plateau to DF and MOR  iN  M T S

Piruna haferniki H. Freeman, 1970  Chisos Skipperling  W TX to Hidalgo  iN  M T S L I Ha

Piruna kemneri H. Freeman, 1990  Kemner's Skipperling  S Mexico (Oaxaca, Puebla)  iN  M T L

Piruna dampfi (E. Bell, 1942)  Violet-dusted Skipperling  SW & Transvolcanic Mexico (GRO, MICH, MEX, VER)  iN  M T S

Piruna gyrans (Plötz, 1884)  Variable Skipperling  W, C & S Mexico to Oaxaca  iN  M T OI S L

Piruna ceracates (Hewitson, 1874)  Veracruz Skipperling  S Mexico (Veracruz & Oaxaca)  iN  M S L

Piruna bailowitzi A. Warren & Gott, 2021  ref.27  iN

Piruna roeveri (L. Miller & J. Miller, 1972)  Roever's Skipperling  E & S Mexico (Coahuila to Oaxaca)  iN  M T1 T2 S L Ha

Piruna temascaltepeca A. Warren & Gott, 2021  ref.27

Piruna brunnea (Scudder, 1872)  Chocolate Skipperling  E and S Mexico to Costa Rica  iN  M S L

Piruna purepecha A. Warren & L. González, 1999  Purepecha Skipperling  SW Mexico (COL, GRO, JAL, MICH, MOR)  iN  M T1 T2 S L

Piruna mullinsi H. Freeman, 1991  Mullins' Skipperling  S Mexico (Oaxaca, Puebla)  iN  M T S L Ha

Piruna sina H. Freeman, 1970  Fine-spotted Skipperling  NW Mexico (DGO, SIN, SON)  iN  M T S L

Piruna uruapana A. Warren & Gott, 2021  ref.27

Piruna chinicuila A. Warren & Gott, 2021  ref.27

Piruna microsticta (Godman, 1900)  Southwest-Mexican Skipperling  [= ajijiciensis], SW Mexico (JAL to CHIS)  iN  M T S L Ha

Piruna penaea (Dyar, 1918)  Hour-glass Skipperling  W Mexico to Oaxaca  iN

Piruna penaea penaea (Dyar, 1918)  TL: Mexico (Gue)  M T S L Ha

Piruna penaea E Mexican segregate ,  S TX (one-time stray), E Mexico to Chiapas  M S L

Piruna aea (Dyar, 1912)  Many-spotted Skipperling    iN

Piruna aea aea (Dyar, 1912)  C & S Mexico to Guatemala  M T S L Ha

Piruna aea mexicana H. Freeman, 1979  [= cingo], NW Mexico to SE AZ  M T S L I

Piruna millerorum Steinhauser, 1991  Millers' Skipperling  NW Mexico (DGO, SIN, SON, CHIH)  M T S L

Piruna maculata H. Freeman, 1970  Sinaloan Skipperling  NW Mexico (Durango & Sinaloa)  M T S

Piruna polingii (W. Barnes, 1900)  Four-spotted Skipperling  C AZ, S NM to Guerrero and Puebla  iN  M T S L1 L2 I Ha G

Piruna jonka Steinhauser, 1991  Oaxacan Skipperling  S Mexico (Oaxaca, Puebla)  iN  M T S L

  Tribe Butleriini

Butleria  W. F. Kirby, 1871  

Butleria paniscoides (Blanchard, 1852)  Paniscoides Skipperling    iN

Butleria paniscoides polyspilus (C. Felder, 1862)  Chile, Argentina (Neuquen, Rio Negro)  M T S

Butleria paniscoides paniscoides (Blanchard, 1852)  Chile (widespread)  M T S

Butleria elwesi Evans, 1939  Elwes' Skipperling  Chile, Argentina (Neuquen, Rio Negro, Chubut)  iN  M T S

Butleria flavomaculatus (Blanchard, 1852)  Blanchard's Skipperling    iN

Butleria flavomaculatus flavomaculatus (Blanchard, 1852)  Chile (widespread)  M S

Butleria flavomaculatus tristriata Bryk, 1944  Chile (Valdivia, Osorno, Llanquihue)  M T

Butleria flavomaculatus valdivianus (Philippi, 1859)  Chile (Chiloe, Coyhaique)  M S

Butleria quilla Evans, 1939  Quilla Skipperling  Chile (widespread), Argentina (Neuquen, Rio Negro)  iN  M T S

Butleria sotoi Reed, 1877  Soto's Skipperling  Chile (Valparaiso, Cachapoal, Curico, Cauquenes, Linares, Nuble, Concepcion)  M T S

Butleria philippii (A. Butler, 1881)  Philipp's Skipperling  Chile (Nuble, Concepcion, Arauco, Araucania, Valdivia, Osorno)  iN  M T

Butleria fruticolens (A. Butler, 1881)  Butler's Skipperling  Chile, Argentina (Neuquen, Rio Negro)  iN  M T S HS

Butleria bissexguttatus (Philippi, 1859)  Herrera's Skipperling  Chile (widespread), Argentina (Neuquen, Rio Negro)  iN  M S

Argopteron  E. Watson, 1893  

Argopteron aureipennis (Blanchard, 1852)  Gold-plated Skipperling  Chile, Argentina  iN  M T S IL L

Argopteron aureum L. Peña, 1968  Gold-washed Skipperling  Chile  M Ma

Argopteron puelmae (Calvert, 1888)  Gold-winged Skipperling  Chile  iN  M S

  Subfamily Hesperiinae

  Tribe Erionotini

Erionota  Mabille, 1878  

Erionota thrax (Linnaeus, 1767)  Banana Skipper    iN

Erionota thrax thrax (Linnaeus, 1767)  Introduced to Hawaii  M S IL

  Tribe Hesperiini, Subtribe Hesperiina

Euphyes  Scudder, 1872  

Euphyes vestris (Boisduval, 1852)  Dun Skipper    iN

Euphyes vestris harbisoni J. Brown & McGuire, 1983  S CA; ref.13  M T1 T2 T3 S L

Euphyes vestris vestris (Boisduval, 1852)  N CA to BC, ID; ref.13  M T S HS L I Ha G

Euphyes vestris osceola (Lintner, 1878)  ref.146  M S

Euphyes vestris metacomet (T. Harris, 1862)  E USA, SE Canada, NE Mexico; univoltine to multivoltine; ref.13  M S L1 L2 L3 L4 I

Euphyes kiowah (Reakirt, 1866)  Dun Skipper  SE WY, CO, NM, AZ, far NW Mexico; univoltine; ref.13

Euphyes kiowah kiowah (Reakirt, 1866)  SE WY, CO, NM, AZ, far NW Mexico; univoltine; ref.13  M T S L

Euphyes kiowah chamuli H. Freeman, 1969  Chiapas & Guatemala; ref.13  iN  M T S

Euphyes bimacula (Grote & Robinson, 1867)  Two-spotted Skipper    iN

Euphyes bimacula illinois (Dodge, 1872)  IL, IA, S SD, NE, NW KS, NE CO  M T S1 S2

Euphyes bimacula bimacula (Grote & Robinson, 1867)  New England and ONT to VA  M S L1 L2 Ha G

Euphyes bimacula arbogasti Gatrelle, 1999  South Carolina (Berkeley Co.)  M T1 T2

Euphyes arpa (Boisduval & Le Conte, [1837])  Palmetto Skipper  FL, S GA, S AL, S MS  iN  M T S L G

Euphyes pilatka (W. H. Edwards, 1867)  Palatka Skipper    iN

Euphyes pilatka pilatka (W. H. Edwards, 1867)  VA to FL, W to MS  M T S L I Ha G

Euphyes pilatka klotsi L. Miller, Harvey & J. Miller, 1985  lower Florida Keys  M T1 T2 S I

Euphyes berryi (E. Bell, 1941)  Berry's Skipper  FL, GA, coastal SC, NC  iN  M T S L Ha G

Euphyes conspicua (W. H. Edwards, 1863)  Black Dash    iN

Euphyes conspicua orono (Scudder, 1872)  New England  M S L

Euphyes conspicua conspicua (W. H. Edwards, 1863)  upper midwest USA (OH, MI, IN, IL, WI)  M S L G

Euphyes conspicua buchholzi (P. Ehrlich & Gillham, 1951)  NE, OK, IA  M T1 T2 S

Euphyes dion (W. H. Edwards, 1879)  Dion Skipper  ONT & NY to FL, W to E TX, OK, KS, NE, SD & ND  iN  M T S L1 L2 L3 L4 G

Euphyes bayensis Shuey, 1989  Bay Skipper  S MS, S LA, SE TX  iN  M T S L

Euphyes dukesi (Lindsey, 1923)  Dukes' Skipper    iN

Euphyes dukesi dukesi (Lindsey, 1923)  AR, SE TX to AL, NC, VA, OH, MI  M T S L1 L2 G

Euphyes dukesi calhouni Shuey, 1996  Peninsular Florida  M T S L Ha

Euphyes cherra Evans, 1955  Cherra Skipper  Venezuela, Bolivia, S Brazil  M T S

Euphyes subferrugineus (Hayward, 1934)  Subferrugineus Skipper  TL: Argentina

Euphyes subferrugineus subferrugineus (Hayward, 1934)  S Brazil & N Argentina

Euphyes subferrugineus biezankoi O. Mielke, 1972  S Brazil (MG to SC)  M T S

Euphyes antra Evans, 1955  Antra Skipper  SE Mexico (OAX) to Peru  iN  M T S L

Euphyes ampa Evans, 1955  Ampa Skipper  S Mexico to Honduras, Nicaragua  M T S

Euphyes canda Steinhauser & A. Warren, 2002  Candelaria Skipper  W Mexico (Nay to Oax); ref.104  iN  M T S L

Euphyes leptosema (Mabille, 1891)  Leptosema Skipper  S Brazil, Paraguay, Argentina  iN  M T S

Euphyes fumata O. Mielke, 1972  Fumata Skipper  SE Brazil (Pr)  iN  M T

Euphyes eberti O. Mielke, 1972  Ebert's Skipper  S Brazil (MG)  M T

Euphyes peneia (Godman, 1900)  Guardpost Skipper  E & W Mexico to Ecuador, Peru & N Brazil, Trinidad  iN  M T S1 S2

Euphyes cornelius (Latreille, [1824])  Cornelius Skipper    iN

Euphyes cornelius agra Evans, 1955  Bahamas  M S L

Euphyes cornelius cornelius (Latreille, [1824])  Cuba  M T S

Euphyes singularis (Herrich-Schäffer, 1865)  Singularis Skipper    iN

Euphyes singularis singularis (Herrich-Schäffer, 1865)  Cuba  M S

Euphyes singularis insolata (A. Butler, 1878)  Jamaica, Hispaniola, Puerto Rico  M T S HS

Mnaseas  Godman, 1901  [includes Arotis as its synonym]; ref.13

Mnaseas mapirica (E. Bell, 1930)  Mapirica Skipper  Bolivia, S Brazil; ref.13  M T S

Mnaseas pandora (Lindsey, 1925)  Pandora Skipper  Peru, N Brazil; ref.13  M T S

Mnaseas derasa (Herrich-Schäffer, 1870)  Derasa Skipper    iN

Mnaseas derasa derasa (Herrich-Schäffer, 1870)  [= infuscata] C to SE Brazil (Brasilia to SP, ES & RJ); ref.13  M T S

Mnaseas derasa brunnea (O. Mielke, 1972)  SE Brazil (Pr, SC)  M T S

Mnaseas derasa tuba (Evans, 1955)  Amazonian Brazil (Para)  M T

Mnaseas bryna (Evans, 1955)  Bryna Skipper  Guyana, French Guiana, N & NE Brazil; ref.13  M T

Mnaseas kayei (E. Bell, 1932)  Kaye's Skipper  Trinidad, Guianas, Venezuela, Brazil (Amapa to MG & ES); ref.13  M T S

Mnaseas sirene (Mabille, 1904)  Sirene Skipper  SE Brazil; ref.13  iN  M T S HS

Mnaseas evansi (O. Mielke, 1972)  Hidden Skipper  French Guiana; ref.13  M T

Mnaseas bias (Plötz, 1882)  no specimen known, S. Amer.  M TI

Mnaseas bicolor (Mabille, 1889)  Dull Skipper  SE Mexico to Panama  M T OI S

Mnaseas macia Evans, 1955  Macia Skipper  Nicaragua, Colombia, Ecuador  M T S

Mnaseas inca E. Bell, 1930  Bolivia; ref.13  M T S

Serdis  Mabille, 1904  

Serdis statius (Plötz, 1883)  Muted Serdis    iN

Serdis statius statius (Plötz, 1883)  Venezuela  M T S

Serdis statius flagrans Mabille, 1904  Ecuador, N Brazil  M T TI S

Serdis statius dives Evans, 1955  Peru, Bolivia  M S

Serdis venezuelae (Westwood, 1852)  Venezuelan Serdis    iN

Serdis venezuelae fractifascia (C. Felder & R. Felder, 1867)  Colombia

Serdis venezuelae venezuelae (Westwood, 1852)  Venezuela, Ecuador  M T

Serdis venezuelae fulla Evans, 1955  Colombia, Ecuador  M S

Serdis viridicans (C. Felder & R. Felder, 1867)  Green-washed Serdis    iN

Serdis viridicans viridicans (C. Felder & R. Felder, 1867)  Colombia  M S

Serdis viridicans kirschi (Weymer, 1890)  Colombia, Ecuador (Zamora)  M T

Serdis viridicans lupus Evans, 1955  Venezuela

Serdis viridicans pandra Evans, 1955  Colombia, Ecuador  M S

Serdis viridicans edith Evans, 1955  Peru (Amazonas)

Serdis viridicans junia Evans, 1955  N Peru  M S

Pompe  Grishin & Lamas, 2022  ref.16

Pompe postpuncta (Draudt, 1923)  Bolivia, S Brazil (MT, Pr), Paraguay, Uruguay; ref.16

Cyclosma  Draudt, 1923  

Cyclosma glamis Evans, 1955  Glamis Skipper  Peru  iN  M T

Cyclosma altama (Schaus, 1902)  Altama Skipper  S Brazil, Paraguay  iN  M T S

Cyclosma abdonides Draudt, 1923  Abdonides Skipper  Peru, Bolivia  iN  M T S G

Caligulana  E. Bell, 1942  

Caligulana caligula (Schaus, 1902)  Caligula Skipper  SE Brazil  iN  M T

Chloeria  Mabille, 1904  

Chloeria psittacina (C. Felder & R. Felder, 1867)    iN  M S

Tirynthoides  E. Bell, 1940  

Tirynthoides lotana (A. Butler, 1870)  Lotana Skipper  Surinam, Venezuela, Colombia, Peru, Brazil  iN  M T

Tirynthoides virilis (N. Riley, 1929)  Virilis Skipper  E Ecuador, Brazil, Paraguay, Argentina  iN  M T S L

Testia  Grishin, 2019  ref.5

Testia mammaea (Hewitson, 1876)  Mammaea Skipper  SE Brazil; ref.13  M T S

Testia potesta (E. Bell, 1941)  Potesta Skipper  Amazonian Peru, Brazil; ref.5  M T S

Oxynthes  Godman, 1900  [includes Oeonus as a subgenus]; ref.140

Oxynthes pyste (Godman, 1900)  Veracruzan Skipper  SE Mexico; ref.140  M T S

Oxynthes edlichi (Plötz, 1886)  Edlich's Skipper  Brazil  M S

Oxynthes egma (Evans, 1955)  Egma Skipper  E Brazil (Espirito Santo); ref.140  M T

Oxynthes trinka (Evans, 1955)  Trinka Skipper  Guyana, Brazil (Am); ref.13  iN  M T

Oxynthes martius (Mabille, 1889)  E & S Mexico to Panama; ref.16  M T OI S L

Oxynthes corusca (Herrich-Schäffer, 1869)  Corusca Skipper  S. Am. (to Bolivia & Brazil); ref.16  iN  M T L

Lindra  Evans, 1955  

Lindra simulius (H. Druce, 1876)  Simulius Skipper  Peru (Cuzco), Bolivia  M S

Lindra procax (Draudt, 1923)  Procax Skipper  Brazil (Mato Grosso), Bolivia  M T G

Lindra boliviana O. Mielke, 1993  Bolivian Skipper  Bolivia  iN  M S

Lindra vanewrighti O. Mielke, 1978  Vane-Wright's Skipper  Peru (Cuzco)  iN  M T S

Lindra brasus (O. Mielke, 1968)  Brasus Skipper  Belize, Panama, Brazil (DF)  M T S

Lindra neroides (Herrich-Schäffer, 1869)  Neroides Skipper  [not Decinea] S Brazil; ref.140

Lindra neroides neroides (Herrich-Schäffer, 1869)  [= ackeryi] S Brazil; ref.140  M T

Lindra neroides huxleyi O. Mielke, 1978  Brazil (Pr); ref.140  M T

Lindra olafmielkei A. Warren, Gott & Dolibaina, 2021  ref.28

Noxys  Grishin, 2022  ref.13

Noxys viricuculla (Hayward, 1951)  Viricuculla Skipper  Peru & Bolivia; ref.13  iN  M S

Metrocles  Godman, 1900  [includes Propertius, Hansa & Chalcone]; ref.13

Metrocles leucogaster Godman, 1900  Whitened Metron    iN

Metrocles leucogaster leucogaster Godman, 1900  Panama, Colombia; ref.13  M OI OD S

Metrocles leucogaster ambrosei (Weeks, 1906)  Venezuela

Metrocles leucogaster leucomelas (Röber, 1925)  Brazil (RS)  M T

Metrocles briquenydan (Weeks, 1901)  Brick Skipper    iN

Metrocles briquenydan briquenydan (Weeks, 1901)  Bolivia; ref.13

Metrocles briquenydan chalcone (Schaus, 1902)  Brazil (RJ); ref.13  M T S

Metrocles briquenydan australis (O. Mielke, 1980)  Brazil (RS)  M T

Metrocles devergens (Draudt, 1923)  Devergens Skipper    iN

Metrocles devergens devergens (Draudt, 1923)  Colombia, Guyana; ref.13  M T G

Metrocles devergens hydra (Evans, 1955)  SE Brazil  M S

Metrocles cotundo (Nicolay, 1980)  Cotundo Skipper  Ecuador; ref.13  M T S

Metrocles santarus (E. Bell, 1940)  Santarus Skipper  SE Brazil; ref.13  M T S

Metrocles scitula (Hayward, 1951)  Brazil; ref.13

Metrocles schrottkyi (Giacomelli, 1911)  Argentina; ref.13

Metrocles nun Grishin, 2024  C Brazil (at least); ref.154  M T

Metrocles argentea (Weeks, 1901)  Weeks' Skipper  Bolivia; ref.13  iN  M T

Metrocles zisa (Plötz, 1882)  Zisa Skipper  S Brazil, N Argentina; ref.13  M S

Metrocles hyboma (Plötz, 1886)  Hyboma Skipper  S Brazil; ref.13  iN  M S

Metrocles propertius (Fabricius, 1793)  Propertius Skipper  Peru, Brazil, Paraguay, Argentina; ref.13  iN  M T

Metrocles dejongi (Austin, 2008)  de Jong's Skipper  Brazil (Rd); ref.120,13

Metrocles phineus (Cramer, 1777)  Phineus Skipper  Guianas, Venezuela, Peru, Brazil (Rd); ref.13  M T

Metron  Godman, 1900  

Metron voranus (Mabille, 1891)  [= tania]; ref.13  M T

Metron chrysogastra (A. Butler, 1870)  Orange-headed Metron    iN

Metron chrysogastra chrysogastra (A. Butler, 1870)  E & W Mexico to Colombia, Venezuela, Trinidad  M T S L

Metron chrysogastra gades Evans, 1955  Peru, Bolivia  M S

Metron chrysogastra hypodesma (Plötz, 1882)  Brazil (Amazonas, Espirito Santo)  M T

Metron noctis (Kaye, 1914)  Kaye's Metron  Trinidad, Ecuador, Brazil, Paraguay, Argentina  M T S

Metron oropa (Hewitson, 1877)  Hewitson's Metron  S Brazil, Paraguay  iN  M T

Metron hypochlora (Draudt, 1923)  Peru; ref.13

Metron hypochlora tomba Evans, 1955  S Brazil (MT)

Metron hypochlora hypochlora (Draudt, 1923)  Peru; ref.13  M S

Metron fasciata (Möschler, 1877)  Suriname; ref.13  M T

Metron zimra (Hewitson, 1877)  Olive Metron  S Mexico (Chia) to Argentina  iN  M S

Phemiades  Hübner, [1819]  [= Libra]; ref.153

Phemiades aligula (Schaus, 1902)  Aligula Skipper    iN

Phemiades aligula aligula (Schaus, 1902)  S Brazil  M T S

Phemiades aligula decia (Hayward, 1948)  SE Brazil, Paraguay, Argentina  M S

Phemiades anatolica (Plötz, 1883)  Anatolian Skipper  SE Brazil  iN  M T S

Phemiades pseudophineus de Jong, 1983  Impostor Skipper  S Brazil (MT)

Phemiades rufescens Steinhauser & Austin, 1993  Rufous Skipper  Chiapas, Mexico and Costa Rica  M T S

Phemiades pohli (E. Bell, 1932)  Pohl's Skipper    iN

Phemiades pohli cidra Evans, 1955  Ecuador

Phemiades pohli pohli (E. Bell, 1932)  S Brazil, Paraguay, Argentina  M T

Phemiades vergens Evans, 1955  Vergens Skipper  Peru (Cosnipata)  M S

Phemiades milvius (Mabille, 1904)  Milvius Skipper  TL: Brazil

Phemiades milvius milor Evans, 1955  Peru  M S

Phemiades milvius milvius (Mabille, 1904)  Guyana, Brazil

Notamblyscirtes  Scott, 2006  

Notamblyscirtes simius (W. H. Edwards, 1881)  Simius Skipper  S SASK, E MT, E WY, W SD, W NE, E CO, N & W TX, NM, C AZ; ref.16  iN  M T S1 S2 L G

Notamblyscirtes durango Scott, 2017  Durango Skipper  SE AZ, COAH, DGO; ref.16  M S L

Holguinia  Evans, 1955  

Holguinia holguin Evans, 1955  Holguin Skipper  Cuba  M T S

Onespa  Steinhauser, 1974  

Onespa nubis Steinhauser, 1974  Nubis Skipper  S Mexico (OAX, CHIS) to El Salvador  iN  M T S L

Onespa gala (Godman, 1900)  Gala Skipper  C Mexico (DF, GRO, MICH, VER)  iN  M S Ha

Onespa brockorum Austin & A. Warren, 2009  Brocks’ Skipper  NW Mexico (Son, Sin, possibly Dgo); ref.105  M T L

Onespa nakamura Austin & A. Warren, 2009  Nakamura’s Skipper  Costa Rica (San Jose Prov.: Bajo la Hondura); ref.105  M T

Neposa  Austin & A. Warren, 2009  

Neposa heras (Godman, 1900)  Heras Skipper  SW Mexico (GRO, OAX)  iN  M T S

Neposa hestia Austin & A. Warren, 2009  Hestia Skipper  SE Mexico (VER); ref.105  iN  M T

Neposa armandoi Austin & A. Warren, 2009  Armando’s Skipper  SW Mexico (OAX); ref.105  iN  M T

Neposa isabelae Austin & A. Warren, 2009  Isabel’s Skipper  Guatemala; ref.105  M T

Buzyges  Godman, 1900  

Buzyges idothea Godman, 1900  Idothea Skipper  Costa Rica  iN  M T S L

Buzyges rolla (Mabille, 1883)  Rolla Skipper  Costa Rica and Panama  M T S L

Buzyges benito (H. Freeman, 1979)  Benito's Skipper  SW Mexico (COL, GRO, JAL, OAX)  iN  M T S

Buzella  Grishin, 2019  ref.5

Buzella mellanaformis (Austin & A. Warren, 2009)  Mellana-like Skipper  Costa Rica; ref.105,5  M T

Librita  Evans, 1955  

Librita librita (Plötz, 1886)  Librita Skipper  W & S Mexico to Guatemala  iN  M T S Ha

Decinea  Evans, 1955  

Decinea huasteca (H. Freeman, 1969)  Huasteca Skipper  E Mexico  iN  M T S L

Decinea colombiana Grishin, 2022  [= Evans' "derisor"]; ref.13  iN  M T S

Decinea denta Evans, 1955  Peru, Bolivia; ref.13

Decinea denta denta Evans, 1955  Peru, Bolivia; ref.13  M T

Decinea denta pruda Evans, 1955  Paraguay  M T

Decinea antus (Mabille, 1895)  SE Brazil; ref.13  M T S

Decinea decinea (Hewitson, 1876)  Decinea Skipper  S Brazil  iN  M T S L

Decinea lydora (Plötz, 1882)  Colombia to Bolivia; ref.140  iN

Decinea lydora lydora (Plötz, 1882)  [= Evans' "neroides"] Colombia, Ecuador; ref.140  M S L

Decinea lydora lyco (Mabille, 1878)  Ecuador, Peru, Bolivia  M T S

Decinea notata Grishin, 2023  Peru; ref.153

Decinea dama (Herrich-Schäffer, 1869)  Dama Skipper  Colombia  iN  M T S

Decinea formosus (Hayward, 1940)  Ecuador; ref.153  M T

Decinea onasima (Hewitson, 1877)  S Brazil, Argentina; ref.153

Decinea onasima boliviensis (E. Bell, 1930)  Bolivia  M T

Decinea onasima onasima (Hewitson, 1877)  TL: Brazil ([RJ])  M T

Decinea zapota Evans, 1955  Zapota Skipper  Guatemala  iN  M T S

Decinea milesi (Weeks, 1901)  Miles' Skipper  Peru, Bolivia  iN  M S

Tirynthia  Godman, 1900  

Tirynthia conflua (Herrich-Schäffer, 1869)  Conflua Skipper  E Mexico to Argentina  iN  M T S L

Tirynthia conda Evans, 1955  Conda Skipper  Ecuador, Bolivia  iN  M S

Quasimellana  Burns, 1994  

Quasimellana mexicana (E. Bell, 1942)  Mexican Mellana  E & W Mexico, Guatemala, Belize  iN  M T S

Quasimellana mulleri (E. Bell, 1942)  Muller's Mellana  NE & W Mexico  M T S

Quasimellana eulogius (Plötz, 1882)  Common Mellana  S TX, E & W Mexico to Venezuela, Guianas, Trinidad, Brazil  iN  M T S L1 L2 L3 L4 Ha

Quasimellana agnesae (E. Bell, 1959)  Coastal Mellana  W Mexico (Nayarit to Guerrero); coastal  M T S L

Quasimellana balsa (E. Bell, 1942)  Sullied Mellana  W Mexico to El Salvador  iN  M T S L

Quasimellana siblinga Burns, 1994  Sibling Mellana  NW to S Mexico (SON, SIN, OAX)  iN  M T S

Quasimellana aurora (E. Bell, 1942)  Bright Mellana  W Mexico to El Salvador  iN  M T S

Quasimellana nayana (E. Bell, 1941)  Nayarit Mellana  W Mexico to Costa Rica, Venezuela  iN  M T S

Quasimellana noka (Evans, 1955)  Noka Mellana  distribution unknown

Quasimellana pazina (Evans, 1955)  Pazina Mellana  Bolivia

Quasimellana antipazina Burns, 1994  Costa Rican Mellana  Costa Rica  M T S

Quasimellana sista (Evans, 1955)  Sista Mellana  Venezuela

Quasimellana andersoni Burns, 1994  Anderson's Mellana  E Mexico to Costa Rica  M T

Quasimellana sethos (Mabille, 1889)  Small Mellana  [Nicaragua], Panama, Colombia, Ecuador, [Venezuela]  M T OI S

Quasimellana myron (Godman, 1900)  Greenish Mellana  E & W Mexico to Panama  M T S

Quasimellana servilius (Möschler, 1883)  Green Mellana  [= verba], E & S Mexico to Colombia, Venezuela, Ecuador, Brazil  iN  M T S

Quasimellana inconspicua (Hayward, 1951)  Inconspicuous Mellana  Ecuador, Peru, Bolivia  iN  M S

Quasimellana angra (Evans, 1955)  Angry Mellana  French Guiana, Venezuela, Ecuador, Peru, N & C Brazil  iN  M S

Quasimellana amicus (E. Bell, 1942)  Amicus Mellana  Ecuador  M T S

Quasimellana fieldi (E. Bell, 1942)  Field's Mellana  E Mexico to Costa Rica  iN  M T S

Quasimellana imperfida Burns, 1994  Imperfida Mellana  Colombia, Bolivia, S Brazil, Paraguay  M T

Quasimellana mielkei Burns, 1994  Mielke's Mellana  Brazil (MG)

Quasimellana nicomedes (Mabille, 1883)  Nicomedes Mellana  SE Brazil  M T

Quasimellana meridiani (Hayward, 1934)  Meridian Mellana  SE Brazil, Paraguay, Argentina  iN  M T

Quasimellana pandora (Hayward, 1940)  Pandora Mellana  Guianas, N & C Brazil, Ecuador, Peru, Bolivia  iN  M S

Atrytone  Scudder, 1872  [includes Problema as a subgenus]; ref.10

Atrytone arogos (Boisduval & Le Conte, [1837])  Arogos Skipper  ref.10  iN

Atrytone arogos arogos (Boisduval & Le Conte, [1837])  NJ to FL, W to LA  M T S L G

Atrytone arogos iowa (Scudder, 1868)  IL, IA, S MN, S ND, SD, SE MT, E WY, E CO, NE, KS, W MO, NW AR, OK to C TX  M S L1 L2 L3 L4 L5

Atrytone bulenta (Boisduval & Le Conte, [1837])  Rare Skipper  brackish coastal marshes, NJ to GA; ref.10  iN  M S L

Atrytone byssus (W. H. Edwards, 1880)  Byssus Skipper  ref.10  iN

Atrytone byssus byssus (W. H. Edwards, 1880)  NC to FL, W to MS  M T S L1 L2 L3 G

Atrytone byssus kumskaka Scudder, 1887  IN to IA, MO, E KS, E OK, W AR, W LA, E TX  M S L

Zariaspes  Godman, 1900  

Zariaspes mys (Hübner, [1808])  Mys Skipper  E Mexico to S Brazil & Paraguay  iN  M T S L

Zariaspes mythecus Godman, 1900  Godman's Skipper  W Mexico (NAY to OAX)  iN  M T S L

Anatrytone  Dyar, 1905  

Anatrytone logan (W. H. Edwards, 1863)  Delaware Skipper    iN

Anatrytone logan logan (W. H. Edwards, 1863)  ME to MN, S to FL & E TX  M T S L1 L2 L3 L4 G

Anatrytone logan lagus (W. H. Edwards, 1881)  SE ALTA & E MT to W & S TX, NL  M T S L1 L2

Anatrytone mazai (H. Freeman, 1969)  De la Maza's Skipper  S TX, E & W Mexico to Costa Rica  iN  M T S L

Anatrytone potosiensis (H. Freeman, 1969)  Potosi Skipper  E Mexico to Honduras  iN  M T S L

Anatrytone mella (Godman, 1900)  Mella Skipper  S Mexico to Colombia & Peru  iN  M T S L

Anatrytone perfida (Möschler, 1879)  Perfida Skipper  Trinidad, Guianas, Venezuela, Colombia, Brazil, Peru, Paraguay, Argentina  iN  M T S

Anatrytone sarah Burns, 1994  Sarah's Skipper  Colombia, Ecuador, Peru, Bolivia  iN  M T S

Anatrytone barbara (R. Williams & E. Bell, 1931)  Barbara's Skipper  Surinam to C Brazil  iN  M T S

Anatrytone flavens (Hayward, 1940)  Flavens Skipper  Argentina  M S

Hylephila  Billberg, 1820  

Hylephila phyleus (Drury, 1773)  Fiery Skipper    iN

Hylephila phyleus phyleus (Drury, 1773)  [= muertovalle] CA to FL (strays far N), S to Argentina; Caribbean; Hawaii  M T S1 S2 IL L1 L2 L3 L4 L5 L6 I G

Hylephila phyleus taxus Evans, 1955  Ecuador (Rio Pastaza, Ambato, 5000-7000')

Hylephila phyleus monticola Hayward, 1939  Ecuador (Bolivar, 9000')  M T

Hylephila phyleus anca Evans, 1955  S Peru (Carabaya, Marcapata)

Hylephila phyleus andina (Staudinger, 1894)  Bolivia (Cocopata, Cochabamba), N Argentina  M T

Hylephila basistrigata (Eaton, 1932)  W Peru (Lima, Trujillo, Tarapaca, etc.) to Chile

Hylephila signata (Blanchard, 1852)  Signata Skipper  Chile  iN  M T S

Hylephila ancora (Plötz, 1883)  Ancora Skipper  Bolivia, Uruguay, Argentina  iN

Hylephila isonira Dyar, 1913  Isonira Skipper    iN

Hylephila isonira isonira Dyar, 1913  Ecuador, Peru, Bolivia  M S

Hylephila isonira mima Evans, 1955  Peru (Arequipa, Arica, Tirapata, Titicaca), Bolivia (La Paz), Chile

Hylephila fasciolata (Blanchard, 1852)  Fasciolata Skipper  Argentina, Chile  iN  M S

Hylephila zapala Evans, 1955  Zapala Skipper  Argentina (Zapala)

Hylephila ignorans (Plötz, 1883)  Ignorans Skipper  Cordillera de Merida, Venezuela  M S

Hylephila adriennae MacNeill & Herrera, 1999  Adrienne's Skipper  Sierra Nevada de Santa Marta, Colombia  iN  M T

Hylephila herrerai MacNeill, 2002  Herrera's Skipper  Chile (Parinacota, Cotacotani, 4500m)

Hylephila pseudoherrerai MacNeill, 2002  False Herrera's Skipper  Peru (Ayacucho, Reserva Nacional de Pampas Galeras, 4000m)  M S

Hylephila pallisteri MacNeill, 2002  Pallister's Skipper  Peru (Cuzco, Ollantaitambo, 9200'), possibly also Abra Acjanacu, 3600m ( ...  iN  M S

Hylephila blancasi MacNeill, 2002  Blancas' Skipper  Peru ("Santia," Lima [Canta], Junin, Huanuco)  M S

Hylephila tentativa MacNeill, 2002  Tentative Skipper  Peru (Ayacucho, vic. Apacheta de Tambo, 4200-4250m)  M S

Hylephila shapiroi MacNeill, 2002  Shapiro's Skipper  Peru (Junin, vic. Abra Anticona, 4843m)

Hylephila galera Evans, 1955  Galera Skipper  Peru (Junin, Galera Pass, 4800m, Tarma, 3000m, Yauli, 4300m)  M S

Hylephila boulleti (Mabille, 1906)  Boullet's Skipper  Argentina (Catamarca, Tucuman), Chile, Bolivia, Peru  iN  M T S

Hylephila rossi MacNeill, 2002  Ross' Skipper  Peru (Puno, 10 mi N Ayaviri; Cuzco-Puno, La Raya, 4318m)  M S

Hylephila peruana Draudt, 1923  Peruana Skipper  Peru (Cuzco, 3500-4500m)  iN  M T S G

Hylephila venustus (Hayward, 1940)  Venustus Skipper  Argentina (Santa Cruz Prov) N to Santiago Prov., Chile  iN  M S

Hylephila kenhaywardi MacNeill, 1999  Ken Hayward's Skipper  Chilean Andes, 3000-4600m (Elqui, Copiapo, El Loa, Parinacota)

Hylephila lamasi MacNeill & Herrera, 1999  Lamas' Skipper  S-central Peru, coastal  iN  M S

Pompeius  Evans, 1955  [includes Appia as its synonym]; ref.12

Pompeius appia (Evans, 1955)  Appia Skipper  Uruguay, Argentina; ref.12  iN  M T

Pompeius amblyspila (Mabille, 1898)  Amblyspila Skipper  [Nicaragua?], Colombia to Argentina  iN  M T S

Pompeius fuscus Grishin, 2023  Brazil; ref.153

Pompeius pompeius (Latreille, [1824])  Pompeius Skipper  E & W Mexico to Argentina  iN  M S L1 L2

Pompeius darina Evans, 1955  S Brazil (MT)

Hedone  Scudder, 1872  may be a subgenus of Polites; ref.10,128

Hedone vibex (Geyer, 1832)  Whirlabout  ref.10,16  iN

Hedone vibex vibex (Geyer, 1832)  NC to FL, W to LA  M S HS L1 L2 L3 G

Hedone vibex brettoides (W. H. Edwards, 1883)  C & NE TX  M T S L

Hedone praeceps Scudder, 1872  Whirlabout  S TX, E & W Mexico to Trinidad, Venezuela, Ecuador; ref.16  iN  M T S L1 L2 L3 L4 I1 I2

Hedone vibicoides (de Jong, 1983)  Surinam; ref.16  M T

Hedone calla (Evans, 1955)  Peru; ref.16

Hedone catilina (Plötz, 1886)  [= stigma] Guianas, Venezuela, Ecuador, Peru, Bolivia, Brazil, A ...; ref.16  iN  M T S

Hedone yunga Grishin, 2023  Yungas Region; ref.20  M T

Hedone mira Grishin & Lamas, 2022  S Peru; ref.16  M T

Hedone miracla Grishin, 2024  coast of central Peru; ref.154  M T

Hedone dictynna (Godman & Salvin, 1896)  Dictynna Skipper  Lesser Antilles; ref.16  iN  M T S L

Hedone bittiae (Lindsey, 1925)  Peru; ref.16  M S

Limochores  Scudder, 1872  may be a subgenus of Polites; ref.10,128

Limochores sonora (Scudder, 1872)  Sonoran Skipper    iN

Limochores sonora N CA coastal segregate (, )  N CA coast  M S L

Limochores sonora siris (W. H. Edwards, 1881)  N OR coast, W WA  M S L

Limochores sonora Willamette Valley segregate (, )  Willamette Valley, W OR  M S

Limochores sonora sonora (Scudder, 1872)  Cascades of BC-WA-OR-CA to Sierra Nevada of CA, S to Baja California  M S L1 L2 L3 L4 Ha G

Limochores sonora flavaventris (Austin, 1998)  NW NV to S-central OR  M T

Limochores sonora longinqua (Austin, 1998)  White Mts. of CA-NV  M T S L

Limochores sonora NE Oregon segregate (, )  NE OR, SE WA  M S

Limochores sonora utahensis (Skinner, 1911)  C ID, W MT, W & S WY, UT, CO  M T S L I H

Limochores mystic (W. H. Edwards, 1863)  Long Dash    iN

Limochores mystic mystic (W. H. Edwards, 1863)  SE Canada to MAN, S to IA, WV, VA  M S L1 L2 L3 L4 L5 Ha

Limochores mystic dacotah (W. H. Edwards, 1871)  S ALTA to NE CO, SW CO, C AZ  M T S L Ha

Limochores mystic nino Grishin, 2023  AZ: Coconino Co.; ref.151  M T

Limochores mystic Pacific northwestern segregate (, )  E WA, BC, W ALTA, N ID, NW MT  M S

Limochores origenes (Fabricius, 1793)  Crossline Skipper  ref.10  iN

Limochores origenes origenes (Fabricius, 1793)  SE Canada, New England to E SD, S to NE TX, C FL  M S Ma L1 L2 L3 L4 L5 L6 I F G

Limochores origenes rhena (W. H. Edwards, 1878)  W ND & E MT to E CO & NE NM  M T S

Limochores pupillus (Plötz, 1882)  Pupiled Skipper  [= puxillius, Evans' "pupillus" is catahorma] NW to S-central Mexico; ref.13  iN  M T S L Ha

Limochores catahorma (Dyar, 1916)  [= Evans' "pupillus"] NW to S-central Mexico; ref.13  iN  M T S L

Polites  Scudder, 1872  [includes Wallengrenia as a subgenus]; ref.10,12,128

Polites themistocles (Latreille, [1824])  Tawny-edged Skipper    iN

Polites themistocles turneri H. Freeman, 1944  S BC, NE WA  M T

Polites themistocles UT segregate ,  UT, N AZ  M S I

Polites themistocles themistocles (Latreille, [1824])  SE Canada, New England to MT, S to NM, S FL, disjunct in N CA  M S L1 L2 L3 L4 L5 G

Polites coras (Cramer, 1775)  Peck's Skipper  [= peckius]; ref.16  iN

Polites coras coras (Cramer, 1775)  SE Canada, New England to NE OR, S BC; univoltine; ref.16  M S L1 L2 G

Polites coras surllano Scott, 2006  southern Great Plains, CO to MO, at least; range poorly defined, bivoltine  M S Hy L

Polites coras AZ White Mts. segregate ,  AZ White Mts.  M S L

Polites mardon (W. H. Edwards, 1881)  Mardon Skipper    iN

Polites mardon mardon (W. H. Edwards, 1881)  WA Puget Sound area & nr. Mt. Adams  M S L G

Polites mardon N CA-S OR coastal segregate ,  NW CA (Del Norte Co.), SW OR (Curry Co.) near coast  M S I

Polites mardon klamathensis Mattoon, J. Emmel & T. Emmel, 1998  SW OR (Jackson & Klamath Cos.) S end Cascades  M T S L Ha

Polites sabuleti (Boisduval, 1852)  Sandhill Skipper    iN

Polites sabuleti margaretae L. Miller & MacNeill, 1969  Baja California Sur  M T S

Polites sabuleti sabuleti (Boisduval, 1852)  TL: USA (Ca)  M T S L1 L2 Ha G

Polites sabuleti channelensis J. Emmel & T. Emmel, 1998  Santa Rosa I., Santa Barbara Co., CA  M T

Polites sabuleti aestivalis J. Emmel, T. Emmel & Mattoon, 1998  NW CA, SW OR  M T S L Ha

Polites sabuleti E-central Cascadian segregate ,  SE OR Cascades  M S

Polites sabuleti tecumseh (F. Grinnell, 1903)  Sierra Nevada, high elevations  M T S L Ha

Polites sabuleti albamontana Austin, 1987  White Mts. of CA-NV  M T S L1 L2

Polites sabuleti genoa (Plötz, 1883)  W NV (vic. Carson City)  M S L Ha

Polites sabuleti alkaliensis Austin, 1987  SE WA, E OR, NE NV to N UT, W CO, NW NM; also CO Front Range  M T S Hy L I Ha

Polites sabuleti sinemaculata Austin, 1987  Humboldt Co., NV  M T S

Polites sabuleti basinensis Austin, 1988  Reese R. Valley, C NV  M T S L Ha

Polites sabuleti nigrescens Austin, 1987  Steptoe Valley, White Pine Co., NV  M T Ha

Polites sabuleti chusca (W. H. Edwards, 1873)  far SE CA, S NV, NW AZ, SW UT  M T S

Polites sabuleti ministigma Scott, 1981  S-central CO (San Luis Valley, upper Arkansas R. Valley)  M T S

Polites norae MacNeill, 1993  Guaymas Skipper  coastal Sonora-Sinaloa, NW Mexico  iN  M T S L I

Polites draco (W. H. Edwards, 1871)  Draco Skipper  Yukon, N BC, Rocky Mts. from W ALTA to N NM  iN  M S L1 L2 L3 Ha G

Polites carus (W. H. Edwards, 1883)  Carus Skipper  far W OK, far SE CO, NM, W TX, AZ, NW Mexico to central Mexico  iN  M S HS L I Ha G

Polites subreticulata (Plötz, 1883)  Subreticulate Skipper  E & W Mexico to Costa Rica  iN  M T OI S L

Polites rhesus (W. H. Edwards, 1878)  Rhesus Skipper    iN

Polites rhesus rhesus (W. H. Edwards, 1878)  S ALTA, SASK, E MT, W ND, W SD, W NE, E WY, E CO, far W OK, NM, C AZ  M T S L1 L2 I Ha G

Polites rhesus otomi A. Warren & Gott, 2021  NW to S-Central Mexico; ref.26  M S Ha

Polites baracoa (Lucas, 1857)  Baracoa Skipper    iN

Polites baracoa baracoa (Lucas, 1857)  S GA, FL, Cuba  M T S1 S2 L G

Polites baracoa loma Evans, 1955  Hispaniola  M S

Polites egeremet (Scudder, 1863)  Northern Broken-dash  SE Canada, New England to C FL, W to E TX, N to SE ND, C MN; ref.12  iN  M S L1 L2 L3 G

Polites otho (J. E. Smith, 1797)  Southern Broken-dash  ref.12  iN

Polites otho otho (J. E. Smith, 1797)  MD to FL, W to TX, E & W Mexico to Costa Rica  M T S L1 L2 L3 L4 L5 L6 L7 L8 G

Polites otho clavus (Erichson, [1849])  [= curassavica], S TX, E & W Mexico to Venezuela, Guianas, Trinidad, N Brazil, Peru  M T S L

Polites sapuca (Evans, 1955)  Paraguay, Argentina; ref.12  M T S

Polites premnas (Wallengren, 1860)  Trinidad, Guianas, Venezuela, Brazil, Peru, Bolivia, Paraguay, A ...; ref.12  iN  M T S HS L

Polites misera (Lucas, 1857)  Cuban Broken-dash  Cuba, Isla de Juventud, N Bahamas; ref.12  iN  M T S

Polites drury (Latreille, [1824])  Drury's Broken-dash  S Bahamas, Turks & Caicos, Hispaniola, P. Rico, Virgin Is.; ref.12  iN  M T S

Polites vesuria (Plötz, 1882)  Jamaican Broken-dash  Jamaica  iN  M S L

Polites ophites (Mabille, 1878)  Fiery Broken-dash  St. Martin, St. Eustatius, Saba; St. Kitts to St. Vincent, Trinidad; ref.12  iN  M T S

Nyctelius  Hayward, 1948  

Nyctelius nyctelius (Latreille, [1824])  Violet-banded Skipper, Nyctelius Skipper    iN

Nyctelius nyctelius nyctelius (Latreille, [1824])  Greater Antilles, Virgin Islands, S Texas to Argentina  M T S L1 L2 G

Nyctelius nyctelius agari (Dillon, 1947)  Lesser Antilles  M S

Nyctelius paranensis (Schaus, 1902)  Parana Skipper  SE Brazil  iN  M T

Conga  Evans, 1955  

Conga chydaea (A. Butler, 1877)  Hidden-ray Skipper  S TX to Argentina; ref.13,16  iN  M T S L

Conga urqua (Schaus, 1902)  Urqua Skipper  SE Brazil, Argentina  iN  M T S

Conga immaculata (E. Bell, 1930)  Sullied Skipper  SE Brazil  iN  M T S

Conga iheringii (Mabille, 1891)  Dark-patched Skipper  SE Brazil  iN  M T

Conga zela (Plötz, 1883)  Zela Skipper  SE Brazil, Uruguay, Argentina  iN  M T

Vernia  Grishin, 2019  ref.5

Vernia clara Grishin, 2023  E & W Mexico to Panama; ref.153  M S L

Vernia dares (Plötz, 1883)  Dares Skipper  Colombia to S Brazil; ref.5  iN  M S

Vernia verna (W. H. Edwards, 1862)  Little Glassywing  ref.5  iN

Vernia verna sequoyah (H. Freeman, 1942)  AR, SE OK, E TX, [NE Mexico]; ref.146  M T1 T2 S L

Vernia verna verna (W. H. Edwards, 1862)  S QUE, ONT, New England to N FL, W to LA, N to MN, SE SD, E NE, NE KS  M T S L1 L2 L3 F G

Atalopedes  Scudder, 1872  [includes Linka]; ref.7

Atalopedes carteri Evans, 1955  Carter's Skipper  New Providence Island, Bahamas  M S

Atalopedes mesogramma (Latreille, [1824])  Mesogramma Skipper    iN

Atalopedes mesogramma mesogramma (Latreille, [1824])  Cuba, New Providence, Andros  M T S

Atalopedes mesogramma apa W. Comstock, 1944  Hispaniola, Puerto Rico  M T S

Atalopedes lina (Plötz, 1883)  Lina Skipper  Colombia, Bolivia; ref.7  M S

Atalopedes campestris (Boisduval, 1852)  Casachem  Baja California to WA, Columbia Basin to W Idaho; ref.16  iN  M T S L Ha G

Atalopedes huron (W. H. Edwards, 1863)  Sachem  E USA (strays far N), E & W Mexico to Ecuador, Venezuela, Brazil; ref.16  iN  M T S L1 L2 L3 L4 L5 L6 L7 L8 L9 L10

Atalopedes flaveola (Mabille, 1891)  Flaveola Skipper  [= clarkei], Venezuela's Caribbean islands  M T

Atalopedes bahiensis (Schaus, 1902)  Bahia Skipper  E Brazil (Ba)  M T

Hesperia  Fabricius, 1793  

Hesperia nabokovi (E. Bell & W. Comstock, 1948)  Nabokov's Skipper  Hispaniola  M T S

Hesperia comma (Linnaeus, 1758)    iN

Hesperia comma lena Korshunov & Gorbunov, 1995  ref.121

Hesperia colorado (Scudder, 1874)  Western Branded Skipper  ref.121  iN

Hesperia colorado colorado (Scudder, 1874)  S-central CO, N NM  M T S L Ha

Hesperia colorado sublima A. Warren & Calhoun, 2015  Rocky Mts. of Colorado, at and above treeline; ref.106,121  M S1 S2 Ha G

Hesperia colorado ochracea Lindsey, 1941  CO Front Range (Larimer to El Paso cos.); ref.121  M T S1 S2 L1 L2 L3 Ha

Hesperia colorado susanae L. Miller, 1962  S-central & SE AZ, SW NM  M T1 T2 S L1 L2

Hesperia colorado mojavensis Austin & McGuire, 1998  Spring Mts., Clark Co., NV  M T1 T2 S

Hesperia colorado idaho (W. H. Edwards, 1883)  W CO to E CA, N to SE BC, MT, WY  M T S L1 L2 L3 L4

Hesperia colorado E Columbia Basin segregate ,  E Columbia Basin, NE OR, SE WA  M S Ab L1 L2 L3 Ha

Hesperia colorado leussleri Lindsey, 1940  SW CA  M T S1 S2 L1 L2 Ha

Hesperia colorado tildeni H. Freeman, 1956  W-central CA (Coast Ranges)  M T S

Hesperia colorado harpalus (W. H. Edwards, 1881)  CA Sierra Nevada  M T S1 S2

Hesperia colorado oregonia (W. H. Edwards, 1883)  N CA-SW OR Cascades, E Siskiyous  M T S L Ha G

Hesperia colorado W Cascades segregate ,  W OR Cascades (from Clackamas to N Jackson cos.)  M S L1 L2 L3 L4 Ha

Hesperia colorado mattoonorum McGuire, 1998  NW CA-SW OR W Siskiyous & outer Coast Range  M S L I Ha

Hesperia colorado N CA coastal segregate ,  Marin Co., CA  M S

Hesperia colorado dodgei (E. Bell, 1927)  Santa Cruz Co., CA  M T S L G

Hesperia colorado hulbirti Lindsey, 1939  Olympic Mts., WA, high elevations  M T1 T2 S

Hesperia colorado manitoba (Scudder, 1874)  N Manitoba to Alaska, S in Canadian Rocky Mts. to N-central WA & N MT  M S

Hesperia colorado borealis Lindsey, 1942  Newfoundland, Labrador, N Quebec  M S L

Hesperia colorado laurentina (Lyman, 1892)  SE Canada, N New England, N MI, WI, MN to E MAN  M S L

Hesperia assiniboia (Lyman, 1892)  Plains Branded Skipper  Peace River BC-ALTA to C MAN, S to MT, ND, far W MN  iN  M S L1 L2

Hesperia juba (Scudder, 1872)  Juba Skipper  S BC to W SD, S to NW NM, N AZ, CA & Baja California  iN  M T S L1 L2 L3 I Ha G

Hesperia lindseyi (W. Holland, 1930)  Lindsey's Skipper    iN

Hesperia lindseyi lindseyi (W. Holland, 1930)  W-central to S CA, far SW AZ  M T S L Ha

Hesperia lindseyi macneilli J. Emmel, T. Emmel & Mattoon, 1998  outer N Coast Ranges, NW CA  M T S

Hesperia lindseyi eldorado J. Emmel, T. Emmel & Mattoon, 1998  W slope Sierra Nevada, CA (Butte to El Dorado cos.)  M T S

Hesperia lindseyi septentrionalis J. Emmel, T. Emmel & Mattoon, 1998  Siskiyou Mts & Modoc Plateau, N CA, S OR, NW NV  M T S L1 L2 Ha

Hesperia lindseyi mccorklei P. Severns & D. Severns, 2005  upper Rogue River drainage, Douglas Co., OR  M T

Hesperia miriamae MacNeill, 1959  Sierra Skipper    iN

Hesperia miriamae miriamae MacNeill, 1959  S Sierra Nevada, CA (Alpine to Tulare cos.), high elevations  M T S

Hesperia miriamae longaevicola McGuire, 1998  CA-NV White Mts., high elevations  M S L Ha

Hesperia nevada (Scudder, 1874)  Nevada Skipper    iN

Hesperia nevada nevada (Scudder, 1874)  ALTA & SASK, Rocky Mts. from MT-ID to N-central NM  M T1 T2 T3 S L1 L2 Ha G

Hesperia nevada N Great Basin & Columbia Basin segregate ,  N Great Basin, OR-ID-NV & Columbia Basin, WA-BC  M S L Ha

Hesperia nevada sierra Austin, J. Emmel, T. Emmel & Mattoon, 1998  S OR (Warner Mts.) to S end Sierra Nevada, CA  M T S L1 L2 Ha

Hesperia woodgatei (R. Williams, 1914)  Apache Skipper    iN

Hesperia woodgatei Hualapai Mts. segregate ,  Hualapai Mts., Mohave Co., AZ  M S

Hesperia woodgatei woodgatei (R. Williams, 1914)  S CO, NM, W TX, C & SE AZ, CHIH; ref.19  M T S L1 L2 Ha G

Hesperia woodgatei NW Mexican segregate ,  NW Mexico (DGO)  M S

Hesperia woodgatei NE Mexican segregate ,  NE Mexico (COAH, NL)  M S L

Hesperia balcones (Grishin, 2023)  Hill Country Skipper  C TX (Travis, Kerr & Bear cos.); ref.19  M S1 S2 L H

Hesperia uncas W. H. Edwards, 1863  Uncas Skipper    iN

Hesperia uncas gilberti MacNeill, 1964  S-central Mexico (DF, MEX, OAX)  M T

Hesperia uncas nadineae A. Warren, Gott & McGuire, 2021  NE Mexico (Coah, NL); ref.29  M S

Hesperia uncas Minnesota segregate ,  Minnesota (Sherburne Co.)  M S

Hesperia uncas tomichi M. Fisher, 2008  Colorado Rockies, high elevations; univoltine  M T S L Ha

Hesperia uncas uncas W. H. Edwards, 1863  SE MAN to S ALTA, S to NE ID, WY,W NE, W KS, W OK, E CO, NM, N & W TX  M T S L1 L2 L3 Ha G

Hesperia uncas lasus (W. H. Edwards, 1884)  W CO, UT, AZ, SW NM  M T S L Ha

Hesperia uncas grandiosa Austin & McGuire, 1998  White River and Big Smokey valleys, Nye Co., NV  M T1 T2 S L Ha

Hesperia uncas fulvapalla Austin & McGuire, 1998  Railroad Valley, Nye Co., NV & Calleo area, Juab Co., UT  M T1 T2 S L Ha

Hesperia uncas reeseorum Austin & McGuire, 1998  Reese River and Mason valleys, Lander Co., NV  M T1 T2 S L Ha

Hesperia uncas terraclivosa Austin & McGuire, 1998  C NV (Toquima, Mts, Schell Creek Range, Humboldt Range) to Alvord Desert, Harney ...  M T1 T2 S1 S2 Ha

Hesperia uncas central Oregon segregate ,  Crook-Deschutes cos., central OR  M S

Hesperia uncas giulianii McGuire, 1998  vic. Adobe Hills, Mono Co., CA  M S L

Hesperia uncas macswaini MacNeill, 1964  CA-NV White Mts., high elevations  M T S L1 L2 I H

Hesperia leonardus T. Harris, 1862  Leonard's Skipper    iN

Hesperia leonardus leonardus T. Harris, 1862  SE Canada, New England to SC, NE GA, W to WI, MO  M T1 T2 S L1 L2 G

Hesperia leonardus midwest blends segregate ,  E SASK, C MN to E OK  M S1 S2 L1 L2 L3 L4

Hesperia leonardus pawnee Dodge, 1874  E CO to E MT, E to SW MN, NW IA  M T S L1 L2 L3 Ha

Hesperia leonardus montana (Skinner, 1911)  S Platte River Canyon, C CO  M T S1 S2 L1 L2 L3 L4 Ha

Hesperia columbia (Scudder, 1872)  Columbian Skipper    iN

Hesperia columbia columbia (Scudder, 1872)  C & S CA  M T S Ab L1 L2 G

Hesperia columbia N Cascades-Siskiyous segregate ,  Siskiyou Mts. & SW Cascades, NW CA, SW OR  M S L Ha

Hesperia pahaska Leussler, 1938  Pahaska Skipper    iN

Hesperia pahaska pahaska Leussler, 1938  SASK, MT & ND, S to W AZ & C NM; ref.151  M S1 S2 L Ha

Hesperia pahaska williamsi Lindsey, 1940  S AZ to W TX, NW Mexico  M T S L1 L2 L3 I Ha

Hesperia pahaska hannawackeri Grishin, 2023  SE UT & SW CO; ref.151

Hesperia pahaska martini MacNeill, 1964  SE CA  M T1 T2 S L

Hesperia pahaska Baja California Norte segregate ,  N Baja California, Mexico  M S

Hesperia viridis (W. H. Edwards, 1883)  Green Skipper  SE WY to E AZ & C TX, NE Mexico  iN  M S1 S2 L1 L2 L3 Ha G

Hesperia metea Scudder, 1863  Cobweb Skipper    iN

Hesperia metea metea Scudder, 1863  New England to OH, WI  M S L G

Hesperia metea intermedia Gatrelle, 2003  S Appalachians (SW VA to N GA)  M T S L

Hesperia metea licinus (W. H. Edwards, 1871)  NE & IL S to TX, MS  M T S L

Hesperia attalus (W. H. Edwards, 1871)  Dotted Skipper    iN

Hesperia attalus attalus (W. H. Edwards, 1871)  S KS to C TX  M T S L1 L2 L3 G

Hesperia attalus nigrescens Gatrelle, 1999  S NC, N SC, NJ?  M T

Hesperia attalus slossonae (Skinner, 1890)  FL to LA  M T S1 S2 L1 L2

Hesperia ottoe W. H. Edwards, 1866  Ottoe Skipper  S MAN, E MT & W ND, S to E CO, W OK, N TX, E to IL, MI  iN  M T S1 S2 HS L G

Hesperia meskei (W. H. Edwards, 1877)  Meske's Skipper    iN

Hesperia meskei meskei (W. H. Edwards, 1877)  NE TX, AR, E OK  M T S L G

Hesperia meskei straton (W. H. Edwards, 1881)  S NC to S FL  M T S L1 L2 L3 L4

Hesperia meskei pinocayo Gatrelle & Minno, 2003  Big Pine Key, Monroe Co., FL (apparently extinct)  M T S

Hesperia sassacus T. Harris, 1862  Indian Skipper    iN

Hesperia sassacus nantahala Gatrelle & Grkovich, 2003  SW VA, W NC, TN, N GA  M T S L

Hesperia sassacus sassacus T. Harris, 1862  SE Canada, New England to MD, W to MI  M S L I Ha G

Hesperia sassacus manitoboides (Fletcher, 1889)  S MAN to WI  M T S L

Hesperia dacotae (Skinner, 1911)  Dakota Skipper  S MAN, ND, E SD, W MN  iN  M T S L G

Pseudocopaeodes  Skinner & R. Williams, 1923  

Pseudocopaeodes eunus (W. H. Edwards, 1881)  Alkali Skipper    iN

Pseudocopaeodes eunus eunus (W. H. Edwards, 1881)  S CA (Kern to San Diego cos.)  M T S L1 L2 I Ha G

Pseudocopaeodes eunus alinea (Scott, 1981)  S NV, far SE CA  M T S

Pseudocopaeodes eunus ash Grishin, 2023  NV: Ash Meadows; ref.151

Pseudocopaeodes eunus flavus Austin & J. Emmel, 1998  Inyo Co., CA, N through Lyon, Mineral, Storey, Washoe, Pershing, Lander and Nye c ...  M T S L1 L2 Ha

Pseudocopaeodes eunus obscurus Austin & J. Emmel, 1998  Carson City, NV, also Honey L., Lassen Co., CA & vic. Mono L. & L. Crowley, Mono ...  M T1 T2 T3

Parachoranthus  L. Miller, 1965  

Parachoranthus magdalia (Herrich-Schäffer, 1863)  Magdalia Skipper  Cuba, [N Andros]  iN  M T S

Neochlodes  Austin & DeVries, 2001  

Neochlodes maculosus Austin & DeVries, 2001  Maculosus Skipper  Costa Rica (Cerro Echandi, 2400m, Puntarenas)  M T

Ochlodes  Scudder, 1872  

Ochlodes batesi (E. Bell, 1935)  Bates' Skipper  Hispaniola  iN  M S

Ochlodes yuma (W. H. Edwards, 1873)  Yuma Skipper    iN

Ochlodes yuma sacramentorum Austin, 1998  CA Central Valley  M T1 T2

Ochlodes yuma yuma (W. H. Edwards, 1873)  far NW NM, N AZ, S UT, S NV, SE CA  M T S L I

Ochlodes yuma lutea Austin, 1998  C & N NV, SE OR  M T1 T2 L Ha

Ochlodes yuma Columbia Basin segregate ,  C Washington (Dry Falls area) to Columbia R. of S WA, N OR  M S

Ochlodes yuma lower Snake River drainage segregate ,  vic. Hells Canyon (Snake R.), NE OR, W ID  M S

Ochlodes yuma scudderi (Skinner, 1899)  WY (Wind R. Cyn.), W CO, NE UT  M T S L I H

Ochlodes yuma anasazi S. Cary & Stanford, 1995  N New Mexico (Rio Grande Gorge)  M T S L1 L2 H

Ochlodes santacruza Scott, 1981  Woodland Skipper  Santa Cruz & Santa Catalina Islds, CA; ref.151

Ochlodes santacruza santacruza Scott, 1981  Santa Cruz Island, CA; ref.151  M T S

Ochlodes santacruza catalina J. Emmel & T. Emmel, 1998  Santa Catalina Island, CA; ref.151  M T L

Ochlodes sylvanoides (Boisduval, 1852)  Woodland Skipper    iN

Ochlodes sylvanoides sylvanoides (Boisduval, 1852)  S BC, W WA, W OR, W CA, N BC (Mexico)  M T S1 S2 L1 L2 L3 Ha G

Ochlodes sylvanoides orecoasta Scott, 1981  immediate coastal habitats from W WA to N CA, variable phenotype  M T S

Ochlodes sylvanoides bonnevilla Scott, 1981  N Great Basin, esp. Elko Co., NV, variable but palest phenotype  M T S L Ha

Ochlodes sylvanoides omnigena Austin, 1998  E CA Sierra Nevada & E OR-WA Cascades to MT, WY, W CO  M T1 T2 L1 L2 L3 I Ha

Ochlodes napa (W. H. Edwards, 1865)  Woodland Skipper  CO Front Range (Larimer to El Paso & Fremont cos.), E WY, W SD

Ochlodes napa napa (W. H. Edwards, 1865)  CO Front Range (Larimer to El Paso & Fremont cos.), E WY, W SD  M S L Ha

Ochlodes napa kaibab Grishin, 2023  N AZ; ref.151

Ochlodes agricola (Boisduval, 1852)  Rural Skipper    iN

Ochlodes agricola Northern segregate ,  NW CA, SW OR  M S L Ha

Ochlodes agricola agricola (Boisduval, 1852)  W CA, N BC (MEX)  M T S L1 L2 Ha G

Ochlodes agricola nemorum (Boisduval, 1852)  W Sierra Nevada, CA  M T S

Ochlodes agricola verus (W. H. Edwards, 1881)  far S Sierra Nevada, to SE (CA); ref.151  M T S L I

Stinga  Evans, 1955  

Stinga morrisoni (W. H. Edwards, 1878)  Morrison’s Skipper  CO Front Range, NM, E AZ, W TX, N to S-central Mexico  iN  M T S L1 L2 L3 L4 L5 I Ha G

Stinga kendamulaza A. Warren & Austin, 2009  Kendamulaza Skipper  S Mexico (TLAX, PUE, OAX); ref.107  M T L Ha

Poanes  Scudder, 1872  

Poanes yehl (Skinner, 1893)  Yehl Skipper  VA to FL, AL to E TX, far E OK, N to S IN, IL  iN  M T S HS L1 L2 L3 G

Poanes aaroni (Skinner, 1890)  Aaron's Skipper    iN

Poanes aaroni aaroni (Skinner, 1890)  NJ to NC  M S G

Poanes aaroni minimus Gatrelle, 1999  S SC  M T L

Poanes aaroni howardi (Skinner, 1896)  FL, S GA, S AL, S MS  M T S1 S2 L

Poanes aaroni bordeloni Gatrelle, 2000  SE TX, SW LA  M T S L

Poanes viator (W. H. Edwards, 1865)  Broad-winged Skipper    iN

Poanes viator viator (W. H. Edwards, 1865)  NE, SE ND, E SD to NY, ONT  M S L G

Poanes viator zizaniae Shapiro, 1971  MA to FL, E to C TX  M S L1 L2 L3 Ha

Poanes massasoit (Scudder, 1863)  Mulberry Wing    iN

Poanes massasoit massasoit (Scudder, 1863)  NH & MA to MD, NY to MN, far SE ND, E SD  M T S1 S2 L1 L2 L3 G

Poanes massasoit chermocki Andersen & Simmons, 1976  vic. New Bridge, Dorchester Co., MD  M T S

Lon  Grishin, 2019  ref.5

Lon hobomok (T. Harris, 1862)  Hobomok Skipper  ref.5  iN

Lon hobomok hobomok (T. Harris, 1862)  SE Canada to SASK & E-central ALTA, S to NE CO, E OK, AR to TN, N AL  M T S L1 L2 L3 Ha G

Lon hobomok monofacies (Gatrelle, 2002)  NC to GA  M T S L1 L2

Lon hobomok wetona (Scott, 1981)  S-central CO, E NM ranges  M T S

Lon zabulon (Boisduval & Le Conte, [1837])  Zabulon Skipper  NE USA S to montane Mexico & Panama; ref.5  iN  M T S L1 L2 L3 L4 L5 L6 F Ha G

Lon co Grishin, 2023  Zabulon Skipper  Mexico to Costa Rica; ref.151  M T S L Ha

Lon ma Grishin, 2023  Zabulon Skipper  Panama; ref.151  M L

Lon taxiles (W. H. Edwards, 1881)  Taxiles Skipper  W SD, W NE, E WY, CO, S ID, UT, AZ, NM, W TX, NW Mexico to DF, OAX; ref.5  iN

Lon taxiles taxiles (W. H. Edwards, 1881)  W SD, W NE, E WY, CO, S ID, UT, AZ, NM, W TX, NW Mexico to DF, OAX  M S L1 L2 I Ha G

Lon taxiles albimaculatus Spomer, 2021  TL: USA (Ne)

Lon azin (Godman, 1900)  Azin Skipper  Colombia; ref.5  iN  M T

Lon macneilli (Burns, 1992)  MacNeill's Skipper  Colombia (Sierra Nevada de Santa Marta); ref.5  M T

Lon inimica (A. Butler & H. Druce, 1872)  Yellow-stained Skipper  E & W Mexico to Colombia, Venezuela; ref.5  iN  M T S L Ha

Lon melane (W. H. Edwards, 1869)  Umber Skipper  ref.5  iN

Lon melane melane (W. H. Edwards, 1869)  CA (widespread W of Sierra Nevada), N Baja California  M T S L G

Lon melane sur Grishin, 2023  Baja California Sur; ref.151  M T S

Lon vitellina (Herrich-Schäffer, 1869)  'Mexican' Umber Skipper  SE AZ, SW NM, W TX to Oaxaca; ref.151  M S L I Ha

Lon chia Grishin, 2023  Chiapas' Umber Skipper  VER (Los Tuxtlas), Chiapas to El Salvador; ref.151  M T S1 S2 L

Lon poa (Evans, 1955)  'Central American' Umber Skipper  Costa Rica & Panama; ref.151  M S

Lon ulphila (Plötz, 1883)  Ulphila Skipper  Mexico, a domen dubium placed here for now; ref.5,139  M TI

Lon monticola (Godman, 1900)  Oyamel Skipper  [likely conspecific with ulphila] W to S-central Mexico; ref.5,139  iN  M T S1 S2 HS L1 L2

Lon niveolimbus (Mabille, 1889)  Snow-fringed Skipper  Chiapas, Guatemala; ref.5  iN  M T OI S1 S2

Evansiella  Hayward, 1948  

Evansiella cordela (Plötz, 1882)  Brazil & Paraguay  iN  M T S

Paratrytone  Godman, 1900  

Paratrytone snowi (W. H. Edwards, 1877)  Snow’s Skipper  SE WY (Laramie Co.), CO Front Range, S CO, NM, E AZ, CHIH, [SON?]  iN  M S L1 L2 Ha G

Paratrytone decepta L. Miller & J. Miller, 1972  Morelos Skipper  NE to S-central Mexico (TAMPS, COAH, DF, MOR, VER)  iN  M T S L

Paratrytone kemneri Steinhauser, 1996  Kemner's Skipper  N to S-central Mexico  M T S

Paratrytone raspa (Evans, 1955)  Raspa Skipper  [= miahua] SW Mexico (MICH to OAX)  iN  M T S L Ha

Paratrytone pilza (Evans, 1955)  Pilza Skipper  S-central Mexico (GTO, DF, TLAX, PUE)  M S

Paratrytone polyclea Godman, 1900  Polyclea Skipper  SW Mexico (GRO, OAX, CHIS)  iN  M S L

Paratrytone rhexenor Godman, 1900  Crazy-spotted Skipper  NW & NE Mexico to OAX, VER  iN  M S

Paratrytone aphractoia Dyar, 1914  Snowball-spotted Skipper  NW & S-central Mexico  iN  M T S L Ha

Paratrytone samenta (Dyar, 1914)  Samenta Skipper  W Mexico (SON to OAX); ref.13  iN  M S L

Paratrytone omiltemensis Steinhauser, 1996  Omiltemi Skipper  SW Mexico (MICH to OAX)  iN  M T S Ha

Megaleas  Godman, 1901  

Megaleas syrna (Godman & Salvin, 1879)  Megaleas Skipper  Costa Rica  M T G

Tisias  Godman, 1901  

Tisias myna (Mabille, 1889)  Myna Ruby-eye  Costa Rica and Panama  iN  M T OI S

Tisias caesena (Hewitson, 1867)  TL: Brazil  M T S

Tisias lesueur (Latreille, [1824])    iN

Tisias lesueur canna Evans, 1955  TL: Peru  M T

Tisias lesueur lesueur (Latreille, [1824])  TL: USA  M S

Tisias carystoides Nicolay, 1980  TL: Ecuador  M T

Xeniades  Godman, 1900  [includes Cravera, Tixe & Putuma]; ref.13,16

Xeniades putumayo (Constantino & Salazar, 2013)  Putumayo Ruby-eye  C Colombia; ref.58,13,16  iN

Xeniades hermoda (Hewitson, 1870)  Quadrate Ruby-eye  Panama to Ecuador; ref.13  M T S

Xeniades quadrata (Herrich-Schäffer, 1869)  ref.13  iN  M T S

Xeniades rinda (Evans, 1955)  Venezuela; ref.13  M T

Xeniades orchamus (Cramer, 1777)  Smear-spotted Skipper  E & W Mexico to Argentina  iN  M T S L

Xeniades difficilis Draudt, 1923  Peru, Bolivia; ref.13  M T L G

Xeniades pteras Godman, 1900  Band-spotted Skipper  S Mexico (Oax, Chis) to Colombia & Venezuela; ref.13  M S L

Xeniades chalestra (Hewitson, 1866)  Band-spotted Skipper    iN

Xeniades chalestra chalestra (Hewitson, 1866)  Trinidad, Guianas, Colombia, Venezuela to Peru, Bolivia, S Brazil, Paraguay  M T S

Xeniades chalestra corna Evans, 1955  SE Brazil  M S

Xeniades victoria Evans, 1955  Victoria Skipper  SE Brazil  iN  M S

Xeniades rara (de Jong, 1983)  Rara Skipper  Surinam, French Guiana; ref.13  M T S

Xeniades laureatus Draudt, 1923  Laureatus Skipper  Bolivia; ref.13  M T S G

Vacerra  Godman, 1900  

Vacerra cervara Steinhauser, 1974  Cervara Skipper  E & W Mexico to Panama  iN  M T S L

Vacerra lachares Godman, 1900  Lachares Skipper  Costa Rica  M S

Vacerra gayra (Dyar, 1918)  Gayra Skipper  E & W Mexico, Belize  iN  M S L Ha

Vacerra egla (Hewitson, 1877)  Egla Skipper  Nicaragua, Panama  M T S L

Vacerra litana (Hewitson, 1866)  Litana Skipper  [= MEX & Central American records for bonfilus aeas], E & W Mexico to Gu ...  iN  M T S L

Vacerra bonfilius (Latreille, [1824])  Bonfilus Skipper    iN

Vacerra bonfilius bonfilius (Latreille, [1824])  Peru, S Brazil, Paraguay  M T

Vacerra bonfilius bonta Evans, 1955  Bolivia

Vacerra caniola (Herrich-Schäffer, 1869)  Caniola Skipper    iN

Vacerra caniola caniola (Herrich-Schäffer, 1869)  Guatemala to Venezuela  M T S L

Vacerra caniola elva Evans, 1955  Peru, Paraguay

Vacerra evansi Hayward, 1938  Blue-haired Skipper  SE Brazil, Paraguay, N Argentina  iN

Vacerra hermesia (Hewitson, 1870)  Hermesia Skipper    iN

Vacerra hermesia hermesia (Hewitson, 1870)  Ecuador  M T S L

Vacerra hermesia cecropterus (Draudt, 1923)  Peru, Bolivia, Argentina  M T S L G

Oligoria  Scudder, 1872  [includes Cobaloides and Jongiana]; ref.13

Oligoria unica (de Jong, 1983)  Unique Skipper  Surinam; ref.13  M T

Oligoria argus (Hayward, 1939)  SE S. Amer.; ref.13

Oligoria locutia (Hewitson, 1876)  Locutia Skipper  SE Brazil; ref.13  iN  M T S

Oligoria maculata (W. H. Edwards, 1865)  Twin-spot Skipper  SE USA (E TX to FL, N to NC)  iN  M T S L1 L2 I G

Oligoria percosius (Godman, 1900)  Double-dotted Skipper  S TX, E & SW Mexico to Guyana; ref.10  iN  M T S1 S2 L

Oligoria rindgei (H. Freeman, 1969)  Rindge’s Skipper  S Mexico; ref.10  iN  M T G

Oligoria obtena Grishin, 2023  Ecuador; ref.153

Oligoria lucifer (Hübner, [1831])  Lucifer Skipper  W Mexico to Argentina; ref.10  iN  M T S L Ha

Oligoria mustea (H. Freeman, 1979)  Muste Skipper  W Mexico (Colima, Chiapas); ref.10  iN  M S

Atrytonopsis  Godman, 1900  

Atrytonopsis quinteri Burns, 2015  Crystal Skipper  Bogue Banks and Bear Islands, NC  iN  M S L1 L2 L3 L4 I Ha

Atrytonopsis loammi (Whitney, 1876)  Loamm's Skipper  FL, SC, formerly far S NC  iN  M T S1 S2 L G

Atrytonopsis hianna (Scudder, 1868)  Dusted Skipper    iN

Atrytonopsis hianna hianna (Scudder, 1868)  New England to SE SASK, S to GA, AR & LA  M S L1 L2 L3 Ha G

Atrytonopsis hianna turneri H. Freeman, 1948  E MT, E WY, E CO, NE NM, ND to N-central TX  M T S L

Atrytonopsis deva (W. H. Edwards, 1877)  Deva Skipper  C & SE AZ, SW NM, NW to S-central Mexico  iN  M T S L Ha G

Atrytonopsis vierecki (Skinner, 1902)  Viereck’s Skipper  SW & SE UT, S CO, AZ, NM, far W OK, N & W TX, N COAH  iN  M T S L1 L2 L3 Ha G

Atrytonopsis lunus (W. H. Edwards, 1884)  Moon-marked Skipper  C & SE AZ, SW NM, SON  iN  M T S L I G

Atrytonopsis frappenda (Dyar, 1920)  Pedregal Skipper  [= zweifeli] Mexico; ref.96  iN  M T S G

Atrytonopsis llorentei A. Warren, 2009  Llorente's Skipper  West-central Mexico (MICH, MEX, SIN); ref.109  M T Ha

Atrytonopsis austinorum A. Warren, 2011  Austins' Skipper  Jalisco, Mexico; ref.110  M T

Atrytonopsis cestus (W. H. Edwards, 1884)  Cestus Skipper  far NW Mexico (SON), SE AZ  iN  M T S L I Ha G

Atrytonopsis python (W. H. Edwards, 1882)  Python Skipper  far S NV, AZ, NM, W TX, CHIH, SON  iN  M T S L G

Atrytonopsis margarita (Skinner, 1913)    iN  M T L

Atrytonopsis pittacus (W. H. Edwards, 1882)  White-barred Skipper    iN

Atrytonopsis pittacus pittacus (W. H. Edwards, 1882)  C & SE AZ, SW NM, W TX, SON  M S Ab L G

Atrytonopsis pittacus Southern segregate ,  SW Mexico (AGS, GRO, OAX)  M S

Atrytonopsis edwardsi W. Barnes & McDunnough, 1916  Sheep Skipper  N-central and NW Mexico, S AZ, SW NM, W TX  iN

Atrytonopsis edwardsi edwardsi W. Barnes & McDunnough, 1916  N-central and NW Mexico, S AZ, SW NM, W TX  iN  M T S L1 L2 I G

Atrytonopsis edwardsi rupilius (Schaus, 1913)  W Mexico; ref.151  M T

Atrytonopsis ovinia (Hewitson, 1866)  Ovinia Skipper  [= zaovinia], S-central Mexico to Costa Rica  iN  M T S L Ha

Thespieus  Godman, 1900  

Thespieus pinda Evans, 1955  Pinda Skipper  Peru (Limbani, Carabaya)  M S

Thespieus inez Nicolay, 1973  Inez' Skipper  Colombia (Arcabuco, 2200m, Dept. Boyaca)  M T

Thespieus tihoneta (Weeks, 1901)  Tihoneta Skipper    iN

Thespieus tihoneta ovallei E. Bell, 1938  Colombia  M T S

Thespieus tihoneta pampa Evans, 1955  Peru

Thespieus tihoneta tihoneta (Weeks, 1901)  Peru, Bolivia, Argentina  M T S G

Thespieus opigena (Hewitson, 1866)  Opigena Skipper  Ecuador, Peru, Bolivia  iN  M T S L

Thespieus guerreronis Dyar, 1913  Mexico (Gue); ref.153  M T

Thespieus dalman (Latreille, [1824])  Chalk-marked Skipper  E & W Mexico to Argentina  iN  M T S L

Thespieus mandal Grishin, 2023  SE Brazil; ref.153

Thespieus ethemides (Burmeister, 1878)  Ethemides Skipper  S Brazil, Paraguay, Argentina  iN  M T S G

Thespieus jora Evans, 1955  Jora Skipper  SE Brazil, Uruguay  iN  M S

Thespieus macareus (Herrich-Schäffer, 1869)  Chestnut-marked Skipper  TX (rare stray), E & W Mexico to Colombia & Venezuela  iN  M T S L G

Thespieus aspernatus Draudt, 1923  Aspernatus Skipper  SE Mexico to Venezuela, S Brazil, Paraguay & Argentina  iN  M T S G

Thespieus castor Hayward, 1948  Castor Skipper  SE Brazil  M S

Thespieus fulvangula (Weymer, 1890)  Fulvangula Skipper  Ecuador, Peru  M T S

Thespieus lutetia (Hewitson, 1866)  Lutetia Skipper  SE Brazil, Argentina  iN  M T S

Thespieus xarippe (A. Butler, 1870)  Xarippe Skipper    iN

Thespieus xarippe hallia Evans, 1955  Colombia, Peru  M S

Thespieus xarippe xarippe (A. Butler, 1870)  S Brazil, Paraguay  M T S

Thespieus xarina Hayward, 1948  Xarina Skipper  S Brazil, Paraguay, Argentina (Colombia records probably in error)  iN  M S

Thespieus othna (A. Butler, 1870)  Othna Skipper    iN

Thespieus othna tinka Evans, 1955  Panama & Colombia  M S

Thespieus othna othna (A. Butler, 1870)  Venezuela, Colombia, Ecuador, Peru, Bolivia  M T S L G

Thespieus thona Evans, 1955  Thona Skipper  Peru   iN  M S

Thespieus argentina Draudt, 1923  Argentine Skipper  S Peru, Bolivia, Argentina  iN  M T S G

Thespieus duidensis E. Bell, 1932  Mt. Duida Skipper  S Venezuela  M T

Thespieus haywardi Evans, 1937  Hayward's Skipper  SE Brazil, Argentina  iN  M S

Thespieus peruviae Lindsey, 1925  Peruvian Skipper  Ecuador, Peru  M S

Thespieus vividus (Mabille, 1891)  Vivid Skipper  SE Brazil  iN  M S

Thespieus hieroglyphica Draudt, 1923  Hieroglyphic Skipper  Peru, Bolivia  M T S G

Thespieus tapayuna J. Zikán, 1938  Tapayuna Skipper  SE Brazil  iN  M S

Thespieus zikani O. Mielke, 1971  Zikan's Skipper  Colombia  M S

Thespieus caraca Evans, 1955  Caraca Skipper  S Brazil  M S

Thespieus maacki Carneiro, Dolibaina, O. Mielke & Casagrande, 2014  Brazil; ref.50

Thespieus catochra (Plötz, 1882)  Catochra Skipper  S Brazil, Argentina  iN  M T S

Thespieus himella (Hewitson, 1868)  Himella Skipper  SE Brazil  iN  M T S

Thespieus abauna J. Zikán, 1938  Abauna Skipper  SE Brazil  M S

Thespieus abatira J. Zikán, 1938  Abatira Skipper  S Brazil  iN  M S

Thespieus matucanae Lindsey, 1925  Matucana Skipper  Peru  M S

Thespieus fassli (Draudt, 1923)  Fassl's Skipper  Peru  iN  M T S G

Thespieus homochromus O. Mielke, 1978  Homochromus Skipper  SE Brazil  iN  M S

  Tribe Hesperiini, Subtribe Moncina

Gracilata  Grishin, 2022  

Gracilata quadrinotata (Mabille, 1889)  Four-marked Skipper  Costa Rica and Panama; ref.13  M T S

Psoralis  Mabille, 1904  [includes Pamba and Saniba]; ref.13

Psoralis pamba (Evans, 1955)  TL: Ecuador  M T

Psoralis idee (Weeks, 1901)  TL: Bolivia  M T S

Psoralis stacara (Schaus, 1902)    iN  M T S

Psoralis magnamacus Grishin, 2023  Panama; ref.153

Psoralis sabina (Plötz, 1882)  S Brazil; ref.13  iN  M T S

Psoralis laska (Evans, 1955)  Brazil; ref.13  M T

Psoralis arva (Evans, 1955)  S Brazil; ref.13  iN  M T

Psoralis umbrata (Erschoff, 1876)    iN  M S L

Psoralis calcarea (Schaus, 1902)  Calcarea Skipper  S Brazil; ref.13  iN

Psoralis calcarea ponda (Evans, 1955)  Costa Rica to Colombia  M T S

Psoralis calcarea calcarea (Schaus, 1902)  Guianas, E & SE Brazil  M T

Psoralis visendus (E. Bell, 1942)  Visendus Skipper  Colombia, Ecuador; ref.13  iN  M T

Hermio  Grishin, 2022  ref.13

Hermio hermione (Schaus, 1913)  Hermione Skipper  S Mexico (Chiapas) to Panama; ref.13  iN  M T S L

Hermio vina (Evans, 1955)  Peru; ref.13  M T

Alychna  Grishin, 2019  ref.5

Alychna victa (Evans, 1955)  Peru; ref.5  iN  M S

Alychna chulumani Medeiros, Dolibaina & O. Mielke, 2020  TL: Bolivia

Alychna degener (Plötz, 1882)  Ecuador; ref.5  iN  M T S

Alychna zenus (E. Bell, 1942)  Ecuador; ref.5  M T

Alychna ayonis Grishin, 2023  Ecuador; ref.153

Alychna exclamationis (Mabille, 1898)  Peru, Bolivia; ref.5  iN  M T S

Alychna venta (Evans, 1955)  Venezuela; ref.5  M T

Alychna argenteus Medeiros, Dolibaina & Casagrande, 2020  TL: Peru

Alychna gota (Evans, 1955)  Colombia; ref.13  M T

Zalomes  E. Bell, 1947  

Zalomes biforis (Weymer, 1890)    iN  M T S

Zalomes naco Steinhauser, 1991  TL: Ecuador  M T

Zalomes cordillera (L. Miller & J. Miller, 1972)  TL: Ecuador  M T

Zalomes allynorum (L. Miller & J. Miller, 1972)  TL: Ecuador  M T

Zalomes merida Evans, 1955  TL: Venezuela  M T

Zalomes illimanensis Evans, 1955    iN  M T

Zalomes banco Steinhauser, 1991  TL: Ecuador  M T

Zalomes conspicua (Hayward, 1942)  TL: Ecuador  M S

Zalomes coto Steinhauser, 1991  TL: Ecuador  M T

Zalomes wanda Evans, 1955  TL: Peru  M T

Wahydra  Steinhauser, 1991  

Wahydra tassa (Evans, 1955)  [= Dalla curia] Colombia; ref.93  M T

Wahydra nieblensis Steinhauser, 1991  TL: Ecuador  M T

Wahydra bella (Hayward, 1939)    iN  M S

Wahydra thisbe (Hayward, 1942)    iN  M T S

Wahydra ekka (Evans, 1955)    iN  M T

Wahydra dores (E. Bell, 1959)    iN  M T

Wahydra vola (Evans, 1955)  TL: Peru  M T

Wahydra shueyi Carneiro, Henao, O. Mielke & Casagrande, 2015  Ecuador; ref.93

Wahydra cuzcona Grishin, 2023  Peru: Cuzco; ref.153

Wahydra kenava (A. Butler, 1870)    iN  M T S

Wahydra curtis Henao, O. Mielke, Carneiro & Casagrande, 2015  Bolivia; ref.93  iN

Wahydra mapiriensis Casagrande, Henao, Carneiro & O. Mielke, 2015  Bolivia; ref.93

Wahydra subhebetis Steinhauser, 1991  TL: Colombia  M T

Wahydra banios Grishin, 2023  Ecuador; ref.153

Wahydra trinitas Henao, O. Mielke, Carneiro & Casagrande, 2015  Colombia; ref.93

Wahydra carneiroi Henao, Gaviria & Salazar, 2017  Colombia; ref.94

Wahydra graslieae A. Warren, Carneiro & Dolibaina, 2018  Ecuador; ref.54  iN

Adlerodea  Hayward, 1940  

Adlerodea lemba Evans, 1955  TL: Brazil (Am)  M T

Adlerodea modesta Hayward, 1940    iN

Adlerodea petrovna (Schaus, 1902)  Petrovna Skipper  S Mexico to S Brazil  M T S

Adlerodea asema (Mabille, 1891)  Honduras Skipper  Honduras; ref.13  iN  M T G

Adlerodea mineira O. Mielke, 1968    iN  M T

Adlerodea subpunctata (Hayward, 1940)  ref.13

Adlerodea subpunctata subpunctata (Hayward, 1940)  Argentina; ref.13  M T S

Adlerodea subpunctata intermedia (de Jong, 1983)  TL: Surinam  M T

Adlerodea subpunctata sempa (Evans, 1955)  TL: Guyana  M T S

Mucia  Godman, 1900  

Mucia zygia (Plötz, 1886)  Black-dotted Skipper  E & W Mexico to Argentina  iN  M T S L

Mucia rusta (Evans, 1955)  ref.140  M T

Ralis  Grishin, 2019  ref.5

Ralis immaculatus (Hayward, 1940)  [= concolor]; ref.5,13  M T

Ralis coyana (Schaus, 1902)  S Brazil; ref.5  M T

Vinius  Godman, 1900  

Vinius phellus (Mabille, 1883)  ref.20  M T S

Vinius exilis (Plötz, 1883)    iN  M T S L

Vinius letis (Plötz, 1883)    iN  M T S

Vinius pulcherrimus Hayward, 1934    iN

Vinius sagitta (Mabille, 1889)  Sagitta Skipper  Nicaragua to upper Amazon  iN  M T OI S

Vinius sophistes (Dyar, 1918)  Mexico; ref.20  M T

Vinius tryhana (Kaye, 1914)  Gold-washed Skipper    iN

Vinius tryhana tryhana (Kaye, 1914)  E Mexico to Costa Rica, Guyana, Trinidad  M T S L I G

Vinius tryhana istria Evans, 1955  TL: Brazil (Pr)  M T

Rhinthon  Godman, 1900  

Rhinthon osca (Plötz, 1882)  Osca Skipper  S TX (as stray); E & SW Mexico to Venezuela & Ecuador, Trinidad  iN  M T OI S L

Rhinthon cubana (Herrich-Schäffer, 1865)  Cuban Skipper  Greater Antilles  iN  M T S

Rhinthon andricus (Mabille, 1895)  S Brazil; ref.20  M S

Rhinthon aqua (Evans, 1955)  Colombia; ref.20  M S

Rhinthon braesia (Hewitson, 1867)    iN  M T S

Rhinthon molion (Godman, 1901)  Blue-based Skipper  E Mexico to Ecuador  iN  M T S HS L

Rhinthon bajula (Schaus, 1902)  S. Brazil; ref.13  iN  M T S

Cynea  Evans, 1955  [includes Quinta and Nycea as subgenera]; ref.13

Cynea trimaculata (Herrich-Schäffer, 1869)  Trimaculata Skipper  Brazil  iN  M T S

Cynea corisana (Plötz, 1882)  Corsiana Skipper  Nicaragua to Venezuela  M S

Cynea bistrigula (Herrich-Schäffer, 1869)  Bistrigula Skipper  Guianas, Venezuela, Brazil, Bolivia  iN  M T S

Cynea nigricola H. Freeman, 1969  Nigricola Skipper  S Mexico  M T S

Cynea megalops (Godman, 1900)  Megalops Skipper  [= Papias nigrans], E Mexico to Colombia & W Ecuador  iN  M T S1 S2

Cynea cynea (Hewitson, 1876)  Cynea Skipper  S Mexico to Venezuela & Brazil  iN  M T S1 S2

Cynea aureofimbra Grishin, 2023  Ecuador; ref.153

Cynea erebina (Möschler, 1879)  [= hycsos] Colombia to Brazil; ref.153

Cynea erebina erebina (Mabille, 1891)  Venezuela, Colombia, Ecuador  M T S

Cynea erebina somba Evans, 1955  Brazil  M T S

Cynea quada Grishin, 2023  Ecuador; ref.153

Cynea cleochares (Mabille, 1891)  Venezuela; ref.153  M T

Cynea diluta (Herrich-Schäffer, 1869)  Diluted Skipper  [= osembo] S Mexico to S Brazil; ref.13  iN  M T S

Cynea anthracinus (Mabille, 1877)  Anthracinus Skipper    iN

Cynea anthracinus anthracinus (Mabille, 1877)  S Mexico, Guatemala  M T S

Cynea anthracinus holomelas (Mabille, 1891)  Costa Rica to Peru, Guianas  M T S

Cynea cyrus (Plötz, 1882)  Cyrus Skipper  TL: Brazil (RJ)

Cynea cyrus hippo Evans, 1955  Trinidad  M T

Cynea cyrus rhino Evans, 1955  French Guiana, Brazil (Amazonas), Peru (Iquitos)  M T

Cynea cyrus cyrus (Plötz, 1882)  SE Brazil  M T

Cynea rope Grishin, 2022  Rope Skipper  Nicaragua; ref.13  M T S

Cynea iquita (E. Bell, 1941)  Iquitos Skipper  Peru (Iquitos), Brazil (Amazonas), Surinam  M T S

Cynea popla Evans, 1955  Popla Skipper  Panama to S Brazil, Guianas, Trinidad  iN  M T1 T2 T3 S

Cynea robba Evans, 1955  Robba Skipper  TL: Guyana

Cynea robba robba Evans, 1955  Guyana, Brazil (Maranham)  M T

Cynea robba nippa Evans, 1955  Brazil (Matto Grosso, DF, Amazonas)  M T S

Cynea melius (Geyer, 1832)  Melius Skipper  SE Brazil  iN  M T S L

Cynea irma (Möschler, 1879)  Fogged Skipper  E Mexico to S Brazil  iN  M T S L

Cynea achirae Grishin, 2023  [MX "cannae"] E & W Mexico; ref.153  M T

Cynea cannae (Herrich-Schäffer, 1869)  Canna Skipper  El Salvador to Argentina; ref.13  iN  M T S1 S2 L I

Onophas  Godman, 1900  

Onophas columbaria (Herrich-Schäffer, 1870)  Blue-glossed Skipper    iN

Onophas columbaria columbaria (Herrich-Schäffer, 1870)  SE Mexico to Bolivia and C Brazil  M T S L

Onophas columbaria distigma E. Bell, 1930  TL: Brazil (SC)  M T S

Onophas columbaria flossites (M. Butler, 1874)  TL: Brazil (Am)  M T

Onophas watsoni E. Bell, 1930  TL: Brazil (SC)  M T S

Chitta  Grishin, 2019  ref.5

Chitta chittara (Schaus, 1902)  S. Amer.; ref.5  M T

Gufa  Grishin, 2022  

Gufa gulala (Schaus, 1902)  Gulala Skipper  Panama to Bolivia & S Brazil; ref.13  M T

Gufa fusca (Hayward, 1940)  S Brazil; ref.13  M T S

Eutus  Grishin, 2022  ref.13

Eutus amazonicus Grishin, 2023  Amazonian region; ref.153

Eutus incus Grishin, 2023  Peru: Cuzco; ref.153

Eutus yesta (Evans, 1955)  Peru; ref.13  M T

Eutus rastaca (Schaus, 1902)  [= brema] S Brazil; ref.13  iN  M T

Eutus mubevensis (E. Bell, 1932)  Paraguay; ref.13  iN  M T S

Eutus septemaculatus Grishin, 2023  Brazil; ref.153

Godmia  Grishin, 2022  ref.13

Godmia chlorocephala (Godman, 1900)  Green-headed Skipper  Panama; ref.13  M T OI S

Godmia viridicapita Grishin, 2023  Ecuador; ref.153

Thoon  Godman, 1900  

Thoon modius (Mabille, 1889)  Moody Skipper  E & W Mexico to upper Amazon and Bolivia  iN  M T OI S L

Halotus  Godman, 1900  

Halotus angellus (Plötz, 1886)  Angellus Skipper  Costa Rica and Panama  iN  M T S L

Halotus jonaveriorum Burns, 1992  John-and-Avery's Skipper  W Mexico  iN  M T S L Ha

Halotus rica (E. Bell, 1942)  Costa Rican Skipper  W Mexico to Costa Rica  iN  M T S L

Curva  Grishin, 2022  ref.16

Curva hyagnis (Godman, 1900)  Hyagnis Skipper  ref.16  iN

Curva hyagnis hyagnis (Godman, 1900)  E Mexico to Costa Rica  M T S L

Curva hyagnis meraca (Hayward, 1947)  TL: Brazil

Curva darienensis (Gaviria, Siewert, O. Mielke & Casagrande, 2018)  Panama; ref.39,123,16

Alerema  Hayward, 1942  

Alerema simplex (E. Bell, 1930)  Simple Skipper  W Mexico and Yucatan to Argentina; ref.13  M T S

Crinifemur  Steinhauser, 2008  

Crinifemur viridans Steinhauser, 2008  Guyana; ref.122  iN  M S

Rhomba  Grishin, 2022  ref.13

Rhomba mirnae (Siewert, Nakamura & O. Mielke, 2014)  Mesoamerica; ref.5,140

Rhomba gertschi (E. Bell, 1937)  Gertsch's Skipper  [= Psoralis panamensis] Panama; ref.123,13  M T

Rhomba pulla Grishin, 2023  Peru:Cuzco; ref.153

Niconiades  Hübner, [1821]  

Niconiades incomptus Austin, 1997  Half-tailed Skipper  E & W Mexico to Costa Rica  iN  M S L

Niconiades xanthaphes Hübner, [1821]  Stub-tailed Skipper  Panama to Uruguay and Paraguay  iN  M T S L

Niconiades gladys Evans, 1955    iN

Niconiades tina Nicolay, 1980  TL: Ecuador  M T

Niconiades yoka Evans, 1955    iN  M S

Niconiades cydia (Hewitson, 1876)    iN  M T

Niconiades linga Evans, 1955    iN  M T S

Niconiades comitana H. Freeman, 1969  Comitana Skipper  SE Mexico (Chiapas and Veracruz)  iN  M T S

Niconiades victoria Grishin, 2023  E Mexico to Guatemala (at least); ref.153  M S L

Niconiades nikko Hayward, 1948  Nikko Skipper  Colombia to S Brazil and Paraguay  iN  M S L

Niconiades loda Evans, 1955  TL: Brazil (MG)

Niconiades caeso (Mabille, 1891)    iN  M T

Niconiades nabona Evans, 1955    iN  M S

Niconiades merenda (Mabille, 1878)    iN  M T

Niconiades centralis O. Mielke, 1967    iN  M T OI

Niconiades bifurcus Anderson, 2008  Bifurcus Skipper  Nicaragua; ref.56  M T

Niconiades derisor (Mabille, 1891)  Derisor Skipper  [= vista] Oaxaca to Ecuador; ref.13  M T S

Niconiades viridis (E. Bell, 1930)    iN  M T S

Niconiades peri (Evans, 1955)  Brazil (Pa); ref.13  iN  M S

Joanna  Evans, 1955  

Joanna boxi Evans, 1955  TL: Guyana  M T

Joanna joanna Evans, 1955  Joanna’s Skipper  E & W Mexico to Panama  iN  M T S L

Joanna clara Steinhauser, 2007  Clara’s Skipper  TL: Brazil (RJ)  M T

Vinpeius  Austin, 1997  

Vinpeius tinga (Evans, 1955)  Freeman's Skipper  E Mexico to El Salvador, [Venezuela?]  iN  M T

Pares  Grishin, 2019  ref.5

Pares viridiceps (Mabille, 1889)  Panama; ref.13  M T OI

Pares maritza (Nicolay, 1980)  Maritza Skipper  Panama to W Ecuador; ref.5  M T S

Pares pares (E. Bell, 1959)  Paraguay; ref.5  iN  M T

Bruna  Evans, 1955  

Bruna bruna Evans, 1955  TL: ?  M T

Paracarystus  Godman, 1900  [= Punta]; ref.154

Paracarystus punta (Evans, 1955)  ref.154  M T

Paracarystus hypargyra (Herrich-Schäffer, 1869)  Hypargyra Skipper  SE Mexico to S Brazil  iN  M T S1 S2 L

Paracarystus menestries (Latreille, [1824])    iN

Paracarystus menestries rona (Hewitson, 1866)  TL: Brazil (Pa)  M T S1 S2 IL

Paracarystus menestries koza (A. Butler, 1870)  TL: Brazil (Pa)  M T

Paracarystus menestries menestries (Latreille, [1824])  TL: Brazil  M T S

Paracarystus evansi Hayward, 1938    iN  M S

Paracarystus ranka (Evans, 1955)  Peru; ref.13  M T

Viridina  Grishin, 2019  ref.5

Viridina viridis (E. Bell, 1942)  Ecuador; ref.5  M T

Viridina viridenex (Weeks, 1901)  Bolivia; ref.5  M S

Viridina subviridis (Hayward, 1940)  Ecuador; ref.5  M T

Moeris  Godman, 1900  

Moeris striga (Geyer, 1832)    iN  M S

Moeris seth O. Mielke, Carneiro & Casagrande, 2015  ref.52  iN

Moeris menopis (Schaus, 1902)    iN  M T S

Moeris strada Evans, 1955    iN  M T1 T2 T3

Moeris stroma Evans, 1955  Flag Skipper  E & W Mexico to Colombia  iN  M T1 T2 S L

Moeris nut O. Mielke, Carneiro & Casagrande, 2015  ref.52

Moeris anna (Mabille, 1898)  SE S. Amer; ref.5  iN  M T S

Tricrista  Grishin, 2019  ref.5

Tricrista canta (Evans, 1955)  Colombia; ref.13  M T S

Tricrista slopa (Evans, 1955)  Guyana; ref.13  M T

Tricrista advena (Draudt, 1923)  ref.5

Tricrista advena advena (Draudt, 1923)  TL: Brazil  M T S G

Tricrista advena jon (Nicolay, 1980)  TL: Ecuador  M T

Tricrista roppai (O. Mielke, 1980)  S Brazil; ref.5  M T

Tricrista cristina (Evans, 1955)  Colombia; ref.5  M T

Tricrista crista (Evans, 1955)  Venezuela; ref.5  M T

Tricrista cristatus (E. Bell, 1930)  S Brazil; ref.5  M T S

Tricrista aethus (Hayward, 1951)  Disjunct Skipper  Nicaragua, Bolivia; ref.13  M T S IL

Tricrista circellata (Plötz, 1882)  S Brazil; ref.13  M T S

Tricrista taxes (Godman, 1900)  Taxes Skipper  Costa Rica and Panama; ref.13  M T S

Lancephallus  Grishin, 2023  ref.153

Lancephallus purpurus Grishin, 2023  Venezuela & The Guianas; ref.153  M T

Vettius  Godman, 1901  

Vettius pica (Herrich-Schäffer, 1869)  Two-toned Skipper  SE Mexico to upper Amazon; ref.154  M T S L

Vettius lafrenaye (Latreille, [1824])  Brazil  iN  M T S

Vettius triangularis (Hübner, [1831])  Triangular Skipper  Panama to Guianas and Amazonas  iN  M S1 S2

Vettius richardi (Weeks, 1906)  Richard's Skipper  Panama, Colombia, Venezuela, Guianas, Brazil, Ecuador  iN  M S IL

Vettius monacha (Plötz, 1882)    iN  M T S

Vettius jabesa (A. Butler, 1870)    iN  M T S

Vettius crispa Evans, 1955  TL: Brazil (Am)  M T

Vettius phyllus (Cramer, 1777)  Phyllus Skipper    iN

Vettius phyllus phyllus (Cramer, 1777)  [= marcus] Panama and Trinidad to C Brazil; ref.80  M T S1 S2 S3 L

Vettius phyllus pura Evans, 1955  TL: Peru  M T1 T2 L

Vettius phyllus yunga Evans, 1955  TL: Bolivia  M T

Vettius prona Evans, 1955  S Brazil; ref.154  M T1 T2 S

Vettius chagres Nicolay, 1973  Chagres Skipper  Panama & Colombia  M T S

Vettius mitsuko Nakahara, Nakamura & MacDonald, 2020  Mitsuko Skipper  Panama; ref.79  iN

Tigasis  Godman, 1900  [includes Arita]; ref.5

Tigasis wellingi (H. Freeman, 1969)  Welling's Skipper  S Mexico; ref.13  M T

Tigasis arita (Schaus, 1902)  Arita Skipper  in S. Amer. to Argentina; ref.5  iN  M T S1 S2 L

Tigasis corope (Herrich-Schäffer, 1869)  Corope Skipper  SE & S Brazil; ref.13  M T S

Tigasis zalates Godman, 1900  Zalates Skipper  Costa Rica to Ecuador  M T S

Tigasis physcoa (Hewitson, 1868)  TL: ?  M T S

Tigasis colomus (E. Bell, 1941)  TL: Colombia  M T

Tigasis perloides (Plötz, 1882)  S Brazil & Paraguay; ref.13  iN  M T S

Tigasis polistion (Schaus, 1902)    iN  M T

Tigasis serra (Evans, 1955)    iN  M T S

Inglorius  Austin, 1997  

Inglorius mediocris Austin, 1997  Mediocre Skipper  Campeche, N Guatemala, Belize  M S

Gallio  Evans, 1955  

Gallio coda (Evans, 1955)  [not Tigasis] Ecuador; ref.154  M T

Gallio garima (Schaus, 1902)  Trinidad; ref.13  iN  M T S

Gallio massarus (E. Bell, 1940)  S Brazil; ref.13  M T S

Gallio carasta (Schaus, 1902)  S Brazil  iN  M T S

Gallio imperatriz Carneiro, Dolibaina, O. Mielke & Casagrande, 2015  Brazil; ref.53

Gallio eti Carneiro, Dolibaina, O. Mielke & Casagrande, 2015  Brazil; ref.53

Gallio furtadoi Carneiro, Dolibaina, O. Mielke & Casagrande, 2015  Brazil; ref.53

Gallio seriatus (Mabille, 1891)  Venezuela; ref.13  iN  M T S

Gallio danius (E. Bell, 1941)  S Brazil; ref.13  M T S

Gallio madius (E. Bell, 1941)  S Brazil; ref.13  M T

Mnasicles  Godman, 1901  [includes Styriodes, Remella & Nausia as subgenera]; ref.13

Mnasicles dedecora (Plötz, 1882)  Dedecora Skipper  [= quaka], S Mexico to Ecuador; ref.13  M S

Mnasicles quota (Evans, 1955)  Guyana; ref.13  M T

Mnasicles lyco (Schaus, 1913)  Lyco Skipper  Costa Rica to Colombia; ref.13  M T S

Mnasicles browni (E. Bell, 1959)  Brown's Skipper  Costa Rica; ref.5,13  M T S L

Mnasicles nausiphanes (Schaus, 1913)  Cloud-forest Skipper  W Mexico to Panama; ref.13  M T S L

Mnasicles ecua Grishin, 2023  Ecuador; ref.153

Mnasicles remus (Fabricius, 1798)  Black-spot Remella  E & W Mexico to S Brazil & Paraguay; ref.13  iN  M T S L

Mnasicles duena (Evans, 1955)  Guatemalan Remella  S Mexico to Guatemala; ref.13  iN  M T S L

Mnasicles rita (Evans, 1955)  Rita’s Remella  E & W Mexico to Panama; ref.13  iN  M T S L

Mnasicles vopiscus (Herrich-Schäffer, 1869)  Cryptic Remella  SE Mexico to N Brazil; ref.13  iN  M T S L

Mnasicles cusillunia (Weeks, 1901)  Bolivia; ref.13  M S L

Mnasicles geta Godman, 1901  Violet-frosted Skipper  E Mexico to Costa Rica  iN  M T S L

Mnasicles hicetaon Godman, 1901  Gray Skipper  W Mexico and Yucatan to Trinidad and Paraguay  iN  M T S

Amblyscirtes  Scudder, 1872  [includes Stomyles, Mastor, Amblyteria & Flor as subgenera]; ref.11,13

Amblyscirtes carolina (Skinner, 1892)  Carolina Roadside-Skipper    iN  M T S L1 L2 G

Amblyscirtes reversa F. Jones, 1926  Reversed Roadside-Skipper    iN  M T S HS L

Amblyscirtes aesculapius (Fabricius, 1793)  Lace-winged Roadside-Skipper    iN  M S L1 L2 L3 L4 G

Amblyscirtes hegon (Scudder, 1863)  Pepper and Salt Skipper  E US  iN  M S L1 L2 L3

Amblyscirtes nemoris (W. H. Edwards, 1864)  E US; ref.150  M L1 L2 I Ha G

Amblyscirtes gelidus Grishin, Patterson & Pavulaan, 2023  MI & WV for now; ref.150  M S

Amblyscirtes matheri Patterson, Pavulaan & Grishin, 2023  SE US; ref.150

Amblyscirtes nysa W. H. Edwards, 1877  Nysa Roadside-Skipper    iN  M T S L1 L2 G

Amblyscirtes alternata (Grote & Robinson, 1867)  Dusky Roadside-Skipper  [= asella] SE US; ref.13  iN  M T S L1 L2 G

Amblyscirtes vialis (W. H. Edwards, 1862)  Common Roadside-Skipper    iN  M T S1 S2 L1 L2 L3 L4 L5 L6 I G

Amblyscirtes linda H. Freeman, 1943  Linda's Roadside-Skipper    iN  M T1 T2 T3 S L1 L2 L3 Ha G

Amblyscirtes aenus W. H. Edwards, 1878  Bronze Roadside-Skipper    iN

Amblyscirtes aenus erna H. Freeman, 1943  NE Mexico  M T1 T2 S HS L

Amblyscirtes aenus aenus W. H. Edwards, 1878  TL: USA (Co)  M T S G

Amblyscirtes aenus megamacula Scott, 1998  NW Mexico  M S HS L1 L2 I

Amblyscirtes aeratus Grishin, 2023  SW Mexico; ref.153  M T

Amblyscirtes fluonia Godman, 1900  Brassy Roadside-Skipper    iN  M T S L

Amblyscirtes cassus W. H. Edwards, 1883  Cassus Roadside-Skipper    iN  M T S L1 L2 I G

Amblyscirtes texanae E. Bell, 1927  Texas Roadside-Skipper    iN  M T1 T2 S L I G

Amblyscirtes belli H. Freeman, 1941  Bell's Roadside-Skipper    iN  M T1 T2 T3 T4 T5 T6 S HS L1 L2 L3 G

Amblyscirtes celia Skinner, 1895  Celia’s Roadside-Skipper    iN  M T S1 S2 L1 L2 I Ha G

Amblyscirtes tolteca Scudder, 1872  Toltec Roadside-Skipper    iN

Amblyscirtes tolteca tolteca Scudder, 1872  S TX (as stray), E & W Mexico to Nicaragua  M S L

Amblyscirtes tolteca prenda Evans, 1955  SE AZ, far NW Mexico  M S L1 L2 I

Amblyscirtes nereus (W. H. Edwards, 1876)  Slaty Roadside-Skipper    iN  M T S HS L1 L2 I G

Amblyscirtes eos (W. H. Edwards, 1871)  Dotted Roadside-Skipper    iN  M T S L1 L2 I G

Amblyscirtes arizonae H. Freeman, 1993  Elissa Roadside-Skipper  USA (Az) to NW Mexico; ref.11  iN  M T S L1 L2 I

Amblyscirtes elissa Godman, 1900  Elissa Roadside-Skipper  SW Mexico; ref.11  iN  M S L Ha

Amblyscirtes oslari (Skinner, 1899)  Oslar’s Roadside-Skipper    iN  M T S L1 L2 G

Amblyscirtes brocki H. Freeman, 1992  Brock’s Roadside-Skipper  TL: Mexico (Son)

Amblyscirtes brocki brocki H. Freeman, 1992  TL: Mexico (Son)  M T S L

Amblyscirtes brocki Southern segregate ,   M T S L1 L2

Amblyscirtes exoteria (Herrich-Schäffer, 1869)  Large Roadside-Skipper    iN

Amblyscirtes exoteria exoteria (Herrich-Schäffer, 1869)  TL: ?  M T S L1 L2 I G

Amblyscirtes exoteria Southern segregate ,   M S L Ha

Amblyscirtes phylace (W. H. Edwards, 1878)  Orange-headed Roadside-Skipper    iN  M T S L I H G

Amblyscirtes fimbriata (Plötz, 1882)  Orange-edged Roadside-Skipper    iN

Amblyscirtes fimbriata fimbriata (Plötz, 1882)  TL: Mexico  M T S L1 L2 I G

Amblyscirtes fimbriata pallida H. Freeman, 1993  TL: Mexico (Mex)  M T L Ha

Amblyscirtes anubis (Godman, 1900)  Half-edged Roadside-Skipper    iN  M T S L

Amblyscirtes chrysoplea Grishin, 2023  SW Mexico; ref.153

Amblyscirtes chrysomisa Grishin, 2023  S Mexico; ref.153  M T

Amblyscirtes repta (Evans, 1955)  W Mexico; ref.153  M T

Amblyscirtes novimmaculatus A. Warren, 1998  Immaculate Roadside-Skipper  SW Mexico; ref.111  iN  M T L Ha

Amblyscirtes patriciae (E. Bell, 1959)  Patricia’s Roadside-Skipper  TL: Guatemala  M T S

Amblyscirtes raphaeli H. Freeman, 1973  Giant Roadside-Skipper  TL: Mexico (Oax)  M T S G

Amblyscirtes folia Godman, 1900  Larger Roadside-Skipper    iN  M T S L I Ha

Amblyscirtes meridus Grishin, 2023  E Mexico to Belize & Nicaragua; ref.153  M T

Amblyscirtes florus (Godman, 1900)  False Roadside-Skipper  W & SW Mexico; ref.13  iN  M T S L I

Vidius  Evans, 1955  

Vidius fraus (Godman, 1900)  Frosty-banded Skipper  E Mexico to Costa Rica; ref.13  iN  M T S L

Vidius dagon (Evans, 1955)  [= felus]; ref.140  M T S

Vidius catocala (Herrich-Schäffer, 1869)  [= dorpa]; ref.140  iN  M T S

Vidius pompeoides Grishin, 2022  Amazonian region; ref.16  M T

Vidius vidius (Mabille, 1891)    iN  M T S

Vidius similis O. Mielke, 1980    iN  M T S

Vidius cocalus (Hayward, 1939)  [= prado]; ref.140  iN  M T S

Vidius obscurior (Hayward, 1934)  Argentina; ref.140

Vidius tanna (de Jong, 1983)  Suriname; ref.151  M T

Rectava  Grishin, 2022  ref.13

Rectava nostra (Evans, 1955)  Brazil  iN

Rectava nostra nostra (Evans, 1955)  Brazil (Pa); ref.13  M T

Rectava nostra roraimae (O. Mielke & Casagrande, 1992)  Brazil (Rr); ref.13  M T

Rectava vorgia (Schaus, 1902)  Brazil (Pr); ref.13  iN  M T S

Rectava chiriquensis Grishin, 2023  Panama; ref.153

Rectava ignarus (E. Bell, 1932)  Bolivia; ref.13  iN  M T L

Rectava sobrinus (Schaus, 1902)  S Brazil; ref.13  M T S

Rectava cascatona (O. Mielke, 1992)  Brazil; ref.140  M T OI

Cobalopsis  Godman, 1900  

Cobalopsis zetus (E. Bell, 1942)  Mexico; ref.13  M T

Cobalopsis autumna (Plötz, 1882)  Autumna Skipper  SE Mexico to S Brazil  iN  M T S

Cobalopsis nero (Herrich-Schäffer, 1869)  Nero Skipper  [= zetus], E & S Mexico to S Brazil & Paraguay  iN  M T S1 S2 L

Cobalopsis similis O. Mielke, 1989  TL: Brazil (Ma)  M T

Cobalopsis monotona O. Mielke, 1989  TL: Brazil (MG)  M T

Cobalopsis valerius (Möschler, 1879)  [= miaba, Evans' "valerius" is ancus] Costa Rica to S Brazil & P ...; ref.13  iN  M T S1 S2

Cobalopsis dictys (Godman, 1900)  Bottom-spotted Skipper  E & S Mexico to Ecuador; ref.13  iN  M T S1 S2 L

Cobalopsis adictys Grishin, 2023  Costa Rica & Panama; ref.153

Artonia  Grishin, 2019  ref.5

Artonia artona (Hewitson, 1868)  White-veined Skipper  Nicaragua to S Brazil; ref.5  iN  M T S L

Lurida  Grishin, 2019  ref.5

Lurida lurida (Herrich-Schäffer, 1869)  ref.5  iN  M T S

Lurida zama (Hayward, 1939)  Argentina; ref.140  iN  M S

Corra  Grishin, 2019  ref.5

Corra conka (Evans, 1955)  Silver-plated Skipper  S Mexico to Panama

Corra conka argentus (H. Freeman, 1969)  S Mexico to El Salvador, at least; ref.140  M T S L Ha

Corra conka conka (Evans, 1955)  S Mexico to Panama  M T1 T2 S L

Corra catargyra (C. Felder & R. Felder, 1867)  ref.140  M T S

Corra coryna (Hewitson, 1866)  Silver-plated Skipper  ref.140  iN  M T S L

Corra xanthus (O. Mielke, 1989)  Brazil; ref.140  M T

Nastra  Evans, 1955  [includes Rayia as subgenus]; ref.16

Nastra celeus (Mabille, 1891)  ref.13  iN

Nastra celeus celeus (Mabille, 1891)  ref.13  M T S

Nastra celeus ochraceus (Biezanko & O. Mielke, 1973)  TL: Brazil (RS)  M T

Nastra celeus vetus (O. Mielke, 1969)  TL: Brazil (Pr)  M T

Nastra ethologus (Hayward, 1934)    iN  M S

Nastra chao (Mabille, 1898)    iN  M T S

Nastra nappa (Evans, 1955)  Brazil; ref.13  M T S

Nastra leucone (Godman, 1900)  Leucone Skipper  E Mexico to Costa Rica  iN  M T S L

Nastra subsordida (Mabille, 1891)  Three-dotted Skipper  [= trimaculata] Honduras to Panama; ref.13  M T S

Nastra leuconoides (Lindsey, 1925)  Peru; ref.16  M S

Nastra lherminier (Latreille, [1824])  Swarthy Skipper  E USA: KS to TX, E to FL, NJ, CT  iN  M S1 S2 L1 L2 L3 Ha G

Nastra fusca (Grote & Robinson, 1867)  Neamathla Skipper  [= neamathla] SE USA (FL, TX), NOT in Mexico or Central America; ref.16  iN  M T S L G

Nastra julia (H. Freeman, 1945)  Julia’s Skipper  [= hoffmanni], far SW USA, C & S TX, E & W Mexico to C. Rica  iN  M T S L1 L2 I

Nastra perigenes (Godman, 1900)  Pale-rayed Skipper  S TX, E & W Mexico to Colombia; ref.10,16  iN  M T S L

Nastra dryas (Hayward, 1940)  Paraguay, Argentina & S Brazil  iN

Lerodea  Scudder, 1872  

Lerodea erythrostictus (Prittwitz, 1868)    iN  M T S

Lerodea gracia Dyar, 1913    iN  M T

Lerodea emba Evans, 1955  TL: Peru  M S

Lerodea sonex Grishin, 2022  [= Evans' "xenos"] Peru  iN  M T

Lerodea eufala (W. H. Edwards, 1869)  Eufala Skipper    iN

Lerodea eufala eufala (W. H. Edwards, 1869)  S & C CA, to S NM, C & S TX-OK to VA-FL (strays N); to Argent.  M T S L1 L2 I1 I2 G

Lerodea eufala concepcionis (Strand, 1921)  TL: Chile  M T

Lerodea arabus (W. H. Edwards, 1882)  Violet-clouded Skipper  S AZ & NW Mexico (incl. BCS); ref.146  iN  M S1 S2 L Ha G

Lerodea dysaules Godman, 1900  Olive-clouded Skipper  [not arabus!] S TX to Costa Rica, W to MX: Sonora; ref.146  M S L

Lerodea casta Hayward, 1942  TL: Ecuador

Lerodea williamsi Hayward, 1942  TL: Ecuador

Fidius  Grishin, 2019  ref.5

Fidius fido (Evans, 1955)  Brazil; ref.5  M T S

Fidius ochraceus (O. Mielke, 1980)  Brazil; ref.5,140  M T

Cymaenes  Scudder, 1872  

Cymaenes tripunctus (Herrich-Schäffer, 1865)  Three-spotted Skipper    iN

Cymaenes tripunctus tripunctus (Herrich-Schäffer, 1865)  S FL, Bahamas, Cuba, Hispaniola, Puerto Rico, Jamaica, etc.  M T S L1 L2 G

Cymaenes tripunctus theogenis (Capronnier, 1874)  Chiapas to S Brazil & Paraguay  M T OI S1 S2

Cymaenes lepta (Hayward, 1939)    iN  M S

Cymaenes isus (Godman, 1900)  Fawn-spotted Skipper  [= Evans' "trebius"] SE AZ (rare stray), S TX, E & W MX to Co ...; ref.13,16  iN  M T S Ma L1 L2 L3 L4

Cymaenes edata (Plötz, 1882)  Venezuela; ref.13  M S

Cymaenes melaporphyrus Grishin, 2023  Mexico to Guatemala (at least); ref.153  M S

Cymaenes laureolus (Schaus, 1913)  Laureolus Skipper  Costa Rica to Ecuador  M T S

Cymaenes lochius (Plötz, 1882)  [Evans' "lochius" is Lerema lucius] Venezuela (at least); ref.13  M TI

Cymaenes loxa Evans, 1955  Brazil; ref.13  M T1 T2

Cymaenes albiventris O. Mielke, 1992  TL: Brazil (Ma)

Cymaenes albiventris albiventris O. Mielke, 1992  TL: Brazil (Ma)  M T

Cymaenes albiventris capixaba O. Mielke, 1992  TL: Brazil (ES)   M T OI

Cymaenes uruba (Plötz, 1886)    iN

Cymaenes uruba uruba (Plötz, 1886)  TL: Brazil  M T S

Cymaenes uruba taberi (Weeks, 1901)  TL: Bolivia  M T S G

Cymaenes alumna (A. Butler, 1877)  Alumna Skipper  SE Mexico to Bolivia & S Brazil  iN  M T S1 S2 L

Cymaenes chapa O. Mielke, 1968  TL: Brazil (DF)  M T

Cymaenes cavalla Evans, 1955    iN  M T S

Cymaenes tripunctata (Latreille, [1824])    iN  M T S

Cymaenes macintyrei (Hayward, 1939)  ref.140  M S

Cymaenes idria Evans, 1955    iN  M T

Cymaenes gisca Evans, 1955    iN  M T1 T2 T3 S L

Cymaenes laza O. Mielke, 1968    iN  M T

Cymaenes mabillei (Giacomelli, 1911)    iN  M S

Cymaenes riba O. Mielke, 1968  TL: Brazil (DF)  M T

Cymaenes chela Evans, 1955  TL: Venezuela  M T S

Cymaenes finca Cock, 1985  TL: Trinidad  M T

Cymaenes modestus (Hayward, 1943)  TL: Peru  M S

Cymaenes incomptus (Hayward, 1934)  ref.140  iN

Cymaenes catarinae (O. Mielke, 1989)  ref.140  M T

Cymaenes lumina (Herrich-Schäffer, 1869)  Misnamed Skipper  [= odilia, = trebius, =corescene] S. Amer.; ref.13,16  iN  M T S

Cymaenes aequatoria (Hayward, 1940)  Ecuador; ref.13  M T S

Cymaenes miqua (Dyar, 1913)  Peru; ref.13  M T

Cymaenes psyllus (Mabille, 1898)  TL: Bolivia  M T S

Cymaenes jamba Evans, 1955  TL: Argentina  M T

Monca  Evans, 1955  

Monca jera (Godman, 1900)  Jera Skipper  SW Mexico: MICH & MOR to CHIS  iN  M T S

Monca crispinus (Plötz, 1882)  Violet-patched Skipper  [= tyrtaeus], S TX, E & W Mexico to Colombia  iN  M T S L

Monca telata (Herrich-Schäffer, 1869)  Telata Skipper  Belize, Guatemala to Guyana, Trinidad and Tobago  iN  M T S L

Monca penda Evans, 1955    iN  M T S

Monca branca Evans, 1955    iN  M T

Monca ornata (Hayward, 1940)    iN  M S

Troyus  A. Warren & Turland, 2012  ref.112,10

Troyus turneri A. Warren & Turland, 2012  Turner's Skipper  Jamaica; ref.112  M T L Ha G

Troyus fabulosus Grishin, 2023  Fabulous Skipper  S TX (sporadic), E & W Mexico to Honduras; ref.19  M S L1 L2

Troyus fantasos (Cramer, 1780)  Fantastic Skipper  Costa Rica to Paraguay; ref.10,19  iN  M S

Troyus onaca (Evans, 1955)  Onaca Skipper  SE Mexico to El Salvador; ref.10  iN  M T1 T2 T3 S L

Troyus aurelius (Plötz, 1882)  Aurelius Skipper  Guatemala to C Brazil; ref.10  iN  M S L

Troyus phyllides (Röber, 1925)  Ex-marcus Skipper  [formerly marcus] S Mexico (Chia) to Argentina; ref.80  iN  M T S1 S2 L

Troyus diversa (Herrich-Schäffer, 1869)  ref.10  iN

Troyus diversa maeon (Mabille, 1891)  Nicaragua to Colombia  M T S

Troyus diversa lyrcea (Plötz, 1882)  TL: Brazil  M S

Troyus diversa donga (Evans, 1955)  TL: Peru  M T

Troyus diversa drona (Evans, 1955)  TL: Bolivia  M T

Troyus diversa diversa (Herrich-Schäffer, 1869)  TL: ?  M S L

Troyus drova (Evans, 1955)  Ecuador; ref.10  iN  M T L

Lerema  Scudder, 1872  [includes Morys and Geia as subgenera]; ref.13

Lerema pattenii Scudder, 1872  Overcast Skipper  [= Evans' "lumina", = bipunctata] Mexico to Colombia; ref.13  iN  M T OI OD S

Lerema accius (J. E. Smith, 1797)  Clouded Skipper  SE USA (strays N); ref.19  iN  M T S HS L1 L2 G

Lerema ochrius Grishin, 2023  Ocherous Skipper  SE AZ, S NM, W & S TX and MX (at least); ref.19  M T S Ma L1 L2 L3 I1 I2

Lerema liris Evans, 1955  Liris Skipper  E & W Mexico to Costa Rica  iN  M T1 T2 S L I H

Lerema ancillaris (A. Butler, 1877)  Plain skipper  Costa Rica and Trinidad to upper Amazon  iN  M T S

Lerema lucius Grishin, 2022  Lucius Skipper  [= Evans' "lochius"] W & E Mexico to Venezuela; ref.13  iN  M T S

Lerema lineosa (Herrich-Schäffer, 1865)  [= aethra]; ref.13  M T S

Lerema micythus (Godman, 1900)  Micythus Skipper  S & E Mexico to Colombia and Ecuador  iN  M T S L

Lerema ecuadorica Grishin, 2023  Ecuador; ref.153

Lerema compta (A. Butler, 1877)    iN  M T S

Lerema venias (E. Bell, 1942)  Venezuela; ref.13  M T

Lerema sinta (Evans, 1955)  TL: Brazil (Pa)  M T

Lerema sobra (O. Mielke, 1968)  TL: Brazil (DF)  M T

Lerema subgrisea (Mabille, 1898)    iN

Lerema subgrisea prada (Evans, 1955)  TL: Venezuela  M T

Lerema subgrisea paradoxa (de Jong, 1983)  TL: Surinam  M T

Lerema subgrisea subgrisea (Mabille, 1898)  TL: Bolivia  M T S

Lerema valda (Evans, 1955)  Valda Skipper  E & W Mexico to Costa Rica; ref.13  iN  M T S

Lerema ancus (Möschler, 1879)  [= Evans' "valerius"]; ref.13  iN  M T S

Lerema lyde (Godman, 1900)  Violet-studded Skipper  S & E Mexico to Costa Rica  iN  M T S L I

Lerema geisa (Möschler, 1879)  Geisa Skipper  Costa Rica to Colombia; ref.13  iN  M T S1 S2

Lerema etelka (Schaus, 1902)  Trinidad to Argentina; ref.13  M T

Lerema yahua (Evans, 1955)  TL: Peru  M T

Contrastia  Grishin, 2022  ref.13

Contrastia distigma (Plötz, 1882)  ref.13  iN  M T S

Vehilius  Godman, 1900  

Vehilius labdacus (Godman, 1900)  [formely a synonym of inca] N. & C. Amer.; ref.13  M T S L

Vehilius inca (Scudder, 1872)  Inca Skipper  [= xenos, Evans' "xenos" is sonex] C. Amer. to Argentina; ref.13  iN  M T S1 S2

Vehilius barrosi O. Mielke, 1968  TL: Brazil (MG)  M T

Vehilius stictomenes (A. Butler, 1877)    iN

Vehilius stictomenes illudens (Mabille, 1891)  E & W Mexico to Panama  M T S L1 L2

Vehilius stictomenes stictomenes (A. Butler, 1877)  TL: Brazil (Pa)  M T S1 S2

Vehilius stictomenes madra Evans, 1955  TL: Ecuador  M T S

Vehilius putus E. Bell, 1941  TL: Peru  M T

Vehilius vetula (Mabille, 1878)  Vetula Skipper  Nicaragua to Bolivia  iN  M T S HS L

Vehilius major de Jong, 1983  TL: Surinam  M T

Vehilius clavicula (Plötz, 1884)    iN  M T S

Vehilius spitzi (O. Mielke, 1967)  ref.140  M OI

Vehilius gorta Evans, 1955  TL: Brazil (Go)  M T

Vehilius tanta (Evans, 1955)  ref.140  M T

Vehilius campestris (O. Mielke, 1980)  ref.140  iN  M T

Vehilius warreni (Weeks, 1901)  Bolivia; ref.13  iN  M S

Vehilius jabre Medeiros, Souza & Kerpel, 2023  NE Brazil; ref.147

Vehilius tinta (Evans, 1955)  ref.140  M T

Vehilius limae (Lindsey, 1925)  Peru; ref.13  iN  M T S

Papias  Godman, 1900  [includes Mnasilus]; ref.13

Papias integra (Mabille, 1891)  S Mexico to Venezuela; ref.13  M T S1 S2 L

Papias subcostulata (Herrich-Schäffer, 1870)  Jungle Skipper  in S. Amer. to S Brazil & Paraguay; ref.13  iN  M S

Papias latonia (Schaus, 1913)  Latonia Skipper  Costa Rica and Panama; ref.13  M T S L

Papias microsema Godman, 1900  Mexico to Costa Rica; ref.16  M T S

Papias phainis Godman, 1900  Somber Skipper  SE Mexico to Peru and Uruguay  iN  M T S1 S2

Papias projectus E. Bell, 1942  TL: Ecuador  M T

Papias monus E. Bell, 1942  ref.16  M T

Papias unicolor (Hayward, 1938)  ref.16

Papias quigua Evans, 1955  TL: Venezuela  M T S

Papias allubita (A. Butler, 1877)  Butler's Skipper  E & W Mexico to S Brazil & Paraguay; ref.13  iN  M T S L

Papias amyrna (Mabille, 1891)  [= guianae] S. Amer.; ref.13,16  M T S

Veadda  Grishin, 2019  ref.5

Veadda veadeira (O. Mielke, 1968)  Brazil; ref.5  M OI OD G

Duroca  Grishin, 2019  ref.5

Duroca caraca (O. Mielke, 1992)  Brazil; ref.140  M T

Duroca duroca (Plötz, 1882)  S. Brazil  iN  M T S HS L

Duroca lenta (Evans, 1955)  S. Brazil; ref.140  M T

Saturnus  Evans, 1955  

Saturnus fartuga (Schaus, 1902)  S Brazil; ref.13  M T S

Saturnus jaguar (Steinhauser, 2008)  Steinhauser's Skipper  Guyana, Nicaragua; ref.122,16  M T

Saturnus metonidia (Schaus, 1902)  S Brazil  iN  M T S

Saturnus saturnus (Fabricius, 1787)    iN

Saturnus saturnus saturnus (Fabricius, 1787)  TL: French Guiana  M S

Saturnus saturnus servus Evans, 1955  TL: Brazil (Pr)  M T

Saturnus obscurus (E. Bell, 1941)  Bell's Skipper  E & S Mexico to Panama; ref.153  M T S L

Saturnus obscurior Grishin, 2023  Costa Rica & Panama; ref.153

Saturnus reticulata (Plötz, 1883)  Venezuela & Trinidad (at least)  iN  M T S

Saturnus tiberius (Möschler, 1883)  ref.153

Saturnus tiberius tiberius (Möschler, 1883)  TL: Surinam  M T S

Saturnus tiberius suffuscus (Hayward, 1940)  ref.153  M S

Saturnus meton (Mabille, 1891)  ref.153  M T S

Saturnus conspicuus (E. Bell, 1941)  ref.153  M T S L

Parphorus  Godman, 1900  

Parphorus kadeni (Evans, 1955)  tentatively in Parphorus, possibly from Venezuela  M T

Parphorus hermieri Grishin, 2022  Amazonian region; ref.16

Parphorus harpe (Steinhauser, 2008)  Peru; ref.122,16  iN

Parphorus decora (Herrich-Schäffer, 1869)  Decora Skipper  SE Mexico to S Brazil  iN  M T S1 S2 L

Parphorus felta Evans, 1955    iN  M T

Parphorus granta Evans, 1955  Granta Skipper  Panama  M T IL

Parphorus oca Evans, 1955  Oca Skipper  Panama to Peru  M T S

Parphorus prosper Evans, 1955  TL: Brazil (Pa)  M T

Parphorus pseudecorus (Hayward, 1934)    iN  M S

Parphorus storax (Mabille, 1891)  Storax Skipper    iN

Parphorus storax storax (Mabille, 1891)  SE Mexico to S Brazil  M T S L

Parphorus storax sorra Evans, 1955  TL: Ecuador  M T1 T2

Picova  Grishin, 2022  ref.13

Picova incompta (Hayward, 1942)  Ecuador; ref.13  M T

Picova steinbachi (E. Bell, 1930)  Bolivia; ref.13  M T S

Cantha  Evans, 1955  

Cantha matuta (Plötz, 1886)  TL: ?

Cantha calva Evans, 1955    iN  M T

Cantha zara (E. Bell, 1941)  TL: Bolivia  M T

Cantha zoirodicta Grishin, 2023  Amazonian Peru and Brazil; ref.153

Cantha meiodicta Grishin, 2023  Amazonian Peru and Brazil; ref.153

Cantha eteocla (Plötz, 1882)  Eteocla Skipper  Costa Rica to Argentina; ref.140  M T S

Cantha buriti (O. Mielke, 1968)  Brazil; ref.140  M T

Cantha roraimae (E. Bell, 1932)  Roraima Skipper  Chiapas and Belize to N Brazil  M T S

Cantha honor Evans, 1955    iN  M T

Cantha ivea Evans, 1955  TL: Brazil (SC)  M T S

Haza  Grishin, 2019  ref.5

Haza hazarma (Hewitson, 1877)  SE S. Amer.; ref.5  iN  M T S

Vistigma  Hayward, 1939  

Vistigma bryanti (Weeks, 1906)  Trinidad, Venezuela; ref.13  iN  M S

Vistigma subviridis (Plötz, 1886)  S Brazil; ref.13

Vistigma xanthobasis Hayward, 1939  Argentina; ref.13  iN  M S

Vistigma virgo (Evans, 1955)  Brazil; ref.13  M T

Vistigma vira (A. Butler, 1870)  Brazil; ref.13  M T S

Vistigma opus (Steinhauser, 2008)  Opus Skipper  Nicaragua, Guyana; ref.122,13  M T

Gemmia  Brockmann & Grishin, 2022  ref.41

Gemmia buechei Brockmann & Grishin, 2022  Peru; ref.41  M T

Phlebodes  Hübner, [1819]  

Phlebodes confixa (A. Butler, 1877)  TL: Brazil (Am)  M T S

Phlebodes meesi de Jong, 1983  TL: Surinam  M T S

Phlebodes notex Evans, 1955  TL: Brazil (Am)  M T

Phlebodes pertinax (Stoll, 1781)  TL: Surinam  M T S

Phlebodes sameda (Herrich-Schäffer, 1869)  TL: ?  M T S

Phlebodes sifax Evans, 1955  Sifax Skipper  Chiapas, Belize, Costa Rica to Guianas, Amazon Basin; ref.153  M T S L

Phlebodes duplex Grishin, 2023  Guatemala; ref.153

Phlebodes campo (E. Bell, 1947)  TL: Brazil (RJ)  M T S

Phlebodes fuldai (E. Bell, 1930)  ref.13  iN  M T S

Phlebodes yalta (Evans, 1955)  S Brazil; ref.154  M T S

Phlebodes torax Evans, 1955  TL: Brazil (RJ)  M T

Dubia  Grishin, 2019  ref.5

Dubia dubia (E. Bell, 1932)  Brazil; ref.5  iN  M T S

Mnasitheus  Godman, 1900  

Mnasitheus cephoides Hayward, 1943  Cephoides Skipper  Belize to S Brazil  M S IL G

Mnasitheus chrysophrys (Mabille, 1891)  Chrysophrys Skipper  SE Mexico to S Brazil & Paraguay  iN  M T OI S

Mnasitheus nella Evans, 1955    iN  M T S

Mnasitheus padus (Evans, 1955)  Peru; ref.13  iN  M T

Mnasitheus forma Evans, 1955  TL: Brazil (Pa)  M T

Mnasitheus nitra Evans, 1955  Nitra Skipper  W Mexico, Guatemala, Belize to S Brazil  iN  M T S

Mnasitheus submetallescens (Hayward, 1940)    iN  M S

Mnasitheus sucova (Schaus, 1902)  Sucova Skipper  [= Cynea fista], Chiapas to S Brazil; ref.13  iN  M T S

Mnasitheus strandi Biezanko & O. Mielke, 1973  TL: Brazil (RS)  M T

Naevolus  Hemming, 1939  

Naevolus orius (Mabille, 1883)  Orius Skipper  E & W Mexico to S Brazil  iN  M T S L

Naevolus naevus Evans, 1955  Ecuador; ref.13  M T

Naevolus brunnescens (Hayward, 1939)  Ecuador; ref.13  M S

Sodalia  Evans, 1955  

Sodalia argyrospila (Mabille, 1877)    iN  M T S L

Sodalia sodalis (A. Butler, 1877)  Sodalis Skipper  Nicaragua to S Brazil  M T S

Sodalia coler (Schaus, 1902)    iN  M T S

Sodalia petiti Gaviria-Ortiz, Dolibaina & A. Warren, 2020  Ecuador; ref.40

Metiscus  Godman, 1900  

Metiscus atheas Godman, 1900  [= Evans' "achelous", = matheri] Mexico to Colombia; ref.13  iN  M T S L

Metiscus goth Grishin, 2022  [= Evans' "infuscata"] Costa Rica; ref.13,16

Metiscus angularis (Möschler, 1877)  [= astur] C. & S. Amer.; ref.42,13  M T S

Metiscus ester (Lemes, O. Mielke & Casagrande, 2020)  Brazil; ref.42

Lychnuchus  Hübner, [1831]  [includes Enosis]; ref.13

Lychnuchus ponka (Evans, 1955)  Amazonian region; ref.153  M T

Lychnuchus blotta (Evans, 1955)  Trinidad; ref.13  M T

Lychnuchus demon (Evans, 1955)  Peru; ref.13  M T1 T2

Lychnuchus immaculata (Hewitson, 1868)  Immaculata Skipper  SE Mexico to Ecuador and Peru; ref.13  iN  M T S L

Lychnuchus aphilos (Herrich-Schäffer, 1869)  ref.13  iN  M T S

Lychnuchus valle Grishin, 2023  W Colombia; ref.153

Lychnuchus topo (Nicolay, 1980)  Ecuador; ref.13  M T

Lychnuchus blenda (Evans, 1955)  Brazil (Am)  M T

Lychnuchus dognini (Mabille, 1889)  Ecuador; ref.13  iN  M T S

Lychnuchus celsus (Fabricius, 1793)    iN  M S

Mit  Grishin, 2022  [includes Rotundia as subgenus]; ref.13

Mit gemignanii (Hayward, 1940)  Argentina; ref.13  M T S

Mit badius (E. Bell, 1930)  Bolivia; ref.13  M T S

Mit schausi (O. Mielke & Casagrande, 2002)  S Brazil; ref.13  iN  M T S

Eutychide  Godman, 1900  

Eutychide paria (Plötz, 1882)  Gold-dot Skipper  SE Mexico to Colombia and upper Amazon  M S

Eutychide ochus Godman, 1900  Ochus Skipper  SE Mexico to Costa Rica; ref.153  M T S

Eutychide ochoides Grishin, 2023  Ecuador & Peru; ref.153

Eutychide rogersi (Kaye, 1914)  [= Evans' "subcordata"] broadly in Amazonian region; ref.153  iN  M T

Eutychide subcordata (Herrich-Schäffer, 1869)  [= olympia] SE & S Brazil; ref.153  M S

Eutychide complana (Herrich-Schäffer, 1869)  Compliant Skipper  E Mexico to Bolivia and upper Amazon  iN  M T S L

Eutychide physcella (Hewitson, 1866)    iN  M T S

Dion  Godman, 1901  

Dion iccius (Evans, 1955)  Guyana; ref.13  M T

Dion gemmatus (A. Butler, 1872)  Gemmed Skipper  Costa Rica and Panama  iN  M T S L

Dion carmenta (Hewitson, 1870)    iN

Dion carmenta carmenta (Hewitson, 1870)  TL: Ecuador  M T L

Dion carmenta acraea (Plötz, 1882)  TL: Colombia  M T

Dion carmenta caucana Salazar & Vargas, [2017]  Colombia; ref.91

Dion carmenta coma Evans, 1955  TL: Ecuador

Dion meda Hewitson, 1877    iN  M T S

Dion bora Grishin, 2023  El Salvador to Panama; ref.153  M S

Dion occida Grishin, 2023  Venezuela, Peru, & Bolivia (at least); ref.153  M S L

Dion agassus (Mabille, 1891)  Amazonian; ref.13,20  M T S L

Dion uza (Hewitson, 1877)  [= pruinosa] SE & S Brazil; ref.13,20  iN  M T S

Venas  Evans, 1955  

Venas caerulans (Mabille, 1878)    iN  M T S

Venas evans (A. Butler, 1877)  TL: Brazil (Pa)  M T S

Eprius  Godman, 1901  [includes Repens as a subgenus]; ref.13

Eprius veleda (Godman, 1901)  Veleda Skipper  E Mexico to Panama  iN  M T S

Eprius veledinus Grishin, 2023  S America; ref.153

Eprius obrepta (Kivirikko, 1936)  Argentina & Brazil; ref.140  M T

Eprius repens (Evans, 1955)  S Brazil; ref.13  M T

Eprius penna (Evans, 1955)  Brazil; ref.13,140  M T

Eprius planus (Weeks, 1901)  Bolivia; ref.13,140  M T S

Peba  O. Mielke, 1968  

Peba verames (Schaus, 1902)    iN  M T

Radiatus  O. Mielke, 1968  

Radiatus panamensis Grishin, 2023  Panama; ref.153

Radiatus bradus O. Mielke, 1968    iN  M OI

Lattus  Grishin, 2022  ref.13

Lattus arabupuana (E. Bell, 1932)  Brazil; ref.13  M T

Lattus minor (O. Mielke, 1967)  Brazil; ref.140  M T OI

Tarmia  Lindsey, 1925  

Tarmia monastica Lindsey, 1925  TL: Peru  M S

Tarmia greeneyi A. Warren, Medeiros, Dolibaina & O. Mielke , 2019  Peru; ref.44

Panca  Evans, 1955  

Panca subpunctuli (Hayward, 1934)    iN  M S

Panca moseri Dolibaina, Carneiro & O. Mielke, 2017  Brazil; ref.48,45  iN

Panca acroleuca (Plötz, 1884)  Brazil; ref.45  M S

Panca mirnae (O. Mielke, Dolibaina, Carneiro & A. Warren, 2015)  Brazil; ref.47,45

Panca mictra (Evans, 1955)  Brazil; ref.45  M T

Panca paulo (E. Bell, 1932)  Brazil; ref.45  M T

Panca puri Medeiros, O. Mielke & Casagrande, 2020  Brazil; ref.45

Panca xavante Medeiros, O. Mielke & Dolibaina, 2020  Brazil; ref.45

Panca satyr (Evans, 1955)  Brazil; ref.45  iN  M T S

Panca tobiasi (O. Mielke, 1992)  Brazil; ref.45  M T

Panca trogon (Evans, 1955)  ref.45  iN  M T1 T2

Panca steinhauseri (Dolibaina & A. Warren, 2015)  ref.47,45  iN

Panca fiedleri (Carneiro, O. Mielke & Casagrande, 2015)  Konrad's Skipper  Brazil (MG, RJ); ref.51,140  iN

Ginungagapus  Carneiro, O. Mielke & Casagrande, 2015  ref.51

Ginungagapus schmithi (E. Bell, 1930)    iN  M T S

Ginungagapus awarreni Carneiro, O. Mielke & Casagrande, 2015  Andy's Skipper  Brazil (Ba, MG); ref.51

Ginungagapus bocus (E. Bell, 1947)  TL: Brazil (RJ)  M T S

Ginungagapus tangerinii Carneiro, O. Mielke & Casagrande, 2015  Tangerini's Skipper  Brazil (MG, RJ); ref.51

Ginungagapus ranesus (Schaus, 1902)    iN  M T S

Lucida  Evans, 1955  

Lucida oebasus (Godman, 1900)  Mottled Skipper  Costa Rica to Ecuador; ref.13  M T S L

Lucida melitaea (Draudt, 1923)  Colombia; ref.13  M T G

Lucida scopas (Mabille, 1891)  Venezuela; ref.13  M T S L

Lucida leopardus (Weeks, 1901)  Bolivia; ref.13  M S

Lucida lucia (Capronnier, 1874)  S Brazil; ref.13  iN  M T S

Eutocus  Godman, 1901  

Eutocus fosca (Evans, 1955)  Brazil; ref.140  M T

Eutocus rogan (Evans, 1955)  Brazil; ref.140  M T

Eutocus brasilia (Carneiro, O. Mielke & Casagrande, 2015)  Brasilia Skipper  Brazil (DF); ref.51,140

Eutocus minimus E. Bell, 1940  TL: Peru  M T

Eutocus facilis (Plötz, 1884)  Facilis Skipper  S Mexico to Ecuador and Bolivia  iN  M T S L

Eutocus quichua (Lindsey, 1925)    iN  M T S

Eutocus fabulinus (Plötz, 1884)  TL: Surinam  M S

Eutocus vetulus (Mabille, 1883)  Vetulus Skipper  Nicaragua to Argentina  iN

Eutocus vetulus vetulus (Mabille, 1883)  [= interpunctata Plötz, = vida Evans] Nicaragua to Bolivia & S Brazil; ref.139  M T1 T2 S L

Eutocus vetulus matildae (Hayward, 1941)  Argentina  iN  M S

Artines  Godman, 1901  

Artines bipunctata O. Mielke, 1968  TL: Brazil (DF)  M T

Artines aquilina (Plötz, 1882)    iN  M T S

Artines angelica Medeiros, 2019  ref.43

Artines focus Evans, 1955    iN  M T S

Artines cofus Medeiros & Dolibaina, 2019  ref.43

Artines donia Medeiros & Carneiro, 2019  ref.43

Artines maya A. Warren & Dolibaina, 2015  ref.47

Artines rica Steinhauser & Austin, 1993  Costa Rican Peacock-Skipper  Costa Rica  iN  M T S

Artines panama Medeiros & Carneiro, 2019  ref.43

Artines aepitus (Geyer, 1832)  Widespread Peacock-Skipper  Panama to S Brazil  iN  M T S L

Artines bamba Medeiros & O. Mielke, 2019  ref.43

Artines delfos Medeiros & Casagrande, 2019  ref.43

Artines litoralis Medeiros & Dolibaina, 2019  ref.43

Artines liege Medeiros, O. Mielke & Casagrande, 2019  ref.43  iN

Artines solange Medeiros & O. Mielke, 2019  ref.43

Phanes  Godman, 1901  

Phanes abaris (Mabille, 1891)    iN  M T S G

Phanes rezia (Plötz, 1882)    iN  M T S

Phanes aletes (Geyer, 1832)  Jeweled Skipper  E & W Mexico to S Brazil  iN  M S1 S2 L

Phanes almoda (Hewitson, 1866)  Almoda Skipper  SE Mexico to S Brazil  iN  M T S L

Phanes espa Evans, 1955  TL: Brazil (RJ)  M T

Pheraeus  Godman, 1900  

Pheraeus odilia (Plötz, 1884)    iN

Pheraeus odilia epidius (Mabille, 1891)  Nicaragua to Colombia  iN  M T S

Pheraeus odilia argus (Draudt, 1923)  TL: Bolivia  M T S G

Pheraeus odilia odilia (Plötz, 1884)  Brazil  iN  M S L

Pheraeus covadonga H. Freeman, 1969  'Eastern' Covadonga Skipper    iN

Pheraeus covadonga covadonga H. Freeman, 1969  E Mexico: Tamaulipas to Chiapas  M T S L

Pheraeus covadonga loxicha Steinhauser, 1991  SW Mexico: Oaxaca  M T S

Pheraeus fastus (Hayward, 1939)    iN  M S

Pheraeus honta Evans, 1955    iN  M T

Pheraeus pulcher Grishin, 2023  Upper Amazonian; ref.153

Pheraeus perpulcher (Hayward, 1934)    iN  M T S

Pheraeus jaruensis Steinhauser, 1991  TL: Brazil (Rd)  M T OI OD G

Pheraeus maria Steinhauser, 1991    iN  M T

Pheraeus unia (A. Butler, 1870)  Confusing Skipper  SE Brazil; probably mislabeled from Hispaniola  iN  M T OI S

Pheraeus argynnis (Plötz, 1884)    iN  M T S

Pheraeus guandu Saraiva, Carneiro, O. Mielke & Casagrande, 2020  ref.55

Pheraeus montes (E. Bell, 1947)    iN  M T S G

Pheraeus rumba Evans, 1955  TL: Brazil

Pheraeus rumba rumba Evans, 1955  TL: Brazil  M T

Pheraeus rumba kofan Steinhauser, 1991  TL: Colombia  M T G

Lamponia  Evans, 1955  

Lamponia elegantula (Herrich-Schäffer, 1869)    iN  M T S

Lamponia lamponia (Hewitson, 1876)    iN  M T S L

Lamponia travassosi (Hayward, 1942)    iN  M S

Lamponia ploetzii (Capronnier, 1874)  Brazil; ref.13  iN

Lamponia ploetzii morretesi (O. Mielke, 1992)  Brazil; ref.13  M L

Lamponia ploetzii ploetzii (Capronnier, 1874)  Brazil; ref.13  M T S L

Molla  Evans, 1955  

Molla molla Evans, 1955    iN  M T S L

Rufocumbre  Dolibaina, O. Mielke & Casagrande, 2017  ref.49

Rufocumbre celioi Dolibaina, O. Mielke & Casagrande, 2017  ref.49

Rufocumbre emeryi Dolibaina, O. Mielke & Casagrande, 2017  ref.49  iN

Rufocumbre eberti (Evans, 1955)    iN  M T

Rufocumbre schneideri Dolibaina, O. Mielke & Casagrande, 2017  ref.49

Rufocumbre lucasi Dolibaina, O. Mielke & Casagrande, 2017  ref.49

Cumbre  Evans, 1955  

Cumbre lamasi Dolibaina, O. Mielke & Casagrande, 2014  Ecuador & Peru; ref.46

Cumbre haywardi Dolibaina, O. Mielke & Casagrande, 2014  Peru to Argentina; ref.46

Cumbre cumbre (Schaus, 1902)    iN  M T S

Cumbre meridionalis (Hayward, 1934)    iN  M S

Tava  Grishin, 2019  ref.5

Tava tavola (Schaus, 1902)  ref.5  M T

Callimormus  Scudder, 1872  

Callimormus juventus Scudder, 1872  Juventus Skipper  S Mexico to Argentina  iN  M T S L

Callimormus diaeses Schaus, 1902  [= "interpunctata" of Evans] S Brazil, Paraguay & Argentina; ref.139  iN  M T

Callimormus alsimo (Möschler, 1883)  Alsimo Skipper  Panama to S Brazil  iN  M T S

Callimormus radiola (Mabille, 1878)  Radiant Skipper    iN

Callimormus radiola radiola (Mabille, 1878)  S Mexico to S Brazil  M S L

Callimormus radiola janna Evans, 1955  TL: Colombia  M T

Callimormus radiola elegans Hayward, 1938  TL: Ecuador  M S L

Callimormus radiola pusillus Hayward, 1934  TL: Argentina  M S

Callimormus corus E. Bell, 1941    iN  M T S

Callimormus simplicius Hayward, 1939    iN  M S

Callimormus saturnus (Herrich-Schäffer, 1869)  Saturnus Skipper  E & W Mexico to S Brazil & Paraguay  iN  M T S1 S2 L1 L2

Callimormus rades Grishin, 2023  Panama & Colombia; ref.153

Callimormus corades (C. Felder, 1862)    iN  M T S

Callimormus rivera (Plötz, 1882)    iN  M T S

Gubrus  Grishin, 2022  ref.13

Gubrus lubens Grishin, 2023  Ecuador; ref.153

Gubrus lugubris (Lindsey, 1925)  Peru; ref.13  iN  M S

Lento  Evans, 1955  

Lento xanthina (Mabille, 1891)  Xanthina Skipper  Costa Rica to Colombia and Venezuela  iN  M T S L

Lento longa Evans, 1955  TL: Peru  M T

Lento ferrago (Plötz, 1884)    iN  M T S

Lento lento (Mabille, 1878)    iN  M T S

Lento lucto Evans, 1955  TL: Peru  M T

Lento imerius (Plötz, 1884)  TL: Brazil  M S

Lento listo Evans, 1955  TL: ?  M T

Lento grosso Steinhauser, 1991  TL: Brazil (MT)  M T

Lento flavocostata (Plötz, 1884)  TL: Brazil (RJ)  M S

Lento lotus (E. Bell, 1947)  TL: Brazil (RJ)  M T

Lento apta Evans, 1955    iN  M T S

Lento lora Evans, 1955    iN  M T

Lento genta Evans, 1955  TL: Brazil (ES)  M T

Lento krexoides (Hayward, 1940)    iN  M S L

Lento ludo Evans, 1955    iN  M T

Lento pyra Evans, 1955  TL: Brazil (ES)  M T

Virga  Evans, 1955  

Virga riparia O. Mielke, 1969  TL: Brazil (SC)  M S

Virga hygrophila O. Mielke, 1969    iN  M S

Virga silvanus (Hayward, 1947)    iN  M T

Virga virginius (Möschler, 1883)  Virginius Skipper  Chiapas to upper Amazon  iN  M T S

Virga clenchi L. Miller, 1970  Clench's Skipper  E Mexico: TAMPS to VER & CAMP  iN  M S L

Virga xantho (Schaus, 1913)  Xantho Skipper  Nicaragua to Ecuador  iN  M T S

Virga austrinus (Hayward, 1934)    iN  M S L

Virga paraiba Nicolay, 1973  TL: Brazil (Pb)  M T

Mnestheus  Godman, 1901  

Mnestheus ittona (A. Butler, 1870)  Ittona Skipper  Costa Rica to Bolivia  iN  M T S L

Mnestheus damma Evans, 1955  TL: Ecuador  M T S

Mnestheus silvaticus Hayward, 1940  Argentina; ref.13  iN  M S

Ludens  Evans, 1955  [includes Levina]; Li19

Ludens labens Grishin, 2023  Panama; ref.153

Ludens ludens (Mabille, 1891)  Ludens Skipper  Costa Rica to Argentina  iN  M T S1 S2

Ludens petrovna (Schaus, 1902)    iN  M T

Ludens levina (Plötz, 1884)    iN  M T S

Rigga  Grishin, 2019  ref.5

Rigga hesia (Hewitson, 1870)  Ecuador; ref.5  M T S

Rigga oeagrus (Godman, 1900)  Tawny-washed Skipper  Costa Rica; ref.5  M T S L

Rigga ira (A. Butler, 1870)  Venezuela; ref.5  iN  M T S

Rigga isa Grishin, 2023  Ecuador; ref.153

Rigga auristriga (Draudt, 1923)  Peru & Bolivia; ref.5  M T S

Rigga sapala (Godman, 1900)  Sapala Skipper  Panama to Peru; ref.5  iN  M T OI S

Rigga paramus (E. Bell, 1947)  Colombia; ref.5  iN  M T S

Rigga spangla (Evans, 1955)  Ecuador; ref.13  iN  M T

Misius  Evans, 1955  

Misius misius (Mabille, 1891)  Misius Skipper  Peru, Brazil (Amazonas)  iN  M T S

  Tribe Hesperiini, Subtribe Anthoptina

Anthoptus  E. Bell, 1942  

Anthoptus insignis (Plötz, 1882)  Immaculate Skipper  E & W Mexico to S Brazil  iN  M S L

Anthoptus inculta (Dyar, 1918)  Inculta Skipper  [= macalpinei] S Mexico possibly to Costa Rica; ref.13  iN  M T S

Anthoptus maracanae (E. Bell, 1934)  Trinidadian Skipper  Panama, Colombia, Trinidad, Tobago, possibly Costa Rica  iN  M T S L

Anthoptus epictetus (Fabricius, 1793)  Trailside Skipper  SE Mexico to Argentina  iN  M T S1 S2 S3 IL L

Corticea  Evans, 1955  

Corticea schwarzi (E. Bell, 1941)  Colombia; ref.13  M T S

Corticea vicinus (Plötz, 1884)  Colombia; ref.13  M TI S

Corticea sylva (Hayward, 1942)  Ecuador; ref.13  M T S

Corticea mendica (Mabille, 1898)    iN  M T S

Corticea lysias (Plötz, 1883)  Lysias Skipper    iN

Corticea lysias lysias (Plötz, 1883)  SE Mexico to Panama  M T OI S

Corticea lysias coxeyi (Hayward, 1942)  TL: Ecuador  M T S

Corticea lysias pena Evans, 1955  TL: [Bolivia]  M T S

Corticea lysias potex Evans, 1955  TL: French Guiana  M T S

Corticea rivula (Mabille, 1891)  TL: Brazil (Am)  M T

Corticea graziellae E. Bell, 1959  TL: Brazil (Pb)  M T

Corticea corticea (Plötz, 1882)  Redundant Skipper  S TX (as stray), E & W Mexico to S Brazil & Paraguay  iN  M T S1 S2 L

Corticea noctis (Plötz, 1882)    iN  M T S

Corticea bella O. Mielke, 1969    iN  M T S

Corticea oblinita (Mabille, 1891)    iN  M T S

Corticea obscura O. Mielke, 1969    iN  M T S

Corticea aidae O. Mielke, 1969  TL: Brazil (RJ)  M T

Corticea immocerinus (Hayward, 1934)    iN  M S

Corticea diamantina O. Mielke, 1992  TL: Brazil (MG)  M T

Corticea similea E. Bell, 1942  Similar Skipper  SW Mexico (MICH, GRO)  M T S

Choranthus  Scudder, 1872  [includes Asbolis & Pyrrhocalles]; ref.13

Choranthus capucinus (Lucas, 1857)  Monk Skipper  Cuba, N Bahamas, S & C Florida; ref.13,20  iN  M T S L I

Choranthus orientis (Skinner, 1920)  Caribbean Skipper  Cuba; ref.13

Choranthus orientis eleutherae (M. Bates, 1934)  Eleuthera and Andros, Bahamas  M S

Choranthus orientis orientis (Skinner, 1920)  Cuba  M T S

Choranthus antiqua (Herrich-Schäffer, 1863)  Caribbean Skipper  Hispaniola; ref.13  iN  M T S1 S2

Choranthus jamaicensis (Schaus, 1902)  Jamaican Skipper  Jamaica; ref.13  iN  M T S1 S2

Choranthus lilliae E. Bell, 1931  Lilly's Skipper  Jamaica; ref.20  iN  M T S G

Choranthus vitellius (Fabricius, 1793)  Vitellius Skipper  Puerto Rico E to Virgin Islands  iN  M T S L

Choranthus haitensis Skinner, 1920  Haitian Skipper  Hispaniola  iN  M T S G

Choranthus schwartzi Gali, 1983  Schwartz's Skipper  Hispaniola  M T S

Choranthus maria Minno, 1990  Maria's Skipper  Hispaniola; status requires verification  M T

Choranthus melissa Gali, 1983  Melissa's Skipper  Hispaniola  M T

Choranthus richmondi L. Miller, 1965  Richmond's Skipper  Great Bahama Bank, N Bahamas  M T G

Choranthus radians (Lucas, 1857)  Radians Skipper  Cuba and Isla de Juventud, Cat I., Great Exuma I.  iN  M T S G

Choranthus borincona (F. Watson, 1937)  Puerto Rican Skipper  Puerto Rico; ref.20  iN  M T S

Racta  Evans, 1955  

Racta apella (Schaus, 1913)  Apella Skipper    iN

Racta apella apella (Schaus, 1913)  Costa Rica to Ecuador  M T S L

Racta apella raza Evans, 1955  Ecuador, Peru, Bolivia  M T

Racta plasma Evans, 1955  Plasma Skipper  Ecuador, Peru  iN  M T

Racta racta Evans, 1955  Racta Skipper  Ecuador, Peru, Bolivia  iN  M T L

Racta chiria Evans, 1955  Chiria Skipper  Peru  iN  M T

Racta dalla Evans, 1955  Dalla-like Skipper  Ecuador, Colombia  M T S

Igapophilus  O. Mielke, 1980  

Igapophilus rufus O. Mielke, 1980    iN  M T

  Tribe Hesperiini, Subtribe Falgina

Flaccilla  Godman, 1901  

Flaccilla aecas (Stoll, 1781)  Aecas Ruby-eye  E & W Mexico to S Brazil  iN  M T S L

Flaccilla lactea Grishin, 2023  Peru:Cuzco; ref.153

Turesis  Godman, 1901  

Turesis tabascoensis H. Freeman, 1979  Tabasco Skipper  SE Mexico  M T

Turesis complanula (Herrich-Schäffer, 1869)  Complanula Skipper  SE Mexico to S Brazil  iN  M T S HS L

Turesis theste Godman, 1901  Theste Skipper  SE Mexico to Costa Rica  M T S

Turesis basta Evans, 1955  TL: Brazil (Pa)  M T

Mnasinous  Godman, 1900  

Mnasinous cinnamomea (Herrich-Schäffer, 1869)  Brazil; ref.13,16  iN  M T

Mnasinous modestus (Godman, 1901)  [= Evans' "ina" from N. Am., true ina is conspecific wi ...; ref.13,Zhang22d  iN  M T S

Mnasinous phaeomelas (Hübner, [1831])  Hubner's Skipper  [= Evans' "ina" from C. & S. Am.] Panama to Brazil & Paraguay; ref.16  iN  M T S

Mnasinous purus (E. Bell, 1940)  Peru; ref.13,16  M T S

Mnasinous ina (Plötz, 1882)  Ina Skipper  [= typhon] SE Mexico to Costa Rica; ref.13,16  iN  M T S

Mnasinous patage Godman, 1900  Black-veined Skipper  SW & E Mexico to Costa Rica; ref.13,16  M T S L

Methion  Godman, 1900  

Methion melas Godman, 1900  Rusty Skipper  S Mexico (VER, CHIS), Guatemala  M T S L

Thargella  Godman, 1900  

Thargella volasus (Godman, 1901)  [= dolor] Belize, Guatemala to Guyana and Peru; ref.13  M T S

Thargella caura (Plötz, 1882)  Round-winged Skipper    iN

Thargella caura caura (Plötz, 1882)  SE Mexico to Bolivia and C Brazil  M T S

Thargella caura occulta (Schaus, 1902)  TL: Brazil (RJ)  M T S

Thargella evansi Biezanko & O. Mielke, 1973    iN  M S

Thargella tristissimus (Schaus, 1902)  Peru; ref.13  M T

Propapias  O. Mielke, 1992  

Propapias sipariana (Kaye, 1925)  Sipariana Skipper  Panama, Venezuela, Trinidad, Guyana, Surinam, Brazil, Ecuador, Argentina  iN  M T1 T2 S

Synapte  Mabille, 1904  

Synapte silius (Latreille, [1824])  Rain-forest Faceted-Skipper  SE Mexico to Argentina  iN  M T OI S1 S2 S3 L

Synapte malitiosa (Herrich-Schäffer, 1865)  Caribbean Faceted-Skipper    iN

Synapte malitiosa malitiosa (Herrich-Schäffer, 1865)  Cuba, Jamaica  M T S

Synapte malitiosa adoceta A. Schwartz & Sommer, 1986  Hispaniola  M T S

Synapte malitiosa antistia (Plötz, 1882)  TL: Brazil (RJ)  M T S

Synapte malitiosa equa Evans, 1955  TL: Bolivia  M T

Synapte malitiosa pericles (Möschler, 1879)  TL: Colombia  M T S

Synapte pecta Evans, 1955  Northern Faceted-Skipper  S TX, E & W Mexico to Costa Rica  iN  M T1 T2 S L

Synapte puma Evans, 1955  Panamanian Faceted-Skipper  Panama  iN  M T S L

Synapte silna Evans, 1955  Southwest-Mexican Faceted-Skipper  SW Mexico (NAY to OAX)  iN  M T S L

Synapte salenus (Mabille, 1883)  Salenus Faceted-Skipper    iN

Synapte salenus salenus (Mabille, 1883)  S TX (rare stray), E Mexico to Colombia  M T S1 S2 L

Synapte salenus osta Evans, 1955  TL: Bolivia  M T

Synapte lutulenta (Herrich-Schäffer, 1869)  Whitened Faceted-Skipper  Costa Rica to Ecuador  iN  M T S L

Synapte shiva Evans, 1955  Faded Faceted-Skipper  W Mexico to Nicaragua  iN  M T1 T2 S L

Synapte syraces (Godman, 1901)  Bold Faceted-Skipper  W Mexico, El Salvador  iN  M T S L I

Koria  Grishin, 2022  

Koria kora (Hewitson, 1877)  S Brazil; ref.13  iN  M T S

Justinia  Evans, 1955  

Justinia norda Evans, 1955  False Saliana  ["phaetusa"], SE Mexico to Colombia  iN  M T S L

Justinia phaetusa (Hewitson, 1866)    iN  M T S

Justinia gava Evans, 1955  TL: Guyana  M T

Justinia justinianus (Latreille, [1824])  Justinianus Skipper  Nicaragua [as J. j. justinianus, but may be J. j. dappa Evans, 1955]  iN

Justinia justinianus cappa Evans, 1955  TL: Ecuador  M T

Justinia justinianus hyperythrus (Kaye, 1914)  TL: Trinidad  M T S

Justinia justinianus dappa Evans, 1955  TL: Brazil (Pa)  M T1 T2 T3

Justinia justinianus justinianus (Latreille, [1824])  TL: Brazil  M T S L

Justinia papaea (Hewitson, 1876)    iN  M T S

Justinia maculata (E. Bell, 1930)  TL: Bolivia  M T S

Justinia septa Evans, 1955  TL: Brazil (MG)  M T

Mnasalcas  Godman, 1900  

Mnasalcas continua (Evans, 1955)  TL: Bolivia

Mnasalcas continua continua (Evans, 1955)  Bolivia; ref.13  M T S

Mnasalcas continua conta (Evans, 1955)  TL: Ecuador  M T

Mnasalcas thymoetes (Hayward, 1942)  Ecuador; ref.13  M T

Mnasalcas boyaca (Nicolay, 1973)  Colombia; ref.13  M T

Mnasalcas simplicissima (Herrich-Schäffer, 1870)  Strange-stigma Skipper  Costa Rica to Bolivia; ref.13  iN  M T S

Mnasalcas ritans (Schaus, 1902)  S Brazil; ref.13  iN  M T S

Barrolla  Grishin, 2019  

Barrolla barroni (Evans, 1955)  Barron's Skipper  [= molla] Ecuador; ref.5  M T

Falga  Mabille, 1898  

Falga sciras Godman, 1901  Sciras Skipper  Chiapas to Costa Rica  iN  M S

Falga athena Grishin, 2023  Panama; ref.153

Falga jeconia (A. Butler, 1870)  Venezuela  iN  M T S

Falga jacta Evans, 1955  Colombia; ref.153  M T

Falga farina Evans, 1955  TL: Bolivia  M T

Falga mirabilis Evans, 1955  Peru & Bolivia; ref.153

Falga mirabilis odol Evans, 1955  Peru; ref.153  M T

Falga mirabilis mirabilis Evans, 1955  Peru & Bolivia; ref.153  M T

Falga ombra Evans, 1955  Ecuador; ref.153  M T L

Falga theoclea (Hewitson, 1870)    iN  M T S

  Tribe Hesperiini, Subtribe Apaustina

Apaustus  Hübner, [1819]  

Apaustus gracilis (C. Felder & R. Felder, 1867)  Delicate Skipper    iN

Apaustus gracilis gracilis (C. Felder & R. Felder, 1867)  Nicaragua to Ecuador  M T S L

Apaustus gracilis smarti Evans, 1955  TL: Guyana  M T S

Apaustus menes (Stoll, 1782)  Menes Skipper  Nicaragua to upper Amazon  iN  M S

Apaustus radiatus Hayward, 1940    iN  M T

Ancyloxypha  C. Felder, 1862  

Ancyloxypha melanoneura C. Felder & R. Felder, 1867    iN  M T

Ancyloxypha nitedula (Burmeister, 1878)    iN  M S

Ancyloxypha aurea Hayward, 1940    iN  M T

Ancyloxypha dryas Hayward, 1942    iN  M S

Ancyloxypha ramba Evans, 1955  TL: Ecuador  M T

Ancyloxypha arene (W. H. Edwards, 1871)  Tropical Least Skipper  SE AZ, SW NM, W & S TX, S to Costa Rica  iN  M T S L I H G

Ancyloxypha numitor (Fabricius, 1793)  Least Skipper  Great Plains, eastward; SE SASK to Quebec, to S TX, far NE MEX  iN  M T S L1 L2 L3 L4 L5 L6 Ha G

Adopaeoides  Godman, 1900  

Adopaeoides prittwitzi (Plötz, 1884)  Sunrise Skipper  SE AZ, SW NM, W TX, S to S-central Mexico  iN  M S L I Ha

Adopaeoides bistriata Godman, 1900  TL: Mexico (Dur)  M T

  Tribe Hesperiini, Subtribe Thymelicina

Thymelicus  Hübner, [1819]  

Thymelicus lineola (Ochsenheimer, 1808)  European Skipper    iN

Thymelicus lineola lineola (Ochsenheimer, 1808)  introduced; Newfoundland to VA & TN, W to MAN, S Rocky Mts.  M S L1 L2 I Ha G

Oarisma  Scudder, 1872  [includes Copaeodes]; ref.10

Oarisma poweshiek (Parker, 1870)  Poweshiek Skipperling  SE MAN, E ND, NE SD, W & S MN, N-central IA, SE WI, S MI  iN  M S G

Oarisma garita (Reakirt, 1866)  Garita Skipperling    iN

Oarisma garita garita (Reakirt, 1866)  SE BC to S MAN (plus Peace Riv.), S through Rockies to AZ & NM  M T S L1 L2 L3 Ha G

Oarisma garita calega (Godman, 1900)  NW (DGO) to S (OAX) Mexico  M T S

Oarisma era Dyar, 1927  Bold-veined Skipperling  NW & NE to central Mexico  iN  M T S Ha

Oarisma nanus (Herrich-Schäffer, 1865)  Nanus Skipperling  Cuba & Isla de Juventud  M T S L

Oarisma stillmani (E. Bell & W. Comstock, 1948)  Stillman's Skipperling  Hispaniola including Isla Catalina; ref.10  M T S

Oarisma eoa (D. Smith, L. Miller & McKenzie, 1991)  Anegada Skipperling  Anegada; ref.10  M OI

Oarisma bruneri E. Bell, 1959  Bruner's Skipperling  Cuba  M T

Oarisma edwardsii (W. Barnes, 1897)  Edwards’ Skipperling  S-central CO, NM, C & E AZ, NW Mexico to S-central Mexico; ref.10  iN  M T S L I G

Oarisma aurantiaca (Hewitson, 1868)  Orange Skipperling  S CA & NV to E TX (strays N), S to Guatemala; ref.10  iN  M T S L1 L2 L3 I F G

Oarisma minima (W. H. Edwards, 1870)  Southern Skipperling  SE AZ, C TX to FL & NC, E & W Mexico to Costa Rica; ref.10  iN  M T S L1 L2 L3 L4 I1 I2 G

Oarisma jean (Evans, 1955)  ref.10  iN  M T S

Oarisma favor (Evans, 1955)  ref.140  M T S

Oarisma boeta (Hewitson, 1870)    iN  M T OI L

Oarisma castanea (O. Mielke, 1969)  Brazil; ref.10  iN  M T S

  Tribe Hesperiini, Subtribe Calpodina

Panoquina  Hemming, 1934  

Panoquina errans (Skinner, 1892)  Wandering Skipper  S CA, W Mexico  iN  M T S L I Ha G

Panoquina panoquinoides (Skinner, 1891)  Obscure Skipper  S FL to S TX, Mexican Gulf Coast to Costa Rica, most West Indies  iN  M T S Ma L1 L2 G

Panoquina eugeon (Godman & Salvin, 1896)  Eugeon Skipper  Grenada, Grenadines; ref.140

Panoquina eugeon eugeon (Godman & Salvin, 1896)  Grenada, Grenadines; ref.140  M T S

Panoquina eugeon minima de Jong, 1983  The Guianas; ref.140  M T1 T2

Panoquina calna Evans, 1955  Peru; ref.140

Panoquina albistriga O. Mielke, 1980  S Brazil; ref.140  M T

Panoquina panoquin (Scudder, 1863)  Salt Marsh Skipper  coastal salt marshes, New England to S TX  iN  M T S L1 L2 Ha G

Panoquina chapada Evans, 1955  TL: Brazil (MT)

Panoquina hecebolus (Scudder, 1872)  Hecebolus Skipper  [= ortygia] S TX, E & S Mexico to Paraguay; ref.16  iN  M T S L

Panoquina ocola (W. H. Edwards, 1863)  Ocola Skipper  Greater Antilles, Texas to Argentina  iN  M T S L1 L2 L3 G

Panoquina lucas (Fabricius, 1793)  Purple-washed Skipper  [= sylvicola, = woodruffi], Gr. & Less. Antilles, S TX to Argentina  iN

Panoquina lucas lucas (Fabricius, 1793)  [= sylvicola, = woodruffi], Gr. & Less. Antilles, S TX to Argentina  M T S L I G Txt

Panoquina lucas distipuncta K. Johnson & Matusik, 1988  [not ocola] SW Dominican Republic; ref.16  M T

Panoquina trix Evans, 1955  [= confusa] Brazil; ref.140  M T

Panoquina bola E. Bell, 1942    iN  M T L

Panoquina peraea (Hewitson, 1866)  TL: Brazil (RJ)  M T

Panoquina corrupta (Herrich-Schäffer, 1865)  Corrupt Skipper  Cuba and Isla de Juventud  iN  M T S

Panoquina nero (Fabricius, 1798)  Nero Skipper  Hispaniola and Puerto Rico  iN  M T S L

Panoquina jay Grishin, 2023  NE Peru; ref.153

Panoquina evansi (H. Freeman, 1946)  Evans’ Skipper  S TX, E & W Mexico to W Panama  iN  M T S L1 L2

Panoquina fusina (Hewitson, 1868)    iN

Panoquina fusina sonta Evans, 1955  Panama & Colombia  M S

Panoquina fusina fusina (Hewitson, 1868)  TL: Brazil (Pa)  M T

Panoquina fusina viola Evans, 1955  [Honduras?, Colombia?], Bolivia, SE Brazil, Paraguay  M S

Panoquina fusina jumbo Evans, 1955  TL: Jamaica

Panoquina pauper (Mabille, 1878)  Pauper Skipper    iN

Panoquina pauper pauper (Mabille, 1878)  SE Mexico to Colombia  M T S L

Panoquina pauper grapte (H. Druce, 1908)  TL: Peru  M T

Panoquina danishi R. Anderson, 2014  Danish's skipper  Panama; ref.57

Panoquina neblina Vargas & Salazar, 2013  Hazy skipper  Choco region in Colombia; ref.58  iN

Panoquina luctuosa (Herrich-Schäffer, 1869)    iN

Panoquina luctuosa luctuosa (Herrich-Schäffer, 1869)  TL: ?  M T G

Panoquina luctuosa fonda Evans, 1955  TL: Ecuador  M T G

Panoquina luctuosa cineas Evans, 1955  TL: Peru  M T G

Panoquina evadnes (Stoll, 1781)  Evadnes Skipper  S Mexico to Argentina  iN  M S L

Zenis  Godman, 1900  

Zenis par Grishin, 2022  Colombia to Peru; ref.16

Zenis hemizona (Dyar, 1918)  Purple-stained Skipper  ref.16  M T S L

Zenis janka Evans, 1955  ref.16

Zenis jebus (Plötz, 1882)    iN

Zenis jebus jebus (Plötz, 1882)  TL: Brazil

Zenis jebus beckeri O. Mielke & Casagrande, 2002  TL: Brazil (ES)  M T

Zenis minos (Latreille, [1824])  Minos Skipper  SE Mexico, Honduras, Costa Rica, S Brazil  iN  M T S

Calpodes  Hübner, [1819]  

Calpodes ethlius (Stoll, 1782)  Brazilian Skipper  SE US (strays far N), E & W MX to Argentina, W. Indies, Galapagos; ref.10  iN  M S L1 L2 I G

Calpodes fusta (Evans, 1955)  Suffused Saliana  E & W Mexico to Brazil & Bolivia  iN  M T1 T2 S L

Calpodes salva (Evans, 1955)  TL: Brazil (Pa)  M T

Calpodes triangularis (Kaye, 1914)  Triangular Saliana  E & S Mexico to Argentina  iN  M T S1 S2 L

Calpodes fischer (Latreille, [1824])  TL: Brazil  M T S

Calpodes nigel (Evans, 1955)    iN  M T1 T2 S

Calpodes hewitsoni (N. Riley, 1926)  Green Saliana  SE Mexico to Bolivia  iN  M T S1 S2

Calpodes chiomara (Hewitson, 1867)  Chiomara Saliana  Panama to Peru & Brazil  iN  M T S

Calpodes longirostris (Sepp, [1840])  Shy Saliana  E & W Mexico to S Brazil  iN  M S L

Calpodes morsa (Evans, 1955)  Morsa Saliana  Taboga Is., Panama, to Guianas & Amazon basin  iN  M T1 T2 S

Calpodes salius (Cramer, 1775)  Sullied Saliana  S Mexico to S Brazil & Uruguay  iN  M S1 S2 L

Calpodes salianus Grishin, 2023  Amazonian Peru; ref.153

Calpodes saladin (Evans, 1955)  Violet-tipped Saliana    iN

Calpodes saladin saladin (Evans, 1955)  S Mexico to Colombia & Peru  M T S

Calpodes saladin catha (Evans, 1955)  TL: Brazil (SC)  M T S

Calpodes saladin culta (Evans, 1955)  TL: Brazil (Pa)  M T S

Calpodes severus (Mabille, 1895)  Dark Saliana  S Mexico to Colombia & Ecuador  iN  M S IL L

Calpodes chocoensis (Salazar & Constantino, 2013)  Choco Megaleas Skipper  [= angelae] Choco region in Colombia; ref.90,16  iN  M T

Calpodes antoninus (Latreille, [1824])  Persistent Saliana  S Mexico to S Brazil  iN  M S L

Calpodes covancae (O. Mielke, 1970)  TL: Brazil (RJ)  M T

Calpodes salona (Evans, 1955)    iN  M T

Calpodes esperi (Evans, 1955)  Perching Saliana    iN

Calpodes esperi esperi (Evans, 1955)  E & W Mexico to S Brazil and Bolivia  M T S1 S2 L

Calpodes esperi soroa (D. Smith & L. Hernández, 1992)  W Cuba  M S L

Calpodes placens (A. Butler, 1874)  Ochreous Saliana  Costa Rica to Colombia & Ecuador  iN  M T S

Calpodes stingo Grishin, 2023  Ecuador; ref.153

Calpodes mamurra (Plötz, 1886)    iN  M T S

Calpodes mathiolus (Herrich-Schäffer, 1869)    iN  M T S

  Tribe Hesperiini, Subtribe Carystina

Corta  Grishin, 2022  

Corta lycortas (Godman, 1900)  Lycortas Skipper  E Mexico, Belize; ref.13  iN  M T S L

Orthos  Evans, 1955  

Orthos orthos (Godman, 1900)  Orthos Skipper  Panama, Trinidad, Peru  iN  M T S

Orthos minka Evans, 1955  Amazonian Brazil; ref.140  M T

Orthos hyalinus (E. Bell, 1930)  SE Brazil, Argentina; ref.140  M T S

Turmosa  Grishin, 2019  

Turmosa camposa (Plötz, 1886)  Brazil; ref.5  iN  M S

Dubiella  Evans, 1936  

Dubiella dubius (Stoll, 1781)  [= disjuncta, = cinica]; ref.13  iN  M T S

Dubiella fiscella (Hewitson, 1877)    iN

Dubiella fiscella fiscella (Hewitson, 1877)  TL: Brazil (Pa)  M T S L

Dubiella fiscella belpa Evans, 1955  S Mexico to Ecuador, Venezuela, Guianas  iN  M T S L

Carystina  Evans, 1955  

Carystina aurifer (Godman & Salvin, 1879)  Golden-spotted Ruby-eye  Costa Rica to W Ecuador  iN  M T S

Carystina mielkei Salazar, Constantino & Vargas, 2013  Mielke's Ruby-eye  W Colombia; ref.58  iN

Carystina pieris (Draudt, 1923)  Peru  iN  M T S G

Carystina lysiteles (Mabille, 1891)  Ecuador & Peru  iN  M T S L

Carystina discors (Plötz, 1882)  ref.13  iN  M TI

Aides  Billberg, 1820  

Aides duma Evans, 1955    iN

Aides duma argyrina Cowan, 1970  [= epitus], Panama to S Brazil  M S1 S2 L

Aides duma duma Evans, 1955  TL: [Argentina]  M T

Aides aestria (Hewitson, 1866)  TL: Brazil (RJ)  M T S

Aides dysoni Godman, 1900  Dyson's Silverpatch  E Mexico to Colombia  iN  M T OI S L

Aides aegita (Hewitson, 1866)  Aegita Silverpatch  [= ocrinus] Panama to Bolivia & Brazil; ref.153  iN  M T S1 S2 S3 L

Aides nobra Grishin, 2023  [= Evans' "ocrinus"] Panama, Colombia, French Guiana; ref.153  M S

Aides brilla (H. Freeman, 1970)  Brilliant Silverpatch  SE Mexico, Guatemala, Belize  iN  M T S L

Aides brino (Stoll, 1781)  Brino Silverpatch  Honduras, Colombia to Guianas & N Brazil  iN  M S

Thracides  Hübner, [1819]  

Thracides joannisii Mabille, 1904  Joannis' Skipper  Panama to Ecuador  M T TI S G

Thracides cilissa (Hewitson, 1867)  Cilissa Skipper  SE Colombia, NE Peru, N Brazil  iN  M T S

Thracides sacrator (Godman & Salvin, 1879)  Colombia; ref.13  iN  M T L G

Thracides polites (Godman & Salvin, 1879)  ref.13  iN

Thracides polites polites (Godman & Salvin, 1879)  Colombia; ref.13  M T

Thracides polites pilla (Evans, 1955)  TL: Peru  M S L

Thracides arcalaus (Stoll, 1782)  Yellow-spotted Ruby-eye  S Mexico to Guianas & Amazon basin  iN  M T S L

Thracides pavo Grishin, 2023  E & W Mexico to Ecuador; ref.153  M S L

Thracides phidon (Cramer, 1779)  Jewel-studded Skipper  Venezuela to S Brazil  iN  M S L

Thracides nanea (Hewitson, 1867)  Nanea Skipper  NE Peru, N Brazil  iN  M T S

Thracides chiricana Röber, 1926  Chiricana Skipper  Costa Rica, Panama  iN  M T S Txt

Thracides nida Evans, 1955  Nida Skipper  Colombia  iN  M OD S L Txt

Thracides panimeron H. Druce, 1908  Panimeron Skipper  Peru, Bolivia  iN  M T S

Thracides thrasea (Hewitson, 1866)  Thrasea Skipper  S Mexico (OAX), Belize, Guianas, Amazon basin to Bolivia  iN  M T OI S I

Thracides cleanthes (Latreille, [1824])  Cleanthes Skipper    iN

Thracides cleanthes binota Evans, 1955  Colombia to Guianas and Trinidad  M S

Thracides cleanthes telmela (Hewitson, 1866)  French Guiana, Ecuador, Peru, N Brazil  M T S

Thracides cleanthes quarta Evans, 1955  Peru (La Merced)

Thracides cleanthes trebla Evans, 1955  Bolivia  M S

Thracides cleanthes cleanthes (Latreille, [1824])  C and S Brazil, Paraguay  M T S

Thracides cleanthes quinta Evans, 1955  E Brazil  M S

Vertica  Evans, 1955  

Vertica umber (Herrich-Schäffer, 1869)    iN  M T

Vertica ibis Evans, 1955  Ibis Skipper  SE Mexico, Bolivia  M T S

Vertica verticalis (Plötz, 1882)    iN

Vertica verticalis verticalis (Plötz, 1882)  TL: Brazil  M T

Vertica verticalis coatepeca (Schaus, 1902)  SE Mexico to Costa Rica  M T S L

Vertica verticalis grandipuncta (Mabille, 1883)  TL: Brazil (Pa)  M T

Vertica subrufescens (Schaus, 1913)  Subrufescens Skipper  SE Mexico to Costa Rica  M S L I

Vertica pudor Evans, 1955  TL: Brazil (SC)  M T S

Vertica brasta (Evans, 1955)  Peru; ref.13  iN

Calvetta  Grishin, 2019  

Calvetta calvina (Hewitson, 1866)  White-rayed Ruby-Eye  ref.13  iN  M T TI S IL L

Cobalus  Hübner, [1819]  

Cobalus virbius (Cramer, 1777)  White-edged Ruby-eye    iN

Cobalus virbius quadrum (Mabille, 1891)  TL: Colombia  M T

Cobalus virbius virbius (Cramer, 1777)  Costa Rica to S Brazil & Paraguay  M T S L

Cobalus virbius hanta Evans, 1955  TL: Brazil (Pe)  M T

Cobalus virbius hersilia (Plötz, 1882)  TL: Brazil (RJ)

Cobalus fidicula (Hewitson, 1877)  White-centered Ruby-eye  S Mexico to Panama  iN  M T S L

Carystus  Hübner, [1819]  [includes Moeros, Argon & Synale]; ref.13

Carystus moeros (Möschler, 1877)  ref.13  iN  M T S

Carystus argus Möschler, 1879  W & S Mexico to Colombia; ref.13,16  M T S L

Carystus lota (Hewitson, 1877)  Argon Skipper  in S. Amer. to Argentina; ref.13,16  iN  M T L

Carystus cynaxa (Hewitson, 1867)  Black-veined Ruby-eye  E & S Mexico to Costa Rica; ref.13  iN  M T S L Ha

Carystus phorcus (Cramer, 1777)  White-patched Ruby-eye    iN

Carystus phorcus phorcus (Cramer, 1777)  E & W Mexico to Argentina; ref.13  M T S1 S2 S3 L

Carystus phorcus claudianus (Latreille, [1824])  TL: Brazil  M T S

Carystus diores (Plötz, 1882)  TL: S. Amer.  M S

Carystus junior Evans, 1955  TL: Brazil (Am)  M T

Carystus ploetzi O. Mielke & Casagrande, 2002  Ploetz's Ruby-eye  E & S Mexico to central Brazil & Peru; ref.13  iN  M T OI S L

Carystus hylaspes (Stoll, 1781)    iN  M T S

Carystus sylvanus (Hayward, 1948)  Argentina; ref.13

Carystus elana (Plötz, 1882)  ref.13  iN

Carystus elana elana (Plötz, 1882)  TL: Brazil

Carystus elana apheka (Kaye, 1918)  TL: Uruguay

Carystus metella (Plötz, 1882)  Brazil; ref.13

Carystus elvira (Plötz, 1882)  TL: S. Amer.

Carystus elvira elvira (Plötz, 1882)  TL: S. Amer.  M TI

Carystus elvira rufoventris Austin & O. Mielke, 2007  ref.140  iN  M T

Carystus superbiens Mabille, 1891    iN  M T S

Carystus cacaulandia Austin & O. Mielke, 2007  TL: Brazil (Rd)  M T

Carystus hocus Evans, 1955  TL: Guyana  M T S

Carystus jolus (Stoll, 1782)    iN  M T S

Carystus periphas Mabille, 1891    iN

Carystus periphas periphas Mabille, 1891  TL: Brazil (Am)  M T S L

Carystus periphas amax Evans, 1955  TL: Colombia  M T

Aroma  Evans, 1955  

Aroma aroma (Hewitson, 1867)  Aroma Skipper  Nicaragua to N Brazil & Peru  iN  M T S L

Aroma henricus (Staudinger, 1876)  Henricus Skipper    iN

Aroma henricus henricus (Staudinger, 1876)  Costa Rica to Peru  M T S L

Aroma henricus balba Evans, 1955  TL: Ecuador

Molo  Godman, 1900  

Molo pelta (Evans, 1955)  Ecuador; ref.5  M S

Molo mango (Guenée, 1865)  Mango Skipper  Costa Rica to C Brazil  iN  M T S1 S2 S3 S4 L

Molo muzo Evans, 1955  Muzo Skipper  Costa Rica to Colombia  M T IL

Tellona  Evans, 1955  

Tellona variegata (Hewitson, 1870)  TL: Ecuador  M T S

Talides  Hübner, [1819]  

Talides cantra Evans, 1955  Cantra Ruby-eye  S Mexico to Colombia, Venezuela & W Ecuador  iN  M S L

Talides sinois Hübner, [1819]    iN  M T

Talides riosa Evans, 1955    iN  M S

Talides laeta Grishin, 2023  Peru:Cuzco; ref.153

Talides alternata E. Bell, 1941  Alternate Ruby-eye  S Mexico to S Brazil  iN  M T S L

Talides hispa Evans, 1955  Hispa Skipper  Costa Rica to W Ecuador, W Brazil  M S IL

Talides eluta Grishin, 2023  Peru & Brazil; ref.153

Talides sergestus (Cramer, 1775)  Sergestus Ruby-eye  SE Mexico to S Brazil  iN  M S L

Talides montezuma Geale & Salazar, [2017]  Colombia; ref.91

Ebusus  Evans, 1955  

Ebusus ebusus (Cramer, 1780)    iN

Ebusus ebusus nigrior L. Miller, 1985  SE Mexico (Veracruz and Chiapas)  M T

Ebusus ebusus ebusus (Cramer, 1780)  TL: Surinam  M T

Zetka  Grishin, 2019  

Zetka zeteki (E. Bell, 1931)  Zetek’s Skipper  Guatemala, Panama; ref.5  M T

Zetka irena (Evans, 1955)  Ecuador; ref.13

Coolus  Grishin, 2019  

Coolus bushi (F. Watson, 1937)  Bush's Skipper  Hispaniola; ref.5  iN  M T

Mielkeus  Grishin, 2019  

Mielkeus tertianus (Herrich-Schäffer, 1869)  Blurry Skipper  SE Mexico to French Guiana and Trinidad; ref.5  iN  M T S

Mielkeus klugi (E. Bell, 1941)  Peru; ref.5  M T S

Mielkeus lucretius (Latreille, [1824])  Brazil; ref.5  iN  M S L

Mielkeus diana (Plötz, 1886)  ref.5  iN

Mielkeus diana felix (Evans, 1955)  TL: Peru  M T

Mielkeus diana diana (Plötz, 1886)  TL: Brazil (SP)  M T S G

Neoxeniades  Hayward, 1938  

Neoxeniades gabina (Godman, 1900)  Gabina Skipper  E Mexico to Costa Rica; ref.13  M T OI

Neoxeniades ethoda (Hewitson, 1866)  Ethoda Skipper  SE Brazil (Espirito Santo to RJ); ref.5  iN  M T S

Neoxeniades angustior Grishin, 2023  SE Brazil (RJ); ref.153

Neoxeniades myra Evans, 1955  TL: Colombia  M S

Neoxeniades parna (Evans, 1955)  [= vixen]; ref.13  M T S

Neoxeniades scipio (Fabricius, 1793)    iN

Neoxeniades scipio posta Evans, 1955  TL: Peru  M S

Neoxeniades scipio fulguratoides (Kaye, 1925)  TL: Trinidad  M T S

Neoxeniades scipio nosa Evans, 1955  TL: Brazil (Ba)  M S

Neoxeniades scipio scipio (Fabricius, 1793)  TL: Indiis  M T S

Neoxeniades tropa Evans, 1955  TL: Colombia

Neoxeniades luda (Hewitson, 1877)  Luda Skipper  W & S Mexico to Colombia  iN  M T S L

Neoxeniades pluviasilva Burns, 2007  Pluviasilva Skipper  Nicaragua to Costa Rica  iN  M T S L

Neoxeniades anchicayensis Steinhauser, 2007  Anchicaya Skipper  TL: Colombia  M S

Neoxeniades turmada (H. Druce, 1912)    iN  M T S

Neoxeniades musarion Hayward, 1938  TL: Brazil  M S

Neoxeniades cincia (Hewitson, 1867)    iN  M T S

Daron  Grishin, 2019  

Daron seron (Godman, 1901)  Seron Skipper  ref.5

Daron seron seron (Godman, 1901)  Honduras, Colombia  M T OI S

Daron seron sexton (Evans, 1955)  TL: Brazil (Pa)

Damas  Godman, 1901  

Damas cervelina (Orellana & Costa, 2019)  Venezuela; ref.90

Damas zea Grishin, 2023  Guyana; ref.153

Damas horridus (E. Bell, 1940)  TL: Peru  M T OD G Txt

Damas immacula Nicolay, 1973  Immaculate Ruby-eye  Panama  iN  M T1 T2 OI OD S G

Damas clavus (Herrich-Schäffer, 1869)  Violet-washed Skipper  E & S Mexico to S Brazil & Bolivia  iN  M T S L G

Tromba  Evans, 1955  

Tromba tromba Evans, 1955  TL: Peru  M S

Tromba xantha Grishin, 2023  Mexico to Nicaragua; ref.153  M S L

Tromba xanthura (Godman, 1901)  Yellow-washed Ruby-eye  Costa Rica to Colombia  iN  M T L

Pseudosarbia  Berg, 1897  

Pseudosarbia flavofasciata Skinner, 1921  TL: Brazil (MT)  M S

Pseudosarbia phoenicicola Berg, 1897    iN  M S

Pyrrhopygopsis  Godman, 1901  

Pyrrhopygopsis romula (H. Druce, 1875)    iN

Pyrrhopygopsis romula romula (H. Druce, 1875)  TL: Colombia  M T S

Pyrrhopygopsis romula tenebricosa (Hewitson, 1876)  TL: Peru  M T IL

Pyrrhopygopsis romula lugubris H. Druce, 1908  TL: Bolivia  M T S G

Pyrrhopygopsis quispica (Plötz, 1886)    iN  M T S IL

Pyrrhopygopsis socrates (Ménétriés, 1855)    iN

Pyrrhopygopsis socrates socrates (Ménétriés, 1855)  TL: Brazil (MG)  M T S

Pyrrhopygopsis socrates orasus (H. Druce, 1876)  TL: Peru  M T OI S L

Pyrrhopygopsis socrates crates Mabille & Boullet, 1912  TL: Bolivia; [Ecuador]; Peru  M T S IL

Pyrrhopygopsis agaricon H. Druce, 1908  Agaricon Skipper    iN

Pyrrhopygopsis agaricon agaricon H. Druce, 1908  E Costa Rica, Colombia  M T S IL

Pyrrhopygopsis agaricon nanta Evans, 1955  TL: Ecuador  M T

Nyctus  Mabille, 1891  

Nyctus crinitus Mabille, 1891  TL: Peru  M T

Nyctus hiarbas (Cramer, 1775)  TL: Surinam  M S

  Tribe Pericharini, Subtribe Pericharina

Oz  Grishin, 2022  

Oz sebastiani (Salazar & Constantino, 2013)  Sebastián's Ruby-eye  Choco region in Colombia; ref.58,13

Oz ozias (Hewitson, 1878)  Ozias Ruby-eye  ref.13  iN

Oz ozias ozina (Evans, 1955)  Ecuador, Peru, Bolivia  M S

Oz ozias ozias (Hewitson, 1878)  S Brazil, Argentina; ref.13  M T L

Orses  Godman, 1901  

Orses cynisca (Swainson, 1821)  Yellow-edged Ruby-eye  E & W Mexico to S Brazil & Paraguay  iN  M T OI S IL L

Orses itea (Swainson, 1821)    iN  M S

Perichares  Scudder, 1872  

Perichares saptine (Godman & Salvin, 1879)  Golden-banded Ruby-eye  S Mexico (Oax Chis) to Costa Rica; ref.13  iN  M T S L

Perichares seneca (Latreille, [1824])    iN

Perichares seneca crotona (Hewitson, 1866)  Panama, Venezuela  M T S

Perichares seneca seneca (Latreille, [1824])  TL: Brazil  M T S

Perichares chima Evans, 1955    iN  M T S

Perichares deceptus (A. Butler & H. Druce, 1872)  Flasher Ruby-eye    iN

Perichares deceptus deceptus (A. Butler & H. Druce, 1872)  El Salvador to Costa Rica  M T S L G

Perichares deceptus drina Evans, 1955  TL: Colombia  M T

Perichares deceptus quadrimaculata Röber, 1925  TL: Colombia

Perichares deceptus fulvimargo (A. Butler, 1873)  TL: Venezuela  M T S

Perichares deceptus luscinia (Plötz, 1882)  TL: Brazil (SC)  M T S L

Perichares colenda (Hewitson, 1866)    iN  M T S

Perichares forbesi Biezanko & O. Mielke, 1973  TL: Brazil (SP)  M T S

Perichares furcata (Mabille, 1891)  S Brazil; ref.5  M T S

Perichares philetes (Gmelin, [1790])  Caribbean Ruby-eye  Greater Antilles  iN  M S Txt

Perichares poaceaphaga Burns, 2008  Grass-feeding Ruby-eye  Costa Rica  M T OD G

Perichares adela (Hewitson, 1867)  Green-backed Ruby-eye  S TX (uncommon), E & W Mexico to S Brazil  iN  M T S L I G Txt

Perichares marmorata Scudder, 1872  TL: Venezuela  M S Txt

Perichares limana Evans, 1955  TL: Peru  M Txt

Perichares aurina Evans, 1955  Evans' Ruby-eye  [Honduras?], S Brazil & Paraguay  iN  M T S Txt

Perichares prestoeaphaga Burns, 2008  Prestoea-feeding Ruby-eye  Costa Rica  M T OD G

Perichares geonomaphaga Burns, 2008  Geonoma-feeding Ruby-eye  Costa Rica  M T OD G

Perichares lotus (A. Butler, 1870)  Lotus Ruby-eye  SE Mexico S Brazil & Paraguay  iN  M T S G

Perichares fura Grishin, 2023  [convergence with furcata] Ecuador; ref.153

Perichares matha Evans, 1955  TL: Ecuador  M T S

Perichares butus (Möschler, 1877)    iN  M T S

Perichares metallica (N. Riley, 1921)  S Brazil; ref.5  M T

Perichares manu (O. Mielke & Casagrande, 2004)  Peru, Brazil; ref.5  iN  M T OI S

Perichares romeroi (O. Mielke & Casagrande, 2004)  Venezuela; ref.5  M T OI

Perichares haworthiana (Swainson, 1821)  Green-banded Ruby-eye  Panama to S Brazil; ref.5  iN  M T OI

Lycas  Godman, 1901  

Lycas godart (Latreille, [1824])    iN

Lycas godart godart (Latreille, [1824])  TL: Brazil  M T

Lycas godart boisduvalii (Ehrmann, 1909)  TL: Venezuela  M T S

Lycas argentea (Hewitson, 1866)  Silvered Ruby-eye  E & W Mexico to Colombia; ref.16  iN  M T S

Lycas devanes (Herrich-Schäffer, 1869)  S.Am. [to Argentina]; ref.16  M T L

Oenides  Mabille, 1904  

Oenides vulpina (C. Felder & R. Felder, 1867)  Colombia; ref.5  iN  M T S L

  Tribe Pericharini, Subtribe Orphina

Orphe  Godman, 1901  

Orphe gerasa (Hewitson, 1867)  Gerasa Skipper  Nicaragua to Amazon basin & Guianas  iN  M T OI S L

Orphe vatinius Godman, 1901  Vatinius Skipper  Costa Rica, Colombia, Venezuela, Guyana, French Guiana, Brazil, Peru, Ec ...  iN  M T S L

Pseudorphe  A. Warren & Dolibaina, 2015  

Pseudorphe pyrex (Evans, 1955)  Colombia, Peru, Brazil; ref.24  iN  M T S

  Tribe Megathymini, Subtribe Carystoidina

Carystoides  Godman, 1901  

Carystoides mexicana H. Freeman, 1969  Mexican Ruby-eye  E Mexico  iN  M T

Carystoides floresi H. Freeman, 1969  Freeman’s Ruby-eye  E Mexico  iN  M T S

Carystoides escalantei H. Freeman, 1969  Escalante's Ruby-eye  E Mexico to Costa Rica  iN  M T S

Carystoides abrahami H. Freeman, 1969  Abraham’s Ruby-eye  E & W Mexico  iN  M T

Carystoides certima (Hewitson, 1866)    iN

Carystoides lila Evans, 1955  Lila Ruby-eye  S Mexico to S Ecuador  iN  M S

Carystoides lebbaeus (Hewitson, 1876)  Lebbaeus Ruby-eye    iN

Carystoides lebbaeus lebbaeus (Hewitson, 1876)  Costa Rica to NW Ecuador  M T S

Carystoides lebbaeus benchos (Weeks, 1909)  TL: Bolivia

Carystoides lebbaeus cundina Evans, 1955  TL: Colombia

Carystoides manta Evans, 1955  TL: Colombia

Carystoides hondura Evans, 1955  Honduran Ruby-eye  Honduras to Costa Rica  iN

Carystoides hondura hondura Evans, 1955  Honduras to Costa Rica  M S

Carystoides hondura pacifica Vargas & Salazar, [2017]  Colombia; ref.91

Carystoides alda (Plötz, 1882)  TL: Brazil

Carystoides basoches (Latreille, [1824])  Basoches Ruby-eye  S Mexico to S Brazil & Paraguay  iN  M T S

Carystoides yenna Evans, 1955    iN  M S

Carystoides noseda (Hewitson, 1866)    iN  M T

Carystoides sicania (Hewitson, 1876)    iN

Carystoides sicania orbius (Godman, 1901)  S Mexico to Ecuador & Guianas  M S L

Carystoides sicania sicania (Hewitson, 1876)  TL: Brazil ([RJ])  M T

Carystoides cathaea (Hewitson, 1866)    iN  M T L G

Carystoides metaensis Vargas & Salazar, [2017]  Colombia; ref.91

Carystoides sylvestris Vargas & Salazar, [2017]  Colombia; ref.91

Carystoides balza Evans, 1955  TL: Ecuador

Carystoides maroma (Möschler, 1877)  Maroma Ruby-eye  Panama to Amazon basin  iN  M T S

Carystoides goliath Grishin, 2023  Colombia; ref.153

  Tribe Megathymini, Subtribe Megathymina

Aegiale  C. Felder & R. Felder, 1860  

Aegiale hesperiaris (Walker, 1856)  Tequila Giant-Skipper; Gusano de Maguey  much of N & C Mexico  iN  M T S1 S2 S3 S4 S5 IL L

Turnerina  H. Freeman, 1959  

Turnerina mejicanus (E. Bell, 1938)  Turner's Giant-Skipper  Durango  iN  M T G

Turnerina hazelae (D. Stallings & J. Turner, 1958)  Guerrero Giant-Skipper  Guerrero & Oaxaca  M T S

Turnerina begoniae J. García & A. Turrent, 2019  Pueblo; ref.59  M T

Agathymus  H. Freeman, 1959  

Agathymus indecisa (A. Butler & H. Druce, 1872)  Guatemalan Giant-Skipper  Chiapas to Costa Rica  iN  M T S

Agathymus escalantei D. Stallings, J. Turner & V. Stallings, 1966  Escalante's Giant-Skipper  Guerrero  iN  M T Ha

Agathymus rethon (Dyar, 1913)  Black Giant-Skipper  Mexico (COL, GRO, JAL, MICH, MOR, NAY, OAX, PUE, VER), El Salvador  iN  M T S L

Agathymus tehuacanus J. García & A. Turrent, 2020  Pueblo; ref.60

Agathymus gentryi Roever, 1998  Gentry's Giant-Skipper  SW AZ (W Pima, C Maricopa, W Pinal, W Yuma), E SB Co., CA  M S

Agathymus baueri (D. Stallings & J. Turner, 1954)  Bauer's Giant-Skipper    iN

Agathymus baueri freemani D. Stallings & J. Turner, 1960  W-C AZ: SW Yavapai Co., S Mohave Co.  M T S

Agathymus baueri baueri (D. Stallings & J. Turner, 1954)  C AZ: E Yavapai, W Gila, NE Maricopa, N Pinal Cos.  M T S L H

Agathymus aryxna (Dyar, 1905)  Arizona Giant-Skipper  SE AZ, SW NM, NE SON  iN  M T S HS L

Agathymus evansi (H. Freeman, 1950)  Huachuca Giant-Skipper  Huachuca Mts., Cochise Co., AZ  iN  M T S HS L1 L2 H

Agathymus belli (H. Freeman, 1955)  Bell's Giant-Skipper  Chihuahua, Durango  M T S

Agathymus ricei D. Stallings, J. Turner & V. Stallings, 1966  Rice's Giant-Skipper  Puebla  M T S1 S2

Agathymus polingi (Skinner, 1905)  Poling's Giant-Skipper  S AZ, SW NM (Hidalgo Co.), NE Sonora  iN  M T S L1 L2 I H Ha

Agathymus neumoegeni (W. H. Edwards, 1882)  Neumoegen’s Giant-Skipper    iN

Agathymus neumoegeni neumoegeni (W. H. Edwards, 1882)  C AZ (Yavapai Co.) to W-C NM (Catron Co.)  M T S L

Agathymus neumoegeni S NM segregate ,  S NM: Organ Mts. (Dona Ana-Otero Cos.) to Sacramento Mts.  M S F Ha

Agathymus neumoegeni carlsbadensis (D. Stallings & J. Turner, 1957)  Guadalupe Mts. (Eddy-Otero Cos., NM; Culberson Co., TX)  M T S

Agathymus neumoegeni judithae (D. Stallings & J. Turner, 1957)  Hueco Mts., Hudspeth Co., TX  M T S

Agathymus neumoegeni diabloensis H. Freeman, 1962  Sierra Diablo, Sierra Blanca, Hudspeth Co., TX  M T S

Agathymus neumoegeni florenceae (D. Stallings & J. Turner, 1957)  Davis Mts., Jeff Davis Co., TX  M T S F Ha

Agathymus neumoegeni mcalpinei (H. Freeman, 1955)  E side of Glass Mts., Brewster Co., TX  M T S F Ha

Agathymus chisosensis (H. Freeman, 1952)  Neumoegen’s Giant-Skipper  Chisos Mts., S Brewster Co., TX  M T S

Agathymus juliae (D. Stallings & J. Turner, 1958)  Julia’s Giant-Skipper  Durango  M T S

Agathymus hoffmanni (H. Freeman, 1952)  Hoffmann’s Giant-Skipper  S-C Mexico (DF, GTO, HGO, MEX, SLP)  iN  M T S

Agathymus alliae (D. Stallings & J. Turner, 1957)  Mojave Giant-Skipper    iN

Agathymus alliae alliae (D. Stallings & J. Turner, 1957)  N Coconino Co., AZ  M T S

Agathymus alliae paiute Roever, 1998  SE CA (SB, Inyo), S NV (Clark), SW UT (Wa), NW AZ  M S L Ha

Agathymus fieldi H. Freeman, 1960  Field's Giant-Skipper  W Mexico: JAL, SON  iN  M T S

Agathymus stephensi (Skinner, 1912)  California Giant-Skipper  S CA (Riverside, E San Diego, W Imperial Cos.), Baja (N)  iN  M T S L1 L2 I H1 H2

Agathymus comstocki (Harbison, 1957)  Comstock's Giant-Skipper  Baja California (N)  iN  M T1 T2 L1 L2 H Ha

Agathymus [n. sp.] Roever, MS  Undescribed Giant-Skipper  Baja California (N)  M L H

Agathymus dawsoni Harbison, 1963  Dawson’s Giant-Skipper  Baja California (N)  M T1 T2 S L1 L2 H Ha

Agathymus mariae (W. Barnes & Benjamin, 1924)  Mary's Giant-Skipper    iN

Agathymus mariae mariae (W. Barnes & Benjamin, 1924)  S NM, trans-Pecos counties, TX  M T S L

Agathymus mariae chinatiensis H. Freeman, 1964  Chinati Mts., Presidio Co., TX  M T S

Agathymus mariae lajitaensis H. Freeman, 1964  S Presidio Co., TX  M T S

Agathymus mariae rindgei H. Freeman, 1964  Val Verde and Kinney Cos., TX  M T S Ha

Agathymus gilberti H. Freeman, 1964  Gilbert's Giant-Skipper  Brewster, Terrell and Val Verde Cos., TX  M T

Agathymus micheneri D. Stallings, J. Turner & V. Stallings, 1961  Michener’s Giant-Skipper  Coahuila  M T S L

Agathymus galeana Grishin, 2023  Mexico (NL); ref.153

Agathymus remingtoni (D. Stallings & J. Turner, 1958)  Remington's Giant-Skipper  E Mexico: HGO, SLP, TAMPS  iN  M T S

Agathymus estelleae (D. Stallings & J. Turner, 1958)  Estelle's Giant-Skipper    iN

Agathymus estelleae valverdiensis H. Freeman, 1966  S-C TX (Terrell, Val Verde, Edwards, Kinney Cos.)  M T S

Agathymus estelleae estelleae (D. Stallings & J. Turner, 1958)  Coahuila, Nuevo Leon  M T S

Stallingsia  H. Freeman, 1959  

Stallingsia maculosus (H. Freeman, 1955)  Manfreda Giant-Skipper  S TX, N NL  iN  M T1 T2 S1 S2 S3 S4 H1 H2 H3

Stallingsia smithi (H. Druce, 1896)  Smith's Giant-Skipper  SW MX (JAL, MICH, GRO, DF, MOR)  M T OI S L

Stallingsia jacki D. Stallings, J. Turner & V. Stallings, 1963  Chiapan Giant-Skipper  Chiapas  M T S

Megathymus  Scudder, 1872  

Megathymus streckeri (Skinner, 1895)  Strecker's Giant-Skipper    iN

Megathymus streckeri streckeri (Skinner, 1895)  NE AZ, NW NM, SW CO, SE UT  M T S L I Ha

Megathymus streckeri S Navajo Co., Arizona segregate ,  S end White Mts., AZ  M S L Ha

Megathymus streckeri leussleri W. Holland, 1931  TX panhandle & E NM, to E MT, W ND, (SE SASK?)  M T S1 S2 L F Ha

Megathymus streckeri texana W. Barnes & McDunnough, 1912  S-C Texas  M T1 T2 L I1 I2 H

Megathymus cofaqui (Strecker, 1876)  Cofaqui Giant-Skipper    iN

Megathymus cofaqui cofaqui (Strecker, 1876)  [= harrisi], Georgia to NC  M T TI S L I1 I2 Ha

Megathymus cofaqui slotteni Gatrelle, 1999  Florida  M T S L I H

Megathymus yuccae (Boisduval & Le Conte, [1837])  Yucca Giant-Skipper    iN

Megathymus yuccae yuccae (Boisduval & Le Conte, [1837])  [= buchholzi], SE USA, E of Mississippi River  M T1 T2 OI S L1 L2 I Ha

Megathymus yuccae reinthali H. Freeman, 1963  [= kendalli], C to NE TX, C & E OK, SW AR, N & C LA  M T1 T2 S L I1 I2 I3 F Ha

Megathymus yuccae wilsonorum D. Stallings & J. Turner, 1958  Tamaulipas & far S TX (lower Rio Grande Valley)  M T S L I

Megathymus yuccae louiseae H. Freeman, 1963  S-C TX (Val Verde, Kinney Cos) to NE Mexico (COAH, NL)  M T S

Megathymus yuccae coloradensis C. Riley, 1877  SE WY, W NE, W KS, E CO, E NM, W OK, TX panhandle  M T1 T2 T3 T4 S1 S2 L I H

Megathymus yuccae reubeni D. Stallings, J. Turner & V. Stallings, 1963  Hueco Mts., Hudspeth Co., TX, to N Chihuahua  M T S

Megathymus yuccae navajo Skinner, 1911  [= arizonae], W CO, S UT, W NM, AZ, N Sonora  M T1 T2 S Ma L1 L2 I1 I2 Ha

Megathymus yuccae martini D. Stallings & J. Turner, 1956  Mohave Desert, S CA, W NV, far NW AZ  M T S L1 L2 Ha

Megathymus yuccae harbisoni J. Emmel & T. Emmel, 1998  Beaumont, Riverside Co. CA, S to Baja California Norte  M T S Ha

Megathymus beulahae D. Stallings & J. Turner, 1958  Broad-banded Giant-Skipper    iN

Megathymus beulahae gayleae D. Stallings, J. Turner & V. Stallings, 1963  COAH, NL, TAMPS, SLP  M T1 T2 S L

Megathymus beulahae beulahae D. Stallings & J. Turner, 1958  Hidalgo  M T1 T2 S

Megathymus ursus Poling, 1902  Ursine Giant-Skipper    iN

Megathymus ursus ursus Poling, 1902  S AZ (Pinal, Pima, Sta. Cruz, Cochise), SW NM (Hidalgo), SON  M T OI S L I Ha

Megathymus ursus deserti R. Wielgus, J. Wielgus & D. Wielgus, 1972  E Maricopa Co., AZ  M T

Megathymus violae D. Stallings & J. Turner, 1956  Ursine Giant-Skipper  C & S NM (E of Rio Grande), W TX, COAH, NL; ref.11  iN  M T S L I Ha

Family HEDYLIDAE Guenée, 1858 – AMERICAN MOTH-BUTTERFLIES

  Subfamily Hedylinae

Macrosoma  Hübner, [1818]  [compiled with help from D. Bettman & G. Lamas]

Macrosoma undescribed-one Lourido, MS  [Central American "tipulata"] Costa Rica to Colombia  M S IL

Macrosoma tipulata Hübner, 1818  Amazonian  iN  M S IL

Macrosoma heliconiaria (Guenée, 1858)  Central America to Amazon  iN  M S IL

Macrosoma albipannosa (Prout, 1916)  Ecuador & Peru  iN  M S IL

Macrosoma semiermis (Prout, 1932)  Mexico to Bolivia  iN  M S IL

Macrosoma pectinogyna Scoble, 1990  Ecuador  iN  M OI OD

Macrosoma leucoplethes (Prout, 1917)  Ecuador  iN  M S IL

Macrosoma subornata (W. Warren, 1904)  Colombia to Peru  iN  M IL

Macrosoma albimacula (W. Warren, 1900)  Ecuador  iN  M IL

Macrosoma rubedinaria (Walker, 1862)  Cuba, Jamaica, Central America to Brazil  iN  M S IL

Macrosoma minutipuncta (Prout, 1916)  Peru  iN  M IL

Macrosoma coscoja (Dognin, 1900)  Ecuador  iN  M S IL

Macrosoma amaculata Scoble, 1990  Central America  iN  M IL

Macrosoma leucophasiata (Thierry-Mieg, 1904)  Ecuador to Bolivia  iN  M S IL

Macrosoma ustrinaria (Herrich-Schäffer, 1854)  Panama to Amazon  iN  M S IL

Macrosoma undescribed-three (Lourido, MS)  Central America  M IL

Macrosoma undescribed-two (Lourido, MS)  Brazil  M IL

Macrosoma albida (Schaus, 1901)  SE Brazil  iN  M S IL

Macrosoma napiaria (Guenée, 1858)  SE Brazil  iN  M S IL

Macrosoma leptosiata (C. Felder & Rogenhofer, 1875)  Amazonian  iN  M S IL

Macrosoma lamellifera (Prout, 1916)  Amazonian  iN  M IL

Macrosoma stabilinota (Prout, 1932)  Amazonian  iN  M S IL

Macrosoma muscerdata (C. Felder & Rogenhofer, 1875)  Costa Rica to Amazon  iN  M S IL

Macrosoma costilunata (Prout, 1916)  Costa Rica to Ecuador  iN  M IL

Macrosoma albistria (Prout, 1916)  S Peru  iN  M IL

Macrosoma satellitiata (Guenée, 1858)  Central America to SE Brazil  iN  M S IL

Macrosoma uniformis (W. Warren, 1904)  Ecuador, Peru & Brazil  iN  M IL

Macrosoma cascaria (Schaus, 1901)  Mexico to Venezuela  iN  M S IL

Macrosoma albifascia (W. Warren, 1904)  Equador, Peru, Bolivia & Brazil (Upper Amazon)  iN  M IL

Macrosoma intermedia (Dognin, 1911)  Costa Rica to Peru  iN  M IL

Macrosoma biapicata (Prout, 1917)  Peru  M OI OD

Macrosoma bahiata (C. Felder & Rogenhofer, 1875)  Central America? to Brazil  iN  M S IL

Macrosoma majormacula Lévêque, 2007  Costa Rica to Amazon  iN  M S IL

Macrosoma lucivittata (Walker, 1863)  Central America to Brazil  iN  M S IL

Macrosoma paularia (Schaus, 1901)  Bolivia, C & SE Brazil  iN  M S IL

Macrosoma klagesi (Prout, 1916)  Amazonian  iN  M S IL

Macrosoma nigrimacula (W. Warren, 1897)  Amazonian  iN  M S IL

Macrosoma interrupta (W. Warren, 1904)  Central America to Ecuador  M S IL

Macrosoma cellulata (Dognin, 1911)  Ecuador & Peru  iN  M IL

Macrosoma undescribed-five (Lourido, MS)  Costa Rica  M IL

Macrosoma conifera (W. Warren, 1897)  Central America to Amazon  iN  M S IL

Macrosoma undescribed-four (Lourido, MS)  Brazil  M IL

Macrosoma zikani (Prout, 1932)  SE Brazil  iN  M IL L

Macrosoma undescribed-six (Lourido, MS)  Brazil  M IL

Macrosoma hedylaria (W. Warren, 1894)  SE & S Brazil  iN  M S IL

Macrosoma hyacinthina (W. Warren, 1905)  Central America to Ecuador  iN  M S IL

Macrosoma undescribed-seven (Lourido, MS)  Brazil  M IL

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